1108 lines
135 KiB
XML
1108 lines
135 KiB
XML
<document id="5334ECC4B611F45007D207F384C38E8C" ID-CLB-Dataset="32217" ID-DOI="10.11646/zootaxa.4317.2.6" ID-GBIF-Dataset="ea3fccc3-ba05-4c8f-958a-61d8c20b1710" ID-ISSN="1175-5326" ID-Zenodo-Dep="884319" ID-ZooBank="E82Ab46E-Cb3A-4513-808F-6C9Cf17Be372" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1504527074084" checkinUser="plazi" docAuthor="Dehling, D. Matthias" docDate="2017" docId="DF4E87F0FFECDE66C3E172FCFCB0FD40" docLanguage="en" docName="zootaxa.4317.2.6.pdf" docOrigin="Zootaxa 4317 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Limnonectes kong Dehling 2017, sp. nov." docType="treatment" docVersion="7" lastPageNumber="307" masterDocId="2377FF88FFE5DE76C376726BFFF7FFE8" masterDocTitle="A new wide-headed Fanged Frog of the Limnonectes kuhlii group (Anura: Dicroglossidae) from western Borneo with a redescription of Rana conspicillata Günther, 1872" masterLastPageNumber="309" masterPageNumber="291" pageNumber="300" updateTime="1698467822354" updateUser="ExternalLinkService">
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<mods:title id="03B54AD3BFF2FB45A55690A2060B42D2">A new wide-headed Fanged Frog of the Limnonectes kuhlii group (Anura: Dicroglossidae) from western Borneo with a redescription of Rana conspicillata Günther, 1872</mods:title>
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<mods:namePart id="3DD054292D08D61E6C8E5DFE438C975D">Dehling, D. Matthias</mods:namePart>
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<mods:title id="CF952F13CAFBAD40765A53DA88FCDB76">Zootaxa</mods:title>
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<mods:part id="0801942179932CF400DD282CFC0EC0C5">
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<mods:date id="01000664F5D343D58A0CC56AA93ECA20">2017</mods:date>
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<mods:number id="C46446E3C21B3DE423508C66CAB133B9">2017-04-04</mods:number>
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<mods:number id="337C29E94B67AEA63EDC8CC7C546877A">4317</mods:number>
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<mods:identifier id="2F003A55D5543DA4BA018CA2F9904286" type="DOI">10.11646/zootaxa.4317.2.6</mods:identifier>
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<treatment id="DF4E87F0FFECDE66C3E172FCFCB0FD40" ID-DOI="http://doi.org/10.5281/zenodo.6507987" ID-GBIF-Taxon="133169967" ID-Zenodo-Dep="6507987" LSID="urn:lsid:plazi:treatment:DF4E87F0FFECDE66C3E172FCFCB0FD40" httpUri="http://treatment.plazi.org/id/DF4E87F0FFECDE66C3E172FCFCB0FD40" lastPageId="16" lastPageNumber="307" pageId="9" pageNumber="300">
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<subSubSection id="1FFD656DFFECDE7FC3E172FCFE2DFF5A" box="[151,474,151,178]" pageId="9" pageNumber="300" type="nomenclature">
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<paragraph id="575836E6FFECDE7FC3E172FCFE2DFF5A" blockId="9.[151,474,151,178]" box="[151,474,151,178]" pageId="9" pageNumber="300">
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<heading id="0C10818AFFECDE7FC3E172FCFE2DFF5A" bold="true" box="[151,474,151,178]" fontSize="11" level="1" pageId="9" pageNumber="300" reason="1">
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||
<emphasis id="6593EAF4FFECDE7FC3E172FCFE2DFF5A" bold="true" box="[151,474,151,178]" pageId="9" pageNumber="300">
|
||
<taxonomicName id="90E74D65FFECDE7FC3E172FCFE81FF59" ID-CoL="6QDL2" authority="Dehling, 2017" authorityName="Dehling" authorityYear="2017" box="[151,374,151,177]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" kingdom="Animalia" order="Anura" pageId="9" pageNumber="300" phylum="Chordata" rank="species" species="kong" status="sp. nov.">
|
||
<emphasis id="6593EAF4FFECDE7FC3E172FCFE81FF59" bold="true" box="[151,374,151,177]" italics="true" pageId="9" pageNumber="300">Limnonectes kong</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="7EA0578FFFECDE7FC20A72F3FE2DFF5A" box="[380,474,152,178]" pageId="9" pageNumber="300" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="1FFD656DFFECDE7FC3E172B5FD3FFED8" pageId="9" pageNumber="300" type="reference_group">
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||
<paragraph id="575836E6FFECDE7FC3E172B5FDBAFF1B" blockId="9.[151,712,221,304]" box="[151,589,221,243]" pageId="9" pageNumber="300">
|
||
<taxonomicName id="90E74D65FFECDE7FC3E172B5FE03FF1B" authority=": Gunther, 1872" authorityName=": Gunther" authorityYear="1872" box="[151,500,221,243]" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="9" pageNumber="300" phylum="Chordata" rank="species" species="conspicillata">
|
||
<emphasis id="6593EAF4FFECDE7FC3E172B5FEA1FF1B" box="[151,342,221,243]" italics="true" pageId="9" pageNumber="300">Rana conspicillata</emphasis>
|
||
:
|
||
<bibRefCitation id="33764B17FFECDE7FC21572B6FE03FF1B" author="Gunther" box="[355,500,221,243]" pageId="9" pageNumber="300" refString="Gunther, A. (1872) On the reptiles and amphibians oF Borneo. P roceedings of the Zoological Society of London, 1872, 586 - 600, pls. xxxv - xl." type="journal article" year="1872">Günther, 1872</bibRefCitation>
|
||
</taxonomicName>
|
||
(partim)
|
||
</paragraph>
|
||
<paragraph id="575836E6FFECDE7FC3E17296FDF3FEFA" blockId="9.[151,712,221,304]" box="[151,516,252,274]" pageId="9" pageNumber="300">
|
||
<taxonomicName id="90E74D65FFECDE7FC3E17296FE5CFEFA" authority=": Gunther, 1874" authorityName=": Gunther" authorityYear="1874" box="[151,427,252,274]" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="9" pageNumber="300" phylum="Chordata" rank="species" species="kuhlii">
|
||
<emphasis id="6593EAF4FFECDE7FC3E17296FEFAFEFA" box="[151,269,252,274]" italics="true" pageId="9" pageNumber="300">Rana kuhlii</emphasis>
|
||
:
|
||
<bibRefCitation id="33764B17FFECDE7FC26C7297FE5CFEFA" author="Gunther" box="[282,427,252,274]" pageId="9" pageNumber="300" refString="Gunther, A. C. L. G. (1874) Reptilia (1872). Zoological Record, 9, 61 - 81." type="journal article" year="1874">Günther, 1874</bibRefCitation>
|
||
</taxonomicName>
|
||
(partim)
|
||
</paragraph>
|
||
<paragraph id="575836E6FFECDE7FC3E17370FD3FFED8" blockId="9.[151,712,221,304]" box="[151,712,282,304]" pageId="9" pageNumber="300">
|
||
<taxonomicName id="90E74D65FFECDE7FC3E17370FD98FED8" ID-CoL="6QDKX" authority=": Matsui et al. 2013" authorityName="Matsui et al." authorityYear="2013" box="[151,623,282,304]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" kingdom="Animalia" order="Anura" pageId="9" pageNumber="300" phylum="Chordata" rank="species" species="conspicillatus">
|
||
<emphasis id="6593EAF4FFECDE7FC3E17370FE5EFED8" box="[151,425,282,304]" italics="true" pageId="9" pageNumber="300">Limnonectes conspicillatus</emphasis>
|
||
:
|
||
<bibRefCitation id="33764B17FFECDE7FC2C17371FD98FED8" author="Matsui" box="[439,623,282,304]" pageId="9" pageNumber="300" refString="Matsui, M., Dubois, A. & Ohler, A. (2013) New replacement name For Rana paradoxa Mocquard, 1890 with designations oF lectotypes For Rana paradoxa and Rana conspicillata Gunther, 1872: both synonymized with Limnonectes kuhlii (Tschudi, 1838) (Dicroglossidae: Dicroglossinae). Asian Herpetological Research, 4 (3), 187 - 189." type="journal article" year="2013">
|
||
Matsui
|
||
<emphasis id="6593EAF4FFECDE7FC1727370FDC7FED8" box="[516,560,282,304]" italics="true" pageId="9" pageNumber="300">et al</emphasis>
|
||
. 2013
|
||
</bibRefCitation>
|
||
</taxonomicName>
|
||
(partim)
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="1FFD656DFFECDE7FC3E17336FEDAFC86" pageId="9" pageNumber="300" type="materials_examined">
|
||
<paragraph id="575836E6FFECDE7FC3E17336FE95FE56" blockId="9.[151,1437,349,2030]" pageId="9" pageNumber="300">
|
||
<emphasis id="6593EAF4FFECDE7FC3E17336FEF3FE9E" bold="true" box="[151,260,349,374]" pageId="9" pageNumber="300">
|
||
<typeStatus id="885C8844FFECDE7FC3E17336FEF3FE9E" box="[151,260,349,374]" pageId="9" pageNumber="300" type="holotype">Holotype</typeStatus>
|
||
</emphasis>
|
||
.
|
||
<collectionCode id="31F6AE23FFECDE7FC2607335FE90FE9D" box="[278,359,350,373]" country="Switzerland" httpUri="http://grbio.org/cool/5hwi-0wgz" name="Naturhistorisches Museum der Burgergemeinde Bern" pageId="9" pageNumber="300">NMBE</collectionCode>
|
||
1059923, adult male, from Kubah National Park (
|
||
<geoCoordinate id="32D35021FFECDE7FC0C87336FBB4FE9E" box="[958,1091,349,374]" direction="north" orientation="latitude" pageId="9" pageNumber="300" precision="1" value="1.6068499">1°36.411’N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="32D35021FFECDE7FC7257335FB05FE9E" box="[1107,1266,350,374]" direction="east" orientation="longitude" pageId="9" pageNumber="300" precision="1" value="110.18998">110°11.399’E</geoCoordinate>
|
||
;
|
||
<quantity id="901F9B03FFECDE7FC6777335FABAFE9D" box="[1281,1357,350,374]" metricMagnitude="2" metricUnit="m" metricValue="2.9" pageId="9" pageNumber="300" unit="m" value="290.0">290 m</quantity>
|
||
a.s.l.), Matang Range,
|
||
<collectingCountry id="2FF07676FFECDE7FC22573EAFE40FE72" box="[339,439,385,410]" name="Malaysia" pageId="9" pageNumber="300">
|
||
<collectingRegion id="9523F804FFECDE7FC22573EAFE40FE72" box="[339,439,385,410]" country="Malaysia" name="Sarawak" pageId="9" pageNumber="300">Sarawak</collectingRegion>
|
||
</collectingCountry>
|
||
,
|
||
<collectingCountry id="2FF07676FFECDE7FC2B073EAFDDAFE72" box="[454,557,385,410]" name="Malaysia" pageId="9" pageNumber="300">Malaysia</collectingCountry>
|
||
(Borneo); ColleCted
|
||
<date id="23591026FFECDE7FC05473EAFBF1FE71" box="[802,1030,385,410]" pageId="9" pageNumber="300" value="2009-09-26">26 September 2009</date>
|
||
by J. Maximilian Dehling and D. Matthias Dehling.
|
||
</paragraph>
|
||
<paragraph id="575836E6FFECDE7FC3B173A2FECFFD7E" blockId="9.[151,1437,349,2030]" pageId="9" pageNumber="300">
|
||
<emphasis id="6593EAF4FFECDE7FC3B173A2FEC8FE0A" bold="true" box="[199,319,457,482]" pageId="9" pageNumber="300">
|
||
<typeStatus id="885C8844FFECDE7FC3B173A2FEC8FE0A" box="[199,319,457,482]" pageId="9" pageNumber="300" type="paratype">Paratypes</typeStatus>
|
||
</emphasis>
|
||
.
|
||
<collectionCode id="31F6AE23FFECDE7FC22373A1FE51FE09" box="[341,422,458,481]" country="Switzerland" httpUri="http://grbio.org/cool/5hwi-0wgz" name="Naturhistorisches Museum der Burgergemeinde Bern" pageId="9" pageNumber="300">NMBE</collectionCode>
|
||
1059917 (juvenile male), 1059918 (adult female), 1059919 (adult male), 1059920 (subadult male), all from Kubah National Park;
|
||
<collectionCode id="31F6AE23FFECDE7FC1B67385FCE6FDED" box="[704,785,494,517]" country="Switzerland" httpUri="http://grbio.org/cool/5hwi-0wgz" name="Naturhistorisches Museum der Burgergemeinde Bern" pageId="9" pageNumber="300">NMBE</collectionCode>
|
||
1059924, 1059926 (two adult males), 1059925 (subadult male), all from Bako National Park (
|
||
<geoCoordinate id="32D35021FFECDE7FC147707AFD4FFDC2" box="[561,696,529,554]" direction="north" orientation="latitude" pageId="9" pageNumber="300" precision="1" value="1.7114">1°42.684’N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="32D35021FFECDE7FC1B2707AFC94FDC2" box="[708,867,529,554]" direction="east" orientation="longitude" pageId="9" pageNumber="300" precision="1" value="110.44615">110°26.769’E</geoCoordinate>
|
||
),
|
||
<collectingCountry id="2FF07676FFECDE7FC001707AFC2DFDC2" box="[887,986,529,554]" name="Malaysia" pageId="9" pageNumber="300">
|
||
<collectingRegion id="9523F804FFECDE7FC001707AFC2DFDC2" box="[887,986,529,554]" country="Malaysia" name="Sarawak" pageId="9" pageNumber="300">Sarawak</collectingRegion>
|
||
</collectingCountry>
|
||
,
|
||
<collectingCountry id="2FF07676FFECDE7FC093707AFBB8FDC2" box="[997,1103,529,554]" name="Malaysia" pageId="9" pageNumber="300">Malaysia</collectingCountry>
|
||
; 1059927–1059929, 1059932 (four adult females), 1059930–1059931 (two adult males), all from Gunung Gading National Park (
|
||
<geoCoordinate id="32D35021FFECDE7FC666705EFA6FFDA6" box="[1296,1432,565,590]" direction="north" orientation="latitude" pageId="9" pageNumber="300" precision="1" value="1.6951001">1°41.706’N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="32D35021FFECDE7FC3E17032FEC0FD9A" box="[151,311,601,626]" direction="east" orientation="longitude" pageId="9" pageNumber="300" precision="1" value="109.84092">109°50.455’E</geoCoordinate>
|
||
),
|
||
<collectingCountry id="2FF07676FFECDE7FC23A7032FE58FD9A" box="[332,431,601,626]" name="Malaysia" pageId="9" pageNumber="300">
|
||
<collectingRegion id="9523F804FFECDE7FC23A7032FE58FD9A" box="[332,431,601,626]" country="Malaysia" name="Sarawak" pageId="9" pageNumber="300">Sarawak</collectingRegion>
|
||
</collectingCountry>
|
||
,
|
||
<collectingCountry id="2FF07676FFECDE7FC2CA7032FDD1FD9A" box="[444,550,601,626]" name="Malaysia" pageId="9" pageNumber="300">Malaysia</collectingCountry>
|
||
; ColleCted by J. M. Dehling and D. M. Dehling between
|
||
<date id="23591026FFECDE7FC7DD7031FEC3FD7E" pageId="9" pageNumber="300" value="2009-09-22" valueMax="2009-10-18" valueMin="2009-09-22">22 September and 18 OCtober 2009</date>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="575836E6FFECDE7FC3B170CAFBE6FCEA" blockId="9.[151,1437,349,2030]" pageId="9" pageNumber="300">
|
||
<emphasis id="6593EAF4FFECDE7FC3B170CAFE4CFD52" bold="true" box="[199,443,673,698]" pageId="9" pageNumber="300">Referred specimens.</emphasis>
|
||
<collectionCode id="31F6AE23FFECDE7FC2B270C9FDEFFD51" box="[452,536,674,697]" pageId="9" pageNumber="300">BMNH</collectionCode>
|
||
1947.2.29.24, adult female, from Matang Range,
|
||
<collectingCountry id="2FF07676FFECDE7FC72170CAFB4DFD52" box="[1111,1210,673,698]" name="Malaysia" pageId="9" pageNumber="300">
|
||
<collectingRegion id="9523F804FFECDE7FC72170CAFB4DFD52" box="[1111,1210,673,698]" country="Malaysia" name="Sarawak" pageId="9" pageNumber="300">Sarawak</collectingRegion>
|
||
</collectingCountry>
|
||
,
|
||
<collectingCountry id="2FF07676FFECDE7FC7B170CAFAD9FD52" box="[1223,1326,673,698]" name="Malaysia" pageId="9" pageNumber="300">Malaysia</collectingCountry>
|
||
(Borneo) (
|
||
<typeStatus id="885C8844FFECDE7FC3E870AEFEC0FD36" box="[158,311,709,734]" pageId="9" pageNumber="300" type="paralectotype">paraleCtotype</typeStatus>
|
||
of
|
||
<taxonomicName id="90E74D65FFECDE7FC22A70ADFDD8FD35" authorityName=": Gunther" authorityYear="1872" box="[348,559,709,733]" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="9" pageNumber="300" phylum="Chordata" rank="species" species="conspicillata">
|
||
<emphasis id="6593EAF4FFECDE7FC22A70ADFDD8FD35" box="[348,559,709,733]" italics="true" pageId="9" pageNumber="300">Rana conspicillata</emphasis>
|
||
</taxonomicName>
|
||
);
|
||
<collectionCode id="31F6AE23FFECDE7FC13070ADFD60FD35" box="[582,663,710,733]" country="Switzerland" httpUri="http://grbio.org/cool/5hwi-0wgz" name="Naturhistorisches Museum der Burgergemeinde Bern" pageId="9" pageNumber="300">NMBE</collectionCode>
|
||
1059921 (subadult female), 1059922 (adult female) from Santubong National Park;
|
||
<collectingCountry id="2FF07676FFECDE7FC2367082FE54FCEA" box="[320,419,745,770]" name="Malaysia" pageId="9" pageNumber="300">
|
||
<collectingRegion id="9523F804FFECDE7FC2367082FE54FCEA" box="[320,419,745,770]" country="Malaysia" name="Sarawak" pageId="9" pageNumber="300">Sarawak</collectingRegion>
|
||
</collectingCountry>
|
||
,
|
||
<collectingCountry id="2FF07676FFECDE7FC2D97082FDEEFCEA" box="[431,537,745,770]" name="Malaysia" pageId="9" pageNumber="300">Malaysia</collectingCountry>
|
||
, ColleCted by J. M. Dehling in
|
||
<date id="23591026FFECDE7FC0067082FBF9FCEA" box="[880,1038,745,770]" pageId="9" pageNumber="300" value="2009-10">OCtober 2009</date>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="575836E6FFECDE7FC3B17166FEDAFC86" blockId="9.[151,1437,349,2030]" pageId="9" pageNumber="300">
|
||
<emphasis id="6593EAF4FFECDE7FC3B17166FE78FCCE" bold="true" box="[199,399,781,806]" pageId="9" pageNumber="300">Molecular data.</emphasis>
|
||
Holotype—NMBE 1059923 (GenBank ACCession #
|
||
<accessionNumber id="48B4AB05FFECDE7FC7557166FB52FCCE" box="[1059,1189,781,806]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MF322505" pageId="9" pageNumber="300">MF322505</accessionNumber>
|
||
); Paratypes—NMBE 1059917 (
|
||
<accessionNumber id="48B4AB05FFECDE7FC26F715AFE6BFCA2" box="[281,412,817,842]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MF322509" pageId="9" pageNumber="300">MF322509</accessionNumber>
|
||
), 1059919 (
|
||
<accessionNumber id="48B4AB05FFECDE7FC134715AFD33FCA2" box="[578,708,817,842]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MF322506" pageId="9" pageNumber="300">MF322506</accessionNumber>
|
||
), 1059920 (
|
||
<accessionNumber id="48B4AB05FFECDE7FC01C715AFC1AFCA2" box="[874,1005,817,842]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MF322510" pageId="9" pageNumber="300">MF322510</accessionNumber>
|
||
), 1059926 (
|
||
<accessionNumber id="48B4AB05FFECDE7FC7E5715AFAE1FCA2" box="[1171,1302,817,842]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MF322507" pageId="9" pageNumber="300">MF322507</accessionNumber>
|
||
), 1059927 (
|
||
<accessionNumber id="48B4AB05FFECDE7FC3E9713EFED6FC86" box="[159,289,853,878]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MF322508" pageId="9" pageNumber="300">MF322508</accessionNumber>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="1FFD656DFFECDE7FC3B17112FEEAFAAA" pageId="9" pageNumber="300" type="diagnosis">
|
||
<paragraph id="575836E6FFECDE7FC3B17112FEEAFAAA" blockId="9.[151,1437,349,2030]" pageId="9" pageNumber="300">
|
||
<emphasis id="6593EAF4FFECDE7FC3B17112FEB4FC7A" bold="true" box="[199,323,889,914]" pageId="9" pageNumber="300">Diagnosis:</emphasis>
|
||
Based on the presenCe of odontoids in the lower jaw (larger in males than in females), the enlarged, wide head, moderately depressed, stout body, short legs, thiCk and wrinkled skin, and the hidden tympanum, the new speCies is plaCed in the
|
||
<taxonomicName id="90E74D65FFECDE7FC2A771A9FDD9FC31" authorityName=": Gunther" authorityYear="1874" box="[465,558,961,985]" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="9" pageNumber="300" phylum="Chordata" rank="species" species="kuhlii">
|
||
<emphasis id="6593EAF4FFECDE7FC2A771A9FDD9FC31" box="[465,558,961,985]" italics="true" pageId="9" pageNumber="300">L. kuhlii</emphasis>
|
||
</taxonomicName>
|
||
group of the genus
|
||
<taxonomicName id="90E74D65FFECDE7FC07C71A9FC60FC31" authorityName="Fitzinger" authorityYear="1843" box="[778,919,962,985]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" kingdom="Animalia" order="Anura" pageId="9" pageNumber="300" phylum="Chordata" rank="genus">
|
||
<emphasis id="6593EAF4FFECDE7FC07C71A9FC60FC31" box="[778,919,962,985]" italics="true" pageId="9" pageNumber="300">Limnonectes</emphasis>
|
||
</taxonomicName>
|
||
. It is distinguished from the other members of the group by the Combination of the following CharaCters: Large body size and marked sexual dimorphism with SVL
|
||
<quantity id="901F9B03FFECDE7FC3A37662FE8CFBC9" box="[213,379,1033,1058]" metricMagnitude="-2" metricUnit="m" metricValue="6.7" metricValueMax="7.63" metricValueMin="5.77" pageId="9" pageNumber="300" unit="mm" value="67.0" valueMax="76.3" valueMin="57.7">57.7–76.3 mm</quantity>
|
||
in adult males and
|
||
<quantity id="901F9B03FFECDE7FC1147662FCFFFBC9" box="[610,776,1033,1058]" metricMagnitude="-2" metricUnit="m" metricValue="5.595" metricValueMax="5.9399999999999995" metricValueMin="5.25" pageId="9" pageNumber="300" unit="mm" value="55.95" valueMax="59.4" valueMin="52.5">52.5–59.4 mm</quantity>
|
||
in adult females; head very large in males and markedly sexually dimorphiC, males (HW/SVL 0.41–0.48, HL/SVL 0.44–0.50) having wider and longer heads than females (HW/SVL 0.38–0.40; HL/SVL HL/SVL 0.39–0.43); head slightly longer than wide (HW/HL 0.93–1.00 in males,
|
||
<quantity id="901F9B03FFECDE7FC3E1761DFED0FB66" box="[151,295,1141,1166]" metricMagnitude="-2" metricUnit="m" metricValue="2.4003" metricValueMax="2.4638" metricValueMin="2.3368" pageId="9" pageNumber="300" unit="in" value="0.945" valueMax="0.97" valueMin="0.92">0.92–0.97 in</quantity>
|
||
females); odontoids in males enlarged with a rounded knob-like tip; vomerine teeth in two highly elevated, large, broad posteromedially direCted ridges, separated from eaCh other by about half length of individual ridge; dorsolateral and transverse postorbital dermal folds absent; narrow, non-moveable dermal fringes between fingers along preaxial and postaxial sides of phalanges of hand; dorsal faCe of thigh and shank sparsely sCattered with enlarged, oval tuberCles; toes not fully webbed, leaving two phalanges on Toe IV and 1.5 phalanges on Toes II and III free.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="1FFD656DFFECDE78C3B17726FE51FD07" lastPageId="14" lastPageNumber="305" pageId="9" pageNumber="300" type="description">
|
||
<paragraph id="575836E6FFECDE7FC3B17726FB12F8B6" blockId="9.[151,1437,349,2030]" pageId="9" pageNumber="300">
|
||
<emphasis id="6593EAF4FFECDE7FC3B17726FDEDFA8E" bold="true" box="[199,538,1357,1382]" pageId="9" pageNumber="300">
|
||
Description of the
|
||
<typeStatus id="885C8844FFECDE7FC2DB7726FDE1FA8E" box="[429,534,1357,1382]" pageId="9" pageNumber="300" type="holotype">holotype</typeStatus>
|
||
.
