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<document id="97E5D90A6628F663F820782BC9FEBC93" ID-CLB-Dataset="27580" ID-DOI="10.11646/zootaxa.4554.1.2" ID-GBIF-Dataset="d41c057e-5b64-4634-84dc-bd5f56a9d789" ID-ISSN="1175-5326" ID-Zenodo-Dep="2623296" ID-ZooBank="986112BE-6ABB-47FE-BA8F-CECEF1C21A5D" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1554270904597" checkinUser="plazi" docAuthor="Carmo, Daniel D. D. &amp; Henriques, Augusto L." docDate="2019" docId="746A87ED7306FF9CFF17D249234C1658" docLanguage="en" docName="zootaxa.4554.1.2.pdf" docOrigin="Zootaxa 4554 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Tabanus macrocerus Carmo &amp; Henriques 2019, sp. n." docType="treatment" docUuid="373C4486-ED69-4885-A9F5-13116C1E7502" docUuidSource="ZooBank" docVersion="10" lastPageNumber="78" masterDocId="8853FF957308FF93FF80D65527531464" masterDocTitle="Taxonomy of Tabanus trivittatus species-group (Diptera: Tabanidae), with description of five new species" masterLastPageNumber="100" masterPageNumber="63" pageNumber="77" updateTime="1698716223463" updateUser="ExternalLinkService">
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<mods:title id="8C836442CD23E0A5D710AD90C2D1A500">Taxonomy of Tabanus trivittatus species-group (Diptera: Tabanidae), with description of five new species</mods:title>
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<mods:namePart id="9E662A367A32594E572DD6923EAF5272">Carmo, Daniel D. D.</mods:namePart>
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<mods:namePart id="1229AB305C5463BE3AC0E23B4759E116">Henriques, Augusto L.</mods:namePart>
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<mods:title id="659AEA0063EDF4AC6BE648CE306A1F82">Zootaxa</mods:title>
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<mods:number id="417BFD602EC8C1A56DC559DE0AECC090">2019-02-08</mods:number>
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<mods:number id="5AAFF7CA843F5A8D8FB5752EBCF64ACA">4554</mods:number>
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<treatment id="746A87ED7306FF9CFF17D249234C1658" ID-DOI="http://doi.org/10.5281/zenodo.5944726" ID-GBIF-Taxon="154534514" ID-Zenodo-Dep="5944726" LSID="urn:lsid:zoobank.org:act:373C4486-ED69-4885-A9F5-13116C1E7502" httpUri="http://treatment.plazi.org/id/746A87ED7306FF9CFF17D249234C1658" lastPageId="15" lastPageNumber="78" pageId="14" pageNumber="77">
<subSubSection id="B4D965707306FF9DFF17D24925B61052" box="[151,741,1052,1078]" pageId="14" pageNumber="77" type="nomenclature">
<paragraph id="FC7C36FB7306FF9DFF17D24925B61052" blockId="14.[151,741,1052,1111]" box="[151,741,1052,1078]" pageId="14" pageNumber="77">
<heading id="A73481977306FF9DFF17D24925B61052" bold="true" box="[151,741,1052,1078]" fontSize="11" level="1" pageId="14" pageNumber="77" reason="1">
<emphasis id="CEB7EAE97306FF9DFF17D24925B61052" bold="true" box="[151,741,1052,1078]" pageId="14" pageNumber="77">
<taxonomicName id="3BC34D787306FF9DFF17D24926C01052" authority="Carmo &amp; Henriques, 2019" authorityName="Carmo &amp; Henriques" authorityYear="2019" box="[151,403,1052,1078]" class="Insecta" family="Tabanidae" genus="Tabanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="77" phylum="Arthropoda" rank="species" species="macrocerus" status="sp. nov.">
<emphasis id="CEB7EAE97306FF9DFF17D24926C01052" bold="true" box="[151,403,1052,1078]" italics="true" pageId="14" pageNumber="77">Tabanus macrocerus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D58457927306FF9DFE1AD249268F1052" box="[410,476,1052,1078]" pageId="14" pageNumber="77" rank="species">sp. n.</taxonomicNameLabel>
