308 lines
28 KiB
XML
308 lines
28 KiB
XML
<document ID-DOI="10.1590/S1679-62252010000400005" ID-ISSN="1982-0224" ID-Zenodo-Dep="5422215" approvalRequired="87" approvalRequired_for_textStreams="77" approvalRequired_for_treatments="10" checkinTime="1630380595706" checkinUser="carolina" docAuthor="Bertaco, Vinicius A. & Malabarba, Luiz R." docDate="2010" docId="03FC9766FFF6D673FC12FE60FAB47A33" docLanguage="en" docName="NeotropIchthyol.2010.63.pdf" docOrigin="Neotropical Ichthyology 8 (4)" docSource="http://www.scielo.br/scielo.php?script=sci_arttext&pid=S1679-62252010000400005&lng=en&tlng=en" docStyle="DocumentStyle:85A40B78B715C4497F837D9D78407CE5.4:NeotropIchthyol.2003-2013.journal_article" docStyleId="85A40B78B715C4497F837D9D78407CE5" docStyleName="NeotropIchthyol.2003-2013.journal_article" docStyleVersion="4" docTitle="Hemibrycon tridens Eigenmann 1922" docType="treatment" docVersion="2" lastPageId="29" masterDocId="FFC5EF1EFFEAD66EFFB2FFB0FFAA7C11" masterDocTitle="A review of the Cis-Andean species of Hemibrycon Günther (Teleostei: Characiformes: Characidae: Stevardiinae), with description of two new species" masterLastPageNumber="737" masterPageNumber="737" pageId="28" pageNumber="737" updateTime="1630688377022" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>A review of the Cis-Andean species of Hemibrycon Günther (Teleostei: Characiformes: Characidae: Stevardiinae), with description of two new species</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Bertaco, Vinicius A.</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Malabarba, Luiz R.</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>Neotropical Ichthyology</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2010</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2010-12-31</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>8</mods:number>
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</mods:detail>
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<mods:detail type="issue">
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<mods:number>4</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>737</mods:start>
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<mods:end>737</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://www.scielo.br/scielo.php?script=sci_arttext&pid=S1679-62252010000400005&lng=en&tlng=en</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">10.1590/S1679-62252010000400005</mods:identifier>
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<mods:identifier type="ISSN">1982-0224</mods:identifier>
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<mods:identifier type="Zenodo-Dep">5422215</mods:identifier>
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</mods:mods>
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<treatment LSID="urn:lsid:plazi:treatment:03FC9766FFF6D673FC12FE60FAB47A33" httpUri="http://treatment.plazi.org/id/03FC9766FFF6D673FC12FE60FAB47A33" lastPageId="29" pageId="28">
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<subSubSection box="[928,1346,464,488]" pageId="28" type="nomenclature">
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<paragraph blockId="28.[928,1346,464,519]" box="[928,1346,464,488]" pageId="28">
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<heading box="[928,1346,464,488]" centered="true" fontSize="10" level="2" pageId="28" reason="2">
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<taxonomicName authority="Eigenmann, 1922" authorityName="Eigenmann" authorityYear="1922" box="[928,1346,464,488]" class="Actinopterygii" family="Characidae" genus="Hemibrycon" kingdom="Animalia" order="Characiformes" pageId="28" phylum="Chordata" rank="species" species="tridens">
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<emphasis bold="true" box="[928,1346,464,488]" pageId="28">
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<emphasis bold="true" box="[928,1146,464,488]" italics="true" pageId="28">Hemibrycon tridens</emphasis>
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Eigenmann, 1922
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="28" type="description">
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<paragraph blockId="28.[928,1346,464,519]" box="[1099,1175,496,519]" pageId="28">
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<figureCitation box="[1099,1175,496,519]" captionStart="Fig" captionStartId="29.[129,168,720,743]" captionTargetBox="[128,1450,257,711]" captionTargetId="figure-657@29.[128,1451,257,711]" captionTargetPageId="29" captionText="Fig. 25. Hemibrycon tridens, CAS 44358, holotype, 51.5 mm SL, male, upper río Ucayali basin, Peru. The black marks are preservation artifacts." figureDoi="http://doi.org/10.5281/zenodo.5422263" httpUri="https://zenodo.org/record/5422263/files/figure.png" pageId="28">
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<emphasis bold="true" box="[1099,1175,496,519]" pageId="28">Fig. 25</emphasis>
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</figureCitation>
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</paragraph>
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<paragraph blockId="28.[816,1458,561,743]" pageId="28">
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<taxonomicName authority="Eigenmann, 1922: 152" authorityName="Eigenmann" authorityPageNumber="152" authorityYear="1922" box="[816,1349,561,584]" class="Actinopterygii" family="Characidae" genus="Hemibrycon" kingdom="Animalia" order="Characiformes" pageId="28" phylum="Chordata" rank="species" species="tridens">
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<emphasis box="[816,1058,561,584]" italics="true" pageId="28">Hemibrycon tridens</emphasis>
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<materialsCitation box="[1071,1349,561,584]" collectionCode="IU" country="Peru" county="Uruhuasi" location="Uruhuasi" municipality="Eigenmann" pageId="28" specimenCount="1" stateProvince="Apurimac" typeStatus="holotype">Eigenmann, 1922: 152</materialsCitation>
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</taxonomicName>
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(species described in key, distribution: extralimital, río
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<collectingRegion box="[1350,1458,593,616]" country="Peru" name="Apurimac" pageId="28">Apurimac</collectingRegion>
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at
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<location LSID="urn:lsid:plazi:treatment:03FC9766FFF6D673FC12FE60FAB47A33:8E8A70ABFFF6D672FCC6FDC1FC707E99" box="[884,986,625,648]" country="Peru" county="Uruhuasi" municipality="Eigenmann" name="Uruhuasi" pageId="28" stateProvince="Apurimac">Uruhuasi</location>
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[
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<location LSID="urn:lsid:plazi:treatment:03FC9766FFF6D673FC12FE60FAB47A33:8E8A70ABFFF6D672FC5EFDC1FBCF7E99" box="[1004,1125,625,648]" country="Peru" county="Uruhuasi" municipality="Eigenmann" name="Curuhuasi" pageId="28" stateProvince="Apurimac">Curuhuasi</location>
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, upper
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<collectingRegion box="[1215,1344,625,648]" country="Peru" name="Ucayali" pageId="28">río Ucayali</collectingRegion>
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drainage,
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<collectingCountry box="[854,911,657,680]" name="Peru" pageId="28">Peru</collectingCountry>
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]). -
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<collectingMunicipality box="[941,1069,657,680]" pageId="28">Eigenmann</collectingMunicipality>
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, 1927: 402 (in key), 403-404 (pl. 96, fig. 3, original description, distribution:
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<collectingCounty box="[1345,1454,688,711]" pageId="28">Uruhuasi</collectingCounty>
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, southeastern
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<collectingCountry box="[1001,1056,720,743]" name="Peru" pageId="28">Peru</collectingCountry>
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;
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<typeStatus box="[1067,1165,720,743]" pageId="28">holotype</typeStatus>
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13723 IU).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="28" type="diagnosis">
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<paragraph blockId="28.[816,1459,779,1122]" pageId="28">
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<emphasis bold="true" box="[816,933,779,802]" pageId="28">Diagnosis.</emphasis>
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<taxonomicName authorityName="Eigenmann" authorityYear="1922" box="[938,1151,779,802]" class="Actinopterygii" family="Characidae" genus="Hemibrycon" kingdom="Animalia" order="Characiformes" pageId="28" phylum="Chordata" rank="species" species="tridens">
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<emphasis box="[938,1151,779,802]" italics="true" pageId="28">Hemibrycon tridens</emphasis>
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</taxonomicName>
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is readily distinguished from all congeners, except
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<taxonomicName authorityName=", Sep" authorityYear="1921" box="[1059,1141,811,834]" class="Actinopterygii" family="Characidae" genus="Hemibrycon" kingdom="Animalia" order="Characiformes" pageId="28" phylum="Chordata" rank="species" species="beni">
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<emphasis box="[1059,1078,811,834]" italics="true" pageId="28">H</emphasis>
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.
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<emphasis box="[1094,1141,811,834]" italics="true" pageId="28">beni</emphasis>
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</taxonomicName>
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, by the number of branched anal-fin rays (17
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<emphasis box="[991,1013,843,866]" italics="true" pageId="28">vs</emphasis>
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. 18-34,
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<figureCitation box="[1097,1160,843,866]" captionStart="Fig" captionStartId="3.[129,168,1924,1947]" captionTargetBox="[133,1445,1416,1903]" captionTargetPageId="3" captionText="Fig. 2. Tukey box plots of number of branched anal-fin rays in Hemibrycon species. Mean represented by thick vertical bar, and 25th and 75th percetiles as lateral borders of box plots. Number in parentheses = N." figureDoi="http://doi.org/10.5281/zenodo.5422219" httpUri="https://zenodo.org/record/5422219/files/figure.png" pageId="28">Fig. 2</figureCitation>
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).
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<taxonomicName authorityName="Eigenmann" authorityYear="1922" box="[1177,1385,843,866]" class="Actinopterygii" family="Characidae" genus="Hemibrycon" kingdom="Animalia" order="Characiformes" pageId="28" phylum="Chordata" rank="species" species="tridens">
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<emphasis box="[1177,1385,843,866]" italics="true" pageId="28">Hemibrycon tridens</emphasis>
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</taxonomicName>
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differs from
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<taxonomicName authorityName=", Sep" authorityYear="1921" box="[875,954,875,898]" class="Actinopterygii" family="Characidae" genus="Hemibrycon" kingdom="Animalia" order="Characiformes" pageId="28" phylum="Chordata" rank="species" species="beni">
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<emphasis box="[875,894,875,898]" italics="true" pageId="28">H</emphasis>
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.
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<emphasis box="[907,954,875,898]" italics="true" pageId="28">beni</emphasis>
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</taxonomicName>
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by the number of lateral line scales (39
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<emphasis box="[1386,1409,875,898]" italics="true" pageId="28">vs</emphasis>
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. 44- 53,
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<figureCitation box="[855,916,907,930]" captionStart="Fig" captionStartId="3.[809,848,1924,1947]" captionText="Fig. 3. Tukey box plots of number of lateral line scales in Hemibrycon species. Mean represented by thick vertical bar, and 25th and 75th percetiles as lateral borders of box plots. Number in parentheses = N." pageId="28">Fig. 3</figureCitation>
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), and by the number of scale rows above and below lateral line (6 and 5
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<emphasis box="[1041,1064,939,962]" italics="true" pageId="28">vs</emphasis>
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. 7-8 and 6-8, respectively). Among remaining Cis-Andean species of
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<taxonomicName baseAuthorityName="Malabarba & Weitzman" baseAuthorityYear="2003" box="[1214,1352,971,994]" class="Actinopterygii" family="Characidae" genus="Hemibrycon" kingdom="Animalia" order="Characiformes" pageId="28" phylum="Chordata" rank="genus">
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<emphasis box="[1214,1352,971,994]" italics="true" pageId="28">Hemibrycon</emphasis>
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</taxonomicName>
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, only
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<taxonomicName class="Actinopterygii" family="Characidae" genus="Hemibrycon" kingdom="Animalia" order="Characiformes" pageId="28" phylum="Chordata" rank="species" species="helleri">
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<emphasis box="[1432,1451,971,994]" italics="true" pageId="28">H</emphasis>
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.
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<emphasis box="[816,888,1003,1026]" italics="true" pageId="28">helleri</emphasis>
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</taxonomicName>
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and
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<taxonomicName box="[947,1118,1003,1026]" class="Actinopterygii" family="Characidae" genus="Hemibrycon" kingdom="Animalia" order="Characiformes" pageId="28" phylum="Chordata" rank="species" species="mikrostiktos">
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<emphasis box="[947,966,1003,1026]" italics="true" pageId="28">H</emphasis>
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.
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<emphasis box="[983,1118,1003,1026]" italics="true" pageId="28">mikrostiktos</emphasis>
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</taxonomicName>
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have lower branched anal-fin rays (19-23 and 18-21). From these species,
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<taxonomicName authorityName="Eigenmann" authorityYear="1922" box="[1279,1384,1035,1058]" class="Actinopterygii" family="Characidae" genus="Hemibrycon" kingdom="Animalia" order="Characiformes" pageId="28" phylum="Chordata" rank="species" species="tridens">
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<emphasis box="[1279,1298,1035,1058]" italics="true" pageId="28">H</emphasis>
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.
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<emphasis box="[1310,1384,1035,1058]" italics="true" pageId="28">tridens</emphasis>
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</taxonomicName>
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differs additionally by the number of lateral line scales (39
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<emphasis box="[1362,1384,1067,1090]" italics="true" pageId="28">vs</emphasis>
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. 40-42 and 42-45, respectively).
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="29" pageId="28" type="description">
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<paragraph blockId="28.[816,1459,1158,2044]" pageId="28">
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<emphasis bold="true" box="[816,953,1158,1181]" pageId="28">Description.</emphasis>
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Morphometric data for
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<taxonomicName authorityName="Eigenmann" authorityYear="1922" box="[1197,1300,1158,1181]" class="Actinopterygii" family="Characidae" genus="Hemibrycon" kingdom="Animalia" order="Characiformes" pageId="28" phylum="Chordata" rank="species" species="tridens">
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<emphasis box="[1197,1216,1158,1181]" italics="true" pageId="28">H</emphasis>
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.
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<emphasis box="[1227,1300,1158,1181]" italics="true" pageId="28">tridens</emphasis>
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</taxonomicName>
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summarized in
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<tableCitation box="[816,900,1190,1213]" captionStart="Table 5" captionStartId="21.[129,191,261,284]" captionText="Table 5. Morphometric data of holotype (H) of Hemibrycon polyodon(BMNH 1858.7.25.41) from Guayaquil, Ecuador, and non-typesfrom tributary of río Pastaza, río Marañon basin (KU 19978, 19992, 20004), and of holotype of Hemibrycon tridens (CAS 44358) from rio Apurimac, upper río Ucayali basin, Curuhuasi, Peru.m, males; f, females.SD = Standard deviation." pageId="28">Table 5</tableCitation>
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. Largest male
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<quantity box="[1063,1161,1190,1213]" metricMagnitude="-2" metricUnit="m" metricValue="5.15" pageId="28" unit="mm" value="51.5">51.5 mm</quantity>
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SL. Body compressed and elongate; greatest body depth anterior to dorsal-fin origin. Dorsal profile of head convex. Dorsal body profile convex from occipital bone to base of last dorsal-fin ray; straight from this point to adipose-fin origin. Ventral profile of head slightly convex. Ventral body profile convex from pectoral-fin origin to pelvic-fin origin, and straight to slightly convex to anal-fin origin. Body profile along anal-fin base posterodorsally slanted. Caudal peduncle elongate, nearly straight to slightly concave along dorsal and ventral margins.
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</paragraph>
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<paragraph blockId="28.[816,1459,1158,2044]" pageId="28">Snout rounded from margin of upper lip to vertical through anterior nostrils. Head relatively small. Mouth sub-terminal, mouth slit nearly at horizontal through inferior margin of eye. Maxillalongandslightlycurved, alignedatangleofapproximately 45º to longitudinal body axis, and its posterior tip extending below orbit, reaching nearly to vertical through center of eye.</paragraph>
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<paragraph blockId="28.[816,1459,1158,2044]" pageId="28">Premaxilla with two tooth rows; outer row with 5 tricuspid teeth with central cusp slightly longer; inner row with 4 tri- to pentacuspids teeth, gradually decreasing in length from first to third teeth and last tooth smallest; central cusp twice or three times longer and broader than other cusps. Maxilla fully toothed with 12 tricuspid teeth, with central cusp longer. Three anteriormost dentary teeth larger, with 5 cusps, followed by medium sized tooth with 3 cusps, and 8 teeth with 1-3 cusps or conical; central cusp in all teeth two to three times longer and broader than other cusps. Cusp tips slightly curved posteriorly and lingually.</paragraph>
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<paragraph blockId="29.[129,1045,180,205]" box="[129,1045,180,205]" pageId="29">
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766 A review of the Cis-Andean species of
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<taxonomicName baseAuthorityName="Malabarba & Weitzman" baseAuthorityYear="2003" box="[925,1045,180,201]" class="Actinopterygii" family="Characidae" genus="Hemibrycon" kingdom="Animalia" order="Characiformes" pageId="29" phylum="Chordata" rank="genus">
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<emphasis box="[925,1045,180,201]" italics="true" pageId="29">Hemibrycon</emphasis>
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</taxonomicName>
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</paragraph>
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<caption ID-DOI="http://doi.org/10.5281/zenodo.5422263" ID-Zenodo-Dep="5422263" httpUri="https://zenodo.org/record/5422263/files/figure.png" pageId="29" startId="29.[129,168,720,743]" targetBox="[128,1450,257,711]" targetPageId="29">
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<paragraph blockId="29.[129,1451,720,775]" pageId="29">
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<emphasis bold="true" box="[129,213,720,743]" pageId="29">Fig. 25.</emphasis>
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<taxonomicName authority=", CAS" authorityName="CAS" box="[219,499,720,743]" class="Actinopterygii" family="Characidae" genus="Hemibrycon" kingdom="Animalia" order="Characiformes" pageId="29" phylum="Chordata" rank="species" species="tridens">
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<emphasis box="[219,434,720,743]" italics="true" pageId="29">Hemibrycon tridens</emphasis>
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, CAS
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</taxonomicName>
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44358, holotype, 51.5 mm SL, male, upper río Ucayali basin, Peru. The black marks are preservation artifacts.
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</paragraph>
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</caption>
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<paragraph blockId="29.[129,772,814,1765]" pageId="29">
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Dorsal-fin rays ii,8, first unbranched ray approximately onehalf length of second ray. Dorsal-fin origin located posterior to middle of SL and posterior to vertical through pelvic-fin origin. Profile of distal margin of dorsal fin slightly concave. Dorsal fin without bony hooks. Adipose-fin located at vertical through insertion posterior to last anal-fin ray.Anal-fin rays iii,17 (
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<figureCitation captionStart="Fig" captionStartId="3.[129,168,1924,1947]" captionTargetBox="[133,1445,1416,1903]" captionTargetPageId="3" captionText="Fig. 2. Tukey box plots of number of branched anal-fin rays in Hemibrycon species. Mean represented by thick vertical bar, and 25th and 75th percetiles as lateral borders of box plots. Number in parentheses = N." figureDoi="http://doi.org/10.5281/zenodo.5422219" httpUri="https://zenodo.org/record/5422219/files/figure.png" pageId="29">Fig. 2</figureCitation>
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). Anal-fin profile slightly nearly straight. Anal-fin origin approximately at vertical through insertion in the last dorsal fin rays.Anal-fin rays of male bearing one pair of small bony hooks along posterolateral border of each segment of lepidotrichia, along last unbranched ray and sixteenth branched rays. Hooks located along anteriormost branch and proximal
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<superScript attach="right" box="[656,664,1165,1179]" fontSize="6" pageId="29">1</superScript>
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/2 to
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<superScript attach="right" box="[717,725,1165,1179]" fontSize="6" pageId="29">2</superScript>
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/3 of each ray. Pectoral-fin rays i,10. Pectoral-fin tip surpassing pelvicfin origin. Pectoral fin without bony hooks. Pelvic-fin rays i,6,i. Pelvic-fin tip nearly reaching anal-fin origin. Pelvic-fin origin located 4 predorsal scales anterior to vertical through dorsalfin origin. Pelvic fin of male bearing one small bony hook per segment of lepidotrichia along ventromedial border of second to seventh branched rays. Caudal fin forked with 19 principal rays without bony hooks; lobes similar in size. Caudal-fin base with a few scales. Dorsal procurrent rays 10 and ventral procurrent rays 10 (
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<typeStatus box="[330,424,1486,1509]" pageId="29">holotype</typeStatus>
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x-ray).
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</paragraph>
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<paragraph blockId="29.[129,772,814,1765]" pageId="29">
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Scales cycloid, moderately large. Lateral line complete with 39 (
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<figureCitation box="[170,235,1550,1573]" captionStart="Fig" captionStartId="3.[809,848,1924,1947]" captionText="Fig. 3. Tukey box plots of number of lateral line scales in Hemibrycon species. Mean represented by thick vertical bar, and 25th and 75th percetiles as lateral borders of box plots. Number in parentheses = N." pageId="29">Fig. 3</figureCitation>
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). Scale rows between dorsal-fin origin and lateral line 6; scale rows between lateral line and pelvic-fin origin 5. Predorsal scales 14, arranged in regular series. Scales rows around caudal peduncle 14.
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</paragraph>
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<paragraph blockId="29.[129,772,814,1765]" pageId="29">
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Precaudal vertebrae 18; caudal vertebrae 22; total vertebrae 40. Supraneurals 5 (
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<typeStatus box="[335,429,1710,1733]" pageId="29">holotype</typeStatus>
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x-ray). Gill rakers on upper limb of outer gill arch 5, on lower limb 10.
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</paragraph>
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<paragraph blockId="29.[129,772,1796,2043]" lastBlockId="29.[809,1451,814,869]" pageId="29">
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<emphasis bold="true" box="[129,302,1796,1819]" pageId="29">Color in alcohol.</emphasis>
|
|
<typeStatus box="[306,400,1796,1819]" pageId="29">Holotype</typeStatus>
|
|
badly preserved. Humeral and caudal peduncle spot not evident. Base of caudal fin and middle rays black pigmented. Distal half of dorsal fin with dark pigmentation, and caudal and anal fin with dark brown pigmentation diffuse. Pectoral and pelvic fins hyaline. Body yellowish. Left side of
|
|
<typeStatus box="[129,221,1956,1979]" pageId="29">holotype</typeStatus>
|
|
has three black marks due to preservation artifacts (
|
|
<figureCitation box="[136,211,1988,2011]" captionStart="Fig" captionStartId="29.[129,168,720,743]" captionTargetBox="[128,1450,257,711]" captionTargetId="figure-657@29.[128,1451,257,711]" captionTargetPageId="29" captionText="Fig. 25. Hemibrycon tridens, CAS 44358, holotype, 51.5 mm SL, male, upper río Ucayali basin, Peru. The black marks are preservation artifacts." figureDoi="http://doi.org/10.5281/zenodo.5422263" httpUri="https://zenodo.org/record/5422263/files/figure.png" pageId="29">Fig. 25</figureCitation>
|
|
). Eigenmann (1927: 404) described the color pattern “A dark lateral band, caudal spot, and middle caudal rays dusky; distal parts of dorsal and anal and outer part of caudal lobes dusky; margin of lower caudal lobe dusky”.
|
|
</paragraph>
|
|
<paragraph blockId="29.[809,1451,910,997]" pageId="29">
|
|
<emphasis bold="true" box="[809,1042,910,933]" pageId="29">Sexual dimorphism.</emphasis>
|
|
Male of
|
|
<taxonomicName box="[1139,1357,910,933]" class="Actinopterygii" family="Characidae" genus="Hemibrycon" kingdom="Animalia" order="Characiformes" pageId="29" phylum="Chordata" rank="species" species="tridens">
|
|
<emphasis box="[1139,1357,910,933]" italics="true" pageId="29">Hemibrycon tridens</emphasis>
|
|
</taxonomicName>
|
|
is easily recognized by the presence of bony hooks on the anal- and pelvic-fin rays. No gill gland was observed on first gill arch.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection pageId="29" type="distribution">
|
|
<paragraph blockId="29.[809,1451,1038,1093]" pageId="29">
|
|
<emphasis bold="true" box="[809,949,1038,1061]" pageId="29">Distribution.</emphasis>
|
|
<taxonomicName box="[953,1156,1038,1061]" class="Actinopterygii" family="Characidae" genus="Hemibrycon" kingdom="Animalia" order="Characiformes" pageId="29" phylum="Chordata" rank="species" species="tridens">
|
|
<emphasis box="[953,1156,1038,1061]" italics="true" pageId="29">Hemibrycon tridens</emphasis>
|
|
</taxonomicName>
|
|
is known from río
|
|
<collectingRegion box="[1342,1448,1038,1061]" country="Peru" name="Apurimac" pageId="29">Apurimac</collectingRegion>
|
|
, upper
|
|
<collectingRegion box="[874,992,1070,1093]" country="Peru" name="Ucayali" pageId="29">río Ucayali</collectingRegion>
|
|
drainage, Curuhuasi,
|
|
<collectingCountry box="[1223,1272,1070,1093]" name="Peru" pageId="29">Peru</collectingCountry>
|
|
(
|
|
<figureCitation box="[1285,1349,1070,1093]" captionStart="Fig" captionStartId="4.[136,175,1739,1762]" captionTargetBox="[145,760,1305,1720]" captionTargetId="figure-936@4.[136,778,1291,1720]" captionTargetPageId="4" captionText="Fig. 4. Map of central and northern South America showing geographic distribution of Hemibrycon beni (1, solid hexagon), H. divisorensis (2, open triangle), H. helleri (3, ellipse), H. huambonicus (4, solid triangles), H. jelskii (5, solid dots), H. inambari (6, rectangles), H. metae (7, solid squares), H. mikrostiktos (8, open square), H. polyodon (9, inverted triangle), H. surinamensis (10, solid diamonds), H. taeniurus (11,Trinidad Island), H.tridens (12, open circle).Some symbols represent more than one lot or locality of specimens. 1-12 = type locality of each species." figureDoi="http://doi.org/10.5281/zenodo.5422221" httpUri="https://zenodo.org/record/5422221/files/figure.png" pageId="29">Fig. 4</figureCitation>
|
|
).
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection pageId="29" type="discussion">
|
|
<paragraph blockId="29.[809,1452,1138,1449]" pageId="29">
|
|
<emphasis bold="true" box="[809,914,1138,1161]" pageId="29">Remarks.</emphasis>
|
|
Eigenmann (1922: 152) proposed
|
|
<taxonomicName box="[1252,1451,1138,1161]" class="Actinopterygii" family="Characidae" genus="Hemibrycon" kingdom="Animalia" order="Characiformes" pageId="29" phylum="Chordata" rank="species" species="tridens">
|
|
<emphasis box="[1252,1451,1138,1161]" italics="true" pageId="29">Hemibrycon tridens</emphasis>
|
|
</taxonomicName>
|
|
based on
|
|
<specimenCount box="[910,1056,1170,1193]" pageId="29" type="generic">one specimen</specimenCount>
|
|
, diagnosed in his key to
|
|
<taxonomicName class="Actinopterygii" family="Characidae" genus="Hemibrycon" kingdom="Animalia" order="Characiformes" pageId="29" phylum="Chordata" rank="species" species="undetermined">
|
|
<emphasis box="[1323,1451,1170,1193]" italics="true" pageId="29">Hemibrycon</emphasis>
|
|
species.
|
|
</taxonomicName>
|
|
Only in 1927, Eigenmann presented a full description, and designated
|
|
<accessionNumber box="[981,1109,1234,1257]" httpUri="http://www.ncbi.nlm.nih.gov/protein/IUM13723" pageId="29">IUM 13723</accessionNumber>
|
|
(now
|
|
<accessionNumber box="[1183,1313,1234,1257]" httpUri="http://www.ncbi.nlm.nih.gov/protein/CAS44358" pageId="29">CAS 44358</accessionNumber>
|
|
) as the type specimen. The
|
|
<typeStatus box="[965,1056,1266,1289]" pageId="29">holotype</typeStatus>
|
|
is the only known specimen. The type locality provided by Eigenmann (1922) for
|
|
<taxonomicName box="[1250,1352,1298,1321]" class="Actinopterygii" family="Characidae" genus="Hemibrycon" kingdom="Animalia" order="Characiformes" pageId="29" phylum="Chordata" rank="species" species="tridens">
|
|
<emphasis box="[1250,1269,1298,1321]" italics="true" pageId="29">H</emphasis>
|
|
.
|
|
<emphasis box="[1280,1352,1298,1321]" italics="true" pageId="29">tridens</emphasis>
|
|
</taxonomicName>
|
|
was “Rio
|
|
<collectingRegion box="[809,914,1330,1353]" country="Peru" name="Apurimac" pageId="29">Apurimac</collectingRegion>
|
|
at Uruhuasi”. According to Hernán Ortega (pers. comm.), the correct type locality of
|
|
<taxonomicName box="[1159,1259,1362,1385]" class="Actinopterygii" family="Characidae" genus="Hemibrycon" kingdom="Animalia" order="Characiformes" pageId="29" phylum="Chordata" rank="species" species="tridens">
|
|
<emphasis box="[1159,1178,1362,1385]" italics="true" pageId="29">H</emphasis>
|
|
.
|
|
<emphasis box="[1188,1259,1362,1385]" italics="true" pageId="29">tridens</emphasis>
|
|
</taxonomicName>
|
|
is the río
|
|
<collectingRegion box="[1349,1451,1362,1385]" country="Peru" name="Apurimac" pageId="29">Apurimac</collectingRegion>
|
|
at Curuhuasi, upper
|
|
<collectingRegion box="[1033,1154,1394,1417]" country="Peru" name="Ucayali" pageId="29">río Ucayali</collectingRegion>
|
|
drainage. He believed that Eigenmann erroneously wrote “Uruhuasi” instead of Curuhuasi.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection pageId="29" type="materials_examined">
|
|
<paragraph blockId="29.[809,1451,1491,1570]" pageId="29">
|
|
<emphasis bold="true" box="[809,1015,1491,1512]" pageId="29">Material examined.</emphasis>
|
|
<materialsCitation accessionNumber="CAS 44358" collectionCode="CAS" collectorName="F. Rosenberg." country="Peru" location="Cusco" pageId="29" specimenCount="1" stateProvince="Cusco" typeStatus="holotype">
|
|
<emphasis bold="true" box="[1024,1125,1491,1512]" pageId="29">
|
|
<typeStatus box="[1024,1121,1491,1512]" pageId="29">Holotype</typeStatus>
|
|
.
|
|
</emphasis>
|
|
<accessionNumber box="[1135,1254,1491,1512]" httpUri="http://www.ncbi.nlm.nih.gov/protein/CAS44358" pageId="29">
|
|
<collectionCode box="[1135,1182,1491,1512]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="29" type="Museum">CAS</collectionCode>
|
|
44358
|
|
</accessionNumber>
|
|
, male (x-ray),
|
|
<quantity metricMagnitude="-2" metricUnit="m" metricValue="5.15" pageId="29" unit="mm" value="51.5">51.5 mm</quantity>
|
|
SL,
|
|
<collectingCountry box="[901,949,1520,1541]" name="Peru" pageId="29">Peru</collectingCountry>
|
|
, 24 leagues from
|
|
<collectingRegion box="[1138,1202,1520,1541]" country="Peru" name="Cusco" pageId="29">Cusco</collectingRegion>
|
|
, Curuhuasi, province of Aymaraes, Department of
|
|
<collectingRegion box="[1064,1164,1549,1570]" country="Peru" name="Apurimac" pageId="29">Apurimac</collectingRegion>
|
|
,
|
|
<collectorName box="[1174,1310,1549,1570]" pageId="29">F. Rosenberg.</collectorName>
|
|
</materialsCitation>
|
|
</paragraph>
|
|
</subSubSection>
|
|
</treatment>
|
|
</document> |