treatments-xml/data/9F/52/87/9F5287886A1CFF8F94EA6F08AC6AFEE4.xml
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<document id="2505D9C6DEEBE668DC90AA68EC9BFE6A" ID-CLB-Dataset="58359" ID-DOI="10.11646/zootaxa.5285.1.7" ID-GBIF-Dataset="77e7fae7-a1e0-43cb-af62-d4e9597f329f" ID-ISSN="1175-5326" ID-Zenodo-Dep="7935889" ID-ZooBank="F2895A5A-A06E-480F-A6C8-061150B57041" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="carolina" checkinTime="1684131653704" checkinUser="plazi" docAuthor="Pati, Sameer Kumar, Bajantri, Parashuram Prabhu &amp; Hegde, Gopalkrishna Dattatraya" docDate="2023" docId="9F5287886A1CFF8F94EA6F08AC6AFEE4" docLanguage="en" docName="zootaxa.5285.1.7.pdf" docOrigin="Zootaxa 5285 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5285.1.7" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Aradhya placida Pati &amp; Bajantri &amp; Hegde 2023, sp. nov." docType="treatment" docUuid="84351375-954D-41EB-BB6C-C175745194AD" docUuidSource="ZooBank" docVersion="8" lastPageNumber="174" masterDocId="636BFFF06A15FF82947D685EA945FFA5" masterDocTitle="A new genus and new species of freshwater crab (Decapoda: Brachyura: Gecarcinucidae) from the Central Western Ghats of India" masterLastPageNumber="175" masterPageNumber="161" pageNumber="170" updateTime="1699505753996" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="84B6644B9ACC9302783EA17818E6DCB3">A new genus and new species of freshwater crab (Decapoda: Brachyura: Gecarcinucidae) from the Central Western Ghats of India</mods:title>
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<mods:namePart id="202AF90EB013806ED0B6CB8918AA6FA8">Hegde, Gopalkrishna Dattatraya</mods:namePart>
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<heading id="4C0C81F26A1CFF8B94EA6F08A881F8D4" bold="true" box="[151,452,1878,1905]" fontSize="11" level="1" pageId="9" pageNumber="170" reason="1">
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<taxonomicName id="D0FB4D1D6A1CFF8B94EA6F08A81AF8D4" ID-CoL="90e8146e-992e-427d-923d-7cc05de0c133" authority="Pati &amp; Bajantri &amp; Hegde, 2023" authorityName="Pati &amp; Bajantri &amp; Hegde" authorityYear="2023" box="[151,351,1878,1905]" class="Malacostraca" family="Gecarcinucidae" genus="Aradhya" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="170" phylum="Arthropoda" rank="species" species="placida" status="sp. nov.">
<emphasis id="258FEA8C6A1CFF8B94EA6F08A81AF8D4" bold="true" box="[151,351,1878,1905]" italics="true" pageId="9" pageNumber="170">Aradhya placida</emphasis>
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<paragraph id="1744369E6A1CFF8B94EA6F25A802F830" blockId="9.[151,452,1878,1977]" box="[151,327,1915,1941]" pageId="9" pageNumber="170">[Peaceful Crab]</paragraph>
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(
<figureCitation id="8FC02A1B6A1CFF8B94E26FC1A84EF81C" box="[159,267,1951,1977]" captionStart-0="FIGURE 2" captionStart-1="FIGURE 3" captionStart-2="FIGURE 4" captionStart-3="FIGURE 5" captionStart-4="FIGURE 6" captionStartId-0="3.[151,250,1943,1967]" captionStartId-1="4.[151,250,1858,1882]" captionStartId-2="5.[151,250,1910,1934]" captionStartId-3="6.[151,250,1943,1967]" captionStartId-4="7.[151,250,1352,1376]" captionTargetBox-0="[192,1395,181,1919]" captionTargetBox-1="[151,1435,181,1834]" captionTargetBox-2="[214,1372,181,1886]" captionTargetBox-3="[203,1384,181,1919]" captionTargetBox-4="[153,1434,419,1327]" captionTargetId-0="figure-19@3.[192,1395,181,1919]" captionTargetId-1="figure-24@4.[151,1435,181,1834]" captionTargetId-2="figure-19@5.[214,1372,181,1886]" captionTargetId-3="figure-24@6.[203,1384,181,1919]" captionTargetId-4="figure-183@7.[151,1436,417,1329]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="5" captionTargetPageId-3="6" captionTargetPageId-4="7" captionText-0="FIGURE 2. Aradhya placida gen. et sp. nov., holotype male (25.12 × 19.85 mm) (ZSI-WRC C.2292). A, overall dorsal view; B, frontal view of cephalothorax; C, overall ventral view. Scale bar = 10 mm." captionText-1="FIGURE 3. Aradhya placida gen. et sp. nov.: paratype male (28.53 × 21.82 mm) (ZSI-WRC C.2294) (A, B); paratype male (23.63 × 19.34 mm) (ZSI-WGRC IR.INV.22823) (C, D); paratype male (23.09 × 18.23 mm) (ZSI-WRC C.2293) (EI). A, C, E, overall dorsal view; B, D, F, overall ventral view; G, third maxillipeds; H, dorsal view of left G1; I, left G2. Scale bars = 10 mm (AF), 2 mm (G, H), 1 mm (I)." captionText-2="FIGURE 4. Aradhya placida gen. et sp. nov.: paratype female (27.03 × 20.76 mm) (ZSI-WRC C.2294) (A); paratype female (26.69 × 20.87 mm) (ZSI-WRC C.2293) (B, DG); paratype female (23.51 × 19.03 mm) (ZSI-WGRC IR.INV.22823) (C). A, B, C, overall dorsal view; D, third maxillipeds; E, dorsal view of pleonal somites 46 and telson; F, ventral view of pleon with pleopods; G, thoracic sternites showing vulvae. Scale bars = 10 mm (AC), 5 mm (EG), 2 mm (D)." captionText-3="FIGURE 5. Aradhya placida gen. et sp. nov., holotype male (25.12 × 19.85 mm) (ZSI-WRC C.2292).A, mouth parts exposed; B, third maxillipeds; C, thoracic sternites with G1 and G2; D, pleon and telson; E, dorsal view of left G1; F, left G2. Scale bars = 5 mm (C, D), 2 mm (A, B, E), 1 mm (F)." captionText-4="FIGURE 6. Aradhya placida gen. et sp. nov., holotype male (25.12 × 19.85 mm) (ZSI-WRC C.2292). A, pleon and telson; B, dorsal view of left G1; C, dorsal view of left G1 ultimate article; D, ventral view of left G1; E, ventral view of left G1 ultimate article; F, left G2. Scale bars = 5 mm (A), 2 mm (B, D), 1 mm (C, E, F)." figureDoi-0="http://doi.org/10.5281/zenodo.7935895" figureDoi-1="http://doi.org/10.5281/zenodo.7935897" figureDoi-2="http://doi.org/10.5281/zenodo.7935901" figureDoi-3="http://doi.org/10.5281/zenodo.7935903" figureDoi-4="http://doi.org/10.5281/zenodo.7935907" httpUri-0="https://zenodo.org/record/7935895/files/figure.png" httpUri-1="https://zenodo.org/record/7935897/files/figure.png" httpUri-2="https://zenodo.org/record/7935901/files/figure.png" httpUri-3="https://zenodo.org/record/7935903/files/figure.png" httpUri-4="https://zenodo.org/record/7935907/files/figure.png" pageId="9" pageNumber="170">Figs. 26</figureCitation>
,
<figureCitation id="8FC02A1B6A1CFF8B956A6FC1A819F81C" box="[279,348,1951,1977]" captionStart="FIGURE 7" captionStartId="11.[151,250,1928,1952]" captionTargetBox="[151,1435,921,1905]" captionTargetId="figure-329@11.[151,1435,921,1905]" captionTargetPageId="11" captionText="FIGURE 7. Colour in life and habitats of Aradhya placida gen. et sp. nov. A, holotype male (25.12 × 19.85 mm) (ZSI-WRC C.2292); B, paratype male (23.09 × 18.23 mm) (ZSI-WRC C.2293); C, a live crab inside the burrow; D, a live crab near the base of a tree; E, trees and boulders at the type locality. This figure is available in colour at Zootaxa online." figureDoi="http://doi.org/10.5281/zenodo.7935910" httpUri="https://zenodo.org/record/7935910/files/figure.png" pageId="9" pageNumber="170">7AD</figureCitation>
)
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<emphasis id="258FEA8C6A1FFF8894EA68C9A801FF15" bold="true" box="[151,324,150,177]" pageId="10" pageNumber="171">Type material.</emphasis>
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, CL
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, CH
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, FW
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(
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C.2292).
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, CL
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, CH
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, FW
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),
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(CW
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, CL
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<quantity id="D0039B7B6A1FFF8895926979AB26FEE4" box="[495,611,295,321]" metricMagnitude="-2" metricUnit="m" metricValue="1.8079999999999998" pageId="10" pageNumber="171" unit="mm" value="18.08">18.08 mm</quantity>
, FW
<quantity id="D0039B7B6A1FFF8896E26979AA42FEE4" box="[671,775,295,321]" metricMagnitude="-3" metricUnit="m" metricValue="6.02" pageId="10" pageNumber="171" unit="mm" value="6.02">6.02 mm</quantity>
), same collection data as for holotype (
<collectionCode id="71EAAE5B6A1FFF8890C06979AC77FEE4" box="[1213,1330,295,321]" pageId="10" pageNumber="171">ZSI-WRC</collectionCode>
C.2293);
</materialsCitation>
<materialsCitation id="A7933CC36A1FFF8894EA6915AA92FEC0" ID-GBIF-Occurrence="4109228301" box="[151,983,330,357]" pageId="10" pageNumber="171" specimenCount="2" specimenCount-adult="1" specimenCount-male="1">
<specimenCount id="01FDFD176A1FFF8894EA6915A98AFEC0" box="[151,207,331,357]" pageId="10" pageNumber="171" type="adult">adult</specimenCount>
<specimenCount id="01FDFD176A1FFF8894AA6915A848FEC0" box="[215,269,331,357]" pageId="10" pageNumber="171" type="male">male</specimenCount>
(CW
<quantity id="D0039B7B6A1FFF88952E6914A88DFEC0" box="[339,456,330,357]" metricMagnitude="-2" metricUnit="m" metricValue="2.853" pageId="10" pageNumber="171" unit="mm" value="28.53">28.53 mm</quantity>
, CL
<quantity id="D0039B7B6A1FFF8895826915AB31FEC0" box="[511,628,331,357]" metricMagnitude="-2" metricUnit="m" metricValue="2.182" pageId="10" pageNumber="171" unit="mm" value="21.82">21.82 mm</quantity>
, CH
<quantity id="D0039B7B6A1FFF8896CD6915AA60FEC0" box="[688,805,331,357]" metricMagnitude="-2" metricUnit="m" metricValue="1.8829999999999998" pageId="10" pageNumber="171" unit="mm" value="18.83">18.83 mm</quantity>
, FW
<quantity id="D0039B7B6A1FFF88971F6915AA8EFEC0" box="[866,971,331,357]" metricMagnitude="-3" metricUnit="m" metricValue="6.37" pageId="10" pageNumber="171" unit="mm" value="6.37">6.37 mm</quantity>
),
</materialsCitation>
<materialsCitation id="A7933CC36A1FFF8897A26915AC18FE2C" ID-GBIF-Occurrence="4109228305" collectingDate="2022-07-11" collectionCode="ZSI-WRC" pageId="10" pageNumber="171" specimenCount="2" specimenCount-adult="1" specimenCount-female="1">
<specimenCount id="01FDFD176A1FFF8897A26915AD52FEC0" box="[991,1047,331,357]" pageId="10" pageNumber="171" type="adult">adult</specimenCount>
<specimenCount id="01FDFD176A1FFF8890626915AD2EFEC0" box="[1055,1131,331,357]" pageId="10" pageNumber="171" type="female">female</specimenCount>
(CW
<quantity id="D0039B7B6A1FFF8890CC6915AC63FEC0" box="[1201,1318,331,357]" metricMagnitude="-2" metricUnit="m" metricValue="2.703" pageId="10" pageNumber="171" unit="mm" value="27.03">27.03 mm</quantity>
, CL
<quantity id="D0039B7B6A1FFF8891206915A980FE2C" metricMagnitude="-2" metricUnit="m" metricValue="2.076" pageId="10" pageNumber="171" unit="mm" value="20.76">20.76 mm</quantity>
, CH
<quantity id="D0039B7B6A1FFF88957C6930A833FE2C" box="[257,374,366,393]" metricMagnitude="-2" metricUnit="m" metricValue="1.85" pageId="10" pageNumber="171" unit="mm" value="18.5">18.50 mm</quantity>
, FW
<quantity id="D0039B7B6A1FFF8895C86931AB5BFE2C" box="[437,542,367,393]" metricMagnitude="-3" metricUnit="m" metricValue="6.41" pageId="10" pageNumber="171" unit="mm" value="6.41">6.41 mm</quantity>
), same collection data as for holotype,
<date id="6345105E6A1FFF8897A76931AD29FE2C" box="[986,1132,367,393]" pageId="10" pageNumber="171" value="2022-07-11">
<collectingDate id="7301E9B66A1FFF8897A76931AD29FE2C" box="[986,1132,367,393]" pageId="10" pageNumber="171" value="2022-07-11">11 July 2022</collectingDate>
</date>
(
<collectionCode id="71EAAE5B6A1FFF8890016931ADB4FE2C" box="[1148,1265,367,393]" pageId="10" pageNumber="171">ZSI-WRC</collectionCode>
C.2294);
</materialsCitation>
<materialsCitation id="A7933CC36A1FFF8891196931AADFFE08" ID-GBIF-Occurrence="4109228306" pageId="10" pageNumber="171" specimenCount="2" specimenCount-adult="1" specimenCount-male="1">
<specimenCount id="01FDFD176A1FFF8891196931ACD9FE2C" box="[1380,1436,367,393]" pageId="10" pageNumber="171" type="adult">adult</specimenCount>
<specimenCount id="01FDFD176A1FFF8894EA69CDA988FE08" box="[151,205,403,429]" pageId="10" pageNumber="171" type="male">male</specimenCount>
(CW
<quantity id="D0039B7B6A1FFF88956969CDA8CCFE08" box="[276,393,403,429]" metricMagnitude="-2" metricUnit="m" metricValue="2.363" pageId="10" pageNumber="171" unit="mm" value="23.63">23.63 mm</quantity>
, CL
<quantity id="D0039B7B6A1FFF8895BC69CDAB73FE08" box="[449,566,403,429]" metricMagnitude="-2" metricUnit="m" metricValue="1.934" pageId="10" pageNumber="171" unit="mm" value="19.34">19.34 mm</quantity>
, CH
<quantity id="D0039B7B6A1FFF88960F69CCABA2FE08" box="[626,743,402,429]" metricMagnitude="-2" metricUnit="m" metricValue="1.567" pageId="10" pageNumber="171" unit="mm" value="15.67">15.67 mm</quantity>
, FW
<quantity id="D0039B7B6A1FFF88975B69CCAACAFE08" box="[806,911,402,429]" metricMagnitude="-3" metricUnit="m" metricValue="5.72" pageId="10" pageNumber="171" unit="mm" value="5.72">5.72 mm</quantity>
),
</materialsCitation>
<materialsCitation id="A7933CC36A1FFF8897DE69CDA803FE50" ID-GBIF-Occurrence="4109228309" collectingDate="2022-07-11" collectionCode="ZSI-WGRC" collectorName="G. D. Hegde" pageId="10" pageNumber="171" specimenCount="2" specimenCount-adult="1" specimenCount-female="1">
<specimenCount id="01FDFD176A1FFF8897DE69CDAA9EFE08" box="[931,987,403,429]" pageId="10" pageNumber="171" type="adult">adult</specimenCount>
<specimenCount id="01FDFD176A1FFF88979E69CDAD6AFE08" box="[995,1071,403,429]" pageId="10" pageNumber="171" type="female">female</specimenCount>
(CW
<quantity id="D0039B7B6A1FFF88900B69CCADAEFE08" box="[1142,1259,402,429]" metricMagnitude="-2" metricUnit="m" metricValue="2.351" pageId="10" pageNumber="171" unit="mm" value="23.51">23.51 mm</quantity>
, CL
<quantity id="D0039B7B6A1FFF88915F69CDACDDFE08" box="[1314,1432,403,429]" metricMagnitude="-2" metricUnit="m" metricValue="1.903" pageId="10" pageNumber="171" unit="mm" value="19.03">19.03 mm</quantity>
, CH
<quantity id="D0039B7B6A1FFF8894BB69E8A879FE74" box="[198,316,438,465]" metricMagnitude="-2" metricUnit="m" metricValue="1.625" pageId="10" pageNumber="171" unit="mm" value="16.25">16.25 mm</quantity>
, FW
<quantity id="D0039B7B6A1FFF88950769E8A8A6FE74" box="[378,483,438,465]" metricMagnitude="-3" metricUnit="m" metricValue="5.69" pageId="10" pageNumber="171" unit="mm" value="5.69">5.69 mm</quantity>
), same collection data as for holotype,
<date id="6345105E6A1FFF8897DC69E9AD73FE74" box="[929,1078,439,465]" pageId="10" pageNumber="171" value="2022-07-11">
<collectingDate id="7301E9B66A1FFF8897DC69E9AD73FE74" box="[929,1078,439,465]" pageId="10" pageNumber="171" value="2022-07-11">11 July 2022</collectingDate>
</date>
, coll.
