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<document id="0463DE2F7436EDB4897A586873C537FB" ID="10.11646/zootaxa.4232.1.6SLASH10169" ID-CLB-Dataset="36544" ID-DOI="10.11646/zootaxa.4232.1.6" ID-GBIF-Dataset="86aa9515-2068-4285-838a-e73c8b8aaeb7" ID-ISSN="1175-5326" ID-Zenodo-Dep="293098" ID-ZooBank="9D76AB7D-0553-4A90-8FA8-C70153B705AE" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1487148507536" checkinUser="plazi" docAuthor="Calcetas, Orlando A. &amp; Adorada, Jessamyn R." docDate="2017" docId="502A6F54FFCEFFE1FF1CFF6BFEE3A58C" docLanguage="en" docName="zootaxa.4232.1.6SLASH10169.pdf" docOrigin="Zootaxa 4232 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Leucopholis reflexa Moser 1924" docType="treatment" docVersion="9" lastPageNumber="90" masterDocId="AC13172CFFCBFFE4FF8BFFFCFF81A116" masterDocTitle="Taxonomic review of the genus Leucopholis Dejean, 1833 (Coleoptera: Scarabaeidae: Melolonthinae: Leucopholini) in the Philippines" masterLastPageNumber="103" masterPageNumber="85" pageNumber="90" updateTime="1698436935605" updateUser="ExternalLinkService">
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<mods:title id="B02A165DBC0F2498C86633EF256C2733">Taxonomic review of the genus Leucopholis Dejean, 1833 (Coleoptera: Scarabaeidae: Melolonthinae: Leucopholini) in the Philippines</mods:title>
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<mods:namePart id="86C13D17C15D6476D53BDC0AA961CC2A">Calcetas, Orlando A.</mods:namePart>
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<mods:date id="DD0C43F9D639AE359A9082CDD4506D12">2017</mods:date>
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<treatment id="502A6F54FFCEFFE1FF1CFF6BFEE3A58C" ID-DOI="http://doi.org/10.5281/zenodo.6000491" ID-GBIF-Taxon="127695500" ID-Zenodo-Dep="6000491" LSID="urn:lsid:plazi:treatment:502A6F54FFCEFFE1FF1CFF6BFEE3A58C" httpUri="http://treatment.plazi.org/id/502A6F54FFCEFFE1FF1CFF6BFEE3A58C" lastPageNumber="90" pageId="5" pageNumber="90">
<subSubSection id="90998DC9FFCEFFE1FF1CFF6BFE8CA1C5" pageId="5" pageNumber="90" type="nomenclature">
<paragraph id="D83CDE42FFCEFFE1FF1CFF6BFDA4A1A4" blockId="5.[151,549,151,211]" box="[151,549,151,178]" pageId="5" pageNumber="90">
<heading id="8374692EFFCEFFE1FF1CFF6BFDA4A1A4" bold="true" box="[151,549,151,178]" fontSize="11" level="1" pageId="5" pageNumber="90" reason="1">
<taxonomicName id="1F83A5C1FFCEFFE1FF1CFF6BFDA4A1A4" authority="Moser, 1924" authorityName="Moser" authorityYear="1924" box="[151,549,151,178]" class="Insecta" family="Melolonthidae" genus="Leucopholis" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="90" phylum="Arthropoda" rank="species" species="reflexa">
<emphasis id="EAF70250FFCEFFE1FF1CFF6BFDA4A1A4" bold="true" box="[151,549,151,178]" pageId="5" pageNumber="90">
<emphasis id="EAF70250FFCEFFE1FF1CFF6BFE07A1A7" bold="true" box="[151,390,151,177]" italics="true" pageId="5" pageNumber="90">Leucopholis reflexa</emphasis>
