238 lines
28 KiB
XML
238 lines
28 KiB
XML
<document id="399148FB03DB40A712AD4B6F34E63361" ID="10.11646/zootaxa.4232.1.6SLASH10169" ID-CLB-Dataset="36544" ID-DOI="10.11646/zootaxa.4232.1.6" ID-GBIF-Dataset="86aa9515-2068-4285-838a-e73c8b8aaeb7" ID-ISSN="1175-5326" ID-Zenodo-Dep="293098" ID-ZooBank="9D76AB7D-0553-4A90-8FA8-C70153B705AE" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1487148507536" checkinUser="plazi" docAuthor="Calcetas, Orlando A. & Adorada, Jessamyn R." docDate="2017" docId="502A6F54FFCEFFECFF1CFB10FC89A1EE" docLanguage="en" docName="zootaxa.4232.1.6SLASH10169.pdf" docOrigin="Zootaxa 4232 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Leucopholis bakeri Moser 1924" docType="treatment" docVersion="9" lastPageNumber="92" masterDocId="AC13172CFFCBFFE4FF8BFFFCFF81A116" masterDocTitle="Taxonomic review of the genus Leucopholis Dejean, 1833 (Coleoptera: Scarabaeidae: Melolonthinae: Leucopholini) in the Philippines" masterLastPageNumber="103" masterPageNumber="85" pageNumber="90" updateTime="1698436935605" updateUser="ExternalLinkService">
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<mods:title id="39D77A8AC354A551BEA05915FA1D9133">Taxonomic review of the genus Leucopholis Dejean, 1833 (Coleoptera: Scarabaeidae: Melolonthinae: Leucopholini) in the Philippines</mods:title>
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<mods:namePart id="D19235B5A6EA7E72848D217E4581A4D0">Calcetas, Orlando A.</mods:namePart>
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<mods:namePart id="5402D20012823780A8CE49F494132896">Adorada, Jessamyn R.</mods:namePart>
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<mods:typeOfResource id="7BA42E9F743701F60D0468609E97E31F">text</mods:typeOfResource>
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<mods:title id="5CC4D20E66FD23FC12712044509A4CA5">Zootaxa</mods:title>
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<mods:part id="F3FE94DB42D4BF55B9E0F19194B40449">
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<mods:date id="704029348070B88C4BB5B32A0D26D36B">2017</mods:date>
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<mods:number id="E809FCCC06B8F811B53FAD26165DAB3C">4232</mods:number>
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<mods:identifier id="C305933693E6171A0537C3A01D05C56A" type="DOI">10.11646/zootaxa.4232.1.6</mods:identifier>
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<treatment id="502A6F54FFCEFFECFF1CFB10FC89A1EE" ID-DOI="http://doi.org/10.5281/zenodo.6000493" ID-GBIF-Taxon="127695498" ID-Zenodo-Dep="6000493" LSID="urn:lsid:plazi:treatment:502A6F54FFCEFFECFF1CFB10FC89A1EE" httpUri="http://treatment.plazi.org/id/502A6F54FFCEFFECFF1CFB10FC89A1EE" lastPageId="8" lastPageNumber="92" pageId="5" pageNumber="90">
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<subSubSection id="90998DC9FFCEFFE1FF1CFB10FE9AA43E" pageId="5" pageNumber="90" type="nomenclature">
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<paragraph id="D83CDE42FFCEFFE1FF1CFB10FDA6A411" blockId="5.[151,551,1260,1320]" box="[151,551,1260,1287]" pageId="5" pageNumber="90">
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<heading id="8374692EFFCEFFE1FF1CFB10FDA6A411" bold="true" box="[151,551,1260,1287]" fontSize="11" level="1" pageId="5" pageNumber="90" reason="1">
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<taxonomicName id="1F83A5C1FFCEFFE1FF1CFB10FDA6A411" authority="Moser , 1924" authorityName="Moser" authorityYear="1924" box="[151,551,1260,1287]" class="Insecta" family="Melolonthidae" genus="Leucopholis" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="90" phylum="Arthropoda" rank="species" species="bakeri">
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<emphasis id="EAF70250FFCEFFE1FF1CFB10FDA6A411" bold="true" box="[151,551,1260,1287]" pageId="5" pageNumber="90">