<collectorName id="BA0E53486A1FFF88900669E9AC46FE74" box="[1147,1283,439,465]" pageId="10" pageNumber="171">G.D. Hegde</collectorName>
(
<collectionCode id="71EAAE5B6A1FFF88916E69E9ACD9FE74" box="[1299,1436,439,465]" pageId="10" pageNumber="171">ZSI-WGRC</collectionCode>
IR.INV.22823).
</materialsCitation>
</paragraph>
<paragraph id="1744369E6A1FFF8894BA69A0A805FD98" blockId="10.[151,1437,150,2013]" pageId="10" pageNumber="171">
<materialsCitation id="A7933CC36A1FFF8894BA69A0A805FD98" ID-GBIF-Occurrence="4109228307" collectionCode="ZSI-WRC" pageId="10" pageNumber="171" specimenCount="9" specimenCount-adult="4" specimenCount-female="1" specimenCount-male="1" specimenCount-subadult="3">
<emphasis id="258FEA8C6A1FFF8894BA69A0A8F9FDBD" bold="true" box="[199,444,510,536]" pageId="10" pageNumber="171">Additional material.</emphasis>
<specimenCount id="01FDFD176A1FFF8895B869A1AB7FFDBC" box="[453,570,511,537]" pageId="10" pageNumber="171" type="subadult">2 subadult</specimenCount>
<specimenCount id="01FDFD176A1FFF88963E69A1ABC2FDBC" box="[579,647,511,537]" pageId="10" pageNumber="171" type="male">males</specimenCount>
,
<specimenCount id="01FDFD176A1FFF8896E969A1ABA6FDBC" box="[660,739,511,537]" pageId="10" pageNumber="171" type="adult">4 adult</specimenCount>
/
<specimenCount id="01FDFD176A1FFF88969169A1AA0FFDBC" box="[748,842,511,537]" pageId="10" pageNumber="171" type="subadult">subadult</specimenCount>
<specimenCount id="01FDFD176A1FFF88972E69A1AAE8FDBC" box="[851,941,511,537]" pageId="10" pageNumber="171" type="female">females</specimenCount>
, same collection data as for holotype (
<collectionCode id="71EAAE5B6A1FFF88911A69A1A992FD98" pageId="10" pageNumber="171">ZSI-WRC</collectionCode>
C.2295).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="5FE165156A1FFF8894BA6A18AB56FDC4" box="[199,531,582,609]" pageId="10" pageNumber="171" type="diagnosis">
<paragraph id="1744369E6A1FFF8894BA6A18AB56FDC4" blockId="10.[151,1437,150,2013]" box="[199,531,582,609]" pageId="10" pageNumber="171">
<emphasis id="258FEA8C6A1FFF8894BA6A18A807FDC5" bold="true" box="[199,322,582,608]" pageId="10" pageNumber="171">Diagnosis.</emphasis>
As for new genus.
</paragraph>
</subSubSection>
<subSubSection id="5FE165156A1FFF8994BA6A34AA9DFC24" lastPageId="11" lastPageNumber="172" pageId="10" pageNumber="171" type="description">
<paragraph id="1744369E6A1FFF8894BA6A34AABAFB04" blockId="10.[151,1437,150,2013]" pageId="10" pageNumber="171">
<emphasis id="258FEA8C6A1FFF8894BA6A34AB73FD20" bold="true" box="[199,566,618,645]" pageId="10" pageNumber="171">
Description of male
<typeStatus id="C840883C6A1FFF8895B56A35AB74FD20" box="[456,561,619,645]" pageId="10" pageNumber="171" type="holotype">holotype</typeStatus>
.
</emphasis>
Carapace transversely ovate, broader than long (CW/CL = 1.3), strongly inflated, deep (
<collectingCountry id="6FEC760E6A1FFF8895396AD1A82EFD0C" box="[324,363,655,681]" name="Switzerland" pageId="10" pageNumber="171">CH</collectingCountry>
/CW = 0.6); dorsal surface generally smooth, glabrous, strongly arched; anterolateral surface strongly inflated in frontal view, smooth; posterolateral surface with distinct, oblique striations; anterolateral margins convex, cristate, lacking distinct serrations, shorter than posterolateral margins; posterolateral margins converging posteriorly, gently concave medially, joining gently concave posterior margin; front strongly deflexed, trapezoidal, with anterior margin narrow (FW/CW = 0.25), smooth, cristate, sinuous in frontal view; frontal medial triangle incomplete, with dorsal margin only, lateral margins indiscernible; epigastric cristae poorly developed, only visible as 2 low, slightly rugose, round protuberances; postorbital cristae poorly developed, only visible laterally as low, smooth protuberances; external orbital angle indistinct; epibranchial tooth very low, blunt; postorbital region flat, with strong posterolateral depression; branchial regions strongly inflated; cervical grooves visible as medial, shallow, rugose depressions, not reaching level of postorbital cristae; mesogastric groove shallow, narrow, short, bifurcated posteriorly; H-shaped groove poorly developed; subhepatic region rugose; suborbital region with low granules posteriorly; pterygostomial region with low granules anteriorly; supraorbital margin sinuous, cristate, smooth; suborbital margin concave, cristate with low granules, continuous with supraorbital margin; epistome with distinct gape between anterior margin and front exposing antennular fossae, posterior margin with distinct, triangular medial lobe and strongly sinuous lateral lobes (
<figureCitation id="8FC02A1B6A1FFF8897616CD9AAE4FB04" box="[796,929,1159,1185]" captionStart="FIGURE 2" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[192,1395,181,1919]" captionTargetId="figure-19@3.[192,1395,181,1919]" captionTargetPageId="3" captionText="FIGURE 2. Aradhya placida gen. et sp. nov., holotype male (25.12 × 19.85 mm) (ZSI-WRC C.2292). A, overall dorsal view; B, frontal view of cephalothorax; C, overall ventral view. Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.7935895" httpUri="https://zenodo.org/record/7935895/files/figure.png" pageId="10" pageNumber="171">Figs. 2AC</figureCitation>
,
<figureCitation id="8FC02A1B6A1FFF8897D16CD8AAB4FB04" box="[940,1009,1158,1185]" captionStart="FIGURE 5" captionStartId="6.[151,250,1943,1967]" captionTargetBox="[203,1384,181,1919]" captionTargetId="figure-24@6.[203,1384,181,1919]" captionTargetPageId="6" captionText="FIGURE 5. Aradhya placida gen. et sp. nov., holotype male (25.12 × 19.85 mm) (ZSI-WRC C.2292).A, mouth parts exposed; B, third maxillipeds; C, thoracic sternites with G1 and G2; D, pleon and telson; E, dorsal view of left G1; F, left G2. Scale bars = 5 mm (C, D), 2 mm (A, B, E), 1 mm (F)." figureDoi="http://doi.org/10.5281/zenodo.7935903" httpUri="https://zenodo.org/record/7935903/files/figure.png" pageId="10" pageNumber="171">5A, B</figureCitation>
).
</paragraph>
<paragraph id="1744369E6A1FFF8894BA6CF5A840FB4C" blockId="10.[151,1437,150,2013]" pageId="10" pageNumber="171">
Eyes small as compared to orbits; eyestalk short, moderately stout, concave medially; cornea small, pigmented (
<figureCitation id="8FC02A1B6A1FFF8894E26C91A9BDFB4C" box="[159,248,1231,1257]" captionStart="FIGURE 2" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[192,1395,181,1919]" captionTargetId="figure-19@3.[192,1395,181,1919]" captionTargetPageId="3" captionText="FIGURE 2. Aradhya placida gen. et sp. nov., holotype male (25.12 × 19.85 mm) (ZSI-WRC C.2292). A, overall dorsal view; B, frontal view of cephalothorax; C, overall ventral view. Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.7935895" httpUri="https://zenodo.org/record/7935895/files/figure.png" pageId="10" pageNumber="171">Fig. 2B</figureCitation>
).
</paragraph>
<paragraph id="1744369E6A1FFF8894BA6CADA840FA40" blockId="10.[151,1437,150,2013]" pageId="10" pageNumber="171">
Antennules short, folded in transversely broad fossae; antennae with flagellum shorter than eyestalks (
<figureCitation id="8FC02A1B6A1FFF8891456CADACCAFAA8" box="[1336,1423,1267,1293]" captionStart="FIGURE 2" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[192,1395,181,1919]" captionTargetId="figure-19@3.[192,1395,181,1919]" captionTargetPageId="3" captionText="FIGURE 2. Aradhya placida gen. et sp. nov., holotype male (25.12 × 19.85 mm) (ZSI-WRC C.2292). A, overall dorsal view; B, frontal view of cephalothorax; C, overall ventral view. Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.7935895" httpUri="https://zenodo.org/record/7935895/files/figure.png" pageId="10" pageNumber="171">Fig. 2B</figureCitation>
). Mandibular palp with 2 articles; terminal article bilobed, anterior lobe narrow, long, posterior lobe broad, short, nearly half length of anterior lobe (
<figureCitation id="8FC02A1B6A1FFF8896576D65ABC2FAF0" box="[554,647,1338,1365]" captionStart="FIGURE 5" captionStartId="6.[151,250,1943,1967]" captionTargetBox="[203,1384,181,1919]" captionTargetId="figure-24@6.[203,1384,181,1919]" captionTargetPageId="6" captionText="FIGURE 5. Aradhya placida gen. et sp. nov., holotype male (25.12 × 19.85 mm) (ZSI-WRC C.2292).A, mouth parts exposed; B, third maxillipeds; C, thoracic sternites with G1 and G2; D, pleon and telson; E, dorsal view of left G1; F, left G2. Scale bars = 5 mm (C, D), 2 mm (A, B, E), 1 mm (F)." figureDoi="http://doi.org/10.5281/zenodo.7935903" httpUri="https://zenodo.org/record/7935903/files/figure.png" pageId="10" pageNumber="171">Fig. 5A</figureCitation>
). First, second maxillipeds each with long flagellum on exopod (
<figureCitation id="8FC02A1B6A1FFF8891126D65A9F8FADC" captionStart="FIGURE 5" captionStartId="6.[151,250,1943,1967]" captionTargetBox="[203,1384,181,1919]" captionTargetId="figure-24@6.[203,1384,181,1919]" captionTargetPageId="6" captionText="FIGURE 5. Aradhya placida gen. et sp. nov., holotype male (25.12 × 19.85 mm) (ZSI-WRC C.2292).A, mouth parts exposed; B, third maxillipeds; C, thoracic sternites with G1 and G2; D, pleon and telson; E, dorsal view of left G1; F, left G2. Scale bars = 5 mm (C, D), 2 mm (A, B, E), 1 mm (F)." figureDoi="http://doi.org/10.5281/zenodo.7935903" httpUri="https://zenodo.org/record/7935903/files/figure.png" pageId="10" pageNumber="171">Fig. 5A</figureCitation>
). Third maxillipeds cover most of buccal cavity when closed; ischium subrectangular, longer than broad, with deep, oblique submedial groove; merus subquadrate, slightly broader than long, sunken on inner side, anterolateral margin rounded; exopod slender, distally narrow, reaching beyond anterolateral corner of ischium, lacking flagellum (
<figureCitation id="8FC02A1B6A1FFF8894E26D95A9BDFA40" box="[159,248,1482,1509]" captionStart="FIGURE 5" captionStartId="6.[151,250,1943,1967]" captionTargetBox="[203,1384,181,1919]" captionTargetId="figure-24@6.[203,1384,181,1919]" captionTargetPageId="6" captionText="FIGURE 5. Aradhya placida gen. et sp. nov., holotype male (25.12 × 19.85 mm) (ZSI-WRC C.2292).A, mouth parts exposed; B, third maxillipeds; C, thoracic sternites with G1 and G2; D, pleon and telson; E, dorsal view of left G1; F, left G2. Scale bars = 5 mm (C, D), 2 mm (A, B, E), 1 mm (F)." figureDoi="http://doi.org/10.5281/zenodo.7935903" httpUri="https://zenodo.org/record/7935903/files/figure.png" pageId="10" pageNumber="171">Fig. 5B</figureCitation>
).
