166 lines
12 KiB
XML
166 lines
12 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.773.24878" ID-GBIF-Dataset="3b67ab3e-bd75-4194-a90c-c12b3dc8332b" ID-Pensoft-Pub="1313-2970-773-1" ID-ZBK="900F9307844A4B9AB428DA97FF3B4B5C" ModsDocAuthor="" ModsDocDate="2018" ModsDocID="1313-2970-773-1" ModsDocOrigin="ZooKeys 773" ModsDocTitle="A molecular and conchological dissection of the “scaly” Georissa of Malaysian Borneo (Gastropoda, Neritimorpha, Hydrocenidae)" checkinTime="1531209184653" checkinUser="pensoft" docAuthor="Khalik, Mohd Zacaery, Hendriks, Kasper, Vermeulen, Jaap J. & Schilthuizen, Menno" docDate="2018" docId="B0798B78B9C62055C2AFD9059605CF99" docLanguage="en" docName="ZooKeys 773: 1-55" docOrigin="ZooKeys 773" docSource="http://dx.doi.org/10.3897/zookeys.773.24878" docTitle="Georissa kinabatanganensis Khalik, Hendriks, Vermeulen & Schilthuizen, 2018, sp. n." docType="treatment" docUuid="A952A54F-D486-4C27-AFCA-8E7020E41ADA" docUuidSource="ZooBank" docVersion="5" lastPageNumber="1" masterDocId="FFACFFA96149FF8D3477FFCCD227FF8E" masterDocTitle="A molecular and conchological dissection of the " scaly " Georissa of Malaysian Borneo (Gastropoda, Neritimorpha, Hydrocenidae)" masterLastPageNumber="55" masterPageNumber="1" pageNumber="1" updateTime="1643514872515" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>A molecular and conchological dissection of the " scaly " Georissa of Malaysian Borneo (Gastropoda, Neritimorpha, Hydrocenidae)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Khalik, Mohd Zacaery</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Hendriks, Kasper</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Vermeulen, Jaap J.</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Schilthuizen, Menno</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2018</mods:date>
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<mods:detail type="volume">
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<mods:number>773</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>55</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.773.24878</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.773.24878</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-773-1</mods:identifier>
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<mods:identifier type="ZBK">900F9307844A4B9AB428DA97FF3B4B5C</mods:identifier>
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<mods:identifier type="ZooBank">900F9307844A4B9AB428DA97FF3B4B5C</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="146053682" LSID="urn:lsid:zoobank.org:act:A952A54F-D486-4C27-AFCA-8E7020E41ADA" httpUri="http://treatment.plazi.org/id/B0798B78B9C62055C2AFD9059605CF99" lastPageNumber="1" pageId="0" pageNumber="1">
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<subSubSection pageId="0" pageNumber="1" type="nomenclature">
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName LSID="http://zoobank.org/A952A54F-D486-4C27-AFCA-8E7020E41ADA" class="Gastropoda" family="Hydrocenidae" genus="Georissa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Georissa kinabatanganensis" order="Cycloneritida" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="kinabatanganensis">Georissa kinabatanganensis</taxonomicName>
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<taxonomicNameLabel pageId="0" pageNumber="1">sp. n.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="type locality">
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<paragraph pageId="0" pageNumber="1">Type locality.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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Bukit Keruak, near Kinabatangan river, Sandakan, Sabah, Malaysia (
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<geoCoordinate direction="north" orientation="latitude" precision="1" value="5.523083">05°31.385'N</geoCoordinate>
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,
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<geoCoordinate direction="east" orientation="longitude" precision="1" value="118.28522">118°17.113'E</geoCoordinate>
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).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="type material">
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<paragraph pageId="0" pageNumber="1">Type material.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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Holotype. Bukit Keruak, near Kinabatangan river, Sandakan, Sabah, Malaysia (
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<geoCoordinate direction="north" orientation="latitude" precision="9" value="5.523">05°31.38'N</geoCoordinate>
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,
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<geoCoordinate direction="east" orientation="longitude" precision="9" value="118.285164">118°17.11'E</geoCoordinate>
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): BOR/MOL 13921 (leg. M Schilthuizen). Paratypes. Bukit Keruak, near Kinabatangan river, Sandakan, Sabah (
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<geoCoordinate direction="north" orientation="latitude" precision="9" value="5.523">05°31.38'N</geoCoordinate>
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,
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<geoCoordinate direction="east" orientation="longitude" precision="9" value="118.285164">118°17.11'E</geoCoordinate>
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): MZU/MOL 17.26 (>50). BOR/MOL 1458, BOR/MOL 11656, BOR/MOL 11665, BOR/MOL 11711, BOR/MOL 13871. Batu Pangi, Sabah: BOR/MOL 1455. Batu Tomanggong, Sabah: BOR/MOL 1456, BOR/MOL 1457, BOR/MOL 10530.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="etymology">
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<paragraph pageId="0" pageNumber="1">Etymology.</paragraph>
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<paragraph pageId="0" pageNumber="1">Named after the district of Kinabatangan, Sabah, Malaysia, where the type locality Bukit Keruak is located.</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="description">
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<paragraph pageId="0" pageNumber="1">Description.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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Protoconch. Color: orange. Sculpture pattern: smooth to meshed - rounded to undefined mesh pattern. Mesh width: 14-21
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<normalizedToken originalValue="µm">µm</normalizedToken>
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. Teleoconch. Color: orange. First whorl: flat and angular at the shoulder. Subsequent whorls: angular, slightly rounded at the periphery, with number of whorls: 2-2
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<normalizedToken originalValue="¼">1/4</normalizedToken>
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. SH: 1.00-1.32 mm, SW: 1.13-1.37 mm, SI: 0.85-0.99. Shell sculpture. Radial sculpture: absent, only weak to strong growth lines are visible. Spiral sculpture: present, and strongly sculpted, with continuous to discontinuous ribbings. Scales: two series of diagonal vertical scales, widely spaced in between, both series are strongly sculpted, broad, and the scales are regularly placed. Aperture. Shape: oval. Basal side: rounded, angular at the columellar region. Parietal side: straight, palatal edge attached to the body whorl. AH: 0.54-0.66 mm, AW: 0.75-0.86 mm, AI: 0.65-0.80. Holotype dimension. SH: 1.00 mm, SW: 1.18 mm, AH: 0.54 mm, AW: 0.78 mm.
