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<document id="D62684B3005F933451F15E109014A902" ID="10.11646/zootaxa.4232.1.6SLASH10169" ID-CLB-Dataset="36544" ID-DOI="10.11646/zootaxa.4232.1.6" ID-GBIF-Dataset="86aa9515-2068-4285-838a-e73c8b8aaeb7" ID-ISSN="1175-5326" ID-Zenodo-Dep="293098" ID-ZooBank="9D76AB7D-0553-4A90-8FA8-C70153B705AE" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1487148507536" checkinUser="plazi" docAuthor="Calcetas, Orlando A. &amp; Adorada, Jessamyn R." docDate="2017" docId="502A6F54FFC0FFE9FF1CFB9FFCB4A515" docLanguage="en" docName="zootaxa.4232.1.6SLASH10169.pdf" docOrigin="Zootaxa 4232 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Leucopholis guevarai Calcetas &amp; Adorada, new species" docType="treatment" docUuid="B77CE70A-E156-4ECA-B4ED-4C441502048E" docUuidSource="ZooBank" docVersion="10" lastPageNumber="97" masterDocId="AC13172CFFCBFFE4FF8BFFFCFF81A116" masterDocTitle="Taxonomic review of the genus Leucopholis Dejean, 1833 (Coleoptera: Scarabaeidae: Melolonthinae: Leucopholini) in the Philippines" masterLastPageNumber="103" masterPageNumber="85" pageNumber="96" updateTime="1698436935605" updateUser="ExternalLinkService">
<mods:mods id="67D0C9BD1311D6B90D47E5978A2B4A41" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="A9A2AD4EF636707AFD39295B227520DB">
<mods:title id="6A87941F997470DD292090F7407E59C6">Taxonomic review of the genus Leucopholis Dejean, 1833 (Coleoptera: Scarabaeidae: Melolonthinae: Leucopholini) in the Philippines</mods:title>
</mods:titleInfo>
<mods:name id="F7598567503848DE84BAA143513C84CE" type="personal">
<mods:role id="917BF6D828728C3C264FB2D17A4D86DF">
<mods:roleTerm id="203684840462FC7EE5C9452E6C5E01D3">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="8B4E071E38293B7B084CBFFE9F8FA000">Calcetas, Orlando A.</mods:namePart>
</mods:name>
<mods:name id="C97C826B0230FD29BC34AE657F047EFB" type="personal">
<mods:role id="91C614EC539EF1E7116FE466CB052A52">
<mods:roleTerm id="CAE16911045CEB83CAA489224339342F">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="480F276DBC75F0A7DB9AB41F7D037F24">Adorada, Jessamyn R.</mods:namePart>
</mods:name>
<mods:typeOfResource id="1F3090CC92C40B82A9CB2709A3FEB990">text</mods:typeOfResource>
<mods:relatedItem id="E7F7B6229F42A1C12C26B088C514606C" type="host">
<mods:titleInfo id="806DBBAB856D0D3ED753F0232EC6DC5C">
<mods:title id="874033BB4987FDF3DC1486576385F15C">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="9883E50E99B8E7129D8C3742154FC670">
<mods:date id="7DE33FA212505A5D6D80299BA3D5400F">2017</mods:date>
<mods:detail id="EF6B5A66EAA9BE128CE936D9A0E8455C" type="volume">
<mods:number id="3664933FE6743B07275C34AFF105CD41">4232</mods:number>
</mods:detail>
<mods:detail id="FFE5D2375D6119D880343B5238143E0C" type="issue">
<mods:number id="F126FFE4A97D0731D0692F2664D5BF27">1</mods:number>
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<mods:start id="43BE1280DC34BC7C627BA1A5D2DBAC50">85</mods:start>
<mods:end id="081B15DA2EDFD8BF3E4382521FC23B25">103</mods:end>
</mods:extent>
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<mods:classification id="5D5531F439C9FDED4EFB1BB810802BA1">journal article</mods:classification>
<mods:identifier id="7C9AF59E887AAC672328D48E5791C328" type="CLB-Dataset">36544</mods:identifier>
<mods:identifier id="59014D243CAA1BB62DB190957AEA9664" type="DOI">10.11646/zootaxa.4232.1.6</mods:identifier>
<mods:identifier id="EB2DF11E0F67ACBF240FADDE058C3B80" type="GBIF-Dataset">86aa9515-2068-4285-838a-e73c8b8aaeb7</mods:identifier>
<mods:identifier id="272E6DBFBBF9F0DFEED1AFD4FBBBA967" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="3CD8E3ED7D05D15A5DC243EC0091EAEF" type="Zenodo-Dep">293098</mods:identifier>
<mods:identifier id="D702C4B212A3B3BDE94B1F220DC3F235" type="ZooBank">9D76AB7D-0553-4A90-8FA8-C70153B705AE</mods:identifier>
</mods:mods>
<treatment id="502A6F54FFC0FFE9FF1CFB9FFCB4A515" ID-DOI="http://doi.org/10.5281/zenodo.6000498" ID-GBIF-Taxon="127695504" ID-Zenodo-Dep="6000498" LSID="urn:lsid:zoobank.org:act:B77CE70A-E156-4ECA-B4ED-4C441502048E" httpUri="http://treatment.plazi.org/id/502A6F54FFC0FFE9FF1CFB9FFCB4A515" lastPageId="13" lastPageNumber="97" pageId="11" pageNumber="96">
<subSubSection id="90998DC9FFC0FFEFFF1CFB9FFE9AA589" pageId="11" pageNumber="96" type="nomenclature">
<paragraph id="D83CDE42FFC0FFEFFF1CFB9FFCC0A568" blockId="11.[151,833,1123,1183]" box="[151,833,1123,1150]" pageId="11" pageNumber="96">
<heading id="8374692EFFC0FFEFFF1CFB9FFCC0A568" bold="true" box="[151,833,1123,1150]" fontSize="11" level="1" pageId="11" pageNumber="96" reason="1">
<emphasis id="EAF70250FFC0FFEFFF1CFB9FFCC0A568" bold="true" box="[151,833,1123,1150]" pageId="11" pageNumber="96">
<taxonomicName id="1F83A5C1FFC0FFEFFF1CFB9FFD24A568" authority="Calcetas &amp; Adorada" authorityName="Calcetas &amp; Adorada" box="[151,677,1123,1150]" class="Insecta" family="Melolonthidae" genus="Leucopholis" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="96" phylum="Arthropoda" rank="species" species="guevarai" status="sp. nov.">
<emphasis id="EAF70250FFC0FFEFFF1CFB9FFE1DA56B" bold="true" box="[151,412,1123,1149]" italics="true" pageId="11" pageNumber="96">Leucopholis guevarai</emphasis>
Calcetas &amp; Adorada
</taxonomicName>
,
<taxonomicNameLabel id="F1C4BF2BFFC0FFEFFD3AFB98FCC0A568" box="[689,833,1124,1150]" pageId="11" pageNumber="96" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="D83CDE42FFC0FFEFFF1CFB7AFE9AA589" blockId="11.[151,833,1123,1183]" box="[151,283,1158,1183]" pageId="11" pageNumber="96">
<figureCitation id="40B8C2C7FFC0FFEFFF1CFB7AFE9AA589" box="[151,283,1158,1183]" captionStart-0="FIGURE 30 – 34" captionStart-1="FIGURE 35" captionStartId-0="12.[151,250,1833,1855]" captionStartId-1="13.[151,250,1104,1126]" captionText-0="FIGURE 30 – 34. Leucopholis guevarai new species. 30, antenna, male; 31, protibia, dorsal aspect; 32, male genitalia, dorsal aspect; 33, male genitalia, ventral aspect; 34, male genitalia, lateral aspect." captionText-1="FIGURE 35. Leucopholis guevarai new species habitus, dorsal aspect." pageId="11" pageNumber="96">Figs. 30–35</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="90998DC9FFC0FFEFFF1CFB32FD17A759" pageId="11" pageNumber="96" type="materials_examined">
<paragraph id="D83CDE42FFC0FFEFFF1CFB32FD17A759" blockId="11.[151,1437,1230,2011]" pageId="11" pageNumber="96">
<emphasis id="EAF70250FFC0FFEFFF1CFB32FE18A5F1" bold="true" box="[151,409,1230,1255]" pageId="11" pageNumber="96">
<typeStatus id="073860E0FFC0FFEFFF1CFB32FE85A5F1" box="[151,260,1230,1255]" pageId="11" pageNumber="96" type="holotype">Holotype</typeStatus>
description.
</emphasis>
Length
<quantity id="1F7B73A7FFC0FFEFFE73FB33FDCAA5F0" box="[504,587,1231,1255]" metricMagnitude="-2" metricUnit="m" metricValue="2.1" pageId="11" pageNumber="96" unit="mm" value="21.0">21 mm</quantity>
, width
<quantity id="1F7B73A7FFC0FFEFFD14FB33FD73A5F0" box="[671,754,1231,1255]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="11" pageNumber="96" unit="mm" value="10.0">10 mm</quantity>
. Body elongate ovoid, uniformally dull dark reddish brown. Dorsum covered with evenly distributed, nearly uniformally sized, recumbent, circular to ovoid, and nonoverlapping scales. Clypeus weakly convex apically and with weakly but distinct median cleft. Clypeus nearly as long as labrum mesally, nearly straight or vertical laterally, and weakly indented mesally. Mentum with anterior margin strongly convex; posterior half with deep median groove or depression, median carina smooth. Antennal club length
<quantity id="1F7B73A7FFC0FFEFFE97FA7FFE27A48C" box="[284,422,1411,1435]" metricMagnitude="-3" metricUnit="m" metricValue="2.2" metricValueMax="2.3" metricValueMin="2.1" pageId="11" pageNumber="96" unit="mm" value="2.2" valueMax="2.3" valueMin="2.1">2.1–2.3 mm</quantity>
, distinctly longer than antenomeres II–VII (
<figureCitation id="40B8C2C7FFC0FFEFFC1DFA7EFC6BA48D" box="[918,1002,1410,1435]" captionStart="FIGURE 30 – 34" captionStartId="12.[151,250,1833,1855]" captionText="FIGURE 30 – 34. Leucopholis guevarai new species. 30, antenna, male; 31, protibia, dorsal aspect; 32, male genitalia, dorsal aspect; 33, male genitalia, ventral aspect; 34, male genitalia, lateral aspect." pageId="11" pageNumber="96">Fig. 30</figureCitation>
), laterally concave or boat-shaped and nearly as wide as clypeus. Pronotum anterior and posterior angle obtuse. Lateral margins weakly to strongly carinate. Anterior and posterior margin smooth. Protibial spur distinctly short and not extending in front of the anterior angle or sinuation (
<figureCitation id="40B8C2C7FFC0FFEFFE45FA12FDA0A711" box="[462,545,1518,1543]" captionStart="FIGURE 30 – 34" captionStartId="12.[151,250,1833,1855]" captionText="FIGURE 30 – 34. Leucopholis guevarai new species. 30, antenna, male; 31, protibia, dorsal aspect; 32, male genitalia, dorsal aspect; 33, male genitalia, ventral aspect; 34, male genitalia, lateral aspect." pageId="11" pageNumber="96">Fig. 31</figureCitation>
). Metaventral process length
<quantity id="1F7B73A7FFC0FFEFFCE6FA13FC47A710" box="[877,966,1519,1543]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" pageId="11" pageNumber="96" unit="mm" value="1.1">1.1 mm</quantity>
. Posterior metatibial with 17–19 spicules. Abdominal sternites covered with ovoid to ovoid-lanceolate, non-overlapping scales. Pygidium covered apically and basally with blunt, spindle-shaped scales.
</paragraph>
</subSubSection>
<subSubSection id="90998DC9FFC0FFEFFF4CF9A6FB2AA65C" pageId="11" pageNumber="96" type="description">
<paragraph id="D83CDE42FFC0FFEFFF4CF9A6FD28A7C9" blockId="11.[151,1437,1230,2011]" pageId="11" pageNumber="96">
<emphasis id="EAF70250FFC0FFEFFF4CF9A6FE85A765" bold="true" box="[199,260,1626,1651]" pageId="11" pageNumber="96">Male</emphasis>
. Genitalia length
<quantity id="1F7B73A7FFC0FFEFFE5DF9A7FD9CA764" box="[470,541,1627,1650]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" pageId="11" pageNumber="96" unit="mm" value="7.0">7 mm</quantity>
. Phallobase dorsal anterior margin nearly straight (
<figureCitation id="40B8C2C7FFC0FFEFFBE6F9A6FB45A765" box="[1133,1220,1626,1651]" captionStart="FIGURE 30 – 34" captionStartId="12.[151,250,1833,1855]" captionText="FIGURE 30 – 34. Leucopholis guevarai new species. 30, antenna, male; 31, protibia, dorsal aspect; 32, male genitalia, dorsal aspect; 33, male genitalia, ventral aspect; 34, male genitalia, lateral aspect." pageId="11" pageNumber="96">Fig. 32</figureCitation>
). Paramere ventral inner margin spindle shaped, narrow towards apex, with thick apical processes. Mid-posterior margin triangulate, apically tapered, posterior junction narrower (
<figureCitation id="40B8C2C7FFC0FFEFFD33F95EFC8EA7AD" box="[696,783,1698,1723]" captionStart="FIGURE 30 – 34" captionStartId="12.[151,250,1833,1855]" captionText="FIGURE 30 – 34. Leucopholis guevarai new species. 30, antenna, male; 31, protibia, dorsal aspect; 32, male genitalia, dorsal aspect; 33, male genitalia, ventral aspect; 34, male genitalia, lateral aspect." pageId="11" pageNumber="96">Fig. 33</figureCitation>
). Dorsal mid-posterior margin bell-shaped and sloping downward towards the lateral angles. (
<figureCitation id="40B8C2C7FFC0FFEFFDC3F93AFD1DA7C9" box="[584,668,1734,1759]" captionStart="FIGURE 30 – 34" captionStartId="12.[151,250,1833,1855]" captionText="FIGURE 30 – 34. Leucopholis guevarai new species. 30, antenna, male; 31, protibia, dorsal aspect; 32, male genitalia, dorsal aspect; 33, male genitalia, ventral aspect; 34, male genitalia, lateral aspect." pageId="11" pageNumber="96">Fig. 34</figureCitation>
).
</paragraph>
<paragraph id="D83CDE42FFC0FFEFFF4CF916FD7CA614" blockId="11.[151,1437,1230,2011]" box="[199,765,1770,1795]" pageId="11" pageNumber="96">
<emphasis id="EAF70250FFC0FFEFFF4CF916FEB4A615" bold="true" box="[199,309,1770,1795]" pageId="11" pageNumber="96">Variation</emphasis>
. Length
<quantity id="1F7B73A7FFC0FFEFFE17F916FDADA614" box="[412,556,1770,1795]" metricMagnitude="-2" metricUnit="m" metricValue="2.175" metricValueMax="2.25" metricValueMin="2.1" pageId="11" pageNumber="96" unit="mm" value="21.75" valueMax="22.5" valueMin="21.0">21–22.5 mm</quantity>
, width
<quantity id="1F7B73A7FFC0FFEFFDF5F917FD78A614" box="[638,761,1771,1795]" metricMagnitude="-2" metricUnit="m" metricValue="1.05" metricValueMax="1.1" metricValueMin="1.0" pageId="11" pageNumber="96" unit="mm" value="10.5" valueMax="11.0" valueMin="10.0">10–11 mm</quantity>
.
</paragraph>
<paragraph id="D83CDE42FFC0FFEFFF4CF8F2FB2AA65C" blockId="11.[151,1437,1230,2011]" pageId="11" pageNumber="96">
<materialsCitation id="68EBD41FFFC0FFEFFF4CF8F2FCF4A631" ID-GBIF-Occurrence="1434654940" box="[199,885,1806,1831]" location="Sulu" pageId="11" pageNumber="96" specimenCount="1" stateProvince="Sulu" typeStatus="holotype">
<emphasis id="EAF70250FFC0FFEFFF4CF8F2FE3EA631" bold="true" box="[199,447,1806,1831]" pageId="11" pageNumber="96">Specimens examined</emphasis>
.
