212 lines
22 KiB
XML
212 lines
22 KiB
XML
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<mods:title id="87B4732FAD55ACCBE188F310D2A7A39A">The empidoid flies (Diptera: Empidoidea, exclusive of Dolichopodidae) of the Russian Arctic islands and Svalbard Archipelago</mods:title>
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<mods:name id="1A21EE69E9536F3C4EAF3207B16FF490" type="personal">
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<mods:namePart id="79EF2E59A54126489F9A08A00A3C57C8">Shamshev, Igor V.</mods:namePart>
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<mods:roleTerm id="24E06A70E5D7E37B02C222553F60102E">Author</mods:roleTerm>
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<mods:namePart id="A6F4E2155611A5C1685DC8375CCC952B">Sinclair, Bradley J.</mods:namePart>
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<mods:nameIdentifier id="9C0FC092A62EA06D697C18154950D664" type="email">bradley.sinclair@canada.ca</mods:nameIdentifier>
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<mods:namePart id="2795DA105657F7016AFDE983389A34BC">Khruleva, Olga A.</mods:namePart>
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<mods:nameIdentifier id="C971D29747664053E2F7354EF83A06D3" type="email">oa-khruleva@mail.ru</mods:nameIdentifier>
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<mods:title id="285FEAE44E78C308B30017BE7C49D5D5">Zootaxa</mods:title>
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<mods:date id="B0C8E581044F212C90091ADD8F4C84A1">2020</mods:date>
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<mods:number id="EA12E45148BB9217DC7553D95E27A8E6">2020-09-07</mods:number>
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<mods:number id="3872AC8B0902E4A2231BB1B0567D0ED1">4848</mods:number>
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<treatment id="7B7E785C64649F1557EEF9B7FC5DE885" ID-DOI="http://doi.org/10.5281/zenodo.4407568" ID-GBIF-Taxon="177136872" ID-Zenodo-Dep="4407568" LSID="urn:lsid:plazi:treatment:7B7E785C64649F1557EEF9B7FC5DE885" httpUri="http://treatment.plazi.org/id/7B7E785C64649F1557EEF9B7FC5DE885" lastPageId="7" lastPageNumber="8" pageId="6" pageNumber="7">
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<subSubSection id="BBCD9AC164649F1457EEF9B7FE24EFFE" box="[151,494,1651,1678]" pageId="6" pageNumber="7" type="nomenclature">
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<paragraph id="F368C94A64649F1457EEF9B7FE24EFFE" blockId="6.[151,494,1651,1678]" box="[151,494,1651,1678]" pageId="6" pageNumber="7">
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<heading id="A8207E2664649F1457EEF9B7FE24EFFE" bold="true" box="[151,494,1651,1678]" fontSize="11" level="2" pageId="6" pageNumber="7" reason="3">
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<taxonomicName id="34D7B2C964649F1457EEF9B7FE24EFFE" ID-CoL="88YPN" authority="Wagner" authorityName="Wagner" box="[151,494,1651,1678]" class="Insecta" family="Brachystomatidae" genus="Heleodromia" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="irwini">
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<emphasis id="C1A3155864649F1457EEF9B7FE24EFFE" bold="true" box="[151,494,1651,1678]" pageId="6" pageNumber="7">
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<emphasis id="C1A3155864649F1457EEF9B7FE4FEFFE" bold="true" box="[151,389,1651,1678]" italics="true" pageId="6" pageNumber="7">Heleodromia irwini</emphasis>
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Wagner
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||
</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="BBCD9AC164649F1457EEF97CFB58EF9F" pageId="6" pageNumber="7" type="reference_group">
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<paragraph id="F368C94A64649F1457EEF97CFBF9EFA0" blockId="6.[151,1170,1720,1775]" box="[151,1075,1720,1745]" pageId="6" pageNumber="7">
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<treatmentCitationGroup id="D3C7EE6464649F1457EEF97CFBF9EFA0" box="[151,1075,1720,1745]" pageId="6" pageNumber="7">
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<treatmentCitation id="7276EF5B64649F1457EEF97CFDD1EFA1" author="Wagner, R." box="[151,539,1720,1745]" page="37" pageId="6" pageNumber="7" year="1985">