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<emphasis id="EAF70250FFCEFFE1FF1CFB10FEFEA410" bold="true" box="[151,383,1260,1286]" italics="true" pageId="5" pageNumber="90">Leucopholis bakeri</emphasis>
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<bibRefCitation id="BC12A3B3FFCEFFE1FE0DFB11FDA6A411" author="Moser" box="[390,551,1261,1287]" pageId="5" pageNumber="90" refString="Moser, J. (1924) New species of melolonthid beetles from the Philippine Islands. Philippine Journal of Science, 25, 349 - 357." type="journal article" year="1924">Moser, 1924</bibRefCitation>
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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<paragraph id="D83CDE42FFCEFFE1FF1CFAF3FE9AA43E" blockId="5.[151,551,1260,1320]" box="[151,283,1295,1320]" pageId="5" pageNumber="90">
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<figureCitation id="40B8C2C7FFCEFFE1FF1CFAF3FE9AA43E" box="[151,283,1295,1320]" captionStart-0="FIGURE 12 – 16" captionStart-1="FIGURE 17 – 21" captionStartId-0="6.[151,250,1897,1919]" captionStartId-1="7.[151,250,1726,1748]" captionTargetBox-0="[259,719,1204,1838]" captionTargetPageId-0="6" captionText-0="FIGURE 12 – 16. Leucopholis reflexa Moser and L. bakeri Moser. 12, Leucopholis reflexa habitus, dorsal aspect; 13, Leucopholis reflexa habitus, dorsal aspect (photograph of the syntype from ZMHB, courtesy of Dr. Joachim Willers); 14. Leucopholis bakeri clypeus and labrum, male, apical aspect; 15. Leucopholis bakeri protibia; 19, Leucopholis bakeri clypeus and labrum, dorsal aspect." captionText-1="FIGURE 17 – 21. Leucopholis bakeri Moser; 16, male genitalia, dorsal aspect; 17, male genitalia, ventral aspect; 18, male genitalia, lateral aspect; 20, habitus, dorsal aspect; 21, habitus, dorsal aspect (photograph of a syntype from ZMHB courtesy of Dr. Joachim Willers)." httpUri-0="https://zenodo.org/record/293162/files/figure.png" pageId="5" pageNumber="90">Figs. 14–21</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="90998DC9FFCEFFE1FF1CFAA4FD98A478" box="[151,537,1368,1390]" pageId="5" pageNumber="90" type="reference_group">
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<paragraph id="D83CDE42FFCEFFE1FF1CFAA4FD98A478" blockId="5.[151,537,1368,1390]" box="[151,537,1368,1390]" pageId="5" pageNumber="90">
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<treatmentCitationGroup id="F893F96CFFCEFFE1FF1CFAA4FD98A478" box="[151,537,1368,1390]" pageId="5" pageNumber="90">
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<taxonomicName id="1F83A5C1FFCEFFE1FF1CFAA4FD94A478" authority="Moser, 1924: 354" authorityName="Moser" authorityPageNumber="354" authorityYear="1924" box="[151,533,1368,1390]" class="Insecta" family="Melolonthidae" genus="Leucopholis" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="90" phylum="Arthropoda" rank="species" species="bakeri">
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<emphasis id="EAF70250FFCEFFE1FF1CFAA4FEDBA478" box="[151,346,1368,1390]" italics="true" pageId="5" pageNumber="90">Leucopholis bakeri</emphasis>
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<treatmentCitation id="5922F853FFCEFFE1FEEBFAA5FD94A478" author="Moser" box="[352,533,1368,1390]" page="354" pageId="5" pageNumber="90" year="1924">
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<bibRefCitation id="BC12A3B3FFCEFFE1FEEBFAA5FE63A478" author="Moser" box="[352,482,1368,1390]" pageId="5" pageNumber="90" refString="Moser, J. (1924) New species of melolonthid beetles from the Philippine Islands. Philippine Journal of Science, 25, 349 - 357." type="journal article" year="1924">Moser, 1924</bibRefCitation>
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: 354
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</treatmentCitation>
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</taxonomicName>
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.