Henriques &amp; Carmo
</emphasis>
</heading>
</paragraph>
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<subSubSection id="B4D965707306FF9DFF17D26B26D21033" box="[151,385,1086,1111]" pageId="14" pageNumber="77" type="description">
<paragraph id="FC7C36FB7306FF9DFF17D26B26D21033" blockId="14.[151,741,1052,1111]" box="[151,385,1086,1111]" pageId="14" pageNumber="77">
(
<figureCitation id="64F82A7E7306FF9DFF1ED26B26461033" box="[158,277,1086,1111]" captionStart="FIGURE 13" captionStartId="15.[151,250,1616,1638]" captionTargetBox="[362,1229,637,1595]" captionTargetId="figure@15.[340,1247,628,1595]" captionTargetPageId="15" captionText="FIGURE 13. Tabanus macrocerus sp. n. Holotype female. A. Dorsal habitus. B. Lateral habitus. C. Head, anterior view. D. Head, lateral view. Scale bars. A, B. 5 mm. C, D. 1 mm." httpUri="https://zenodo.org/record/2623324/files/figure.png" pageId="14" pageNumber="77">Figures 13</figureCitation>
AD, 36)
</paragraph>
</subSubSection>
<subSubSection id="B4D965707306FF9DFF17D2D3246B114B" pageId="14" pageNumber="77" type="diagnosis">
<paragraph id="FC7C36FB7306FF9DFF17D2D3246B114B" blockId="14.[151,1437,1158,2014]" pageId="14" pageNumber="77">
<emphasis id="CEB7EAE97306FF9DFF17D2D3265910FB" bold="true" box="[151,266,1158,1183]" pageId="14" pageNumber="77">Diagnosis</emphasis>
. A brownish species with postpedicel long, about two times the style length. Notopleuron integument paler than adjacent scutum. All coxae, mid and hind femora yellowish. Abdominal median dorsal stripe a series of yellow connected triangles, sometimes apparently even. Dorsolateral stripes a series of disconnected dashes on tergites 1-4, sometimes barely visible on tergite 5. Abdomen ventrally banded and without a median black stripe, if present, inconspicuous and not reaching the sternite margin.
</paragraph>
</subSubSection>
<subSubSection id="B4D965707306FF9CFF47D36F23AD149C" lastPageId="15" lastPageNumber="78" pageId="14" pageNumber="77" type="materials_examined">
<paragraph id="FC7C36FB7306FF9DFF47D36F2503124F" blockId="14.[151,1437,1158,2014]" pageId="14" pageNumber="77">
<emphasis id="CEB7EAE97306FF9DFF47D36F26C01137" bold="true" box="[199,403,1338,1363]" pageId="14" pageNumber="77">
<typeStatus id="237888597306FF9DFF47D36F26671137" box="[199,308,1338,1363]" pageId="14" pageNumber="77" type="holotype">Holotype</typeStatus>
female.
</emphasis>
Length
<quantity id="3B3B9B1E7306FF9DFE74D36E251C1136" box="[500,591,1339,1363]" metricMagnitude="-3" metricUnit="m" metricValue="9.2" pageId="14" pageNumber="77" unit="mm" value="9.2">9.2 mm</quantity>
, wing
<quantity id="3B3B9B1E7306FF9DFD1DD36E25AB1136" box="[669,760,1339,1363]" metricMagnitude="-3" metricUnit="m" metricValue="7.9" pageId="14" pageNumber="77" unit="mm" value="7.9">7.9 mm</quantity>
. Eyes bare. Frons pruinescent yellow, divergent above, F.I. 7.5, D.I. 1.96. Callus brown, higher than wide. Median callus black surrounded by dark pruinescence and hairs. Subcallus inflated, dark brown, paler in middle. Clypeus and gena pale gray with white hairs, except for brownish pruinescence and hairs on upper gena near antennal base. First antennal segment pale brown, dark pruinescent above, second and third segments orange, pale pruinescent, the style only slightly darker. Palpus with yellowish integument, gray pruinescence and black hairs, white hairs on basal half, mainly on anterior and posterior margins. Proboscis black, dark gray pruinescent.