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<taxonomicName id="34D7B2C964649F1457EEF97CFDD1EFA1" ID-CoL="88YPN" authority="Wagner, 1985: 37" authorityName="Wagner" authorityPageNumber="37" authorityYear="1985" box="[151,539,1720,1745]" class="Insecta" family="Brachystomatidae" genus="Heleodromia" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="irwini">
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||
<emphasis id="C1A3155864649F1457EEF97CFE95EFA0" box="[151,351,1720,1744]" italics="true" pageId="6" pageNumber="7">Heleodromia irwini</emphasis>
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<materialsCitation id="43BFC31764649F14561CF97DFDD1EFA1" ID-GBIF-Occurrence="3026890365" box="[357,539,1720,1745]" country="United Kingdom" location="Moray" pageId="6" pageNumber="7" specimenCount="1" stateProvince="Moray">
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<bibRefCitation id="9746B4BB64649F14561CF97DFDD1EFA1" author="Wagner, R." box="[357,539,1720,1745]" pageId="6" pageNumber="7" pagination="33 - 43" refId="ref65559" refString="Wagner, R. (1985) A revision of the genus Heleodromia (Diptera, Empididae) in Europe. Aquatic Insects, 7, 33 - 43. https: // doi. org / 10.1080 / 01650428509361196" type="journal article" year="1985">Wagner, 1985: 37</bibRefCitation>
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</materialsCitation>
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</taxonomicName>
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</treatmentCitation>
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.
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<typeStatus id="2C6C77E864649F14555DF97DFD9CEFA1" box="[548,598,1721,1745]" pageId="6" pageNumber="7">Type</typeStatus>
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locality:
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<collectingCountry id="8BC089DA64649F1455CFF97CFCDBEFA0" box="[694,785,1720,1744]" name="United Kingdom" pageId="6" pageNumber="7">Scotland</collectingCountry>
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,
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<collectingRegion id="311307A864649F145462F97DFCAAEFA1" box="[795,864,1721,1745]" country="United Kingdom" name="Moray" pageId="6" pageNumber="7">Moray</collectingRegion>
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, Dorback Burn,
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<collectingCountry id="8BC089DA64649F14537EF97DFBE5EFA0" box="[1031,1071,1721,1744]" name="United Kingdom" pageId="6" pageNumber="7">UK</collectingCountry>
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.
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</treatmentCitationGroup>
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</paragraph>
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<paragraph id="F368C94A64649F1457EEF913FB58EF9F" blockId="6.[151,1170,1720,1775]" box="[151,1170,1751,1775]" pageId="6" pageNumber="7">
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Other references:
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<bibRefCitation id="9746B4BB64649F145634F913FD8CEF9F" author="Sinclair, B. J. & Brooks, S. E. & Cumming, J. M. & Coovert, G. A." box="[333,582,1751,1775]" pageId="6" pageNumber="7" pagination="629 - 651" refId="ref65374" refString="Sinclair, B. J., Brooks, S. E., Cumming, J. M. & Coovert, G. A. (2011) Revision of the Nearctic species of Heleodromia (Diptera: Empidoidea: Brachystomatidae). The Canadian Entomologist, 143, 629 - 651. https: // doi. org / 10.4039 / n 11 - 036" type="journal article" year="2011">
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Sinclair
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||
<emphasis id="C1A3155864649F1456DBF91CFE04EF9F" box="[418,462,1751,1775]" italics="true" pageId="6" pageNumber="7">et al</emphasis>
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., 2011: 641
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||
</bibRefCitation>
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(revision, redescription);
|
||
<bibRefCitation id="9746B4BB64649F145434F913FBD3EF9F" author="Shamshev, I. V." box="[845,1049,1751,1775]" pageId="6" pageNumber="7" pagination="3 - 183" refId="ref65045" refString="Shamshev, I. V. (2016) An annotated checklist of empidoid flies (Diptera: Empidoidea, except Dolichopodidae) of Russia. Proceedings of the Russian Entomological Society, 87, 3 - 183." type="journal article" year="2016">Shamshev, 2016: 17</bibRefCitation>
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(checklist).