|
||
</emphasis>
|
||
Adult male (
|
||
<figureCitation id="CFDC2A63FFECDE7FC1C17726FD08FA8E" box="[695,767,1357,1382]" captionStart="FIGURE 5" captionStartId="10.[151,250,1253,1275]" captionTargetBox="[151,1436,625,1232]" captionTargetId="figure@10.[151,1436,625,1232]" captionTargetPageId="10" captionText="FIGURE 5. Dorsal view (leFt) and ventral view (right) oF the holotype oF Limnonectes kong sp. nov. (NMBE 1059923, Field no. JMD 368, adult male). Scale bar represents 10 mm." httpUri="https://zenodo.org/record/884329/files/figure.png" pageId="9" pageNumber="300">Fig. 5</figureCitation>
|
||
); body large (SVL
|
||
<quantity id="901F9B03FFECDE7FC0937726FBB8FA8D" box="[997,1103,1357,1382]" metricMagnitude="-2" metricUnit="m" metricValue="6.83" pageId="9" pageNumber="300" unit="mm" value="68.3">68.3 mm</quantity>
|
||
;
|
||
<tableCitation id="1A65035DFFECDE7FC7287726FB41FA8E" box="[1118,1206,1357,1382]" captionStart="TABLE 2" captionStartId="7.[125,197,516,535]" captionText="TABLE 2. Measurements of the type series of Limnonectes kong sp. nov., the female paralectotype of L. conspicillatus re-assigned to L. kong, and the adult type specimens Limnonectes conspicillatus excluding the re-assigned specimens. For abbreviations see Material & Methods." pageId="9" pageNumber="300">Table 2</tableCitation>
|
||
), very stout, widest posterior to head, strongly tapering to groin; head massively enlarged, very long and wide (HL/SUL 0.48, HW/ SUL 0.48), as long as wide (HW/HL 1.00); snout relatively short (SL/HL 0.34), aCuminate in dorsal view, rounded in profile, slightly projeCting beyond lower jaw, little wider than long (SL/EE 0.96); Canthus rostralis moderately expressed between eye and nostril, straight-lined; loreal region grooved; nostrils rounded, direCted dorsolaterally; situated Closer to tip of snout than to eye (EN/NS 1.20), separated from eaCh other by distanCe smaller than distanCe between eye and nostril (NN/EN 0.90); eyes direCted anterolaterally, protruding, small (ED/HL 0.24); eye diameter muCh shorter than snout length (ED/SL 0.71); interorbital distanCe muCh wider than upper eyelid (IO/EW 1.60) and wider than internarial distanCe (IO/NN 1.30); tympanum ConCealed under thiCk layer of skin, not disCernible; upper jaw with dentition; teeth on premaxilla muCh larger than those on maxilla, forming dense, transverse row; Choanae rounded; vomerine teeth in two highly elevated, large, broad posteromedially direCted ridges, separated from eaCh other by about half length of individual ridge (
|
||
<figureCitation id="CFDC2A63FFECDE7FC1BE74B2FCE6F91A" box="[712,785,1753,1778]" captionStart="FIGURE 6" captionStartId="10.[151,250,1757,1779]" captionTargetBox="[151,1436,1356,1736]" captionTargetId="figure@10.[151,1436,1356,1736]" captionTargetPageId="10" captionText="FIGURE 6. Frontal view oF the mandibula (leFt) and ventral view oF the vomer region (right) oF the holotype oF Limnonectes kong sp. nov. (NMBE 1059923, adult male). Not to scale." httpUri="https://zenodo.org/record/884331/files/figure.png" pageId="9" pageNumber="300">Fig. 6</figureCitation>
|
||
); mandibular symphysis thiCkened to large bony knob; odontoid proCess present Close to mandibular symphysis on both sides, direCted dorsally, large, long (OL/MH 2.1; OL/SVL 0.08) with blunt, rounded tip, fitting into deep rounded Cavities in upper jaw when mouth Closed (
|
||
<figureCitation id="CFDC2A63FFECDE7FC63C754AFA7AF8D2" box="[1354,1421,1825,1850]" captionStart="FIGURE 6" captionStartId="10.[151,250,1757,1779]" captionTargetBox="[151,1436,1356,1736]" captionTargetId="figure@10.[151,1436,1356,1736]" captionTargetPageId="10" captionText="FIGURE 6. Frontal view oF the mandibula (leFt) and ventral view oF the vomer region (right) oF the holotype oF Limnonectes kong sp. nov. (NMBE 1059923, adult male). Not to scale." httpUri="https://zenodo.org/record/884331/files/figure.png" pageId="9" pageNumber="300">Fig. 6</figureCitation>
|
||
); tongue Comparatively short and broad, Covered with small pustules, most densely in anterior third.
|
||
</paragraph>
|
||
<paragraph id="575836E6FFECDE7FC3B17502FC30F806" blockId="9.[151,1437,349,2030]" pageId="9" pageNumber="300">
|
||
Dorsal side of trunk, head, and limbs rugose; posterior half of trunk and dorsal faCe of thigh and shank sparsely sCattered with enlarged, oval tuberCles (
|
||
<figureCitation id="CFDC2A63FFECDE7FC11175E6FD4EF84E" box="[615,697,1933,1958]" captionStart="FIGURE 4" captionStartId="6.[151,250,1929,1951]" captionTargetBox="[168,1422,1192,1922]" captionTargetId="figure@6.[168,1422,1192,1922]" captionTargetPageId="6" captionText="FIGURE 4. Dorsal view oF the shanks oF the preserved lectotype oF Limnonectes conspicillatus (BMNH 1947.2. 29.20, adult male; leFt); the preserved holotype oF Limnonectes kong sp. nov. (NMBE 1059923, adult male; centre); and a preserved paratype oF L. kong sp. nov. (NMBE 1059924, adult male; right)." httpUri="https://zenodo.org/record/884327/files/figure.png" pageId="9" pageNumber="300">Figs. 4</figureCitation>
|
||
and
|
||
<figureCitation id="CFDC2A63FFECDE7FC18175E6FCF0F84E" box="[759,775,1933,1958]" captionStart="FIGURE 5" captionStartId="10.[151,250,1253,1275]" captionTargetBox="[151,1436,625,1232]" captionTargetId="figure@10.[151,1436,625,1232]" captionTargetPageId="10" captionText="FIGURE 5. Dorsal view (leFt) and ventral view (right) oF the holotype oF Limnonectes kong sp. nov. (NMBE 1059923, Field no. JMD 368, adult male). Scale bar represents 10 mm." httpUri="https://zenodo.org/record/884329/files/figure.png" pageId="9" pageNumber="300">5</figureCitation>
|
||
); supratympaniC fold ConspiCuous, Curved, running from posterior Corner of eye to anterior edge of fore limb insertion; ventral side with dense, weak, low wrinkles; transversal dermal fold on Chest between anterior edges of arm insertion.
|
||
</paragraph>
|
||
<paragraph id="575836E6FFEFDE7CC3B172FCFDE0FDD5" blockId="10.[151,1436,151,573]" pageId="10" pageNumber="301">
|
||
Forelimbs stout, short (ARM/SVL 0.42); hand relatively short (HND/SVL 0.23); tips of fingers rounded, swollen palmarly (
|
||
<figureCitation id="CFDC2A63FFEFDE7CC20572D7FE4AFF3C" box="[371,445,188,213]" captionStart="FIGURE 7" captionStartId="11.[151,250,1900,1922]" captionTargetBox="[151,1436,445,1878]" captionTargetId="figure@11.[151,1436,445,1878]" captionTargetPageId="11" captionText="FIGURE 7. Palmar view oF right hand (leFt) and plantar view oF right Foot (right) oF the holotype (NMBE 1059923, adult male) oF Limnonectes kong sp. nov. (NMBE 1059923, adult male). Scale bar represents 10 mm." httpUri="https://zenodo.org/record/884333/files/figure.png" pageId="10" pageNumber="301">Fig. 7</figureCitation>
|
||
); relative length of fingers: II <I <IV <III; subartiCular tuberCles rounded, well developed, numbering one on Fingers I and II, two on Fingers III and IV; tuberCles on Fingers I and II and proximal tuberCles on Fingers III and IV muCh larger than distal ones, highly elevated (
|
||
<figureCitation id="CFDC2A63FFEFDE7CC775736FFBBFFEF4" box="[1027,1096,260,285]" captionStart="FIGURE 7" captionStartId="11.[151,250,1900,1922]" captionTargetBox="[151,1436,445,1878]" captionTargetId="figure@11.[151,1436,445,1878]" captionTargetPageId="11" captionText="FIGURE 7. Palmar view oF right hand (leFt) and plantar view oF right Foot (right) oF the holotype (NMBE 1059923, adult male) oF Limnonectes kong sp. nov. (NMBE 1059923, adult male). Scale bar represents 10 mm." httpUri="https://zenodo.org/record/884333/files/figure.png" pageId="10" pageNumber="301">Fig. 7</figureCitation>
|
||
); fingers without webbing but with narrow dermal fringes between fingers along phalanges from subartiCular tuberCles to disCs, most ConspiCuous on Fingers II and III, not moveable (
|
||
<figureCitation id="CFDC2A63FFEFDE7CC1457327FD8FFE8C" box="[563,632,332,357]" captionStart="FIGURE 7" captionStartId="11.[151,250,1900,1922]" captionTargetBox="[151,1436,445,1878]" captionTargetId="figure@11.[151,1436,445,1878]" captionTargetPageId="11" captionText="FIGURE 7. Palmar view oF right hand (leFt) and plantar view oF right Foot (right) oF the holotype (NMBE 1059923, adult male) oF Limnonectes kong sp. nov. (NMBE 1059923, adult male). Scale bar represents 10 mm." httpUri="https://zenodo.org/record/884333/files/figure.png" pageId="10" pageNumber="301">Fig. 7</figureCitation>
|
||
); thenar tuberCle small, about two-fifth length and about one-sixth area of metaCarpal of Finger I in palmar view, oval, low; inner palmar tuberCle subequal in shape and size to thenar tuberCle, Covering proximal part of metaCarpals of Fingers II and III; outer palmar tuberCle somewhat narrower, but equal in length to thenar and inner palmar tuberCle, slightly more prominent, Covering proximal part of metaCarpal of Finger IV (
|
||
<figureCitation id="CFDC2A63FFEFDE7CC24473B7FE81FE1C" box="[306,374,476,501]" captionStart="FIGURE 7" captionStartId="11.[151,250,1900,1922]" captionTargetBox="[151,1436,445,1878]" captionTargetId="figure@11.[151,1436,445,1878]" captionTargetPageId="11" captionText="FIGURE 7. Palmar view oF right hand (leFt) and plantar view oF right Foot (right) oF the holotype (NMBE 1059923, adult male) oF Limnonectes kong sp. nov. (NMBE 1059923, adult male). Scale bar represents 10 mm." httpUri="https://zenodo.org/record/884333/files/figure.png" pageId="10" pageNumber="301">Fig. 7</figureCitation>
|
||
); nuptial pad as large and distinCt glandular swelling on dorsal side of Finger I, with extremely minute asperities as regularly, widely sCattered roundish, flat glandular knobs on dorsal side of metaCarpal and preaxial side of proximal phalanx.
