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<document id="D001C19B7E186C65449E19DAF2274123" ID-DOI="10.5281/zenodo.4508002" ID-ISSN="2345-7600" ID-Zenodo-Dep="4508002" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1612542801306" checkinUser="carolina" docAuthor="Wu, Jie &amp; Zhou, Hong-zhang" docDate="2006" docId="03F987CD62692A39E85EF9DEA5F1CA3A" docLanguage="en" docName="RafflesBZool.54.2.215-223.pdf" docOrigin="Raffles Bulletin of Zoology 54 (2)" docStyle="DocumentStyle:C6443723295B2E4C6F22B9B23F08089C.5:RafflesBZool.2006-2013.journal_article" docStyleId="C6443723295B2E4C6F22B9B23F08089C" docStyleName="RafflesBZool.2006-2013.journal_article" docStyleVersion="5" docTitle="Borolinus bisetifer Wu &amp; Zhou 2006, new species" docType="treatment" docVersion="4" lastPageNumber="218" masterDocId="FFC0FFB562682A3AE95EFF8AA421CC62" masterDocTitle="Description Of A New Species Of Borolinus Bernhauer (Coleoptera: Staphylinidae: Osoriinae) From Yunnan, China, With Two New Records From Southern China" masterLastPageNumber="223" masterPageNumber="215" pageNumber="216" updateTime="1698867385935" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="A4D7F8FAC7515D2B5555C595F4B3B20E">Description Of A New Species Of Borolinus Bernhauer (Coleoptera: Staphylinidae: Osoriinae) From Yunnan, China, With Two New Records From Southern China</mods:title>
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<mods:namePart id="F69B5DCAED4EBAC8CA22E15C9F03A4BC">Wu, Jie</mods:namePart>
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<mods:namePart id="EA19D83A33274DC9D4BCFAB1EDBEDC2E">Zhou, Hong-zhang</mods:namePart>
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<mods:typeOfResource id="918AA9A49945D11969E09F5B1D0CF71C">text</mods:typeOfResource>
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<mods:title id="CF58DFC056DA6573936DBDCAB974EE11">Raffles Bulletin of Zoology</mods:title>
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<mods:part id="00AA6F384A86741361A90386A06324A9">
<mods:date id="FF8553C332446B03E0AAA3066F82306E">2006</mods:date>
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<mods:number id="9B791F7441F74C271E2B58B0772B744C">2006-08-31</mods:number>
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<mods:number id="AF6A87FEB3A1080E0D761E0998B6FABB">54</mods:number>
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<treatment id="03F987CD62692A39E85EF9DEA5F1CA3A" LSID="urn:lsid:plazi:treatment:03F987CD62692A39E85EF9DEA5F1CA3A" httpUri="http://treatment.plazi.org/id/03F987CD62692A39E85EF9DEA5F1CA3A" lastPageId="3" lastPageNumber="218" pageId="1" pageNumber="216">
<subSubSection id="C34A655062692A3BE85EF9DEA643CA0E" box="[256,610,1620,1644]" pageId="1" pageNumber="216" type="nomenclature">
<paragraph id="8BEF36DB62692A3BE85EF9DEA643CA0E" blockId="1.[256,610,1620,1675]" box="[256,610,1620,1644]" pageId="1" pageNumber="216">
<heading id="D0A781B762692A3BE85EF9DEA643CA0E" bold="true" box="[256,610,1620,1644]" centered="true" fontSize="10" level="2" pageId="1" pageNumber="216" reason="2">
<emphasis id="B924EAC962692A3BE85EF9DEA643CA0E" bold="true" box="[256,610,1620,1644]" pageId="1" pageNumber="216">
<taxonomicName id="4C504D5862692A3BE85EF9DEA5EFCA0E" ID-CoL="MKSB" authority="Wu &amp; Zhou, 2006" authorityName="Wu &amp; Zhou" authorityYear="2006" box="[256,462,1620,1644]" class="Insecta" family="Staphylinidae" genus="Borolinus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="216" phylum="Arthropoda" rank="species" species="bisetifer" status="sp. nov.">
<emphasis id="B924EAC962692A3BE85EF9DEA5EFCA0E" bold="true" box="[256,462,1620,1644]" italics="true" pageId="1" pageNumber="216">Borolinus bisetifer</emphasis>
</taxonomicName>
,
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<paragraph id="8BEF36DB62692A3BE82FF9FEA5D1CAE9" blockId="1.[256,610,1620,1675]" box="[369,496,1652,1675]" pageId="1" pageNumber="216">
(
<figureCitation id="136B2A5E62692A3BE827F9FEA5C6CAE9" box="[377,487,1652,1675]" captionStart="Figs" captionStartId="2.[137,181,2035,2056]" captionTargetBox="[131,1468,262,1974]" captionTargetId="figure-87@2.[113,1593,220,2000]" captionTargetPageId="2" captionText="Figs. 1-11. Borolinus bisetifer, new species: 1, head and pronotum in dorsal view; 2, aedeagus in dorsal view; 3, aedeagus in lateral view; 4, 9th abdominal sternite of male; 5, left half of 9th abdominal sternite of female; 6, mandibles; 7, 9th and 10th abdominal tergites of male; 8, labrum; 9, mentum; 10, maxilla; 11, ligula and labial palpi. Scale bars: Figs. 1, 6 = 1 mm; Figs. 2-5, 7-10, 11 = 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.4508004" httpUri="https://zenodo.org/record/4508004/files/figure.png" pageId="1" pageNumber="216">Figs. 1-11</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C34A655062692A3BE92FF939A6A2CB17" pageId="1" pageNumber="216" type="materials_examined">