<bibRefCitation id="BC12A3B3FFCEFFE1FE05FF64FDA4A1A4" author="Moser" box="[398,549,152,178]" pageId="5" pageNumber="90" refString="Moser, J. (1924) New species of melolonthid beetles from the Philippine Islands. Philippine Journal of Science, 25, 349 - 357." type="journal article" year="1924">Moser, 1924</bibRefCitation>
</emphasis>
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</paragraph>
<paragraph id="D83CDE42FFCEFFE1FF1CFF46FE8CA1C5" blockId="5.[151,549,151,211]" box="[151,269,186,211]" pageId="5" pageNumber="90">
<figureCitation id="40B8C2C7FFCEFFE1FF1CFF46FE8CA1C5" box="[151,269,186,211]" captionStart-0="FIGURE 6 11" captionStart-1="FIGURE 12 16" captionStartId-0="4.[151,250,1938,1960]" captionStartId-1="6.[151,250,1897,1919]" captionTargetBox-0="[241,536,1099,1756]" captionTargetBox-1="[259,719,1204,1838]" captionTargetId-0="figure@4.[241,536,1099,1756]" captionTargetPageId-0="4" captionTargetPageId-1="6" captionText-0="FIGURE 6 11. Leucopholis reflexa Moser. 6, clypeus, male, dorsal aspect; 7, clypeus, male, apical aspect; 8, male genitalia, dorsal aspect; 9, genitalia, male, ventral aspect; 10, genitalia, male, lateral aspect; 11, clypeus, female, apical aspect." captionText-1="FIGURE 12 16. Leucopholis reflexa Moser and L. bakeri Moser. 12, Leucopholis reflexa habitus, dorsal aspect; 13, Leucopholis reflexa habitus, dorsal aspect (photograph of the syntype from ZMHB, courtesy of Dr. Joachim Willers); 14. Leucopholis bakeri clypeus and labrum, male, apical aspect; 15. Leucopholis bakeri protibia; 19, Leucopholis bakeri clypeus and labrum, dorsal aspect." httpUri-0="https://zenodo.org/record/293099/files/figure.png" httpUri-1="https://zenodo.org/record/293162/files/figure.png" pageId="5" pageNumber="90">Figs. 613</figureCitation>
</paragraph>
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<subSubSection id="90998DC9FFCEFFE1FF1CFEFEFD9FA00E" box="[151,542,258,280]" pageId="5" pageNumber="90" type="reference_group">
<paragraph id="D83CDE42FFCEFFE1FF1CFEFEFD9FA00E" blockId="5.[151,542,258,280]" box="[151,542,258,280]" pageId="5" pageNumber="90">
<treatmentCitationGroup id="F893F96CFFCEFFE1FF1CFEFEFD9FA00E" box="[151,542,258,280]" pageId="5" pageNumber="90">
<taxonomicName id="1F83A5C1FFCEFFE1FF1CFEFEFD9BA00E" authority="Moser, 1924: 354" authorityName="Moser" authorityPageNumber="354" authorityYear="1924" box="[151,538,258,280]" class="Insecta" family="Melolonthidae" genus="Leucopholis" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="90" phylum="Arthropoda" rank="species" species="reflexa">
<emphasis id="EAF70250FFCEFFE1FF1CFEFEFEDFA00E" box="[151,350,258,280]" italics="true" pageId="5" pageNumber="90">Leucopholis reflexa</emphasis>
<treatmentCitation id="5922F853FFCEFFE1FEEFFEFFFD9BA00E" author="Moser" box="[356,538,258,280]" page="354" pageId="5" pageNumber="90" year="1924">
<bibRefCitation id="BC12A3B3FFCEFFE1FEEFFEFFFE66A00E" author="Moser" box="[356,487,258,280]" pageId="5" pageNumber="90" refString="Moser, J. (1924) New species of melolonthid beetles from the Philippine Islands. Philippine Journal of Science, 25, 349 - 357." type="journal article" year="1924">Moser, 1924</bibRefCitation>
: 354
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.
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<subSubSection id="90998DC9FFCEFFE1FF1CFEB9FC44A2F0" pageId="5" pageNumber="90" type="description">
<paragraph id="D83CDE42FFCEFFE1FF1CFEB9FDE3A320" blockId="5.[151,1437,325,1178]" pageId="5" pageNumber="90">
<emphasis id="EAF70250FFCEFFE1FF1CFEB9FEA8A048" bold="true" box="[151,297,325,350]" pageId="5" pageNumber="90">Description.</emphasis>
Male: length
<quantity id="1F7B73A7FFCEFFE1FE4DFEB9FDEDA04B" box="[454,620,325,350]" metricMagnitude="-2" metricUnit="m" metricValue="3.05" metricValueMax="3.25" metricValueMin="2.85" pageId="5" pageNumber="90" unit="mm" value="30.5" valueMax="32.5" valueMin="28.5">28.532.5 mm</quantity>
, width 14.0
<quantity id="1F7B73A7FFCEFFE1FC8BFEB9FCE7A04B" box="[768,870,325,350]" metricMagnitude="-2" metricUnit="m" metricValue="1.65" pageId="5" pageNumber="90" unit="mm" value="16.5">16.5 mm</quantity>
; female length 33.0 mm, width
<quantity id="1F7B73A7FFCEFFE1FB45FEB9FAB4A04B" box="[1230,1333,325,350]" metricMagnitude="-2" metricUnit="m" metricValue="1.65" pageId="5" pageNumber="90" unit="mm" value="16.5">16.5 mm</quantity>
. Clypeus anterior margin wide and strongly convex mesally, without median cleft, weakly indented and distinctly longer than labrum (
<figureCitation id="40B8C2C7FFCEFFE1FEB9FE71FE20A0B0" box="[306,417,397,422]" captionStart="FIGURE 6 11" captionStartId="4.[151,250,1938,1960]" captionTargetBox="[241,536,1099,1756]" captionTargetId="figure@4.[241,536,1099,1756]" captionTargetPageId="4" captionText="FIGURE 6 11. Leucopholis reflexa Moser. 6, clypeus, male, dorsal aspect; 7, clypeus, male, apical aspect; 8, male genitalia, dorsal aspect; 9, genitalia, male, ventral aspect; 10, genitalia, male, lateral aspect; 11, clypeus, female, apical aspect." httpUri="https://zenodo.org/record/293099/files/figure.png" pageId="5" pageNumber="90">Figs. 67</figureCitation>
). Antennal club length
<quantity id="1F7B73A7FFCEFFE1FD3BFE72FCBBA0B3" box="[688,826,398,422]" metricMagnitude="-3" metricUnit="m" metricValue="1.75" metricValueMax="1.8" metricValueMin="1.7" pageId="5" pageNumber="90" unit="mm" value="1.75" valueMax="1.8" valueMin="1.7">1.71.8 mm</quantity>
and nearly as long as antennomeres IIVII. Anterior margin of mentum with pronounced median cleft and with ovoid-shaped depressions medially, posterolateral margin bordered with lateral carina. Pronotum anterior angle rounded, posterior angle obtuse and lobed in both sexes. Metaventral process length
<quantity id="1F7B73A7FFCEFFE1FD9CFE05FD20A307" box="[535,673,505,529]" metricMagnitude="-3" metricUnit="m" metricValue="2.05" metricValueMax="2.3" metricValueMin="1.8" pageId="5" pageNumber="90" unit="mm" value="2.05" valueMax="2.3" valueMin="1.8">1.82.3 mm</quantity>
. Protibial spur extending in front of the anterior angle in both sexes. Posterior metatibial with 1826 spicules.
</paragraph>
<paragraph id="D83CDE42FFCEFFE1FF4CFDBDFDF8A3FC" blockId="5.[151,1437,325,1178]" pageId="5" pageNumber="90">
<emphasis id="EAF70250FFCEFFE1FF4CFDBDFE85A34C" bold="true" box="[199,260,577,602]" pageId="5" pageNumber="90">Male</emphasis>
. Genitalia length
<quantity id="1F7B73A7FFCEFFE1FE51FDBDFDB6A34F" box="[474,567,577,602]" metricMagnitude="-3" metricUnit="m" metricValue="9.6" pageId="5" pageNumber="90" unit="mm" value="9.6">9.6 mm</quantity>
. Phallobase dorsal anterior margin with narrow V-shaped margin mesally. Paramere ventral inner posterior margin inverted, elongate, and spindle-shaped; parameres parallel laterally, apically and basally pointed, mid-posterior margin triangulate and posterior angle rounded. Dorsal mid-posterior margin bell-shaped, sloping downwards to the lateral posterior angle, inner posterior margin cup-shaped and inner anterior margin ovoid-shaped (
<figureCitation id="40B8C2C7FFCEFFE1FE7AFD2DFDEAA3FC" box="[497,619,721,746]" captionStart="FIGURE 6 11" captionStartId="4.[151,250,1938,1960]" captionTargetBox="[241,536,1099,1756]" captionTargetId="figure@4.[241,536,1099,1756]" captionTargetPageId="4" captionText="FIGURE 6 11. Leucopholis reflexa Moser. 6, clypeus, male, dorsal aspect; 7, clypeus, male, apical aspect; 8, male genitalia, dorsal aspect; 9, genitalia, male, ventral aspect; 10, genitalia, male, lateral aspect; 11, clypeus, female, apical aspect." httpUri="https://zenodo.org/record/293099/files/figure.png" pageId="5" pageNumber="90">Figs. 810</figureCitation>
).
</paragraph>
<paragraph id="D83CDE42FFCEFFE1FF4CFD09FE6FA26C" blockId="5.[151,1437,325,1178]" pageId="5" pageNumber="90">
<emphasis id="EAF70250FFCEFFE1FF4CFD09FE9FA218" bold="true" box="[199,286,757,782]" pageId="5" pageNumber="90">Female</emphasis>
. Clypeus weakly convex, weakly recurved, strongly indented laterally and distinctly narrower than labrum (
<figureCitation id="40B8C2C7FFCEFFE1FF7EFCE5FEC6A224" box="[245,327,793,818]" captionStart="FIGURE 6 11" captionStartId="4.[151,250,1938,1960]" captionTargetBox="[241,536,1099,1756]" captionTargetId="figure@4.[241,536,1099,1756]" captionTargetPageId="4" captionText="FIGURE 6 11. Leucopholis reflexa Moser. 6, clypeus, male, dorsal aspect; 7, clypeus, male, apical aspect; 8, male genitalia, dorsal aspect; 9, genitalia, male, ventral aspect; 10, genitalia, male, lateral aspect; 11, clypeus, female, apical aspect." httpUri="https://zenodo.org/record/293099/files/figure.png" pageId="5" pageNumber="90">Fig. 11</figureCitation>
). Antennal club length
<quantity id="1F7B73A7FFCEFFE1FDC0FCE6FD31A227" box="[587,688,794,818]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" pageId="5" pageNumber="90" unit="mm" value="1.3">1.30 mm</quantity>
, shorter than antennomeres IIVII. Pronotum anterior angle obtuse and posterior angle at right angle. Hypomeron less exposed dorsally. Metaventral process length
<quantity id="1F7B73A7FFCEFFE1FB58FCC2FAADA243" box="[1235,1324,830,854]" metricMagnitude="-3" metricUnit="m" metricValue="2.3" pageId="5" pageNumber="90" unit="mm" value="2.3">2.3 mm</quantity>
. Posterior metatibia with 3033 spicules.
</paragraph>
<paragraph id="D83CDE42FFCEFFE1FF4CFC79FC44A2F0" blockId="5.[151,1437,325,1178]" pageId="5" pageNumber="90">
<materialsCitation id="68EBD41FFFCEFFE1FF4CFC79FCC5A288" ID-GBIF-Occurrence="1434655029" box="[199,836,901,926]" collectionCode="LUZON" country="Philippines" location="Specimens" pageId="5" pageNumber="90" specimenCount="2" specimenCount-male="2">
<emphasis id="EAF70250FFCEFFE1FF4CFC79FE41A288" bold="true" box="[199,448,901,926]" pageId="5" pageNumber="90">
<location id="DD5C8899FFCEFFE1FF4CFC79FEC4A288" LSID="urn:lsid:plazi:treatment:502A6F54FFCEFFE1FF1CFF6BFEE3A58C:DD5C8899FFCEFFE1FF4CFC79FEC4A288" box="[199,325,901,926]" country="Philippines" name="Specimens" pageId="5" pageNumber="90">Specimens</location>
examined
</emphasis>
.
<emphasis id="EAF70250FFCEFFE1FE44FC79FD0DA288" bold="true" box="[463,652,901,926]" pageId="5" pageNumber="90">
<collectingCountry id="A0949ED2FFCEFFE1FE44FC79FD07A288" box="[463,646,901,926]" name="Philippines" pageId="5" pageNumber="90">PHILIPPINES</collectingCountry>
:
</emphasis>
<specimenCount id="CE8515CBFFCEFFE1FD1FFC7AFD4EA288" box="[660,719,902,926]" pageId="5" pageNumber="90" type="male">2♂♂</specimenCount>
;
<collectionCode id="BE924687FFCEFFE1FD57FC79FCC5A288" box="[732,836,901,926]" pageId="5" pageNumber="90">
<emphasis id="EAF70250FFCEFFE1FD57FC79FCC5A288" bold="true" box="[732,836,901,926]" pageId="5" pageNumber="90">LUZON</emphasis>
</collectionCode>
</materialsCitation>
:
<materialsCitation id="68EBD41FFFCEFFE1FCDAFC79FEF0A2D4" ID-GBIF-Occurrence="1434655033" collectingDate="1975-05" country="Philippines" location="Mount Mabalagbag" municipality="Santa Cruz" pageId="5" pageNumber="90" specimenCount="1" specimenCount-male="1" stateProvince="Marinduque">
<collectingRegion id="1A4710A0FFCEFFE1FCDAFC79FC6DA288" box="[849,1004,901,926]" country="Philippines" name="Marinduque" pageId="5" pageNumber="90">Marinduque</collectingRegion>
:
<specimenCount id="CE8515CBFFCEFFE1FC72FC7AFB9FA288" box="[1017,1054,902,926]" pageId="5" pageNumber="90" type="male">1♂</specimenCount>
,
<collectingMunicipality id="38584438FFCEFFE1FBA7FC7AFB2AA288" box="[1068,1195,902,926]" pageId="5" pageNumber="90">Santa Cruz</collectingMunicipality>
,
<location id="DD5C8899FFCEFFE1FB33FC7AFA19A288" LSID="urn:lsid:plazi:treatment:502A6F54FFCEFFE1FF1CFF6BFEE3A58C:DD5C8899FFCEFFE1FB33FC7AFA19A288" box="[1208,1432,901,926]" country="Philippines" municipality="Santa Cruz" name="Mount Mabalagbag" pageId="5" pageNumber="90" stateProvince="Marinduque">Mount Mabalagbag</location>
,
<collectingDate id="BC79016AFFCEFFE1FF1CFC55FF6BA2D4" box="[151,234,937,962]" pageId="5" pageNumber="90" value="1975-05">
<date id="AC3DF882FFCEFFE1FF1CFC55FF6BA2D4" box="[151,234,937,962]" pageId="5" pageNumber="90" value="1975-05">v-1975</date>
</collectingDate>
.
<emphasis id="EAF70250FFCEFFE1FF7CFC55FEF0A2D4" bold="true" box="[247,369,937,962]" pageId="5" pageNumber="90">VISAYAS</emphasis>
</materialsCitation>
:
<materialsCitation id="68EBD41FFFCEFFE1FEF4FC55FA17A2D4" ID-GBIF-Occurrence="1434655028" box="[383,1430,937,962]" collectingDate="1952-06-02" collectorName="C. Yaptengco" country="Philippines" location="Tayasan" pageId="5" pageNumber="90" specimenCount="5" specimenCount-female="3" specimenCount-male="2" stateProvince="Negros Oriental">
<collectingRegion id="1A4710A0FFCEFFE1FEF4FC55FDC7A2D4" box="[383,582,937,962]" country="Philippines" name="Negros Oriental" pageId="5" pageNumber="90">Negros Oriental</collectingRegion>
:
<specimenCount id="CE8515CBFFCEFFE1FDDFFC56FD0FA2D4" box="[596,654,938,962]" pageId="5" pageNumber="90" type="female">2♀♀</specimenCount>
,
<location id="DD5C8899FFCEFFE1FD16FC56FD7AA2D4" LSID="urn:lsid:plazi:treatment:502A6F54FFCEFFE1FF1CFF6BFEE3A58C:DD5C8899FFCEFFE1FD16FC56FD7AA2D4" box="[669,763,938,962]" country="Philippines" name="Tayasan" pageId="5" pageNumber="90" stateProvince="Negros Oriental">Tayasan</location>