|
||
</paragraph>
|
||
<caption id="0398666EFFEFDE7CC3E1768EFD31FAF2" httpUri="https://zenodo.org/record/884329/files/figure.png" pageId="10" pageNumber="301" targetBox="[151,1436,625,1232]" targetPageId="10">
|
||
<paragraph id="575836E6FFEFDE7CC3E1768EFD31FAF2" blockId="10.[151,1435,1253,1306]" pageId="10" pageNumber="301">
|
||
<emphasis id="6593EAF4FFEFDE7CC3E1768EFEE3FB13" bold="true" box="[151,276,1253,1275]" pageId="10" pageNumber="301">FIGURE 5.</emphasis>
|
||
Dorsal view (leFt) and ventral view (right) oF the holotype oF
|
||
<taxonomicName id="90E74D65FFEFDE7CC0E2768CFBBBFB13" authority="Dehling, 2017" authorityName="Dehling" authorityYear="2017" box="[916,1100,1253,1276]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" kingdom="Animalia" order="Anura" pageId="10" pageNumber="301" phylum="Chordata" rank="species" species="kong" status="sp. nov.">
|
||
<emphasis id="6593EAF4FFEFDE7CC0E2768CFBBBFB13" box="[916,1100,1253,1276]" italics="true" pageId="10" pageNumber="301">Limnonectes kong</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="6593EAF4FFEFDE7CC722768DFB51FB13" bold="true" box="[1108,1190,1254,1275]" pageId="10" pageNumber="301">
|
||
<taxonomicNameLabel id="7EA0578FFFEFDE7CC722768DFB51FB13" box="[1108,1190,1254,1275]" pageId="10" pageNumber="301" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(NMBE 1059923, Field no. JMD 368, adult male). Scale bar represents 10 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="0398666EFFEFDE7CC3E174B6FD15F8FA" httpUri="https://zenodo.org/record/884331/files/figure.png" pageId="10" pageNumber="301" targetBox="[151,1436,1356,1736]" targetPageId="10">
|
||
<paragraph id="575836E6FFEFDE7CC3E174B6FD15F8FA" blockId="10.[151,1436,1757,1810]" pageId="10" pageNumber="301">
|
||
<emphasis id="6593EAF4FFEFDE7CC3E174B6FEE4F91A" bold="true" box="[151,275,1757,1779]" pageId="10" pageNumber="301">FIGURE 6.</emphasis>
|
||
Frontal view oF the mandibula (leFt) and ventral view oF the vomer region (right) oF the holotype oF
|
||
<taxonomicName id="90E74D65FFEFDE7CC66A74B5FF3FF8FA" authority="Dehling, 2017" authorityName="Dehling" authorityYear="2017" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" kingdom="Animalia" order="Anura" pageId="10" pageNumber="301" phylum="Chordata" rank="species" species="kong" status="sp. nov.">
|
||
<emphasis id="6593EAF4FFEFDE7CC66A74B5FF3FF8FA" italics="true" pageId="10" pageNumber="301">Limnonectes kong</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="6593EAF4FFEFDE7CC3B87497FEE8F8F9" bold="true" box="[206,287,1788,1809]" pageId="10" pageNumber="301">
|
||
<taxonomicNameLabel id="7EA0578FFFEFDE7CC3B87497FEE8F8F9" box="[206,287,1788,1809]" pageId="10" pageNumber="301" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(NMBE 1059923, adult male). Not to scale.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="575836E6FFEFDE7DC3B17555FE35FE60" blockId="10.[151,1436,1854,2023]" lastBlockId="11.[151,1436,151,392]" lastPageId="11" lastPageNumber="302" pageId="10" pageNumber="301">
|
||
Hindlimbs sturdy, moderately long (LEG/SVL 1.42); tibiofibula Comparatively short (TFL/SVL 0.45), subequal in length to thigh (TFL/THL 0.97); heels not meeting eaCh other when knees flexed and thighs held perpendiCularly to median plane; distinCt, low transverse dermal fold at heel; prominent dermal fold along distal three-fourth of preaxial side of tarsus to proximal end of inner metatarsal tuberCle (
|
||
<figureCitation id="CFDC2A63FFEFDE7CC74B75C1FB75F82B" box="[1085,1154,1962,1987]" captionStart="FIGURE 7" captionStartId="11.[151,250,1900,1922]" captionTargetBox="[151,1436,445,1878]" captionTargetId="figure@11.[151,1436,445,1878]" captionTargetPageId="11" captionText="FIGURE 7. Palmar view oF right hand (leFt) and plantar view oF right Foot (right) oF the holotype (NMBE 1059923, adult male) oF Limnonectes kong sp. nov. (NMBE 1059923, adult male). Scale bar represents 10 mm." httpUri="https://zenodo.org/record/884333/files/figure.png" pageId="10" pageNumber="301">Fig. 7</figureCitation>
|
||
); foot length about equal to tibiofibula length (FOT/TFL 1.01); relative length of toes: I <II <V <III <IV; tips of toes enlarged to small disCs, without CirCummarginal groove; subartiCular tuberCles well developed, rounded, highly elevated, numbering one on Toes I and II, two on Toes III and V, and three on Toe IV; proximal subartiCular tuberCles larger and more prominent than distal ones; pedal webbing formula
|
||
<date id="23591026FFEEDE7DC19372B4FD0DFF10" box="[741,762,223,248]" pageId="11" pageNumber="302">
|
||
<emphasis id="6593EAF4FFEEDE7DC19372B4FD07FF10" bold="true" box="[741,752,223,248]" pageId="11" pageNumber="302">I</emphasis>
|
||
1
|
||
</date>
|
||
/1.5
|
||
<date id="23591026FFEEDE7DC05F72B4FCBEFF10" box="[809,841,223,248]" pageId="11" pageNumber="302">
|
||
<emphasis id="6593EAF4FFEEDE7DC05F72B4FCC8FF10" bold="true" box="[809,831,223,248]" pageId="11" pageNumber="302">II</emphasis>
|
||
1
|
||
</date>
|
||
/1.5
|
||
<date id="23591026FFEEDE7DC00172B4FC55FF10" box="[887,930,223,248]" pageId="11" pageNumber="302">
|
||
<emphasis id="6593EAF4FFEEDE7DC00172B4FC6FFF10" bold="true" box="[887,920,223,248]" pageId="11" pageNumber="302">III</emphasis>
|
||
1
|
||
</date>
|
||
/2
|
||
<date id="23591026FFEEDE7DC0CA72B4FC1DFF10" box="[956,1002,223,248]" pageId="11" pageNumber="302">
|
||
<emphasis id="6593EAF4FFEEDE7DC0CA72B4FC2CFF10" bold="true" box="[956,987,223,248]" pageId="11" pageNumber="302">IV</emphasis>
|
||
2
|
||
</date>
|
||
/1+
|
||
<emphasis id="6593EAF4FFEEDE7DC77872B4FBD5FF10" bold="true" box="[1038,1058,223,248]" pageId="11" pageNumber="302">V</emphasis>
|
||
(
|
||
<figureCitation id="CFDC2A63FFEEDE7DC74272B4FB8DFF10" box="[1076,1146,223,248]" captionStart="FIGURE 7" captionStartId="11.[151,250,1900,1922]" captionTargetBox="[151,1436,445,1878]" captionTargetId="figure@11.[151,1436,445,1878]" captionTargetPageId="11" captionText="FIGURE 7. Palmar view oF right hand (leFt) and plantar view oF right Foot (right) oF the holotype (NMBE 1059923, adult male) oF Limnonectes kong sp. nov. (NMBE 1059923, adult male). Scale bar represents 10 mm." httpUri="https://zenodo.org/record/884333/files/figure.png" pageId="11" pageNumber="302">Fig. 7</figureCitation>
|
||
); dermal flap on preaxial side of Toe I and postaxial side of Toe V from distal edges of metatarsal tuberCles to proximal ends of disCs (
|
||
<figureCitation id="CFDC2A63FFEEDE7DC619736FFF50FEA8" captionStart="FIGURE 7" captionStartId="11.[151,250,1900,1922]" captionTargetBox="[151,1436,445,1878]" captionTargetId="figure@11.[151,1436,445,1878]" captionTargetPageId="11" captionText="FIGURE 7. Palmar view oF right hand (leFt) and plantar view oF right Foot (right) oF the holotype (NMBE 1059923, adult male) oF Limnonectes kong sp. nov. (NMBE 1059923, adult male). Scale bar represents 10 mm." httpUri="https://zenodo.org/record/884333/files/figure.png" pageId="11" pageNumber="302">Fig. 7</figureCitation>
|
||
); inner metatarsal tuberCle very prominent, large and very elongate (length:
|
||
<quantity id="901F9B03FFEEDE7DC08C7343FBA3FEA8" box="[1018,1108,296,320]" metricMagnitude="-3" metricUnit="m" metricValue="4.1" pageId="11" pageNumber="302" unit="mm" value="4.1">4.1 mm</quantity>
|
||
), about three-fourth length of Metatarsal I and slightly less than two-fifths length of Toe I; outer metatarsal tuberCle very small and low, rounded, hardly disCernible (
|
||
<figureCitation id="CFDC2A63FFEEDE7DC2197304FE43FE60" box="[367,436,367,392]" captionStart="FIGURE 7" captionStartId="11.[151,250,1900,1922]" captionTargetBox="[151,1436,445,1878]" captionTargetId="figure@11.[151,1436,445,1878]" captionTargetPageId="11" captionText="FIGURE 7. Palmar view oF right hand (leFt) and plantar view oF right Foot (right) oF the holotype (NMBE 1059923, adult male) oF Limnonectes kong sp. nov. (NMBE 1059923, adult male). Scale bar represents 10 mm." httpUri="https://zenodo.org/record/884333/files/figure.png" pageId="11" pageNumber="302">Fig. 7</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="0398666EFFEEDE7DC3E17507FBD5F848" httpUri="https://zenodo.org/record/884333/files/figure.png" pageId="11" pageNumber="302" targetBox="[151,1436,445,1878]" targetPageId="11">
|
||
<paragraph id="575836E6FFEEDE7DC3E17507FBD5F848" blockId="11.[151,1436,1900,1952]" pageId="11" pageNumber="302">
|
||
<emphasis id="6593EAF4FFEEDE7DC3E17507FEE5F869" bold="true" box="[151,274,1900,1922]" pageId="11" pageNumber="302">FIGURE 7.</emphasis>
|
||
Palmar view oF right hand (leFt) and plantar view oF right Foot (right) oF the holotype (NMBE 1059923, adult male) oF
|
||
<taxonomicName id="90E74D65FFEEDE7DC3C475E0FE9EF848" authority="Dehling, 2017" authorityName="Dehling" authorityYear="2017" box="[178,361,1930,1952]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" kingdom="Animalia" order="Anura" pageId="11" pageNumber="302" phylum="Chordata" rank="species" species="kong" status="sp. nov.">
|
||
<emphasis id="6593EAF4FFEEDE7DC3C475E0FE9EF848" box="[178,361,1930,1952]" italics="true" pageId="11" pageNumber="302">Limnonectes kong</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="6593EAF4FFEEDE7DC20675E0FE37F848" bold="true" box="[368,448,1931,1952]" pageId="11" pageNumber="302">
|
||
<taxonomicNameLabel id="7EA0578FFFEEDE7DC20675E0FE37F848" box="[368,448,1931,1952]" pageId="11" pageNumber="302" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(NMBE 1059923, adult male). Scale bar represents 10 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="0398666EFFE9DE7AC3E171B0FD6FFBE7" httpUri="https://zenodo.org/record/884335/files/figure.png" pageId="12" pageNumber="303" targetBox="[151,1436,193,965]" targetPageId="12">
|
||
<paragraph id="575836E6FFE9DE7AC3E171B0FD6FFBE7" blockId="12.[151,1436,987,1039]" pageId="12" pageNumber="303">
|
||
<emphasis id="6593EAF4FFE9DE7AC3E171B0FEE2FC18" bold="true" box="[151,277,987,1009]" pageId="12" pageNumber="303">FIGURE 8.</emphasis>
|
||
(A) Ventral view oF the euthanized holotype oF
|
||
<taxonomicName id="90E74D65FFE9DE7AC06571B7FC3AFC19" authority="Dehling, 2017" authorityName="Dehling" authorityYear="2017" box="[787,973,987,1009]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" kingdom="Animalia" order="Anura" pageId="12" pageNumber="303" phylum="Chordata" rank="species" species="kong" status="sp. nov.">
|
||
<emphasis id="6593EAF4FFE9DE7AC06571B7FC3AFC19" box="[787,973,987,1009]" italics="true" pageId="12" pageNumber="303">Limnonectes kong</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="6593EAF4FFE9DE7AC0A071B0FBDEFC18" bold="true" box="[982,1065,987,1008]" pageId="12" pageNumber="303">
|
||
<taxonomicNameLabel id="7EA0578FFFE9DE7AC0A071B0FBDEFC18" box="[982,1065,987,1008]" pageId="12" pageNumber="303" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(NMBE 1059923, adult male); (B) detail oF stomach, showing encapsuled nematodes.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="575836E6FFE9DE7AC3B17657FB3BFB75" blockId="12.[151,1437,1084,2009]" pageId="12" pageNumber="303">
|
||
<emphasis id="6593EAF4FFE9DE7AC3B17657FE5DFBBD" bold="true" box="[199,426,1084,1109]" pageId="12" pageNumber="303">Colouration in life.</emphasis>
|
||
Dorsal side dark grey, loosely sCattered tuberCles with a light grey tip; flanks and sides of legs lightened to brownish light grey; ventral side greyish white; ventral side of head light grey with whitish retiCulated pattern (
|
||
<figureCitation id="CFDC2A63FFE9DE7AC20776EFFE42FB75" box="[369,437,1156,1181]" captionStart="FIGURE 8" captionStartId="12.[151,250,987,1009]" captionTargetBox="[151,1436,193,965]" captionTargetId="figure@12.[151,1436,193,965]" captionTargetPageId="12" captionText="FIGURE 8. (A) Ventral view oF the euthanized holotype oF Limnonectes kong sp. nov. (NMBE 1059923, adult male); (B) detail oF stomach, showing encapsuled nematodes." httpUri="https://zenodo.org/record/884335/files/figure.png" pageId="12" pageNumber="303">Fig. 8</figureCitation>
|
||
). Palmar side of metaCarpals and plantar side of metatarsals dark grey.
|
||
</paragraph>
|
||
<paragraph id="575836E6FFE9DE7AC3B176C3FDD5FB0D" blockId="12.[151,1437,1084,2009]" pageId="12" pageNumber="303">
|
||
<emphasis id="6593EAF4FFE9DE7AC3B176C3FDECFB29" bold="true" box="[199,539,1192,1217]" pageId="12" pageNumber="303">Colouration in preservative.</emphasis>
|
||
Pattern and lightness of Colouration very similar to state in life but Colour has turned from grey to brown (
|
||
<figureCitation id="CFDC2A63FFE9DE7AC2B976A7FDE3FB0D" box="[463,532,1228,1253]" captionStart="FIGURE 5" captionStartId="10.[151,250,1253,1275]" captionTargetBox="[151,1436,625,1232]" captionTargetId="figure@10.[151,1436,625,1232]" captionTargetPageId="10" captionText="FIGURE 5. Dorsal view (leFt) and ventral view (right) oF the holotype oF Limnonectes kong sp. nov. (NMBE 1059923, Field no. JMD 368, adult male). Scale bar represents 10 mm." httpUri="https://zenodo.org/record/884329/files/figure.png" pageId="12" pageNumber="303">Fig. 5</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="575836E6FFE9DE7AC3B1769BFD2AF97D" blockId="12.[151,1437,1084,2009]" pageId="12" pageNumber="303">
|
||
<emphasis id="6593EAF4FFE9DE7AC3B1769BFECBFAE1" bold="true" box="[199,316,1264,1289]" pageId="12" pageNumber="303">Variation.</emphasis>
|
||
The male paratypes are very similar in overall size (SVL
|
||
<quantity id="901F9B03FFE9DE7AC0B5769BFB90FAE0" box="[963,1127,1264,1289]" metricMagnitude="-2" metricUnit="m" metricValue="6.7" metricValueMax="7.63" metricValueMin="5.77" pageId="12" pageNumber="303" unit="mm" value="67.0" valueMax="76.3" valueMin="57.7">57.7–76.3 mm</quantity>
|
||
) and head proportion (HW/ SVL 0.41–0.47, HL/SVL 0.44–0.50) to the holotype. The females are smaller (SVL
|
||
<quantity id="901F9B03FFE9DE7AC70F777FFAD5FAC5" box="[1145,1314,1300,1325]" metricMagnitude="-2" metricUnit="m" metricValue="5.595" metricValueMax="5.9399999999999995" metricValueMin="5.25" pageId="12" pageNumber="303" unit="mm" value="55.95" valueMax="59.4" valueMin="52.5">52.5–59.4 mm</quantity>
|
||
) and have shorter and narrower heads (HW/SVL 0.38–0.40; HL/SVL HL/SVL 0.39–0.43) (
|
||
<figureCitation id="CFDC2A63FFE9DE7AC75B7753FB84FAB8" box="[1069,1139,1336,1361]" captionStart="FIGURE 9" captionStartId="13.[151,250,1780,1802]" captionTargetBox="[151,1436,805,1769]" captionTargetId="figure@13.[151,1436,805,1769]" captionTargetPageId="13" captionText="FIGURE 9. Limnonectes kong sp. nov.: (A) Female paratype (NMBE 1059918) calling From a cavity in the ground, Matang, Sarawak; (B) Female paratype (NMBE 1059928) perched on a rock along a stream, Gading NP, Sarawak; (C) lateral view and (D) dorsal view oF a male paratype (NMBE 1059924), Bako NP, Sarawak." httpUri="https://zenodo.org/record/884337/files/figure.png" pageId="12" pageNumber="303">Fig. 9</figureCitation>
|
||
). The odontoids are muCh larger in adult males (OL/SVL 0.053–0.087) than in females (OL/SVL 0.036–0.044). Extend of webbing does not differ between the sexes and is generally very Conservative without a marked deviation from the extent observed in the holotype. The tympanum is disCernible only in the smallest paratype (NMBE 1059917, juvenile male). The granulation on the surfaCe of the posterior trunk and the hindlimbs is most prominently developed in the largest males and in adult females (
|
||
<figureCitation id="CFDC2A63FFE9DE7AC2947787FDDDF9ED" box="[482,554,1516,1541]" captionStart="FIGURE 9" captionStartId="13.[151,250,1780,1802]" captionTargetBox="[151,1436,805,1769]" captionTargetId="figure@13.[151,1436,805,1769]" captionTargetPageId="13" captionText="FIGURE 9. Limnonectes kong sp. nov.: (A) Female paratype (NMBE 1059918) calling From a cavity in the ground, Matang, Sarawak; (B) Female paratype (NMBE 1059928) perched on a rock along a stream, Gading NP, Sarawak; (C) lateral view and (D) dorsal view oF a male paratype (NMBE 1059924), Bako NP, Sarawak." httpUri="https://zenodo.org/record/884337/files/figure.png" pageId="12" pageNumber="303">Fig. 9</figureCitation>
|
||
). The surfaCe of subadult males is muCh smoother. This suggests that these tuberCles are developed in fully mature speCimens only. Colouration varies between speCimens and some are light grey, other light brown dorsally in life, with darker grey or brown pattern Consisting of transverse interorbital bars, stripes on the limbs and along the supratympaniC fold (
|
||
<figureCitation id="CFDC2A63FFE9DE7AC06C7433FC95F998" box="[794,866,1624,1649]" captionStart="FIGURE 9" captionStartId="13.[151,250,1780,1802]" captionTargetBox="[151,1436,805,1769]" captionTargetId="figure@13.[151,1436,805,1769]" captionTargetPageId="13" captionText="FIGURE 9. Limnonectes kong sp. nov.: (A) Female paratype (NMBE 1059918) calling From a cavity in the ground, Matang, Sarawak; (B) Female paratype (NMBE 1059928) perched on a rock along a stream, Gading NP, Sarawak; (C) lateral view and (D) dorsal view oF a male paratype (NMBE 1059924), Bako NP, Sarawak." httpUri="https://zenodo.org/record/884337/files/figure.png" pageId="12" pageNumber="303">Fig. 9</figureCitation>
|
||
). Two paratypes have a ConspiCuous longitudinal yellowish line on the baCk along the spine (
|
||
<figureCitation id="CFDC2A63FFE9DE7AC10B7417FD37F97D" box="[637,704,1660,1685]" captionStart="FIGURE 9" captionStartId="13.[151,250,1780,1802]" captionTargetBox="[151,1436,805,1769]" captionTargetId="figure@13.[151,1436,805,1769]" captionTargetPageId="13" captionText="FIGURE 9. Limnonectes kong sp. nov.: (A) Female paratype (NMBE 1059918) calling From a cavity in the ground, Matang, Sarawak; (B) Female paratype (NMBE 1059928) perched on a rock along a stream, Gading NP, Sarawak; (C) lateral view and (D) dorsal view oF a male paratype (NMBE 1059924), Bako NP, Sarawak." httpUri="https://zenodo.org/record/884337/files/figure.png" pageId="12" pageNumber="303">Fig. 9</figureCitation>
|
||
b).
|
||
</paragraph>
|
||
<paragraph id="575836E6FFE9DE7BC3B174CBFDFDFF10" blockId="12.[151,1437,1084,2009]" lastBlockId="13.[151,1437,151,752]" lastPageId="13" lastPageNumber="304" pageId="12" pageNumber="303">
|
||
<emphasis id="6593EAF4FFE9DE7AC3B174CBFDE4F951" bold="true" box="[199,531,1696,1721]" pageId="12" pageNumber="303">Morphological comparison.</emphasis>
|
||
Based on the inComplete toe webbing,
|
||
<taxonomicName id="90E74D65FFE9DE7AC0AA74CAFBC4F950" authorityName="Dehling" authorityYear="2017" box="[988,1075,1696,1720]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" kingdom="Animalia" order="Anura" pageId="12" pageNumber="303" phylum="Chordata" rank="species" species="kong">
|
||
<emphasis id="6593EAF4FFE9DE7AC0AA74CAFBC4F950" box="[988,1075,1696,1720]" italics="true" pageId="12" pageNumber="303">L. kong</emphasis>
|
||
</taxonomicName>
|
||
is easily distinguished from all desCribed Bornean speCies of the
|
||
<taxonomicName id="90E74D65FFE9DE7AC16A74AEFD8AF934" authorityName=": Gunther" authorityYear="1874" box="[540,637,1732,1756]" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="12" pageNumber="303" phylum="Chordata" rank="species" species="kuhlii">
|
||
<emphasis id="6593EAF4FFE9DE7AC16A74AEFD8AF934" box="[540,637,1732,1756]" italics="true" pageId="12" pageNumber="303">L. kuhlii</emphasis>
|
||
</taxonomicName>
|
||
group with fully-webbed toes, i.e.