</paragraph>
<paragraph id="FC7C36FB7306FF9DFF47D06226F513F2" blockId="14.[151,1437,1158,2014]" pageId="14" pageNumber="77">
Scutum and scutellum grayish brown pruinescent, recumbent golden hairs and somewhat erect black hairs mixed, integument black, except supra-alar area brown. Notopleuron paler, brown with golden hairs on upper half and black on bottom half. Pleuron with black integument, largely yellowish pruinescent and white hairs, except upper surface of katepisternum with yellowish integument and anepisternum largely with yellowish hairs. All coxae with integument yellowish and white hairs. Fore femur blackish with black hairs. Fore tibia heavy bicolored, the basal 2/3 white with white hairs, the remainder blackish with black hairs. Fore tarsus blackish with black hairs. Mid and hind femora yellowish with white hairs, mid femur with black hairs on anterior border. Mid tibia yellowish with white hairs, black hairs at apex. Hind tibia yellowish with white hairs, black hairs at posterior border and apex. Mid and hind tarsi brown with black hairs. Wing lightly infuscated on r
<subScript id="604734BE7306FF9DFBFAD13023D11317" attach="left" box="[1146,1154,1893,1907]" fontSize="6" pageId="14" pageNumber="77">1</subScript>
and r
<subScript id="604734BE7306FF9DFB42D13023891317" attach="left" box="[1218,1242,1893,1907]" fontSize="6" pageId="14" pageNumber="77">2+3</subScript>
cells with brown veins, stigma yellowish.
</paragraph>
<paragraph id="FC7C36FB7306FF9CFF47D1F423AD149C" blockId="14.[151,1437,1158,2014]" lastBlockId="15.[151,1436,151,573]" lastPageId="15" lastPageNumber="78" pageId="14" pageNumber="77">Abdomen brown, darkening to the apex. Median stripe series of continuous yellowish triangles of pruinescence and hairs on tergites 16. Dorsolateral stripesseries of disconnected yellowish pruinescent dashes on tergites 14, barely visible on tergite 5. On second tergite, the dash is largest and prominent. Sides of tergites paler and white haired. Sternites pale brown largely white hairs, but black hairs on anterior margin of sternites 35, more abundant on sternite 6 and wholly on sternite 7. Anterior border of sternites 26 whitish pruinescent.</paragraph>
</subSubSection>
<subSubSection id="B4D965707307FF9CFF47D75126D41579" box="[199,391,260,285]" pageId="15" pageNumber="78" type="description">
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<emphasis id="CEB7EAE97307FF9CFF47D75126581579" bold="true" box="[199,267,260,285]" pageId="15" pageNumber="78">Male.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="B4D965707307FF9CFF47D77225FB155B" box="[199,680,295,320]" pageId="15" pageNumber="78" type="discussion">
<paragraph id="FC7C36FB7307FF9CFF47D77225FB155B" blockId="15.[151,1436,151,573]" box="[199,680,295,320]" pageId="15" pageNumber="78">
<emphasis id="CEB7EAE97307FF9CFF47D772261C1524" bold="true" box="[199,335,295,320]" pageId="15" pageNumber="78">Comments.</emphasis>
See under
<taxonomicName id="3BC34D787307FF9CFE4CD77C25361524" authority="Carmo &amp; Henriques, 2019" authorityName="Carmo &amp; Henriques" authorityYear="2019" box="[460,613,295,320]" class="Insecta" family="Tabanidae" genus="Tabanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="15" pageNumber="78" phylum="Arthropoda" rank="species" species="mackerrasi" status="sp. nov.">
<emphasis id="CEB7EAE97307FF9CFE4CD77C25361524" box="[460,613,295,320]" italics="true" pageId="15" pageNumber="78">T. mackerrasi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D58457927307FF9CFDECD77D25FB155B" box="[620,680,296,319]" pageId="15" pageNumber="78" rank="species">sp. n.</taxonomicNameLabel>
</paragraph>
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<emphasis id="CEB7EAE97307FF9CFF47D719263C1501" bold="true" box="[199,367,332,357]" pageId="15" pageNumber="78">Type material</emphasis>
.