, 6.0 m,
<collectingDate id="BC79016AFFCEFFE1FCD7FC55FC4CA2D4" box="[860,973,937,962]" pageId="5" pageNumber="90" value="1952-06-02">
<date id="AC3DF882FFCEFFE1FCD7FC55FC4CA2D4" box="[860,973,937,962]" pageId="5" pageNumber="90" value="1952-06-02">2-vi-1952</date>
</collectingDate>
,
<collectorName id="7576BB94FFCEFFE1FC50FC56FBF9A2D4" box="[987,1144,938,962]" pageId="5" pageNumber="90">C. Yaptengco</collectorName>
;
<emphasis id="EAF70250FFCEFFE1FB0DFC55FAAEA2D4" bold="true" box="[1158,1327,937,962]" pageId="5" pageNumber="90">MINDANAO:</emphasis>
<specimenCount id="CE8515CBFFCEFFE1FAB2FC56FAFDA2D4" box="[1337,1404,938,962]" pageId="5" pageNumber="90" type="male">2♂♂</specimenCount>
<specimenCount id="CE8515CBFFCEFFE1FAF7FC56FA17A2D4" box="[1404,1430,938,962]" pageId="5" pageNumber="90" type="female">1♀</specimenCount>
</materialsCitation>
;
<materialsCitation id="68EBD41FFFCEFFE1FF1CFC31FE1CA2F0" ID-GBIF-Occurrence="1434655025" box="[151,413,973,998]" country="Philippines" location="Misamis Oriental" pageId="5" pageNumber="90" specimenCount="1" specimenCount-male="1" stateProvince="Misamis Oriental">
<emphasis id="EAF70250FFCEFFE1FF1CFC31FEF0A2F0" bold="true" box="[151,369,973,998]" pageId="5" pageNumber="90">
<collectingRegion id="1A4710A0FFCEFFE1FF1CFC31FEEDA2F0" box="[151,364,973,998]" country="Philippines" name="Misamis Oriental" pageId="5" pageNumber="90">Misamis Oriental</collectingRegion>
:
</emphasis>
<specimenCount id="CE8515CBFFCEFFE1FEF3FC32FE1CA2F0" box="[376,413,974,998]" pageId="5" pageNumber="90" type="male">1♂</specimenCount>
</materialsCitation>
,
<materialsCitation id="68EBD41FFFCEFFE1FE22FC32FC44A2F0" ID-GBIF-Occurrence="1434655022" box="[425,965,973,998]" collectingDate="1932-12-05" collectorName="H. L. Philipps." country="Philippines" location="Cagayan de Oro City" pageId="5" pageNumber="90" specimenCount="1" stateProvince="Cagayan de Oro City">
<collectingRegion id="1A4710A0FFCEFFE1FE22FC32FD16A2F0" box="[425,663,973,998]" country="Philippines" name="Cagayan de Oro" pageId="5" pageNumber="90">
Cagayan de
<collectingRegion id="1A4710A0FFCEFFE1FDBFFC32FD16A2F0" box="[564,663,973,998]" inferredBySuffix="true" pageId="5" pageNumber="90">Oro City</collectingRegion>
</collectingRegion>
,
<collectingDate id="BC79016AFFCEFFE1FD28FC31FC9DA2F0" box="[675,796,973,998]" pageId="5" pageNumber="90" value="1932-12-05">
<date id="AC3DF882FFCEFFE1FD28FC31FC9DA2F0" box="[675,796,973,998]" pageId="5" pageNumber="90" value="1932-12-05">5-xii-1932</date>
</collectingDate>
,
<collectorName id="7576BB94FFCEFFE1FCACFC32FC44A2F0" box="[807,965,973,998]" pageId="5" pageNumber="90">H.L. Philipps.</collectorName>
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="90998DC9FFCEFFE1FF4CFC0DFCABA51C" box="[199,810,1009,1034]" pageId="5" pageNumber="90" type="distribution">
<paragraph id="D83CDE42FFCEFFE1FF4CFC0DFCABA51C" blockId="5.[151,1437,325,1178]" box="[199,810,1009,1034]" pageId="5" pageNumber="90">
<emphasis id="EAF70250FFCEFFE1FF4CFC0DFED8A51C" bold="true" box="[199,345,1009,1034]" pageId="5" pageNumber="90">Distribution</emphasis>
.
