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<document id="1B77C50D159725E1A080EE0B73C22EFB" ID-CLB-Dataset="27580" ID-DOI="10.11646/zootaxa.4554.1.2" ID-GBIF-Dataset="d41c057e-5b64-4634-84dc-bd5f56a9d789" ID-ISSN="1175-5326" ID-Zenodo-Dep="2623296" ID-ZooBank="986112BE-6ABB-47FE-BA8F-CECEF1C21A5D" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1554270904597" checkinUser="plazi" docAuthor="Carmo, Daniel D. D. &amp; Henriques, Augusto L." docDate="2019" docId="746A87ED731BFF87FF17D7C124E416CC" docLanguage="en" docName="zootaxa.4554.1.2.pdf" docOrigin="Zootaxa 4554 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Tabanus platycerus Fairchild" docType="treatment" docVersion="9" lastPageNumber="83" masterDocId="8853FF957308FF93FF80D65527531464" masterDocTitle="Taxonomy of Tabanus trivittatus species-group (Diptera: Tabanidae), with description of five new species" masterLastPageNumber="100" masterPageNumber="63" pageNumber="82" updateTime="1698716223463" updateUser="ExternalLinkService">
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<mods:title id="7EDF5E0A07C961499334714414F1D818">Taxonomy of Tabanus trivittatus species-group (Diptera: Tabanidae), with description of five new species</mods:title>
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<treatment id="746A87ED731BFF87FF17D7C124E416CC" ID-DOI="http://doi.org/10.5281/zenodo.5944734" ID-GBIF-Taxon="154534523" ID-Zenodo-Dep="5944734" LSID="urn:lsid:plazi:treatment:746A87ED731BFF87FF17D7C124E416CC" httpUri="http://treatment.plazi.org/id/746A87ED731BFF87FF17D7C124E416CC" lastPageId="20" lastPageNumber="83" pageId="19" pageNumber="82">
<subSubSection id="B4D96570731BFF80FF17D7C126A815CA" box="[151,507,404,430]" pageId="19" pageNumber="82" type="nomenclature">
<paragraph id="FC7C36FB731BFF80FF17D7C126A815CA" blockId="19.[151,507,404,463]" box="[151,507,404,430]" pageId="19" pageNumber="82">
<heading id="A7348197731BFF80FF17D7C126A815CA" bold="true" box="[151,507,404,430]" fontSize="11" level="1" pageId="19" pageNumber="82" reason="1">
<taxonomicName id="3BC34D78731BFF80FF17D7C126A815CA" ID-CoL="9HRSH" authority="Fairchild" authorityName="Fairchild" box="[151,507,404,430]" class="Insecta" family="Tabanidae" genus="Tabanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="19" pageNumber="82" phylum="Arthropoda" rank="species" species="platycerus">
<emphasis id="CEB7EAE9731BFF80FF17D7C126A815CA" bold="true" box="[151,507,404,430]" pageId="19" pageNumber="82">
<emphasis id="CEB7EAE9731BFF80FF17D7C126D215CA" bold="true" box="[151,385,404,430]" italics="true" pageId="19" pageNumber="82">Tabanus platycerus</emphasis>
Fairchild
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="B4D96570731BFF80FF17D7E3262E15AB" box="[151,381,438,463]" pageId="19" pageNumber="82" type="description">
<paragraph id="FC7C36FB731BFF80FF17D7E3262E15AB" blockId="19.[151,507,404,463]" box="[151,381,438,463]" pageId="19" pageNumber="82">
(
<figureCitation id="64F82A7E731BFF80FF1ED7E3264615AB" box="[158,277,438,463]" captionStart="FIGURE 17" captionStartId="19.[151,250,1889,1911]" captionTargetBox="[355,1236,658,1821]" captionTargetId="figure@19.[340,1247,638,1837]" captionTargetPageId="19" captionText="FIGURE 17. Tabanus platycerus. Paratype female. A. Dorsal habitus. B. Lateral habitus. C. Head, anterior view. D. Head, lateral view. E. Labels. Scale bars. A, B. 5 mm. C, D. 1 mm." httpUri="https://zenodo.org/record/2623332/files/figure.png" pageId="19" pageNumber="82">Figures 17</figureCitation>