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</paragraph>
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</subSubSection>
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<subSubSection id="BBCD9AC164649F1557EEF8D9FC1BE989" lastPageId="7" lastPageNumber="8" pageId="6" pageNumber="7" type="materials_examined">
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<paragraph id="F368C94A64649F1457EEF8D9FCCDEE2C" blockId="6.[151,1437,1821,2028]" pageId="6" pageNumber="7">
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<emphasis id="C1A3155864649F1457EEF8D9FE48EE47" bold="true" box="[151,386,1821,1847]" pageId="6" pageNumber="7">Material examined.</emphasis>
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<materialsCitation id="43BFC31764649F1456F2F8D9FCC9EE2C" ID-GBIF-Occurrence="3026890653" collectingDate="2016-07-12" collectingDateMax="2016-07-16" collectingDateMin="2016-07-12" collectionCode="YPT, LFV, ZIN" country="Russia" latitude="71.01667" location="river Neozhidannaya" longLatPrecision="973" longitude="179.15" municipality="Wrangel Island" pageId="6" pageNumber="7" specimenCode="BT 4" specimenCount="1" specimenCount-male="1" stateProvince="Chukotka">
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<emphasis id="C1A3155864649F1456F2F8D9FC43EE47" bold="true" box="[395,905,1821,1847]" pageId="6" pageNumber="7">
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<collectingCountry id="8BC089DA64649F1456F2F8D9FE38EE47" box="[395,498,1821,1847]" name="Russia" pageId="6" pageNumber="7">RUSSIA</collectingCountry>
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.
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||
<collectingRegion id="311307A864649F145578F8D9FDB3EE47" box="[513,633,1821,1847]" country="Russia" name="Chukotka" pageId="6" pageNumber="7">Chukotka</collectingRegion>
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AO (
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||
<collectingMunicipality id="130C533064649F1455C3F8D9FCB6EE47" box="[698,892,1821,1847]" pageId="6" pageNumber="7">Wrangel Island</collectingMunicipality>
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):
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||
</emphasis>
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||
middle flow of
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||
<location id="F6089F9164649F14533DF8D9FAF8EE47" LSID="urn:lsid:plazi:treatment:7B7E785C64649F1557EEF9B7FC5DE885:F6089F9164649F14533DF8D9FAF8EE47" box="[1092,1330,1821,1847]" country="Russia" latitude="71.01667" longLatPrecision="973" longitude="179.15" municipality="Wrangel Island" name="river Neozhidannaya" pageId="6" pageNumber="7" stateProvince="Chukotka">river Neozhidannaya</location>
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,
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||
<geoCoordinate id="96E3AF8D64649F145246F8DBFA56EE4A" box="[1343,1436,1823,1850]" degrees="71" direction="north" minutes="01" orientation="latitude" pageId="6" pageNumber="7" precision="925" value="71.01667">71°01′N</geoCoordinate>