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collected by
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&amp;
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female:
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,
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,
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,
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45,
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04°21
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51
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S
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50°01
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W
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<collectingDate id="9839E9D37307FF9CFDCAD73A259715EC" box="[586,708,367,392]" pageId="15" pageNumber="78" value="2001-11-27">27.xi.2001</collectingDate>
</date>
, biting in horse (
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);
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: same data of the
<typeStatus id="237888597307FF9CFA94D73A222515EC" box="[1300,1398,367,392]" pageId="15" pageNumber="78" type="holotype">holotype</typeStatus>
(
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,
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<collectionCode id="9AD2AE3E7307FF9CFEA1D7C1263915C8" box="[289,362,404,428]" country="USA" httpUri="http://grbio.org/cool/jekt-x82m" name="Florida State Collection of Arthropods, The Museum of Entomology" pageId="15" pageNumber="78" type="Museum">FSCA</collectionCode>
,
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<collectionCode id="9AD2AE3E7307FF9CFE32D7C126AC15C8" box="[434,511,404,428]" country="Brazil" httpUri="http://grbio.org/cool/90vz-swjg" name="Museu Paraense Emilio Goeldi" pageId="15" pageNumber="78">MPEG</collectionCode>
,
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<collectionCode id="9AD2AE3E7307FF9CFDC7D7C125F615C8" box="[583,677,404,428]" pageId="15" pageNumber="78">CEUFT</collectionCode>
);
<emphasis id="CEB7EAE97307FF9CFD3CD7C125A115C9" box="[700,754,404,429]" italics="true" pageId="15" pageNumber="78">idem</emphasis>
,
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<collectingDate id="9839E9D37307FF9CFC84D7C1242E15C9" box="[772,893,404,429]" pageId="15" pageNumber="78" value="2001-11-29">29.xi.2001</collectingDate>
</date>
(
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<collectionCode id="9AD2AE3E7307FF9CFC31D7C1234015C8" box="[945,1043,404,428]" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="15" pageNumber="78">MZUSP</collectionCode>
),
<collectingCounty id="151D4E777307FF9CFBA9D7C0225D15C9" box="[1065,1294,404,429]" pageId="15" pageNumber="78">Novo Repartimento</collectingCounty>
,
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0 8,
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<geoCoordinate id="99F7503C7307FF9CFEA3D7E226E715B4" box="[291,436,439,464]" direction="west" orientation="longitude" pageId="15" pageNumber="78" precision="15" value="-49.906944">49°5425″W</geoCoordinate>
,
<date id="887D103B7307FF9CFE38D7E2257F15B4" box="[440,556,439,464]" pageId="15" pageNumber="78" value="2001-11-28">
<collectingDate id="9839E9D37307FF9CFE38D7E2257F15B4" box="[440,556,439,464]" pageId="15" pageNumber="78" value="2001-11-28">28.xi.2001</collectingDate>
</date>
,