A–E, 33)
</paragraph>
</subSubSection>
<subSubSection id="B4D96570731BFF80FF17D7AB267C1636" pageId="19" pageNumber="82" type="reference_group">
<paragraph id="FC7C36FB731BFF80FF17D7AB267C1636" blockId="19.[151,1436,510,594]" pageId="19" pageNumber="82">
<treatmentCitationGroup id="DCD311D5731BFF80FF17D7AB267C1636" pageId="19" pageNumber="82">
<taxonomicName id="3BC34D78731BFF80FF17D7AB25661670" ID-CoL="9HRSH" authority="Fairchild, 1976: 251" authorityName="Fairchild" authorityPageNumber="251" authorityYear="1976" box="[151,565,510,532]" class="Insecta" family="Tabanidae" genus="Tabanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="19" pageNumber="82" phylum="Arthropoda" rank="species" species="platycerus">
<emphasis id="CEB7EAE9731BFF80FF17D7AB260E1670" box="[151,349,510,532]" italics="true" pageId="19" pageNumber="82">Tabanus platycerus</emphasis>
<treatmentCitation id="7D6210EA731BFF80FEE4D7AB25661670" author="Fairchild, G. B." box="[356,565,510,532]" page="251" pageId="19" pageNumber="82" year="1976">
<bibRefCitation id="98524B0A731BFF80FEE4D7AB25511670" author="Fairchild, G. B." box="[356,514,510,532]" pageId="19" pageNumber="82" pagination="237 - 261" refId="ref17120" refString="Fairchild, G. B. (1976) Notes on Neotropical Tabanidae (Dipt.) XVI. The Tabanus trivittatus complex. Studia Entomologica, 19 (1 - 4), 237 - 261." type="journal article" year="1976">Fairchild, 1976</bibRefCitation>
: 251
</treatmentCitation>
</taxonomicName>
, fig. 9;
<treatmentCitation id="7D6210EA731BFF80FD06D7AB24061670" author="Fairchild, G. B." box="[646,853,510,532]" page="119" pageId="19" pageNumber="82" year="1986">
<bibRefCitation id="98524B0A731BFF80FD06D7AB24761670" author="Fairchild, G. B." box="[646,805,510,532]" pageId="19" pageNumber="82" pagination="1 - 139" refId="ref17242" refString="Fairchild, G. B. (1986) The Tabanidae of Panama. Contibutions of the American Entomological Institute, 22 (3), 1 - 139." type="journal article" year="1986">Fairchild, 1986</bibRefCitation>
: 119
</treatmentCitation>
(
<collectingCountry id="84D4766B731BFF80FCE3D7AB24E61670" box="[867,949,510,532]" name="Panama" pageId="19" pageNumber="82">Panama</collectingCountry>
spp.);
<treatmentCitation id="7D6210EA731BFF80FC7CD7AB22161670" author="Henriques, A. L. &amp; Gorayeb, I. S." box="[1020,1349,510,532]" page="19" pageId="19" pageNumber="82" year="1993">
<bibRefCitation id="98524B0A731BFF80FC7CD7AB22721670" author="Henriques, A. L. &amp; Gorayeb, I. S." box="[1020,1313,510,532]" pageId="19" pageNumber="82" pagination="1 - 23" refId="ref17768" refString="Henriques, A. L. &amp; Gorayeb, I. S. (1993) A colecao de Tabanidae Insecta: (Diptera) do Museu Paraense Emilio Goeldi, Belem, Para, Brasil. Goeldiana, Zoologia, 20, 1 - 23." type="journal article" year="1993">Henriques &amp; Gorayeb, 1993</bibRefCitation>
: 19
</treatmentCitation>
(MPEG spp., in part);
<treatmentCitation id="7D6210EA731BFF80FED5D44825D01657" author="Fairchild, G. B. &amp; Burger J. F." box="[341,643,541,563]" page="146" pageId="19" pageNumber="82" year="1994">