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</treatmentCitationGroup>
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</paragraph>
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||
</subSubSection>
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<subSubSection id="90998DC9FFCEFFE3FF1CFA66FF6EA68C" lastPageId="7" lastPageNumber="92" pageId="5" pageNumber="90" type="description">
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<paragraph id="D83CDE42FFCEFFE1FF1CFA66FA85A670" blockId="5.[151,1437,1434,2038]" pageId="5" pageNumber="90">
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<emphasis id="EAF70250FFCEFFE1FF1CFA66FEA8A4A5" bold="true" box="[151,297,1434,1459]" pageId="5" pageNumber="90">Description.</emphasis>
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Male: length
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<quantity id="1F7B73A7FFCEFFE1FE44FA66FDB8A4A4" box="[463,569,1434,1459]" metricMagnitude="-2" metricUnit="m" metricValue="2.25" pageId="5" pageNumber="90" unit="mm" value="22.5">22.5 mm</quantity>
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, width 11.0 mm; female: length
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<quantity id="1F7B73A7FFCEFFE1FC31FA66FBA5A4A4" box="[954,1060,1434,1459]" metricMagnitude="-2" metricUnit="m" metricValue="2.51" pageId="5" pageNumber="90" unit="mm" value="25.1">25.1 mm</quantity>
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, width
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<quantity id="1F7B73A7FFCEFFE1FBF6FA67FB69A4A4" box="[1149,1256,1435,1459]" metricMagnitude="-2" metricUnit="m" metricValue="1.23" pageId="5" pageNumber="90" unit="mm" value="12.3">12.3 mm</quantity>
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. Dorsal surface uniformally dark reddish brown. Clypeus anterior margin without median cleft in both sexes, weakly convex anteriorly, nearly as long as labrum in both sexes (
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<figureCitation id="40B8C2C7FFCEFFE1FD75FA19FCCBA4E8" box="[766,842,1509,1534]" captionStart="FIGURE 12 – 16" captionStartId="6.[151,250,1897,1919]" captionTargetBox="[259,719,1204,1838]" captionTargetPageId="6" captionText="FIGURE 12 – 16. Leucopholis reflexa Moser and L. bakeri Moser. 12, Leucopholis reflexa habitus, dorsal aspect; 13, Leucopholis reflexa habitus, dorsal aspect (photograph of the syntype from ZMHB, courtesy of Dr. Joachim Willers); 14. Leucopholis bakeri clypeus and labrum, male, apical aspect; 15. Leucopholis bakeri protibia; 19, Leucopholis bakeri clypeus and labrum, dorsal aspect." httpUri="https://zenodo.org/record/293162/files/figure.png" pageId="5" pageNumber="90">Fig.14</figureCitation>
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||
), inclined backward at approximately 60 0, weakly indented laterally. Head with dorsal surface rugosely punctate and covered with recumbent circular to ovoid scales. Antennal club length
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||
<quantity id="1F7B73A7FFCEFFE1FE07F9D2FE51A753" box="[396,464,1582,1606]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="5" pageNumber="90" unit="mm" value="2.0">2 mm</quantity>
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, longer than antenomeres II–VII. Pronotum anterior margin thickly margined mesally and thinly margined laterally. Anterior pronotal angle obtuse and rounded. Posterior pronotal angle square, with apical lobe; surface covered with recumbent, circular to ovoid, apically pointed, basally rounded or truncate scales; surface also with sparse apically and basally blunt spindle to lanceolate scales adjacent to posterior margin. Protibial spur extending in front of anterior angle of protibia in both sexes (
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<figureCitation id="40B8C2C7FFCEFFE1FC56F941FBB1A7C0" box="[989,1072,1725,1750]" captionStart="FIGURE 12 – 16" captionStartId="6.[151,250,1897,1919]" captionTargetBox="[259,719,1204,1838]" captionTargetPageId="6" captionText="FIGURE 12 – 16. Leucopholis reflexa Moser and L. bakeri Moser. 12, Leucopholis reflexa habitus, dorsal aspect; 13, Leucopholis reflexa habitus, dorsal aspect (photograph of the syntype from ZMHB, courtesy of Dr. Joachim Willers); 14. Leucopholis bakeri clypeus and labrum, male, apical aspect; 15. Leucopholis bakeri protibia; 19, Leucopholis bakeri clypeus and labrum, dorsal aspect." httpUri="https://zenodo.org/record/293162/files/figure.png" pageId="5" pageNumber="90">Fig. 15</figureCitation>
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). Metaventral process length
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<quantity id="1F7B73A7FFCEFFE1FAF2F942FF47A7EF" metricMagnitude="-3" metricUnit="m" metricValue="1.4" pageId="5" pageNumber="90" unit="mm" value="1.4">1.4 mm</quantity>
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. Metaventrite densely setose, scales ovoid to lanceolate. Posterior metatibia with 15–15 spicules. Anterior margin of elytra with distinct groove at anterior angle and terminating at the middle, covered with scales identical to those on pronotum. Abdominal sternites covered with basally round or truncate ovoid scales. Abdominal sternite V laterally on the same plane with the other segments. Pygidium covered with spindle-shaped scales.
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</paragraph>
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<paragraph id="D83CDE42FFCEFFE2FF4CF88DFB71A1C3" blockId="5.[151,1437,1434,2038]" lastBlockId="6.[151,1437,151,395]" lastPageId="6" lastPageNumber="91" pageId="5" pageNumber="90">
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<emphasis id="EAF70250FFCEFFE1FF4CF88DFE85A69C" bold="true" box="[199,260,1905,1930]" pageId="5" pageNumber="90">Male</emphasis>
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. Genitalia length
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<quantity id="1F7B73A7FFCEFFE1FE59F88EFD96A69F" box="[466,535,1906,1930]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" pageId="5" pageNumber="90" unit="mm" value="8.0">8 mm</quantity>
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. Phallobase dorsal anterior margin with narrow V-shaped margin mesally (
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<figureCitation id="40B8C2C7FFCEFFE1FAE4F88DFF37A6B8" captionStart="FIGURE 12 – 16" captionStartId="6.[151,250,1897,1919]" captionTargetBox="[259,719,1204,1838]" captionTargetPageId="6" captionText="FIGURE 12 – 16. Leucopholis reflexa Moser and L. bakeri Moser. 12, Leucopholis reflexa habitus, dorsal aspect; 13, Leucopholis reflexa habitus, dorsal aspect (photograph of the syntype from ZMHB, courtesy of Dr. Joachim Willers); 14. Leucopholis bakeri clypeus and labrum, male, apical aspect; 15. Leucopholis bakeri protibia; 19, Leucopholis bakeri clypeus and labrum, dorsal aspect." httpUri="https://zenodo.org/record/293162/files/figure.png" pageId="5" pageNumber="90">Fig. 16</figureCitation>
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). Paramere ventral inner posterior margin elongate, spindle-shaped, and bordered by membranous integument (
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<figureCitation id="40B8C2C7FFCEFFE1FF14F845FF79A6C4" box="[159,248,1977,2002]" captionStart="FIGURE 17 – 21" captionStartId="7.[151,250,1726,1748]" captionText="FIGURE 17 – 21. Leucopholis bakeri Moser; 16, male genitalia, dorsal aspect; 17, male genitalia, ventral aspect; 18, male genitalia, lateral aspect; 20, habitus, dorsal aspect; 21, habitus, dorsal aspect (photograph of a syntype from ZMHB courtesy of Dr. Joachim Willers)." pageId="5" pageNumber="90">Fig. 17</figureCitation>
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). Mid-posterior margin of parameres triangulate and apically round. Dorsal mid-posterior margin of parameres bell shaped. Inner anterior margin of parameres spindle shaped. Parameres laterally with longitudinal groove on mesal half to the dorsum and bounded by carinae on both sides. Lateral groove wider sub-apically and mesally narrow, posterior margin subcircle (
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<figureCitation id="40B8C2C7FFCDFFE2FD0EFF40FD59A1C2" box="[645,728,188,213]" captionStart="FIGURE 17 – 21" captionStartId="7.[151,250,1726,1748]" captionText="FIGURE 17 – 21. Leucopholis bakeri Moser; 16, male genitalia, dorsal aspect; 17, male genitalia, ventral aspect; 18, male genitalia, lateral aspect; 20, habitus, dorsal aspect; 21, habitus, dorsal aspect (photograph of a syntype from ZMHB courtesy of Dr. Joachim Willers)." pageId="6" pageNumber="91">Fig. 18</figureCitation>
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). Apical processes wider and lobe-like apically.