</paragraph>
<paragraph id="1744369E6A1FFF8894BA6DB1A89EF918" blockId="10.[151,1437,150,2013]" pageId="10" pageNumber="171">
Chelipeds generally smooth, longer than carapace, unequal, left chela larger, sparsely punctate (
<figureCitation id="8FC02A1B6A1FFF88916D6DB1ACCAF9AC" box="[1296,1423,1519,1545]" captionStart="FIGURE 2" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[192,1395,181,1919]" captionTargetId="figure-19@3.[192,1395,181,1919]" captionTargetPageId="3" captionText="FIGURE 2. Aradhya placida gen. et sp. nov., holotype male (25.12 × 19.85 mm) (ZSI-WRC C.2292). A, overall dorsal view; B, frontal view of cephalothorax; C, overall ventral view. Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.7935895" httpUri="https://zenodo.org/record/7935895/files/figure.png" pageId="10" pageNumber="171">Fig. 2A, C</figureCitation>
). Major chela with merus/basis-ischium joint distinct, structures moveable; merus rugose on outer surface, lacking subterminal spine, ventral margins tuberculated; carpus generally smooth, gently inflated, inner distal major tooth short, broad, blunt, triangular, sub-basal tooth small, low, blunt; palm as long as high, generally smooth; dactylus gently curved, slender, longer than upper margin of palm, generally smooth; fingers with distinct gape when closed, teeth low, blunt (
<figureCitation id="8FC02A1B6A1FFF8895296EFDA88BF918" box="[340,462,1699,1725]" captionStart="FIGURE 2" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[192,1395,181,1919]" captionTargetId="figure-19@3.[192,1395,181,1919]" captionTargetPageId="3" captionText="FIGURE 2. Aradhya placida gen. et sp. nov., holotype male (25.12 × 19.85 mm) (ZSI-WRC C.2292). A, overall dorsal view; B, frontal view of cephalothorax; C, overall ventral view. Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.7935895" httpUri="https://zenodo.org/record/7935895/files/figure.png" pageId="10" pageNumber="171">Fig. 2A, C</figureCitation>
).
</paragraph>
<paragraph id="1744369E6A1FFF8894BA6E99A9AFF830" blockId="10.[151,1437,150,2013]" pageId="10" pageNumber="171">
Ambulatory legs slender, long, P3 longest, ca. 2.0× CL, longer than minor cheliped; merus/basis-ischium joint (P2P5) distinct, structures moveable; merus (P2P5) elongated, equal in length to combined length of carpus and propodus, slightly rugose on outer surface, lacking subdistal spine; carpus (P2P4) with medial crista on outer surface; propodus (P2P5) with small, sharp chitinous spines on anterior and posterior margins; dactylus (P2P5) recurved, longer than propodus, with short, sharp chitinous spines on margins, distal chitinous part very short (
<figureCitation id="8FC02A1B6A1FFF8891126F09A998F830" captionStart="FIGURE 2" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[192,1395,181,1919]" captionTargetId="figure-19@3.[192,1395,181,1919]" captionTargetPageId="3" captionText="FIGURE 2. Aradhya placida gen. et sp. nov., holotype male (25.12 × 19.85 mm) (ZSI-WRC C.2292). A, overall dorsal view; B, frontal view of cephalothorax; C, overall ventral view. Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.7935895" httpUri="https://zenodo.org/record/7935895/files/figure.png" pageId="10" pageNumber="171">Fig. 2A, C</figureCitation>
).
</paragraph>
<paragraph id="1744369E6A1FFF8994BA6FC1ABACFEB8" blockId="10.[151,1437,150,2013]" lastBlockId="11.[151,1437,150,897]" lastPageId="11" lastPageNumber="172" pageId="10" pageNumber="171">
Thoracic sternites punctate, setose anteriorly, glabrous posteriorly; s1 and s2 separated by cristate margin of s2; s2/s3 cristate, incomplete, not reaching edge of sternum; s3/s4 deep, broad, complete, reaching edge of sternum; s4/s5, s5/s6, s6/s7, s7/s8 shallow, narrow, medially interrupted; s7 with longitudinal medial groove; s8 completely covered by pleon, broad medially, with longitudinal medial groove (
<figureCitation id="8FC02A1B6A1EFF8997F168E5AAABFF70" box="[908,1006,187,213]" captionStart="FIGURE 2" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[192,1395,181,1919]" captionTargetId="figure-19@3.[192,1395,181,1919]" captionTargetPageId="3" captionText="FIGURE 2. Aradhya placida gen. et sp. nov., holotype male (25.12 × 19.85 mm) (ZSI-WRC C.2292). A, overall dorsal view; B, frontal view of cephalothorax; C, overall ventral view. Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.7935895" httpUri="https://zenodo.org/record/7935895/files/figure.png" pageId="11" pageNumber="172">Figs. 2C</figureCitation>
,
<figureCitation id="8FC02A1B6A1EFF89978468E4AD58FF70" box="[1017,1053,186,213]" captionStart="FIGURE 5" captionStartId="6.[151,250,1943,1967]" captionTargetBox="[203,1384,181,1919]" captionTargetId="figure-24@6.[203,1384,181,1919]" captionTargetPageId="6" captionText="FIGURE 5. Aradhya placida gen. et sp. nov., holotype male (25.12 × 19.85 mm) (ZSI-WRC C.2292).A, mouth parts exposed; B, third maxillipeds; C, thoracic sternites with G1 and G2; D, pleon and telson; E, dorsal view of left G1; F, left G2. Scale bars = 5 mm (C, D), 2 mm (A, B, E), 1 mm (F)." figureDoi="http://doi.org/10.5281/zenodo.7935903" httpUri="https://zenodo.org/record/7935903/files/figure.png" pageId="11" pageNumber="172">5C</figureCitation>
). Pleonal locking mechanism with low tubercles on submedial part of s5 (
<figureCitation id="8FC02A1B6A1EFF8996246881ABF1FF5C" box="[601,692,222,249]" captionStart="FIGURE 5" captionStartId="6.[151,250,1943,1967]" captionTargetBox="[203,1384,181,1919]" captionTargetId="figure-24@6.[203,1384,181,1919]" captionTargetPageId="6" captionText="FIGURE 5. Aradhya placida gen. et sp. nov., holotype male (25.12 × 19.85 mm) (ZSI-WRC C.2292).A, mouth parts exposed; B, third maxillipeds; C, thoracic sternites with G1 and G2; D, pleon and telson; E, dorsal view of left G1; F, left G2. Scale bars = 5 mm (C, D), 2 mm (A, B, E), 1 mm (F)." figureDoi="http://doi.org/10.5281/zenodo.7935903" httpUri="https://zenodo.org/record/7935903/files/figure.png" pageId="11" pageNumber="172">Fig. 5C</figureCitation>
). Sternopleonal cavity deep, narrow, short, reaching anteriorly to level of mid-length of cheliped coxae (
<figureCitation id="8FC02A1B6A1EFF899637695DABE9FEB8" box="[586,684,259,285]" captionStart="FIGURE 2" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[192,1395,181,1919]" captionTargetId="figure-19@3.[192,1395,181,1919]" captionTargetPageId="3" captionText="FIGURE 2. Aradhya placida gen. et sp. nov., holotype male (25.12 × 19.85 mm) (ZSI-WRC C.2292). A, overall dorsal view; B, frontal view of cephalothorax; C, overall ventral view. Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.7935895" httpUri="https://zenodo.org/record/7935895/files/figure.png" pageId="11" pageNumber="172">Figs. 2C</figureCitation>
,
<figureCitation id="8FC02A1B6A1EFF8996C5695CAB99FEB8" box="[696,732,258,285]" captionStart="FIGURE 5" captionStartId="6.[151,250,1943,1967]" captionTargetBox="[203,1384,181,1919]" captionTargetId="figure-24@6.[203,1384,181,1919]" captionTargetPageId="6" captionText="FIGURE 5. Aradhya placida gen. et sp. nov., holotype male (25.12 × 19.85 mm) (ZSI-WRC C.2292).A, mouth parts exposed; B, third maxillipeds; C, thoracic sternites with G1 and G2; D, pleon and telson; E, dorsal view of left G1; F, left G2. Scale bars = 5 mm (C, D), 2 mm (A, B, E), 1 mm (F)." figureDoi="http://doi.org/10.5281/zenodo.7935903" httpUri="https://zenodo.org/record/7935903/files/figure.png" pageId="11" pageNumber="172">5C</figureCitation>
).
</paragraph>
<paragraph id="1744369E6A1EFF8994BA6979A9BCFD98" blockId="11.[151,1437,150,897]" pageId="11" pageNumber="172">
Pleon narrow, triangular, with concave lateral margins; pleonal somites 16 progressively longer; pleonal somite 1 rectangular, shortest, narrower than pleonal somite 2; pleonal somite 2 trapezoidal, narrower than pleonal somite 3; pleonal somite 3 trapezoidal, broadest among pleonal somites, with strongly convex lateral margins; pleonal somite 4 trapezoidal, with gently concave lateral margins; pleonal somite 5 trapezoidal, with concave lateral margins; pleonal somite 6 trapezoidal, slightly broader than long (proximal width ca. 1.3× medial length), longest among pleonal somites, shorter than telson, with gently concave lateral margins (
<figureCitation id="8FC02A1B6A1EFF8990396985ADECFE50" box="[1092,1193,475,501]" captionStart="FIGURE 2" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[192,1395,181,1919]" captionTargetId="figure-19@3.[192,1395,181,1919]" captionTargetPageId="3" captionText="FIGURE 2. Aradhya placida gen. et sp. nov., holotype male (25.12 × 19.85 mm) (ZSI-WRC C.2292). A, overall dorsal view; B, frontal view of cephalothorax; C, overall ventral view. Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.7935895" httpUri="https://zenodo.org/record/7935895/files/figure.png" pageId="11" pageNumber="172">Figs. 2C</figureCitation>
,
<figureCitation id="8FC02A1B6A1EFF8990CA6984AD98FE50" box="[1207,1245,474,501]" captionStart="FIGURE 5" captionStartId="6.[151,250,1943,1967]" captionTargetBox="[203,1384,181,1919]" captionTargetId="figure-24@6.[203,1384,181,1919]" captionTargetPageId="6" captionText="FIGURE 5. Aradhya placida gen. et sp. nov., holotype male (25.12 × 19.85 mm) (ZSI-WRC C.2292).A, mouth parts exposed; B, third maxillipeds; C, thoracic sternites with G1 and G2; D, pleon and telson; E, dorsal view of left G1; F, left G2. Scale bars = 5 mm (C, D), 2 mm (A, B, E), 1 mm (F)." figureDoi="http://doi.org/10.5281/zenodo.7935903" httpUri="https://zenodo.org/record/7935903/files/figure.png" pageId="11" pageNumber="172">5D</figureCitation>
). Telson narrow, elongated (medial length ca. 1.4× proximal width), with gently concave lateral margins, apex broad, rounded (
<figureCitation id="8FC02A1B6A1EFF89911969A1A9FFFD98" captionStart="FIGURE 2" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[192,1395,181,1919]" captionTargetId="figure-19@3.[192,1395,181,1919]" captionTargetPageId="3" captionText="FIGURE 2. Aradhya placida gen. et sp. nov., holotype male (25.12 × 19.85 mm) (ZSI-WRC C.2292). A, overall dorsal view; B, frontal view of cephalothorax; C, overall ventral view. Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.7935895" httpUri="https://zenodo.org/record/7935895/files/figure.png" pageId="11" pageNumber="172">Figs. 2C</figureCitation>
,
<figureCitation id="8FC02A1B6A1EFF8994BB6A7CA9AEFD98" box="[198,235,546,573]" captionStart="FIGURE 5" captionStartId="6.[151,250,1943,1967]" captionTargetBox="[203,1384,181,1919]" captionTargetId="figure-24@6.[203,1384,181,1919]" captionTargetPageId="6" captionText="FIGURE 5. Aradhya placida gen. et sp. nov., holotype male (25.12 × 19.85 mm) (ZSI-WRC C.2292).A, mouth parts exposed; B, third maxillipeds; C, thoracic sternites with G1 and G2; D, pleon and telson; E, dorsal view of left G1; F, left G2. Scale bars = 5 mm (C, D), 2 mm (A, B, E), 1 mm (F)." figureDoi="http://doi.org/10.5281/zenodo.7935903" httpUri="https://zenodo.org/record/7935903/files/figure.png" pageId="11" pageNumber="172">5D</figureCitation>
).