|
||
<taxonomicName id="90E74D65FFE9DE7AC76074AEFABDF934" box="[1046,1354,1732,1756]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" kingdom="Animalia" order="Anura" pageId="12" pageNumber="303" phylum="Chordata" rank="species" species="conspicillatus">
|
||
<emphasis id="6593EAF4FFE9DE7AC76074AEFABDF934" box="[1046,1354,1732,1756]" italics="true" pageId="12" pageNumber="303">Limnonectes conspicillatus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<emphasis id="6593EAF4FFE9DE7AC6F374AEFEE0F8E8" italics="true" pageId="12" pageNumber="303">
|
||
<taxonomicName id="90E74D65FFE9DE7AC6F374AEFEE4F8E8" authorityName="Matsui, Dubois & Ohler" authorityYear="2013" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="12" pageNumber="303" phylum="Chordata" rank="species" species="mocquardi">L. mocquardi</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
Adult females of
|
||
<taxonomicName id="90E74D65FFE9DE7AC29B7482FDB3F8E8" authorityName="Dehling" authorityYear="2017" box="[493,580,1768,1792]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" kingdom="Animalia" order="Anura" pageId="12" pageNumber="303" phylum="Chordata" rank="species" species="kong">
|
||
<emphasis id="6593EAF4FFE9DE7AC29B7482FDB3F8E8" box="[493,580,1768,1792]" italics="true" pageId="12" pageNumber="303">L. kong</emphasis>
|
||
</taxonomicName>
|
||
(SVL
|
||
<quantity id="901F9B03FFE9DE7AC1E17483FCC8F8E8" box="[663,831,1768,1793]" metricMagnitude="-2" metricUnit="m" metricValue="5.595" metricValueMax="5.9399999999999995" metricValueMin="5.25" pageId="12" pageNumber="303" unit="mm" value="55.95" valueMax="59.4" valueMin="52.5">52.5–59.4 mm</quantity>
|
||
) are smaller than females of
|
||
<taxonomicName id="90E74D65FFE9DE7AC7E17482FAA3F8E8" box="[1175,1364,1768,1792]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" kingdom="Animalia" order="Anura" pageId="12" pageNumber="303" phylum="Chordata" rank="species" species="conspicillatus">
|
||
<emphasis id="6593EAF4FFE9DE7AC7E17482FAA3F8E8" box="[1175,1364,1768,1792]" italics="true" pageId="12" pageNumber="303">L. conspicillatus</emphasis>
|
||
</taxonomicName>
|
||
(SVL 64.9–69.0 mm) and are usually larger than females of
|
||
<taxonomicName id="90E74D65FFE9DE7AC1847566FC70F8CC" authorityName="Matsui, Dubois & Ohler" authorityYear="2013" box="[754,903,1804,1828]" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="12" pageNumber="303" phylum="Chordata" rank="species" species="mocquardi">
|
||
<emphasis id="6593EAF4FFE9DE7AC1847566FC70F8CC" box="[754,903,1804,1828]" italics="true" pageId="12" pageNumber="303">L. mocquardi</emphasis>
|
||
</taxonomicName>
|
||
(SVL 36.6–53.0 mm;
|
||
<bibRefCitation id="33764B17FFE9DE7AC7F27567FAA5F8CD" author="Matsui" box="[1156,1362,1804,1829]" pageId="12" pageNumber="303" refString="Matsui, M., Dubois, A. & Ohler, A. (2013) New replacement name For Rana paradoxa Mocquard, 1890 with designations oF lectotypes For Rana paradoxa and Rana conspicillata Gunther, 1872: both synonymized with Limnonectes kuhlii (Tschudi, 1838) (Dicroglossidae: Dicroglossinae). Asian Herpetological Research, 4 (3), 187 - 189." type="journal article" year="2013">
|
||
Matsui
|
||
<emphasis id="6593EAF4FFE9DE7AC7AE7566FAFFF8CC" box="[1240,1288,1804,1828]" italics="true" pageId="12" pageNumber="303">et al</emphasis>
|
||
. 2013
|
||
</bibRefCitation>
|
||
; JMD, unpubl. data). Males of
|
||
<taxonomicName id="90E74D65FFE9DE7AC2DF755AFE09F8A0" authorityName="Dehling" authorityYear="2017" box="[425,510,1840,1864]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" kingdom="Animalia" order="Anura" pageId="12" pageNumber="303" phylum="Chordata" rank="species" species="kong">
|
||
<emphasis id="6593EAF4FFE9DE7AC2DF755AFE09F8A0" box="[425,510,1840,1864]" italics="true" pageId="12" pageNumber="303">L. kong</emphasis>
|
||
</taxonomicName>
|
||
(SVL
|
||
<quantity id="901F9B03FFE9DE7AC138755BFD02F8A0" box="[590,757,1840,1865]" metricMagnitude="-2" metricUnit="m" metricValue="6.7" metricValueMax="7.63" metricValueMin="5.77" pageId="12" pageNumber="303" unit="mm" value="67.0" valueMax="76.3" valueMin="57.7">57.7–76.3 mm</quantity>
|
||
) are usually larger than males of
|
||
<taxonomicName id="90E74D65FFE9DE7AC700755AFAFAF8A0" authorityName="Matsui, Dubois & Ohler" authorityYear="2013" box="[1142,1293,1840,1864]" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="12" pageNumber="303" phylum="Chordata" rank="species" species="mocquardi">
|
||
<emphasis id="6593EAF4FFE9DE7AC700755AFAFAF8A0" box="[1142,1293,1840,1864]" italics="true" pageId="12" pageNumber="303">L. mocquardi</emphasis>
|
||
</taxonomicName>
|
||
(SVL
|
||
<quantity id="901F9B03FFE9DE7AC62A755BFEF4F884" metricMagnitude="-2" metricUnit="m" metricValue="5.24" metricValueMax="5.82" metricValueMin="4.66" pageId="12" pageNumber="303" unit="mm" value="52.4" valueMax="58.2" valueMin="46.6">46.6– 58.2 mm</quantity>
|
||
;
|
||
<bibRefCitation id="33764B17FFE9DE7AC265753FFE06F885" author="Matsui" box="[275,497,1876,1901]" pageId="12" pageNumber="303" refString="Matsui, M., Dubois, A. & Ohler, A. (2013) New replacement name For Rana paradoxa Mocquard, 1890 with designations oF lectotypes For Rana paradoxa and Rana conspicillata Gunther, 1872: both synonymized with Limnonectes kuhlii (Tschudi, 1838) (Dicroglossidae: Dicroglossinae). Asian Herpetological Research, 4 (3), 187 - 189." type="journal article" year="2013">
|
||
Matsui
|
||
<emphasis id="6593EAF4FFE9DE7AC21B753EFE54F884" box="[365,419,1876,1900]" italics="true" pageId="12" pageNumber="303">et al</emphasis>
|
||
. 2013
|
||
</bibRefCitation>
|
||
; JMD, unpubl. data). In addition,
|
||
<taxonomicName id="90E74D65FFE9DE7AC0FD753EFC13F884" authorityName="Dehling" authorityYear="2017" box="[907,996,1876,1900]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" kingdom="Animalia" order="Anura" pageId="12" pageNumber="303" phylum="Chordata" rank="species" species="kong">
|
||
<emphasis id="6593EAF4FFE9DE7AC0FD753EFC13F884" box="[907,996,1876,1900]" italics="true" pageId="12" pageNumber="303">L. kong</emphasis>
|
||
</taxonomicName>
|
||
differs from
|
||
<taxonomicName id="90E74D65FFE9DE7AC7F1753EFAB2F884" box="[1159,1349,1876,1900]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" kingdom="Animalia" order="Anura" pageId="12" pageNumber="303" phylum="Chordata" rank="species" species="conspicillatus">
|
||
<emphasis id="6593EAF4FFE9DE7AC7F1753EFAB2F884" box="[1159,1349,1876,1900]" italics="true" pageId="12" pageNumber="303">L. conspicillatus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="90E74D65FFE9DE7AC6F3753EFEF8F878" authorityName="Matsui, Dubois & Ohler" authorityYear="2013" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="12" pageNumber="303" phylum="Chordata" rank="species" species="mocquardi">
|
||
<emphasis id="6593EAF4FFE9DE7AC6F3753EFEF8F878" italics="true" pageId="12" pageNumber="303">L. mocquardi</emphasis>
|
||
</taxonomicName>
|
||
in the tympanum being indisCernible (vs. ususally disCernible in
|
||
<taxonomicName id="90E74D65FFE9DE7AC0887512FB4FF878" box="[1022,1208,1912,1936]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" kingdom="Animalia" order="Anura" pageId="12" pageNumber="303" phylum="Chordata" rank="species" species="conspicillatus">
|
||
<emphasis id="6593EAF4FFE9DE7AC0887512FB4FF878" box="[1022,1208,1912,1936]" italics="true" pageId="12" pageNumber="303">L. conspicillatus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="90E74D65FFE9DE7AC7827512FA7CF878" authorityName="Matsui, Dubois & Ohler" authorityYear="2013" box="[1268,1419,1912,1936]" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="12" pageNumber="303" phylum="Chordata" rank="species" species="mocquardi">
|
||
<emphasis id="6593EAF4FFE9DE7AC7827512FA7CF878" box="[1268,1419,1912,1936]" italics="true" pageId="12" pageNumber="303">L. mocquardi</emphasis>
|
||
</taxonomicName>
|
||
), the vomerine ridge being short and separated from eaCh other by half length of individual ridge (vs. longer and separated by less than width of individual ridge in
|
||
<taxonomicName id="90E74D65FFE9DE7AC19B75AAFC5DF830" box="[749,938,1984,2008]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" kingdom="Animalia" order="Anura" pageId="12" pageNumber="303" phylum="Chordata" rank="species" species="conspicillatus">
|
||
<emphasis id="6593EAF4FFE9DE7AC19B75AAFC5DF830" box="[749,938,1984,2008]" italics="true" pageId="12" pageNumber="303">L. conspicillatus</emphasis>
|
||
</taxonomicName>
|
||
; shorter and separated by a little less than individual length in
|
||
<taxonomicName id="90E74D65FFE8DE7BC2F772F2FDEEFF58" authorityName="Matsui, Dubois & Ohler" authorityYear="2013" box="[385,537,152,176]" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="13" pageNumber="304" phylum="Chordata" rank="species" species="mocquardi">
|
||
<emphasis id="6593EAF4FFE8DE7BC2F772F2FDEEFF58" box="[385,537,152,176]" italics="true" pageId="13" pageNumber="304">L. mocquardi</emphasis>
|
||
</taxonomicName>
|
||
), the middle fingers having very narrow, non-moveable dermal fringes on the phalanges only (vs. somewhat broader fringes in
|
||
<taxonomicName id="90E74D65FFE8DE7BC1BA72D6FC93FF3C" authorityName="Matsui, Dubois & Ohler" authorityYear="2013" box="[716,868,188,212]" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="13" pageNumber="304" phylum="Chordata" rank="species" species="mocquardi">
|
||
<emphasis id="6593EAF4FFE8DE7BC1BA72D6FC93FF3C" box="[716,868,188,212]" italics="true" pageId="13" pageNumber="304">L. mocquardi</emphasis>
|
||
</taxonomicName>
|
||
; broad, moveable flaps present on phalanges and metaCarpals in
|
||
<taxonomicName id="90E74D65FFE8DE7BC236728AFE0DFF10" box="[320,506,224,248]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" kingdom="Animalia" order="Anura" pageId="13" pageNumber="304" phylum="Chordata" rank="species" species="conspicillatus">
|
||
<emphasis id="6593EAF4FFE8DE7BC236728AFE0DFF10" box="[320,506,224,248]" italics="true" pageId="13" pageNumber="304">L. conspicillatus</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="575836E6FFE8DE7BC3B1736FFDC8FD18" blockId="13.[151,1437,151,752]" pageId="13" pageNumber="304">
|
||
Of the speCies of the
|
||
<taxonomicName id="90E74D65FFE8DE7BC2B5736EFDD3FEF4" authorityName=": Gunther" authorityYear="1874" box="[451,548,260,284]" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="13" pageNumber="304" phylum="Chordata" rank="species" species="kuhlii">
|
||
<emphasis id="6593EAF4FFE8DE7BC2B5736EFDD3FEF4" box="[451,548,260,284]" italics="true" pageId="13" pageNumber="304">L. kuhlii</emphasis>
|
||
</taxonomicName>
|
||
group from Borneo with inComplete webbing,
|
||
<taxonomicName id="90E74D65FFE8DE7BC73C736EFEDEFEA8" authority="Inger, Boeadi & Taufik, 1996" authorityName="Inger, Boeadi & Taufik" authorityYear="1996" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" kingdom="Animalia" order="Anura" pageId="13" pageNumber="304" phylum="Chordata" rank="species" species="asperatus">
|
||
<emphasis id="6593EAF4FFE8DE7BC73C736EFB2FFEF4" box="[1098,1240,261,284]" italics="true" pageId="13" pageNumber="304">L. asperatus</emphasis>
|
||
Inger, Boeadi & Taufik, 1996
|
||
</taxonomicName>
|
||
(SVL of female
|
||
<quantity id="901F9B03FFE8DE7BC283734CFDBEFED7" box="[501,585,295,320]" metricMagnitude="-2" metricUnit="m" metricValue="3.6" pageId="13" pageNumber="304" unit="mm" value="36.0">36 mm</quantity>
|
||
, adult males
|
||
<quantity id="901F9B03FFE8DE7BC1917343FC95FEA8" box="[743,866,296,320]" metricMagnitude="-2" metricUnit="m" metricValue="3.6" metricValueMax="3.8" metricValueMin="3.4" pageId="13" pageNumber="304" unit="mm" value="36.0" valueMax="38.0" valueMin="34.0">34-38 mm</quantity>
|
||
),
|
||
<taxonomicName id="90E74D65FFE8DE7BC00E7342FA6BFEA8" authority="Matsui, Nishikawa & Eto, 2014" authorityName="Matsui, Nishikawa & Eto" authorityYear="2014" box="[888,1436,295,320]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" kingdom="Animalia" order="Anura" pageId="13" pageNumber="304" phylum="Chordata" rank="species" species="cintalubang">
|
||
<emphasis id="6593EAF4FFE8DE7BC00E7342FBE8FEA8" box="[888,1055,296,320]" italics="true" pageId="13" pageNumber="304">L. cintalubang</emphasis>
|
||
<bibRefCitation id="33764B17FFE8DE7BC75C734CFA6BFEA8" author="Matsui" box="[1066,1436,295,320]" pageId="13" pageNumber="304" refString="Matsui, M., Nishikawa, K. & Eto, K. (2014 b) A new burrow-utilising Fanged Frog From Sarawak, East Malaysia (Anura: Dicroglossidae). The Raffles Bulletin of Zoology, 62, 679 - 687." type="journal article" year="2014">Matsui, Nishikawa & Eto, 2014</bibRefCitation>
|
||
</taxonomicName>
|
||
(females
|
||
<quantity id="901F9B03FFE8DE7BC27F7327FE78FE8C" box="[265,399,332,356]" metricMagnitude="-2" metricUnit="m" metricValue="3.75" metricValueMax="4.3" metricValueMin="3.2" pageId="13" pageNumber="304" unit="mm" value="37.5" valueMax="43.0" valueMin="32.0">32–43 mm</quantity>
|
||
, male
|
||
<quantity id="901F9B03FFE8DE7BC29B7327FDBCFE8C" box="[493,587,332,357]" metricMagnitude="-2" metricUnit="m" metricValue="4.5" pageId="13" pageNumber="304" unit="mm" value="45.0">45 mm</quantity>
|
||
),
|
||
<taxonomicName id="90E74D65FFE8DE7BC11F7326FD28FE8C" box="[617,735,332,356]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" kingdom="Animalia" order="Anura" pageId="13" pageNumber="304" phylum="Chordata" rank="species" species="hikidai">
|
||
<emphasis id="6593EAF4FFE8DE7BC11F7326FD28FE8C" box="[617,735,332,356]" italics="true" pageId="13" pageNumber="304">L. hikidai</emphasis>
|
||
</taxonomicName>
|
||
(females
|
||
<quantity id="901F9B03FFE8DE7BC0157327FBE4FE8C" box="[867,1043,332,356]" metricMagnitude="-2" metricUnit="m" metricValue="3.7099999999999995" metricValueMax="4.04" metricValueMin="3.38" pageId="13" pageNumber="304" unit="mm" value="37.099999999999994" valueMax="40.4" valueMin="33.8">33.8–40.4 mm</quantity>
|
||
, males
|
||
<quantity id="901F9B03FFE8DE7BC70A7327FAD9FE8C" box="[1148,1326,332,357]" metricMagnitude="-2" metricUnit="m" metricValue="3.345" metricValueMax="3.6799999999999997" metricValueMin="3.01" pageId="13" pageNumber="304" unit="mm" value="33.45" valueMax="36.8" valueMin="30.1">30.1–36.8 mm</quantity>
|
||
), and
|
||
<taxonomicName id="90E74D65FFE8DE7BC6F37326FE2AFE60" authority="Matsui, 2015" authorityName="Matsui" authorityYear="2015" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" kingdom="Animalia" order="Anura" pageId="13" pageNumber="304" phylum="Chordata" rank="species" species="sinuatodorsalis">
|
||
<emphasis id="6593EAF4FFE8DE7BC6F37326FEB3FE60" italics="true" pageId="13" pageNumber="304">L. sinuatodorsalis</emphasis>
|
||
<bibRefCitation id="33764B17FFE8DE7BC23C7304FE2AFE60" author="Matsui" box="[330,477,367,392]" pageId="13" pageNumber="304" refString="Matsui, M. (2015) A new species oF Limnonectes From the border oF East Kalimantan and Sarawak, Borneo Island (Anura, Dicroglossidae). Current Herpetology, 34, 120 - 127. https: // doi. org / 10.5358 / hsj. 34.120" type="journal article" year="2015">Matsui, 2015</bibRefCitation>
|
||
</taxonomicName>
|
||
(females
|
||
<quantity id="901F9B03FFE8DE7BC13D731BFD32FE6F" box="[587,709,368,392]" metricMagnitude="-2" metricUnit="m" metricValue="3.6" metricValueMax="3.8" metricValueMin="3.4" pageId="13" pageNumber="304" unit="mm" value="36.0" valueMax="38.0" valueMin="34.0">34–38 mm</quantity>
|
||
, males
|
||
<quantity id="901F9B03FFE8DE7BC06E731BFC66FE60" box="[792,913,368,392]" metricMagnitude="-2" metricUnit="m" metricValue="3.0" metricValueMax="3.2" metricValueMin="2.8" pageId="13" pageNumber="304" unit="mm" value="30.0" valueMax="32.0" valueMin="28.0">28–32 mm</quantity>
|
||
) are muCh smaller than
|
||
<taxonomicName id="90E74D65FFE8DE7BC7EA731AFB07FE60" authorityName="Dehling" authorityYear="2017" box="[1180,1264,368,392]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" kingdom="Animalia" order="Anura" pageId="13" pageNumber="304" phylum="Chordata" rank="species" species="kong">
|
||
<emphasis id="6593EAF4FFE8DE7BC7EA731AFB07FE60" box="[1180,1264,368,392]" italics="true" pageId="13" pageNumber="304">L. kong</emphasis>
|
||
</taxonomicName>
|
||
(females
|
||
<quantity id="901F9B03FFE8DE7BC62B7304FF08FE44" metricMagnitude="-2" metricUnit="m" metricValue="5.595" metricValueMax="5.9399999999999995" metricValueMin="5.25" pageId="13" pageNumber="304" unit="mm" value="55.95" valueMax="59.4" valueMin="52.5">52.5– 59.4 mm</quantity>
|
||
, males
|
||
<quantity id="901F9B03FFE8DE7BC22173FFFE0AFE44" box="[343,509,404,429]" metricMagnitude="-2" metricUnit="m" metricValue="6.7" metricValueMax="7.63" metricValueMin="5.77" pageId="13" pageNumber="304" unit="mm" value="67.0" valueMax="76.3" valueMin="57.7">57.7–76.3 mm</quantity>
|
||
) and laCk the massive enlargement of the head in adult males (
|
||
<bibRefCitation id="33764B17FFE8DE7BC7A573FFFA60FE45" author="Inger" box="[1235,1431,404,429]" pageId="13" pageNumber="304" refString="Inger, R. F., Boeadi & TauFik, A. (1996) New species oF ranid Frogs (Amphibia: Anura) From Central Kalimantan, Borneo. The Raffles Bulletin of Zoology, 44, 363 - 369." type="journal article" year="1996">
|
||
Inger
|
||
<emphasis id="6593EAF4FFE8DE7BC66F73FEFABCFE44" box="[1305,1355,404,428]" italics="true" pageId="13" pageNumber="304">et al</emphasis>
|
||
. 1996
|
||
</bibRefCitation>
|
||
; Matsui & Nishikawa 2014;
|
||
<bibRefCitation id="33764B17FFE8DE7BC2B873DCFD5CFE38" author="Matsui" box="[462,683,439,464]" pageId="13" pageNumber="304" refString="Matsui, M., Nishikawa, K. & Eto, K. (2014 b) A new burrow-utilising Fanged Frog From Sarawak, East Malaysia (Anura: Dicroglossidae). The Raffles Bulletin of Zoology, 62, 679 - 687." type="journal article" year="2014" yearSuffix="b">
|
||
Matsui
|
||
<emphasis id="6593EAF4FFE8DE7BC15573D2FDA4FE38" box="[547,595,440,464]" italics="true" pageId="13" pageNumber="304">et al</emphasis>
|
||
. 2014b
|
||
</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="33764B17FFE8DE7BC1C073DCFCA5FE38" author="Dehling" box="[694,850,439,464]" pageId="13" pageNumber="304" refString="Dehling, J. M. (2014) Eine neue FangzahnFroschart der Gattung Limnonectes (Anura: Dicroglossidae) vom Gunung Lawit, Malaiische Halbinsel / A new species oF Fanged Frog oF the genus Limnonectes (Anura: Dicroglossidae) From Gunung Lawit, Peninsular Malaysia. Sauria, 36, 17 - 30." type="journal article" year="2014">Dehling 2014</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="33764B17FFE8DE7BC02873DCFC07FE38" author="Matsui" box="[862,1008,439,464]" pageId="13" pageNumber="304" refString="Matsui, M. (2015) A new species oF Limnonectes From the border oF East Kalimantan and Sarawak, Borneo Island (Anura, Dicroglossidae). Current Herpetology, 34, 120 - 127. https: // doi. org / 10.5358 / hsj. 34.120" type="journal article" year="2015">Matsui 2015</bibRefCitation>
|
||
). The new speCies differs in the extent of webbing from all fully webbed speCies from the Sunda region, i.e.
|
||
<taxonomicName id="90E74D65FFE8DE7BC0AA73B6FB42FE1C" baseAuthorityName="TsChudi" baseAuthorityYear="1838" box="[988,1205,476,500]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" kingdom="Animalia" order="Anura" pageId="13" pageNumber="304" phylum="Chordata" rank="species" species="kuhlii">
|
||
<emphasis id="6593EAF4FFE8DE7BC0AA73B6FB42FE1C" box="[988,1205,476,500]" italics="true" pageId="13" pageNumber="304">Limnonectes kuhlii</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="90E74D65FFE8DE7BC7BE73B6FE42FDF0" authority="Matsui, Belabut & Ahmad, 2014" authorityName="Matsui, Belabut & Ahmad" authorityYear="2014" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" kingdom="Animalia" order="Anura" pageId="13" pageNumber="304" phylum="Chordata" rank="species" species="selatan">
|
||
<emphasis id="6593EAF4FFE8DE7BC7BE73B6FACCFE1C" box="[1224,1339,476,500]" italics="true" pageId="13" pageNumber="304">L. selatan</emphasis>
|
||
<bibRefCitation id="33764B17FFE8DE7BC63173B7FE42FDF0" author="Matsui" pageId="13" pageNumber="304" refString="Matsui, M., Belabut, D. M. & Ahmad, N. (2014 a) Two new species oF Fanged Frogs From Peninsular Malaysia (Anura: Dicroglossidae). Zootaxa, 3881 (1), 75 - 93. https: // doi. org / 10.11646 / zootaxa. 3881.1.6" type="journal article" year="2014">Matsui, Belabut & Ahmad, 2014</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="90E74D65FFE8DE7BC2B4706AFDBAFDF0" box="[450,589,512,536]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" kingdom="Animalia" order="Anura" pageId="13" pageNumber="304" phylum="Chordata" rank="species" species="sisikdagu">
|
||
<emphasis id="6593EAF4FFE8DE7BC2B4706AFDBAFDF0" box="[450,589,512,536]" italics="true" pageId="13" pageNumber="304">L. sisikdagu</emphasis>
|
||
</taxonomicName>
|
||
MCLeod,
|
||
<bibRefCitation id="33764B17FFE8DE7BC1B3706BFB57FDF0" author="Horner" box="[709,1184,511,536]" pageId="13" pageNumber="304" refString="McLeod, D. S., Horner, S. J., Husted, C., Barley, A. & Iskandar, D. (2011) " Same-same, but diFFerent ": an unusual new species oF the Limnonectes kuhlii complex From west Sumatra (Anura: Dicroglossidae). Zootaxa, 2883, 52 - 64." type="journal article" year="2011">Horner, Husted, Barley & Iskandar, 2011</bibRefCitation>
|
||
, and
|
||
<taxonomicName id="90E74D65FFE8DE7BC797706AFE59FDD4" authority="Matsui, Belabut & Ahmad, 2014" authorityName="Matsui, Belabut & Ahmad" authorityYear="2014" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" kingdom="Animalia" order="Anura" pageId="13" pageNumber="304" phylum="Chordata" rank="species" species="utara">
|
||
<emphasis id="6593EAF4FFE8DE7BC797706AFACAFDF0" box="[1249,1341,513,536]" italics="true" pageId="13" pageNumber="304">L. utara</emphasis>
|
||
<bibRefCitation id="33764B17FFE8DE7BC6317394FE59FDD4" author="Matsui" pageId="13" pageNumber="304" refString="Matsui, M., Belabut, D. M. & Ahmad, N. (2014 a) Two new species oF Fanged Frogs From Peninsular Malaysia (Anura: Dicroglossidae). Zootaxa, 3881 (1), 75 - 93. https: // doi. org / 10.11646 / zootaxa. 3881.1.6" type="journal article" year="2014">Matsui, Belabut & Ahmad, 2014</bibRefCitation>
|
||
</taxonomicName>
|
||
(MCLeod
|
||
<emphasis id="6593EAF4FFE8DE7BC151704EFDAEFDD4" box="[551,601,548,572]" italics="true" pageId="13" pageNumber="304">et al</emphasis>
|
||
. 2011;
|
||
<bibRefCitation id="33764B17FFE8DE7BC1C6704FFC66FDD4" author="Matsui" box="[688,913,548,573]" pageId="13" pageNumber="304" refString="Matsui, M., Belabut, D. M. & Ahmad, N. (2014 a) Two new species oF Fanged Frogs From Peninsular Malaysia (Anura: Dicroglossidae). Zootaxa, 3881 (1), 75 - 93. https: // doi. org / 10.11646 / zootaxa. 3881.1.6" type="journal article" year="2014" yearSuffix="a">
|
||
Matsui
|
||
<emphasis id="6593EAF4FFE8DE7BC070704EFCCFFDD4" box="[774,824,548,572]" italics="true" pageId="13" pageNumber="304">et al</emphasis>
|
||
. 2014a
|
||
</bibRefCitation>
|
||
).