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<emphasis id="B924EAC962692A3BE92FF939A54DCAAA" bold="true" box="[113,364,1715,1736]" pageId="1" pageNumber="216">
<emphasis id="B924EAC962692A3BE92FF939A570CAAA" bold="true" box="[113,337,1715,1736]" italics="true" pageId="1" pageNumber="216">Material examined.</emphasis>
</emphasis>
<materialsCitation id="3B383C8662692A3BE824F939A48BCB60" collectingDate="2004-02-10" collectionCode="IOZ-, CAS" collectorName="Jie Wu" country="China" elevation="810" location="Menglun" municipality="Xishuangbanna" pageId="1" pageNumber="216" specimenCount="1" stateProvince="Yunnan">
Holotype- male,
<collectingCountry id="F347764B62692A3BEB61F939A6A3CAAA" box="[575,642,1715,1736]" name="China" pageId="1" pageNumber="216">China</collectingCountry>
,
<collectingRegion id="4994F83962692A3BEBCBF939A6CDCAAA" box="[661,748,1715,1736]" country="China" name="Yunnan" pageId="1" pageNumber="216">Yunnan</collectingRegion>
,
<collectingMunicipality id="6B8BACA162692A3BE92FF95AA528CA87" box="[113,265,1744,1765]" pageId="1" pageNumber="216">Xishuangbanna</collectingMunicipality>
,
<location id="8E8F600062692A3BE84CF95AA54ACA87" LSID="urn:lsid:plazi:treatment:03F987CD62692A39E85EF9DEA5F1CA3A:8E8F600062692A3BE84CF95AA54ACA87" box="[274,363,1744,1765]" country="China" municipality="Xishuangbanna" name="Menglun" pageId="1" pageNumber="216" stateProvince="Yunnan">Menglun</location>
,
<quantity id="4CA89B3E62692A3BE828F95AA594CA87" box="[374,437,1744,1765]" metricMagnitude="2" metricUnit="m" metricValue="8.1" pageId="1" pageNumber="216" unit="m" value="810.0">
<elevation id="007DD1E862692A3BE828F95AA594CA87" box="[374,437,1744,1765]" metricMagnitude="2" metricUnit="m" metricValue="8.1" pageId="1" pageNumber="216" unit="m" value="810.0">810 m</elevation>
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,
<date id="FFEE101B62692A3BE8E1F95AA61BCA87" box="[447,570,1744,1765]" pageId="1" pageNumber="216" value="2004-02-10">
<collectingDate id="EFAAE9F362692A3BE8E1F95AA61BCA87" box="[447,570,1744,1765]" pageId="1" pageNumber="216" value="2004-02-10">10 Feb.2004</collectingDate>
</date>
,
<collectorName id="26A5530D62692A3BEB1AF95AA6A9CA87" box="[580,648,1744,1765]" pageId="1" pageNumber="216">Jie Wu</collectorName>
coll.(
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<collectionCode id="ED41AE1E62692A3BE92FF967A485CB60" box="[113,164,1773,1794]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="1" pageNumber="216" type="Museum">CAS</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BEF36DB62692A3BE92FF8ADA6A2CB17" blockId="1.[113,752,1830,1909]" pageId="1" pageNumber="216">
<materialsCitation id="3B383C8662692A3BE92FF8ADA48BCB17" collectingDate="2004-02-10" collectionCode="IOZ-, CAS" collectorName="Jie Wu &amp; Jiantao Yin" country="China" elevation="810" location="Menglun" municipality="Xishuangbanna" pageId="1" pageNumber="216" specimenCount="6" specimenCount-female="5" specimenCount-male="1" stateProvince="Yunnan" typeStatus="paratype">
<typeStatus id="54EB887962692A3BE92FF8ADA4F0CB5E" box="[113,209,1831,1852]" pageId="1" pageNumber="216" type="paratype">Paratypes</typeStatus>
.
<emphasis id="B924EAC962692A3BE983F8ACA4C8CB59" bold="true" box="[221,233,1830,1851]" pageId="1" pageNumber="216"></emphasis>
<specimenCount id="9D56FD5262692A3BE9ACF8ADA516CB5E" box="[242,311,1831,1852]" pageId="1" pageNumber="216" type="male">1 male</specimenCount>
,
<specimenCount id="9D56FD5262692A3BE81AF8ADA584CB5E" box="[324,421,1831,1852]" pageId="1" pageNumber="216" type="female">5 females</specimenCount>
,
<collectingCountry id="F347764B62692A3BE8ECF8ADA5CFCB5E" box="[434,494,1831,1852]" name="China" pageId="1" pageNumber="216">China</collectingCountry>
,
<collectingRegion id="4994F83962692A3BE8A4F8ADA669CB5E" box="[506,584,1831,1852]" country="China" name="Yunnan" pageId="1" pageNumber="216">Yunnan</collectingRegion>
,
<collectingMunicipality id="6B8BACA162692A3BEB0AF8ADA6CDCB5E" box="[596,748,1831,1852]" pageId="1" pageNumber="216">Xishuangbanna</collectingMunicipality>
,
<location id="8E8F600062692A3BE92FF8C9A4EBCB3A" LSID="urn:lsid:plazi:treatment:03F987CD62692A39E85EF9DEA5F1CA3A:8E8F600062692A3BE92FF8C9A4EBCB3A" box="[113,202,1859,1880]" country="China" municipality="Xishuangbanna" name="Menglun" pageId="1" pageNumber="216" stateProvince="Yunnan">Menglun</location>
,
<quantity id="4CA89B3E62692A3BE988F8C9A536CB3A" box="[214,279,1859,1880]" metricMagnitude="2" metricUnit="m" metricValue="8.1" pageId="1" pageNumber="216" unit="m" value="810.0">
<elevation id="007DD1E862692A3BE988F8C9A536CB3A" box="[214,279,1859,1880]" metricMagnitude="2" metricUnit="m" metricValue="8.1" pageId="1" pageNumber="216" unit="m" value="810.0">810 m</elevation>
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,
<date id="FFEE101B62692A3BE87DF8C9A581CB3A" box="[291,416,1859,1880]" pageId="1" pageNumber="216" value="2004-02-10">
<collectingDate id="EFAAE9F362692A3BE87DF8C9A581CB3A" box="[291,416,1859,1880]" pageId="1" pageNumber="216" value="2004-02-10">10 Feb.2004</collectingDate>
</date>
,
<collectorName id="26A5530D62692A3BE8F2F8C9A5D3CB3A" box="[428,498,1859,1880]" pageId="1" pageNumber="216">Jie Wu</collectorName>
&amp;
<collectorName id="26A5530D62692A3BEB4BF8C9A6A4CB3A" box="[533,645,1859,1880]" pageId="1" pageNumber="216">Jiantao Yin</collectorName>
coll.(
<collectionCode id="ED41AE1E62692A3BEBE3F8C9A6D1CB3A" box="[701,752,1859,1880]" pageId="1" pageNumber="216">IOZ-</collectionCode>
<collectionCode id="ED41AE1E62692A3BE92FF8EAA482CB17" box="[113,163,1888,1909]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="1" pageNumber="216" type="Museum">CAS</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B383C8662692A3BE9EBF8EAA65ECB17" box="[181,639,1888,1909]" collectingDate="2004-02-10" collectionCode="ZRC" collectorName="Jie Wu &amp; Jiantao Yin" country="China" elevation="810" location="Menglun" municipality="Xishuangbanna" pageId="1" pageNumber="216" specimenCount="1" specimenCount-male="1" stateProvince="Yunnan" typeStatus="paratype">
<specimenCount id="9D56FD5262692A3BE9EBF8EAA4DBCB17" box="[181,250,1888,1909]" pageId="1" pageNumber="216" type="male">1 male</specimenCount>
, same data as in other
<typeStatus id="54EB887962692A3BE884F8EAA617CB17" box="[474,566,1888,1909]" pageId="1" pageNumber="216" type="paratype">paratypes</typeStatus>
(
<collectionCode id="ED41AE1E62692A3BEB1AF8EAA659CB17" box="[580,632,1888,1909]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="1" pageNumber="216">ZRC</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C34A655062692A39E92FF817A4F9C81A" lastPageId="3" lastPageNumber="218" pageId="1" pageNumber="216" type="description">
<paragraph id="8BEF36DB62692A3BE92FF817A6B8C476" blockId="1.[113,752,1949,2068]" pageId="1" pageNumber="216">
<emphasis id="B924EAC962692A3BE92FF817A52DCBD7" bold="true" box="[113,268,1949,1973]" pageId="1" pageNumber="216">
<emphasis id="B924EAC962692A3BE92FF817A4D6CBD7" bold="true" box="[113,247,1949,1973]" italics="true" pageId="1" pageNumber="216">Description.</emphasis>
</emphasis>
Body not so depressed as in other species of the genus. Head dark red. Pronotum and elytra red. Abdomen almost dark red except three black terminal segments. Femora, tibiae and tarsi reddish. Antenna dark red.
</paragraph>
<paragraph id="8BEF36DB62692A3BEA72FF4BA18DCE5A" blockId="1.[812,1452,193,568]" pageId="1" pageNumber="216">
Head (
<figureCitation id="136B2A5E62692A3BEA2DFF4BA791CCBA" box="[883,944,193,216]" captionStart="Figs" captionStartId="2.[137,181,2035,2056]" captionTargetBox="[131,1468,262,1974]" captionTargetId="figure-87@2.[113,1593,220,2000]" captionTargetPageId="2" captionText="Figs. 1-11. Borolinus bisetifer, new species: 1, head and pronotum in dorsal view; 2, aedeagus in dorsal view; 3, aedeagus in lateral view; 4, 9th abdominal sternite of male; 5, left half of 9th abdominal sternite of female; 6, mandibles; 7, 9th and 10th abdominal tergites of male; 8, labrum; 9, mentum; 10, maxilla; 11, ligula and labial palpi. Scale bars: Figs. 1, 6 = 1 mm; Figs. 2-5, 7-10, 11 = 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.4508004" httpUri="https://zenodo.org/record/4508004/files/figure.png" pageId="1" pageNumber="216">Fig. 1</figureCitation>
) transverse, in front furnished with two parallel inner lateral teeth, which are distinctly projecting and pointed, almost reaching apical 2/3 of mandibles viewed from above; frontal depression between inner lateral teeth, deep, slightly longer than wide; outer lateral tooth situated at outer base of inner lateral tooth, triangularly pointed and broadly depressed at its base; lateral lobe situated between eye and outer lateral tooth, not gibbous. Vertex smooth, not sulcate, only in middle of base with an indistinct longitudinal fovea; along lateral and postero-lateral margin with some scattered punctures and long golden pubescence. Eyes glabrous, moderately convex. Clypeus steeply elongate, medially with one pair of long setae.
</paragraph>
<paragraph id="8BEF36DB62692A3BEA72FDEBA117C87A" blockId="1.[812,1452,609,1048]" pageId="1" pageNumber="216">
Labrum (
<figureCitation id="136B2A5E62692A3BEAC4FDEBA7C3CE1A" box="[922,994,609,632]" captionStart="Figs" captionStartId="2.[137,181,2035,2056]" captionTargetBox="[131,1468,262,1974]" captionTargetId="figure-87@2.[113,1593,220,2000]" captionTargetPageId="2" captionText="Figs. 1-11. Borolinus bisetifer, new species: 1, head and pronotum in dorsal view; 2, aedeagus in dorsal view; 3, aedeagus in lateral view; 4, 9th abdominal sternite of male; 5, left half of 9th abdominal sternite of female; 6, mandibles; 7, 9th and 10th abdominal tergites of male; 8, labrum; 9, mentum; 10, maxilla; 11, ligula and labial palpi. Scale bars: Figs. 1, 6 = 1 mm; Figs. 2-5, 7-10, 11 = 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.4508004" httpUri="https://zenodo.org/record/4508004/files/figure.png" pageId="1" pageNumber="216">Fig. 8</figureCitation>
) transverse, with two long and ciliate submembranous lobes projecting anteriorly, anterior margin furnished with 6 pairs of setae of various length, behind them in middle with two additional pairs of long setae. Mandibles extremely protruding (
<figureCitation id="136B2A5E62692A3BED1DFD6BA0B0CE9A" box="[1091,1169,737,760]" captionStart="Figs" captionStartId="2.[137,181,2035,2056]" captionTargetBox="[131,1468,262,1974]" captionTargetId="figure-87@2.[113,1593,220,2000]" captionTargetPageId="2" captionText="Figs. 1-11. Borolinus bisetifer, new species: 1, head and pronotum in dorsal view; 2, aedeagus in dorsal view; 3, aedeagus in lateral view; 4, 9th abdominal sternite of male; 5, left half of 9th abdominal sternite of female; 6, mandibles; 7, 9th and 10th abdominal tergites of male; 8, labrum; 9, mentum; 10, maxilla; 11, ligula and labial palpi. Scale bars: Figs. 1, 6 = 1 mm; Figs. 2-5, 7-10, 11 = 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.4508004" httpUri="https://zenodo.org/record/4508004/files/figure.png" pageId="1" pageNumber="216">Fig. 6</figureCitation>
), apical tooth pointed, immediately behind which with two sharp teeth on dorsal side and one sharp tooth on inner side. Inner lobe of maxillae (
<figureCitation id="136B2A5E62692A3BEA6AFCCBA7A1CF3A" box="[820,896,833,856]" captionStart="Figs" captionStartId="2.[137,181,2035,2056]" captionTargetBox="[131,1468,262,1974]" captionTargetId="figure-87@2.[113,1593,220,2000]" captionTargetPageId="2" captionText="Figs. 1-11. Borolinus bisetifer, new species: 1, head and pronotum in dorsal view; 2, aedeagus in dorsal view; 3, aedeagus in lateral view; 4, 9th abdominal sternite of male; 5, left half of 9th abdominal sternite of female; 6, mandibles; 7, 9th and 10th abdominal tergites of male; 8, labrum; 9, mentum; 10, maxilla; 11, ligula and labial palpi. Scale bars: Figs. 1, 6 = 1 mm; Figs. 2-5, 7-10, 11 = 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.4508004" httpUri="https://zenodo.org/record/4508004/files/figure.png" pageId="1" pageNumber="216">Fig. 10</figureCitation>
) with one slim and pointed tooth at apex, behind this tooth remaining inner side with dense ciliae in transverse rows; outer lobe hatchet-shaped, anterior half with dense ciliae in rows.
<collectingRegion id="4994F83962692A3BEAD4FC2BA7C5CFDA" box="[906,996,929,952]" country="Myanmar" name="Chin" pageId="1" pageNumber="216">Mentum</collectingRegion>
(
<figureCitation id="136B2A5E62692A3BEAADFC2BA012CFDA" box="[1011,1075,929,952]" captionStart="Figs" captionStartId="2.[137,181,2035,2056]" captionTargetBox="[131,1468,262,1974]" captionTargetId="figure-87@2.[113,1593,220,2000]" captionTargetPageId="2" captionText="Figs. 1-11. Borolinus bisetifer, new species: 1, head and pronotum in dorsal view; 2, aedeagus in dorsal view; 3, aedeagus in lateral view; 4, 9th abdominal sternite of male; 5, left half of 9th abdominal sternite of female; 6, mandibles; 7, 9th and 10th abdominal tergites of male; 8, labrum; 9, mentum; 10, maxilla; 11, ligula and labial palpi. Scale bars: Figs. 1, 6 = 1 mm; Figs. 2-5, 7-10, 11 = 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.4508004" httpUri="https://zenodo.org/record/4508004/files/figure.png" pageId="1" pageNumber="216">Fig. 9</figureCitation>
) trapeziform and setose, depressed laterally, in middle of front margin with a broad protruding denticle, apically ciliated. Paraglossa and glossa (
<figureCitation id="136B2A5E62692A3BEC0CFC6BA182CF9A" box="[1362,1443,993,1016]" captionStart="Figs" captionStartId="2.[137,181,2035,2056]" captionTargetBox="[131,1468,262,1974]" captionTargetId="figure-87@2.[113,1593,220,2000]" captionTargetPageId="2" captionText="Figs. 1-11. Borolinus bisetifer, new species: 1, head and pronotum in dorsal view; 2, aedeagus in dorsal view; 3, aedeagus in lateral view; 4, 9th abdominal sternite of male; 5, left half of 9th abdominal sternite of female; 6, mandibles; 7, 9th and 10th abdominal tergites of male; 8, labrum; 9, mentum; 10, maxilla; 11, ligula and labial palpi. Scale bars: Figs. 1, 6 = 1 mm; Figs. 2-5, 7-10, 11 = 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.4508004" httpUri="https://zenodo.org/record/4508004/files/figure.png" pageId="1" pageNumber="216">Fig. 11</figureCitation>
) ciliate anteriorly, not separated from each other.
</paragraph>
<paragraph id="8BEF36DB62692A3BEA72FBCBA799C93A" blockId="1.[812,1452,1089,1368]" pageId="1" pageNumber="216">
Antennae long, posteriorly reaching shoulder of elytra, weakly geniculate, 1
<superScript id="7C259B9362692A3BEAE7FBD5A7E2C80F" attach="left" box="[953,963,1119,1133]" fontSize="6" pageId="1" pageNumber="216">st</superScript>
segment baculiform, apically emarginate to hold 2
<superScript id="7C259B9362692A3BEA30FBF5A75CC8EF" attach="left" box="[878,893,1151,1165]" fontSize="6" pageId="1" pageNumber="216">nd</superScript>
segment, about as long as 2
<superScript id="7C259B9362692A3BEDC1FBF5A096C8FA" attach="none" box="[1183,1207,1151,1176]" fontSize="6" pageId="1" pageNumber="216">nd-</superScript>
4
<superScript id="7C259B9362692A3BED9BFBF5A0F0C8EF" attach="right" box="[1221,1233,1151,1165]" fontSize="6" pageId="1" pageNumber="216">th</superScript>
segments combined, 2
<superScript id="7C259B9362692A3BEA67FB15A769C8CF" attach="left" box="[825,840,1183,1197]" fontSize="6" pageId="1" pageNumber="216">nd</superScript>
smallest, almost quadrate, 3
<superScript id="7C259B9362692A3BED27FB15A0A7C8CF" attach="left" box="[1145,1158,1183,1197]" fontSize="6" pageId="1" pageNumber="216">rd</superScript>
long and narrowed at base, twice as long as 2
<superScript id="7C259B9362692A3BEA81FB35A7CFC8AF" attach="none" box="[991,1006,1215,1229]" fontSize="6" pageId="1" pageNumber="216">nd</superScript>
, 4
<superScript id="7C259B9362692A3BED59FB35A03BC8BA" attach="none" box="[1031,1050,1215,1240]" fontSize="6" pageId="1" pageNumber="216">th-</superScript>
6
<superScript id="7C259B9362692A3BED76FB35A015C8AF" attach="left" box="[1064,1076,1215,1229]" fontSize="6" pageId="1" pageNumber="216">th</superScript>
slightly elongate, 7
<superScript id="7C259B9362692A3BEDA2FB35A131C8BA" attach="none" box="[1276,1296,1215,1240]" fontSize="6" pageId="1" pageNumber="216">th-</superScript>
9
<superScript id="7C259B9362692A3BEC43FB35A108C8AF" attach="left" box="[1309,1321,1215,1229]" fontSize="6" pageId="1" pageNumber="216">th</superScript>
moniliform, 10
<superScript id="7C259B9362692A3BEA19FB55A772C88F" attach="left" box="[839,851,1247,1261]" fontSize="6" pageId="1" pageNumber="216">th</superScript>
slightly transverse, 11
<superScript id="7C259B9362692A3BED1CFB55A06FC88F" attach="left" box="[1090,1102,1247,1261]" fontSize="6" pageId="1" pageNumber="216">th</superScript>
conical, bluntly pointed, about 2 times as long as 10
<superScript id="7C259B9362692A3BEAAAFB75A021C96F" attach="none" box="[1012,1024,1279,1293]" fontSize="6" pageId="1" pageNumber="216">th</superScript>
. In addition to usual setae, segments 6
<superScript id="7C259B9362692A3BECC8FB75A18AC97A" attach="none" box="[1430,1451,1279,1304]" fontSize="6" pageId="1" pageNumber="216">th-</superScript>
11
<superScript id="7C259B9362692A3BEA19FA95A772C94F" attach="left" box="[839,851,1311,1325]" fontSize="6" pageId="1" pageNumber="216">th</superScript>
with tomentose pubscence, gradually becoming denser toward apex.
</paragraph>
<paragraph id="8BEF36DB62692A3BEA72FA0BA7CECABA" blockId="1.[812,1452,1409,1752]" pageId="1" pageNumber="216">Pronotum transverse, its frontal margin regularly emarginate, sides parallel, median longitudinal sulcus narrow, almost covering whole length of pronotum, disc of pronotum with finely vermiform microsculpture and on each side of sulcus with a distinct puncture; lateral line continuing, but turning upward at anterior angle, vanishing before reaching middle, basally bent dorsad at hind angle, forming curved ridge, continuing along posterior constriction and reaching basal line; sides of pronotum, along lateral line, with moderately dense setiferous punctures and golden setae, especially denser at posterior angle.</paragraph>
<paragraph id="8BEF36DB62692A3BEA72F88BA754CB3A" blockId="1.[812,1452,1793,1880]" pageId="1" pageNumber="216">Protibiae externally with 13-15 denticles, which gradually becoming shorter basally. Middle and hind tibiae densely setose.</paragraph>
<paragraph id="8BEF36DB62692A3BEA72F80BA163CB9A" blockId="1.[812,1451,1921,2040]" pageId="1" pageNumber="216">Elytra almost quadrate, disc with distinctly coriaceous micropunctures, slightly expanding laterally at posterior part, posterior margin weakly emarginate, small setiferous punctures moderately dense along lateral margin.</paragraph>
<caption id="DF2F6653626A2A38E9D7F879A111C420" ID-DOI="http://doi.org/10.5281/zenodo.4508004" ID-Zenodo-Dep="4508004" httpUri="https://zenodo.org/record/4508004/files/figure.png" pageId="2" pageNumber="217" startId="2.[137,181,2035,2056]" targetBox="[131,1468,262,1974]" targetPageId="2">
<paragraph id="8BEF36DB626A2A38E9D7F879A111C420" blockId="2.[137,1475,2035,2114]" pageId="2" pageNumber="217">
Figs. 1-11.
<taxonomicName id="4C504D58626A2A38E9A6F87EA58DC46B" authority="Wu &amp; Zhou, 2006" authorityName="Wu &amp; Zhou" authorityYear="2006" box="[248,428,2036,2057]" class="Insecta" family="Staphylinidae" genus="Borolinus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="217" phylum="Arthropoda" rank="species" species="bisetifer" status="sp. nov.">
<emphasis id="B924EAC9626A2A38E9A6F87EA58DC46B" box="[248,428,2036,2057]" italics="true" pageId="2" pageNumber="217">Borolinus bisetifer</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A21757B2626A2A38E8E7F879A610C46A" box="[441,561,2035,2056]" pageId="2" pageNumber="217" rank="species">new species</taxonomicNameLabel>
: 1, head and pronotum in dorsal view; 2, aedeagus in dorsal view; 3, aedeagus in lateral view; 4, 9
<superScript id="7C259B93626A2A38E9F0F785A498C479" attach="left" box="[174,185,2063,2075]" fontSize="5" pageId="2" pageNumber="217">th</superScript>
abdominal sternite of male; 5, left half of 9
<superScript id="7C259B93626A2A38EB01F785A64BC479" attach="left" box="[607,618,2063,2075]" fontSize="5" pageId="2" pageNumber="217">th</superScript>
abdominal sternite of female; 6, mandibles; 7, 9
<superScript id="7C259B93626A2A38ED61F785A06BC479" attach="left" box="[1087,1098,2063,2075]" fontSize="5" pageId="2" pageNumber="217">th</superScript>
and 10
<superScript id="7C259B93626A2A38EDCFF785A0BDC479" attach="left" box="[1169,1180,2063,2075]" fontSize="5" pageId="2" pageNumber="217">th</superScript>
abdominal tergites of male; 8, labrum; 9, mentum; 10, maxilla; 11, ligula and labial palpi. Scale bars: Figs. 1, 6 = 1 mm; Figs. 2-5, 7-10, 11 = 0.25 mm.
</paragraph>
</caption>
<paragraph id="8BEF36DB626B2A39E92FFF4BA49DCD1A" blockId="3.[113,752,191,376]" pageId="3" pageNumber="218">
Abdomen cylindrical, weakly broadened to 7
<superScript id="7C259B93626B2A39EB0DFF35A67ECCAF" attach="left" box="[595,607,191,205]" fontSize="6" pageId="3" pageNumber="218">th</superScript>
segment, 3
<superScript id="7C259B93626B2A39EB84FF35A6D1CCBA" attach="none" box="[730,752,191,216]" fontSize="6" pageId="3" pageNumber="218">rd-</superScript>
6
<superScript id="7C259B93626B2A39E920FF55A4ABCC8F" attach="left" box="[126,138,223,237]" fontSize="6" pageId="3" pageNumber="218">th</superScript>
segments with anterior and posterior row of long golden setae, these setae denser at lateral side, but gradually becoming sparser medially, 7
<superScript id="7C259B93626B2A39E8C1FE95A58ACD4F" attach="left" box="[415,427,287,301]" fontSize="6" pageId="3" pageNumber="218">th</superScript>
and 8
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irregularly covered with long golden setae, but in middle of 7
<superScript id="7C259B93626B2A39E8A8FEB5A623CD2F" attach="left" box="[502,514,319,333]" fontSize="6" pageId="3" pageNumber="218">th</superScript>
with a broad glabrous region.
</paragraph>
<paragraph id="8BEF36DB626B2A39E92FFE2BA4E3CF3A" blockId="3.[113,752,417,856]" pageId="3" pageNumber="218">
<emphasis id="B924EAC9626B2A39E92FFE2BA48ECDDA" box="[113,175,417,440]" italics="true" pageId="3" pageNumber="218">Male.</emphasis>
Aedeagus (
<figureCitation id="136B2A5E626B2A39E877FE2BA55ECDDA" box="[297,383,417,440]" captionStart="Figs" captionStartId="2.[137,181,2035,2056]" captionTargetBox="[131,1468,262,1974]" captionTargetId="figure-87@2.[113,1593,220,2000]" captionTargetPageId="2" captionText="Figs. 1-11. Borolinus bisetifer, new species: 1, head and pronotum in dorsal view; 2, aedeagus in dorsal view; 3, aedeagus in lateral view; 4, 9th abdominal sternite of male; 5, left half of 9th abdominal sternite of female; 6, mandibles; 7, 9th and 10th abdominal tergites of male; 8, labrum; 9, mentum; 10, maxilla; 11, ligula and labial palpi. Scale bars: Figs. 1, 6 = 1 mm; Figs. 2-5, 7-10, 11 = 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.4508004" httpUri="https://zenodo.org/record/4508004/files/figure.png" pageId="3" pageNumber="218">Fig. 2, 3</figureCitation>
) with median lobe slightly bulbous at base, curved behind basal orifice, posterior part baculiform, rounded at apex and weakly sclerotized on dorsal side; basal orifice roundly emarginate above, parameres short, thin and pointed, with basal part protruding dorsally and connecting each other to form a characteristic triangular structure, apically emarginate. Ninth tergite (
<figureCitation id="136B2A5E626B2A39E8CDFDEBA5F4CE1A" box="[403,469,609,632]" captionStart="Figs" captionStartId="2.[137,181,2035,2056]" captionTargetBox="[131,1468,262,1974]" captionTargetId="figure-87@2.[113,1593,220,2000]" captionTargetPageId="2" captionText="Figs. 1-11. Borolinus bisetifer, new species: 1, head and pronotum in dorsal view; 2, aedeagus in dorsal view; 3, aedeagus in lateral view; 4, 9th abdominal sternite of male; 5, left half of 9th abdominal sternite of female; 6, mandibles; 7, 9th and 10th abdominal tergites of male; 8, labrum; 9, mentum; 10, maxilla; 11, ligula and labial palpi. Scale bars: Figs. 1, 6 = 1 mm; Figs. 2-5, 7-10, 11 = 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.4508004" httpUri="https://zenodo.org/record/4508004/files/figure.png" pageId="3" pageNumber="218">Fig. 7</figureCitation>
) sclerotized with left and right plates separated, each with 8 or 9 setae of various length. Tenth tergite almost membraneous except for sclerotized apical part, posterior margin emarginate in middle, with 3 pairs of setae. Ninth sternite (
<figureCitation id="136B2A5E626B2A39E8FBFD6BA5C2CE9A" box="[421,483,737,760]" captionStart="Figs" captionStartId="2.[137,181,2035,2056]" captionTargetBox="[131,1468,262,1974]" captionTargetId="figure-87@2.[113,1593,220,2000]" captionTargetPageId="2" captionText="Figs. 1-11. Borolinus bisetifer, new species: 1, head and pronotum in dorsal view; 2, aedeagus in dorsal view; 3, aedeagus in lateral view; 4, 9th abdominal sternite of male; 5, left half of 9th abdominal sternite of female; 6, mandibles; 7, 9th and 10th abdominal tergites of male; 8, labrum; 9, mentum; 10, maxilla; 11, ligula and labial palpi. Scale bars: Figs. 1, 6 = 1 mm; Figs. 2-5, 7-10, 11 = 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.4508004" httpUri="https://zenodo.org/record/4508004/files/figure.png" pageId="3" pageNumber="218">Fig. 4</figureCitation>
) composed of two plates, apical plate sclerotized except for membraneous apex, with two long setae, basal plate less sclerotized, broadened in middle.
</paragraph>
<paragraph id="8BEF36DB626B2A39E92FFC0BA5A5CF9A" blockId="3.[113,752,897,1016]" pageId="3" pageNumber="218">
<emphasis id="B924EAC9626B2A39E92FFC0BA4EFCFFA" box="[113,206,897,920]" italics="true" pageId="3" pageNumber="218">Female.</emphasis>
Ninth sternite (
<figureCitation id="136B2A5E626B2A39E8D1FC0BA5F9CFFA" box="[399,472,897,920]" captionStart="Figs" captionStartId="2.[137,181,2035,2056]" captionTargetBox="[131,1468,262,1974]" captionTargetId="figure-87@2.[113,1593,220,2000]" captionTargetPageId="2" captionText="Figs. 1-11. Borolinus bisetifer, new species: 1, head and pronotum in dorsal view; 2, aedeagus in dorsal view; 3, aedeagus in lateral view; 4, 9th abdominal sternite of male; 5, left half of 9th abdominal sternite of female; 6, mandibles; 7, 9th and 10th abdominal tergites of male; 8, labrum; 9, mentum; 10, maxilla; 11, ligula and labial palpi. Scale bars: Figs. 1, 6 = 1 mm; Figs. 2-5, 7-10, 11 = 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.4508004" httpUri="https://zenodo.org/record/4508004/files/figure.png" pageId="3" pageNumber="218">Fig. 5</figureCitation>
) with hemisternite less sclerotized, anterior margin not strongly rounded, second gonocoxite strongly sclerotized except for membraneous apex, with one stout seta.
</paragraph>
<paragraph id="8BEF36DB626B2A39E92FFBABA4F9C81A" blockId="3.[113,752,1056,1144]" pageId="3" pageNumber="218">
Measurements.