<collectingCountry id="A0949ED2FFCEFFE1FEECFC0DFE64A51C" box="[359,485,1009,1034]" name="Philippines" pageId="5" pageNumber="90">Philippines</collectingCountry>
(Luzon, Visayas, Mindanao)
</paragraph>
</subSubSection>
<subSubSection id="90998DC9FFCEFFE1FF4CFBE9FEE3A58C" pageId="5" pageNumber="90" type="discussion">
<paragraph id="D83CDE42FFCEFFE1FF4CFBE9FEE3A58C" blockId="5.[151,1437,325,1178]" pageId="5" pageNumber="90">
<emphasis id="EAF70250FFCEFFE1FF4CFBE9FEB5A538" bold="true" box="[199,308,1045,1070]" pageId="5" pageNumber="90">Remarks</emphasis>
. Externally similar to
<taxonomicName id="1F83A5C1FFCEFFE1FDC8FBEAFD41A53B" box="[579,704,1046,1069]" class="Insecta" family="Melolonthidae" genus="Leucopholis" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="90" phylum="Arthropoda" rank="species" species="irrorata">
<emphasis id="EAF70250FFCEFFE1FDC8FBEAFD41A53B" box="[579,704,1046,1069]" italics="true" pageId="5" pageNumber="90">L. irrorata</emphasis>
</taxonomicName>
(
<figureCitation id="40B8C2C7FFCEFFE1FD5FFBE9FCADA538" box="[724,812,1045,1070]" captionStart="FIGURE 12 16" captionStartId="6.[151,250,1897,1919]" captionTargetBox="[259,719,1204,1838]" captionTargetPageId="6" captionText="FIGURE 12 16. Leucopholis reflexa Moser and L. bakeri Moser. 12, Leucopholis reflexa habitus, dorsal aspect; 13, Leucopholis reflexa habitus, dorsal aspect (photograph of the syntype from ZMHB, courtesy of Dr. Joachim Willers); 14. Leucopholis bakeri clypeus and labrum, male, apical aspect; 15. Leucopholis bakeri protibia; 19, Leucopholis bakeri clypeus and labrum, dorsal aspect." httpUri="https://zenodo.org/record/293162/files/figure.png" pageId="5" pageNumber="90">Fig. 12</figureCitation>
). Phallobase of male genitalia distinctly shorter than parameres. The specimens examined from the UNSM, NMM, and IRRI-AC match the digital photographs of one of Mosers
<typeStatus id="073860E0FFCEFFE1FE96FBA2FEFEA560" box="[285,383,1118,1142]" pageId="5" pageNumber="90" type="syntype">syntypes</typeStatus>
from the ZMHB (provided by Dr. Joachim Willers) (
<figureCitation id="40B8C2C7FFCEFFE1FC67FBA1FBC3A560" box="[1004,1090,1117,1142]" captionStart="FIGURE 12 16" captionStartId="6.[151,250,1897,1919]" captionTargetBox="[259,719,1204,1838]" captionTargetPageId="6" captionText="FIGURE 12 16. Leucopholis reflexa Moser and L. bakeri Moser. 12, Leucopholis reflexa habitus, dorsal aspect; 13, Leucopholis reflexa habitus, dorsal aspect (photograph of the syntype from ZMHB, courtesy of Dr. Joachim Willers); 14. Leucopholis bakeri clypeus and labrum, male, apical aspect; 15. Leucopholis bakeri protibia; 19, Leucopholis bakeri clypeus and labrum, dorsal aspect." httpUri="https://zenodo.org/record/293162/files/figure.png" pageId="5" pageNumber="90">Fig. 13</figureCitation>
). The clypeal character of the species is distinct.
</paragraph>
</subSubSection>
</treatment>
</document>