</paragraph>
<paragraph id="1744369E6A1EFF8994BA6A19ABCCFC9C" blockId="11.[151,1437,150,897]" pageId="11" pageNumber="172">
G1 moderately stout, long, tip reaching pleonal locking structure
<emphasis id="258FEA8C6A1EFF8997F76A19AA89FDC5" box="[906,972,583,608]" italics="true" pageId="11" pageNumber="172">in situ</emphasis>
; flexible zone small; ultimate article stout, conical, short, ca. 0.3× length of penultimate article, gently curved outwards at angle of ca. 15° from longitudinal axis, tip broad, tubular, dorsal flap absent; penultimate article stout, broad at proximal half, abruptly narrow at distal half, outer margin strongly convex, shelf-like at basal half, inner margin characteristically convex distally; groove for G2 marginal (
<figureCitation id="8FC02A1B6A1EFF8995266A89A8F9FD54" box="[347,444,726,753]" captionStart="FIGURE 5" captionStartId="6.[151,250,1943,1967]" captionTargetBox="[203,1384,181,1919]" captionTargetId="figure-24@6.[203,1384,181,1919]" captionTargetPageId="6" captionText="FIGURE 5. Aradhya placida gen. et sp. nov., holotype male (25.12 × 19.85 mm) (ZSI-WRC C.2292).A, mouth parts exposed; B, third maxillipeds; C, thoracic sternites with G1 and G2; D, pleon and telson; E, dorsal view of left G1; F, left G2. Scale bars = 5 mm (C, D), 2 mm (A, B, E), 1 mm (F)." figureDoi="http://doi.org/10.5281/zenodo.7935903" httpUri="https://zenodo.org/record/7935903/files/figure.png" pageId="11" pageNumber="172">Figs. 5E</figureCitation>
,
<figureCitation id="8FC02A1B6A1EFF8995B56A89AB49FD54" box="[456,524,727,753]" captionStart="FIGURE 6" captionStartId="7.[151,250,1352,1376]" captionTargetBox="[153,1434,419,1327]" captionTargetId="figure-183@7.[151,1436,417,1329]" captionTargetPageId="7" captionText="FIGURE 6. Aradhya placida gen. et sp. nov., holotype male (25.12 × 19.85 mm) (ZSI-WRC C.2292). A, pleon and telson; B, dorsal view of left G1; C, dorsal view of left G1 ultimate article; D, ventral view of left G1; E, ventral view of left G1 ultimate article; F, left G2. Scale bars = 5 mm (A), 2 mm (B, D), 1 mm (C, E, F)." figureDoi="http://doi.org/10.5281/zenodo.7935907" httpUri="https://zenodo.org/record/7935907/files/figure.png" pageId="11" pageNumber="172">6BE</figureCitation>
). G2 distinctly shorter than G1, ca. 0.6× length of G1; ultimate article very short, ca. 0.2× length of penultimate article, straight, narrow, with acute tip; penultimate article much stouter at proximal third, appearing subtriangular (
<figureCitation id="8FC02A1B6A1EFF89958E6B41AB15FC9C" box="[499,592,798,825]" captionStart="FIGURE 5" captionStartId="6.[151,250,1943,1967]" captionTargetBox="[203,1384,181,1919]" captionTargetId="figure-24@6.[203,1384,181,1919]" captionTargetPageId="6" captionText="FIGURE 5. Aradhya placida gen. et sp. nov., holotype male (25.12 × 19.85 mm) (ZSI-WRC C.2292).A, mouth parts exposed; B, third maxillipeds; C, thoracic sternites with G1 and G2; D, pleon and telson; E, dorsal view of left G1; F, left G2. Scale bars = 5 mm (C, D), 2 mm (A, B, E), 1 mm (F)." figureDoi="http://doi.org/10.5281/zenodo.7935903" httpUri="https://zenodo.org/record/7935903/files/figure.png" pageId="11" pageNumber="172">Figs. 5F</figureCitation>
,
<figureCitation id="8FC02A1B6A1EFF8996266B41AB39FC9C" box="[603,636,799,825]" captionStart="FIGURE 6" captionStartId="7.[151,250,1352,1376]" captionTargetBox="[153,1434,419,1327]" captionTargetId="figure-183@7.[151,1436,417,1329]" captionTargetPageId="7" captionText="FIGURE 6. Aradhya placida gen. et sp. nov., holotype male (25.12 × 19.85 mm) (ZSI-WRC C.2292). A, pleon and telson; B, dorsal view of left G1; C, dorsal view of left G1 ultimate article; D, ventral view of left G1; E, ventral view of left G1 ultimate article; F, left G2. Scale bars = 5 mm (A), 2 mm (B, D), 1 mm (C, E, F)." figureDoi="http://doi.org/10.5281/zenodo.7935907" httpUri="https://zenodo.org/record/7935907/files/figure.png" pageId="11" pageNumber="172">6F</figureCitation>
).
</paragraph>
<paragraph id="1744369E6A1EFF8994BA6B1CAA9DFC24" blockId="11.[151,1437,150,897]" pageId="11" pageNumber="172">
<emphasis id="258FEA8C6A1EFF8994BA6B1CA830FCF9" bold="true" box="[199,373,834,860]" pageId="11" pageNumber="172">Colour in life.</emphasis>
Crabs have a dark brown dorsal surface of the carapace (
<figureCitation id="8FC02A1B6A1EFF89905E6B1DAD88FCF8" box="[1059,1229,835,861]" captionStart="FIGURE 7" captionStartId="11.[151,250,1928,1952]" captionTargetBox="[151,1435,921,1905]" captionTargetId="figure-329@11.[151,1435,921,1905]" captionTargetPageId="11" captionText="FIGURE 7. Colour in life and habitats of Aradhya placida gen. et sp. nov. A, holotype male (25.12 × 19.85 mm) (ZSI-WRC C.2292); B, paratype male (23.09 × 18.23 mm) (ZSI-WRC C.2293); C, a live crab inside the burrow; D, a live crab near the base of a tree; E, trees and boulders at the type locality. This figure is available in colour at Zootaxa online." figureDoi="http://doi.org/10.5281/zenodo.7935910" httpUri="https://zenodo.org/record/7935910/files/figure.png" pageId="11" pageNumber="172">Fig. 7A, B, D</figureCitation>
). Their chelipeds, ambulatory legs and ventral surface of the carapace are paler (
<figureCitation id="8FC02A1B6A1EFF8997326B39AA89FC24" box="[847,972,871,897]" captionStart="FIGURE 7" captionStartId="11.[151,250,1928,1952]" captionTargetBox="[151,1435,921,1905]" captionTargetId="figure-329@11.[151,1435,921,1905]" captionTargetPageId="11" captionText="FIGURE 7. Colour in life and habitats of Aradhya placida gen. et sp. nov. A, holotype male (25.12 × 19.85 mm) (ZSI-WRC C.2292); B, paratype male (23.09 × 18.23 mm) (ZSI-WRC C.2293); C, a live crab inside the burrow; D, a live crab near the base of a tree; E, trees and boulders at the type locality. This figure is available in colour at Zootaxa online." figureDoi="http://doi.org/10.5281/zenodo.7935910" httpUri="https://zenodo.org/record/7935910/files/figure.png" pageId="11" pageNumber="172">Fig. 7AD</figureCitation>
).
</paragraph>
</subSubSection>
<caption id="438466166A1EFF8994EA6FD6ADE7F84D" ID-DOI="http://doi.org/10.5281/zenodo.7935910" ID-Zenodo-Dep="7935910" httpUri="https://zenodo.org/record/7935910/files/figure.png" pageId="11" pageNumber="172" startId="11.[151,250,1928,1952]" targetBox="[151,1435,921,1905]" targetPageId="11" targetType="figure">
<paragraph id="1744369E6A1EFF8994EA6FD6ADE7F84D" blockId="11.[151,1436,1928,2025]" pageId="11" pageNumber="172">
<emphasis id="258FEA8C6A1EFF8994EA6FD6A851F805" bold="true" box="[151,276,1928,1952]" pageId="11" pageNumber="172">FIGURE 7.</emphasis>
Colour in life and habitats of
<taxonomicName id="D0FB4D1D6A1EFF8996346FD6ABB6F805" authority="Pati &amp; Bajantri &amp; Hegde, 2023" authorityName="Pati &amp; Bajantri &amp; Hegde" authorityYear="2023" box="[585,755,1928,1952]" class="Malacostraca" family="Gecarcinucidae" genus="Aradhya" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="172" phylum="Arthropoda" rank="species" species="placida" status="gen. et sp. nov.">
<emphasis id="258FEA8C6A1EFF8996346FD6ABB6F805" box="[585,755,1928,1952]" italics="true" pageId="11" pageNumber="172">Aradhya placida</emphasis>
</taxonomicName>
<emphasis id="258FEA8C6A1EFF8996876FD6AADCF805" bold="true" box="[762,921,1928,1952]" pageId="11" pageNumber="172">
<taxonomicNameLabel id="3EBC57F76A1EFF8996876FD6AADCF805" box="[762,921,1928,1952]" pageId="11" pageNumber="172" rank="species">gen. et sp. nov.</taxonomicNameLabel>
</emphasis>
A, holotype male (25.12 × 19.85 mm) (ZSI-WRC C.2292); B, paratype male (23.09 × 18.23 mm) (ZSI-WRC C.2293); C, a live crab inside the burrow; D, a live crab near the base of a tree; E, trees and boulders at the type locality. This figure is available in colour at
<emphasis id="258FEA8C6A1EFF89907F6F8FAD10F84D" box="[1026,1109,2001,2024]" italics="true" pageId="11" pageNumber="172">Zootaxa</emphasis>
online.
</paragraph>
</caption>
<subSubSection id="5FE165156A19FF8F94BA68C9A99FFF5C" lastPageId="13" lastPageNumber="174" pageId="12" pageNumber="173" type="etymology">
<paragraph id="1744369E6A19FF8E94BA68C9AD18FF70" blockId="12.[151,1437,151,2013]" pageId="12" pageNumber="173">
<emphasis id="258FEA8C6A19FF8E94BA68C9A80BFF14" bold="true" box="[199,334,151,177]" pageId="12" pageNumber="173">Etymology.</emphasis>
The species epithet is from the Latin for calm or peaceful, which alludes to the behaviour of these crabs. The species name is treated here as a Latin adjective in the nominative singular.
</paragraph>
<paragraph id="1744369E6A19FF8E94BA6881A847FEB8" blockId="12.[151,1437,151,2013]" pageId="12" pageNumber="173">The proposed common name of the new species is “Peaceful Crab” in English and “Shantiyuta Edi” in Kannada.</paragraph>
<paragraph id="1744369E6A19FF8E94BA6979ABAFFC9C" blockId="12.[151,1437,151,2013]" pageId="12" pageNumber="173">
<emphasis id="258FEA8C6A19FF8E94BA6979A803FEE4" bold="true" box="[199,326,295,321]" pageId="12" pageNumber="173">
<typeStatus id="C840883C6A19FF8E94BA6979A804FEE4" box="[199,321,295,321]" pageId="12" pageNumber="173" type="paratype">Paratypes</typeStatus>
.