<bibRefCitation id="98524B0A731BFF80FED5D44825001657" author="Fairchild, G. B. &amp; Burger J. F." box="[341,595,541,563]" pageId="19" pageNumber="82" pagination="1 - 249" refId="ref17334" refString="Fairchild, G. B. &amp; Burger J. F. (1994) A catalog of the Tabanidae (Diptera) of the Americas south of the United States. Memoirs of the American Entomological Institute, 55, 1 - 249." type="journal article" year="1994">Fairchild &amp; Burger,1994</bibRefCitation>
: 146
</treatmentCitation>
(cat.);
<treatmentCitation id="7D6210EA731BFF80FD4BD44824C81657" author="Henriques, A. L." box="[715,923,541,563]" page="88" pageId="19" pageNumber="82" year="1997">
<bibRefCitation id="98524B0A731BFF80FD4BD44824241657" author="Henriques, A. L." box="[715,887,541,563]" pageId="19" pageNumber="82" pagination="57 - 99" refId="ref17585" refString="Henriques, A. L. (1997) A colecao de Tabanidae (Insecta-Diptera) do Instituto Nacional de Pesquisas da Amazonia (INPA), Manaus, Amazonas, Brasil. Boletim do Museu paraense Emilio Goeldi, serie Zoologia, 11 (1), 57 - 99. [&quot; 1995 &quot;]" type="journal article" year="1997">Henriques, 1997</bibRefCitation>
: 88
</treatmentCitation>
(INPA spp., in part);
<treatmentCitation id="7D6210EA731BFF80FBF7D44827BE1636" author="Coscaron, S. &amp; Papavero, N." page="139" pageId="19" pageNumber="82" year="2009">
<bibRefCitation id="98524B0A731BFF80FBF7D44822CB1657" author="Coscaron, S. &amp; Papavero, N." box="[1143,1432,541,563]" pageId="19" pageNumber="82" pagination="1 - 199" refId="ref16375" refString="Coscaron, S. &amp; Papavero, N. (2009) Catalogue of Neotropical Diptera. Tabanidae. Neotropical Diptera, 16, 1 - 199." type="journal article" year="2009">Coscarón &amp; Papavero, 2009</bibRefCitation>
: 139
</treatmentCitation>
(cat.).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<caption id="A8BC6673731BFF80FF17D13425A913F2" httpUri="https://zenodo.org/record/2623332/files/figure.png" pageId="19" pageNumber="82" startId="19.[151,250,1889,1911]" targetBox="[355,1236,658,1821]" targetPageId="19">
<paragraph id="FC7C36FB731BFF80FF17D13425A913F2" blockId="19.[151,1436,1889,1942]" pageId="19" pageNumber="82">
<emphasis id="CEB7EAE9731BFF80FF17D13426481312" bold="true" box="[151,283,1889,1911]" pageId="19" pageNumber="82">FIGURE 17</emphasis>
.
<emphasis id="CEB7EAE9731BFF80FEAAD13426AA1313" box="[298,505,1889,1911]" italics="true" pageId="19" pageNumber="82">
<taxonomicName id="3BC34D78731BFF80FEAAD13426A61313" box="[298,501,1889,1911]" class="Insecta" family="Tabanidae" genus="Tabanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="19" pageNumber="82" phylum="Arthropoda" rank="species" species="platycerus">Tabanus platycerus</taxonomicName>
.
</emphasis>
Paratype female. A. Dorsal habitus. B. Lateral habitus. C. Head, anterior view. D. Head, lateral view. E. Labels. Scale bars. A, B. 5 mm. C, D. 1 mm.
</paragraph>
</caption>
<subSubSection id="B4D96570731CFF87FF17D6C223C8149C" pageId="20" pageNumber="83" type="diagnosis">
<paragraph id="FC7C36FB731CFF87FF17D6C223C8149C" blockId="20.[151,1437,151,680]" pageId="20" pageNumber="83">
<emphasis id="CEB7EAE9731CFF87FF17D6C2265914D4" bold="true" box="[151,266,151,176]" pageId="20" pageNumber="83">Diagnosis</emphasis>
. Notopleuron integument lighter than adjacent scutum. All coxae, mid and hind femora brownish yellow, pruinescent gray, white haired, hind coxae sometimes with black areas. Middorsal abdominal stripe series of slender connected triangles through tergite 6; dorsolateral disconnected through tergite 4.