<location id="F91C60207307FF9CFDBDD7ED25CB15B4" LSID="urn:lsid:plazi:treatment:746A87ED7306FF9CFF17D249234C1658:F91C60207307FF9CFDBDD7ED25CB15B4" box="[573,664,440,464]" country="Brazil" county="Novo Repartimento" latitude="-4.9833336" longLatPrecision="1" longitude="-48.133335" municipality="Vicinal" name="Tucurui" pageId="15" pageNumber="78" stateProvince="Maranhao">Tucuruí</location>
,
<location id="F91C60207307FF9CFD27D7ED243415B4" LSID="urn:lsid:plazi:treatment:746A87ED7306FF9CFF17D249234C1658:F91C60207307FF9CFD27D7ED243415B4" box="[679,871,439,464]" country="Brazil" county="Novo Repartimento" latitude="-4.9833336" longLatPrecision="1" longitude="-48.133335" municipality="Vicinal" name="Faz. do Senador" pageId="15" pageNumber="78" stateProvince="Maranhao">Faz. do Senador</location>
,
<geoCoordinate id="99F7503C7307FF9CFCF6D7E224AE15B4" box="[886,1021,439,464]" direction="south" orientation="latitude" pageId="15" pageNumber="78" precision="15" value="-3.9966667">
03°59
<emphasis id="CEB7EAE97307FF9CFC39D7E2249215B4" bold="true" box="[953,961,439,464]" pageId="15" pageNumber="78"></emphasis>
48
<emphasis id="CEB7EAE97307FF9CFC5DD7E224BF15B4" bold="true" box="[989,1004,439,464]" pageId="15" pageNumber="78"></emphasis>
S
</geoCoordinate>
,
<geoCoordinate id="99F7503C7307FF9CFB8ED7E223F315AB" box="[1038,1184,439,464]" direction="west" orientation="longitude" pageId="15" pageNumber="78" precision="1" value="-49.75">
49°45
<emphasis id="CEB7EAE97307FF9CFBD2D7E2230915B4" bold="true" box="[1106,1114,439,464]" pageId="15" pageNumber="78"></emphasis>
0 3
<emphasis id="CEB7EAE97307FF9CFBF5D7E223D715B4" bold="true" box="[1141,1156,439,464]" pageId="15" pageNumber="78"></emphasis>
W
</geoCoordinate>
,
<date id="887D103B7307FF9CFB2FD7E2227A15B4" box="[1199,1321,439,464]" pageId="15" pageNumber="78" value="2001-11-30">
<collectingDate id="9839E9D37307FF9CFB2FD7E2227A15B4" box="[1199,1321,439,464]" pageId="15" pageNumber="78" value="2001-11-30">30.xi.2001</collectingDate>
</date>
, biting in horse (
<specimenCount id="EAC5FD727307FF9CFF66D78926471590" box="[230,276,476,500]" pageId="15" pageNumber="78" type="female">2 ♀</specimenCount>
<collectionCode id="9AD2AE3E7307FF9CFE9ED78926311590" box="[286,354,476,500]" country="Brazil" httpUri="http://grbio.org/cool/rm88-499z" name="Instituto Nacional de Pesquisas da Amazonia" pageId="15" pageNumber="78">INPA</collectionCode>
);
<location id="F91C60207307FF9CFEF8D78926861591" LSID="urn:lsid:plazi:treatment:746A87ED7306FF9CFF17D249234C1658:F91C60207307FF9CFEF8D78926861591" box="[376,469,476,501]" country="Brazil" county="Novo Repartimento" latitude="-4.9833336" longLatPrecision="1" longitude="-48.133335" municipality="Vicinal" name="Jacunda" pageId="15" pageNumber="78" stateProvince="Maranhao">Jacundá</location>
,
<location id="F91C60207307FF9CFE63D78825391590" LSID="urn:lsid:plazi:treatment:746A87ED7306FF9CFF17D249234C1658:F91C60207307FF9CFE63D78825391590" box="[483,618,477,500]" country="Brazil" county="Novo Repartimento" latitude="-4.9833336" longLatPrecision="1" longitude="-48.133335" municipality="Vicinal" name="Porto Novo" pageId="15" pageNumber="78" stateProvince="Maranhao">Porto Novo</location>
,
<date id="887D103B7307FF9CFDF9D78925A01591" box="[633,755,476,501]" pageId="15" pageNumber="78" value="2001-11-30">
<collectingDate id="9839E9D37307FF9CFDF9D78925A01591" box="[633,755,476,501]" pageId="15" pageNumber="78" value="2001-11-30">30.xi.2001</collectingDate>
</date>
, biting in horse (