|
||
<taxonomicName id="90E74D65FFE8DE7BC0D2704EFABBFDD4" authority="Dehling, 2014" authorityName="Dehling" authorityYear="2014" box="[932,1356,548,573]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" kingdom="Animalia" order="Anura" pageId="13" pageNumber="304" phylum="Chordata" rank="species" species="deinodon">
|
||
<emphasis id="6593EAF4FFE8DE7BC0D2704EFB55FDD4" box="[932,1186,548,572]" italics="true" pageId="13" pageNumber="304">Limnonectes deinodon</emphasis>
|
||
<bibRefCitation id="33764B17FFE8DE7BC7DD704FFABBFDD4" author="Dehling" box="[1195,1356,548,573]" pageId="13" pageNumber="304" refString="Dehling, J. M. (2014) Eine neue FangzahnFroschart der Gattung Limnonectes (Anura: Dicroglossidae) vom Gunung Lawit, Malaiische Halbinsel / A new species oF Fanged Frog oF the genus Limnonectes (Anura: Dicroglossidae) From Gunung Lawit, Peninsular Malaysia. Sauria, 36, 17 - 30." type="journal article" year="2014">Dehling, 2014</bibRefCitation>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="90E74D65FFE8DE7BC6F3704EFEF0FD88" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" kingdom="Animalia" order="Anura" pageId="13" pageNumber="304" phylum="Chordata" rank="species" species="khasianus">
|
||
<emphasis id="6593EAF4FFE8DE7BC6F3704EFEF0FD88" italics="true" pageId="13" pageNumber="304">L. khasianus</emphasis>
|
||
</taxonomicName>
|
||
have less extensive webbing on the toes (
|
||
<emphasis id="6593EAF4FFE8DE7BC184702CFD0AFD88" bold="true" box="[754,765,583,608]" pageId="13" pageNumber="304">I</emphasis>
|
||
1.75/2+
|
||
<emphasis id="6593EAF4FFE8DE7BC025702CFC9EFD88" bold="true" box="[851,873,583,608]" pageId="13" pageNumber="304">II</emphasis>
|
||
1.5/
|
||
<date id="23591026FFE8DE7BC0E0702CFC3BFD88" box="[918,972,583,608]" bridgedPair="-" pageId="13" pageNumber="304">
|
||
3-
|
||
<emphasis id="6593EAF4FFE8DE7BC0DD702CFC3BFD88" bold="true" box="[939,972,583,608]" pageId="13" pageNumber="304">III</emphasis>
|
||
</date>
|
||
1.75/3
|
||
<date id="23591026FFE8DE7BC765702CFBC9FD88" box="[1043,1086,583,608]" pageId="13" pageNumber="304">
|
||
<emphasis id="6593EAF4FFE8DE7BC765702CFBC5FD88" bold="true" box="[1043,1074,583,608]" pageId="13" pageNumber="304">IV</emphasis>
|
||
3
|
||
</date>
|
||
/1.5
|
||
<emphasis id="6593EAF4FFE8DE7BC71D702CFB88FD88" bold="true" box="[1131,1151,583,608]" pageId="13" pageNumber="304">V</emphasis>
|
||
and
|
||
<emphasis id="6593EAF4FFE8DE7BC7CD702CFB31FD88" bold="true" box="[1211,1222,583,608]" pageId="13" pageNumber="304">I</emphasis>
|
||
1.25/2
|
||
<date id="23591026FFE8DE7BC67A702CFADBFD88" box="[1292,1324,583,608]" pageId="13" pageNumber="304">
|
||
<emphasis id="6593EAF4FFE8DE7BC67A702CFAD5FD88" bold="true" box="[1292,1314,583,608]" pageId="13" pageNumber="304">II</emphasis>
|
||
1
|
||
</date>
|
||
+/2
|
||
<date id="23591026FFE8DE7BC623702CFA77FD88" box="[1365,1408,583,608]" pageId="13" pageNumber="304">
|
||
<emphasis id="6593EAF4FFE8DE7BC623702CFA81FD88" bold="true" box="[1365,1398,583,608]" pageId="13" pageNumber="304">III</emphasis>
|
||
1
|
||
</date>
|
||
+/ 2
|
||
<date id="23591026FFE8DE7BC3D37007FF24FD6C" box="[165,211,620,645]" pageId="13" pageNumber="304">
|
||
<emphasis id="6593EAF4FFE8DE7BC3D37007FF33FD6D" bold="true" box="[165,196,620,645]" pageId="13" pageNumber="304">IV</emphasis>
|
||
2
|
||
</date>
|
||
/1+
|
||
<emphasis id="6593EAF4FFE8DE7BC3817007FEFFFD6D" bold="true" box="[247,264,620,645]" pageId="13" pageNumber="304">V</emphasis>
|
||
, respeCtively, vs.
|
||
<date id="23591026FFE8DE7BC2AE7007FE1AFD6D" box="[472,493,620,645]" pageId="13" pageNumber="304">
|
||
<emphasis id="6593EAF4FFE8DE7BC2AE7007FE14FD6D" bold="true" box="[472,483,620,645]" pageId="13" pageNumber="304">I</emphasis>
|
||
1
|
||
</date>
|
||
/1.5
|
||
<date id="23591026FFE8DE7BC16A7007FDCCFD6D" box="[540,571,620,645]" pageId="13" pageNumber="304">
|
||
<emphasis id="6593EAF4FFE8DE7BC16A7007FDC5FD6D" bold="true" box="[540,562,620,645]" pageId="13" pageNumber="304">II</emphasis>
|
||
1
|
||
</date>
|
||
/1.5
|
||
<date id="23591026FFE8DE7BC11C7007FD62FD6C" box="[618,661,620,645]" pageId="13" pageNumber="304">
|
||
<emphasis id="6593EAF4FFE8DE7BC11C7007FD7CFD6D" bold="true" box="[618,651,620,645]" pageId="13" pageNumber="304">III</emphasis>
|
||
1
|
||
</date>
|
||
/2
|
||
<date id="23591026FFE8DE7BC1D87007FD2BFD6C" box="[686,732,620,645]" pageId="13" pageNumber="304">
|
||
<emphasis id="6593EAF4FFE8DE7BC1D87007FD3AFD6D" bold="true" box="[686,717,620,645]" pageId="13" pageNumber="304">IV</emphasis>
|
||
2
|
||
</date>
|
||
/1+
|
||
<emphasis id="6593EAF4FFE8DE7BC0777007FCE2FD6D" bold="true" box="[769,789,620,645]" pageId="13" pageNumber="304">V</emphasis>
|
||
in
|
||
<taxonomicName id="90E74D65FFE8DE7BC0487006FC63FD6C" authorityName="Dehling" authorityYear="2017" box="[830,916,620,644]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" kingdom="Animalia" order="Anura" pageId="13" pageNumber="304" phylum="Chordata" rank="species" species="kong">
|
||
<emphasis id="6593EAF4FFE8DE7BC0487006FC63FD6C" box="[830,916,620,644]" italics="true" pageId="13" pageNumber="304">L. kong</emphasis>
|
||
</taxonomicName>
|
||
), are muCh smaller (SVL
|
||
<quantity id="901F9B03FFE8DE7BC7B37007FA9DFD6C" box="[1221,1386,620,645]" metricMagnitude="-2" metricUnit="m" metricValue="4.05" metricValueMax="4.68" metricValueMin="3.42" pageId="13" pageNumber="304" unit="mm" value="40.5" valueMax="46.8" valueMin="34.2">34.2–46.8 mm</quantity>
|
||
and
|
||
<quantity id="901F9B03FFE8DE7BC3E170E4FF09FD4F" box="[151,254,655,680]" metricMagnitude="-2" metricUnit="m" metricValue="4.65" pageId="13" pageNumber="304" unit="mm" value="46.5">46.5 mm</quantity>
|
||
, respeCtively, in males;
|
||
<quantity id="901F9B03FFE8DE7BC17B70E4FD46FD4F" box="[525,689,655,680]" metricMagnitude="-2" metricUnit="m" metricValue="4.145" metricValueMax="4.61" metricValueMin="3.6799999999999997" pageId="13" pageNumber="304" unit="mm" value="41.45" valueMax="46.1" valueMin="36.8">36.8–46.1 mm</quantity>
|
||
and
|
||
<quantity id="901F9B03FFE8DE7BC19F70E4FC78FD4F" box="[745,911,655,680]" metricMagnitude="-2" metricUnit="m" metricValue="3.4549999999999996" metricValueMax="3.65" metricValueMin="3.26" pageId="13" pageNumber="304" unit="mm" value="34.55" valueMax="36.5" valueMin="32.6">32.6–36.5 mm</quantity>
|
||
, respeCtively, in females, vs.
|
||
<quantity id="901F9B03FFE8DE7BC7AD70E4FA88FD4F" box="[1243,1407,655,680]" metricMagnitude="-2" metricUnit="m" metricValue="6.7" metricValueMax="7.63" metricValueMin="5.77" pageId="13" pageNumber="304" unit="mm" value="67.0" valueMax="76.3" valueMin="57.7">57.7–76.3 mm</quantity>
|
||
in males and
|
||
<quantity id="901F9B03FFE8DE7BC26670DFFE43FD24" box="[272,436,692,717]" metricMagnitude="-2" metricUnit="m" metricValue="5.595" metricValueMax="5.9399999999999995" metricValueMin="5.25" pageId="13" pageNumber="304" unit="mm" value="55.95" valueMax="59.4" valueMin="52.5">52.5–59.4 mm</quantity>
|
||
in females of
|
||
<taxonomicName id="90E74D65FFE8DE7BC12170DEFD5CFD24" authorityName="Dehling" authorityYear="2017" box="[599,683,692,716]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" kingdom="Animalia" order="Anura" pageId="13" pageNumber="304" phylum="Chordata" rank="species" species="kong">
|
||
<emphasis id="6593EAF4FFE8DE7BC12170DEFD5CFD24" box="[599,683,692,716]" italics="true" pageId="13" pageNumber="304">L. kong</emphasis>
|
||
</taxonomicName>
|
||
), and the odontoid proCesses of their males have a pointed tip (vs. blunt, rounded odontoid tip in males).
|
||
</paragraph>
|
||
<caption id="0398666EFFE8DE7BC3E1749FFC7FF8A0" httpUri="https://zenodo.org/record/884337/files/figure.png" pageId="13" pageNumber="304" targetBox="[151,1436,805,1769]" targetPageId="13">
|
||
<paragraph id="575836E6FFE8DE7BC3E1749FFC7FF8A0" blockId="13.[151,1436,1780,1864]" pageId="13" pageNumber="304">
|
||
<emphasis id="6593EAF4FFE8DE7BC3E1749FFEE4F8E1" bold="true" box="[151,275,1780,1802]" pageId="13" pageNumber="304">FIGURE 9.</emphasis>
|
||
<taxonomicName id="90E74D65FFE8DE7BC26D749EFE24F8E2" authority="Dehling, 2017" authorityName="Dehling" authorityYear="2017" box="[283,467,1780,1802]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" kingdom="Animalia" order="Anura" pageId="13" pageNumber="304" phylum="Chordata" rank="species" species="kong" status="sp. nov.">
|
||
<emphasis id="6593EAF4FFE8DE7BC26D749EFE24F8E2" box="[283,467,1780,1802]" italics="true" pageId="13" pageNumber="304">Limnonectes kong</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="6593EAF4FFE8DE7BC2AC749EFDDCF8E2" bold="true" box="[474,555,1781,1802]" pageId="13" pageNumber="304">
|
||
<taxonomicNameLabel id="7EA0578FFFE8DE7BC2AC749EFDDCF8E2" box="[474,555,1781,1802]" pageId="13" pageNumber="304" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
: (A) Female paratype (NMBE 1059918) calling From a cavity in the ground, Matang, Sarawak; (B) Female paratype (NMBE 1059928) perched on a rock along a stream, Gading NP, Sarawak; (C) lateral view and (D) dorsal view oF a male paratype (NMBE 1059924), Bako NP, Sarawak.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="575836E6FFE8DE78C3B17504FC01FE60" blockId="13.[151,1437,1903,2036]" lastBlockId="14.[151,1437,151,1040]" lastPageId="14" lastPageNumber="305" pageId="13" pageNumber="304">
|
||
<emphasis id="6593EAF4FFE8DE7BC3B17504FE0AF860" bold="true" box="[199,509,1903,1928]" pageId="13" pageNumber="304">Molecular differentiation.</emphasis>
|
||
The sequenCes we obtained from the
|
||
<typeStatus id="885C8844FFE8DE7BC0DF751BFC2EF860" box="[937,985,1904,1928]" pageId="13" pageNumber="304">type</typeStatus>
|
||
speCimens showed no or little variation between eaCh other (0.0–0.6% pairwise divergenCe). Comparison of the sequenCes with those deposited in GenBank Corroborated that
|
||
<taxonomicName id="90E74D65FFE8DE7BC2A375D3FDDCF827" authorityName="Dehling" authorityYear="2017" box="[469,555,1975,1999]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" kingdom="Animalia" order="Anura" pageId="13" pageNumber="304" phylum="Chordata" rank="species" species="kong">
|
||
<emphasis id="6593EAF4FFE8DE7BC2A375D3FDDCF827" box="[469,555,1975,1999]" italics="true" pageId="13" pageNumber="304">L. kong</emphasis>
|
||
</taxonomicName>
|
||
is neither ConspeCifiC with
|
||
<taxonomicName id="90E74D65FFE8DE7BC01C75D3FC3EF827" baseAuthorityName="TsChudi" baseAuthorityYear="1838" box="[874,969,1975,1999]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" kingdom="Animalia" order="Anura" pageId="13" pageNumber="304" phylum="Chordata" rank="species" species="kuhlii">
|
||
<emphasis id="6593EAF4FFE8DE7BC01C75D3FC3EF827" box="[874,969,1975,1999]" italics="true" pageId="13" pageNumber="304">L. kuhlii</emphasis>
|
||
</taxonomicName>
|
||
from
|
||
<collectingRegion id="9523F804FFE8DE7BC76775D3FBB2F827" box="[1041,1093,1976,1999]" country="Indonesia" name="Jawa Timur" pageId="13" pageNumber="304">Java</collectingRegion>
|
||
, nor with any other desCribed speCies from the Sunda region for whiCh sequenCes are available (
|
||
<tableCitation id="1A65035DFFE8DE7BC0F175B0FC29F81B" box="[903,990,2011,2036]" captionStart="TABLE 3" captionStartId="14.[151,239,1080,1103]" captionText="TABLE 3. Uncorrected p distance between Limnonectes kong (holotype, NMBE 1059923, Kubah National Park, Malaysia, GenBank accession number MF 322505) and species of the Limnonectes kuhlii group from the Sunda region, based on 583 base pairs of mitochondrial 16 S ribosomal RNA. Quotation marks around the species names indicate that the specimen is wrongly assigned to that nominal species." pageId="13" pageNumber="304">Table 3</tableCitation>
|
||
). The interspeCifiC divergenCe between desCribed speCies of the
|
||
<taxonomicName id="90E74D65FFEBDE78C2C772F2FDE7FF58" baseAuthorityName="TsChudi" baseAuthorityYear="1838" box="[433,528,152,176]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" kingdom="Animalia" order="Anura" pageId="14" pageNumber="305" phylum="Chordata" rank="species" species="kuhlii">
|
||
<emphasis id="6593EAF4FFEBDE78C2C772F2FDE7FF58" box="[433,528,152,176]" italics="true" pageId="14" pageNumber="305">L. kuhlii</emphasis>
|
||
</taxonomicName>
|
||
group, for whiCh moleCular data are available, is 6.5% or higher (Matsui et al. 2010, 2014a, 2014b). The sequenCe of
|
||
<taxonomicName id="90E74D65FFEBDE78C13C72D6FD69FF3C" authorityName="Dehling" authorityYear="2017" box="[586,670,188,212]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" kingdom="Animalia" order="Anura" pageId="14" pageNumber="305" phylum="Chordata" rank="species" species="kong">
|
||
<emphasis id="6593EAF4FFEBDE78C13C72D6FD69FF3C" box="[586,670,188,212]" italics="true" pageId="14" pageNumber="305">L. kong</emphasis>
|
||
</taxonomicName>
|
||
was most similar to those of the sympatriC
|
||
<taxonomicName id="90E74D65FFEBDE78C7F272D6FB18FF3C" box="[1156,1263,188,212]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" kingdom="Animalia" order="Anura" pageId="14" pageNumber="305" phylum="Chordata" rank="species" species="hikidai">
|
||
<emphasis id="6593EAF4FFEBDE78C7F272D6FB18FF3C" box="[1156,1263,188,212]" italics="true" pageId="14" pageNumber="305">L. hikidai</emphasis>
|
||
</taxonomicName>
|
||
(7.7%) and “
|
||
<emphasis id="6593EAF4FFEBDE78C6F372D6FEE3FF1F" italics="true" pageId="14" pageNumber="305">
|
||
<taxonomicName id="90E74D65FFEBDE78C6F372D6FEFBFF1F" authorityName="Inger, Boeadi & Taufik" authorityYear="1996" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" kingdom="Animalia" order="Anura" pageId="14" pageNumber="305" phylum="Chordata" rank="species" species="asperatus">L. asperatus</taxonomicName>
|
||
”
|
||
</emphasis>
|
||
(6.1%) from Bukit Raya National Park in
|
||
<collectingRegion id="9523F804FFEBDE78C18072B4FC8BFF10" box="[758,892,223,248]" country="Indonesia" name="Kalimantan Utara" pageId="14" pageNumber="305">Kalimantan</collectingRegion>
|
||
. The identities of the speCimens from whiCh the sequenCes labelled “
|
||
<taxonomicName id="90E74D65FFEBDE78C20B736EFDFEFEF4" authorityName="Inger, Boeadi & Taufik" authorityYear="1996" box="[381,521,261,284]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" kingdom="Animalia" order="Anura" pageId="14" pageNumber="305" phylum="Chordata" rank="species" species="asperatus">
|
||
<emphasis id="6593EAF4FFEBDE78C20B736EFDFEFEF4" box="[381,521,261,284]" italics="true" pageId="14" pageNumber="305">L. asperatus</emphasis>
|
||
</taxonomicName>
|
||
” stem need to be reassessed, as they show different levels of divergenCe from
|
||
<taxonomicName id="90E74D65FFEBDE78C6F3736EFF3AFEA8" authorityName="Dehling" authorityYear="2017" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" kingdom="Animalia" order="Anura" pageId="14" pageNumber="305" phylum="Chordata" rank="species" species="kong">
|
||
<emphasis id="6593EAF4FFEBDE78C6F3736EFF3AFEA8" italics="true" pageId="14" pageNumber="305">L. kong</emphasis>
|
||
</taxonomicName>
|
||
and from eaCh other. Some of them probably stem from misidentified speCimens. A sequenCe from Serasan Island labelled “
|
||
<taxonomicName id="90E74D65FFEBDE78C2397326FE2DFE8C" authorityName="Inger, Boeadi & Taufik" authorityYear="1996" box="[335,474,333,356]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" kingdom="Animalia" order="Anura" pageId="14" pageNumber="305" phylum="Chordata" rank="species" species="asperatus">
|
||
<emphasis id="6593EAF4FFEBDE78C2397326FE2DFE8C" box="[335,474,333,356]" italics="true" pageId="14" pageNumber="305">L. asperatus</emphasis>
|
||
</taxonomicName>
|
||
” shows only 1.8% divergenCe from
|
||
<taxonomicName id="90E74D65FFEBDE78C0027326FC3FFE8C" authorityName="Dehling" authorityYear="2017" box="[884,968,332,356]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" kingdom="Animalia" order="Anura" pageId="14" pageNumber="305" phylum="Chordata" rank="species" species="kong">
|
||
<emphasis id="6593EAF4FFEBDE78C0027326FC3FFE8C" box="[884,968,332,356]" italics="true" pageId="14" pageNumber="305">L. kong</emphasis>
|
||
</taxonomicName>
|
||
. This speCimen Could in faCt belong to
|
||
<emphasis id="6593EAF4FFEBDE78C6F37326FF23FE60" italics="true" pageId="14" pageNumber="305">
|
||
<taxonomicName id="90E74D65FFEBDE78C6F37326FF27FE60" authorityName="Dehling" authorityYear="2017" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" kingdom="Animalia" order="Anura" pageId="14" pageNumber="305" phylum="Chordata" rank="species" species="kong">L. kong</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
but this should be Confirmed through a morphologiCal re-examination..