</paragraph>
</subSubSection>
<subSubSection id="B4D96570731CFF87FF47D75126D41579" box="[199,391,260,285]" pageId="20" pageNumber="83" type="description">
<paragraph id="FC7C36FB731CFF87FF47D75126D41579" blockId="20.[151,1437,151,680]" box="[199,391,260,285]" pageId="20" pageNumber="83">
<emphasis id="CEB7EAE9731CFF87FF47D75126581579" bold="true" box="[199,267,260,285]" pageId="20" pageNumber="83">Male.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="B4D96570731CFF87FF47D7722666163B" pageId="20" pageNumber="83" type="discussion">
<paragraph id="FC7C36FB731CFF87FF47D7722666163B" blockId="20.[151,1437,151,680]" pageId="20" pageNumber="83">
<emphasis id="CEB7EAE9731CFF87FF47D772261B1524" bold="true" box="[199,328,295,320]" pageId="20" pageNumber="83">Comments</emphasis>
.
<bibRefCitation id="98524B0A731CFF87FED7D772255C1524" author="Fairchild, G. B." box="[343,527,295,320]" pageId="20" pageNumber="83" pagination="237 - 261" refId="ref17120" refString="Fairchild, G. B. (1976) Notes on Neotropical Tabanidae (Dipt.) XVI. The Tabanus trivittatus complex. Studia Entomologica, 19 (1 - 4), 237 - 261." type="journal article" year="1976">Fairchild (1976)</bibRefCitation>
described this species based on specimens from
<collectingCountry id="84D4766B731CFF87FBC0D77D23EF1524" box="[1088,1212,295,320]" name="Costa Rica" pageId="20" pageNumber="83">Costa Rica</collectingCountry>
and
<collectingCountry id="84D4766B731CFF87FB75D77D2202155B" box="[1269,1361,296,319]" name="Panama" pageId="20" pageNumber="83">Panama</collectingCountry>
. Since then, this species has been registered east of the Andes, by
<bibRefCitation id="98524B0A731CFF87FCCAD71922551501" author="Henriques, A. L. &amp; Gorayeb, I. S." box="[842,1286,332,357]" pageId="20" pageNumber="83" pagination="1 - 23" refId="ref17768" refString="Henriques, A. L. &amp; Gorayeb, I. S. (1993) A colecao de Tabanidae Insecta: (Diptera) do Museu Paraense Emilio Goeldi, Belem, Para, Brasil. Goeldiana, Zoologia, 20, 1 - 23." type="journal article" year="1993">
Henriques and Gorayeb (1993;
<collectingCountry id="84D4766B731CFF87FB36D71923AC1501" box="[1206,1279,332,357]" name="Brazil" pageId="20" pageNumber="83">Brazil</collectingCountry>
)
</bibRefCitation>
,
<bibRefCitation id="98524B0A731CFF87FA94D71926F215EC" author="Cardenas, R. E. &amp; Vieira, R. M." pageId="20" pageNumber="83" pagination="153 - 156" refId="ref16048" refString="Cardenas, R. E. &amp; Vieira, R. M. (2005) Nuevas citas de Tabanidos (Diptera: Tabanidae) para Ecuador. Boletin Sociedad Entomologica Aragonesa, 36, 153 - 156" type="journal article" year="2005">
Cárdenas &amp; Vieira (2005;
<collectingCountry id="84D4766B731CFF87FEB7D73A26CA15EC" box="[311,409,367,392]" name="Ecuador" pageId="20" pageNumber="83">Ecuador</collectingCountry>
)
</bibRefCitation>
,
<bibRefCitation id="98524B0A731CFF87FE2ED73A258A15EC" author="Cardenas, R. E. &amp; Buestan J. &amp; Dangles, O." box="[430,729,367,392]" pageId="20" pageNumber="83" pagination="511 - 528" refId="ref16086" refString="Cardenas, R. E., Buestan J. &amp; Dangles, O. (2009) Diversity and distribution models of horse flies (Diptera: Tabanidae) from Ecuador. Annales de la Societe Entomologique de France (n. s.), 45 (4), 511 - 528 + appendix: 31 pp." type="book chapter" year="2009">
Cárdenas (2009;
<collectingCountry id="84D4766B731CFF87FDF3D73A258115EC" box="[627,722,367,392]" name="Ecuador" pageId="20" pageNumber="83">Ecuador</collectingCountry>
)
</bibRefCitation>
and