<emphasis id="B924EAC9626B2A39E84AFBAAA500C85A" bold="true" box="[276,289,1056,1080]" pageId="3" pageNumber="218"></emphasis>
Body length:
<quantity id="4CA89B3E626B2A39E8EFFBABA63FC85A" box="[433,542,1057,1080]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" metricValueMax="1.6" metricValueMin="1.4" pageId="3" pageNumber="218" unit="mm" value="15.0" valueMax="16.0" valueMin="14.0">14-16 mm</quantity>
; HL:
<quantity id="4CA89B3E626B2A39EB08FBABA694C85A" box="[598,693,1057,1080]" metricMagnitude="-4" metricUnit="m" metricValue="7.8" pageId="3" pageNumber="218" unit="mm" value="0.78">0.78 mm</quantity>
; HW:
<quantity id="4CA89B3E626B2A39E92FFBCBA4F5C83A" box="[113,212,1089,1112]" metricMagnitude="-3" metricUnit="m" metricValue="1.62" pageId="3" pageNumber="218" unit="mm" value="1.62">1.62 mm</quantity>
; PL:
<quantity id="4CA89B3E626B2A39E850FBCBA550C83A" box="[270,369,1089,1112]" metricMagnitude="-3" metricUnit="m" metricValue="1.92" pageId="3" pageNumber="218" unit="mm" value="1.92">1.92 mm</quantity>
; PW:
<quantity id="4CA89B3E626B2A39E8EDFBCBA637C83A" box="[435,534,1089,1112]" metricMagnitude="-3" metricUnit="m" metricValue="2.46" pageId="3" pageNumber="218" unit="mm" value="2.46">2.46 mm</quantity>
; EL:
<quantity id="4CA89B3E626B2A39EB0FFBCBA695C83A" box="[593,692,1089,1112]" metricMagnitude="-3" metricUnit="m" metricValue="2.34" pageId="3" pageNumber="218" unit="mm" value="2.34">2.34 mm</quantity>
; EW:
<quantity id="4CA89B3E626B2A39E92FFBEBA4F5C81A" box="[113,212,1121,1144]" metricMagnitude="-3" metricUnit="m" metricValue="2.52" pageId="3" pageNumber="218" unit="mm" value="2.52">2.52 mm</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="C34A6550626B2A39E92FFB2BA573C89A" pageId="3" pageNumber="218" type="etymology">
<paragraph id="8BEF36DB626B2A39E92FFB2BA573C89A" blockId="3.[113,752,1184,1272]" pageId="3" pageNumber="218">
<emphasis id="B924EAC9626B2A39E92FFB2BA4DEC8DA" bold="true" box="[113,255,1184,1209]" pageId="3" pageNumber="218">
<emphasis id="B924EAC9626B2A39E92FFB2BA4CDC8DB" bold="true" box="[113,236,1185,1209]" italics="true" pageId="3" pageNumber="218">Etymology.</emphasis>
</emphasis>
The species name is a combination of the Latin prefix “
<emphasis id="B924EAC9626B2A39E99DFB4BA4C0C8BA" box="[195,225,1217,1240]" italics="true" pageId="3" pageNumber="218">bi-</emphasis>
” and the adjective “
<emphasis id="B924EAC9626B2A39E8E7FB4BA5DDC8BA" box="[441,508,1217,1240]" italics="true" pageId="3" pageNumber="218">setifer</emphasis>
” to indicate the pair of setae on the clypeus.
</paragraph>
</subSubSection>
<subSubSection id="C34A6550626B2A39E92FFAABA49ACA7A" pageId="3" pageNumber="218" type="discussion">
<paragraph id="8BEF36DB626B2A39E92FFAABA49ACA7A" blockId="3.[113,752,1312,1560]" pageId="3" pageNumber="218">
<emphasis id="B924EAC9626B2A39E92FFAABA4CBC95A" bold="true" box="[113,234,1312,1337]" pageId="3" pageNumber="218">
<emphasis id="B924EAC9626B2A39E92FFAABA4F9C95B" bold="true" box="[113,216,1313,1337]" italics="true" pageId="3" pageNumber="218">Remarks.</emphasis>
</emphasis>
This new species can be easily distinguished from
<taxonomicName id="4C504D58626B2A39E92FFACBA4D8C93A" box="[113,249,1345,1368]" class="Insecta" family="Staphylinidae" genus="Borolinus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="218" phylum="Arthropoda" rank="species" species="semirufus">
<emphasis id="B924EAC9626B2A39E92FFACBA4D8C93A" box="[113,249,1345,1368]" italics="true" pageId="3" pageNumber="218">B. semirufus</emphasis>
</taxonomicName>
by having one pair of outer lateral teeth, and from other Chinese species by the reddish coloration and the clypeus having only one pair of long setae. This species is similar to
<taxonomicName id="4C504D58626B2A39E982FA2BA5C4C9DA" authority="Fauvel, 1895" authorityName="Fauvel" authorityYear="1895" box="[220,485,1441,1464]" class="Insecta" family="Staphylinidae" genus="Borolinus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="218" phylum="Arthropoda" rank="species" species="antilope">
<emphasis id="B924EAC9626B2A39E982FA2BA571C9DA" box="[220,336,1441,1464]" italics="true" pageId="3" pageNumber="218">B. antilope</emphasis>
<bibRefCitation id="EFC14B2A626B2A39E809FA2BA5C4C9DA" author="Fauvel, A" box="[343,485,1441,1464]" pageId="3" pageNumber="218" pagination="180 - 286" refId="ref4846" refString="Fauvel, A., 1895. Staphylinides nouveaux de l'Inde et de la Malaisie. Revue d'Entomologie, 14: 180 - 286." type="journal article" year="1895">Fauvel, 1895</bibRefCitation>
</taxonomicName>
, but can be distinguished by the more reddish head and abdomen, and by the pronotum having two distinct punctures on each side of the median sulcus.
</paragraph>
</subSubSection>
<subSubSection id="C34A6550626B2A39E92FF9CBA5F1CA3A" box="[113,464,1601,1625]" pageId="3" pageNumber="218" type="distribution">
<paragraph id="8BEF36DB626B2A39E92FF9CBA5F1CA3A" blockId="3.[113,464,1601,1625]" box="[113,464,1601,1625]" pageId="3" pageNumber="218">
<emphasis id="B924EAC9626B2A39E92FF9CBA533CA3A" box="[113,274,1601,1625]" italics="true" pageId="3" pageNumber="218">
<emphasis id="B924EAC9626B2A39E92FF9CBA4DCCA3B" bold="true" box="[113,253,1601,1625]" italics="true" pageId="3" pageNumber="218">Distribution.</emphasis>
</emphasis>
<collectingCountry id="F347764B626B2A39E844F9CBA57BCA3A" box="[282,346,1601,1624]" name="China" pageId="3" pageNumber="218">China</collectingCountry>
(
<collectingRegion id="4994F839626B2A39E834F9CBA5E5CA3A" box="[362,452,1601,1624]" country="China" name="Yunnan" pageId="3" pageNumber="218">Yunnan</collectingRegion>
).
</paragraph>
</subSubSection>
</treatment>
</document>