|
||
</paragraph>
|
||
<paragraph id="575836E6FFEBDE78C3B173FFFE51FD07" blockId="14.[151,1437,151,1040]" pageId="14" pageNumber="305">
|
||
The sequenCe of
|
||
<taxonomicName id="90E74D65FFEBDE78C2FC73FEFE28FE44" authorityName="Dehling" authorityYear="2017" box="[394,479,404,428]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" kingdom="Animalia" order="Anura" pageId="14" pageNumber="305" phylum="Chordata" rank="species" species="kong">
|
||
<emphasis id="6593EAF4FFEBDE78C2FC73FEFE28FE44" box="[394,479,404,428]" italics="true" pageId="14" pageNumber="305">L. kong</emphasis>
|
||
</taxonomicName>
|
||
is identiCal with one of the 17 Bornean lineages reCently identified by
|
||
<bibRefCitation id="33764B17FFEBDE78C67A73FFFF13FE38" pageId="14" pageNumber="305" refString="Matsui, M., Kuraishi, N., Eto, K., Hamidy, A., Nishikawa, K., Shimada, T., Yambun, P., Vairappan, C. S. & Hossman, M. Y. B. (2016) Unusually high genetic diversity in the Bornean Limnonectes kuhlii - like Fanged Frogs (Anura: Dicroglossidae). Molecular Phylogenetics and Evolution, 102, 305 - 319. https: // doi. org / 10.1016 / j. ympev. 2016.06.009" type="journal article">
|
||
Matsui
|
||
<emphasis id="6593EAF4FFEBDE78C61473FEFA63FE44" box="[1378,1428,404,428]" italics="true" pageId="14" pageNumber="305">et al</emphasis>
|
||
. (2016)
|
||
</bibRefCitation>
|
||
, i.e. “lineage 8”. None of these 17 lineages has so far been attributed to either
|
||
<taxonomicName id="90E74D65FFEBDE78C7FB73D2FABCFE38" box="[1165,1355,440,464]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" kingdom="Animalia" order="Anura" pageId="14" pageNumber="305" phylum="Chordata" rank="species" species="conspicillatus">
|
||
<emphasis id="6593EAF4FFEBDE78C7FB73D2FABCFE38" box="[1165,1355,440,464]" italics="true" pageId="14" pageNumber="305">L. conspicillatus</emphasis>
|
||
</taxonomicName>
|
||
nor
|
||
<taxonomicName id="90E74D65FFEBDE78C6F373D2FEF8FE1C" authorityName="Matsui, Dubois & Ohler" authorityYear="2013" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="14" pageNumber="305" phylum="Chordata" rank="species" species="mocquardi">
|
||
<emphasis id="6593EAF4FFEBDE78C6F373D2FEF8FE1C" italics="true" pageId="14" pageNumber="305">L. mocquardi</emphasis>
|
||
</taxonomicName>
|
||
. However, “lineage 6” of
|
||
<bibRefCitation id="33764B17FFEBDE78C12473B7FCB6FE1D" box="[594,833,476,501]" pageId="14" pageNumber="305" refString="Matsui, M., Kuraishi, N., Eto, K., Hamidy, A., Nishikawa, K., Shimada, T., Yambun, P., Vairappan, C. S. & Hossman, M. Y. B. (2016) Unusually high genetic diversity in the Bornean Limnonectes kuhlii - like Fanged Frogs (Anura: Dicroglossidae). Molecular Phylogenetics and Evolution, 102, 305 - 319. https: // doi. org / 10.1016 / j. ympev. 2016.06.009" type="journal article">
|
||
Matsui
|
||
<emphasis id="6593EAF4FFEBDE78C1DA73B6FD1DFE1C" box="[684,746,476,500]" italics="true" pageId="14" pageNumber="305">et al.</emphasis>
|
||
(2016)
|
||
</bibRefCitation>
|
||
from Matang, whiCh is identiCal with “’
|
||
<emphasis id="6593EAF4FFEBDE78C65B73B6FA6BFE1C" box="[1325,1436,476,500]" italics="true" pageId="14" pageNumber="305">L. 'kuhlii'</emphasis>
|
||
Matang” of Matsui
|
||
<emphasis id="6593EAF4FFEBDE78C205706AFE54FDF0" box="[371,419,512,536]" italics="true" pageId="14" pageNumber="305">et al</emphasis>
|
||
. (2010, 2014) and Matsui & Nishikawa (2014) Could be referable to
|
||
<emphasis id="6593EAF4FFEBDE78C7DC706AFA9BFDF0" box="[1194,1388,512,536]" italics="true" pageId="14" pageNumber="305">
|
||
<taxonomicName id="90E74D65FFEBDE78C7DC706AFA92FDF0" box="[1194,1381,512,536]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" kingdom="Animalia" order="Anura" pageId="14" pageNumber="305" phylum="Chordata" rank="species" species="conspicillatus">L. conspicillatus</taxonomicName>
|
||
;
|
||
</emphasis>
|
||
and “’
|
||
<emphasis id="6593EAF4FFEBDE78C3DA704EFEE0FDD4" box="[172,279,548,572]" italics="true" pageId="14" pageNumber="305">L. 'kuhlii'</emphasis>
|
||
Kinabalu” of Matsui
|
||
<emphasis id="6593EAF4FFEBDE78C17A704EFDCBFDD4" box="[524,572,548,572]" italics="true" pageId="14" pageNumber="305">et al</emphasis>
|
||
. (2010) and Matsui & Nishikawa (2014) Could be referable to
|
||
<emphasis id="6593EAF4FFEBDE78C789704EFA6AFDD4" box="[1279,1437,548,572]" italics="true" pageId="14" pageNumber="305">
|
||
<taxonomicName id="90E74D65FFEBDE78C789704EFA6FFDD4" authorityName="Matsui, Dubois & Ohler" authorityYear="2013" box="[1279,1432,548,572]" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="14" pageNumber="305" phylum="Chordata" rank="species" species="mocquardi">L. mocquardi</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
This needs to be Confirmed by morphologiCal examinations. SequenCes of these lineages differ by 7.1–7.7% and 9.3%, respeCtively from sequenCes of
|
||
<taxonomicName id="90E74D65FFEBDE78C13F7006FD69FD6C" authorityName="Dehling" authorityYear="2017" box="[585,670,620,644]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" kingdom="Animalia" order="Anura" pageId="14" pageNumber="305" phylum="Chordata" rank="species" species="kong">
|
||
<emphasis id="6593EAF4FFEBDE78C13F7006FD69FD6C" box="[585,670,620,644]" italics="true" pageId="14" pageNumber="305">L. kong</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<tableCitation id="1A65035DFFEBDE78C1D87007FCF1FD6C" box="[686,774,620,645]" captionStart="TABLE 3" captionStartId="14.[151,239,1080,1103]" captionText="TABLE 3. Uncorrected p distance between Limnonectes kong (holotype, NMBE 1059923, Kubah National Park, Malaysia, GenBank accession number MF 322505) and species of the Limnonectes kuhlii group from the Sunda region, based on 583 base pairs of mitochondrial 16 S ribosomal RNA. Quotation marks around the species names indicate that the specimen is wrongly assigned to that nominal species." pageId="14" pageNumber="305">Table 3</tableCitation>
|
||
).
|
||
<bibRefCitation id="33764B17FFEBDE78C06B7007FBF7FD6D" box="[797,1024,620,645]" pageId="14" pageNumber="305" refString="Matsui, M., Kuraishi, N., Eto, K., Hamidy, A., Nishikawa, K., Shimada, T., Yambun, P., Vairappan, C. S. & Hossman, M. Y. B. (2016) Unusually high genetic diversity in the Bornean Limnonectes kuhlii - like Fanged Frogs (Anura: Dicroglossidae). Molecular Phylogenetics and Evolution, 102, 305 - 319. https: // doi. org / 10.1016 / j. ympev. 2016.06.009" type="journal article">
|
||
Matsui
|
||
<emphasis id="6593EAF4FFEBDE78C0057006FC51FD6C" box="[883,934,620,644]" italics="true" pageId="14" pageNumber="305">et al</emphasis>
|
||
. (2016)
|
||
</bibRefCitation>
|
||
provided a phylogenetiC analysis of different lineages that are present on Borneo and their relationship to speCies from outside Borneo. SinCe
|
||
<taxonomicName id="90E74D65FFEBDE78C63170FAFA6CFD40" authorityName="Dehling" authorityYear="2017" box="[1351,1435,656,680]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" kingdom="Animalia" order="Anura" pageId="14" pageNumber="305" phylum="Chordata" rank="species" species="kong">
|
||
<emphasis id="6593EAF4FFEBDE78C63170FAFA6CFD40" box="[1351,1435,656,680]" italics="true" pageId="14" pageNumber="305">L. kong</emphasis>
|
||
</taxonomicName>
|
||
does not represent an additional lineage, we refer to this resourCe for an analysis and depiCtion of the phylogenetiC relationships of
|
||
<taxonomicName id="90E74D65FFEBDE78C23D70B2FE68FD18" authorityName="Dehling" authorityYear="2017" box="[331,415,728,752]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" kingdom="Animalia" order="Anura" pageId="14" pageNumber="305" phylum="Chordata" rank="species" species="kong">
|
||
<emphasis id="6593EAF4FFEBDE78C23D70B2FE68FD18" box="[331,415,728,752]" italics="true" pageId="14" pageNumber="305">L. kong</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="1FFD656DFFEBDE66C3B17097FE90FDFF" lastPageId="16" lastPageNumber="307" pageId="14" pageNumber="305" type="distribution">
|
||
<paragraph id="575836E6FFEBDE78C3B17097FD4FFBF8" blockId="14.[151,1437,151,1040]" pageId="14" pageNumber="305">
|
||
<emphasis id="6593EAF4FFEBDE78C3B17097FE97FCFD" bold="true" box="[199,352,764,789]" pageId="14" pageNumber="305">Distribution.</emphasis>
|
||
The
|
||
<typeStatus id="885C8844FFEBDE78C2ED7096FE3CFCFD" box="[411,459,765,789]" pageId="14" pageNumber="305">type</typeStatus>
|
||
speCimens were ColleCted in Kubah, Gading, and Bako National Parks, and speCimens referred to
|
||
<taxonomicName id="90E74D65FFEBDE78C264714AFE92FCD0" authorityName="Dehling" authorityYear="2017" box="[274,357,800,824]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" kingdom="Animalia" order="Anura" pageId="14" pageNumber="305" phylum="Chordata" rank="species" species="kong">
|
||
<emphasis id="6593EAF4FFEBDE78C264714AFE92FCD0" box="[274,357,800,824]" italics="true" pageId="14" pageNumber="305">L. kong</emphasis>
|
||
</taxonomicName>
|
||
based on morphologiCal CharaCters were ColleCted in Santubong National Park. All loCalities are in western
|
||
<collectingCountry id="2FF07676FFEBDE78C26E712FFE8FFCB5" box="[280,376,836,861]" name="Malaysia" pageId="14" pageNumber="305">
|
||
<collectingRegion id="9523F804FFEBDE78C26E712FFE8FFCB5" box="[280,376,836,861]" country="Malaysia" name="Sarawak" pageId="14" pageNumber="305">Sarawak</collectingRegion>
|
||
</collectingCountry>
|
||
(
|
||
<figureCitation id="CFDC2A63FFEBDE78C2FD712FFE15FCB4" box="[395,482,836,861]" captionStart="FIGURE 10" captionStartId="15.[151,250,1084,1106]" captionTargetBox="[157,1433,209,1053]" captionTargetId="figure@15.[151,1436,193,1062]" captionTargetPageId="15" captionText="FIGURE 10. Collection localities oF Limnonectes kong sp. nov. in western Sarawak, Borneo at (1) Gunung Gading, (2) Kubah National Park, Matang, (3), Santubong National Park, and (4) Bako National Park." httpUri="https://zenodo.org/record/884339/files/figure.png" pageId="14" pageNumber="305">Fig. 10</figureCitation>
|
||
).
|
||
<bibRefCitation id="33764B17FFEBDE78C28C712FFD15FCB5" box="[506,738,836,861]" pageId="14" pageNumber="305" refString="Matsui, M., Kuraishi, N., Eto, K., Hamidy, A., Nishikawa, K., Shimada, T., Yambun, P., Vairappan, C. S. & Hossman, M. Y. B. (2016) Unusually high genetic diversity in the Bornean Limnonectes kuhlii - like Fanged Frogs (Anura: Dicroglossidae). Molecular Phylogenetics and Evolution, 102, 305 - 319. https: // doi. org / 10.1016 / j. ympev. 2016.06.009" type="journal article">
|
||
Matsui
|
||
<emphasis id="6593EAF4FFEBDE78C124712EFD71FCB4" box="[594,646,836,860]" italics="true" pageId="14" pageNumber="305">et al</emphasis>
|
||
. (2016)
|
||
</bibRefCitation>
|
||
listed sequenCes obtained from speCimens from Samunsam, Santu Bong, and Lanjak Entimau in western
|
||
<collectingCountry id="2FF07676FFEBDE78C1FA710CFD18FC68" box="[652,751,871,896]" name="Malaysia" pageId="14" pageNumber="305">
|
||
<collectingRegion id="9523F804FFEBDE78C1FA710CFD18FC68" box="[652,751,871,896]" country="Malaysia" name="Sarawak" pageId="14" pageNumber="305">Sarawak</collectingRegion>
|
||
</collectingCountry>
|
||
, whiCh show little divergenCe (<1.0%) from the sequenCes of
|
||
<taxonomicName id="90E74D65FFEBDE78C3E171E6FF1CFC4C" authorityName="Dehling" authorityYear="2017" box="[151,235,908,932]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" kingdom="Animalia" order="Anura" pageId="14" pageNumber="305" phylum="Chordata" rank="species" species="kong">
|
||
<emphasis id="6593EAF4FFEBDE78C3E171E6FF1CFC4C" box="[151,235,908,932]" italics="true" pageId="14" pageNumber="305">L. kong</emphasis>
|
||
</taxonomicName>
|
||
and are thus very likely ConspeCifiC. A sequenCe from Serasan National Park might indiCate that the speCies also oCCurs on the Natuna Islands,
|
||
<collectingCountry id="2FF07676FFEBDE78C15E71C4FD6EFC20" box="[552,665,943,968]" name="Indonesia" pageId="14" pageNumber="305">Indonesia</collectingCountry>
|
||
. This island is loCated about
|
||
<quantity id="901F9B03FFEBDE78C09E71DBFBC8FC20" box="[1000,1087,943,968]" metricMagnitude="5" metricUnit="m" metricValue="1.2" pageId="14" pageNumber="305" unit="km" value="120.0">120 km</quantity>
|
||
and
|
||
<quantity id="901F9B03FFEBDE78C70F71C4FB27FC20" box="[1145,1232,943,968]" metricMagnitude="5" metricUnit="m" metricValue="1.6" pageId="14" pageNumber="305" unit="km" value="160.0">160 km</quantity>
|
||
from Gading and Kubah National Parks, respeCtively, and less than
|
||
<quantity id="901F9B03FFEBDE78C19271BFFCD8FC05" box="[740,815,980,1005]" metricMagnitude="4" metricUnit="m" metricValue="7.0" pageId="14" pageNumber="305" unit="km" value="70.0">70 km</quantity>
|
||
off the Bornean Coast and was ConneCted to Borneo throughout most of the PleistoCene (
|
||
<bibRefCitation id="33764B17FFEBDE78C15A719CFD5BFBF8" author="Voris" box="[556,684,1015,1040]" pageId="14" pageNumber="305" refString="Voris, H. K. (2000) Maps oF Pleistocene sea levels in Southeast Asia: shorelines, river systems and time durations. Journal of Biogeography, 27, 1153 - 1167. https: // doi. org / 10.1046 / j. 1365 - 2699.2000.00489. x" type="journal article" year="2000">Voris 2000</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="0398666EFFEBDE78C3E17653FCF7FB47" pageId="14" pageNumber="305">
|
||
<paragraph id="575836E6FFEBDE78C3E17653FCF7FB47" blockId="14.[151,1436,1079,1199]" pageId="14" pageNumber="305">
|
||
<emphasis id="6593EAF4FFEBDE78C3E17653FEF8FBA7" bold="true" box="[151,271,1080,1103]" pageId="14" pageNumber="305">TABLE 3.</emphasis>
|
||
Uncorrected p distance between
|
||
<taxonomicName id="90E74D65FFEBDE78C1FD7653FCA6FBA7" authorityName="Dehling" authorityYear="2017" box="[651,849,1079,1103]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" kingdom="Animalia" order="Anura" pageId="14" pageNumber="305" phylum="Chordata" rank="species" species="kong">
|
||
<emphasis id="6593EAF4FFEBDE78C1FD7653FCA6FBA7" box="[651,849,1079,1103]" italics="true" pageId="14" pageNumber="305">Limnonectes kong</emphasis>
|
||
</taxonomicName>
|
||
(holotype, NMBE 1059923, Kubah National Park, Malaysia, GenBank accession number
|
||
<accessionNumber id="48B4AB05FFEBDE78C14C7633FD44FB87" box="[570,691,1112,1135]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MF322505" pageId="14" pageNumber="305">MF322505</accessionNumber>
|
||
) and species of the
|
||
<taxonomicName id="90E74D65FFEBDE78C0FC7633FBA1FB87" baseAuthorityName="TsChudi" baseAuthorityYear="1838" box="[906,1110,1111,1135]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" kingdom="Animalia" order="Anura" pageId="14" pageNumber="305" phylum="Chordata" rank="species" species="kuhlii">
|
||
<emphasis id="6593EAF4FFEBDE78C0FC7633FBA1FB87" box="[906,1110,1111,1135]" italics="true" pageId="14" pageNumber="305">Limnonectes kuhlii</emphasis>
|
||
</taxonomicName>
|
||
group from the Sunda region, based on 583 base pairs of mitochondrial 16S ribosomal RNA. Quotation marks around the species names indicate that the specimen is wrongly assigned to that nominal species.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="575836E6FFEBDE78C3E976A1FABFF912" pageId="14" pageNumber="305">
|
||
<table id="25E7C446FFEB2189C3E976A1FA82F912" box="[159,1397,1226,1786]" gridcols="6" gridrows="14" pageId="14" pageNumber="305">
|
||
<tr id="E9D734A4FFEB2189C3E976A1FA82FB08" box="[159,1397,1226,1248]" gridrow="0" pageId="14" pageNumber="305">
|
||
<th id="AA065DD8FFEB2189C3E976A1FEC2FB08" box="[159,309,1226,1248]" gridcol="0" gridrow="0" pageId="14" pageNumber="305">species</th>
|
||
<th id="AA065DD8FFEB2189C20576A1FD67FB08" box="[371,656,1226,1248]" gridcol="1" gridrow="0" pageId="14" pageNumber="305">locality</th>
|
||
<th id="AA065DD8FFEB2189C18A76A1FCA9FB08" box="[764,862,1226,1248]" gridcol="2" gridrow="0" pageId="14" pageNumber="305">country</th>
|
||
<th id="AA065DD8FFEB2189C0EA76A1FBCFFB08" box="[924,1080,1226,1248]" gridcol="3" gridrow="0" pageId="14" pageNumber="305">voucher no.</th>
|
||
<th id="AA065DD8FFEB2189C71E76A1FB29FB08" box="[1128,1246,1226,1248]" gridcol="4" gridrow="0" pageId="14" pageNumber="305">GenBank #</th>
|
||
<th id="AA065DD8FFEB2189C67976A1FA82FB08" box="[1295,1397,1226,1248]" gridcol="5" gridrow="0" pageId="14" pageNumber="305">p distance</th>
|
||
</tr>
|
||
<tr id="E9D734A4FFEB2189C3E9769FFA82FAE2" box="[159,1397,1268,1290]" gridrow="1" pageId="14" pageNumber="305">
|
||
<th id="AA065DD8FFEB2189C3E9769FFEC2FAE2" box="[159,309,1268,1290]" gridcol="0" gridrow="1" pageId="14" pageNumber="305">