<bibRefCitation id="98524B0A731CFF87FC95D73A233615EC" author="Henriques, A. L." box="[789,1125,367,392]" pageId="20" pageNumber="83" pagination="151 - 186" refId="ref17724" refString="Henriques, A. L. (2016) Tabanidae (Diptera) of American Museum of Natural History Collection. Zootaxa, 4137 (2), 151 - 186. https: // doi. org / 10.11646 / zootaxa. 4137.2.1" type="journal article" year="2016">
Henriques (2016;
<collectingCountry id="84D4766B731CFF87FC64D73A230F15EC" box="[996,1116,367,392]" name="Venezuela" pageId="20" pageNumber="83">Venezuela</collectingCountry>
)
</bibRefCitation>
. We also examined several specimens from the Amazon Basin at the INPA and Museu Goeldi collections identified as
<emphasis id="CEB7EAE9731CFF87FB52D7C0223E15C9" box="[1234,1389,404,429]" italics="true" pageId="20" pageNumber="83">
<taxonomicName id="3BC34D78731CFF87FB52D7C0223A15C9" box="[1234,1385,404,429]" class="Insecta" family="Tabanidae" genus="Tabanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="20" pageNumber="83" phylum="Arthropoda" rank="species" species="platycerus">T. platycerus</taxonomicName>
.
</emphasis>
All specimens from the collections had lateral abdominal stripes present beyond segment four, reaching the fifth segment and sometimes with vestiges on the sixth, a character not seen in the
<typeStatus id="23788859731CFF87FBADD788230E1591" box="[1069,1117,477,501]" pageId="20" pageNumber="83">type</typeStatus>
series of
<emphasis id="CEB7EAE9731CFF87FB53D788223D1591" box="[1235,1390,476,501]" italics="true" pageId="20" pageNumber="83">
<taxonomicName id="3BC34D78731CFF87FB53D788223A1591" box="[1235,1385,476,501]" class="Insecta" family="Tabanidae" genus="Tabanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="20" pageNumber="83" phylum="Arthropoda" rank="species" species="platycerus">T. platycerus</taxonomicName>
,
</emphasis>
but present in the very similar
<taxonomicName id="3BC34D78731CFF87FE48D454252F167C" authorityName="Gorayeb &amp; Rafael. The" authorityYear="1984" box="[456,636,511,536]" class="Insecta" family="Tabanidae" genus="Tabanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="20" pageNumber="83" phylum="Arthropoda" rank="species" species="sextriangulus">
<emphasis id="CEB7EAE9731CFF87FE48D454252F167C" box="[456,636,511,536]" italics="true" pageId="20" pageNumber="83">T. sextriangulus</emphasis>
</taxonomicName>
. We believe that this might be the case for all species registers east of the Andes, therefore
<taxonomicName id="3BC34D78731CFF87FE08D47025491659" box="[392,538,548,573]" class="Insecta" family="Tabanidae" genus="Tabanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="20" pageNumber="83" phylum="Arthropoda" rank="species" species="platycerus">
<emphasis id="CEB7EAE9731CFF87FE08D47025491659" box="[392,538,548,573]" italics="true" pageId="20" pageNumber="83">T. platycerus</emphasis>
</taxonomicName>
distribution is restricted to Central America, while citations above refers to
<taxonomicName id="3BC34D78731CFF87FA07D470267D1604" authorityName="Gorayeb &amp; Rafael. The" authorityYear="1984" class="Insecta" family="Tabanidae" genus="Tabanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="20" pageNumber="83" phylum="Arthropoda" rank="species" species="sextriangulus">
<emphasis id="CEB7EAE9731CFF87FA07D470267D1604" italics="true" pageId="20" pageNumber="83">T. sextriangulus</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="B4D96570731CFF87FF47D43924E416CC" pageId="20" pageNumber="83" type="materials_examined">
<paragraph id="FC7C36FB731CFF87FF47D43924E416CC" blockId="20.[151,1437,151,680]" pageId="20" pageNumber="83">
<emphasis id="CEB7EAE9731CFF87FF47D43926FE16E1" bold="true" box="[199,429,620,645]" pageId="20" pageNumber="83">Material examined</emphasis>
.