</emphasis>
The
<typeStatus id="C840883C6A19FF8E95016979A899FEE4" box="[380,476,295,321]" pageId="12" pageNumber="173" type="paratype">paratype</typeStatus>
males of
<taxonomicName id="D0FB4D1D6A19FF8E963B6979AA45FEE4" authority="Pati &amp; Bajantri &amp; Hegde, 2023" authorityName="Pati &amp; Bajantri &amp; Hegde" authorityYear="2023" box="[582,768,295,321]" class="Malacostraca" family="Gecarcinucidae" genus="Aradhya" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="173" phylum="Arthropoda" rank="species" species="placida" status="sp. nov.">
<emphasis id="258FEA8C6A19FF8E963B6979AA45FEE4" box="[582,768,295,321]" italics="true" pageId="12" pageNumber="173">Aradhya placida</emphasis>
</taxonomicName>
<emphasis id="258FEA8C6A19FF8E977B6979AA1BFEE4" bold="true" box="[774,862,295,321]" pageId="12" pageNumber="173">
<taxonomicNameLabel id="3EBC57F76A19FF8E977B6979AA1BFEE4" box="[774,862,295,321]" pageId="12" pageNumber="173" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
possess most of the character states as those present in the
<typeStatus id="C840883C6A19FF8E94A26915A804FEC0" box="[223,321,331,357]" pageId="12" pageNumber="173" type="holotype">holotype</typeStatus>
except for the following differences. One of the male
<typeStatus id="C840883C6A19FF8E97D56915AD56FEC0" box="[936,1043,331,357]" pageId="12" pageNumber="173" type="paratype">paratypes</typeStatus>
(ZSI-WGRC IR.INV.22823) has a proportionally narrower carapace, CW/CL = 1.2 (
<figureCitation id="8FC02A1B6A19FF8E96C16931AA54FE2C" box="[700,785,367,393]" captionStart="FIGURE 3" captionStartId="4.[151,250,1858,1882]" captionTargetBox="[151,1435,181,1834]" captionTargetId="figure-24@4.[151,1435,181,1834]" captionTargetPageId="4" captionText="FIGURE 3. Aradhya placida gen. et sp. nov.: paratype male (28.53 × 21.82 mm) (ZSI-WRC C.2294) (A, B); paratype male (23.63 × 19.34 mm) (ZSI-WGRC IR.INV.22823) (C, D); paratype male (23.09 × 18.23 mm) (ZSI-WRC C.2293) (EI). A, C, E, overall dorsal view; B, D, F, overall ventral view; G, third maxillipeds; H, dorsal view of left G1; I, left G2. Scale bars = 10 mm (AF), 2 mm (G, H), 1 mm (I)." figureDoi="http://doi.org/10.5281/zenodo.7935897" httpUri="https://zenodo.org/record/7935897/files/figure.png" pageId="12" pageNumber="173">Fig. 3C</figureCitation>
) (vs. CW/CL =
<quantity id="D0039B7B6A19FF8E97BF6931AD44FE2C" box="[962,1025,367,393]" metricMagnitude="-2" metricUnit="m" metricValue="3.302" pageId="12" pageNumber="173" unit="in" value="1.3">1.3 in</quantity>
the
<typeStatus id="C840883C6A19FF8E90526931ADD4FE2C" box="[1071,1169,367,393]" pageId="12" pageNumber="173" type="holotype">holotype</typeStatus>
and the remaining male
<typeStatus id="C840883C6A19FF8E94EA69CDA843FE08" box="[151,262,403,429]" pageId="12" pageNumber="173" type="paratype">paratypes</typeStatus>
;
<figureCitation id="8FC02A1B6A19FF8E956E69CDA832FE08" box="[275,375,403,429]" captionStart="FIGURE 2" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[192,1395,181,1919]" captionTargetId="figure-19@3.[192,1395,181,1919]" captionTargetPageId="3" captionText="FIGURE 2. Aradhya placida gen. et sp. nov., holotype male (25.12 × 19.85 mm) (ZSI-WRC C.2292). A, overall dorsal view; B, frontal view of cephalothorax; C, overall ventral view. Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.7935895" httpUri="https://zenodo.org/record/7935895/files/figure.png" pageId="12" pageNumber="173">Figs. 2A</figureCitation>
,
<figureCitation id="8FC02A1B6A19FF8E95F969CDA88EFE08" box="[388,459,403,429]" captionStart="FIGURE 3" captionStartId="4.[151,250,1858,1882]" captionTargetBox="[151,1435,181,1834]" captionTargetId="figure-24@4.[151,1435,181,1834]" captionTargetPageId="4" captionText="FIGURE 3. Aradhya placida gen. et sp. nov.: paratype male (28.53 × 21.82 mm) (ZSI-WRC C.2294) (A, B); paratype male (23.63 × 19.34 mm) (ZSI-WGRC IR.INV.22823) (C, D); paratype male (23.09 × 18.23 mm) (ZSI-WRC C.2293) (EI). A, C, E, overall dorsal view; B, D, F, overall ventral view; G, third maxillipeds; H, dorsal view of left G1; I, left G2. Scale bars = 10 mm (AF), 2 mm (G, H), 1 mm (I)." figureDoi="http://doi.org/10.5281/zenodo.7935897" httpUri="https://zenodo.org/record/7935897/files/figure.png" pageId="12" pageNumber="173">3A, E</figureCitation>
). The cervical grooves of the smaller male
<typeStatus id="C840883C6A19FF8E97C769CDAD60FE08" box="[954,1061,403,429]" pageId="12" pageNumber="173" type="paratype">paratypes</typeStatus>
(ZSI-WRC C.2293; ZSI-WGRC IR.INV.22823) are visible as smooth depressions (
<figureCitation id="8FC02A1B6A19FF8E96B569E9AA79FE74" box="[712,828,439,465]" captionStart="FIGURE 3" captionStartId="4.[151,250,1858,1882]" captionTargetBox="[151,1435,181,1834]" captionTargetId="figure-24@4.[151,1435,181,1834]" captionTargetPageId="4" captionText="FIGURE 3. Aradhya placida gen. et sp. nov.: paratype male (28.53 × 21.82 mm) (ZSI-WRC C.2294) (A, B); paratype male (23.63 × 19.34 mm) (ZSI-WGRC IR.INV.22823) (C, D); paratype male (23.09 × 18.23 mm) (ZSI-WRC C.2293) (EI). A, C, E, overall dorsal view; B, D, F, overall ventral view; G, third maxillipeds; H, dorsal view of left G1; I, left G2. Scale bars = 10 mm (AF), 2 mm (G, H), 1 mm (I)." figureDoi="http://doi.org/10.5281/zenodo.7935897" httpUri="https://zenodo.org/record/7935897/files/figure.png" pageId="12" pageNumber="173">Fig. 3C, E</figureCitation>
) as compared to the rugose depressions of the cervical grooves of the
<typeStatus id="C840883C6A19FF8E95426985A8E4FE50" box="[319,417,475,501]" pageId="12" pageNumber="173" type="holotype">holotype</typeStatus>
and the largest
<typeStatus id="C840883C6A19FF8E962F6985ABF7FE50" box="[594,690,475,501]" pageId="12" pageNumber="173" type="paratype">paratype</typeStatus>
male (
<figureCitation id="8FC02A1B6A19FF8E96826985AA26FE50" box="[767,867,475,501]" captionStart="FIGURE 2" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[192,1395,181,1919]" captionTargetId="figure-19@3.[192,1395,181,1919]" captionTargetPageId="3" captionText="FIGURE 2. Aradhya placida gen. et sp. nov., holotype male (25.12 × 19.85 mm) (ZSI-WRC C.2292). A, overall dorsal view; B, frontal view of cephalothorax; C, overall ventral view. Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.7935895" httpUri="https://zenodo.org/record/7935895/files/figure.png" pageId="12" pageNumber="173">Figs. 2A</figureCitation>
,
<figureCitation id="8FC02A1B6A19FF8E97126985AAD0FE50" box="[879,917,475,501]" captionStart="FIGURE 3" captionStartId="4.[151,250,1858,1882]" captionTargetBox="[151,1435,181,1834]" captionTargetId="figure-24@4.[151,1435,181,1834]" captionTargetPageId="4" captionText="FIGURE 3. Aradhya placida gen. et sp. nov.: paratype male (28.53 × 21.82 mm) (ZSI-WRC C.2294) (A, B); paratype male (23.63 × 19.34 mm) (ZSI-WGRC IR.INV.22823) (C, D); paratype male (23.09 × 18.23 mm) (ZSI-WRC C.2293) (EI). A, C, E, overall dorsal view; B, D, F, overall ventral view; G, third maxillipeds; H, dorsal view of left G1; I, left G2. Scale bars = 10 mm (AF), 2 mm (G, H), 1 mm (I)." figureDoi="http://doi.org/10.5281/zenodo.7935897" httpUri="https://zenodo.org/record/7935897/files/figure.png" pageId="12" pageNumber="173">3A</figureCitation>
). While the flagellum on the TME is absent in the
<typeStatus id="C840883C6A19FF8E94BD69A1A867FDBC" box="[192,290,511,537]" pageId="12" pageNumber="173" type="holotype">holotype</typeStatus>
(
<figureCitation id="8FC02A1B6A19FF8E954C69A1A8CCFDBC" box="[305,393,510,537]" captionStart="FIGURE 5" captionStartId="6.[151,250,1943,1967]" captionTargetBox="[203,1384,181,1919]" captionTargetId="figure-24@6.[203,1384,181,1919]" captionTargetPageId="6" captionText="FIGURE 5. Aradhya placida gen. et sp. nov., holotype male (25.12 × 19.85 mm) (ZSI-WRC C.2292).A, mouth parts exposed; B, third maxillipeds; C, thoracic sternites with G1 and G2; D, pleon and telson; E, dorsal view of left G1; F, left G2. Scale bars = 5 mm (C, D), 2 mm (A, B, E), 1 mm (F)." figureDoi="http://doi.org/10.5281/zenodo.7935903" httpUri="https://zenodo.org/record/7935903/files/figure.png" pageId="12" pageNumber="173">Fig. 5B</figureCitation>
), all the male
<typeStatus id="C840883C6A19FF8E965B69A1ABD4FDBC" box="[550,657,511,537]" pageId="12" pageNumber="173" type="paratype">paratypes</typeStatus>
possess a short flagellum on the said structure, which can reach up to half the width of the merus (
<figureCitation id="8FC02A1B6A19FF8E95A76A7DAB71FD98" box="[474,564,547,573]" captionStart="FIGURE 3" captionStartId="4.[151,250,1858,1882]" captionTargetBox="[151,1435,181,1834]" captionTargetId="figure-24@4.[151,1435,181,1834]" captionTargetPageId="4" captionText="FIGURE 3. Aradhya placida gen. et sp. nov.: paratype male (28.53 × 21.82 mm) (ZSI-WRC C.2294) (A, B); paratype male (23.63 × 19.34 mm) (ZSI-WGRC IR.INV.22823) (C, D); paratype male (23.09 × 18.23 mm) (ZSI-WRC C.2293) (EI). A, C, E, overall dorsal view; B, D, F, overall ventral view; G, third maxillipeds; H, dorsal view of left G1; I, left G2. Scale bars = 10 mm (AF), 2 mm (G, H), 1 mm (I)." figureDoi="http://doi.org/10.5281/zenodo.7935897" httpUri="https://zenodo.org/record/7935897/files/figure.png" pageId="12" pageNumber="173">Fig. 3G</figureCitation>
). The flagellum may be also vestigial or entirely missing in one of the TMEs of the male
<typeStatus id="C840883C6A19FF8E95616A19A8C2FDC4" box="[284,391,583,609]" pageId="12" pageNumber="173" type="paratype">paratypes</typeStatus>
(
<figureCitation id="8FC02A1B6A19FF8E95EB6A19A8B5FDC4" box="[406,496,583,609]" captionStart="FIGURE 3" captionStartId="4.[151,250,1858,1882]" captionTargetBox="[151,1435,181,1834]" captionTargetId="figure-24@4.[151,1435,181,1834]" captionTargetPageId="4" captionText="FIGURE 3. Aradhya placida gen. et sp. nov.: paratype male (28.53 × 21.82 mm) (ZSI-WRC C.2294) (A, B); paratype male (23.63 × 19.34 mm) (ZSI-WGRC IR.INV.22823) (C, D); paratype male (23.09 × 18.23 mm) (ZSI-WRC C.2293) (EI). A, C, E, overall dorsal view; B, D, F, overall ventral view; G, third maxillipeds; H, dorsal view of left G1; I, left G2. Scale bars = 10 mm (AF), 2 mm (G, H), 1 mm (I)." figureDoi="http://doi.org/10.5281/zenodo.7935897" httpUri="https://zenodo.org/record/7935897/files/figure.png" pageId="12" pageNumber="173">Fig. 3G</figureCitation>
). The length of the ambulatory legs varied considerably in the largest male
<typeStatus id="C840883C6A19FF8E91416A19ACD9FDC4" box="[1340,1436,583,609]" pageId="12" pageNumber="173" type="paratype">paratype</typeStatus>
(ZSI-WRC C.2294), in which the P3 length is about 2.9 times CL (
<figureCitation id="8FC02A1B6A19FF8E97FF6A35AABCFD20" box="[898,1017,619,645]" captionStart="FIGURE 3" captionStartId="4.[151,250,1858,1882]" captionTargetBox="[151,1435,181,1834]" captionTargetId="figure-24@4.[151,1435,181,1834]" captionTargetPageId="4" captionText="FIGURE 3. Aradhya placida gen. et sp. nov.: paratype male (28.53 × 21.82 mm) (ZSI-WRC C.2294) (A, B); paratype male (23.63 × 19.34 mm) (ZSI-WGRC IR.INV.22823) (C, D); paratype male (23.09 × 18.23 mm) (ZSI-WRC C.2293) (EI). A, C, E, overall dorsal view; B, D, F, overall ventral view; G, third maxillipeds; H, dorsal view of left G1; I, left G2. Scale bars = 10 mm (AF), 2 mm (G, H), 1 mm (I)." figureDoi="http://doi.org/10.5281/zenodo.7935897" httpUri="https://zenodo.org/record/7935897/files/figure.png" pageId="12" pageNumber="173">Fig. 3A, B</figureCitation>
) (vs. P3 length of about 2.0 times CL in the
<typeStatus id="C840883C6A19FF8E94A16AD1A87BFD0C" box="[220,318,655,681]" pageId="12" pageNumber="173" type="holotype">holotype</typeStatus>
and up to 2.4 times CL in the smaller male
<typeStatus id="C840883C6A19FF8E975F6AD1AAD4FD0C" box="[802,913,655,681]" pageId="12" pageNumber="173" type="paratype">paratypes</typeStatus>
;
<figureCitation id="8FC02A1B6A19FF8E97E16AD1AD5BFD0C" box="[924,1054,655,681]" captionStart="FIGURE 2" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[192,1395,181,1919]" captionTargetId="figure-19@3.[192,1395,181,1919]" captionTargetPageId="3" captionText="FIGURE 2. Aradhya placida gen. et sp. nov., holotype male (25.12 × 19.85 mm) (ZSI-WRC C.2292). A, overall dorsal view; B, frontal view of cephalothorax; C, overall ventral view. Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.7935895" httpUri="https://zenodo.org/record/7935895/files/figure.png" pageId="12" pageNumber="173">Figs. 2A, C</figureCitation>
,
<figureCitation id="8FC02A1B6A19FF8E90546AD1AD2EFD0C" box="[1065,1131,655,681]" captionStart="FIGURE 3" captionStartId="4.[151,250,1858,1882]" captionTargetBox="[151,1435,181,1834]" captionTargetId="figure-24@4.[151,1435,181,1834]" captionTargetPageId="4" captionText="FIGURE 3. Aradhya placida gen. et sp. nov.: paratype male (28.53 × 21.82 mm) (ZSI-WRC C.2294) (A, B); paratype male (23.63 × 19.34 mm) (ZSI-WGRC IR.INV.22823) (C, D); paratype male (23.09 × 18.23 mm) (ZSI-WRC C.2293) (EI). A, C, E, overall dorsal view; B, D, F, overall ventral view; G, third maxillipeds; H, dorsal view of left G1; I, left G2. Scale bars = 10 mm (AF), 2 mm (G, H), 1 mm (I)." figureDoi="http://doi.org/10.5281/zenodo.7935897" httpUri="https://zenodo.org/record/7935897/files/figure.png" pageId="12" pageNumber="173">3CF</figureCitation>
). The lateral margins of the pleonal somite 4 are relatively more concave in the male
<typeStatus id="C840883C6A19FF8E97596AEDAACAFD68" box="[804,911,691,717]" pageId="12" pageNumber="173" type="paratype">paratypes</typeStatus>
(
<figureCitation id="8FC02A1B6A19FF8E97DD6AEDAD5EFD68" box="[928,1051,691,717]" captionStart="FIGURE 3" captionStartId="4.[151,250,1858,1882]" captionTargetBox="[151,1435,181,1834]" captionTargetId="figure-24@4.[151,1435,181,1834]" captionTargetPageId="4" captionText="FIGURE 3. Aradhya placida gen. et sp. nov.: paratype male (28.53 × 21.82 mm) (ZSI-WRC C.2294) (A, B); paratype male (23.63 × 19.34 mm) (ZSI-WGRC IR.INV.22823) (C, D); paratype male (23.09 × 18.23 mm) (ZSI-WRC C.2293) (EI). A, C, E, overall dorsal view; B, D, F, overall ventral view; G, third maxillipeds; H, dorsal view of left G1; I, left G2. Scale bars = 10 mm (AF), 2 mm (G, H), 1 mm (I)." figureDoi="http://doi.org/10.5281/zenodo.7935897" httpUri="https://zenodo.org/record/7935897/files/figure.png" pageId="12" pageNumber="173">Fig. 3B, D</figureCitation>
) than those of the
<typeStatus id="C840883C6A19FF8E908C6AEDAC16FD68" box="[1265,1363,691,717]" pageId="12" pageNumber="173" type="holotype">holotype</typeStatus>
(
<figureCitation id="8FC02A1B6A19FF8E91196AEDA9FFFD54" captionStart="FIGURE 2" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[192,1395,181,1919]" captionTargetId="figure-19@3.[192,1395,181,1919]" captionTargetPageId="3" captionText="FIGURE 2. Aradhya placida gen. et sp. nov., holotype male (25.12 × 19.85 mm) (ZSI-WRC C.2292). A, overall dorsal view; B, frontal view of cephalothorax; C, overall ventral view. Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.7935895" httpUri="https://zenodo.org/record/7935895/files/figure.png" pageId="12" pageNumber="173">Figs. 2C</figureCitation>
,
<figureCitation id="8FC02A1B6A19FF8E94BB6A88A9AEFD54" box="[198,235,726,753]" captionStart="FIGURE 5" captionStartId="6.[151,250,1943,1967]" captionTargetBox="[203,1384,181,1919]" captionTargetId="figure-24@6.[203,1384,181,1919]" captionTargetPageId="6" captionText="FIGURE 5. Aradhya placida gen. et sp. nov., holotype male (25.12 × 19.85 mm) (ZSI-WRC C.2292).A, mouth parts exposed; B, third maxillipeds; C, thoracic sternites with G1 and G2; D, pleon and telson; E, dorsal view of left G1; F, left G2. Scale bars = 5 mm (C, D), 2 mm (A, B, E), 1 mm (F)." figureDoi="http://doi.org/10.5281/zenodo.7935903" httpUri="https://zenodo.org/record/7935903/files/figure.png" pageId="12" pageNumber="173">5D</figureCitation>
,
<figureCitation id="8FC02A1B6A19FF8E948B6A89A859FD54" box="[246,284,727,753]" captionStart="FIGURE 6" captionStartId="7.[151,250,1352,1376]" captionTargetBox="[153,1434,419,1327]" captionTargetId="figure-183@7.[151,1436,417,1329]" captionTargetPageId="7" captionText="FIGURE 6. Aradhya placida gen. et sp. nov., holotype male (25.12 × 19.85 mm) (ZSI-WRC C.2292). A, pleon and telson; B, dorsal view of left G1; C, dorsal view of left G1 ultimate article; D, ventral view of left G1; E, ventral view of left G1 ultimate article; F, left G2. Scale bars = 5 mm (A), 2 mm (B, D), 1 mm (C, E, F)." figureDoi="http://doi.org/10.5281/zenodo.7935907" httpUri="https://zenodo.org/record/7935907/files/figure.png" pageId="12" pageNumber="173">6A</figureCitation>
). The telson in the smallest male
<typeStatus id="C840883C6A19FF8E96EF6A89ABB7FD54" box="[658,754,727,753]" pageId="12" pageNumber="173" type="paratype">paratype</typeStatus>
(ZSI-WRC C.2293) is slightly shorter (medial length ca. 1.3 times the proximal width) (
<figureCitation id="8FC02A1B6A19FF8E95B56AA5AB5EFCB0" box="[456,539,763,789]" captionStart="FIGURE 3" captionStartId="4.[151,250,1858,1882]" captionTargetBox="[151,1435,181,1834]" captionTargetId="figure-24@4.[151,1435,181,1834]" captionTargetPageId="4" captionText="FIGURE 3. Aradhya placida gen. et sp. nov.: paratype male (28.53 × 21.82 mm) (ZSI-WRC C.2294) (A, B); paratype male (23.63 × 19.34 mm) (ZSI-WGRC IR.INV.22823) (C, D); paratype male (23.09 × 18.23 mm) (ZSI-WRC C.2293) (EI). A, C, E, overall dorsal view; B, D, F, overall ventral view; G, third maxillipeds; H, dorsal view of left G1; I, left G2. Scale bars = 10 mm (AF), 2 mm (G, H), 1 mm (I)." figureDoi="http://doi.org/10.5281/zenodo.7935897" httpUri="https://zenodo.org/record/7935897/files/figure.png" pageId="12" pageNumber="173">Fig. 3F</figureCitation>
) than the telson in the
<typeStatus id="C840883C6A19FF8E97676AA5AA39FCB0" box="[794,892,763,789]" pageId="12" pageNumber="173" type="holotype">holotype</typeStatus>
and larger
<typeStatus id="C840883C6A19FF8E97876AA5AD1FFCB0" box="[1018,1114,763,789]" pageId="12" pageNumber="173" type="paratype">paratype</typeStatus>
males (medial length ca. 1.4 times the proximal width) (
<figureCitation id="8FC02A1B6A19FF8E95B46B41AB6EFC9C" box="[457,555,799,825]" captionStart="FIGURE 2" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[192,1395,181,1919]" captionTargetId="figure-19@3.[192,1395,181,1919]" captionTargetPageId="3" captionText="FIGURE 2. Aradhya placida gen. et sp. nov., holotype male (25.12 × 19.85 mm) (ZSI-WRC C.2292). A, overall dorsal view; B, frontal view of cephalothorax; C, overall ventral view. Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.7935895" httpUri="https://zenodo.org/record/7935895/files/figure.png" pageId="12" pageNumber="173">Figs. 2C</figureCitation>
,
<figureCitation id="8FC02A1B6A19FF8E964A6B41AB39FC9C" box="[567,636,799,825]" captionStart="FIGURE 3" captionStartId="4.[151,250,1858,1882]" captionTargetBox="[151,1435,181,1834]" captionTargetId="figure-24@4.[151,1435,181,1834]" captionTargetPageId="4" captionText="FIGURE 3. Aradhya placida gen. et sp. nov.: paratype male (28.53 × 21.82 mm) (ZSI-WRC C.2294) (A, B); paratype male (23.63 × 19.34 mm) (ZSI-WGRC IR.INV.22823) (C, D); paratype male (23.09 × 18.23 mm) (ZSI-WRC C.2293) (EI). A, C, E, overall dorsal view; B, D, F, overall ventral view; G, third maxillipeds; H, dorsal view of left G1; I, left G2. Scale bars = 10 mm (AF), 2 mm (G, H), 1 mm (I)." figureDoi="http://doi.org/10.5281/zenodo.7935897" httpUri="https://zenodo.org/record/7935897/files/figure.png" pageId="12" pageNumber="173">3B, D</figureCitation>
,
<figureCitation id="8FC02A1B6A19FF8E96FA6B40ABE9FC9C" box="[647,684,798,825]" captionStart="FIGURE 5" captionStartId="6.[151,250,1943,1967]" captionTargetBox="[203,1384,181,1919]" captionTargetId="figure-24@6.[203,1384,181,1919]" captionTargetPageId="6" captionText="FIGURE 5. Aradhya placida gen. et sp. nov., holotype male (25.12 × 19.85 mm) (ZSI-WRC C.2292).A, mouth parts exposed; B, third maxillipeds; C, thoracic sternites with G1 and G2; D, pleon and telson; E, dorsal view of left G1; F, left G2. Scale bars = 5 mm (C, D), 2 mm (A, B, E), 1 mm (F)." figureDoi="http://doi.org/10.5281/zenodo.7935903" httpUri="https://zenodo.org/record/7935903/files/figure.png" pageId="12" pageNumber="173">5D</figureCitation>
,
<figureCitation id="8FC02A1B6A19FF8E96C56B41AB98FC9C" box="[696,733,799,825]" captionStart="FIGURE 6" captionStartId="7.[151,250,1352,1376]" captionTargetBox="[153,1434,419,1327]" captionTargetId="figure-183@7.[151,1436,417,1329]" captionTargetPageId="7" captionText="FIGURE 6. Aradhya placida gen. et sp. nov., holotype male (25.12 × 19.85 mm) (ZSI-WRC C.2292). A, pleon and telson; B, dorsal view of left G1; C, dorsal view of left G1 ultimate article; D, ventral view of left G1; E, ventral view of left G1 ultimate article; F, left G2. Scale bars = 5 mm (A), 2 mm (B, D), 1 mm (C, E, F)." figureDoi="http://doi.org/10.5281/zenodo.7935907" httpUri="https://zenodo.org/record/7935907/files/figure.png" pageId="12" pageNumber="173">6A</figureCitation>
).
</paragraph>
<paragraph id="1744369E6A19FF8E94BA6B1DAB41FB04" blockId="12.[151,1437,151,2013]" pageId="12" pageNumber="173">
All
<specimenCount id="01FDFD176A19FF8E948C6B1DA83FFCF8" box="[241,378,835,861]" pageId="12" pageNumber="173" type="female">three female</specimenCount>
<typeStatus id="C840883C6A19FF8E95FD6B1DA8AEFCF8" box="[384,491,835,861]" pageId="12" pageNumber="173" type="paratype">paratypes</typeStatus>
share most of the non-sexual character states as those described in the
<typeStatus id="C840883C6A19FF8E908A6B1DAC1CFCF8" box="[1271,1369,835,861]" pageId="12" pageNumber="173" type="holotype">holotype</typeStatus>
male. They differ only in the following features. The smallest female
<typeStatus id="C840883C6A19FF8E97166B39AA8EFC24" box="[875,971,871,897]" pageId="12" pageNumber="173" type="paratype">paratype</typeStatus>
(ZSI-WGRC IR.INV.22823) possesses a proportionally narrower carapace, CW/CL = 1.2 (
<figureCitation id="8FC02A1B6A19FF8E96BA6BD5AA64FC00" box="[711,801,907,933]" captionStart="FIGURE 4" captionStartId="5.[151,250,1910,1934]" captionTargetBox="[214,1372,181,1886]" captionTargetId="figure-19@5.[214,1372,181,1886]" captionTargetPageId="5" captionText="FIGURE 4. Aradhya placida gen. et sp. nov.: paratype female (27.03 × 20.76 mm) (ZSI-WRC C.2294) (A); paratype female (26.69 × 20.87 mm) (ZSI-WRC C.2293) (B, DG); paratype female (23.51 × 19.03 mm) (ZSI-WGRC IR.INV.22823) (C). A, B, C, overall dorsal view; D, third maxillipeds; E, dorsal view of pleonal somites 46 and telson; F, ventral view of pleon with pleopods; G, thoracic sternites showing vulvae. Scale bars = 10 mm (AC), 5 mm (EG), 2 mm (D)." figureDoi="http://doi.org/10.5281/zenodo.7935901" httpUri="https://zenodo.org/record/7935901/files/figure.png" pageId="12" pageNumber="173">Fig. 4C</figureCitation>
). The posterolateral depression on the postorbital region is shallow in the female
<typeStatus id="C840883C6A19FF8E95D76BF1AB50FC6C" box="[426,533,943,969]" pageId="12" pageNumber="173" type="paratype">paratypes</typeStatus>
(
<figureCitation id="8FC02A1B6A19FF8E96586BF1ABD8FC6C" box="[549,669,943,969]" captionStart="FIGURE 4" captionStartId="5.[151,250,1910,1934]" captionTargetBox="[214,1372,181,1886]" captionTargetId="figure-19@5.[214,1372,181,1886]" captionTargetPageId="5" captionText="FIGURE 4. Aradhya placida gen. et sp. nov.: paratype female (27.03 × 20.76 mm) (ZSI-WRC C.2294) (A); paratype female (26.69 × 20.87 mm) (ZSI-WRC C.2293) (B, DG); paratype female (23.51 × 19.03 mm) (ZSI-WGRC IR.INV.22823) (C). A, B, C, overall dorsal view; D, third maxillipeds; E, dorsal view of pleonal somites 46 and telson; F, ventral view of pleon with pleopods; G, thoracic sternites showing vulvae. Scale bars = 10 mm (AC), 5 mm (EG), 2 mm (D)." figureDoi="http://doi.org/10.5281/zenodo.7935901" httpUri="https://zenodo.org/record/7935901/files/figure.png" pageId="12" pageNumber="173">Fig. 4B, C</figureCitation>
) against the deep posterolateral depression on the postorbital region of the
<typeStatus id="C840883C6A19FF8E94A06B8DA87AFC48" box="[221,319,979,1005]" pageId="12" pageNumber="173" type="holotype">holotype</typeStatus>
(
<figureCitation id="8FC02A1B6A19FF8E95336B8DA8EDFC48" box="[334,424,979,1005]" captionStart="FIGURE 2" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[192,1395,181,1919]" captionTargetId="figure-19@3.[192,1395,181,1919]" captionTargetPageId="3" captionText="FIGURE 2. Aradhya placida gen. et sp. nov., holotype male (25.12 × 19.85 mm) (ZSI-WRC C.2292). A, overall dorsal view; B, frontal view of cephalothorax; C, overall ventral view. Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.7935895" httpUri="https://zenodo.org/record/7935895/files/figure.png" pageId="12" pageNumber="173">Fig. 2A</figureCitation>
). All the female
<typeStatus id="C840883C6A19FF8E96226B8DAB8FFC48" box="[607,714,979,1005]" pageId="12" pageNumber="173" type="paratype">paratypes</typeStatus>
have the cervical grooves being smooth and narrow (
<figureCitation id="8FC02A1B6A19FF8E91616B8DACD1FC48" box="[1308,1428,979,1005]" captionStart="FIGURE 4" captionStartId="5.[151,250,1910,1934]" captionTargetBox="[214,1372,181,1886]" captionTargetId="figure-19@5.[214,1372,181,1886]" captionTargetPageId="5" captionText="FIGURE 4. Aradhya placida gen. et sp. nov.: paratype female (27.03 × 20.76 mm) (ZSI-WRC C.2294) (A); paratype female (26.69 × 20.87 mm) (ZSI-WRC C.2293) (B, DG); paratype female (23.51 × 19.03 mm) (ZSI-WGRC IR.INV.22823) (C). A, B, C, overall dorsal view; D, third maxillipeds; E, dorsal view of pleonal somites 46 and telson; F, ventral view of pleon with pleopods; G, thoracic sternites showing vulvae. Scale bars = 10 mm (AC), 5 mm (EG), 2 mm (D)." figureDoi="http://doi.org/10.5281/zenodo.7935901" httpUri="https://zenodo.org/record/7935901/files/figure.png" pageId="12" pageNumber="173">Fig. 4AC</figureCitation>
) (vs. cervical grooves rugose and relatively broader in the
<typeStatus id="C840883C6A19FF8E97656BA9AA3BFBB4" box="[792,894,1015,1041]" pageId="12" pageNumber="173" type="holotype">holotype</typeStatus>
;
<figureCitation id="8FC02A1B6A19FF8E97F56BA9AAA4FBB4" box="[904,993,1015,1041]" captionStart="FIGURE 2" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[192,1395,181,1919]" captionTargetId="figure-19@3.[192,1395,181,1919]" captionTargetPageId="3" captionText="FIGURE 2. Aradhya placida gen. et sp. nov., holotype male (25.12 × 19.85 mm) (ZSI-WRC C.2292). A, overall dorsal view; B, frontal view of cephalothorax; C, overall ventral view. Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.7935895" httpUri="https://zenodo.org/record/7935895/files/figure.png" pageId="12" pageNumber="173">Fig. 2A</figureCitation>
). The flagellum on the TME is missing, vestigial, or short in the female
<typeStatus id="C840883C6A19FF8E95816C45AB22FB90" box="[508,615,1051,1077]" pageId="12" pageNumber="173" type="paratype">paratypes</typeStatus>
(
<figureCitation id="8FC02A1B6A19FF8E960B6C45AB97FB90" box="[630,722,1051,1077]" captionStart="FIGURE 4" captionStartId="5.[151,250,1910,1934]" captionTargetBox="[214,1372,181,1886]" captionTargetId="figure-19@5.[214,1372,181,1886]" captionTargetPageId="5" captionText="FIGURE 4. Aradhya placida gen. et sp. nov.: paratype female (27.03 × 20.76 mm) (ZSI-WRC C.2294) (A); paratype female (26.69 × 20.87 mm) (ZSI-WRC C.2293) (B, DG); paratype female (23.51 × 19.03 mm) (ZSI-WGRC IR.INV.22823) (C). A, B, C, overall dorsal view; D, third maxillipeds; E, dorsal view of pleonal somites 46 and telson; F, ventral view of pleon with pleopods; G, thoracic sternites showing vulvae. Scale bars = 10 mm (AC), 5 mm (EG), 2 mm (D)." figureDoi="http://doi.org/10.5281/zenodo.7935901" httpUri="https://zenodo.org/record/7935901/files/figure.png" pageId="12" pageNumber="173">Fig. 4D</figureCitation>
). The chelipeds of the female
<typeStatus id="C840883C6A19FF8E905E6C45ADCBFB90" box="[1059,1166,1051,1077]" pageId="12" pageNumber="173" type="paratype">paratypes</typeStatus>
are subequal or slightly unequal (
<figureCitation id="8FC02A1B6A19FF8E957C6C61A838FBFC" box="[257,381,1087,1113]" captionStart="FIGURE 4" captionStartId="5.[151,250,1910,1934]" captionTargetBox="[214,1372,181,1886]" captionTargetId="figure-19@5.[214,1372,181,1886]" captionTargetPageId="5" captionText="FIGURE 4. Aradhya placida gen. et sp. nov.: paratype female (27.03 × 20.76 mm) (ZSI-WRC C.2294) (A); paratype female (26.69 × 20.87 mm) (ZSI-WRC C.2293) (B, DG); paratype female (23.51 × 19.03 mm) (ZSI-WGRC IR.INV.22823) (C). A, B, C, overall dorsal view; D, third maxillipeds; E, dorsal view of pleonal somites 46 and telson; F, ventral view of pleon with pleopods; G, thoracic sternites showing vulvae. Scale bars = 10 mm (AC), 5 mm (EG), 2 mm (D)." figureDoi="http://doi.org/10.5281/zenodo.7935901" httpUri="https://zenodo.org/record/7935901/files/figure.png" pageId="12" pageNumber="173">Fig. 4A, C</figureCitation>
) as compared to the noticeably unequal chelipeds of the
<typeStatus id="C840883C6A19FF8E90716C61AD2BFBFC" box="[1036,1134,1087,1113]" pageId="12" pageNumber="173" type="holotype">holotype</typeStatus>
and
<typeStatus id="C840883C6A19FF8E90D56C61AC4DFBFC" box="[1192,1288,1087,1113]" pageId="12" pageNumber="173" type="paratype">paratype</typeStatus>
males (
<figureCitation id="8FC02A1B6A19FF8E91196C61A998FBD8" captionStart="FIGURE 2" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[192,1395,181,1919]" captionTargetId="figure-19@3.[192,1395,181,1919]" captionTargetPageId="3" captionText="FIGURE 2. Aradhya placida gen. et sp. nov., holotype male (25.12 × 19.85 mm) (ZSI-WRC C.2292). A, overall dorsal view; B, frontal view of cephalothorax; C, overall ventral view. Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.7935895" httpUri="https://zenodo.org/record/7935895/files/figure.png" pageId="12" pageNumber="173">Figs. 2A, C</figureCitation>
,
<figureCitation id="8FC02A1B6A19FF8E94946C3DA868FBD8" box="[233,301,1123,1149]" captionStart="FIGURE 3" captionStartId="4.[151,250,1858,1882]" captionTargetBox="[151,1435,181,1834]" captionTargetId="figure-24@4.[151,1435,181,1834]" captionTargetPageId="4" captionText="FIGURE 3. Aradhya placida gen. et sp. nov.: paratype male (28.53 × 21.82 mm) (ZSI-WRC C.2294) (A, B); paratype male (23.63 × 19.34 mm) (ZSI-WGRC IR.INV.22823) (C, D); paratype male (23.09 × 18.23 mm) (ZSI-WRC C.2293) (EI). A, C, E, overall dorsal view; B, D, F, overall ventral view; G, third maxillipeds; H, dorsal view of left G1; I, left G2. Scale bars = 10 mm (AF), 2 mm (G, H), 1 mm (I)." figureDoi="http://doi.org/10.5281/zenodo.7935897" httpUri="https://zenodo.org/record/7935897/files/figure.png" pageId="12" pageNumber="173">3AF</figureCitation>
). The ambulatory legs of the female
<typeStatus id="C840883C6A19FF8E96B76C3DAA70FBD8" box="[714,821,1123,1149]" pageId="12" pageNumber="173" type="paratype">paratypes</typeStatus>
are not so long (P3 length ca. 2.3 times CL) reflecting sexual dimorphism (
<figureCitation id="8FC02A1B6A19FF8E95016CD9A8BDFB04" box="[380,504,1159,1185]" captionStart="FIGURE 4" captionStartId="5.[151,250,1910,1934]" captionTargetBox="[214,1372,181,1886]" captionTargetId="figure-19@5.[214,1372,181,1886]" captionTargetPageId="5" captionText="FIGURE 4. Aradhya placida gen. et sp. nov.: paratype female (27.03 × 20.76 mm) (ZSI-WRC C.2294) (A); paratype female (26.69 × 20.87 mm) (ZSI-WRC C.2293) (B, DG); paratype female (23.51 × 19.03 mm) (ZSI-WGRC IR.INV.22823) (C). A, B, C, overall dorsal view; D, third maxillipeds; E, dorsal view of pleonal somites 46 and telson; F, ventral view of pleon with pleopods; G, thoracic sternites showing vulvae. Scale bars = 10 mm (AC), 5 mm (EG), 2 mm (D)." figureDoi="http://doi.org/10.5281/zenodo.7935901" httpUri="https://zenodo.org/record/7935901/files/figure.png" pageId="12" pageNumber="173">Fig. 4AC</figureCitation>
).
</paragraph>
<paragraph id="1744369E6A19FF8E94BA6CF5AAA1FAF0" blockId="12.[151,1437,151,2013]" pageId="12" pageNumber="173">
The pleon and telson of the female
<typeStatus id="C840883C6A19FF8E96146CF5AB91FB60" box="[617,724,1195,1221]" pageId="12" pageNumber="173" type="paratype">paratypes</typeStatus>
form an ovate outline and cover the thoracic sternum except for s1s3 and lateral edges when closed (
<figureCitation id="8FC02A1B6A19FF8E960E6C91AB88FB4C" box="[627,717,1231,1257]" captionStart="FIGURE 4" captionStartId="5.[151,250,1910,1934]" captionTargetBox="[214,1372,181,1886]" captionTargetId="figure-19@5.[214,1372,181,1886]" captionTargetPageId="5" captionText="FIGURE 4. Aradhya placida gen. et sp. nov.: paratype female (27.03 × 20.76 mm) (ZSI-WRC C.2294) (A); paratype female (26.69 × 20.87 mm) (ZSI-WRC C.2293) (B, DG); paratype female (23.51 × 19.03 mm) (ZSI-WGRC IR.INV.22823) (C). A, B, C, overall dorsal view; D, third maxillipeds; E, dorsal view of pleonal somites 46 and telson; F, ventral view of pleon with pleopods; G, thoracic sternites showing vulvae. Scale bars = 10 mm (AC), 5 mm (EG), 2 mm (D)." figureDoi="http://doi.org/10.5281/zenodo.7935901" httpUri="https://zenodo.org/record/7935901/files/figure.png" pageId="12" pageNumber="173">Fig. 4E</figureCitation>
). The pleonal somite 1 is the shortest; pleonal somites 25 are progressively longer; and pleonal somite 6 is the longest, much broader than long, subequal in length to the telson, with almost straight to convex lateral margins (
<figureCitation id="8FC02A1B6A19FF8E96D86D49ABB8FA94" box="[677,765,1303,1329]" captionStart="FIGURE 4" captionStartId="5.[151,250,1910,1934]" captionTargetBox="[214,1372,181,1886]" captionTargetId="figure-19@5.[214,1372,181,1886]" captionTargetPageId="5" captionText="FIGURE 4. Aradhya placida gen. et sp. nov.: paratype female (27.03 × 20.76 mm) (ZSI-WRC C.2294) (A); paratype female (26.69 × 20.87 mm) (ZSI-WRC C.2293) (B, DG); paratype female (23.51 × 19.03 mm) (ZSI-WGRC IR.INV.22823) (C). A, B, C, overall dorsal view; D, third maxillipeds; E, dorsal view of pleonal somites 46 and telson; F, ventral view of pleon with pleopods; G, thoracic sternites showing vulvae. Scale bars = 10 mm (AC), 5 mm (EG), 2 mm (D)." figureDoi="http://doi.org/10.5281/zenodo.7935901" httpUri="https://zenodo.org/record/7935901/files/figure.png" pageId="12" pageNumber="173">Fig. 4E</figureCitation>
). The female telson is subtriangular, broader than long, with almost straight lateral margins, and the apex is broad and straight (
<figureCitation id="8FC02A1B6A19FF8E97FD6D65AA9DFAF0" box="[896,984,1339,1365]" captionStart="FIGURE 4" captionStartId="5.[151,250,1910,1934]" captionTargetBox="[214,1372,181,1886]" captionTargetId="figure-19@5.[214,1372,181,1886]" captionTargetPageId="5" captionText="FIGURE 4. Aradhya placida gen. et sp. nov.: paratype female (27.03 × 20.76 mm) (ZSI-WRC C.2294) (A); paratype female (26.69 × 20.87 mm) (ZSI-WRC C.2293) (B, DG); paratype female (23.51 × 19.03 mm) (ZSI-WGRC IR.INV.22823) (C). A, B, C, overall dorsal view; D, third maxillipeds; E, dorsal view of pleonal somites 46 and telson; F, ventral view of pleon with pleopods; G, thoracic sternites showing vulvae. Scale bars = 10 mm (AC), 5 mm (EG), 2 mm (D)." figureDoi="http://doi.org/10.5281/zenodo.7935901" httpUri="https://zenodo.org/record/7935901/files/figure.png" pageId="12" pageNumber="173">Fig. 4E</figureCitation>
).
</paragraph>
<paragraph id="1744369E6A19FF8E94BA6D01A880F988" blockId="12.[151,1437,151,2013]" pageId="12" pageNumber="173">
The pleopod 2 of the female
<typeStatus id="C840883C6A19FF8E96796D01AB2AFADC" box="[516,623,1375,1401]" pageId="12" pageNumber="173" type="paratype">paratypes</typeStatus>
has an endopod being slender, longer than the exopod, with long setae on the article joints and margins; the exopod of the pleopod 2 is slender but relatively broader than the endopod, shorter than the endopod, with long setae over the entire structure (
<figureCitation id="8FC02A1B6A19FF8E97546DF9AA3BFA64" box="[809,894,1447,1473]" captionStart="FIGURE 4" captionStartId="5.[151,250,1910,1934]" captionTargetBox="[214,1372,181,1886]" captionTargetId="figure-19@5.[214,1372,181,1886]" captionTargetPageId="5" captionText="FIGURE 4. Aradhya placida gen. et sp. nov.: paratype female (27.03 × 20.76 mm) (ZSI-WRC C.2294) (A); paratype female (26.69 × 20.87 mm) (ZSI-WRC C.2293) (B, DG); paratype female (23.51 × 19.03 mm) (ZSI-WGRC IR.INV.22823) (C). A, B, C, overall dorsal view; D, third maxillipeds; E, dorsal view of pleonal somites 46 and telson; F, ventral view of pleon with pleopods; G, thoracic sternites showing vulvae. Scale bars = 10 mm (AC), 5 mm (EG), 2 mm (D)." figureDoi="http://doi.org/10.5281/zenodo.7935901" httpUri="https://zenodo.org/record/7935901/files/figure.png" pageId="12" pageNumber="173">Fig. 4F</figureCitation>
). The pleopods 35 of the female
<typeStatus id="C840883C6A19FF8E908B6DF9AC24FA64" box="[1270,1377,1447,1473]" pageId="12" pageNumber="173" type="paratype">paratypes</typeStatus>
have an endopod being slender, longer than the exopod, with long setae on the article joints and margins; the exopod of each of the pleopods 35 is conspicuously stouter than the endopod, shorter than the endopod, with long setae on the margins only (
<figureCitation id="8FC02A1B6A19FF8E951F6E4DA8FDF988" box="[354,440,1555,1581]" captionStart="FIGURE 4" captionStartId="5.[151,250,1910,1934]" captionTargetBox="[214,1372,181,1886]" captionTargetId="figure-19@5.[214,1372,181,1886]" captionTargetPageId="5" captionText="FIGURE 4. Aradhya placida gen. et sp. nov.: paratype female (27.03 × 20.76 mm) (ZSI-WRC C.2294) (A); paratype female (26.69 × 20.87 mm) (ZSI-WRC C.2293) (B, DG); paratype female (23.51 × 19.03 mm) (ZSI-WGRC IR.INV.22823) (C). A, B, C, overall dorsal view; D, third maxillipeds; E, dorsal view of pleonal somites 46 and telson; F, ventral view of pleon with pleopods; G, thoracic sternites showing vulvae. Scale bars = 10 mm (AC), 5 mm (EG), 2 mm (D)." figureDoi="http://doi.org/10.5281/zenodo.7935901" httpUri="https://zenodo.org/record/7935901/files/figure.png" pageId="12" pageNumber="173">Fig. 4F</figureCitation>
).
</paragraph>
<paragraph id="1744369E6A19FF8E94BA6E69AB0EF918" blockId="12.[151,1437,151,2013]" pageId="12" pageNumber="173">
The vulvae of the female
<typeStatus id="C840883C6A19FF8E959A6E69AB17F9F4" box="[487,594,1591,1617]" pageId="12" pageNumber="173" type="paratype">paratypes</typeStatus>
are located close to each other (VD/SW = ca. 0.1) on s6; each vulva opens inwards, subovate, large, occupying ca. 0.6 times the length of s6, touching s5/s6, covered by the soft membranous operculum, the outer margin with a straight, gently projecting sternal vulvar cover, and the inner margin with a strongly concave, raised rim (
<figureCitation id="8FC02A1B6A19FF8E95996EFDAB7AF918" box="[484,575,1699,1725]" captionStart="FIGURE 4" captionStartId="5.[151,250,1910,1934]" captionTargetBox="[214,1372,181,1886]" captionTargetId="figure-19@5.[214,1372,181,1886]" captionTargetPageId="5" captionText="FIGURE 4. Aradhya placida gen. et sp. nov.: paratype female (27.03 × 20.76 mm) (ZSI-WRC C.2294) (A); paratype female (26.69 × 20.87 mm) (ZSI-WRC C.2293) (B, DG); paratype female (23.51 × 19.03 mm) (ZSI-WGRC IR.INV.22823) (C). A, B, C, overall dorsal view; D, third maxillipeds; E, dorsal view of pleonal somites 46 and telson; F, ventral view of pleon with pleopods; G, thoracic sternites showing vulvae. Scale bars = 10 mm (AC), 5 mm (EG), 2 mm (D)." figureDoi="http://doi.org/10.5281/zenodo.7935901" httpUri="https://zenodo.org/record/7935901/files/figure.png" pageId="12" pageNumber="173">Fig. 4G</figureCitation>
).
</paragraph>
<paragraph id="1744369E6A19FF8F94BA6E98A99FFF5C" blockId="12.[151,1437,151,2013]" lastBlockId="13.[151,1436,151,321]" lastPageId="13" lastPageNumber="174" pageId="12" pageNumber="173">
<emphasis id="258FEA8C6A19FF8E94BA6E98A8D5F944" bold="true" box="[199,400,1734,1761]" pageId="12" pageNumber="173">Ecological notes.</emphasis>
<taxonomicName id="D0FB4D1D6A19FF8E95E46E99AB13F944" authority="Pati &amp; Bajantri &amp; Hegde, 2023" authorityName="Pati &amp; Bajantri &amp; Hegde" authorityYear="2023" box="[409,598,1735,1761]" class="Malacostraca" family="Gecarcinucidae" genus="Aradhya" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="173" phylum="Arthropoda" rank="species" species="placida" status="sp. nov.">
<emphasis id="258FEA8C6A19FF8E95E46E99AB13F944" box="[409,598,1735,1761]" italics="true" pageId="12" pageNumber="173">Aradhya placida</emphasis>
</taxonomicName>
<emphasis id="258FEA8C6A19FF8E96226E99ABFFF944" bold="true" box="[607,698,1735,1761]" pageId="12" pageNumber="173">
<taxonomicNameLabel id="3EBC57F76A19FF8E96226E99ABFFF944" box="[607,698,1735,1761]" pageId="12" pageNumber="173" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
was found in the evergreen forest of the Central Western Ghats. Crabs of this new species were collected from the earthen burrows mixed with gravel (ca.
<quantity id="D0039B7B6A19FF8E90E66EB5AC4EF8A0" box="[1179,1291,1771,1797]" metricMagnitude="-1" metricUnit="m" metricValue="4.5" metricValueMax="6.0" metricValueMin="3.0" pageId="12" pageNumber="173" unit="cm" value="45.0" valueMax="60.0" valueMin="30.0">3060 cm</quantity>
depth) close to the base of large trees and boulders (
<figureCitation id="8FC02A1B6A19FF8E96316F51AB84F88C" box="[588,705,1807,1833]" captionStart="FIGURE 7" captionStartId="11.[151,250,1928,1952]" captionTargetBox="[151,1435,921,1905]" captionTargetId="figure-329@11.[151,1435,921,1905]" captionTargetPageId="11" captionText="FIGURE 7. Colour in life and habitats of Aradhya placida gen. et sp. nov. A, holotype male (25.12 × 19.85 mm) (ZSI-WRC C.2292); B, paratype male (23.09 × 18.23 mm) (ZSI-WRC C.2293); C, a live crab inside the burrow; D, a live crab near the base of a tree; E, trees and boulders at the type locality. This figure is available in colour at Zootaxa online." figureDoi="http://doi.org/10.5281/zenodo.7935910" httpUri="https://zenodo.org/record/7935910/files/figure.png" pageId="12" pageNumber="173">Fig. 7CE</figureCitation>
) (https://youtu.be/3A4jgXlvMMU). Their burrows are at least
<quantity id="D0039B7B6A19FF8E91FD6F51A9FCF8E8" metricMagnitude="-1" metricUnit="m" metricValue="6.0" pageId="12" pageNumber="173" unit="cm" value="60.0">60 cm</quantity>
apart from each other. These crabs are probably adapted to semi-terrestrial life as their burrows are clearly away from water sources. The nearest stream is approximately
<quantity id="D0039B7B6A19FF8E97676F09AA38F8D4" box="[794,893,1879,1905]" metricMagnitude="1" metricUnit="m" metricValue="4.5" metricValueMax="6.0" metricValueMin="3.0" pageId="12" pageNumber="173" unit="m" value="45.0" valueMax="60.0" valueMin="30.0">3060 m</quantity>
away from those burrows. About 19 individuals were seen within an area of a radius of
<quantity id="D0039B7B6A19FF8E961F6F25ABDAF830" box="[610,671,1915,1941]" metricMagnitude="1" metricUnit="m" metricValue="2.0" pageId="12" pageNumber="173" unit="m" value="20.0">20 m</quantity>
, most of which were found within the burrows; and the remaining crabs were at a short distance from the burrows. These observations were made between 14.30 to 21.00 hrs., which indicate the crepuscular or nocturnal habit of the species. Crabs of
<taxonomicName id="D0FB4D1D6A19FF8E97096F9DAD6BF878" authority="Pati &amp; Bajantri &amp; Hegde, 2023" authorityName="Pati &amp; Bajantri &amp; Hegde" authorityYear="2023" box="[884,1070,1987,2013]" class="Malacostraca" family="Gecarcinucidae" genus="Aradhya" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="173" phylum="Arthropoda" rank="species" species="placida" status="sp. nov.">
<emphasis id="258FEA8C6A19FF8E97096F9DAD6BF878" box="[884,1070,1987,2013]" italics="true" pageId="12" pageNumber="173">Aradhya placida</emphasis>
</taxonomicName>
<emphasis id="258FEA8C6A19FF8E904E6F9DADC9F878" bold="true" box="[1075,1164,1987,2013]" pageId="12" pageNumber="173">
<taxonomicNameLabel id="3EBC57F76A19FF8E904E6F9DADC9F878" box="[1075,1164,1987,2013]" pageId="12" pageNumber="173" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
are mostly active during the monsoon (June-September) and emerge out of their burrows when there is light rain. It is noteworthy to report the behaviour of these crabs as they are non-aggressive and quite calm or peaceful, unlike other Indian gecarcinucid crabs.
</paragraph>
</subSubSection>
<subSubSection id="5FE165156A18FF8F94BA695CAC6AFEE4" pageId="13" pageNumber="174" type="description">
<paragraph id="1744369E6A18FF8F94BA695CAC6AFEE4" blockId="13.[151,1436,151,321]" pageId="13" pageNumber="174">
<emphasis id="258FEA8C6A18FF8F94BA695CAB42FEB9" bold="true" box="[199,519,258,284]" pageId="13" pageNumber="174">Geographical distribution.</emphasis>
<taxonomicName id="D0FB4D1D6A18FF8F966D695DAB88FEB8" authority="Pati &amp; Bajantri &amp; Hegde, 2023" authorityName="Pati &amp; Bajantri &amp; Hegde" authorityYear="2023" box="[528,717,259,285]" class="Malacostraca" family="Gecarcinucidae" genus="Aradhya" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="174" phylum="Arthropoda" rank="species" species="placida" status="sp. nov.">
<emphasis id="258FEA8C6A18FF8F966D695DAB88FEB8" box="[528,717,259,285]" italics="true" pageId="13" pageNumber="174">Aradhya placida</emphasis>
</taxonomicName>
<emphasis id="258FEA8C6A18FF8F96AA695DAA77FEB8" bold="true" box="[727,818,259,285]" pageId="13" pageNumber="174">
<taxonomicNameLabel id="3EBC57F76A18FF8F96AA695DAA77FEB8" box="[727,818,259,285]" pageId="13" pageNumber="174" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
<materialsCitation id="A7933CC36A18FF8F9746695DAC6FFEE4" ID-GBIF-Occurrence="4109228304" country="India" county="Central Western Ghats" location="Bare" municipality="Uttara Kannada district" pageId="13" pageNumber="174" specimenCount="1" stateProvince="Karnataka">
is currently known only from the
<typeStatus id="C840883C6A18FF8F90BC695DADB4FEB8" box="[1217,1265,259,285]" pageId="13" pageNumber="174">type</typeStatus>
locality (
<location id="122460456A18FF8F9122695DACDDFEB8" LSID="urn:lsid:plazi:treatment:9F5287886A1CFF8F94EA6F08AC6AFEE4:122460456A18FF8F9122695DACDDFEB8" box="[1375,1432,259,285]" country="India" county="Central Western Ghats" municipality="Uttara Kannada district" name="Bare" pageId="13" pageNumber="174" stateProvince="Karnataka">Bare</location>
,
<collectingMunicipality id="F720ACE46A18FF8F94EA6979A8E5FEE4" box="[151,416,295,321]" pageId="13" pageNumber="174">Uttara Kannada district</collectingMunicipality>
,
<collectingRegion id="D53FF87C6A18FF8F95D76979AB19FEE4" box="[426,604,295,321]" country="India" name="Karnataka" pageId="13" pageNumber="174">Karnataka state</collectingRegion>
), which is located in the
<collectingCounty id="FE254E126A18FF8F970F6979AD2BFEE4" box="[882,1134,295,321]" pageId="13" pageNumber="174">Central Western Ghats</collectingCounty>
of
<collectingCountry id="6FEC760E6A18FF8F90E96979AD8BFEE4" box="[1172,1230,295,321]" name="India" pageId="13" pageNumber="174">India</collectingCountry>
(
<figureCitation id="8FC02A1B6A18FF8F90A16979AC5BFEE4" box="[1244,1310,295,321]" captionStart="FIGURE 1" captionStartId="1.[151,250,1864,1888]" captionTargetBox="[155,1431,567,1835]" captionTargetId="figure-209@1.[151,1436,561,1841]" captionTargetPageId="1" captionText="FIGURE 1. Map of southern India, showing the Central- and Southern Western Ghats, and the distribution of Aradhya gen. nov. and related genera. The elevation profile of the Central- and Southern Western Ghats is shown in the graph below; deep valleys/gaps are indicated by down arrows; the graph includes only the northernmost, southernmost, highest-elevation, and lowest-elevation records of each genus. This figure is available in colour at Zootaxa online." figureDoi="http://doi.org/10.5281/zenodo.7935891" httpUri="https://zenodo.org/record/7935891/files/figure.png" pageId="13" pageNumber="174">Fig. 1</figureCitation>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>