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||
</paragraph>
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||
<paragraph id="D83CDE42FFCDFFE2FF4CFF1EFDD6A09D" blockId="6.[151,1437,151,395]" pageId="6" pageNumber="91">
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||
<emphasis id="EAF70250FFCDFFE2FF4CFF1EFE9FA1ED" bold="true" box="[199,286,226,251]" pageId="6" pageNumber="91">Female</emphasis>
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. Clypeus strongly convex apically, strongly recurved at approximately 45 0 and strongly indented laterally (
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<figureCitation id="40B8C2C7FFCDFFE2FE88FEFAFED6A009" box="[259,343,262,287]" captionStart="FIGURE 17 – 21" captionStartId="7.[151,250,1726,1748]" captionText="FIGURE 17 – 21. Leucopholis bakeri Moser; 16, male genitalia, dorsal aspect; 17, male genitalia, ventral aspect; 18, male genitalia, lateral aspect; 20, habitus, dorsal aspect; 21, habitus, dorsal aspect (photograph of a syntype from ZMHB courtesy of Dr. Joachim Willers)." pageId="6" pageNumber="91">Fig. 19</figureCitation>
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||
). Antennal club length
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<quantity id="1F7B73A7FFCDFFE2FDEAFEFBFD3BA009" box="[609,698,263,287]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" pageId="6" pageNumber="91" unit="mm" value="1.1">1.1 mm</quantity>
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, shorter than antenomeres II–VII. Metaventral process length
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<quantity id="1F7B73A7FFCDFFE2FAF2FEFBFF47A054" metricMagnitude="-3" metricUnit="m" metricValue="1.4" pageId="6" pageNumber="91" unit="mm" value="1.4">1.4 mm</quantity>
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||
. Metaventrite sparsely setose and mostly covered with elongate, lanceolate scales. Posterior metatibial with 21–22 spicules. Scutellum twice as wide as long. Abdominal sternite V mesally and posteriorly elevated and not on the same plane with the other segments.
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||
</paragraph>
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||
<paragraph id="D83CDE42FFCDFFE3FC9EF8ECFF6EA68C" blockId="6.[789,1249,1792,1850]" lastBlockId="7.[151,1436,1849,2018]" lastPageId="7" lastPageNumber="92" pageId="6" pageNumber="91">
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||
<materialsCitation id="68EBD41FFFCDFFE3FC9EF8ECFA9FA644" ID-GBIF-Occurrence="1434655024" collectorName="Dapitan, C. F." lastPageId="7" lastPageNumber="92" location="Zamboanga Del Norte" pageId="6" pageNumber="91" specimenCount="19" specimenCount-male="1" stateProvince="Zamboanga Del Norte">
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19 15 16 17
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<specimenCount id="CE8515CBFFCCFFE3FBAEFC97FEC4A644" pageId="7" pageNumber="92" type="generic">
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||
18
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||
<emphasis id="EAF70250FFCCFFE3FF4CF8C5FEC4A644" bold="true" box="[199,325,1849,1874]" pageId="7" pageNumber="92">Specimens</emphasis>
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||
</specimenCount>
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||
examined:
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<emphasis id="EAF70250FFCCFFE3FE6CF8C5FC50A644" bold="true" box="[487,977,1849,1874]" pageId="7" pageNumber="92">
|
||
MINDANAO:
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<collectingRegion id="1A4710A0FFCCFFE3FD2EF8C5FC4DA644" box="[677,972,1849,1874]" country="Philippines" name="Zamboanga del Norte" pageId="7" pageNumber="92">
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||
Zamboanga
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<collectingRegion id="1A4710A0FFCCFFE3FCCCF8C5FC4DA644" box="[839,972,1849,1874]" inferredBySuffix="true" pageId="7" pageNumber="92">Del Norte</collectingRegion>
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</collectingRegion>
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:
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</emphasis>
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<specimenCount id="CE8515CBFFCCFFE3FC6EF8C6FB88A644" box="[997,1033,1850,1874]" pageId="7" pageNumber="92" type="male">1♂</specimenCount>
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,
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<collectorName id="7576BB94FFCCFFE3FBA8F8C5FB45A644" box="[1059,1220,1849,1874]" pageId="7" pageNumber="92">Dapitan, C.F.</collectorName>
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||
Baker
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||
</materialsCitation>
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||
;
|
||
<materialsCitation id="68EBD41FFFCCFFE3FABCF8C5FF6EA68C" ID-GBIF-Occurrence="1434655027" collectingDate="2004-09-15" collectorName="Calamba & Mamalad & E. Ending." location="Misamis Occidental" pageId="7" pageNumber="92" specimenCount="1" specimenCount-female="1" stateProvince="Misamis Occidental">
|
||
<emphasis id="EAF70250FFCCFFE3FABCF8C5FEA0A660" bold="true" pageId="7" pageNumber="92">
|
||
<collectingRegion id="1A4710A0FFCCFFE3FABCF8C5FE9CA660" country="Philippines" name="Misamis Occidental" pageId="7" pageNumber="92">Misamis Occidental</collectingRegion>
|
||
:
|
||
</emphasis>
|
||
<specimenCount id="CE8515CBFFCCFFE3FEA7F8A2FECEA660" box="[300,335,1886,1910]" pageId="7" pageNumber="92" type="female">1♀</specimenCount>
|
||
,
|
||
<collectorName id="7576BB94FFCCFFE3FEEBF8A1FE47A660" box="[352,454,1885,1910]" pageId="7" pageNumber="92">Calamba</collectorName>
|
||
,
|
||
<collectorName id="7576BB94FFCCFFE3FE5DF8A1FDC2A660" box="[470,579,1885,1910]" pageId="7" pageNumber="92">Mamalad</collectorName>
|
||
,
|
||
<taxonomicName id="1F83A5C1FFCCFFE3FDDAF8A2FC39A660" authority="Merrill" authorityName="Merrill" box="[593,952,1885,1910]" class="Liliopsida" family="Pandanaceae" genus="Freycinetia" kingdom="Plantae" order="Pandanales" pageId="7" pageNumber="92" phylum="Tracheophyta" rank="species" species="negrosensis">
|
||
<emphasis id="EAF70250FFCCFFE3FDDAF8A2FCDFA663" box="[593,862,1886,1909]" italics="true" pageId="7" pageNumber="92">Freycinetia negrosensis</emphasis>
|
||
Merrill
|
||
</taxonomicName>
|
||
, mixed dipterocarp forest,
|
||
<collectingDate id="BC79016AFFCCFFE3FB7DF8A1FAF4A660" box="[1270,1397,1885,1910]" pageId="7" pageNumber="92" value="2004-09-15">
|
||
<date id="AC3DF882FFCCFFE3FB7DF8A1FAF4A660" box="[1270,1397,1885,1910]" pageId="7" pageNumber="92" value="2004-09-15">15-ix-2004</date>
|
||
</collectingDate>
|
||
,
|
||
<collectorName id="7576BB94FFCCFFE3FA0FF8A2FF6EA68C" pageId="7" pageNumber="92">E. Ending.</collectorName>
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="8CFC8ECAFFCDFFE2FF1CF895FE20A6CD" httpUri="https://zenodo.org/record/293162/files/figure.png" pageId="6" pageNumber="91" targetBox="[259,719,1204,1838]" targetPageId="6">
|
||
<paragraph id="D83CDE42FFCDFFE2FF1CF895FE20A6CD" blockId="6.[151,1436,1897,2011]" pageId="6" pageNumber="91">
|
||
<emphasis id="EAF70250FFCDFFE2FF1CF895FECDA669" bold="true" box="[151,332,1897,1919]" pageId="6" pageNumber="91">FIGURE 12–16.</emphasis>
|
||
<taxonomicName id="1F83A5C1FFCDFFE2FED1F895FDF7A669" authority="Moser" authorityName="Moser" box="[346,630,1897,1919]" class="Insecta" family="Melolonthidae" genus="Leucopholis" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="91" phylum="Arthropoda" rank="species" species="reflexa">
|
||
<emphasis id="EAF70250FFCDFFE2FED1F895FDA9A669" box="[346,552,1897,1919]" italics="true" pageId="6" pageNumber="91">Leucopholis reflexa</emphasis>
|
||
Moser
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="1F83A5C1FFCDFFE2FD3EF896FCEDA669" authority="Moser." authorityName="Moser." box="[693,876,1897,1919]" class="Insecta" family="Melolonthidae" genus="Leucopholis" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="91" phylum="Arthropoda" rank="species" species="bakeri">
|
||
<emphasis id="EAF70250FFCDFFE2FD3EF896FD42A669" box="[693,707,1898,1919]" italics="true" pageId="6" pageNumber="91">L</emphasis>
|
||
.
|
||
<emphasis id="EAF70250FFCDFFE2FD5CF895FC99A669" box="[727,792,1897,1919]" italics="true" pageId="6" pageNumber="91">bakeri</emphasis>
|
||
Moser.
|
||
</taxonomicName>
|
||
12,
|
||
<taxonomicName id="1F83A5C1FFCDFFE2FC2EF895FBF2A669" box="[933,1139,1897,1919]" class="Insecta" family="Melolonthidae" genus="Leucopholis" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="91" phylum="Arthropoda" rank="species" species="reflexa">
|
||
<emphasis id="EAF70250FFCDFFE2FC2EF895FBF2A669" box="[933,1139,1897,1919]" italics="true" pageId="6" pageNumber="91">Leucopholis reflexa</emphasis>
|
||
</taxonomicName>
|
||
habitus, dorsal aspect; 13,
|
||
<taxonomicName id="1F83A5C1FFCDFFE2FF1CF874FEE0A688" box="[151,353,1928,1950]" class="Insecta" family="Melolonthidae" genus="Leucopholis" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="91" phylum="Arthropoda" rank="species" species="reflexa">
|
||
<emphasis id="EAF70250FFCDFFE2FF1CF874FEE0A688" box="[151,353,1928,1950]" italics="true" pageId="6" pageNumber="91">Leucopholis reflexa</emphasis>
|
||
</taxonomicName>
|
||
habitus, dorsal aspect (photograph of the syntype from ZMHB, courtesy of Dr. Joachim Willers); 14.
|
||
<taxonomicName id="1F83A5C1FFCDFFE2FF1CF85AFEDAA6AA" box="[151,347,1958,1980]" class="Insecta" family="Melolonthidae" genus="Leucopholis" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="91" phylum="Arthropoda" rank="species" species="bakeri">
|
||
<emphasis id="EAF70250FFCDFFE2FF1CF85AFEDAA6AA" box="[151,347,1958,1980]" italics="true" pageId="6" pageNumber="91">Leucopholis bakeri</emphasis>
|
||
</taxonomicName>
|
||
clypeus and labrum, male, apical aspect; 15.
|
||
<taxonomicName id="1F83A5C1FFCDFFE2FCB9F85AFC76A6AA" box="[818,1015,1958,1980]" class="Insecta" family="Melolonthidae" genus="Leucopholis" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="91" phylum="Arthropoda" rank="species" species="bakeri">
|
||
<emphasis id="EAF70250FFCDFFE2FCB9F85AFC76A6AA" box="[818,1015,1958,1980]" italics="true" pageId="6" pageNumber="91">Leucopholis bakeri</emphasis>
|
||
</taxonomicName>
|
||
protibia; 19,
|
||
<taxonomicName id="1F83A5C1FFCDFFE2FB08F85AFAC6A6AA" box="[1155,1351,1958,1980]" class="Insecta" family="Melolonthidae" genus="Leucopholis" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="91" phylum="Arthropoda" rank="species" species="bakeri">
|
||
<emphasis id="EAF70250FFCDFFE2FB08F85AFAC6A6AA" box="[1155,1351,1958,1980]" italics="true" pageId="6" pageNumber="91">Leucopholis bakeri</emphasis>
|
||
</taxonomicName>
|
||
clypeus and labrum, dorsal aspect.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="8CFC8ECAFFCCFFE3FF1CF942FEF0A604" pageId="7" pageNumber="92">
|
||
<paragraph id="D83CDE42FFCCFFE3FF1CF942FEF0A604" blockId="7.[151,1436,1726,1810]" pageId="7" pageNumber="92">
|
||
<emphasis id="EAF70250FFCCFFE3FF1CF942FEC8A7C2" bold="true" box="[151,329,1726,1748]" pageId="7" pageNumber="92">FIGURE 17–21.</emphasis>
|
||
<taxonomicName id="1F83A5C1FFCCFFE3FED9F942FDE7A7C2" authority="Moser" authorityName="Moser" box="[338,614,1726,1748]" class="Insecta" family="Melolonthidae" genus="Leucopholis" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="92" phylum="Arthropoda" rank="species" species="bakeri">
|
||
<emphasis id="EAF70250FFCCFFE3FED9F942FD98A7C2" box="[338,537,1726,1748]" italics="true" pageId="7" pageNumber="92">Leucopholis bakeri</emphasis>
|
||
Moser
|
||
</taxonomicName>
|
||
; 16, male genitalia, dorsal aspect; 17, male genitalia, ventral aspect; 18, male genitalia, lateral aspect; 20, habitus, dorsal aspect; 21, habitus, dorsal aspect (photograph of a syntype from ZMHB courtesy of Dr. Joachim Willers).
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="90998DC9FFCCFFE3FF4CF859FDF9A6A8" box="[199,632,1957,1982]" pageId="7" pageNumber="92" type="distribution">
|
||
<paragraph id="D83CDE42FFCCFFE3FF4CF859FDF9A6A8" blockId="7.[151,1436,1849,2018]" box="[199,632,1957,1982]" pageId="7" pageNumber="92">
|
||
<emphasis id="EAF70250FFCCFFE3FF4CF859FED8A6A8" bold="true" box="[199,345,1957,1982]" pageId="7" pageNumber="92">Distribution</emphasis>
|
||
.
|
||
<collectingCountry id="A0949ED2FFCCFFE3FEECF859FE64A6A8" box="[359,485,1957,1982]" name="Philippines" pageId="7" pageNumber="92">Philippines</collectingCountry>
|
||
(Mindanao).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="90998DC9FFCCFFECFF4CF835FC89A1EE" lastPageId="8" lastPageNumber="93" pageId="7" pageNumber="92" type="discussion">
|
||
<paragraph id="D83CDE42FFCCFFECFF4CF835FC89A1EE" blockId="7.[151,1436,1849,2018]" lastBlockId="8.[151,1436,151,248]" lastPageId="8" lastPageNumber="93" pageId="7" pageNumber="92">
|
||
<emphasis id="EAF70250FFCCFFE3FF4CF835FEB5A6F4" bold="true" box="[199,308,1993,2018]" pageId="7" pageNumber="92">Remarks</emphasis>
|
||
. Externally similar to
|
||
<taxonomicName id="1F83A5C1FFCCFFE3FDBAF836FD32A6F7" box="[561,691,1994,2017]" class="Insecta" family="Melolonthidae" genus="Leucopholis" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="92" phylum="Arthropoda" rank="species" species="guevarai">
|
||
<emphasis id="EAF70250FFCCFFE3FDBAF836FD32A6F7" box="[561,691,1994,2017]" italics="true" pageId="7" pageNumber="92">L. guevarai</emphasis>
|
||
</taxonomicName>
|
||
, differing in the uniform size of the scales on the elytra and in the phallobase of male genitalia being distinctly longer than parameres, which is the opposite of
|
||
<taxonomicName id="1F83A5C1FFC3FFECFB21FF65FAADA1A6" box="[1194,1324,153,176]" class="Insecta" family="Melolonthidae" genus="Leucopholis" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="93" phylum="Arthropoda" rank="species" species="guevarai">
|
||
<emphasis id="EAF70250FFC3FFECFB21FF65FAADA1A6" box="[1194,1324,153,176]" italics="true" pageId="8" pageNumber="93">L. guevarai</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="40B8C2C7FFC3FFECFAB0FF6BFA11A1A6" box="[1339,1424,151,176]" captionStart="FIGURE 17 – 21" captionStartId="7.[151,250,1726,1748]" captionText="FIGURE 17 – 21. Leucopholis bakeri Moser; 16, male genitalia, dorsal aspect; 17, male genitalia, ventral aspect; 18, male genitalia, lateral aspect; 20, habitus, dorsal aspect; 21, habitus, dorsal aspect (photograph of a syntype from ZMHB courtesy of Dr. Joachim Willers)." pageId="8" pageNumber="93">Fig. 20</figureCitation>
|
||
). The specimens examined from the UNSM and NMM were match the digital photographs of Moser’s
|
||
<typeStatus id="073860E0FFC3FFECFA83FF41FADEA1C3" box="[1288,1375,189,213]" pageId="8" pageNumber="93" type="syntype">syntype</typeStatus>
|
||
from the ZMHB (provided by Dr. Joachim Willers) (
|
||
<figureCitation id="40B8C2C7FFC3FFECFD2CFF23FD7BA1EE" box="[679,762,223,248]" captionStart="FIGURE 17 – 21" captionStartId="7.[151,250,1726,1748]" captionText="FIGURE 17 – 21. Leucopholis bakeri Moser; 16, male genitalia, dorsal aspect; 17, male genitalia, ventral aspect; 18, male genitalia, lateral aspect; 20, habitus, dorsal aspect; 21, habitus, dorsal aspect (photograph of a syntype from ZMHB courtesy of Dr. Joachim Willers)." pageId="8" pageNumber="93">Fig. 21</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |