treatments-xml/data/1C/42/58/1C425869E88C5CBFAF75137953446DBA.xml

<document ID-DOI="http://dx.doi.org/10.3897/zookeys.955.51983" ID-GBIF-Dataset="45a98afc-af5a-4734-93d1-205e1ac3694b" ID-PMC="PMC7423785" ID-Pensoft-Pub="1313-2970-955-1" ID-Pensoft-UUID="F7018CF19C6158309D57C83A2F5E9478" ID-PubMed="32855592" ID-ZooBank="C8EF4A4B6F054621B9B5AE47FEE7C217" ModsDocID="1313-2970-955-1" checkinTime="1596691600214" checkinUser="pensoft" docAuthor="Grego, Jozef, Mumladze, Levan, Falniowski, Andrzej, Osikowski, Artur, Rysiewska, Aleksandra, Palatov, Dimitry M. &amp; Hofman, Sebastian" docDate="2020" docId="1C425869E88C5CBFAF75137953446DBA" docLanguage="en" docName="ZooKeys 955: 1-77" docOrigin="ZooKeys 955" docSource="http://dx.doi.org/10.3897/zookeys.955.51983" docTitle="Hausdorfenia shareula Grego &amp; Mumladze 2020, sp. nov." docType="treatment" docUuid="29BDD264-C578-49D8-945E-F4835070D996" docUuidSource="ZooBank" docVersion="5" id="F7018CF19C6158309D57C83A2F5E9478" lastPageNumber="1" masterDocId="F7018CF19C6158309D57C83A2F5E9478" masterDocTitle="Revealing the stygobiotic and crenobiotic molluscan biodiversity hotspot in Caucasus: Part I. The phylogeny of stygobiotic Sadlerianinae Szarowska, 2006 (Mollusca, Gastropoda, Hydrobiidae) from Georgia with descriptions of five new genera and twenty-one new species" masterLastPageNumber="77" masterPageNumber="1" pageNumber="1" updateTime="1668169198162" updateUser="ExternalLinkService">
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<mods:titleInfo>
<mods:title>Revealing the stygobiotic and crenobiotic molluscan biodiversity hotspot in Caucasus: Part I. The phylogeny of stygobiotic Sadlerianinae Szarowska, 2006 (Mollusca, Gastropoda, Hydrobiidae) from Georgia with descriptions of five new genera and twenty-one new species</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Grego, Jozef</mods:namePart>
<mods:affiliation>Horna Micina 219, 97401, Banska Bystrica, Slovakia</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Mumladze, Levan</mods:namePart>
<mods:affiliation>Institute of Zoology, Ilia State University, Kakutsa Cholokashvili Ave 3 / 5, Tbilisi 0162, Georgia</mods:affiliation>
<mods:nameIdentifier type="orcid">https://orcid.org/0000-0002-2172-6973</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Falniowski, Andrzej</mods:namePart>
<mods:affiliation>Department of Malacology, Institute of Zoology and Biomedical Research, Jagiellonian University, ul. Gronostajowa 9, 30 - 387, Krakow, Poland</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Osikowski, Artur</mods:namePart>
<mods:affiliation>Department of Animal Reproduction, Anatomy and Genomics, University of Agriculture in Krakow, al. Mickiewicza 24 / 28, 30 - 059, Krakow, Poland</mods:affiliation>
<mods:nameIdentifier type="orcid">https://orcid.org/0000-0001-6646-2687</mods:nameIdentifier>
<mods:nameIdentifier type="email">a.osikowski@urk.edu.pl</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Rysiewska, Aleksandra</mods:namePart>
<mods:affiliation>Department of Malacology, Institute of Zoology and Biomedical Research, Jagiellonian University, ul. Gronostajowa 9, 30 - 387, Krakow, Poland</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Palatov, Dimitry M.</mods:namePart>
<mods:affiliation>Department of Hydrobiology, Biological Faculty, Moscow State University, 1 - 12 Leninskie Gory, 119991, Moscow, Russian Federation</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Hofman, Sebastian</mods:namePart>
<mods:affiliation>Department of Comparative Anatomy, Institute of Zoology and Biomedical Research, Jagiellonian University, ul. Gronostajowa 9, 30 - 387, Krakow, Poland</mods:affiliation>
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<mods:title>ZooKeys</mods:title>
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<mods:date>2020</mods:date>
<mods:detail type="volume">
<mods:number>955</mods:number>
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<mods:start>1</mods:start>
<mods:end>77</mods:end>
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<mods:url>http://dx.doi.org/10.3897/zookeys.955.51983</mods:url>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.955.51983</mods:identifier>
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<treatment ID-GBIF-Taxon="166096059" LSID="urn:lsid:zoobank.org:act:29BDD264-C578-49D8-945E-F4835070D996" httpUri="http://treatment.plazi.org/id/1C425869E88C5CBFAF75137953446DBA" lastPageNumber="1" pageId="0" pageNumber="1">
<subSubSection pageId="0" pageNumber="1" type="nomenclature">
<paragraph pageId="0" pageNumber="1">
<taxonomicName LSID="1C425869-E88C-5CBF-AF75-137953446DBA" authority="Grego &amp; Mumladze" authorityName="Grego &amp; Mumladze" authorityYear="2020" class="Gastropoda" family="Hydrobiidae" genus="Hausdorfenia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hausdorfenia shareula" order="Littorinimorpha" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="shareula" status="sp. nov.">Hausdorfenia shareula Grego &amp; Mumladze</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="1">sp. nov.</taxonomicNameLabel>
<figureCitation captionStart="Plate 20" captionStartId="F36" captionText="Plate 20. 1, 2 Pontohoratia vinarskii sp. nov., Samegrelo, Chkhorotsku, Letsurtsume, Letsurtsume Cave: 1 holotype 2 paratype 3 P. pichkhaiai sp. nov., Samegrelo, Mukhuri, Shisha Spring, holotype 4 Hausdorfenia pseudohauffenia sp. nov., Racha, Zemo Krikhi, Krikhula Spring, holotype 5 H. shareula sp. nov., Racha, Nikortsmintha, Tsivtskala 2 Spring near power station in the Shareula valley, holotype 6 P. mapeli sp. nov., Samegrelo, Mukhuri, Kanti, Mapeli Spring, holotype 7 H. pseudohauffenia sp. nov., Racha, Zemo Krikhi, Krikhula Spring; aberrant specimen 8, 9 P. vinarskii sp. nov., Samegrelo, Chkhorotsku, Letsurtsume, Letsurtsume Cave: 8 morphotype 9 live specimen with marked carbonate incrustations. The numbers correspond to individuals, and the letters represent the different views of the same individual. Photograph J. Grego." figureDoi="10.3897/zookeys.955.51983.plate20" httpUri="https://binary.pensoft.net/fig/439187" pageId="0" pageNumber="1">Plate 20 (5)</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="type locality">
<paragraph pageId="0" pageNumber="1">Type locality.</paragraph>
<paragraph pageId="0" pageNumber="1">
Georgia • Racha, Nikortsminda, Tsivtskala 2 Spring on left bank of the Shareula River near the power station; 
<geoCoordinate degrees="42" direction="north" minutes="28" orientation="latitude" precision="15" seconds="18" value="42.47167">42°28'18&quot;N</geoCoordinate>
, 
<geoCoordinate degrees="43" direction="east" minutes="03" orientation="longitude" precision="15" seconds="54" value="43.065">43°03'54&quot;E</geoCoordinate>
; 1084 m a.s.l.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="materials_examined">
<paragraph pageId="0" pageNumber="1">Material.</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Holotype</emphasis>
: Georgia • 1 adult, dry; type locality; 06 May 2018; J. Grego, L. Mumladze and M. 
<normalizedToken originalValue="Olšavský">Olsavsky</normalizedToken>
leg.; ISU FM-T012-H. 
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Other material</emphasis>
: Georgia • Fragmented shells; Racha, Nikorsminda, Shareula River Head (Shareula Cave); 
<geoCoordinate degrees="42" direction="north" minutes="28" orientation="latitude" precision="15" seconds="12" value="42.47">42°28'12&quot;N</geoCoordinate>
, 
<geoCoordinate degrees="43" direction="east" minutes="04" orientation="longitude" precision="15" seconds="4" value="43.067776">43°04'4&quot;E</geoCoordinate>
; 1105 m a.s.l.; date; 20.08.2017, J. Grego leg.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="diagnosis">
<paragraph pageId="0" pageNumber="1">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="1">
The new taxon significantly differs from all congeners by its flat shape with spire hidden in apertural profile and its planorboid coiling, a unique feature within the southwestern Caucasus stygobiotic 
<taxonomicName authorityName="Grego &amp; Mumladze" authorityYear="2020" class="Gastropoda" lsidName="" pageId="0" pageNumber="1" rank="class">Gastropoda</taxonomicName>
. Measurement comparison of 
<taxonomicName authorityName="Vinarski, Palatov &amp; Gloeer" authorityYear="2014" class="Gastropoda" family="Hydrobiidae" genus="Pontohoratia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pontohoratia" order="Littorinimorpha" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Pontohoratia</emphasis>
</taxonomicName>
and 
<taxonomicName authorityName="Grego &amp; Mumladze" authorityYear="2020" class="Gastropoda" family="Hydrobiidae" genus="Hausdorfenia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hausdorfenia" order="Littorinimorpha" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Hausdorfenia</emphasis>
</taxonomicName>
species is given in Table 
<tableCitation captionStart="Table 9" captionStartId="T9" captionText="Table 9. Measurement comparison of species in genera Pontohoratia Vinarski, Palatov &amp; Gloeer, 2014 and Hausdorfenia gen. nov." httpUri="http://table.plazi.org/id/D2DE94C54096E9E42D3318D268AEC2C2" pageId="0" pageNumber="1" tableUuid="D2DE94C54096E9E42D3318D268AEC2C2">9</tableCitation>
.
</paragraph>
<caption ID-Table-UUID="D2DE94C54096E9E42D3318D268AEC2C2" httpUri="http://table.plazi.org/id/D2DE94C54096E9E42D3318D268AEC2C2" pageId="0" pageNumber="1" start="Table 9" startId="T9">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Table 9.</emphasis>
Measurement comparison of species in genera 
<taxonomicName authorityName="Vinarski, Palatov &amp; Gloeer" authorityYear="2014" class="Gastropoda" family="Hydrobiidae" genus="Pontohoratia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pontohoratia" order="Littorinimorpha" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Pontohoratia</emphasis>
</taxonomicName>
Vinarski, Palatov &amp; 
<normalizedToken originalValue="Glöer">Gloeer</normalizedToken>
, 2014 and 
<taxonomicName authorityName="Grego &amp; Mumladze" authorityYear="2020" class="Gastropoda" family="Hydrobiidae" genus="Hausdorfenia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hausdorfenia" order="Littorinimorpha" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Hausdorfenia</emphasis>
</taxonomicName>
gen. nov.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="1">
<table pageId="0" pageNumber="1">
<tr pageId="0" pageNumber="1">
<th colspan="1" pageId="0" pageNumber="1" rowspan="2">
<emphasis italics="true" pageId="0" pageNumber="1">Genus species</emphasis>
</th>
<th colspan="1" pageId="0" pageNumber="1" rowspan="1">H</th>
<th colspan="1" pageId="0" pageNumber="1" rowspan="1">W</th>
<th colspan="1" pageId="0" pageNumber="1" rowspan="1">BH</th>
<th colspan="1" pageId="0" pageNumber="1" rowspan="1">BW</th>
<th colspan="1" pageId="0" pageNumber="1" rowspan="1">AH</th>
<th colspan="1" pageId="0" pageNumber="1" rowspan="1">AW</th>
<th colspan="1" pageId="0" pageNumber="1" rowspan="1">CA</th>
<th colspan="1" pageId="0" pageNumber="1" rowspan="2">H/W</th>
<th colspan="1" pageId="0" pageNumber="1" rowspan="2">AH / AW</th>
<th colspan="1" pageId="0" pageNumber="1" rowspan="2">W / BW</th>
<th colspan="1" pageId="0" pageNumber="1" rowspan="2">H/BH</th>
<th colspan="1" pageId="0" pageNumber="1" rowspan="2">H/AH</th>
<th colspan="1" pageId="0" pageNumber="1" rowspan="2">W / AW</th>
<th colspan="1" pageId="0" pageNumber="1" rowspan="2">H/(W- BW)</th>
</tr>
<tr pageId="0" pageNumber="1">
<th colspan="1" pageId="0" pageNumber="1" rowspan="1">mm</th>
<th colspan="1" pageId="0" pageNumber="1" rowspan="1">mm</th>
<th colspan="1" pageId="0" pageNumber="1" rowspan="1">mm</th>
<th colspan="1" pageId="0" pageNumber="1" rowspan="1">mm</th>
<th colspan="1" pageId="0" pageNumber="1" rowspan="1">mm</th>
<th colspan="1" pageId="0" pageNumber="1" rowspan="1">mm</th>
<th colspan="1" pageId="0" pageNumber="1" rowspan="1">deg.</th>
</tr>
<tr pageId="0" pageNumber="1">
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
<taxonomicName authorityName="Grego &amp; Mumladze" authorityYear="2020" class="Gastropoda" family="Hydrobiidae" genus="Pontohoratia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pontohoratia vinarskii" order="Littorinimorpha" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="vinarskii">
<emphasis italics="true" pageId="0" pageNumber="1">Pontohoratia vinarskii</emphasis>
</taxonomicName>
sp. nov. 
<emphasis bold="true" pageId="0" pageNumber="1">Holotype</emphasis>
LT
</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1.08</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1.47</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">0.87</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1.00</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">0.63</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">0.55</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">-20</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">0.73</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1.15</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1.47</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1.24</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1.71</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">2.67</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">2.30</td>
</tr>
<tr pageId="0" pageNumber="1">
<td colspan="1" pageId="0" pageNumber="1" rowspan="2">
<taxonomicName authorityName="Grego &amp; Mumladze" authorityYear="2020" class="Gastropoda" family="Hydrobiidae" genus="Pontohoratia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pontohoratia vinarskii" order="Littorinimorpha" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="vinarskii">
<emphasis italics="true" pageId="0" pageNumber="1">Pontohoratia vinarskii</emphasis>
</taxonomicName>
sp. nov. 
<emphasis bold="true" pageId="0" pageNumber="1">Paratype</emphasis>
LT
</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1.11</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1.42</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">0.95</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1.00</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">0.66</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">0.53</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">-24</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">0.78</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1.25</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1.42</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1.17</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1.68</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">2.68</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">2.64</td>
</tr>
<tr pageId="0" pageNumber="1">
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1.46</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1.50</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1.13</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1.08</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">0.75</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">0.63</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">30</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">0.97</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1.19</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1.39</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1.29</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1.95</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">2.38</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">3.48</td>
</tr>
<tr pageId="0" pageNumber="1">
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
<taxonomicName authorityName="Grego &amp; Mumladze" authorityYear="2020" class="Gastropoda" family="Hydrobiidae" genus="Pontohoratia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pontohoratia mapeli" order="Littorinimorpha" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="mapeli">
<emphasis italics="true" pageId="0" pageNumber="1">Pontohoratia mapeli</emphasis>
</taxonomicName>
sp. nov. 
<emphasis bold="true" pageId="0" pageNumber="1">Holotype</emphasis>
LT
</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">0.57</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1.38</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">0.55</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">0.95</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">0.42</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">0.45</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">-40</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">0.41</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">0.93</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1.45</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1.04</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1.36</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">3.07</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1.33</td>
</tr>
<tr pageId="0" pageNumber="1">
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
<taxonomicName authorityName="Grego &amp; Mumladze" authorityYear="2020" class="Gastropoda" family="Hydrobiidae" genus="Pontohoratia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pontohoratia pichkhaiai" order="Littorinimorpha" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="pichkhaiai">
<emphasis italics="true" pageId="0" pageNumber="1">Pontohoratia pichkhaiai</emphasis>
</taxonomicName>
sp. nov. 
<emphasis bold="true" pageId="0" pageNumber="1">Holotype</emphasis>
LT
</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">0.87</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1.42</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">0.66</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1.00</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">0.50</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">0.50</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">-45</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">0.61</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1.00</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1.42</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1.32</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1.74</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">2.84</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">2.07</td>
</tr>
<tr pageId="0" pageNumber="1">
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
<taxonomicName authorityName="Grego &amp; Mumladze" authorityYear="2020" class="Gastropoda" family="Hydrobiidae" genus="Hausdorfenia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hausdorfenia pseudohauffenia" order="Littorinimorpha" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="pseudohauffenia">
<emphasis italics="true" pageId="0" pageNumber="1">Hausdorfenia pseudohauffenia</emphasis>
</taxonomicName>
sp. nov. 
<emphasis bold="true" pageId="0" pageNumber="1">Holotype</emphasis>
LT
</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">0.82</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1.68</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">0.74</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1.11</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">0.53</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">0.58</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">-48</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">0.49</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">0.91</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1.51</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1.11</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1.55</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">2.90</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1.44</td>
</tr>
<tr pageId="0" pageNumber="1">
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
<taxonomicName genus="Husdorfenia" lsidName="Husdorfenia shareula" pageId="0" pageNumber="1" rank="species" species="shareula">
<emphasis italics="true" pageId="0" pageNumber="1">Husdorfenia shareula</emphasis>
</taxonomicName>
sp. nov. 
<emphasis bold="true" pageId="0" pageNumber="1">Paratype</emphasis>
LT
</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">0.53</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1.34</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">0.50</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">0.82</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">0.48</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">0.42</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">-57</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">0.40</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1.14</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1.63</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1.06</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1.10</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">3.19</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1.02</td>
</tr>
</table>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="description">
<paragraph pageId="0" pageNumber="1">Description.</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Shell</emphasis>
: planispiral, discoid with planorboid (slightly hyperstrophic) coiling and 1.34 mm in diameter. Descending 
<normalizedToken originalValue="2¼">21/4</normalizedToken>
whorls separated by a deep suture. Umbilicus very widely expanding. Shell colour milky white, surface smooth with very faint axial growth lines. Aperture circular, and its labral periphery is oblique to the columellar axis. It attached to the whole length of the adjacent body whorl by a narrow suture. Protoconch pitted over whole surface.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Operculum</emphasis>
: not known.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Animal body</emphasis>
: not known.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Holotype measurements</emphasis>
: H-0.53 mm; W-1.34 mm; BH-0.50 mm; BW-0.82 mm; AH-0.48 mm; AW-0.42 mm; CA: -57°.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Anatomy</emphasis>
: not known.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="etymology">
<paragraph pageId="0" pageNumber="1">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="1">
Name derived from the name of the Shareula River (მდინარე 
<normalizedToken originalValue="შარეულა">შარეულა</normalizedToken>
), left tributary of the Rioni River, in which valley and a nearby tributary the new taxon was found.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="habitat">
<paragraph pageId="0" pageNumber="1">Habitat.</paragraph>
<paragraph pageId="0" pageNumber="1">The intact empty shell was found in sandy sediment at the spring head in a small cave. The supposed habitat is stygobiotic.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="distribution">
<paragraph pageId="0" pageNumber="1">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="1">Except the type locality few similar fragments were found at the Shareula River Head (entrance of Shareula Cave).</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="conservation status">
<paragraph pageId="0" pageNumber="1">Conservation status.</paragraph>
<paragraph pageId="0" pageNumber="1">The number of known locations (2) is no more than 5 and EOO is smaller than 20 km2. There is no reason to suppose that AOO, EOO, number of locations, number of subpopulations or the number or mature individuals are declining however due to its extremely small EOO we assessed as Vulnerable (VU) D2.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="remarks">
<paragraph pageId="0" pageNumber="1">Remarks.</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Grego &amp; Mumladze" authorityYear="2020" class="Gastropoda" family="Hydrobiidae" genus="Hausdorfenia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hausdorfenia pseudohauffenia" order="Littorinimorpha" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="pseudohauffenia">
<emphasis italics="true" pageId="0" pageNumber="1">Hausdorfenia pseudohauffenia</emphasis>
</taxonomicName>
sp. nov. and 
<taxonomicName lsidName="P. shareula" pageId="0" pageNumber="1" rank="species" species="shareula">
<emphasis italics="true" pageId="0" pageNumber="1">P. shareula</emphasis>
</taxonomicName>
sp. nov. display shell features different from other members of the genus, as well as a characteristic operculum with a peg (at least in the former taxon). Both represent a new genus different from 
<taxonomicName authorityName="Vinarski, Palatov &amp; Gloeer" authorityYear="2014" class="Gastropoda" family="Hydrobiidae" genus="Pontohoratia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pontohoratia" order="Littorinimorpha" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Pontohoratia</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>