<materialsCitation id="4CAB3CA6731CFF87FE3FD43824F916E0" ID-GBIF-Occurrence="2234227507" box="[447,938,620,645]" collectionCode="FSCA" pageId="20" pageNumber="83" specimenCount="5" specimenCount-female="5" typeStatus="paratype">
<typeStatus id="23788859731CFF87FE3FD438257C16E1" box="[447,559,621,645]" pageId="20" pageNumber="83" type="paratype">Paratypes</typeStatus>
, label as fig. 17 E (
<specimenCount id="EAC5FD72731CFF87FC9ED439241F16E0" box="[798,844,620,645]" pageId="20" pageNumber="83" type="female">5 ♀</specimenCount>
<collectionCode id="9AD2AE3E731CFF87FCD7D43924F116E0" box="[855,930,620,644]" country="USA" httpUri="http://grbio.org/cool/jekt-x82m" name="Florida State Collection of Arthropods, The Museum of Entomology" pageId="20" pageNumber="83" type="Museum">FSCA</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="4CAB3CA6731CFF87FC38D43927A116CC" ID-GBIF-Occurrence="2234227385" collectionCode="MPEG" country="Panama" location="Bocas del Toro" pageId="20" pageNumber="83" specimenCode="INPA 2" specimenCount="1" specimenCount-female="1" stateProvince="Bocas del Toro" typeStatus="paratype">
<collectingCountry id="84D4766B731CFF87FC38D439236116E0" box="[952,1074,620,644]" name="Panama" pageId="20" pageNumber="83">PANAMA</collectingCountry>
,
<emphasis id="CEB7EAE9731CFF87FBC1D43823A316E0" box="[1089,1264,620,645]" italics="true" pageId="20" pageNumber="83">
<collectingRegion id="3E07F819731CFF87FBC1D43823A316E0" box="[1089,1264,620,645]" country="Panama" name="Bocas del Toro" pageId="20" pageNumber="83">Bocas del Toro</collectingRegion>
</emphasis>
(
<specimenCount id="EAC5FD72731CFF87FA87D439224916E0" box="[1287,1306,620,644]" pageId="20" pageNumber="83" type="female">♀</specimenCount>
<specimenCode id="AC659E80731CFF87FAA4D439222F16E0" box="[1316,1404,620,644]" collectionCode="INPA" country="Brazil" httpUri="http://grbio.org/cool/rm88-499z" name="Instituto Nacional de Pesquisas da Amazonia" pageId="20" pageNumber="83">INPA 2</specimenCode>
♀
<collectionCode id="9AD2AE3E731CFF87FF17D4C527B916CC" box="[151,234,656,680]" country="Brazil" httpUri="http://grbio.org/cool/90vz-swjg" name="Museu Paraense Emilio Goeldi" pageId="20" pageNumber="83">MPEG</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="4CAB3CA6731CFF87FF7CD4DA24E016CC" ID-GBIF-Occurrence="2234227472" box="[252,947,655,680]" collectionCode="MCZ" country="Panama" pageId="20" pageNumber="83" specimenCount="1" stateProvince="Bocas del Toro" typeStatus="holotype">
<typeStatus id="23788859731CFF87FF7CD4DA263716CC" box="[252,356,655,680]" pageId="20" pageNumber="83" type="holotype">Holotype</typeStatus>
examined by photos, available at the
<collectionCode id="9AD2AE3E731CFF87FC8BD4C5241416CC" box="[779,839,656,680]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" lsid="urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="20" pageNumber="83" type="Museum">MCZ</collectionCode>
database
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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