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<geoCoordinate id="96E3AF8D64649F1457EEF887FEC9EE2E" box="[151,259,1859,1886]" degrees="179" direction="east" minutes="09" orientation="longitude" pageId="6" pageNumber="7" precision="925" value="179.15">179°09′E</geoCoordinate>
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,
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||
<collectionCode id="95C6518F64649F145674F886FE89EE2C" box="[269,323,1858,1884]" pageId="6" pageNumber="7">YPT</collectionCode>
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||
,
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||
<specimenCode id="A371613164649F145634F886FE42EE2B" box="[333,392,1857,1884]" collectionCode="BT" pageId="6" pageNumber="7">BT 4</specimenCode>
|
||
,
|
||
<date id="8769EF8A64649F1456EDF885FD8AEE2B" box="[404,576,1857,1883]" pageId="6" pageNumber="7" value="2016-07-12" valueMax="2016-07-16" valueMin="2016-07-12">
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<collectingDate id="972D166264649F1456EDF885FD8AEE2B" box="[404,576,1857,1883]" pageId="6" pageNumber="7" value="2016-07-12" valueMax="2016-07-16" valueMin="2016-07-12">12–16.vii.2016</collectingDate>
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||
</date>
|
||
,
|
||
<collectionCode id="95C6518F64649F145532F886FDB5EE2C" box="[587,639,1858,1884]" pageId="6" pageNumber="7">LFV</collectionCode>
|
||
(
|
||
<specimenCount id="E5D102C364649F1455EBF885FD76EE2C" box="[658,700,1857,1884]" pageId="6" pageNumber="7" type="male">1 ♂</specimenCount>
|
||
,
|
||
<collectionCode id="95C6518F64649F1455B1F886FD31EE2C" box="[712,763,1858,1884]" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="6" pageNumber="7" type="Museum">ZIN</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="F368C94A64649F1557BEF8A1FC1BE989" blockId="6.[151,1437,1821,2028]" lastBlockId="7.[151,1436,151,501]" lastPageId="7" lastPageNumber="8" pageId="6" pageNumber="7">
|
||
<emphasis id="C1A3155864649F1457BEF8A1FE97EE0F" bold="true" box="[199,349,1893,1919]" pageId="6" pageNumber="7">Recognition.</emphasis>
|
||
Small greyish delicate flies (body length
|
||
<quantity id="342F64AF64649F145450F8A1FC50EEF0" box="[809,922,1893,1920]" metricMagnitude="-3" metricUnit="m" metricValue="3.25" metricValueMax="3.5" metricValueMin="3.0" pageId="6" pageNumber="7" unit="mm" value="3.25" valueMax="3.5" valueMin="3.0">3–3.5 mm</quantity>
|
||
) with very slender legs almost lacking distinct setae, eyes dichoptic in both sexes, separated by broad V-shaped flattened frons; legs black to reddish brown, “knees” often somewhat paler; habitually very similar to
|
||
<taxonomicName id="34D7B2C964649F14544BF86AFC6DEEB7" baseAuthorityName="Melander" baseAuthorityYear="1902" box="[818,935,1965,1992]" class="Insecta" family="Brachystomatidae" genus="Heleodromia" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="pullata">
|
||
<emphasis id="C1A3155864649F14544BF86AFC8CEEB7" box="[818,838,1966,1991]" italics="true" pageId="6" pageNumber="7">H</emphasis>
|
||
.