|
||
"
|
||
<taxonomicName id="90E74D65FFEBDE78C3DF769EFF09FAE2" baseAuthorityName="TsChudi" baseAuthorityYear="1838" box="[169,254,1268,1290]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" kingdom="Animalia" order="Anura" pageId="14" pageNumber="305" phylum="Chordata" rank="species" species="kuhlii">
|
||
<emphasis id="6593EAF4FFEBDE78C3DF769EFF09FAE2" box="[169,254,1268,1290]" italics="true" pageId="14" pageNumber="305">L. kuhlii</emphasis>
|
||
</taxonomicName>
|
||
“
|
||
</th>
|
||
<td id="AA065DD8FFEB2189C205769FFD67FAE2" box="[371,656,1268,1290]" gridcol="1" gridrow="1" pageId="14" pageNumber="305">Matang, Sarawak</td>
|
||
<td id="AA065DD8FFEB2189C18A769FFCA9FAE2" box="[764,862,1268,1290]" gridcol="2" gridrow="1" pageId="14" pageNumber="305">Malaysia</td>
|
||
<td id="AA065DD8FFEB2189C0EA769FFBCFFAE2" box="[924,1080,1268,1290]" gridcol="3" gridrow="1" pageId="14" pageNumber="305">KUHE 12025</td>
|
||
<td id="AA065DD8FFEB2189C71E769FFB29FAE2" box="[1128,1246,1268,1290]" gridcol="4" gridrow="1" pageId="14" pageNumber="305">
|
||
<accessionNumber id="48B4AB05FFEBDE78C71F769FFB2FFAE2" box="[1129,1240,1268,1290]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/AB526322" pageId="14" pageNumber="305">AB526322</accessionNumber>
|
||
</td>
|
||
<td id="AA065DD8FFEB2189C679769FFA82FAE2" box="[1295,1397,1268,1290]" gridcol="5" gridrow="1" pageId="14" pageNumber="305">0.071</td>
|
||
</tr>
|
||
<tr id="E9D734A4FFEB2189C3E97776FA82FADB" box="[159,1397,1309,1331]" gridrow="2" pageId="14" pageNumber="305">
|
||
<th id="AA065DD8FFEB2189C3E97776FEC2FADB" box="[159,309,1309,1331]" gridcol="0" gridrow="2" pageId="14" pageNumber="305">
|
||
<taxonomicName id="90E74D65FFEBDE78C3E97775FEF7FADB" box="[159,256,1309,1331]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" kingdom="Animalia" order="Anura" pageId="14" pageNumber="305" phylum="Chordata" rank="species" species="hikidai">
|
||
<emphasis id="6593EAF4FFEBDE78C3E97775FEF7FADB" box="[159,256,1309,1331]" italics="true" pageId="14" pageNumber="305">L. hikidai</emphasis>
|
||
</taxonomicName>
|
||
</th>
|
||
<td id="AA065DD8FFEB2189C2057776FD67FADB" box="[371,656,1309,1331]" gridcol="1" gridrow="2" pageId="14" pageNumber="305">Matang, Sarawak</td>
|
||
<td id="AA065DD8FFEB2189C18A7776FCA9FADB" box="[764,862,1309,1331]" gridcol="2" gridrow="2" pageId="14" pageNumber="305">Malaysia</td>
|
||
<td id="AA065DD8FFEB2189C0EA7776FBCFFADB" box="[924,1080,1309,1331]" gridcol="3" gridrow="2" pageId="14" pageNumber="305">KUHE 10654</td>
|
||
<td id="AA065DD8FFEB2189C71E7776FB29FADB" box="[1128,1246,1309,1331]" gridcol="4" gridrow="2" pageId="14" pageNumber="305">
|
||
<accessionNumber id="48B4AB05FFEBDE78C71F7776FB21FADB" box="[1129,1238,1309,1331]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/AB971130" pageId="14" pageNumber="305">AB971130</accessionNumber>
|
||
</td>
|
||
<td id="AA065DD8FFEB2189C6797776FA82FADB" box="[1295,1397,1309,1331]" gridcol="5" gridrow="2" pageId="14" pageNumber="305">0.077</td>
|
||
</tr>
|
||
<tr id="E9D734A4FFEB2189C3E9772DFA82FAB5" box="[159,1397,1350,1373]" gridrow="3" pageId="14" pageNumber="305">
|
||
<th id="AA065DD8FFEB2189C3E9772DFEC2FAB5" box="[159,309,1350,1373]" gridcol="0" gridrow="3" pageId="14" pageNumber="305">
|
||
<taxonomicName id="90E74D65FFEBDE78C3E97723FEC4FAB4" authorityName="Matsui, Nishikawa & Eto" authorityYear="2014" box="[159,307,1350,1373]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" kingdom="Animalia" order="Anura" pageId="14" pageNumber="305" phylum="Chordata" rank="species" species="cintalubang">
|
||
<emphasis id="6593EAF4FFEBDE78C3E97723FEC4FAB4" box="[159,307,1350,1373]" italics="true" pageId="14" pageNumber="305">L. cintalubang</emphasis>
|
||
</taxonomicName>
|
||
</th>
|
||
<td id="AA065DD8FFEB2189C205772DFD67FAB5" box="[371,656,1350,1373]" gridcol="1" gridrow="3" pageId="14" pageNumber="305">not stated</td>
|
||
<td id="AA065DD8FFEB2189C18A772DFCA9FAB5" box="[764,862,1350,1373]" gridcol="2" gridrow="3" pageId="14" pageNumber="305">Malaysia</td>
|
||
<td id="AA065DD8FFEB2189C0EA772DFBCFFAB5" box="[924,1080,1350,1373]" gridcol="3" gridrow="3" pageId="14" pageNumber="305">KUHE 47815</td>
|
||
<td id="AA065DD8FFEB2189C71E772DFB29FAB5" box="[1128,1246,1350,1373]" gridcol="4" gridrow="3" pageId="14" pageNumber="305">
|
||
<accessionNumber id="48B4AB05FFEBDE78C71F772DFB2FFAB4" box="[1129,1240,1350,1372]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/AB981409" pageId="14" pageNumber="305">AB981409</accessionNumber>
|
||
</td>
|
||
<td id="AA065DD8FFEB2189C679772DFA82FAB5" box="[1295,1397,1350,1373]" gridcol="5" gridrow="3" pageId="14" pageNumber="305">0.143</td>
|
||
</tr>
|
||
<tr id="E9D734A4FFEB2189C3E9771BFA82FA6E" box="[159,1397,1392,1414]" gridrow="4" pageId="14" pageNumber="305">
|
||
<th id="AA065DD8FFEB2189C3E9771BFEC2FA6E" box="[159,309,1392,1414]" gridcol="0" gridrow="4" pageId="14" pageNumber="305">
|
||
“
|
||
<taxonomicName id="90E74D65FFEBDE78C3DC771AFF08FA6E" baseAuthorityName="TsChudi" baseAuthorityYear="1838" box="[170,255,1392,1414]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" kingdom="Animalia" order="Anura" pageId="14" pageNumber="305" phylum="Chordata" rank="species" species="kuhlii">
|
||
<emphasis id="6593EAF4FFEBDE78C3DC771AFF08FA6E" box="[170,255,1392,1414]" italics="true" pageId="14" pageNumber="305">L. kuhlii</emphasis>
|
||
</taxonomicName>
|
||
“
|
||
</th>
|
||
<td id="AA065DD8FFEB2189C205771BFD67FA6E" box="[371,656,1392,1414]" gridcol="1" gridrow="4" pageId="14" pageNumber="305">Kinabalu, Sabah</td>
|
||
<td id="AA065DD8FFEB2189C18A771BFCA9FA6E" box="[764,862,1392,1414]" gridcol="2" gridrow="4" pageId="14" pageNumber="305">Malaysia</td>
|
||
<td id="AA065DD8FFEB2189C0EA771BFBCFFA6E" box="[924,1080,1392,1414]" gridcol="3" gridrow="4" pageId="14" pageNumber="305">BORN 22645</td>
|
||
<td id="AA065DD8FFEB2189C71E771BFB29FA6E" box="[1128,1246,1392,1414]" gridcol="4" gridrow="4" pageId="14" pageNumber="305">
|
||
<accessionNumber id="48B4AB05FFEBDE78C71F771BFB2FFA6E" box="[1129,1240,1392,1414]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/AB526323" pageId="14" pageNumber="305">AB526323</accessionNumber>
|
||
</td>
|
||
<td id="AA065DD8FFEB2189C679771BFA82FA6E" box="[1295,1397,1392,1414]" gridcol="5" gridrow="4" pageId="14" pageNumber="305">0.093</td>
|
||
</tr>
|
||
<tr id="E9D734A4FFEB2189C3E977F2FA82FA47" box="[159,1397,1433,1455]" gridrow="5" pageId="14" pageNumber="305">
|
||
<th id="AA065DD8FFEB2189C3E977F2FEC2FA47" box="[159,309,1433,1455]" gridcol="0" gridrow="5" pageId="14" pageNumber="305">
|
||
<taxonomicName id="90E74D65FFEBDE78C3E977F1FF03FA47" baseAuthorityName="TsChudi" baseAuthorityYear="1838" box="[159,244,1433,1455]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" kingdom="Animalia" order="Anura" pageId="14" pageNumber="305" phylum="Chordata" rank="species" species="kuhlii">
|
||
<emphasis id="6593EAF4FFEBDE78C3E977F1FF03FA47" box="[159,244,1433,1455]" italics="true" pageId="14" pageNumber="305">L. kuhlii</emphasis>
|
||
</taxonomicName>
|
||
</th>
|
||
<td id="AA065DD8FFEB2189C20577F2FD67FA47" box="[371,656,1433,1455]" gridcol="1" gridrow="5" pageId="14" pageNumber="305">Central Java, Purwerojo</td>
|
||
<td id="AA065DD8FFEB2189C18A77F2FCA9FA47" box="[764,862,1433,1455]" gridcol="2" gridrow="5" pageId="14" pageNumber="305">Indonesia</td>
|
||
<td id="AA065DD8FFEB2189C0EA77F2FBCFFA47" box="[924,1080,1433,1455]" gridcol="3" gridrow="5" pageId="14" pageNumber="305">GMU unnumb</td>
|
||
<td id="AA065DD8FFEB2189C71E77F2FB29FA47" box="[1128,1246,1433,1455]" gridcol="4" gridrow="5" pageId="14" pageNumber="305">
|
||
<accessionNumber id="48B4AB05FFEBDE78C71F77F2FB2FFA47" box="[1129,1240,1433,1455]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/AB526316" pageId="14" pageNumber="305">AB526316</accessionNumber>
|
||
</td>
|
||
<td id="AA065DD8FFEB2189C67977F2FA82FA47" box="[1295,1397,1433,1455]" gridcol="5" gridrow="5" pageId="14" pageNumber="305">0.098</td>
|
||
</tr>
|
||
<tr id="E9D734A4FFEB2189C3E977A9FA82FA30" box="[159,1397,1474,1496]" gridrow="6" pageId="14" pageNumber="305">
|
||
<th id="AA065DD8FFEB2189C3E977A9FEC2FA30" box="[159,309,1474,1496]" gridcol="0" gridrow="6" pageId="14" pageNumber="305">
|
||
<taxonomicName id="90E74D65FFEBDE78C3E977A8FF03FA30" baseAuthorityName="TsChudi" baseAuthorityYear="1838" box="[159,244,1474,1496]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" kingdom="Animalia" order="Anura" pageId="14" pageNumber="305" phylum="Chordata" rank="species" species="kuhlii">
|
||
<emphasis id="6593EAF4FFEBDE78C3E977A8FF03FA30" box="[159,244,1474,1496]" italics="true" pageId="14" pageNumber="305">L. kuhlii</emphasis>
|
||
</taxonomicName>
|
||
</th>
|
||
<td id="AA065DD8FFEB2189C20577A9FD67FA30" box="[371,656,1474,1496]" gridcol="1" gridrow="6" pageId="14" pageNumber="305">Java</td>
|
||
<td id="AA065DD8FFEB2189C18A77A9FCA9FA30" box="[764,862,1474,1496]" gridcol="2" gridrow="6" pageId="14" pageNumber="305">Indonesia</td>
|
||
<td id="AA065DD8FFEB2189C0EA77A9FBCFFA30" box="[924,1080,1474,1496]" gridcol="3" gridrow="6" pageId="14" pageNumber="305">not stated</td>
|
||
<td id="AA065DD8FFEB2189C71E77A9FB29FA30" box="[1128,1246,1474,1496]" gridcol="4" gridrow="6" pageId="14" pageNumber="305">
|
||
<accessionNumber id="48B4AB05FFEBDE78C71F77A9FB22FA30" box="[1129,1237,1474,1496]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/AF183138" pageId="14" pageNumber="305">AF183138</accessionNumber>
|
||
</td>
|
||
<td id="AA065DD8FFEB2189C67977A9FA82FA30" box="[1295,1397,1474,1496]" gridcol="5" gridrow="6" pageId="14" pageNumber="305">0.120</td>
|
||
</tr>
|
||
<tr id="E9D734A4FFEB2189C3E97787FA82F9EA" box="[159,1397,1516,1538]" gridrow="7" pageId="14" pageNumber="305">
|
||
<th id="AA065DD8FFEB2189C3E97787FEC2F9EA" box="[159,309,1516,1538]" gridcol="0" gridrow="7" pageId="14" pageNumber="305">
|
||
<taxonomicName id="90E74D65FFEBDE78C3E97786FEEDF9EA" box="[159,282,1516,1538]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" kingdom="Animalia" order="Anura" pageId="14" pageNumber="305" phylum="Chordata" rank="species" species="sisikdagu">
|
||
<emphasis id="6593EAF4FFEBDE78C3E97786FEEDF9EA" box="[159,282,1516,1538]" italics="true" pageId="14" pageNumber="305">L. sisikdagu</emphasis>
|
||
</taxonomicName>
|
||
</th>
|
||
<td id="AA065DD8FFEB2189C2057787FD67F9EA" box="[371,656,1516,1538]" gridcol="1" gridrow="7" pageId="14" pageNumber="305">Sumatra</td>
|
||
<td id="AA065DD8FFEB2189C18A7787FCA9F9EA" box="[764,862,1516,1538]" gridcol="2" gridrow="7" pageId="14" pageNumber="305">Indonesia</td>
|
||
<td id="AA065DD8FFEB2189C0EA7787FBCFF9EA" box="[924,1080,1516,1538]" gridcol="3" gridrow="7" pageId="14" pageNumber="305">RMBR 393</td>
|
||
<td id="AA065DD8FFEB2189C71E7787FB29F9EA" box="[1128,1246,1516,1538]" gridcol="4" gridrow="7" pageId="14" pageNumber="305">
|
||
<accessionNumber id="48B4AB05FFEBDE78C71F7787FB2AF9EA" box="[1129,1245,1516,1538]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/HM067244" pageId="14" pageNumber="305">HM067244</accessionNumber>
|
||
</td>
|
||
<td id="AA065DD8FFEB2189C6797787FA82F9EA" box="[1295,1397,1516,1538]" gridcol="5" gridrow="7" pageId="14" pageNumber="305">0.096</td>
|
||
</tr>
|
||
<tr id="E9D734A4FFEB2189C3E9747EFA82F9C3" box="[159,1397,1557,1579]" gridrow="8" pageId="14" pageNumber="305">
|
||
<th id="AA065DD8FFEB2189C3E9747EFEC2F9C3" box="[159,309,1557,1579]" gridcol="0" gridrow="8" pageId="14" pageNumber="305">
|
||
“
|
||
<taxonomicName id="90E74D65FFEBDE78C3DC747DFEE3F9C3" baseAuthorityName="Boulenger" baseAuthorityYear="1882" box="[170,276,1557,1579]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" kingdom="Animalia" order="Anura" pageId="14" pageNumber="305" phylum="Chordata" rank="species" species="laticeps">
|
||
<emphasis id="6593EAF4FFEBDE78C3DC747DFEE3F9C3" box="[170,276,1557,1579]" italics="true" pageId="14" pageNumber="305">L. laticeps</emphasis>
|
||
</taxonomicName>
|
||
“
|
||
</th>
|
||
<td id="AA065DD8FFEB2189C205747EFD67F9C3" box="[371,656,1557,1579]" gridcol="1" gridrow="8" pageId="14" pageNumber="305">not stated</td>
|
||
<td id="AA065DD8FFEB2189C18A747EFCA9F9C3" box="[764,862,1557,1579]" gridcol="2" gridrow="8" pageId="14" pageNumber="305">Malaysia</td>
|
||
<td id="AA065DD8FFEB2189C0EA747EFBCFF9C3" box="[924,1080,1557,1579]" gridcol="3" gridrow="8" pageId="14" pageNumber="305">not stated</td>
|
||
<td id="AA065DD8FFEB2189C71E747EFB29F9C3" box="[1128,1246,1557,1579]" gridcol="4" gridrow="8" pageId="14" pageNumber="305">
|
||
<accessionNumber id="48B4AB05FFEBDE78C71F747EFB2FF9C3" box="[1129,1240,1557,1579]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/AB277306" pageId="14" pageNumber="305">AB277306</accessionNumber>
|
||
</td>
|
||
<td id="AA065DD8FFEB2189C679747EFA82F9C3" box="[1295,1397,1557,1579]" gridcol="5" gridrow="8" pageId="14" pageNumber="305">0.15</td>
|
||
</tr>
|
||
<tr id="E9D734A4FFEB2189C3E97455FA82F9BD" box="[159,1397,1598,1621]" gridrow="9" pageId="14" pageNumber="305">
|
||
<th id="AA065DD8FFEB2189C3E97455FEC2F9BD" box="[159,309,1598,1621]" gridcol="0" gridrow="9" pageId="14" pageNumber="305">
|
||
“
|
||
<taxonomicName id="90E74D65FFEBDE78C3DC742BFEDDF9BC" box="[170,298,1598,1621]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" kingdom="Animalia" order="Anura" pageId="14" pageNumber="305" phylum="Chordata" rank="species" species="khasianus">
|
||
<emphasis id="6593EAF4FFEBDE78C3DC742BFEDDF9BC" box="[170,298,1598,1621]" italics="true" pageId="14" pageNumber="305">L. khasianus</emphasis>
|
||
</taxonomicName>
|
||
“
|
||
</th>
|
||
<td id="AA065DD8FFEB2189C2057455FD67F9BD" box="[371,656,1598,1621]" gridcol="1" gridrow="9" pageId="14" pageNumber="305">Bala Narathiwat</td>
|
||
<td id="AA065DD8FFEB2189C18A7455FCA9F9BD" box="[764,862,1598,1621]" gridcol="2" gridrow="9" pageId="14" pageNumber="305">Thailand</td>
|
||
<td id="AA065DD8FFEB2189C0EA7455FBCFF9BD" box="[924,1080,1598,1621]" gridcol="3" gridrow="9" pageId="14" pageNumber="305">KUHE 23158</td>
|
||
<td id="AA065DD8FFEB2189C71E7455FB29F9BD" box="[1128,1246,1598,1621]" gridcol="4" gridrow="9" pageId="14" pageNumber="305">
|
||
<accessionNumber id="48B4AB05FFEBDE78C71F7455FB20F9BC" box="[1129,1239,1598,1620]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/AB981414" pageId="14" pageNumber="305">AB981414</accessionNumber>
|
||
</td>
|
||
<td id="AA065DD8FFEB2189C6797455FA82F9BD" box="[1295,1397,1598,1621]" gridcol="5" gridrow="9" pageId="14" pageNumber="305">0.175</td>
|
||
</tr>
|
||
<tr id="E9D734A4FFEB2189C3E97403FA82F996" box="[159,1397,1640,1662]" gridrow="10" pageId="14" pageNumber="305">
|
||
<th id="AA065DD8FFEB2189C3E97403FEC2F996" box="[159,309,1640,1662]" gridcol="0" gridrow="10" pageId="14" pageNumber="305">
|
||
<taxonomicName id="90E74D65FFEBDE78C3E97402FEF1F996" box="[159,262,1641,1662]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" kingdom="Animalia" order="Anura" pageId="14" pageNumber="305" phylum="Chordata" rank="species" species="jarujini">
|
||
<emphasis id="6593EAF4FFEBDE78C3E97402FEF1F996" box="[159,262,1641,1662]" italics="true" pageId="14" pageNumber="305">L. jarujini</emphasis>
|
||
</taxonomicName>
|
||
</th>
|
||
<td id="AA065DD8FFEB2189C2057403FD67F996" box="[371,656,1640,1662]" gridcol="1" gridrow="10" pageId="14" pageNumber="305">Kaeng Krachan Phetchaburi</td>
|
||
<td id="AA065DD8FFEB2189C18A7403FCA9F996" box="[764,862,1640,1662]" gridcol="2" gridrow="10" pageId="14" pageNumber="305">Thailand</td>
|
||
<td id="AA065DD8FFEB2189C0EA7403FBCFF996" box="[924,1080,1640,1662]" gridcol="3" gridrow="10" pageId="14" pageNumber="305">KUHE 20101</td>
|
||
<td id="AA065DD8FFEB2189C71E7403FB29F996" box="[1128,1246,1640,1662]" gridcol="4" gridrow="10" pageId="14" pageNumber="305">
|
||
<accessionNumber id="48B4AB05FFEBDE78C71F7403FB20F996" box="[1129,1239,1640,1662]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/AB558942" pageId="14" pageNumber="305">AB558942</accessionNumber>
|
||
</td>
|
||
<td id="AA065DD8FFEB2189C6797403FA82F996" box="[1295,1397,1640,1662]" gridcol="5" gridrow="10" pageId="14" pageNumber="305">0.149</td>
|
||
</tr>
|
||
<tr id="E9D734A4FFEB2189C3E974FAFA82F94F" box="[159,1397,1681,1703]" gridrow="11" pageId="14" pageNumber="305">
|
||
<th id="AA065DD8FFEB2189C3E974FAFEC2F94F" box="[159,309,1681,1703]" gridcol="0" gridrow="11" pageId="14" pageNumber="305">
|
||
<taxonomicName id="90E74D65FFEBDE78C3E974F9FEEAF94F" authorityName="Inger, Boeadi & Taufik" authorityYear="1996" box="[159,285,1682,1703]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" kingdom="Animalia" order="Anura" pageId="14" pageNumber="305" phylum="Chordata" rank="species" species="asperatus">
|
||
<emphasis id="6593EAF4FFEBDE78C3E974F9FEEAF94F" box="[159,285,1682,1703]" italics="true" pageId="14" pageNumber="305">L. asperatus</emphasis>
|
||
</taxonomicName>
|
||
</th>
|
||
<td id="AA065DD8FFEB2189C20574FAFD67F94F" box="[371,656,1681,1703]" gridcol="1" gridrow="11" pageId="14" pageNumber="305">Bukit Raya NP, Kalimantan</td>
|
||
<td id="AA065DD8FFEB2189C18A74FAFCA9F94F" box="[764,862,1681,1703]" gridcol="2" gridrow="11" pageId="14" pageNumber="305">Indonesia</td>
|
||
<td id="AA065DD8FFEB2189C0EA74FAFBCFF94F" box="[924,1080,1681,1703]" gridcol="3" gridrow="11" pageId="14" pageNumber="305">RMBR 1193</td>
|
||
<td id="AA065DD8FFEB2189C71E74FAFB29F94F" box="[1128,1246,1681,1703]" gridcol="4" gridrow="11" pageId="14" pageNumber="305">
|
||
<accessionNumber id="48B4AB05FFEBDE78C71E74FAFB29F94F" box="[1128,1246,1681,1703]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/HM067241" pageId="14" pageNumber="305">HM067241</accessionNumber>
|
||
</td>
|
||
<td id="AA065DD8FFEB2189C67974FAFA82F94F" box="[1295,1397,1681,1703]" gridcol="5" gridrow="11" pageId="14" pageNumber="305">0.061</td>
|
||
</tr>
|
||
<tr id="E9D734A4FFEB2189C3E974D1FA82F938" box="[159,1397,1722,1744]" gridrow="12" pageId="14" pageNumber="305">
|
||
<th id="AA065DD8FFEB2189C3E974D1FEC2F938" box="[159,309,1722,1744]" gridcol="0" gridrow="12" pageId="14" pageNumber="305">
|
||
<taxonomicName id="90E74D65FFEBDE78C3E974D0FEEAF938" authorityName="Inger, Boeadi & Taufik" authorityYear="1996" box="[159,285,1723,1744]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" kingdom="Animalia" order="Anura" pageId="14" pageNumber="305" phylum="Chordata" rank="species" species="asperatus">
|
||
<emphasis id="6593EAF4FFEBDE78C3E974D0FEEAF938" box="[159,285,1723,1744]" italics="true" pageId="14" pageNumber="305">L. asperatus</emphasis>
|
||
</taxonomicName>
|
||
</th>
|
||
<td id="AA065DD8FFEB2189C20574D1FD67F938" box="[371,656,1722,1744]" gridcol="1" gridrow="12" pageId="14" pageNumber="305">Kalimantan, Palangkaraya</td>
|
||
<td id="AA065DD8FFEB2189C18A74D1FCA9F938" box="[764,862,1722,1744]" gridcol="2" gridrow="12" pageId="14" pageNumber="305">Indonesia</td>
|
||
<td id="AA065DD8FFEB2189C0EA74D1FBCFF938" box="[924,1080,1722,1744]" gridcol="3" gridrow="12" pageId="14" pageNumber="305">FMNH 252416</td>
|
||
<td id="AA065DD8FFEB2189C71E74D1FB29F938" box="[1128,1246,1722,1744]" gridcol="4" gridrow="12" pageId="14" pageNumber="305">
|
||
<accessionNumber id="48B4AB05FFEBDE78C71F74D1FB22F938" box="[1129,1237,1722,1744]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/AF183128" pageId="14" pageNumber="305">AF183128</accessionNumber>
|
||
</td>
|
||
<td id="AA065DD8FFEB2189C67974D1FA82F938" box="[1295,1397,1722,1744]" gridcol="5" gridrow="12" pageId="14" pageNumber="305">0.167</td>
|
||
</tr>
|
||
<tr id="E9D734A4FFEB2189C3E9748FFA82F912" box="[159,1397,1764,1786]" gridrow="13" pageId="14" pageNumber="305">
|
||
<th id="AA065DD8FFEB2189C3E9748FFEC2F912" box="[159,309,1764,1786]" gridcol="0" gridrow="13" pageId="14" pageNumber="305">
|
||
“
|
||
<taxonomicName id="90E74D65FFEBDE78C3DC748EFED0F912" authorityName="Inger, Boeadi & Taufik" authorityYear="1996" box="[170,295,1765,1786]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" kingdom="Animalia" order="Anura" pageId="14" pageNumber="305" phylum="Chordata" rank="species" species="asperatus">
|
||
<emphasis id="6593EAF4FFEBDE78C3DC748EFED0F912" box="[170,295,1765,1786]" italics="true" pageId="14" pageNumber="305">L. asperatus</emphasis>
|
||
</taxonomicName>
|
||
“
|
||
</th>
|
||
<td id="AA065DD8FFEB2189C205748FFD67F912" box="[371,656,1764,1786]" gridcol="1" gridrow="13" pageId="14" pageNumber="305">Serasan Island</td>
|
||
<td id="AA065DD8FFEB2189C18A748FFCA9F912" box="[764,862,1764,1786]" gridcol="2" gridrow="13" pageId="14" pageNumber="305">Indonesia</td>
|
||
<td id="AA065DD8FFEB2189C0EA748FFBCFF912" box="[924,1080,1764,1786]" gridcol="3" gridrow="13" pageId="14" pageNumber="305">BJE 170</td>
|
||
<td id="AA065DD8FFEB2189C71E748FFB29F912" box="[1128,1246,1764,1786]" gridcol="4" gridrow="13" pageId="14" pageNumber="305">
|
||
<accessionNumber id="48B4AB05FFEBDE78C71F748FFB2AF912" box="[1129,1245,1764,1786]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/HM067283" pageId="14" pageNumber="305">HM067283</accessionNumber>
|
||
</td>
|
||
<td id="AA065DD8FFEB2189C679748FFA82F912" box="[1295,1397,1764,1786]" gridcol="5" gridrow="13" pageId="14" pageNumber="305">0.018</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<paragraph id="575836E6FFEBDE78C3B17552FBF9F80A" blockId="14.[151,1436,1849,2018]" pageId="14" pageNumber="305">
|
||
<emphasis id="6593EAF4FFEBDE78C3B17552FEDCF8BA" bold="true" box="[199,299,1849,1874]" pageId="14" pageNumber="305">Ecology.</emphasis>
|
||
The
|
||
<typeStatus id="885C8844FFEBDE78C2077551FE56F8BA" box="[369,417,1850,1874]" pageId="14" pageNumber="305">type</typeStatus>
|
||
speCimens were ColleCted from Calm seCtions of small streams (width 0.3–1.0 m) at elevations between 10 and
|
||
<quantity id="901F9B03FFEBDE78C2A07535FDD7F89D" box="[470,544,1886,1910]" metricMagnitude="2" metricUnit="m" metricValue="4.0" pageId="14" pageNumber="305" unit="m" value="400.0">400 m</quantity>
|
||
a.s.l., where they sat in shallow water. Upon disseCtion, we found the male
|
||
<typeStatus id="885C8844FFEBDE78C3E175EAFF0EF872" box="[151,249,1921,1946]" pageId="14" pageNumber="305" type="holotype">holotype</typeStatus>
|
||
to be heavily infested with many speCimens of an unidentified speCies of nematode whiCh were found enCapsuled on the surfaCe of the stomaCh (
|
||
<figureCitation id="CFDC2A63FFEBDE78C1FB75CEFD22F856" box="[653,725,1957,1982]" captionStart="FIGURE 8" captionStartId="12.[151,250,987,1009]" captionTargetBox="[151,1436,193,965]" captionTargetId="figure@12.[151,1436,193,965]" captionTargetPageId="12" captionText="FIGURE 8. (A) Ventral view oF the euthanized holotype oF Limnonectes kong sp. nov. (NMBE 1059923, adult male); (B) detail oF stomach, showing encapsuled nematodes." httpUri="https://zenodo.org/record/884335/files/figure.png" pageId="14" pageNumber="305">Fig. 8</figureCitation>
|
||
). StomaCh and intestines of the
|
||
<typeStatus id="885C8844FFEBDE78C72775CEFB44F856" box="[1105,1203,1957,1982]" pageId="14" pageNumber="305" type="holotype">holotype</typeStatus>
|
||
Contained Chitinous remains of beetle head Capsules, body plates, and the hind legs of large loCusts.
|
||
</paragraph>
|
||
<caption id="0398666EFFEADE79C3E17657FC2BFB98" httpUri="https://zenodo.org/record/884339/files/figure.png" pageId="15" pageNumber="306" targetBox="[157,1433,209,1053]" targetPageId="15">
|
||
<paragraph id="575836E6FFEADE79C3E17657FC2BFB98" blockId="15.[151,1436,1084,1136]" pageId="15" pageNumber="306">
|
||
<emphasis id="6593EAF4FFEADE79C3E17657FEE8FBB9" bold="true" box="[151,287,1084,1106]" pageId="15" pageNumber="306">FIGURE 10.</emphasis>
|
||
Collection localities oF
|
||
<taxonomicName id="90E74D65FFEADE79C1647656FD3EFBBA" authority="Dehling, 2017" authorityName="Dehling" authorityYear="2017" box="[530,713,1084,1106]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" kingdom="Animalia" order="Anura" pageId="15" pageNumber="306" phylum="Chordata" rank="species" species="kong" status="sp. nov.">
|
||
<emphasis id="6593EAF4FFEADE79C1647656FD3EFBBA" box="[530,713,1084,1106]" italics="true" pageId="15" pageNumber="306">Limnonectes kong</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="6593EAF4FFEADE79C1B97657FCD7FBB9" bold="true" box="[719,800,1084,1105]" pageId="15" pageNumber="306">
|
||
<taxonomicNameLabel id="7EA0578FFFEADE79C1B97657FCD7FBB9" box="[719,800,1084,1105]" pageId="15" pageNumber="306" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
in western Sarawak, Borneo at (1) Gunung Gading, (2) Kubah National Park, Matang, (3), Santubong National Park, and (4) Bako National Park.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="0398666EFFEADE79C3E175FBFB46F80B" httpUri="https://zenodo.org/record/884341/files/figure.png" pageId="15" pageNumber="306" targetBox="[151,1436,1186,1914]" targetPageId="15">
|
||
<paragraph id="575836E6FFEADE79C3E175FBFB46F80B" blockId="15.[151,1436,1936,2019]" pageId="15" pageNumber="306">
|
||
<emphasis id="6593EAF4FFEADE79C3E175FBFEE9F84D" bold="true" box="[151,286,1936,1958]" pageId="15" pageNumber="306">FIGURE 11.</emphasis>
|
||
Ventral view oF the opened abdomina showing the testes (yellow circles) in male type specimens oF
|
||
<taxonomicName id="90E74D65FFEADE79C66A75FAFF3FF82C" authority="Dehling, 2017" authorityName="Dehling" authorityYear="2017" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" kingdom="Animalia" order="Anura" pageId="15" pageNumber="306" phylum="Chordata" rank="species" species="kong" status="sp. nov.">
|
||
<emphasis id="6593EAF4FFEADE79C66A75FAFF3FF82C" italics="true" pageId="15" pageNumber="306">Limnonectes kong</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="6593EAF4FFEADE79C3B875C4FEE9F82C" bold="true" box="[206,286,1967,1988]" pageId="15" pageNumber="306">
|
||
<taxonomicNameLabel id="7EA0578FFFEADE79C3B875C4FEE9F82C" box="[206,286,1967,1988]" pageId="15" pageNumber="306" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
: (A) Holotype, NMBE 1059923, adult male, SVL 68.3 mm; (B) paratype, NMBE 1059924, adult male, 76.3 mm; and (C) paratype, NMBE 1059925, possible subadult male, SVL 53.1 mm. Scale bars represent 10 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="575836E6FFF5DE66C3B172FCFBDFFEA8" blockId="16.[151,1437,151,680]" pageId="16" pageNumber="307">
|
||
Adult females ColleCted in September and OCtober Contained enlarged ova whiCh had a dark, almost blaCk animal pole and a Cream-Coloured vegetative pole. One
|
||
<typeStatus id="885C8844FFF5DE66C07872D6FC99FF3D" box="[782,878,189,213]" pageId="16" pageNumber="307" type="paratype">paratype</typeStatus>
|
||
(NMBE 1059918) was found in a small Cavity at the stream edge (
|
||
<figureCitation id="CFDC2A63FFF5DE66C21472B4FE5EFF10" box="[354,425,223,248]" captionStart="FIGURE 9" captionStartId="13.[151,250,1780,1802]" captionTargetBox="[151,1436,805,1769]" captionTargetId="figure@13.[151,1436,805,1769]" captionTargetPageId="13" captionText="FIGURE 9. Limnonectes kong sp. nov.: (A) Female paratype (NMBE 1059918) calling From a cavity in the ground, Matang, Sarawak; (B) Female paratype (NMBE 1059928) perched on a rock along a stream, Gading NP, Sarawak; (C) lateral view and (D) dorsal view oF a male paratype (NMBE 1059924), Bako NP, Sarawak." httpUri="https://zenodo.org/record/884337/files/figure.png" pageId="16" pageNumber="307">Fig. 9</figureCitation>
|
||
a). The female repeatedly emitted a snarling, drawn-out, low-frequenCy Call that lasted between 0.5 and 3 seConds. A seCond speCimen Called simultaneously in the same way from farther away but we were unable to loCate it and determine whether it was a male or female speCimen.
|
||
</paragraph>
|
||
<paragraph id="575836E6FFF5DE66C3B17327FE90FDFF" blockId="16.[151,1437,151,680]" pageId="16" pageNumber="307">
|
||
The testes of the largest male speCimens, inCluding the holotype, whiCh exhibit the most strongly developed seCondary sexual CharaCteristiCs (large head, large odontoid proCesses, tuberCles on posterior half of body), are relatively small (length x width
|
||
<quantity id="901F9B03FFF5DE66C28C73FFFDA7FE44" box="[506,592,404,428]" metricMagnitude="-3" metricUnit="m" metricValue="3.4" pageId="16" pageNumber="307" unit="mm" value="3.4">3.4 mm</quantity>
|
||
x
|
||
<quantity id="901F9B03FFF5DE66C11C73FFFD37FE44" box="[618,704,404,429]" metricMagnitude="-3" metricUnit="m" metricValue="1.6" pageId="16" pageNumber="307" unit="mm" value="1.6">1.6 mm</quantity>
|
||
[holotype], 4.8 x
|
||
<quantity id="901F9B03FFF5DE66C0F373FFFC2CFE44" box="[901,987,404,428]" metricMagnitude="-3" metricUnit="m" metricValue="2.2" pageId="16" pageNumber="307" unit="mm" value="2.2">2.2 mm</quantity>
|
||
[paratype NMBE 1059924,
|
||
<figureCitation id="CFDC2A63FFF5DE66C66E73FFFAACFE44" box="[1304,1371,404,429]" captionStart="FIGURE 9" captionStartId="13.[151,250,1780,1802]" captionTargetBox="[151,1436,805,1769]" captionTargetId="figure@13.[151,1436,805,1769]" captionTargetPageId="13" captionText="FIGURE 9. Limnonectes kong sp. nov.: (A) Female paratype (NMBE 1059918) calling From a cavity in the ground, Matang, Sarawak; (B) Female paratype (NMBE 1059928) perched on a rock along a stream, Gading NP, Sarawak; (C) lateral view and (D) dorsal view oF a male paratype (NMBE 1059924), Bako NP, Sarawak." httpUri="https://zenodo.org/record/884337/files/figure.png" pageId="16" pageNumber="307">Fig. 9</figureCitation>
|
||
];
|
||
<figureCitation id="CFDC2A63FFF5DE66C60673FFFF43FE38" captionStart="FIGURE 11" captionStartId="15.[151,250,1936,1958]" captionTargetBox="[151,1436,1186,1914]" captionTargetId="figure@15.[151,1436,1186,1914]" captionTargetPageId="15" captionText="FIGURE 11. Ventral view oF the opened abdomina showing the testes (yellow circles) in male type specimens oF Limnonectes kong sp. nov.: (A) Holotype, NMBE 1059923, adult male, SVL 68.3 mm; (B) paratype, NMBE 1059924, adult male, 76.3 mm; and (C) paratype, NMBE 1059925, possible subadult male, SVL 53.1 mm. Scale bars represent 10 mm." httpUri="https://zenodo.org/record/884341/files/figure.png" pageId="16" pageNumber="307">Fig. 11</figureCitation>
|
||
). A smaller male (NMBE 1059925) that we determined to be “subadult” based on SVL, relative small head size, and small size of odontoid proCesses, has similarly sized testes (
|
||
<quantity id="901F9B03FFF5DE66C0F473B7FC2BFE1C" box="[898,988,476,500]" metricMagnitude="-3" metricUnit="m" metricValue="3.2" pageId="16" pageNumber="307" unit="mm" value="3.2">3.2 mm</quantity>
|
||
x
|
||
<quantity id="901F9B03FFF5DE66C08873B7FBADFE1C" box="[1022,1114,476,500]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" pageId="16" pageNumber="307" unit="mm" value="1.3">1.3 mm</quantity>
|
||
;
|
||
<figureCitation id="CFDC2A63FFF5DE66C71F73B7FB4BFE1C" box="[1129,1212,476,501]" captionStart="FIGURE 11" captionStartId="15.[151,250,1936,1958]" captionTargetBox="[151,1436,1186,1914]" captionTargetId="figure@15.[151,1436,1186,1914]" captionTargetPageId="15" captionText="FIGURE 11. Ventral view oF the opened abdomina showing the testes (yellow circles) in male type specimens oF Limnonectes kong sp. nov.: (A) Holotype, NMBE 1059923, adult male, SVL 68.3 mm; (B) paratype, NMBE 1059924, adult male, 76.3 mm; and (C) paratype, NMBE 1059925, possible subadult male, SVL 53.1 mm. Scale bars represent 10 mm." httpUri="https://zenodo.org/record/884341/files/figure.png" pageId="16" pageNumber="307">Fig. 11</figureCitation>
|
||
), and might in faCt already be mature.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="1FFD656DFFF5DE66C3B1704FFCB0FD40" pageId="16" pageNumber="307" type="etymology">
|
||
<paragraph id="575836E6FFF5DE66C3B1704FFCB0FD40" blockId="16.[151,1437,151,680]" pageId="16" pageNumber="307">
|
||
<emphasis id="6593EAF4FFF5DE66C3B1704FFEB9FDD5" bold="true" box="[199,334,548,573]" pageId="16" pageNumber="307">Etymology.</emphasis>
|
||
The speCies epithet is an allusion to the pronounCed sexual dimorphism exhibited by the new speCies. It resembles the male-biased dimorphism in great apes whiCh is most pronounCed in gorillas (
|
||
<emphasis id="6593EAF4FFF5DE66C6737023FAAFFD88" box="[1285,1368,584,608]" italics="true" pageId="16" pageNumber="307">Gorilla</emphasis>
|
||
spp.). SinCe the new speCies is also among the largest of the
|
||
<taxonomicName id="90E74D65FFF5DE66C1857006FCA6FD6C" baseAuthorityName="TsChudi" baseAuthorityYear="1838" box="[755,849,620,644]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" kingdom="Animalia" order="Anura" pageId="16" pageNumber="307" phylum="Chordata" rank="species" species="kuhlii">
|
||
<emphasis id="6593EAF4FFF5DE66C1857006FCA6FD6C" box="[755,849,620,644]" italics="true" pageId="16" pageNumber="307">L. kuhlii</emphasis>
|
||
</taxonomicName>
|
||
group on Borneo we name it after the fiCtional giant gorilla. The name is used as an invariable noun in apposition.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |