190 lines
21 KiB
XML
190 lines
21 KiB
XML
<document id="16CC0F50F466816A3DCBEFE243995D4A" ID-DOI="http://dx.doi.org/10.3897/BDJ.4.e8150" ID-PMC="PMC5019016" ID-Pensoft-Pub="1314-2828--8150" ID-PubMed="27651730" ID-ZooBank="262DB958242246B692E61675C3C07DB1" ModsDocAuthor="" ModsDocDate="2016" ModsDocID="1314-2828--e8150" ModsDocOrigin="Biodiversity Data Journal " ModsDocTitle="A taxonomic monograph of the assassin bug genus Zelus Fabricius (Hemiptera: Reduviidae): 71 species based on 10,000 specimens" checkinTime="1553126057039" checkinUser="pensoft" docAuthor="Zhang, Guanyang, Hart, Elwood R & Weirauch, Christiane" docDate="2016" docId="3D4B65549AC924C39B43887A85A2144F" docLanguage="en" docName="BiodivDatJour 4: e8150" docOrigin="Biodiversity Data Journal 4" docSource="http://dx.doi.org/10.3897/BDJ.4.e8150" docTitle="Zelus means Fabricius 1803" docType="treatment" docVersion="5" lastPageNumber="8150" masterDocId="570F0E795632FF88A15573625E50C567" masterDocTitle="A taxonomic monograph of the assassin bug genus Zelus Fabricius (Hemiptera: Reduviidae): 71 species based on 10,000 specimens" masterLastPageNumber="8150" masterPageNumber="8150" pageNumber="8150" updateTime="1701316348929" updateUser="plazi">
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<mods:mods id="E3CF6D8BE4C1380F2542A4D52D4D15E9" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="AA29874D5AC342B10B168CCE2B934AB5">
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<mods:title id="A6EE3844B78DAED3BA84DA5B7E43C0D0">A taxonomic monograph of the assassin bug genus Zelus Fabricius (Hemiptera: Reduviidae): 71 species based on 10,000 specimens</mods:title>
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</mods:titleInfo>
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<mods:name id="8719C74982FDF6619C44951FBB8F1A26" type="personal">
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<mods:role id="4412D424E45D432D1F1FE970EF2038B0">
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<mods:roleTerm id="FED6BCFD027A4A084DDB7227C7C8674F">Author</mods:roleTerm>
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<mods:namePart id="EE61842D618BBF537BA6324C493F786E">Zhang, Guanyang</mods:namePart>
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<mods:roleTerm id="E0B59D1AF293C26194C7147B7AC52C00">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="6FD4CE26BCB21FB81923003FDC073001">Hart, Elwood R</mods:namePart>
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</mods:name>
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<mods:name id="3180A2037260986F6CD255013B836417" type="personal">
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<mods:roleTerm id="2B2D8D33C0BCF2759CCF4821B3FE0346">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="2CFABBD77A3533B43E149ABC0A9778E6">Weirauch, Christiane</mods:namePart>
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<mods:typeOfResource id="1D9DFE3659D6F561D8E124AFAD31C320">text</mods:typeOfResource>
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<mods:titleInfo id="23EBD0B6ECED4225B2F38BA8CB8CDAB5">
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<mods:title id="91140BF295AD6D9084A8B5D40AB52D65">Biodiversity Data Journal</mods:title>
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<mods:part id="D31DC574F2C10B7DD8F264865E9D7434">
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<mods:date id="8BEA3B6F307789742876ABEEFB75E2E3">2016</mods:date>
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<mods:detail id="38599E77092E741496CDAC00A4131C2F" type="volume">
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<mods:number id="AAA012F6F319610858F4A28DB010972B">4</mods:number>
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</mods:detail>
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<mods:start id="3248807CCAA8BD64EC92BA60A57FDC9F">8150</mods:start>
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<mods:url id="CA8EC7D8BEA04875F029A6B502952C46">http://dx.doi.org/10.3897/BDJ.4.e8150</mods:url>
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<mods:classification id="0A99F955FC9599E402196F1E80AEB17F">journal article</mods:classification>
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<mods:identifier id="8837513F61CDFBA5A97A22D251864CBE" type="DOI">http://dx.doi.org/10.3897/BDJ.4.e8150</mods:identifier>
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<mods:identifier id="89722C93E0D38EFCFC772D3F4685A821" type="Pensoft-Pub">1314-2828--8150</mods:identifier>
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</mods:mods>
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<treatment id="3D4B65549AC924C39B43887A85A2144F" LSID="urn:lsid:plazi:treatment:3D4B65549AC924C39B43887A85A2144F" httpUri="http://treatment.plazi.org/id/3D4B65549AC924C39B43887A85A2144F" lastPageNumber="8150" pageId="0" pageNumber="8150">
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<subSubSection id="A7993DBD0790AEA76D1E0FFD1316144F" pageId="0" pageNumber="8150" type="nomenclature">
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<paragraph id="9196484F87E52129CE1AA1BD5B66FF41" pageId="0" pageNumber="8150">
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<taxonomicName id="1A82B16337AE0384782232743F311FC3" ID-CoL="5D25G" authority="Fabricius, 1803" authorityName="Fabricius" authorityYear="1803" class="Insecta" family="Reduviidae" genus="Zelus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zelus means" order="Hemiptera" pageId="0" pageNumber="8150" phylum="Arthropoda" rank="species" species="means">Zelus means Fabricius, 1803</taxonomicName>
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</paragraph>
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</subSubSection>
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<subSubSection id="42B86E8ED2567893C8C3C976FB824D6D" pageId="0" pageNumber="8150" type="reference_group">
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<paragraph id="62041CBAFDA7B33798A028C83547727E" pageId="0" pageNumber="8150">
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<taxonomicName id="5E50BD99A87BA586D84E5862CFB6211D" class="Insecta" family="Reduviidae" genus="Zelus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zelus means" order="Hemiptera" pageId="0" pageNumber="8150" phylum="Arthropoda" rank="species" species="means">Zelus means</taxonomicName>
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Fabricius, 1803, p. 282. orig. descr.;
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<normalizedToken id="D20FECC6E15D397B53081390D5284967" originalValue="Stål">Stal</normalizedToken>
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, 1868, p. 107, descr.;
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<normalizedToken id="0BB39EC919F30B8329DADCD490437861" originalValue="Stål">Stal</normalizedToken>
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, 1872, p. 89, cat.; Walker, 1873, p. 134, cat; Lethierry and Severin, 1896, p.l62., cat.; Champion, 1898, p. 254, note; Wygodzinsky, 1949a, p. 49, checklist; Maldonado, 1990, p. 329, cat.
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</paragraph>
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<paragraph id="2E3DB52CFA6478C387A6306A1F9B6135" pageId="0" pageNumber="8150">
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<taxonomicName id="80E0A9401AC7C998D7ADE8D6E049F324" class="Insecta" family="Reduviidae" genus="Zelus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zelus means" order="Hemiptera" pageId="0" pageNumber="8150" phylum="Arthropoda" rank="species" species="means">Zelus means</taxonomicName>
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Eccagoras trimaculicollis
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<normalizedToken id="F4F2FA14DFDF821D13BE60065DFB6B74" originalValue="Stål">Stal</normalizedToken>
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1855
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</paragraph>
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<paragraph id="CE1AB82426706BBEB3CCA3742A4F57BE" pageId="0" pageNumber="8150">
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<taxonomicName id="F15FC5A5956AF8DBE02AE7D7DC37286B" class="Insecta" family="Reduviidae" genus="Zelus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zelus trimaculicollis" order="Hemiptera" pageId="0" pageNumber="8150" phylum="Arthropoda" rank="species" species="trimaculicollis">Zelus trimaculicollis</taxonomicName>
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<normalizedToken id="BEC68CF59632717A87028C3BF9614484" originalValue="Stål">Stal</normalizedToken>
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, 1866, p. 298, descr.;
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<normalizedToken id="294A8A2FDA861419A0697A30EF2FE0E8" originalValue="Stål">Stal</normalizedToken>
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, 1872, p. 89, cat.; Lethierry and Severin, 1896, p. 153, cat.; Champion, 1898, p. 254, note; Wygodzinsky, 1949a, p. 50, checklist; Maldonado, 1990, p. 331, cat. syn. nov. (current study).
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</paragraph>
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<paragraph id="811F1313B740458E19D69FD5598CE0F1" pageId="0" pageNumber="8150">
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<taxonomicName id="21612C096E9E12C41454DA6C1DC8D7C6" class="Insecta" family="Reduviidae" genus="Zelus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zelus trimaculatus" order="Hemiptera" pageId="0" pageNumber="8150" phylum="Arthropoda" rank="species" species="trimaculatus">Zelus trimaculatus</taxonomicName>
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Champion, 1898, p. 254, orig. descr.; Wygodzinsky, 1949a, p. 50, checklist; Maldonado, 1990, p. 331, cat. syn. nov. (current study).
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</paragraph>
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</subSubSection>
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<subSubSection id="3F4919A3211786490CED7DDD84B03D01" pageId="0" pageNumber="8150" type="materials_examined">
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<paragraph id="4BC46138893AC7C647ECDF520A49D1AD" pageId="0" pageNumber="8150">Materials</paragraph>
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<paragraph id="D68118D34477DB78BAD41E1A88B02CB8" pageId="0" pageNumber="8150">
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<materialsCitation id="D6FE11E54DA7E1C994AEAD7FBFE92CA5" collectionCode="ZMUC" collectorName="Schmidt" location="South America (American Meridonali)" pageId="0" pageNumber="8150" specimenCount="1" typeStatus="Lectotype">
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Type status:
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<typeStatus id="F44E52FAC644D5F399D8057FAD8FAAB7" pageId="0" pageNumber="8150">Lectotype</typeStatus>
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. Occurrence: occurrenceRemarks: Lectotype of Zelusmeans Fabricius, 1803 (New Designation by Zhang, Har & Weirauch, 2016). Bears the following labels: Type / Z.means in Am. Mer. Schmidt; recordedBy:
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<collectorName id="FFEFF2C753F5CA92B325B436B94FB980" pageId="0" pageNumber="8150">Schmidt</collectorName>
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; Taxon: scientificName: Zelusmeans; family: Reduviidae; genus: Zelus; scientificNameAuthorship: Fabricius, 1803; Location: locality:
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<location id="187611324F3DAF6CCB5AD7B740FCDF23" LSID="urn:lsid:plazi:treatment:3D4B65549AC924C39B43887A85A2144F:187611324F3DAF6CCB5AD7B740FCDF23" name="South America (American Meridonali)" pageId="0" pageNumber="8150">South America (American Meridonali)</location>
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; Record Level: institutionCode:
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<collectionCode id="1112533D3036556837F2055F913AA043" pageId="0" pageNumber="8150">ZMUC</collectionCode>
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</materialsCitation>
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<materialsCitation id="FADF86F8E666A5BE2A41E39297B3F0B2" collectingDate="No date provided" collectionCode="NHRS" country="BRAZIL" location="Unknown" pageId="0" pageNumber="8150" specimenCode="UCR _ ENT 00041011" specimenCount="1" typeStatus="Other material">
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Type status:
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<typeStatus id="ED12545A08A1415BE5E4FAFADBF25EC9" pageId="0" pageNumber="8150">Other material</typeStatus>
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. Occurrence: catalogNumber:
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<specimenCode id="68F020130A06FD73B98B842B8BBA0E77" pageId="0" pageNumber="8150">UCR_ENT 00041011</specimenCode>
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; occurrenceRemarks: Lectotype of Zelustrimaculicollis Stal 1866 (New Designation by Zhang, Hart & Weirauch, 2016), now synonym of Zelusmeans Fabricius 1803. Verbatim label info: Brasil / Typus / trimaculicollis Stal / Lectotype Zelustrimaculicollis Stal / designated by E.R.Hart / Zelusmeans Fabricius det. E.R. Hart 1972 / NHRS-GULI 000000350; sex:
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<specimenType id="F6A58551E10DF250F71336C8A809D90C" pageId="0" pageNumber="8150">Adult Female</specimenType>
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; otherCatalogNumbers: NHRS-GULI 000000350; Taxon: scientificName: Zelusmeans; family: Reduviidae; genus: Zelus; scientificNameAuthorship: Fabricius, 1803; Location: country:
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<collectingCountry id="D7E40B7BCDA5114F67F25F139F9B26AA" pageId="0" pageNumber="8150">BRAZIL</collectingCountry>
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; stateProvince: unknown; locality:
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<location id="ED65FCDD2B6C7710FAFCBEF4E40F5674" LSID="urn:lsid:plazi:treatment:3D4B65549AC924C39B43887A85A2144F:ED65FCDD2B6C7710FAFCBEF4E40F5674" country="BRAZIL" name="Unknown" pageId="0" pageNumber="8150">Unknown</location>
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; Identification: identifiedBy:
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<determinerName id="2C2C83497CFCEEDEA87BFF7A469BCA71" pageId="0" pageNumber="8150">G. Zhang</determinerName>
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; dateIdentified: 2012; Event: eventDate:
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<collectingDate id="3FAD6FAC1D435C49C997B7C1294FEC1B" pageId="0" pageNumber="8150">No date provided</collectingDate>
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; Record Level: institutionCode:
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<collectionCode id="35C0519A43F9FFF760184E5CD1F8BEAD" pageId="0" pageNumber="8150">NHRS</collectionCode>
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</materialsCitation>
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<materialsCitation id="516CC3C94A24CCA10B743A2D6B89D41A" collectingDate="No date provided" collectionCode="NHRS" country="BRAZIL" location="Unknown" pageId="0" pageNumber="8150" specimenCount="1" typeStatus="Other material">
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Type status:
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<typeStatus id="618112815BB3414EA83D67AA8323B9C2" pageId="0" pageNumber="8150">Other material</typeStatus>
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. Occurrence: occurrenceRemarks: Paralectotype of Zelustrimaculicollis Stal 1866 (New Designation by Zhang, Hart & Weirauch 2016), junior synonym of Zelusmeans Fabricius 1803. Verbatim label info: Brasil / Stal / Paratypus; sex:
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<specimenType id="3809A105032B7DBAD6D5C7A30DAB31D2" pageId="0" pageNumber="8150">Adult Female</specimenType>
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; Taxon: scientificName: Zelusmeans; family: Reduviidae; genus: Zelus; scientificNameAuthorship: Fabricius, 1803; Location: country:
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<collectingCountry id="1F8EB2E93C6785F7E67F4C7142F5CE61" pageId="0" pageNumber="8150">BRAZIL</collectingCountry>
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; stateProvince: unknown; locality:
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<location id="6B337E62FB73EA3EE1C4DA9FDB89C337" LSID="urn:lsid:plazi:treatment:3D4B65549AC924C39B43887A85A2144F:6B337E62FB73EA3EE1C4DA9FDB89C337" country="BRAZIL" name="Unknown" pageId="0" pageNumber="8150">Unknown</location>
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; Event: eventDate:
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<collectingDate id="7EA0AC84547047F5B928F2077DEF0353" pageId="0" pageNumber="8150">No date provided</collectingDate>
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; Record Level: institutionCode:
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<collectionCode id="EAC2AC7CB19558EAC934145C689DFF19" pageId="0" pageNumber="8150">NHRS</collectionCode>
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</materialsCitation>
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<materialsCitation id="7C637EB5C44285C038D62E34A5709842" collectingDate="No date provided" collectionCode="NHRS" country="BRAZIL" location="Unknown" pageId="0" pageNumber="8150" specimenCount="1" typeStatus="Other material">
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Type status:
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<typeStatus id="598FDCF61E203095ED14877B53BC2070" pageId="0" pageNumber="8150">Other material</typeStatus>
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. Occurrence: occurrenceRemarks: Paralectotype of Zelustrimaculicollis Stal 1866 (New Designation by Zhang, Hart & Weirauch, 2016), junior synonym of Zelusmeans Fabricius 1803. Verbatim label info: Brasil / Paratypus; sex:
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<specimenType id="8AD2E4769F632CD30F13005ED5F07784" pageId="0" pageNumber="8150">Adult Female</specimenType>
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; Taxon: scientificName: Zelusmeans; family: Reduviidae; genus: Zelus; scientificNameAuthorship: Fabricius, 1803; Location: country:
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<collectingCountry id="8D4AA6DF2711193E616F9FADEFD09EA2" pageId="0" pageNumber="8150">BRAZIL</collectingCountry>
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; stateProvince: unknown; locality:
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<location id="CA8D474832679132E1EDB7AE223C54E5" LSID="urn:lsid:plazi:treatment:3D4B65549AC924C39B43887A85A2144F:CA8D474832679132E1EDB7AE223C54E5" country="BRAZIL" name="Unknown" pageId="0" pageNumber="8150">Unknown</location>
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; Event: eventDate:
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<collectingDate id="135D8820584F6A14747F2661D19492D1" pageId="0" pageNumber="8150">No date provided</collectingDate>
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; Record Level: institutionCode:
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<collectionCode id="7A39EF16CDE971F89495763F03ACF96F" pageId="0" pageNumber="8150">NHRS</collectionCode>
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</materialsCitation>
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<materialsCitation id="2ABCE3E56128647D49CFE469C24F8A24" collectingDate="V. de Chiriqui" collectionCode="BMNH" country="PANAMA" location="PANAMA" pageId="0" pageNumber="8150" specimenCount="1" typeStatus="Other material">
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Type status:
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<typeStatus id="1AD36F2696840ADCDCF591486F613C36" pageId="0" pageNumber="8150">Other material</typeStatus>
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. Occurrence: occurrenceRemarks: Holotype of Zelustrimaculatus Champion 1898, junior synonym of Zelusmeans Fabricius 1803. Bears the following labels: Type/B.C.A. Rhyn. II. Zelustrimaculatus Ch / Sp. figured / V. de Chiriqui, 25-4000 ft., Champion; sex:
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<specimenType id="B2C67E06D6C3BC270A4F60A8978B450E" pageId="0" pageNumber="8150">Adult Female</specimenType>
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; Taxon: scientificName: Zelusmeans; family: Reduviidae; genus: Zelus; scientificNameAuthorship: Fabricius, 1803; Location: country:
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<collectingCountry id="C911CFFED9CFB2C1AE4A4E41107E2822" pageId="0" pageNumber="8150">PANAMA</collectingCountry>
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; stateProvince: Chiriqui; Event: eventDate:
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<collectingDate id="8F1BF372BBA3EC0A29713BB86FCEE39D" pageId="0" pageNumber="8150">V. de Chiriqui</collectingDate>
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; Record Level: institutionCode:
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<collectionCode id="4BB6F6506853873CF8C07D11275ECEB7" pageId="0" pageNumber="8150">BMNH</collectionCode>
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</materialsCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="007AAF994C1BD87A3F41A898D45CFEC0" pageId="0" pageNumber="8150" type="description">
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<paragraph id="56F0C6E4748BA51E3E0A6F398DD647E3" pageId="0" pageNumber="8150">Description</paragraph>
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<paragraph id="7296DED48FC93E2AF78282533FE52E94" pageId="0" pageNumber="8150">Figs 136, 137</paragraph>
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<paragraph id="84B0878176DA6DFF7B00CBFA60665EC2" pageId="0" pageNumber="8150">Male: Unknown.</paragraph>
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<paragraph id="3EF4DDD330126E3A95C37EAD8A4A7DAB" pageId="0" pageNumber="8150">Female: (Fig. 136) Large, total length 16.94-19.31 mm (mean 18.29 mm, Suppl. material 2); slender. COLORATION: Reddish, orangish, mixed with black; pattern variable. VESTITURE: Densely setose. Head with short recumbent setae and some scattered longer setae. Anterior pronotal lobe with erect spine-like setae, short to moderate in length; posterior pronotal lobe with short, erect, spine-like setae. Abdomen with short, recumbent and short to moderately long, erect setae. STRUCTURE: Head: Stout. Postocular lobe long; in dorsal view anteriorly gradually narrowing, posterior portion constant, slightly narrower. Eye smallish; lateral margin only slightly wider than postocular lobe; dorsal and ventral margins removed from surfaces of head. Labium: I: II: III = 1: 1.5: 0.4. Basiflagellomere diameter larger than that of pedicel. Thorax: Anterolateral angle rounded, without projection; medial longitudinal sulcus evident throughout, deepening posteriorly. Posterior pronotal lobe with rugulose surface; disc distinctly elevated above humeral angle; humeral angle rounded, without projection. Scutellum moderately long; apex blunt, not projected. Legs: Moderately robust. Hemelytron: Surpassing apex of abdomen by about twice length of abdominal segment seven; quadrate cell large and broad; Cu and M of cubital cell subparallel.</paragraph>
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</subSubSection>
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<subSubSection id="8204FCD896855ACAE633C182CC7A6B6F" pageId="0" pageNumber="8150" type="diagnosis">
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<paragraph id="E4C3A604139BD73F8E80FDC9B9504957" pageId="0" pageNumber="8150">Diagnosis</paragraph>
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<paragraph id="F173C56F17AEDB43038C131783A1D9C9" pageId="0" pageNumber="8150">
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The humeral angle rounded; the pronotum with spine-like setae; and the colors usually consisting of yellow, orange, red and black. The anterior pronotal lobe is rather small, margins not laterally expanded, nearly continuous with lateral margins of posterior lobe, dorsally nearly flat, not bulging. The small anterior lobe together with the regularly sized posterior lobe gives the pronotum a triangular appearance. Most similar to
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<taxonomicName id="35B6B6FE3EC2DB50B534F6D6B65D0EF6" lsidName="Z. fuliginatus" pageId="0" pageNumber="8150" rank="species" species="fuliginatus">Z. fuliginatus</taxonomicName>
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, but the postocular lobe is shorter.
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</paragraph>
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</subSubSection>
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<subSubSection id="BD2CD80C9630C0D3AF9920E0646CD193" pageId="0" pageNumber="8150" type="distribution">
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<paragraph id="7C8E33EA37E31DA6DD247DB6B7A26872" pageId="0" pageNumber="8150">Distribution</paragraph>
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<paragraph id="05032879CFE869951F36E28698E9C8B6" pageId="0" pageNumber="8150">Southern Central America and South America (Fig. 137). Countries with records: Bolivia, Brazil, Costa Rica, Colombia, Ecuador and Peru.</paragraph>
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</subSubSection>
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<subSubSection id="AC1B48904D849DF12553D519EA52C4D8" pageId="0" pageNumber="8150" type="taxon discussion">
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<paragraph id="4DC139C5E2CF476B7CFD326F979BC580" pageId="0" pageNumber="8150">Taxon discussion</paragraph>
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<paragraph id="06D13537DE0704669894DDE3F66BC1D7" pageId="0" pageNumber="8150">
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On the basis of the spine-like setae on the head and pronotum and the rounded humeral angle,
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<taxonomicName id="0789964FC3FE7C79948DC8C6F6282380" class="Insecta" family="Reduviidae" genus="Zelus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zelus means" order="Hemiptera" pageId="0" pageNumber="8150" phylum="Arthropoda" rank="species" species="means">Zelus means</taxonomicName>
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would be placed in either the
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<taxonomicName id="EE9F608C35E1A138B1388525CB902E02" class="Insecta" family="Reduviidae" genus="Zelus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zelus longipes" order="Hemiptera" pageId="0" pageNumber="8150" phylum="Arthropoda" rank="species" species="longipes">Zelus longipes</taxonomicName>
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species group or the
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<taxonomicName id="9E310028BCE0F4CDE9D27DFFDB39BAB9" class="Insecta" family="Reduviidae" genus="Zelus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zelus vagans" order="Hemiptera" pageId="0" pageNumber="8150" phylum="Arthropoda" rank="species" species="vagans">Zelus vagans</taxonomicName>
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species group. Due to its close resemblance to
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<taxonomicName id="CDF20287117D189B485A017EC5E3609E" lsidName="Z. fuliginatus" pageId="0" pageNumber="8150" rank="species" species="fuliginatus">Z. fuliginatus</taxonomicName>
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, this species is most likely part of the
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<taxonomicName id="A6684E603C5D6ECCCE7CB862D9CC639C" class="Insecta" family="Reduviidae" genus="Zelus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zelus vagans" order="Hemiptera" pageId="0" pageNumber="8150" phylum="Arthropoda" rank="species" species="vagans">Zelus vagans</taxonomicName>
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species group.
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</paragraph>
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<paragraph id="CE870947A6949C60ABF6AD85599EC972" pageId="0" pageNumber="8150">This species appears to have a highly variable color pattern in all areas of its distribution. The clavus and corium range from entirely brownish-black to almost entirely light yellowish-brown. The dorsum of the pronotal lobe may be entirely brownish-black, but is usually variably patterned brownish-black and reddish-brown.</paragraph>
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</subSubSection>
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</treatment>
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</document> |