|
||
<emphasis id="C1A3155864649F145421F869FC6DEEB7" box="[856,935,1965,1991]" italics="true" pageId="6" pageNumber="7">pullata</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="6BECD5CF64649F1454C3F869FBC8EEB7" box="[954,1026,1965,1991]" captionStart="FIGURES 2–5" captionStartId="8.[151,264,1640,1664]" captionTargetBox="[153,1435,183,1614]" captionTargetId="figure-20@8.[151,1437,181,1616]" captionTargetPageId="8" captionText="FIGURES 2–5. Brachystomatidae (2) and Empididae, Clinocerinae (3–5). 2. Heleodromia pullata (Melander), male habitus, lateral view; 3. Clinocera nivalis (Zetterstedt), male habitus, lateral view; 4. Trichoclinocera lapponica (Ringdahl), male habitus, lateral view; 5. same species, male postabdomen, lateral view." figureDoi="http://doi.org/10.5281/zenodo.4406991" httpUri="https://zenodo.org/record/4406991/files/figure.png" pageId="6" pageNumber="7">Fig. 2</figureCitation>
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||
, see below). Male terminalia large, reniform; tergite 8 not broader than sternite 8, only produced medially as a shallow bilobed to forked projection and lacking posterolateral projection (
|
||
<bibRefCitation id="9746B4BB64659F155539FF53FC98E9C1" author="Sinclair, B. J. & Brooks, S. E. & Cumming, J. M. & Coovert, G. A." box="[576,850,151,177]" pageId="7" pageNumber="8" pagination="629 - 651" refId="ref65374" refString="Sinclair, B. J., Brooks, S. E., Cumming, J. M. & Coovert, G. A. (2011) Revision of the Nearctic species of Heleodromia (Diptera: Empidoidea: Brachystomatidae). The Canadian Entomologist, 143, 629 - 651. https: // doi. org / 10.4039 / n 11 - 036" type="journal article" year="2011">
|
||
Sinclair
|
||
<emphasis id="C1A3155864659F1555D9FF53FD18E9C1" box="[672,722,151,177]" italics="true" pageId="7" pageNumber="8">et al</emphasis>
|
||
. 2011: 641
|
||
</bibRefCitation>
|
||
, fig. 10); epandrial lobe slender, finger-like, bearing apical seta; surstylus slender, finger-like, with scattered setae, with small preapical thumb-like projection medially; apex with short dentiform process; phallus with slender, needle-like apex.
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||
</paragraph>
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||
</subSubSection>
|
||
<subSubSection id="BBCD9AC164659F1557BEFEC6FC5DE885" pageId="7" pageNumber="8" type="distribution">
|
||
<paragraph id="F368C94A64659F1557BEFEC6FDB4E8A1" blockId="7.[151,1436,151,501]" pageId="7" pageNumber="8">
|
||
<emphasis id="C1A3155864659F1557BEFEC6FEABE86C" bold="true" box="[199,353,258,284]" pageId="7" pageNumber="8">Distribution.</emphasis>
|
||
Holarctic. The distribution of this species appears to be represented by widely disjunct populations exhibiting boreo-alpine connections in Europe and North America (
|
||
<bibRefCitation id="9746B4BB64659F1554D8FEE3FBEBE831" author="Plant, A." box="[929,1057,294,321]" pageId="7" pageNumber="8" pagination="263" refId="ref64561" refString="Plant, A. (2005) Heleodromia irwini Wagner (Diptera: Empididae), an English boreo-alpine relict? British Journal of Entomology and Natural History, 18, 263." type="journal article" year="2005">Plant 2005</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="9746B4BB64659F155356FEE3FB38E831" author="Plant, A. R. & Jonassen, T. & Grootaert, P. & Meyer, H. & Pollet, M. & Drake, M." box="[1071,1266,295,321]" pageId="7" pageNumber="8" pagination="852 - 868" refId="ref64594" refString="Plant, A. R., Jonassen, T., Grootaert, P., Meyer, H., Pollet, M. & Drake, M. (2017) The arrow points north-endemic areas and post-Devensian assembly of the British Empidoidea fauna (Insecta: Diptera). Biological Journal of the Linnean Society, 120, 852 - 868. https: // doi. org / 10.1093 / biolinnean / blw 011" type="journal article" year="2017">
|
||
Plant
|
||
<emphasis id="C1A3155864659F15530AFEE3FB6DE831" box="[1139,1191,295,321]" italics="true" pageId="7" pageNumber="8">et al</emphasis>
|
||
. 2017
|
||
</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="9746B4BB64659F155278FEE3FF19E815" author="Sinclair, B. J. & Brooks, S. E. & Cumming, J. M. & Coovert, G. A." pageId="7" pageNumber="8" pagination="629 - 651" refId="ref65374" refString="Sinclair, B. J., Brooks, S. E., Cumming, J. M. & Coovert, G. A. (2011) Revision of the Nearctic species of Heleodromia (Diptera: Empidoidea: Brachystomatidae). The Canadian Entomologist, 143, 629 - 651. https: // doi. org / 10.4039 / n 11 - 036" type="journal article" year="2011">
|
||
Sinclair
|
||
<emphasis id="C1A3155864659F155218FEE3FA5FE831" box="[1377,1429,295,321]" italics="true" pageId="7" pageNumber="8">et al</emphasis>
|
||
. 2011
|
||
</bibRefCitation>
|
||
). Our record of
|
||
<taxonomicName id="34D7B2C964659F1556FEFE8FFE26E814" box="[391,492,331,357]" class="Insecta" family="Brachystomatidae" genus="Heleodromia" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="irwini">
|
||
<emphasis id="C1A3155864659F1556FEFE8FFE51E814" box="[391,411,331,356]" italics="true" pageId="7" pageNumber="8">H</emphasis>
|
||
.
|
||
<emphasis id="C1A3155864659F1556D0FE8FFE26E814" box="[425,492,331,356]" italics="true" pageId="7" pageNumber="8">irwini</emphasis>
|
||
</taxonomicName>
|
||
from Wrangel Island may support this suggestion. In Eurasia, this species is known from
|
||
<collectingCountry id="8BC089DA64659F1557AFFEABFEE1E8F9" box="[214,299,367,393]" name="Austria" pageId="7" pageNumber="8">Austria</collectingCountry>
|
||
,
|
||
<collectingCountry id="8BC089DA64659F155640FEABFE68E8F9" box="[313,418,367,393]" name="Germany" pageId="7" pageNumber="8">Germany</collectingCountry>
|
||
,
|
||
<collectingCountry id="8BC089DA64659F1556D6FEABFE29E8F9" box="[431,483,367,393]" name="Italy" pageId="7" pageNumber="8">Italy</collectingCountry>
|
||
,
|
||
<collectingCountry id="8BC089DA64659F155688FEABFDB0E8F9" box="[497,634,367,393]" name="Switzerland" pageId="7" pageNumber="8">Switzerland</collectingCountry>
|
||
,
|
||
<collectingCountry id="8BC089DA64659F1555F1FEABFD20E8F9" box="[648,746,367,393]" name="United Kingdom" pageId="7" pageNumber="8">Scotland</collectingCountry>
|
||
and the Far East of
|
||
<collectingCountry id="8BC089DA64659F1554A3FEABFBEFE8F9" box="[986,1061,367,393]" name="Russia" pageId="7" pageNumber="8">Russia</collectingCountry>
|
||
(Amurskaya and Magadanskaya Provinces); in North America, it is recorded from western part of
|
||
<collectingCountry id="8BC089DA64659F15540BFE57FC0CE8DD" box="[882,966,403,429]" name="Canada" pageId="7" pageNumber="8">Canada</collectingCountry>
|
||
and
|
||
<collectingCountry id="8BC089DA64659F155485FE57FBF2E8DD" box="[1020,1080,403,429]" name="United States of America" pageId="7" pageNumber="8">USA</collectingCountry>
|
||
. It seems that
|
||
<taxonomicName id="34D7B2C964659F1553AFFE57FAF1E8DC" box="[1238,1339,403,428]" class="Insecta" family="Brachystomatidae" genus="Heleodromia" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="irwini">
|
||
<emphasis id="C1A3155864659F1553AFFE57FAF1E8DC" box="[1238,1339,403,428]" italics="true" pageId="7" pageNumber="8">H. irwini</emphasis>
|
||
</taxonomicName>
|
||
is found here for the first time from the tundra zone.
|
||
</paragraph>
|
||
<paragraph id="F368C94A64659F1557BEFE1EFC5DE885" blockId="7.[151,1436,151,501]" box="[199,919,474,501]" pageId="7" pageNumber="8">
|
||
<emphasis id="C1A3155864659F1557BEFE1EFEE0E884" bold="true" box="[199,298,474,500]" pageId="7" pageNumber="8">Habitat.</emphasis>
|
||
The species was collected in wetlands in a river valley.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |