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<document id="548D10BF8D3504BC8002459A34A32424" ID-CLB-Dataset="300009" ID-DOI="10.32225/bkpmnh.2021.50_55" ID-GBIF-Dataset="70224806-ca38-47ca-bd8a-63da3ac532f9" ID-ISSN="2189-6720" ID-Zenodo-Dep="12774639" IM.bibliography_approvedBy="juliana" IM.illustrations_approvedBy="juliana" IM.materialsCitations_approvedBy="juliana" IM.metadata_approvedBy="juliana" IM.taxonomicNames_approvedBy="juliana" IM.treatmentCitations_approvedBy="juliana" IM.treatments_approvedBy="juliana" checkinTime="1721260925551" checkinUser="felipe" docAuthor="Watanabe, Kyohei" docDate="2021" docId="03F987931B51A702FE8C62BCFEB3770E" docLanguage="en" docName="BullKanagawaPrefectMus.50.55-136.pdf" docOrigin="Bulletin of the Kanagawa Prefectural Museum (Natural Science) 50" docSource="https://doi.org/10.32225/bkpmnh.2021.50_55" docStyle="DocumentStyle:020B98DC1285EBBD02620724ECB44EDE.1:BullKanagawaPrefectMus.2013-.journal_article.english" docStyleId="020B98DC1285EBBD02620724ECB44EDE" docStyleName="BullKanagawaPrefectMus.2013-.journal_article.english" docStyleVersion="1" docTitle="Surculus japonicus Watanabe, 2021, sp. nov." docType="treatment" docUuid="3F7E36B5-FA84-4824-AC61-B1E9E1FAC0C7" docUuidSource="ZooBank" docVersion="9" lastPageNumber="77" masterDocId="FFC0FFEB1B45A714FFCA6633FFD77447" masterDocTitle="Taxonomic and Zoogeographic Study of the Japanese Phygadeuontinae (Hymenoptera, Ichneumonidae), with Descriptions of 17 New Species" masterLastPageNumber="136" masterPageNumber="55" pageNumber="75" updateTime="1721329445643" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
<mods:mods id="5C229020A4C76884D6067B3A95708FC0" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="6330FF5C9A36F800103EC1781503236E">
<mods:title id="585DE2852B6955ED73D474AB7EB8B35F">Taxonomic and Zoogeographic Study of the Japanese Phygadeuontinae (Hymenoptera, Ichneumonidae), with Descriptions of 17 New Species</mods:title>
</mods:titleInfo>
<mods:name id="2EE30BE36C81A481BBD16977E1F9142D" type="personal">
<mods:role id="54DC68EFBC98E6DDCBEDA3653DD2AD38">
<mods:roleTerm id="5FE290DEF328403EC8D4B78C56ECA4AA">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="77DCFBF255BC2B9479E2BC0850CE6292">Watanabe, Kyohei</mods:namePart>
<mods:affiliation id="621CB80369DD95A69698266B2B74ABBA">Kanagawa Prefectural Museum of Natural History, 499 Iryuda, Odawara, Kanagawa 250 – 0031, Japan</mods:affiliation>
<mods:nameIdentifier id="7126C41283803AFAE848648296804027" type="email">watanabe-k@nh‛kanagawa-museum‛jp</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="07D6F9BD0450CA13F6C44BFCA3B0BC6B">text</mods:typeOfResource>
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<mods:title id="9E3DA9AF8206E31390B93A2BA9D510D5">Bulletin of the Kanagawa Prefectural Museum (Natural Science)</mods:title>
</mods:titleInfo>
<mods:part id="E1B69C79F7D98EA412B185823E0B78C5">
<mods:date id="32B7831E4E028E9F325D8F406528C82D">2021</mods:date>
<mods:detail id="4CD135E14B3F361AEA4A872D79851A19" type="pubDate">
<mods:number id="949D095E9A3532F177F54D237B0B1293">2021-03-30</mods:number>
</mods:detail>
<mods:detail id="E23F68CC3DF41C1211ABC3B241E54E6A" type="volume">
<mods:number id="D832694B46A50F357D69CB4991C7B884">50</mods:number>
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<mods:location id="3EFF8F2154B47503993A89E5C74054D5">
<mods:url id="AF2856CB8704686DFCA2300E3D951E09">https://doi.org/10.32225/bkpmnh.2021.50_55</mods:url>
</mods:location>
<mods:classification id="3C2E4D5DCEEB591FD4A5D47E7EE5E9BA">journal article</mods:classification>
<mods:identifier id="22CFD18619FA9296263CA4BB5395B8E5" type="CLB-Dataset">300009</mods:identifier>
<mods:identifier id="3AAD3986FBDFEFC2DABF55127C537F11" type="DOI">10.32225/bkpmnh.2021.50_55</mods:identifier>
<mods:identifier id="A88729D71B06786A42EB8089BFE64871" type="GBIF-Dataset">70224806-ca38-47ca-bd8a-63da3ac532f9</mods:identifier>
<mods:identifier id="854763DA8A124EA458C79CA51D8E6C73" type="ISSN">2189-6720</mods:identifier>
<mods:identifier id="058F0C7DA7ACADE4242E46A1A28ADCE6" type="Zenodo-Dep">12774639</mods:identifier>
<mods:identifier id="BFA2B01F1A4AA47C9C0208F4E183BD2A" type="ZooBank">DD695C62-DC2F-4D32-8D33-52648E28661D</mods:identifier>
</mods:mods>
<treatment id="03F987931B51A702FE8C62BCFEB3770E" ID-DOI="http://doi.org/10.5281/zenodo.12773486" ID-GBIF-Taxon="235610675" ID-Zenodo-Dep="12773486" LSID="urn:lsid:zoobank.org:act:3F7E36B5-FA84-4824-AC61-B1E9E1FAC0C7" httpUri="http://treatment.plazi.org/id/03F987931B51A702FE8C62BCFEB3770E" lastPageId="22" lastPageNumber="77" pageId="20" pageNumber="75">
<subSubSection id="C34A650E1B51A700FE8C62BCFDAA70EF" box="[326,637,1167,1192]" pageId="20" pageNumber="75" type="nomenclature">
<paragraph id="8BEF36851B51A700FE8C62BCFDAA70EF" blockId="20.[240,723,1167,1269]" box="[326,637,1167,1192]" pageId="20" pageNumber="75">
<heading id="D0A781E91B51A700FE8C62BCFDAA70EF" box="[326,637,1167,1192]" centered="true" fontSize="10" level="2" pageId="20" pageNumber="75" reason="6">
<emphasis id="B924EA971B51A700FE8C62BCFDAA70EF" bold="true" box="[326,637,1167,1192]" pageId="20" pageNumber="75">
<taxonomicName id="4C504D061B51A700FE8C62BCFDC870E0" authorityName="Watanabe" authorityYear="2021" box="[326,543,1167,1191]" class="Insecta" family="Ichneumonidae" genus="Surculus" kingdom="Animalia" order="Hymenoptera" pageId="20" pageNumber="75" phylum="Arthropoda" rank="species" species="japonicus" status="sp. nov.">
<emphasis id="B924EA971B51A700FE8C62BCFDC870E0" bold="true" box="[326,543,1167,1191]" italics="true" pageId="20" pageNumber="75">Surculus japonicus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21757EC1B51A700FDEF62A3FDAA70EF" box="[549,637,1168,1192]" pageId="20" pageNumber="75" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C34A650E1B51A700FF3A6284FD047089" box="[240,723,1206,1231]" pageId="20" pageNumber="75" type="vernacular_names">
<paragraph id="8BEF36851B51A700FF3A6284FD047089" blockId="20.[240,723,1167,1269]" box="[240,723,1206,1231]" pageId="20" pageNumber="75">(New SJN: Hosomi-mame-togari-himebachi)</paragraph>
</subSubSection>
<subSubSection id="C34A650E1B51A700FECD62EEFD6A70B2" box="[263,701,1245,1269]" pageId="20" pageNumber="75" type="description">
<paragraph id="8BEF36851B51A700FECD62EEFD6A70B2" blockId="20.[240,723,1167,1269]" box="[263,701,1245,1269]" pageId="20" pageNumber="75">
(
<figureCitation id="136B2A001B51A700FEC562EEFE4170B2" box="[271,406,1245,1269]" captionStart="Fig" captionStartId="21.[153,186,1985,2007]" captionTargetBox="[151,1398,998,1961]" captionTargetId="figure-368@21.[151,1398,996,1961]" captionTargetPageId="21" captionText="Fig. 18. Surculus japonicus sp. nov., KPM-NK 81861, holotype, female ― A, lateral habitus; B, head, frontal view; C, head, mesosoma, and metasoma, dorsal view; D, head and mesosoma, lateral view; E, head, mesoscutum and scutellum, dorsal view; F, wings; G, propodeum,dorsal view." figureDoi="http://doi.org/10.5281/zenodo.12774677" httpUri="https://zenodo.org/record/12774677/files/figure.png" pageId="20" pageNumber="75">Figs 18A–G</figureCitation>
,
<figureCitation id="136B2A001B51A700FE6B62EEFE0670B2" box="[417,465,1245,1269]" captionStart="Fig" captionStartId="72.[190,223,1065,1087]" captionTargetBox="[195,1428,161,998]" captionTargetId="figure-227@72.[189,1436,149,1010]" captionTargetPageId="72" captionText="Fig. 62. Bathythrix kuwanae Viereck, 1912 (A, KPM-NK 81718), Ba. margaretae Sawoniewicz, 1980 (B, KPM-NK 55017), Ba. thomsoni (Kerrich, 1942) (C, KPM-NK 81675), Diaglyptidea conformis (Gmelin, 1790) (D, KPM-NK 81645), Nipponaetes haeussleri (Uchida, 1933) (E, KPM-NK 81832), Acrolyta japonica sp. nov. (F, N, KPM-NK 54996, holotype), Acr. rufocincta (Gravenhorst, 1829) (G, KPM-NK 81732), Surculus japonicus sp. nov. (H, KPM-NK 81861, holotype), Isadelphus nigrus sp. nov. (I, KPM-NK 82003, holotype), Lochetica japonica sp. nov. (J, KPM-NK 81989, holotype), Megacara similis Sheng, 1999 (K, KPM-NK 81837), Gnotus chionops (Gravenhorst, 1829) (L, KPM-NK 82100), Gn. striatus (Uchida, 1930) (M, KPM-NK 82060), and Acr. flavicoxis Sheng &amp; Sun, 2014 (O, KPM-NK 81837), females from Japan ― A–C, head, frontal view; D, E, head, lateral view; F–K, anterior margin of clypeus; L, M, mandible; N, O, basal part of antenna, lateral view." figureDoi="http://doi.org/10.5281/zenodo.12774781" httpUri="https://zenodo.org/record/12774781/files/figure.png" pageId="20" pageNumber="75">62H</figureCitation>
,
<figureCitation id="136B2A001B51A700FE1662EEFDD170B2" box="[476,518,1245,1269]" captionStart="Fig" captionStartId="73.[151,185,989,1011]" captionTargetBox="[168,1379,161,949]" captionTargetId="figure-5@73.[151,1398,151,958]" captionTargetPageId="73" captionText="Fig. 63. Acrolyta japonica sp. nov. (A, KPM-NK 54996, holotype), Dichrogaster liostylus (Thomson, 1885) (B, KPM-NK 55074), Dic. nitida Sheng &amp; Sun, 2014 (C, KPM-NK 81908), Bathythrix kuwanae Viereck, 1912 (D, KPM-NK 81718), Retalia japonica Kusigemati, 1985 (E, KPM-NK 81676), Surculus japonicus sp. nov. (F, KPM-NK 81861, holotype), Bentyra ryukyuana sp. nov. (G, KPM-NK 81887, holotype), Hemiteles japonicus sp. nov. (H, KPM-NK 55021, holotype), and H. maculipterus sp. nov. (I, KPM-NK 55025, holotype), females from Japan ― A, median part of collar, dorsal view; B, C, lateral part of pronotum; D–G, mesoscutum and scutellum,dorsal view; H, I, hind tarsus, lateral view." figureDoi="http://doi.org/10.5281/zenodo.12774783" httpUri="https://zenodo.org/record/12774783/files/figure.png" pageId="20" pageNumber="75">63F</figureCitation>
,
<figureCitation id="136B2A001B51A700FDDB62EEFD9570B2" box="[529,578,1245,1269]" captionStart="Fig" captionStartId="73.[151,185,1926,1948]" captionTargetBox="[162,1384,1211,1880]" captionTargetId="figure-109@73.[151,1398,1199,1899]" captionTargetPageId="73" captionText="Fig. 64. Surculus japonicus sp. nov. (A, KPM-NK 81861, holotype), Paraphylax elegans sp. nov. (B, KPM-NK 55078, holotype), Hemiteles japonicus sp. nov. (C, KPM-NK 55021, holotype), H. maculipterus sp. nov. (D, KPM-NK 55026, paratype), Charitopes gastricus (Holmgren, 1868) (E, KPM-NK 81963), Tropistes shimizui sp. nov. (F, KPM-NK 81833, holotype), Uchidella okamotoi (Uchida, 1936) (G, KPM-NK 82073), and Uchidella toichii sp. nov. (H, KPM-NK 82011, holotype), females from Japan ― A–H, propodeum,dorsal view." figureDoi="http://doi.org/10.5281/zenodo.12774785" httpUri="https://zenodo.org/record/12774785/files/figure.png" pageId="20" pageNumber="75">64A</figureCitation>
,
<figureCitation id="136B2A001B51A700FD8662EEFDAB70B2" box="[588,636,1245,1269]" captionStart="Fig" captionStartId="74.[191,224,1111,1133]" captionTargetBox="[193,1432,157,1080]" captionTargetId="figure-134@74.[189,1438,151,1085]" captionTargetPageId="74" captionText="Fig.65.Acrolyta japonica sp. nov. (A, KPM-NK 54996, holotype), Diaglyptidea conformis (Gmelin,1790) (B, KPM-NK 81650), Micraris ryukyuensis sp. nov. (C, KPM-NK 81873, holotype), Bathythrix kuwanae Viereck, 1912 (D, KPM-NK 81718), Bentyra ryukyuana sp. nov. (E, KPM-NK 81887, holotype), Chirotica matsukemushii (Matsumura, 1926) (F, KPM-NK 81906), Paraphylax politus sp. nov. (G, KPM-NK 55079, holotype), Pa. transstriatus sp. nov. (H, KPM-NK 55080, holotype), Pa. yakushimensis sp. nov. (I, KPM-NK 55081, holotype), Pa. yambarensis sp. nov. (J, KPM-NK 55085, holotype), Ethelurgus sodalis kumatai Kusigemati, 1983 (K, KPM- NK 81938), Aclastus etorofuensis (Uchida, 1936) (L, KPM-NK 81976), Hemiteles kuro sp. nov. (M, KPM-NK 55024, holotype), H. yamatonis sp. nov. (O, KPM-NK 55033, paratype), Isadelphus nigrus sp. nov. (P, KPM-NK 82003, holotype), Lochetica japonica sp. nov. (Q, KPM-NK 81989, holotype), Mastrus molestae (Uchida, 1933) (R, KPM-NK 81987), Ma.takadai Momoi, 1970 (R, KPM-NK 81979), Gnotus chionops (Gravenhorst, 1829) (L, KPM-NK 82104), Gn. striatus (Uchida, 1930) (M, KPM-NK 82061), Orthizema semanotae Sheng &amp; Sun, 2014 (V, KPM-NK 82047), Nipponaetes haeussleri (Uchida, 1933) (W, KPM-NK 81832), and Surculus japonicus sp. nov. (X, KPM-NK 81861, holotype), females from Japan ― A-W, carinae around area superomedia of propodeum, dorsal view; X, T I, lateral view." figureDoi="http://doi.org/10.5281/zenodo.12774787" httpUri="https://zenodo.org/record/12774787/files/figure.png" pageId="20" pageNumber="75">65X</figureCitation>
,
<figureCitation id="136B2A001B51A700FD4D62EEFD6270B2" box="[647,693,1245,1269]" captionStart="Fig" captionStartId="75.[151,184,1580,1602]" captionTargetBox="[161,1381,163,1549]" captionTargetId="figure-5@75.[151,1398,151,1553]" captionTargetPageId="75" captionText="Fig. 66. Acrolyta japonica sp. nov. (A, KPM-NK 54996, holotype), Micraris ryukyuensis sp. nov. (B, KPM-NK 81873, holotype), Surculus japonicus sp. nov. (C, KPM-NK 81861, holotype), Bentyra ryukyuana sp. nov. (D, KPM-NK 81887, holotype), Paraphylax elegans sp. nov. (E, KPM-NK 55078, holotype), Pa. politus sp. nov. (F, KPM-NK 55079, holotype), Pa. transstriatus sp. nov. (G, KPM-NK 55080, holotype), Pa.yakushimensis sp. nov. (H, KPM-NK 55081, holotype), Pa. yambarensis sp. nov. (I, KPM-NK 55085, holotype), Hemiteles japonicus sp. nov. (J, KPM-NK 55021, holotype),H. kuro sp. nov. (K, KPM-NK 55024, holotype), H.maculipterus sp. nov. (L, KPM-NK 55025, holotype), H. yamatonis sp. nov. (M, KPM-NK 55032, holotype), Brachypimpla latipetiolator (Uchida, 1935) (N, O, KPM-NK 81973), Isadelphus nigrus sp. nov. (P, KPM-NK 82003, holotype), Lochetica japonica sp. nov. (Q, KPM-NK 81989, holotype), Mastrus molestae (Uchida, 1933) (R, KPM-NK 81987), Megacara similis Sheng, 1999 (S, KPM-NK 81835), Tropistes shimizui sp. nov. (T, KPM-NK 81833, holotype), and Uchidella toichii sp. nov. (U, KPM-NK 82011, holotype), females from Japan ― A-N, P-U, apical part of ovipositor, lateral view (A, C-G, I-K, N, Q-U, left side; B, H, L, M, P, right side); O, apex of ovipositor sheath, lateral view." figureDoi="http://doi.org/10.5281/zenodo.12774789" httpUri="https://zenodo.org/record/12774789/files/figure.png" pageId="20" pageNumber="75">66C</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C34A650E1B51A700FF1F6318FD4B7104" box="[213,668,1323,1347]" pageId="20" pageNumber="75" type="etymology">
<paragraph id="8BEF36851B51A700FF1F6318FD4B7104" blockId="20.[189,775,1323,1503]" box="[213,668,1323,1347]" pageId="20" pageNumber="75">
<emphasis id="B924EA971B51A700FF1F6318FE9E7104" bold="true" box="[213,329,1323,1347]" pageId="20" pageNumber="75">Etymology</emphasis>
. The specific name is from
<collectingCountry id="F34776151B51A700FD96631FFD4F7104" box="[604,664,1324,1347]" name="Japan" pageId="20" pageNumber="75">Japan</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C34A650E1B51A700FF1F6361FBB576CF" pageId="20" pageNumber="75" type="materials_examined">
<paragraph id="8BEF36851B51A700FF1F6361FBB576CF" blockId="20.[189,775,1323,1503]" lastBlockId="20.[850,1437,157,1894]" pageId="20" pageNumber="75">
<emphasis id="B924EA971B51A700FF1F6361FE85712D" bold="true" box="[213,338,1362,1386]" pageId="20" pageNumber="75">Type series</emphasis>
.
<materialsCitation id="3B383CD81B51A700FEAB6361FE3E71FF" ID-GBIF-Occurrence="4906826634" collectingDate="2007-08-04" collectorName="K. Watanabe" country="Japan" location="Hakkaisan" municipality="Mt. Ontakesan" pageId="20" pageNumber="75" specimenCode="KPM-NK 81861" specimenCount="1" stateProvince="Nagano" typeStatus="holotype">
<emphasis id="B924EA971B51A700FEAB6361FE11712D" bold="true" box="[353,454,1362,1386]" pageId="20" pageNumber="75">
<typeStatus id="54EB88271B51A700FEAB6361FE11712D" box="[353,454,1362,1386]" pageId="20" pageNumber="75" type="holotype">Holotype</typeStatus>
</emphasis>
:
<specimenCode id="DBF69EFE1B51A700FE1C6360FD5D712D" box="[470,650,1362,1386]" collectionCode="KPM-NK" pageId="20" pageNumber="75">KPM-NK 81861</specimenCode>
, F,
<collectingCountry id="F34776151B51A700FD7B6361FCD5712D" box="[689,770,1362,1386]" name="Japan" pageId="20" pageNumber="75">JAPAN</collectingCountry>
,
<collectingRegion id="4994F8671B51A700FF776349FE9E71D6" box="[189,329,1401,1426]" country="Japan" name="Nagano" pageId="20" pageNumber="75">Nagano Pref.</collectingRegion>
, Outaki Vil.,
<collectingMunicipality id="6B8BACFF1B51A700FE196349FDB971D6" box="[467,622,1401,1425]" pageId="20" pageNumber="75">Mt. Ontakesan</collectingMunicipality>
,
<location id="8E8F605E1B51A700FDB3634AFD3071D6" LSID="urn:lsid:plazi:treatment:03F987931B51A702FE8C62BCFEB3770E:8E8F605E1B51A700FDB3634AFD3071D6" box="[633,743,1401,1425]" country="Japan" municipality="Mt. Ontakesan" name="Hakkaisan" pageId="20" pageNumber="75" stateProvince="Nagano">Hakkaisan</location>
,
<date id="FFEE10451B51A700FD39634AFEFC71FF" pageId="20" pageNumber="75" value="2007-08-04">
<collectingDate id="EFAAE9AD1B51A700FD39634AFEFC71FF" pageId="20" pageNumber="75" value="2007-08-04">4. VIII. 2007</collectingDate>
</date>
,
<collectorName id="26A553531B51A700FEFD6393FE6B71FF" box="[311,444,1440,1464]" pageId="20" pageNumber="75">K. Watanabe</collectorName>
leg.
</materialsCitation>
<materialsCitation id="3B383CD81B51A700FE3A6393FCD57198" ID-GBIF-Occurrence="4906826565" collectionCode="KPM-NK" country="Japan" location="Kinunuma" pageId="20" pageNumber="75" specimenCode="KPM-NK 81872" specimenCount="1" stateProvince="Tochigi" typeStatus="paratype">
<emphasis id="B924EA971B51A700FE3A6393FD8B71FF" bold="true" box="[496,604,1440,1464]" pageId="20" pageNumber="75">
<typeStatus id="54EB88271B51A700FE3A6393FD8B71FF" box="[496,604,1440,1464]" pageId="20" pageNumber="75" type="paratype">Paratypes</typeStatus>
</emphasis>
:
<collectingCountry id="F34776151B51A700FDA16393FD6C71FF" box="[619,699,1440,1464]" name="Japan" pageId="20" pageNumber="75">JAPAN</collectingCountry>
,
<specimenCode id="DBF69EFE1B51A700FD0C6393FEF87198" collectionCode="KPM-NK" pageId="20" pageNumber="75">KPM-NK 81872</specimenCode>
, F,
<collectingRegion id="4994F8671B51A700FE9C63F4FE357198" box="[342,482,1479,1503]" country="Japan" name="Tochigi" pageId="20" pageNumber="75">Tochigi Pref.</collectingRegion>
, Kuriyama Vil.,
<location id="8E8F605E1B51A700FD5863F4FCD57198" LSID="urn:lsid:plazi:treatment:03F987931B51A702FE8C62BCFEB3770E:8E8F605E1B51A700FD5863F4FCD57198" box="[658,770,1479,1503]" country="Japan" name="Kinunuma" pageId="20" pageNumber="75" stateProvince="Tochigi">Kinunuma</location>
</materialsCitation>
,
<materialsCitation id="3B383CD81B51A700FC9866AEFBD37544" ID-GBIF-Occurrence="4906826504" collectingDate="2004-08-01" collectingDateMax="2004-08-14" collectingDateMin="2004-08-01" collectorName="H. Makihara" county="Nagawa Town" location="Utsukushimatsu" municipality="Daimon" pageId="20" pageNumber="75" specimenCode="KPM-NK 81863" specimenCount="1" stateProvince="Nagano">
<date id="FFEE10451B51A700FC9866AEFBCD74F2" box="[850,1050,157,181]" pageId="20" pageNumber="75" value="2004-08-01" valueMax="2004-08-14" valueMin="2004-08-01">
<collectingDate id="EFAAE9AD1B51A700FC9866AEFBCD74F2" box="[850,1050,157,181]" pageId="20" pageNumber="75" value="2004-08-01" valueMax="2004-08-14" valueMin="2004-08-01">1–14. VIII. 2004</collectingDate>
</date>
,
<collectorName id="26A553531B51A700FBE066AEFB6B74F2" box="[1066,1212,157,181]" pageId="20" pageNumber="75">H. Makihara</collectorName>
leg. (MsT);
<specimenCode id="DBF69EFE1B51A700FA9D66AEFC1F749B" collectionCode="KPM-NK" pageId="20" pageNumber="75">KPM-NK 81863</specimenCode>
, F,
<collectingRegion id="4994F8671B51A700FC3866F7FB52749B" box="[1010,1157,196,220]" country="Japan" name="Nagano" pageId="20" pageNumber="75">Nagano Pref.</collectingRegion>
,
<collectingCounty id="628E4E091B51A700FB5866F7FAE5749C" box="[1170,1330,196,220]" pageId="20" pageNumber="75">Nagawa Town</collectingCounty>
,
<collectingMunicipality id="6B8BACFF1B51A700FAF566F7FA40749B" box="[1343,1431,196,220]" pageId="20" pageNumber="75">Daimon</collectingMunicipality>
,
<location id="8E8F605E1B51A700FC9866D8FBD37544" LSID="urn:lsid:plazi:treatment:03F987931B51A702FE8C62BCFEB3770E:8E8F605E1B51A700FC9866D8FBD37544" box="[850,1028,235,259]" county="Nagawa Town" municipality="Daimon" name="Utsukushimatsu" pageId="20" pageNumber="75" stateProvince="Nagano">Utsukushimatsu</location>
</materialsCitation>
,
<materialsCitation id="3B383CD81B51A700FBDB66D8FC6F7516" ID-GBIF-Occurrence="4906826601" collectingDate="2011-08-26" collectorName="S. Fujie" county="Outaki Vil." location="Tanohara" municipality="Mt. Ontakesan" pageId="20" pageNumber="75" specimenCode="KPM-NK 81862" specimenCount="1" stateProvince="Nagano">
<date id="FFEE10451B51A700FBDB66D8FB647544" box="[1041,1203,235,259]" pageId="20" pageNumber="75" value="2011-08-26">
<collectingDate id="EFAAE9AD1B51A700FBDB66D8FB647544" box="[1041,1203,235,259]" pageId="20" pageNumber="75" value="2011-08-26">26. VIII. 2011</collectingDate>
</date>
,
<collectorName id="26A553531B51A700FB0A66D8FAC07544" box="[1216,1303,235,259]" pageId="20" pageNumber="75">S. Fujie</collectorName>
leg.;
<specimenCode id="DBF69EFE1B51A700FA9366D8FC15756D" collectionCode="KPM-NK" pageId="20" pageNumber="75">KPM-NK 81862</specimenCode>
, F,
<collectingRegion id="4994F8671B51A700FC226721FBA3756D" box="[1000,1140,274,298]" country="Japan" name="Nagano" pageId="20" pageNumber="75">Nagano Pref.</collectingRegion>
,
<collectingCounty id="628E4E091B51A700FBB46721FB24756D" box="[1150,1267,274,298]" pageId="20" pageNumber="75">Outaki Vil.</collectingCounty>
,
<collectingMunicipality id="6B8BACFF1B51A700FB346721FA4F756D" box="[1278,1432,274,298]" pageId="20" pageNumber="75">Mt. Ontakesan</collectingMunicipality>
,
<location id="8E8F605E1B51A700FC98670AFC6F7516" LSID="urn:lsid:plazi:treatment:03F987931B51A702FE8C62BCFEB3770E:8E8F605E1B51A700FC98670AFC6F7516" box="[850,952,313,337]" county="Outaki Vil." municipality="Mt. Ontakesan" name="Tanohara" pageId="20" pageNumber="75" stateProvince="Nagano">Tanohara</location>
</materialsCitation>
,
<materialsCitation id="3B383CD81B51A700FC0C670AFA40753F" ID-GBIF-Occurrence="4906826567" collectingDate="2007-08-08" collectorName="K. Watanabe" location="Mt. Ontake-san" municipality="Otaki Vil." pageId="20" pageNumber="75" specimenCode="KPM-NK 81864" specimenCount="1" stateProvince="Nagano">
<date id="FFEE10451B51A700FC0C670AFB8F7516" box="[966,1112,313,337]" pageId="20" pageNumber="75" value="2007-08-08">
<collectingDate id="EFAAE9AD1B51A700FC0C670AFB8F7516" box="[966,1112,313,337]" pageId="20" pageNumber="75" value="2007-08-08">8. VIII. 2007</collectingDate>
</date>
,
<collectorName id="26A553531B51A700FBAC670AFB267516" box="[1126,1265,313,337]" pageId="20" pageNumber="75">K. Watanabe</collectorName>
leg.;
<specimenCode id="DBF69EFE1B51A700FAF9670AFC4E753F" collectionCode="KPM-NK" pageId="20" pageNumber="75">KPM-NK 81864</specimenCode>
, F,
<collectingRegion id="4994F8671B51A700FC0F6753FB8B753F" box="[965,1116,352,376]" country="Japan" name="Nagano" pageId="20" pageNumber="75">Nagano Pref.</collectingRegion>
,
<collectingMunicipality id="6B8BACFF1B51A700FBA36753FB0D753F" box="[1129,1242,352,378]" pageId="20" pageNumber="75">Ōtaki Vil.</collectingMunicipality>
,
<location id="8E8F605E1B51A700FB226753FA40753F" LSID="urn:lsid:plazi:treatment:03F987931B51A702FE8C62BCFEB3770E:8E8F605E1B51A700FB226753FA40753F" box="[1256,1431,352,376]" municipality="Otaki Vil." name="Mt. Ontake-san" pageId="20" pageNumber="75" stateProvince="Nagano">Mt. Ontake-san</location>
</materialsCitation>
,
<materialsCitation id="3B383CD81B51A700FC9867B5FC1475AB" ID-GBIF-Occurrence="4906826464" collectingDate="2015-06-13" collectingDateMax="2015-06-25" collectingDateMin="2015-06-13" collectorName="S. Shimizu" county="City" location="Jyuroudani" municipality="Arimine" pageId="20" pageNumber="75" specimenCode="KPM-NK 81860, 81865, 81866, 3" specimenCount="1" stateProvince="Toyama Pref., Toyama">
<date id="FFEE10451B51A700FC9867B5FC2F75D9" box="[850,1016,390,414]" pageId="20" pageNumber="75" value="2015-06-13" valueMax="2015-06-25" valueMin="2015-06-13">
<collectingDate id="EFAAE9AD1B51A700FC9867B5FC2F75D9" box="[850,1016,390,414]" pageId="20" pageNumber="75" value="2015-06-13" valueMax="2015-06-25" valueMin="2015-06-13">13–25. VI. 2015</collectingDate>
</date>
,
<collectorName id="26A553531B51A700FBCB67B4FBA775D9" box="[1025,1136,390,415]" pageId="20" pageNumber="75">S. Shimizu</collectorName>
leg. (MsT);
<specimenCode id="DBF69EFE1B51A700FB2667B4FA4E75D9" box="[1260,1433,390,414]" collectionCode="KPM-NK" pageId="20" pageNumber="75">KPM-NK 81860</specimenCode>
,
<specimenCode id="DBF69EFE1B51A700FC98679EFC437582" box="[850,916,429,453]" collectionCode="KPM-NK" pageId="20" pageNumber="75">81865</specimenCode>
,
<specimenCode id="DBF69EFE1B51A700FC6A679EFC357582" box="[928,994,429,453]" collectionCode="KPM-NK" pageId="20" pageNumber="75">81866</specimenCode>
,
<specimenCode id="DBF69EFE1B51A700FC27679DFC2D7581" box="[1005,1018,430,454]" collectionCode="KPM-NK" pageId="20" pageNumber="75">3</specimenCode>
F,
<collectingRegion id="4994F8671B51A700FBD6679DFAD37581" box="[1052,1284,429,454]" isEnumeration="true" pageId="20" pageNumber="75">Toyama Pref., Toyama</collectingRegion>
<collectingCounty id="628E4E091B51A700FAC6679EFAEF7582" box="[1292,1336,429,453]" pageId="20" pageNumber="75">City</collectingCounty>
,
<collectingMunicipality id="6B8BACFF1B51A700FA88679EFA4E7582" box="[1346,1433,429,453]" pageId="20" pageNumber="75">Arimine</collectingMunicipality>
,
<location id="8E8F605E1B51A700FC9867E7FC1475AB" LSID="urn:lsid:plazi:treatment:03F987931B51A702FE8C62BCFEB3770E:8E8F605E1B51A700FC9867E7FC1475AB" box="[850,963,468,492]" county="City" municipality="Arimine" name="Jyuroudani" pageId="20" pageNumber="75" stateProvince="Toyama Pref., Toyama">Jyuroudani</location>
</materialsCitation>
,
<materialsCitation id="3B383CD81B51A700FC0667E7FA9B7654" ID-GBIF-Occurrence="4906826411" collectingDate="2009-09-08" collectingDateMax="2009-09-15" collectingDateMin="2009-09-08" collectorName="M. Watanabe" county="City" location="Jyuroudani" municipality="Arimine" pageId="20" pageNumber="75" specimenCode="KPM-NK 81858, 81859, 81867-81869, 5" specimenCount="1" stateProvince="Toyama Pref., Toyama">
<date id="FFEE10451B51A700FC0667E7FBB375AA" box="[972,1124,468,493]" pageId="20" pageNumber="75" value="2009-09-08" valueMax="2009-09-15" valueMin="2009-09-08">
<collectingDate id="EFAAE9AD1B51A700FC0667E7FBB375AA" box="[972,1124,468,493]" pageId="20" pageNumber="75" value="2009-09-08" valueMax="2009-09-15" valueMin="2009-09-08">8–15. IX. 2009</collectingDate>
</date>
,
<collectorName id="26A553531B51A700FBA667E6FB2775AB" box="[1132,1264,468,492]" pageId="20" pageNumber="75">M. Watanabe</collectorName>
<emphasis id="B924EA971B51A700FB3C67E6FAF075AB" box="[1270,1319,468,492]" italics="true" pageId="20" pageNumber="75">et al.</emphasis>
leg. (MsT);
<specimenCode id="DBF69EFE1B51A700FC9867CFFBD67654" box="[850,1025,507,531]" collectionCode="KPM-NK" pageId="20" pageNumber="75">KPM-NK 81858</specimenCode>
,
<specimenCode id="DBF69EFE1B51A700FBC167C8FB9A7654" box="[1035,1101,507,531]" collectionCode="KPM-NK" pageId="20" pageNumber="75">81859</specimenCode>
,
<specimenCode id="DBF69EFE1B51A700FB9D67C8FB337654" box="[1111,1252,507,531]" collectionCode="KPM-NK" pageId="20" pageNumber="75">81867–81869</specimenCode>
,
<specimenCode id="DBF69EFE1B51A700FB2567C8FB2B7654" box="[1263,1276,507,531]" collectionCode="KPM-NK" pageId="20" pageNumber="75">5</specimenCode>
F, ditto
</materialsCitation>
,
<materialsCitation id="3B383CD81B51A700FA9D67C8FB627626" ID-GBIF-Occurrence="4906826569" collectingDate="2009-09-15" collectingDateMax="2009-09-22" collectingDateMin="2009-09-15" county="City" location="Inonedani" municipality="Arimine" pageId="20" pageNumber="75" specimenCode="KPM-NK 81870, 81871, 2" specimenCount="1" stateProvince="Toyama Pref., Toyama">
<date id="FFEE10451B51A700FA9D67C8FC62767D" pageId="20" pageNumber="75" value="2009-09-15" valueMax="2009-09-22" valueMin="2009-09-15">
<collectingDate id="EFAAE9AD1B51A700FA9D67C8FC62767D" pageId="20" pageNumber="75" value="2009-09-15" valueMax="2009-09-22" valueMin="2009-09-15">15–22. IX. 2009</collectingDate>
</date>
;
<specimenCode id="DBF69EFE1B51A700FC086410FBAF767D" box="[962,1144,546,570]" collectionCode="KPM-NK" pageId="20" pageNumber="75">KPM-NK 81870</specimenCode>
,
<specimenCode id="DBF69EFE1B51A700FB4E6411FB1F767D" box="[1156,1224,546,570]" collectionCode="KPM-NK" pageId="20" pageNumber="75">81871</specimenCode>
,
<specimenCode id="DBF69EFE1B51A700FB1F6411FB35767D" box="[1237,1250,546,570]" collectionCode="KPM-NK" pageId="20" pageNumber="75">2</specimenCode>
F,
<collectingRegion id="4994F8671B51A700FACC6410FC717625" isEnumeration="true" pageId="20" pageNumber="75">Toyama Pref., Toyama</collectingRegion>
<collectingCounty id="628E4E091B51A700FC64647AFC0B7626" box="[942,988,585,609]" pageId="20" pageNumber="75">City</collectingCounty>
,
<collectingMunicipality id="6B8BACFF1B51A700FC2C647AFBE87626" box="[998,1087,585,609]" pageId="20" pageNumber="75">Arimine</collectingMunicipality>
,
<location id="8E8F605E1B51A700FB81647AFB627626" LSID="urn:lsid:plazi:treatment:03F987931B51A702FE8C62BCFEB3770E:8E8F605E1B51A700FB81647AFB627626" box="[1099,1205,585,609]" county="City" municipality="Arimine" name="Inonedani" pageId="20" pageNumber="75" stateProvince="Toyama Pref., Toyama">Inonedani</location>
</materialsCitation>
,
<date id="FFEE10451B51A700FB0B647AFAA57626" box="[1217,1394,585,609]" pageId="20" pageNumber="75" value="2009-09-15" valueMax="2009-09-22" valueMin="2009-09-15">15–22. IX. 2009</date>
, M. Watanabe
<emphasis id="B924EA971B51A700FC706442FC3276CF" box="[954,997,624,648]" italics="true" pageId="20" pageNumber="75">et al</emphasis>
. leg. (MsT).
</paragraph>
</subSubSection>
<subSubSection id="C34A650E1B51A702FCA164A4FE5775DA" lastPageId="22" lastPageNumber="77" pageId="20" pageNumber="75" type="description">
<paragraph id="8BEF36851B51A700FCA164A4FB3C7691" blockId="20.[850,1437,157,1894]" pageId="20" pageNumber="75">
<emphasis id="B924EA971B51A700FCA164A4FC3176E8" bold="true" box="[875,998,663,687]" pageId="20" pageNumber="75">Description</emphasis>
. Female (n=15). Body length 4.6–7.1 (HT: 5.7) mm. Body covered with silver setae.
</paragraph>
<paragraph id="8BEF36851B51A700FCA164D6FBDD7097" blockId="20.[850,1437,157,1894]" pageId="20" pageNumber="75">
Head. Clypeus 1.8 times as wide as maximum length, its anterior margin rounded with a median broad subtruncate convexity (
<figureCitation id="136B2A001B51A700FC0E6500FBF3770C" box="[964,1060,819,843]" captionStart="Fig" captionStartId="72.[190,223,1065,1087]" captionTargetBox="[195,1428,161,998]" captionTargetId="figure-227@72.[189,1436,149,1010]" captionTargetPageId="72" captionText="Fig. 62. Bathythrix kuwanae Viereck, 1912 (A, KPM-NK 81718), Ba. margaretae Sawoniewicz, 1980 (B, KPM-NK 55017), Ba. thomsoni (Kerrich, 1942) (C, KPM-NK 81675), Diaglyptidea conformis (Gmelin, 1790) (D, KPM-NK 81645), Nipponaetes haeussleri (Uchida, 1933) (E, KPM-NK 81832), Acrolyta japonica sp. nov. (F, N, KPM-NK 54996, holotype), Acr. rufocincta (Gravenhorst, 1829) (G, KPM-NK 81732), Surculus japonicus sp. nov. (H, KPM-NK 81861, holotype), Isadelphus nigrus sp. nov. (I, KPM-NK 82003, holotype), Lochetica japonica sp. nov. (J, KPM-NK 81989, holotype), Megacara similis Sheng, 1999 (K, KPM-NK 81837), Gnotus chionops (Gravenhorst, 1829) (L, KPM-NK 82100), Gn. striatus (Uchida, 1930) (M, KPM-NK 82060), and Acr. flavicoxis Sheng &amp; Sun, 2014 (O, KPM-NK 81837), females from Japan ― A–C, head, frontal view; D, E, head, lateral view; F–K, anterior margin of clypeus; L, M, mandible; N, O, basal part of antenna, lateral view." figureDoi="http://doi.org/10.5281/zenodo.12774781" httpUri="https://zenodo.org/record/12774781/files/figure.png" pageId="20" pageNumber="75">Fig. 62H</figureCitation>
). Face polished, punctate (
<figureCitation id="136B2A001B51A700FAF96500FA47770C" box="[1331,1424,819,843]" captionStart="Fig" captionStartId="21.[153,186,1985,2007]" captionTargetBox="[151,1398,998,1961]" captionTargetId="figure-368@21.[151,1398,996,1961]" captionTargetPageId="21" captionText="Fig. 18. Surculus japonicus sp. nov., KPM-NK 81861, holotype, female ― A, lateral habitus; B, head, frontal view; C, head, mesosoma, and metasoma, dorsal view; D, head and mesosoma, lateral view; E, head, mesoscutum and scutellum, dorsal view; F, wings; G, propodeum,dorsal view." figureDoi="http://doi.org/10.5281/zenodo.12774677" httpUri="https://zenodo.org/record/12774677/files/figure.png" pageId="20" pageNumber="75">Fig. 18B</figureCitation>
). MSL 0.8 times as long as BWM. Frons, gena, and vertex polished, with fine punctures. OOL longer than POL. Occipital carina complete, its lower end joined with hypostomal carina distant from mandibular base. Upper tooth of mandible almost as long as lower tooth. Base of mandible slightly convex. Antenna with 18–19 (HT: 18) flagellomeres, subapical part not widened, all segments longer than wide. FL III 6.0–7.0 (HT: 7.0) times as long as maximum depth in lateral view. Apex of flagellum with some robust setae.
</paragraph>
<paragraph id="8BEF36851B51A701FCA162D3FEB576EA" blockId="20.[850,1437,157,1894]" lastBlockId="21.[151,738,155,958]" lastPageId="21" lastPageNumber="76" pageId="20" pageNumber="75">
Mesosoma 1.7 times as long as maximum depth in lateral view. Upper side of collar with a median carina that is not much stronger than surrounding carinae. Lateral part of pronotum largely smooth except for some longitudinal striae medially (
<figureCitation id="136B2A001B51A700FC326348FB8F71D4" box="[1016,1112,1403,1427]" captionStart="Fig" captionStartId="21.[153,186,1985,2007]" captionTargetBox="[151,1398,998,1961]" captionTargetId="figure-368@21.[151,1398,996,1961]" captionTargetPageId="21" captionText="Fig. 18. Surculus japonicus sp. nov., KPM-NK 81861, holotype, female ― A, lateral habitus; B, head, frontal view; C, head, mesosoma, and metasoma, dorsal view; D, head and mesosoma, lateral view; E, head, mesoscutum and scutellum, dorsal view; F, wings; G, propodeum,dorsal view." figureDoi="http://doi.org/10.5281/zenodo.12774677" httpUri="https://zenodo.org/record/12774677/files/figure.png" pageId="20" pageNumber="75">Fig. 18D</figureCitation>
). Epomia long and strong (
<figureCitation id="136B2A001B51A700FAB96348FC5671FD" captionStart="Fig" captionStartId="21.[153,186,1985,2007]" captionTargetBox="[151,1398,998,1961]" captionTargetId="figure-368@21.[151,1398,996,1961]" captionTargetPageId="21" captionText="Fig. 18. Surculus japonicus sp. nov., KPM-NK 81861, holotype, female ― A, lateral habitus; B, head, frontal view; C, head, mesosoma, and metasoma, dorsal view; D, head and mesosoma, lateral view; E, head, mesoscutum and scutellum, dorsal view; F, wings; G, propodeum,dorsal view." figureDoi="http://doi.org/10.5281/zenodo.12774677" httpUri="https://zenodo.org/record/12774677/files/figure.png" pageId="20" pageNumber="75">Fig. 18D</figureCitation>
). Mesoscutum polished, punctate except for postero-median rugulose area (
<figureCitation id="136B2A001B51A700FB8E63FAFB7371A6" box="[1092,1188,1481,1505]" captionStart="Fig" captionStartId="21.[153,186,1985,2007]" captionTargetBox="[151,1398,998,1961]" captionTargetId="figure-368@21.[151,1398,996,1961]" captionTargetPageId="21" captionText="Fig. 18. Surculus japonicus sp. nov., KPM-NK 81861, holotype, female ― A, lateral habitus; B, head, frontal view; C, head, mesosoma, and metasoma, dorsal view; D, head and mesosoma, lateral view; E, head, mesoscutum and scutellum, dorsal view; F, wings; G, propodeum,dorsal view." figureDoi="http://doi.org/10.5281/zenodo.12774677" httpUri="https://zenodo.org/record/12774677/files/figure.png" pageId="20" pageNumber="75">Fig. 18E</figureCitation>
). Notaulus long, fading our near center of mesoscutum (
<figureCitation id="136B2A001B51A700FB5E63C3FB26724F" box="[1172,1265,1520,1544]" captionStart="Fig" captionStartId="21.[153,186,1985,2007]" captionTargetBox="[151,1398,998,1961]" captionTargetId="figure-368@21.[151,1398,996,1961]" captionTargetPageId="21" captionText="Fig. 18. Surculus japonicus sp. nov., KPM-NK 81861, holotype, female ― A, lateral habitus; B, head, frontal view; C, head, mesosoma, and metasoma, dorsal view; D, head and mesosoma, lateral view; E, head, mesoscutum and scutellum, dorsal view; F, wings; G, propodeum,dorsal view." figureDoi="http://doi.org/10.5281/zenodo.12774677" httpUri="https://zenodo.org/record/12774677/files/figure.png" pageId="20" pageNumber="75">Figs 18E</figureCitation>
,
<figureCitation id="136B2A001B51A700FB3163C3FAF1724F" box="[1275,1318,1520,1544]" captionStart="Fig" captionStartId="73.[151,185,989,1011]" captionTargetBox="[168,1379,161,949]" captionTargetId="figure-5@73.[151,1398,151,958]" captionTargetPageId="73" captionText="Fig. 63. Acrolyta japonica sp. nov. (A, KPM-NK 54996, holotype), Dichrogaster liostylus (Thomson, 1885) (B, KPM-NK 55074), Dic. nitida Sheng &amp; Sun, 2014 (C, KPM-NK 81908), Bathythrix kuwanae Viereck, 1912 (D, KPM-NK 81718), Retalia japonica Kusigemati, 1985 (E, KPM-NK 81676), Surculus japonicus sp. nov. (F, KPM-NK 81861, holotype), Bentyra ryukyuana sp. nov. (G, KPM-NK 81887, holotype), Hemiteles japonicus sp. nov. (H, KPM-NK 55021, holotype), and H. maculipterus sp. nov. (I, KPM-NK 55025, holotype), females from Japan ― A, median part of collar, dorsal view; B, C, lateral part of pronotum; D–G, mesoscutum and scutellum,dorsal view; H, I, hind tarsus, lateral view." figureDoi="http://doi.org/10.5281/zenodo.12774783" httpUri="https://zenodo.org/record/12774783/files/figure.png" pageId="20" pageNumber="75">63F</figureCitation>
). Scutellum punctate, without a lateral longitudinal carina except for base. Mesopleuron smooth except for areas below subtegular ridge and speculum with longitudinal striae (
<figureCitation id="136B2A001B51A700FC9060B8FC6B72E4" box="[858,956,1675,1699]" captionStart="Fig" captionStartId="21.[153,186,1985,2007]" captionTargetBox="[151,1398,998,1961]" captionTargetId="figure-368@21.[151,1398,996,1961]" captionTargetPageId="21" captionText="Fig. 18. Surculus japonicus sp. nov., KPM-NK 81861, holotype, female ― A, lateral habitus; B, head, frontal view; C, head, mesosoma, and metasoma, dorsal view; D, head and mesosoma, lateral view; E, head, mesoscutum and scutellum, dorsal view; F, wings; G, propodeum,dorsal view." figureDoi="http://doi.org/10.5281/zenodo.12774677" httpUri="https://zenodo.org/record/12774677/files/figure.png" pageId="20" pageNumber="75">Fig. 18D</figureCitation>
). The impression that is just below speculum with a pit, which situated in the short horizontal groove that joins mesopleural suture. Posterior transverse carina of mesosternum largely absent in front of mid coxae. Metapleuron largely punctate, with a complete juxtacoxal carina. Anterior and posterior transverse carinae of propodeum complete (
<figureCitation id="136B2A001B50A701FEB266A8FE0274F4" box="[376,469,155,179]" captionStart="Fig" captionStartId="73.[151,185,1926,1948]" captionTargetBox="[162,1384,1211,1880]" captionTargetId="figure-109@73.[151,1398,1199,1899]" captionTargetPageId="73" captionText="Fig. 64. Surculus japonicus sp. nov. (A, KPM-NK 81861, holotype), Paraphylax elegans sp. nov. (B, KPM-NK 55078, holotype), Hemiteles japonicus sp. nov. (C, KPM-NK 55021, holotype), H. maculipterus sp. nov. (D, KPM-NK 55026, paratype), Charitopes gastricus (Holmgren, 1868) (E, KPM-NK 81963), Tropistes shimizui sp. nov. (F, KPM-NK 81833, holotype), Uchidella okamotoi (Uchida, 1936) (G, KPM-NK 82073), and Uchidella toichii sp. nov. (H, KPM-NK 82011, holotype), females from Japan ― A–H, propodeum,dorsal view." figureDoi="http://doi.org/10.5281/zenodo.12774785" httpUri="https://zenodo.org/record/12774785/files/figure.png" pageId="21" pageNumber="76">Fig. 64A</figureCitation>
).Anterior section of lateral longitudinal carina of propodeum absent (
<figureCitation id="136B2A001B50A701FD8A66F1FD48749D" box="[576,671,194,218]" captionStart="Fig" captionStartId="73.[151,185,1926,1948]" captionTargetBox="[162,1384,1211,1880]" captionTargetId="figure-109@73.[151,1398,1199,1899]" captionTargetPageId="73" captionText="Fig. 64. Surculus japonicus sp. nov. (A, KPM-NK 81861, holotype), Paraphylax elegans sp. nov. (B, KPM-NK 55078, holotype), Hemiteles japonicus sp. nov. (C, KPM-NK 55021, holotype), H. maculipterus sp. nov. (D, KPM-NK 55026, paratype), Charitopes gastricus (Holmgren, 1868) (E, KPM-NK 81963), Tropistes shimizui sp. nov. (F, KPM-NK 81833, holotype), Uchidella okamotoi (Uchida, 1936) (G, KPM-NK 82073), and Uchidella toichii sp. nov. (H, KPM-NK 82011, holotype), females from Japan ― A–H, propodeum,dorsal view." figureDoi="http://doi.org/10.5281/zenodo.12774785" httpUri="https://zenodo.org/record/12774785/files/figure.png" pageId="21" pageNumber="76">Fig. 64A</figureCitation>
). Area basalis distinct (
<figureCitation id="136B2A001B50A701FE8966DAFE717546" box="[323,422,233,257]" captionStart="Fig" captionStartId="73.[151,185,1926,1948]" captionTargetBox="[162,1384,1211,1880]" captionTargetId="figure-109@73.[151,1398,1199,1899]" captionTargetPageId="73" captionText="Fig. 64. Surculus japonicus sp. nov. (A, KPM-NK 81861, holotype), Paraphylax elegans sp. nov. (B, KPM-NK 55078, holotype), Hemiteles japonicus sp. nov. (C, KPM-NK 55021, holotype), H. maculipterus sp. nov. (D, KPM-NK 55026, paratype), Charitopes gastricus (Holmgren, 1868) (E, KPM-NK 81963), Tropistes shimizui sp. nov. (F, KPM-NK 81833, holotype), Uchidella okamotoi (Uchida, 1936) (G, KPM-NK 82073), and Uchidella toichii sp. nov. (H, KPM-NK 82011, holotype), females from Japan ― A–H, propodeum,dorsal view." figureDoi="http://doi.org/10.5281/zenodo.12774785" httpUri="https://zenodo.org/record/12774785/files/figure.png" pageId="21" pageNumber="76">Fig. 64A</figureCitation>
). Area superomedia indistinct laterally (
<figureCitation id="136B2A001B50A701FF346723FEB3756F" box="[254,356,272,296]" captionStart="Fig" captionStartId="21.[153,186,1985,2007]" captionTargetBox="[151,1398,998,1961]" captionTargetId="figure-368@21.[151,1398,996,1961]" captionTargetPageId="21" captionText="Fig. 18. Surculus japonicus sp. nov., KPM-NK 81861, holotype, female ― A, lateral habitus; B, head, frontal view; C, head, mesosoma, and metasoma, dorsal view; D, head and mesosoma, lateral view; E, head, mesoscutum and scutellum, dorsal view; F, wings; G, propodeum,dorsal view." figureDoi="http://doi.org/10.5281/zenodo.12774677" httpUri="https://zenodo.org/record/12774677/files/figure.png" pageId="21" pageNumber="76">Figs 18G</figureCitation>
,
<figureCitation id="136B2A001B50A701FEB86723FE74756F" box="[370,419,272,296]" captionStart="Fig" captionStartId="73.[151,185,1926,1948]" captionTargetBox="[162,1384,1211,1880]" captionTargetId="figure-109@73.[151,1398,1199,1899]" captionTargetPageId="73" captionText="Fig. 64. Surculus japonicus sp. nov. (A, KPM-NK 81861, holotype), Paraphylax elegans sp. nov. (B, KPM-NK 55078, holotype), Hemiteles japonicus sp. nov. (C, KPM-NK 55021, holotype), H. maculipterus sp. nov. (D, KPM-NK 55026, paratype), Charitopes gastricus (Holmgren, 1868) (E, KPM-NK 81963), Tropistes shimizui sp. nov. (F, KPM-NK 81833, holotype), Uchidella okamotoi (Uchida, 1936) (G, KPM-NK 82073), and Uchidella toichii sp. nov. (H, KPM-NK 82011, holotype), females from Japan ― A–H, propodeum,dorsal view." figureDoi="http://doi.org/10.5281/zenodo.12774785" httpUri="https://zenodo.org/record/12774785/files/figure.png" pageId="21" pageNumber="76">64A</figureCitation>
). Anterior part of propodeum punctate. Median and posterior parts of propodeum longitudinally, obliquely, and irregularly rugulose. Propodeum without projections (
<figureCitation id="136B2A001B50A701FE2267B6FD9C75DA" box="[488,587,389,413]" captionStart="Fig" captionStartId="21.[153,186,1985,2007]" captionTargetBox="[151,1398,998,1961]" captionTargetId="figure-368@21.[151,1398,996,1961]" captionTargetPageId="21" captionText="Fig. 18. Surculus japonicus sp. nov., KPM-NK 81861, holotype, female ― A, lateral habitus; B, head, frontal view; C, head, mesosoma, and metasoma, dorsal view; D, head and mesosoma, lateral view; E, head, mesoscutum and scutellum, dorsal view; F, wings; G, propodeum,dorsal view." figureDoi="http://doi.org/10.5281/zenodo.12774677" httpUri="https://zenodo.org/record/12774677/files/figure.png" pageId="21" pageNumber="76">Figs 18G</figureCitation>
,
<figureCitation id="136B2A001B50A701FD9C67B6FD5175DA" box="[598,646,389,413]" captionStart="Fig" captionStartId="73.[151,185,1926,1948]" captionTargetBox="[162,1384,1211,1880]" captionTargetId="figure-109@73.[151,1398,1199,1899]" captionTargetPageId="73" captionText="Fig. 64. Surculus japonicus sp. nov. (A, KPM-NK 81861, holotype), Paraphylax elegans sp. nov. (B, KPM-NK 55078, holotype), Hemiteles japonicus sp. nov. (C, KPM-NK 55021, holotype), H. maculipterus sp. nov. (D, KPM-NK 55026, paratype), Charitopes gastricus (Holmgren, 1868) (E, KPM-NK 81963), Tropistes shimizui sp. nov. (F, KPM-NK 81833, holotype), Uchidella okamotoi (Uchida, 1936) (G, KPM-NK 82073), and Uchidella toichii sp. nov. (H, KPM-NK 82011, holotype), females from Japan ― A–H, propodeum,dorsal view." figureDoi="http://doi.org/10.5281/zenodo.12774785" httpUri="https://zenodo.org/record/12774785/files/figure.png" pageId="21" pageNumber="76">64A</figureCitation>
). Length of fore wing 4.4–5.6 (HT: 4.9) mm. Areolet absent (
<figureCitation id="136B2A001B50A701FD7D679FFF1575AC" captionStart="Fig" captionStartId="21.[153,186,1985,2007]" captionTargetBox="[151,1398,998,1961]" captionTargetId="figure-368@21.[151,1398,996,1961]" captionTargetPageId="21" captionText="Fig. 18. Surculus japonicus sp. nov., KPM-NK 81861, holotype, female ― A, lateral habitus; B, head, frontal view; C, head, mesosoma, and metasoma, dorsal view; D, head and mesosoma, lateral view; E, head, mesoscutum and scutellum, dorsal view; F, wings; G, propodeum,dorsal view." figureDoi="http://doi.org/10.5281/zenodo.12774677" httpUri="https://zenodo.org/record/12774677/files/figure.png" pageId="21" pageNumber="76">Fig. 18F</figureCitation>
). Vein 2m-cu of fore wing with two bullae (
<figureCitation id="136B2A001B50A701FDB767E0FD0175AC" box="[637,726,467,491]" captionStart="Fig" captionStartId="21.[153,186,1985,2007]" captionTargetBox="[151,1398,998,1961]" captionTargetId="figure-368@21.[151,1398,996,1961]" captionTargetPageId="21" captionText="Fig. 18. Surculus japonicus sp. nov., KPM-NK 81861, holotype, female ― A, lateral habitus; B, head, frontal view; C, head, mesosoma, and metasoma, dorsal view; D, head and mesosoma, lateral view; E, head, mesoscutum and scutellum, dorsal view; F, wings; G, propodeum,dorsal view." figureDoi="http://doi.org/10.5281/zenodo.12774677" httpUri="https://zenodo.org/record/12774677/files/figure.png" pageId="21" pageNumber="76">Fig. 18F</figureCitation>
). Nervellus inclivous, intercepted near the middle (
<figureCitation id="136B2A001B50A701FDB767C9FD017655" box="[637,726,506,530]" captionStart="Fig" captionStartId="21.[153,186,1985,2007]" captionTargetBox="[151,1398,998,1961]" captionTargetId="figure-368@21.[151,1398,996,1961]" captionTargetPageId="21" captionText="Fig. 18. Surculus japonicus sp. nov., KPM-NK 81861, holotype, female ― A, lateral habitus; B, head, frontal view; C, head, mesosoma, and metasoma, dorsal view; D, head and mesosoma, lateral view; E, head, mesoscutum and scutellum, dorsal view; F, wings; G, propodeum,dorsal view." figureDoi="http://doi.org/10.5281/zenodo.12774677" httpUri="https://zenodo.org/record/12774677/files/figure.png" pageId="21" pageNumber="76">Fig. 18F</figureCitation>
). Hind femur 4.8–5.3 (HT: 5.3) times as long as maximum depth in lateral view. Hind TS I: II: III: IV: V = 2.0: 0.9–0.95 (HT: 0.9): 0.6: 0.25–0.3 (HT: 0.3): 0.4–0.5 (HT: 0.4). Tarsal claws simple.
</paragraph>
<caption id="DF2F660D1B51A700FC9861E0FC7D7C6F" ID-DOI="http://doi.org/10.5281/zenodo.12774675" ID-Zenodo-Dep="12774675" httpUri="https://zenodo.org/record/12774675/files/figure.png" pageId="20" pageNumber="75" startId="20.[850,884,2003,2025]" targetBox="[189,813,1563,2098]" targetPageId="20" targetType="figure">
<paragraph id="8BEF36851B51A700FC9861E0FC7D7C6F" blockId="20.[850,1436,2003,2088]" pageId="20" pageNumber="75">
Fig. 17.
<taxonomicName id="4C504D061B51A700FC6961E0FB3973AE" authority="Kusigemati, 1985" authorityName="Kusigemati" authorityYear="1985" box="[931,1262,2003,2025]" class="Insecta" family="Ichneumonidae" genus="Retalia" kingdom="Animalia" order="Hymenoptera" pageId="20" pageNumber="75" phylum="Arthropoda" rank="species" species="japonica">
<emphasis id="B924EA971B51A700FC6961E0FBE973AE" box="[931,1086,2003,2025]" italics="true" pageId="20" pageNumber="75">Retalia japonica</emphasis>
<bibRefCitation id="EFC14B741B51A700FB8F61E0FB3973AE" author="Kusigemati, K." box="[1093,1262,2003,2025]" pageId="20" pageNumber="75" pagination="220 - 228" refId="ref57169" refString="Kusigemati, K., 1985. Three new species of Retalia Seyrig from Formosa and Japan (Hymenoptera: Ichneumonidae). Memoirs of the Kagoshima University, Research Center for the South Pacific, 6: 220 - 228." type="journal article" year="1985">Kusigemati, 1985</bibRefCitation>
</taxonomicName>
, KPM-NK 81677, female from Japan ― A, lateral habitus; B, head, frontal view.
</paragraph>
</caption>
<paragraph id="8BEF36851B50A701FF65648EFB507583" blockId="21.[151,738,155,958]" lastBlockId="21.[813,1400,155,958]" pageId="21" pageNumber="76">
Metasoma. T I slender, 2.35–2.6 (HT: 2.6) times as long as maximum width, largely longitudinally striated except for smooth posterior margin, its sternite ending at far behind of spiracle, its spiracle situated near the basal 0.45. Median dorsal carina of T I present except for posterior part absent. Dorsolateral carina of T I complete. T II to T VI wider than long. T II rugose-punctate anterior 0.75 except for smooth area around thyridium, posterior 0.25 smooth. Laterotergite of T II narrow and separated from T II. T III rugose-punctate except for posterior 0.25 smooth. T IV to T VII covered with fine punctures except for smooth posterior margin. Apex of T VIII protruding and its apex subtruncate. Ovipositor sheath 1.43–1.5 (HT: 1.5) times as long as hind tibia. Ovipositor straight, with a weak nodus and ventral teeth (
<figureCitation id="136B2A001B50A701FBEA679FFBAC7583" box="[1056,1147,428,452]" captionStart="Fig" captionStartId="75.[151,184,1580,1602]" captionTargetBox="[161,1381,163,1549]" captionTargetId="figure-5@75.[151,1398,151,1553]" captionTargetPageId="75" captionText="Fig. 66. Acrolyta japonica sp. nov. (A, KPM-NK 54996, holotype), Micraris ryukyuensis sp. nov. (B, KPM-NK 81873, holotype), Surculus japonicus sp. nov. (C, KPM-NK 81861, holotype), Bentyra ryukyuana sp. nov. (D, KPM-NK 81887, holotype), Paraphylax elegans sp. nov. (E, KPM-NK 55078, holotype), Pa. politus sp. nov. (F, KPM-NK 55079, holotype), Pa. transstriatus sp. nov. (G, KPM-NK 55080, holotype), Pa.yakushimensis sp. nov. (H, KPM-NK 55081, holotype), Pa. yambarensis sp. nov. (I, KPM-NK 55085, holotype), Hemiteles japonicus sp. nov. (J, KPM-NK 55021, holotype),H. kuro sp. nov. (K, KPM-NK 55024, holotype), H.maculipterus sp. nov. (L, KPM-NK 55025, holotype), H. yamatonis sp. nov. (M, KPM-NK 55032, holotype), Brachypimpla latipetiolator (Uchida, 1935) (N, O, KPM-NK 81973), Isadelphus nigrus sp. nov. (P, KPM-NK 82003, holotype), Lochetica japonica sp. nov. (Q, KPM-NK 81989, holotype), Mastrus molestae (Uchida, 1933) (R, KPM-NK 81987), Megacara similis Sheng, 1999 (S, KPM-NK 81835), Tropistes shimizui sp. nov. (T, KPM-NK 81833, holotype), and Uchidella toichii sp. nov. (U, KPM-NK 82011, holotype), females from Japan ― A-N, P-U, apical part of ovipositor, lateral view (A, C-G, I-K, N, Q-U, left side; B, H, L, M, P, right side); O, apex of ovipositor sheath, lateral view." figureDoi="http://doi.org/10.5281/zenodo.12774789" httpUri="https://zenodo.org/record/12774789/files/figure.png" pageId="21" pageNumber="76">Fig. 66C</figureCitation>
).
</paragraph>
<paragraph id="8BEF36851B50A702FC8F67E0FE077531" blockId="21.[813,1400,155,958]" lastBlockId="22.[189,776,155,841]" lastPageId="22" lastPageNumber="77" pageId="21" pageNumber="76">
Coloration (
<figureCitation id="136B2A001B50A701FC0A67E0FB9275AC" box="[960,1093,467,491]" captionStart="Fig" captionStartId="21.[153,186,1985,2007]" captionTargetBox="[151,1398,998,1961]" captionTargetId="figure-368@21.[151,1398,996,1961]" captionTargetPageId="21" captionText="Fig. 18. Surculus japonicus sp. nov., KPM-NK 81861, holotype, female ― A, lateral habitus; B, head, frontal view; C, head, mesosoma, and metasoma, dorsal view; D, head and mesosoma, lateral view; E, head, mesoscutum and scutellum, dorsal view; F, wings; G, propodeum,dorsal view." figureDoi="http://doi.org/10.5281/zenodo.12774677" httpUri="https://zenodo.org/record/12774677/files/figure.png" pageId="21" pageNumber="76">Figs 18A–G</figureCitation>
). Body (excluding wings and legs) black to blackish brown. Scape, pedicel, and base of FL I reddish brown (but dorsal part of scape usually darkened). Mandible yellowish brown except for darkened apex. Palpi and tegula yellow. Metasomal tergites more or less tinged with reddish brown. T I except for posterior margin and apex of metasoma usually black or darkened. T II sometimes partly darkened medially. Membranous part of metasomal sternites yellowish brown. Sclerotized part of metasomal sternites dark brown. Ovipositor reddish brown. Wings hyaline. Veins and pterostigma brown to blackish brown. Fore and mid legs whitish yellow to yellowish brown. Mid femur, tibia, and tarsus partly tinged with blackish brown. Hind coxa black, sometimes narrowly tinged with yellowish brown. Hind trochanter and trochantellus whitish yellow. Hind femur, tibia, and tarsus reddish brown to dark reddish brown. Base and apical parts of hind tibia and tarsus more or less darkened. Tibial spurs whitish yellow.
</paragraph>
<caption id="DF2F660D1B50A701FF5361F2FE4B7C51" ID-DOI="http://doi.org/10.5281/zenodo.12774677" ID-Zenodo-Dep="12774677" httpUri="https://zenodo.org/record/12774677/files/figure.png" pageId="21" pageNumber="76" startId="21.[153,186,1985,2007]" targetBox="[151,1398,998,1961]" targetPageId="21" targetType="figure">
<paragraph id="8BEF36851B50A701FF5361F2FE4B7C51" blockId="21.[153,1401,1985,2070]" pageId="21" pageNumber="76">
Fig. 18.
<taxonomicName id="4C504D061B50A701FF2C61F2FE457391" authorityName="Watanabe" authorityYear="2021" box="[230,402,1985,2006]" class="Insecta" family="Ichneumonidae" genus="Surculus" kingdom="Animalia" order="Hymenoptera" pageId="21" pageNumber="76" phylum="Arthropoda" rank="species" species="japonicus" status="sp. nov.">
<emphasis id="B924EA971B50A701FF2C61F2FE457391" box="[230,402,1985,2006]" italics="true" pageId="21" pageNumber="76">Surculus japonicus</emphasis>
</taxonomicName>
<emphasis id="B924EA971B50A701FE5D61F2FE377391" bold="true" box="[407,480,1985,2006]" pageId="21" pageNumber="76">
<taxonomicNameLabel id="A21757EC1B50A701FE5D61F2FE377391" box="[407,480,1985,2006]" pageId="21" pageNumber="76" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, KPM-NK 81861, holotype, female ― A, lateral habitus; B, head, frontal view; C, head, mesosoma, and metasoma, dorsal view; D, head and mesosoma, lateral view; E, head, mesoscutum and scutellum, dorsal view; F, wings; G, propodeum, dorsal view.
</paragraph>
</caption>
<paragraph id="8BEF36851B53A702FF1F67B6FE5775DA" blockId="22.[189,776,155,841]" box="[213,384,389,413]" pageId="22" pageNumber="77">Male. Unknown.</paragraph>
</subSubSection>
<subSubSection id="C34A650E1B53A702FF1F6798FDD07583" box="[213,519,427,452]" pageId="22" pageNumber="77" type="distribution">
<paragraph id="8BEF36851B53A702FF1F6798FDD07583" blockId="22.[189,776,155,841]" box="[213,519,427,452]" pageId="22" pageNumber="77">
<emphasis id="B924EA971B53A702FF1F6798FE807584" bold="true" box="[213,343,427,451]" pageId="22" pageNumber="77">Distribution</emphasis>
.
<collectingCountry id="F34776151B53A702FEA9679FFE4A7584" box="[355,413,428,451]" name="Japan" pageId="22" pageNumber="77">Japan</collectingCountry>
(Honshu).
</paragraph>
</subSubSection>
<subSubSection id="C34A650E1B53A702FF1F67E1FE6C75AC" box="[213,443,466,491]" pageId="22" pageNumber="77" type="biology_ecology">
<paragraph id="8BEF36851B53A702FF1F67E1FE6C75AC" blockId="22.[189,776,155,841]" box="[213,443,466,491]" pageId="22" pageNumber="77">
<emphasis id="B924EA971B53A702FF1F67E1FE9375AD" bold="true" box="[213,324,466,490]" pageId="22" pageNumber="77">Bionomics</emphasis>
. Unknown.
</paragraph>
</subSubSection>
<subSubSection id="C34A650E1B53A702FF1F67C9FEB3770E" pageId="22" pageNumber="77" type="discussion">
<paragraph id="8BEF36851B53A702FF1F67C9FEB3770E" blockId="22.[189,776,155,841]" pageId="22" pageNumber="77">
<emphasis id="B924EA971B53A702FF1F67C9FEEA7655" bold="true" box="[213,317,506,530]" pageId="22" pageNumber="77">Remarks</emphasis>
. This species resembles
<taxonomicName id="4C504D061B53A702FD8567C9FD0B7655" authorityName="Townes" authorityYear="1970" box="[591,732,506,530]" class="Insecta" family="Ichneumonidae" genus="Surculus" kingdom="Animalia" order="Hymenoptera" pageId="22" pageNumber="77" phylum="Arthropoda" rank="species" species="oculatus">
<emphasis id="B924EA971B53A702FD8567C9FD0B7655" box="[591,732,506,530]" italics="true" pageId="22" pageNumber="77">Sur. oculatus</emphasis>
</taxonomicName>
but it can be distinguished by the nervellus intercepted near the middle (near the posterior end in
<taxonomicName id="4C504D061B53A702FD83647BFD077618" authorityName="Townes" authorityYear="1970" box="[585,720,583,607]" class="Insecta" family="Ichneumonidae" genus="Surculus" kingdom="Animalia" order="Hymenoptera" pageId="22" pageNumber="77" phylum="Arthropoda" rank="species" species="oculatus">
<emphasis id="B924EA971B53A702FD83647BFD077618" box="[585,720,583,607]" italics="true" pageId="22" pageNumber="77">Sur. oculatus</emphasis>
</taxonomicName>
), the sternaulus absent in posterior 0.4 of mesopleuron (nearly complete in
<taxonomicName id="4C504D061B53A702FEF564A5FE1376EA" authorityName="Townes" authorityYear="1970" box="[319,452,661,685]" class="Insecta" family="Ichneumonidae" genus="Surculus" kingdom="Animalia" order="Hymenoptera" pageId="22" pageNumber="77" phylum="Arthropoda" rank="species" species="oculatus">
<emphasis id="B924EA971B53A702FEF564A5FE1376EA" box="[319,452,661,685]" italics="true" pageId="22" pageNumber="77">Sur. oculatus</emphasis>
</taxonomicName>
), the anterior transverse carina of propodeum complete (absent medially in
<taxonomicName id="4C504D061B53A702FDB2648EFD2F7693" authorityName="Townes" authorityYear="1970" box="[632,760,700,724]" class="Insecta" family="Ichneumonidae" genus="Surculus" kingdom="Animalia" order="Hymenoptera" pageId="22" pageNumber="77" phylum="Arthropoda" rank="species" species="oculatus">
<emphasis id="B924EA971B53A702FDB2648EFD2F7693" box="[632,760,700,724]" italics="true" pageId="22" pageNumber="77">Sur. oculatus</emphasis>
</taxonomicName>
), the area basalis of propodeum present (absent in
<taxonomicName id="4C504D061B53A702FD1764D7FEC67765" authorityName="Townes" authorityYear="1970" class="Insecta" family="Ichneumonidae" genus="Surculus" kingdom="Animalia" order="Hymenoptera" pageId="22" pageNumber="77" phylum="Arthropoda" rank="species" species="oculatus">
<emphasis id="B924EA971B53A702FD1764D7FEC67765" italics="true" pageId="22" pageNumber="77">Sur. oculatus</emphasis>
</taxonomicName>
), and the vein 3rs-m of fore wing absent (present in
<taxonomicName id="4C504D061B53A702FF1C6502FE82770E" authorityName="Townes" authorityYear="1970" box="[214,341,817,841]" class="Insecta" family="Ichneumonidae" genus="Surculus" kingdom="Animalia" order="Hymenoptera" pageId="22" pageNumber="77" phylum="Arthropoda" rank="species" species="oculatus">
<emphasis id="B924EA971B53A702FF1C6502FE82770E" box="[214,341,817,841]" italics="true" pageId="22" pageNumber="77">Sur. oculatus</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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