<emphasis id="EAF70250FFC0FFEFFE46F8F2FCF4A631" bold="true" box="[461,885,1806,1831]" pageId="11" pageNumber="96">
<typeStatus id="073860E0FFC0FFEFFE46F8F2FDBBA631" box="[461,570,1806,1831]" pageId="11" pageNumber="96" type="holotype">Holotype</typeStatus>
male: MINDANAO:
<collectingRegion id="1A4710A0FFC0FFEFFCB0F8F2FCF4A631" box="[827,885,1806,1831]" country="Philippines" name="Sulu" pageId="11" pageNumber="96">Sulu</collectingRegion>
</emphasis>
</materialsCitation>
:
<materialsCitation id="68EBD41FFFC0FFEFFC09F8F2FA9AA631" ID-GBIF-Occurrence="1434654922" box="[898,1307,1806,1831]" collectingDate="1963-03-28" collectorName="O. Grino" location="Basilan" pageId="11" pageNumber="96" specimenCount="1" stateProvince="Basilan" typeStatus="holotype">
<collectingRegion id="1A4710A0FFC0FFEFFC09F8F2FC57A631" box="[898,982,1806,1831]" country="Philippines" name="Basilan" pageId="11" pageNumber="96">Basilan</collectingRegion>
Isand,
<collectingDate id="BC79016AFFC0FFEFFBA3F8F2FB29A631" box="[1064,1192,1806,1831]" pageId="11" pageNumber="96" value="1963-03-28">
<date id="AC3DF882FFC0FFEFFBA3F8F2FB29A631" box="[1064,1192,1806,1831]" pageId="11" pageNumber="96" value="1963-03-28">28-iii-1963</date>
</collectingDate>
,
<collectorName id="7576BB94FFC0FFEFFB3FF8F3FA9AA631" box="[1204,1307,1806,1831]" pageId="11" pageNumber="96">O. Grino</collectorName>
</materialsCitation>
.
<materialsCitation id="68EBD41FFFC0FFEFFAADF8F2FB26A65C" ID-GBIF-Occurrence="1434654941" collectingDate="1963-03-28" collectionCode="UPLB" collectorName="O. Grino. The" country="Iceland" location="Basilan" pageId="11" pageNumber="96" specimenCount="1" specimenCount-male="1" stateProvince="Basilan" typeStatus="paratype">
<emphasis id="EAF70250FFC0FFEFFAADF8F2FA1DA631" bold="true" box="[1318,1436,1806,1831]" pageId="11" pageNumber="96">
<typeStatus id="073860E0FFC0FFEFFAADF8F2FA16A631" box="[1318,1431,1806,1831]" pageId="11" pageNumber="96" type="paratype">Paratype</typeStatus>
:
</emphasis>
<specimenCount id="CE8515CBFFC0FFEFFF1CF8CFFF3AA65D" box="[151,187,1843,1867]" pageId="11" pageNumber="96" type="male">1♂</specimenCount>
,
<collectingRegion id="1A4710A0FFC0FFEFFF43F8CEFE9DA65D" box="[200,284,1842,1867]" country="Philippines" name="Basilan" pageId="11" pageNumber="96">Basilan</collectingRegion>
<collectingCountry id="A0949ED2FFC0FFEFFEA8F8CEFEEBA65D" box="[291,362,1842,1867]" name="Iceland" pageId="11" pageNumber="96">Island</collectingCountry>
,
<collectingDate id="BC79016AFFC0FFEFFEFEF8CEFE77A65D" box="[373,502,1842,1867]" pageId="11" pageNumber="96" value="1963-03-28">
<date id="AC3DF882FFC0FFEFFEFEF8CEFE77A65D" box="[373,502,1842,1867]" pageId="11" pageNumber="96" value="1963-03-28">28-iii-1963</date>
</collectingDate>
,
<collectorName id="7576BB94FFC0FFEFFD8AF8CFFD1EA65D" box="[513,671,1842,1867]" pageId="11" pageNumber="96">O. Grino. The</collectorName>
<typeStatus id="073860E0FFC0FFEFFD2EF8CEFC86A65D" box="[677,775,1842,1867]" pageId="11" pageNumber="96" type="holotype">holotype</typeStatus>
and
<typeStatus id="073860E0FFC0FFEFFCB5F8CFFC1FA65D" box="[830,926,1843,1867]" pageId="11" pageNumber="96" type="paratype">paratype</typeStatus>
are deposited at
<collectionCode id="BE924687FFC0FFEFFBD7F8CFFB26A65C" box="[1116,1191,1843,1866]" country="Philippines" httpUri="http://grbio.org/cool/134h-pm2f" name="Museum of Natural History, University of the Philippines" pageId="11" pageNumber="96">UPLB</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="90998DC9FFC0FFEFFF4CF8AAFDF1A679" box="[199,624,1878,1903]" pageId="11" pageNumber="96" type="distribution">
<paragraph id="D83CDE42FFC0FFEFFF4CF8AAFDF1A679" blockId="11.[151,1437,1230,2011]" box="[199,624,1878,1903]" pageId="11" pageNumber="96">
<emphasis id="EAF70250FFC0FFEFFF4CF8AAFED8A679" bold="true" box="[199,345,1878,1903]" pageId="11" pageNumber="96">Distribution</emphasis>
.
<collectingCountry id="A0949ED2FFC0FFEFFEECF8AAFE64A679" box="[359,485,1878,1903]" name="Philippines" pageId="11" pageNumber="96">Philippines</collectingCountry>
(Mindanao)
</paragraph>
</subSubSection>
<subSubSection id="90998DC9FFC0FFEFFF4CF886FC93A6CD" pageId="11" pageNumber="96" type="etymology">
<paragraph id="D83CDE42FFC0FFEFFF4CF886FC93A6CD" blockId="11.[151,1437,1230,2011]" pageId="11" pageNumber="96">
<emphasis id="EAF70250FFC0FFEFFF4CF886FEC7A685" bold="true" box="[199,326,1914,1939]" pageId="11" pageNumber="96">Etymology</emphasis>
. This new species is named and honorably dedicated to Don Pedro Guevara. He was also a monumental figure to the approval of the Tydings-McDuffie Law and a delegate in the Constitutional Convention of 1934, which framed the 1935 Philippine Constitution.
</paragraph>
</subSubSection>
<subSubSection id="90998DC9FFC7FFE9FF4CFF6BFCB4A515" lastPageId="13" lastPageNumber="98" pageId="12" pageNumber="97" type="discussion">
<paragraph id="D83CDE42FFC7FFE8FF4CFF6BFE81A1EE" blockId="12.[151,1436,151,248]" pageId="12" pageNumber="97">
<emphasis id="EAF70250FFC7FFE8FF4CFF6BFEB5A1A6" bold="true" box="[199,308,151,176]" pageId="12" pageNumber="97">Remarks</emphasis>
. Externally similar to
<taxonomicName id="1F83A5C1FFC7FFE8FDA6FF65FD13A1A6" box="[557,658,151,176]" class="Insecta" family="Melolonthidae" genus="Leucopholis" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="97" phylum="Arthropoda" rank="species" species="bakeri">
<emphasis id="EAF70250FFC7FFE8FDA6FF65FD13A1A6" box="[557,658,151,176]" italics="true" pageId="12" pageNumber="97">L. bakeri</emphasis>
</taxonomicName>
but can be distinguished by the body being more elongate than ovate, the male antennal club being distinctly longer and boat-shaped, and the phallobase being shorter than the parameres (
<figureCitation id="40B8C2C7FFC7FFE8FF14FF23FF72A1EE" box="[159,243,223,248]" captionStart="FIGURE 35" captionStartId="13.[151,250,1104,1126]" captionText="FIGURE 35. Leucopholis guevarai new species habitus, dorsal aspect." pageId="12" pageNumber="97">Fig. 35</figureCitation>
).
</paragraph>
<paragraph id="D83CDE42FFC7FFE9FEC4F934FCB4A515" blockId="12.[335,1144,1736,1778]" lastBlockId="13.[761,821,985,1027]" lastPageId="13" lastPageNumber="98" pageId="12" pageNumber="97">32 33 34 35</paragraph>
</subSubSection>
</treatment>
</document>
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