<specimenCount id="EAC5FD727307FF9CFC3ED78924821590" box="[958,977,476,500]" pageId="15" pageNumber="78" type="female"></specimenCount>
<collectionCode id="9AD2AE3E7307FF9CFC5BD789234C1590" box="[987,1055,476,500]" country="Brazil" httpUri="http://grbio.org/cool/rm88-499z" name="Instituto Nacional de Pesquisas da Amazonia" pageId="15" pageNumber="78">INPA</collectionCode>
),
<emphasis id="CEB7EAE97307FF9CFBB4D78923F91591" box="[1076,1194,476,501]" italics="true" pageId="15" pageNumber="78">
<collectingRegion id="3E07F8197307FF9CFBB4D78923F91591" box="[1076,1194,476,501]" country="Brazil" name="Maranhao" pageId="15" pageNumber="78">Maranhão</collectingRegion>
</emphasis>
,
<location id="F91C60207307FF9CFB3CD78927BA1673" LSID="urn:lsid:plazi:treatment:746A87ED7306FF9CFF17D249234C1658:F91C60207307FF9CFB3CD78927BA1673" country="Brazil" county="Novo Repartimento" latitude="-4.9833336" longLatPrecision="1" longitude="-48.133335" municipality="Vicinal" name="Sao Pedro da Agua Branca" pageId="15" pageNumber="78" stateProvince="Maranhao">São Pedro da Água Branca</location>
,
<geoCoordinate id="99F7503C7307FF9CFF74D7AA262F167C" box="[244,380,511,536]" direction="south" orientation="latitude" pageId="15" pageNumber="78" precision="1" value="-4.9833336">
04°59
<emphasis id="CEB7EAE97307FF9CFEB7D7AA266C167C" bold="true" box="[311,319,511,536]" pageId="15" pageNumber="78"></emphasis>
0 5
<emphasis id="CEB7EAE97307FF9CFEDBD7AA2639167C" bold="true" box="[347,362,511,536]" pageId="15" pageNumber="78"></emphasis>
S
</geoCoordinate>
,
<geoCoordinate id="99F7503C7307FF9CFE08D45525491673" box="[392,538,511,536]" direction="west" orientation="longitude" pageId="15" pageNumber="78" precision="1" value="-48.133335">
48°08
<emphasis id="CEB7EAE97307FF9CFE4BD7AA2680167C" bold="true" box="[459,467,511,536]" pageId="15" pageNumber="78"></emphasis>
0 3
<emphasis id="CEB7EAE97307FF9CFE6FD7AA26AD167C" bold="true" box="[495,510,511,536]" pageId="15" pageNumber="78"></emphasis>
W
</geoCoordinate>
,
<date id="887D103B7307FF9CFDA4D7AA25F5167C" box="[548,678,511,536]" pageId="15" pageNumber="78" value="2001-12-07">
<collectingDate id="9839E9D37307FF9CFDA4D7AA25F5167C" box="[548,678,511,536]" pageId="15" pageNumber="78" value="2001-12-07">07.xii.2001</collectingDate>
</date>
,
<collectingMethod id="25824EEC7307FF9CFD32D7AA24681673" box="[690,827,511,536]" pageId="15" pageNumber="78">malaise trap</collectingMethod>
(
<specimenCount id="EAC5FD727307FF9CFCCCD455240C167C" box="[844,863,512,536]" pageId="15" pageNumber="78" type="female"></specimenCount>
<collectionCode id="9AD2AE3E7307FF9CFCE7D45524EE167C" box="[871,957,512,536]" pageId="15" pageNumber="78">CZMA</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="B4D965707307FF9CFF4ED471234C1658" box="[206,1055,548,573]" pageId="15" pageNumber="78" type="etymology">
<paragraph id="FC7C36FB7307FF9CFF4ED471234C1658" blockId="15.[151,1436,151,573]" box="[206,1055,548,573]" pageId="15" pageNumber="78">
<emphasis id="CEB7EAE97307FF9CFF4ED47126061659" bold="true" box="[206,341,548,573]" pageId="15" pageNumber="78">Etymology.</emphasis>
From Greek,
<emphasis id="CEB7EAE97307FF9CFE71D47125101659" box="[497,579,548,573]" italics="true" pageId="15" pageNumber="78">makros</emphasis>
(large) +
<emphasis id="CEB7EAE97307FF9CFD32D47125BD1659" box="[690,750,548,573]" italics="true" pageId="15" pageNumber="78">keros</emphasis>
(horn, relative to antenna).
</paragraph>
</subSubSection>
</treatment>
</document>