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</paragraph>
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<caption pageId="0" pageNumber="1">
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<paragraph pageId="0" pageNumber="1">
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Figure 16.
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<taxonomicName class="Gastropoda" family="Hydrocenidae" genus="Georissa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Georissa kinabatanganensis" order="Cycloneritida" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="kinabatanganensis">Georissa kinabatanganensis</taxonomicName>
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sp. n.
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<normalizedToken originalValue="A–C">A-C</normalizedToken>
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Holotype: BOR/MOL 13921
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<normalizedToken originalValue="D–K">D-K</normalizedToken>
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Paratypes: MZU/MOL 17.26. A, D Shell apertural view B Shell side view C Shell rear view
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<normalizedToken originalValue="E–F">E-F</normalizedToken>
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Shell cross-section from 3D model
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<normalizedToken originalValue="G–H">G-H</normalizedToken>
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Operculum frontal and ventral view I Shell top view J Protoconch side view K Close up of protoconch from top at 1000
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<normalizedToken originalValue="×">x</normalizedToken>
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magnification. Scale bars: 500
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<normalizedToken originalValue="µm">µm</normalizedToken>
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(
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<normalizedToken originalValue="A–I">A-I</normalizedToken>
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); 200
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<normalizedToken originalValue="µm">µm</normalizedToken>
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(J); 10
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<normalizedToken originalValue="µm">µm</normalizedToken>
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(K).
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="cross diagnosis">
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<paragraph pageId="0" pageNumber="1">Cross diagnosis.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName class="Gastropoda" family="Hydrocenidae" genus="Georissa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Georissa kinabatanganensis" order="Cycloneritida" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="kinabatanganensis">Georissa kinabatanganensis</taxonomicName>
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has less variation in shell sculpture compared with
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<taxonomicName genus="G." lsidName="G. hosei" pageId="0" pageNumber="1" rank="species" species="hosei">G. hosei</taxonomicName>
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and
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<taxonomicName genus="G." lsidName="G. scalinella" pageId="0" pageNumber="1" rank="species" species="scalinella">G. scalinella</taxonomicName>
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.
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<taxonomicName genus="G." lsidName="G. kinabatanganensis" pageId="0" pageNumber="1" rank="species" species="kinabatanganensis">G. kinabatanganensis</taxonomicName>
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has two series of acutely projected scales on the whorls. In some cases, the second scale series is weaker than the first, and creates a series of nodular structures at the periphery. Often the shell is wider than high, which gives it a flattened appearance. In addition,
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<taxonomicName genus="G." lsidName="G. kinabatanganensis" pageId="0" pageNumber="1" rank="species" species="kinabatanganensis">G. kinabatanganensis</taxonomicName>
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has widely spaced between the scale series, similar to
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<taxonomicName genus="G." lsidName="G. muluensis" pageId="0" pageNumber="1" rank="species" species="muluensis">G. muluensis</taxonomicName>
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.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="distribution">
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<paragraph pageId="0" pageNumber="1">Distribution.</paragraph>
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<paragraph pageId="0" pageNumber="1">Known from Bukit Keruak, Batu Tomanggong, and Pangi, in the region of Kinabatangan, Sabah.</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="molecular analysis">
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<paragraph pageId="0" pageNumber="1">Molecular analysis.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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RAxML and Bayesian analyses show
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<taxonomicName genus="G." lsidName="G. kinabatanganensis" pageId="0" pageNumber="1" rank="species" species="kinabatanganensis">G. kinabatanganensis</taxonomicName>
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(16S: n = 6; CO1: n = 6) forming a clade with 97% BS and 97% PP and as a sister clade to
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<taxonomicName genus="G." lsidName="G. sepulutensis" pageId="0" pageNumber="1" rank="species" species="sepulutensis">G. sepulutensis</taxonomicName>
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.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="discussion">
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<paragraph pageId="0" pageNumber="1">Discussion.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName class="Gastropoda" family="Hydrocenidae" genus="Georissa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Georissa kinabatanganensis" order="Cycloneritida" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="kinabatanganensis">Georissa kinabatanganensis</taxonomicName>
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is the only species in "scaly group"
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<taxonomicName class="Gastropoda" family="Hydrocenidae" genus="Georissa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Georissa" order="Cycloneritida" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">Georissa</taxonomicName>
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to have a flat shell habitus, all examined individuals have a shell that is broader than high.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |