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<document id="F55026A74547FF36565FAD98767ECE18" ID-CLB-Dataset="6279" ID-DOI="10.5852/ejt.2021.751.1381" ID-GBIF-Dataset="4e3af683-d8a6-40de-991b-c4584d66b8cc" ID-ISSN="2118-9773" ID-Zenodo-Dep="4906007" ID-ZooBank="1D27EA41-2396-463A-B1B2-0CCF79C9F3D2" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1623068023717" checkinUser="felipe" docAuthor="Zamani, Alireza, Chatzaki, Maria, Esyunin, Sergei L. &amp; Marusik, Yuri M." docDate="2021" docId="03F987850365EF7D1C447CB5FC286E5A" docLanguage="en" docName="ejt-751_68-114.pdf" docOrigin="European Journal of Taxonomy 751 (1)" docSource="http://dx.doi.org/10.5852/ejt.2021.751.1381" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Nomisia ameretatae Zamani &amp; Chatzaki &amp; Esyunin &amp; Marusik 2021, sp. nov." docType="treatment" docUuid="91468CB5-29A9-40E4-9224-18814812AC2C" docUuidSource="ZooBank" docVersion="9" lastPageNumber="96" masterDocId="FFC0FFFD037FEF611E347F73FFC16D41" masterDocTitle="One new genus and nineteen new species of ground spiders (Araneae: Gnaphosidae) from Iran, with other taxonomic considerations" masterLastPageNumber="114" masterPageNumber="68" pageNumber="94" updateTime="1699000119853" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="EF03C88637E8F16CF406F1B6C07B083B">One new genus and nineteen new species of ground spiders (Araneae: Gnaphosidae) from Iran, with other taxonomic considerations</mods:title>
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<mods:namePart id="81FD20F561D29150594A0B59FAE8531E">Zamani, Alireza</mods:namePart>
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<mods:affiliation id="0CF2512CE228CC6A5ECAE023AA8AA680">Zoological Museum, Biodiversity Unit, University of Turku, FI- 20014, Finland. Department of Molecular Biology and Genetics, Democritus University of Thrace, Dragana, 68100 Alexandroupolis, Greece. Perm State University, Bukireva Street 15, Perm, 614600, Russia. Institute for Biological Problems of the North RAS, Portovaya Str. 18, Magadan, 685000, Russia. Department of Zoology &amp; Entomology, University of the Free State, Bloemfontein 9300, South Africa.</mods:affiliation>
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<mods:namePart id="72706CAC38E4633D5E25CE76A623CA2E">Chatzaki, Maria</mods:namePart>
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<mods:namePart id="9453B199B2963B826AD19F3DA2E8CC64">Esyunin, Sergei L.</mods:namePart>
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<mods:namePart id="93817F777682F381DD4B7799F0487155">Marusik, Yuri M.</mods:namePart>
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<heading id="D0A781FF0365EF7B1C447CB5FC056EA0" box="[624,964,966,993]" centered="true" fontSize="11" level="2" pageId="26" pageNumber="94" reason="2">
<taxonomicName id="4C504D100365EF7B1C447CB5FCA36EA0" ID-CoL="98YW2" authority="Zamani &amp; Chatzaki &amp; Esyunin &amp; Marusik, 2021" authorityName="Zamani &amp; Chatzaki &amp; Esyunin &amp; Marusik" authorityYear="2021" box="[624,866,966,993]" class="Arachnida" family="Gnaphosidae" genus="Nomisia" kingdom="Animalia" order="Araneae" pageId="26" pageNumber="94" phylum="Arthropoda" rank="species" species="ameretatae" status="sp. nov.">
<emphasis id="B924EA810365EF7B1C447CB5FCA36EA0" bold="true" box="[624,866,966,993]" italics="true" pageId="26" pageNumber="94">Nomisia ameretatae</emphasis>
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<taxonomicNameLabel id="A21757FA0365EF7B1D5D7CB4FC056EA0" box="[873,964,967,993]" pageId="26" pageNumber="94" rank="species">sp. nov.</taxonomicNameLabel>
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urn:lsid:zoobank.org:act:
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<paragraph id="8BEF36930365EF7B1CC47B7DFC826968" blockId="26.[375,1212,966,1065]" box="[752,835,1038,1065]" pageId="26" pageNumber="94">
<figureCitation id="136B2A160365EF7B1CC47B7DFC826968" box="[752,835,1038,1065]" captionStart="Fig" captionStartId="27.[189,232,1892,1918]" captionTargetBox="[230,1367,266,1854]" captionTargetId="figure-14@27.[220,1367,265,1854]" captionTargetPageId="27" captionText="Fig. 17. Nomisia ameretatae sp. nov., holotype, ♂ (MMUE). AB. Habitus, dorsal and ventral views. CD. Bulb, dorsal and retrolateral views. EG. Palp, ventral, retroventral and retrolateral views. H. Chelicera, posterior view. IK. Tibial apophysis, dorsal, retrolateral and ventral views. Scale bars: 0.2 mm, unless stated otherwise." figureDoi="http://doi.org/10.5281/zenodo.4906057" httpUri="https://zenodo.org/record/4906057/files/figure.png" pageId="26" pageNumber="94">Fig. 17</figureCitation>
</paragraph>
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<paragraph id="8BEF36930365EF7B1E897B22FEF7692A" blockId="26.[189,310,1105,1131]" box="[189,310,1105,1131]" pageId="26" pageNumber="94">
<heading id="D0A781FF0365EF7B1E897B22FEF7692A" bold="true" box="[189,310,1105,1131]" fontSize="11" level="3" pageId="26" pageNumber="94" reason="3">
<emphasis id="B924EA810365EF7B1E897B22FEF7692A" bold="true" box="[189,310,1105,1131]" pageId="26" pageNumber="94">Diagnosis</emphasis>
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The new species is most similar to
<taxonomicName id="4C504D100365EF7B1C6F7BF2FBAA69DB" authority="(O. Pickard-Cambridge, 1872)" baseAuthorityName="O. Pickard-Cambridge" baseAuthorityYear="1872" box="[603,1131,1152,1178]" class="Arachnida" family="Gnaphosidae" genus="Nomisia" kingdom="Animalia" order="Araneae" pageId="26" pageNumber="94" phylum="Arthropoda" rank="species" species="ripariensis">
<emphasis id="B924EA810365EF7B1C6F7BF2FD3C69DB" box="[603,765,1153,1178]" italics="true" pageId="26" pageNumber="94">N. ripariensis</emphasis>
(O. Pickard-Cambridge, 1872)
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and
<taxonomicName id="4C504D100365EF7B1A927BF2FF3869FF" authority="Chatzaki, 2010" authorityName="Chatzaki" authorityYear="2010" class="Arachnida" family="Gnaphosidae" genus="Echemus" kingdom="Animalia" order="Araneae" pageId="26" pageNumber="94" phylum="Arthropoda" rank="species" species="levyi">
<emphasis id="B924EA810365EF7B1A927BF2FB3E69DB" box="[1190,1279,1152,1178]" italics="true" pageId="26" pageNumber="94">N. levyi</emphasis>
<bibRefCitation id="EFC14B620365EF7B1B337BF3FF3869FF" author="Chatzaki M." pageId="26" pageNumber="94" pagination="1 - 22" refId="ref17040" refString="Chatzaki M. 2010. A revision of the genus Nomisia in Greece and neighboring regions with the description of two new species. Zootaxa 2501 (1): 1 - 22. https: // doi. org / 10.11646 / zootaxa. 2501.1.1" type="journal article" year="2010">Chatzaki, 2010</bibRefCitation>
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by having a similar tibial apophysis with a triangular protrusion and the tegulum extending prolaterally. It differs from them by the embolic division composed of a pike-like, straight embolus and a subtriangular terminal apophysis behind it (vs broad at base of embolus and with a claw-like terminal apophysis in
<taxonomicName id="4C504D100365EF7B1F627A62FE37686B" baseAuthorityName="O. Pickard-Cambridge" baseAuthorityYear="1872" box="[342,502,1297,1322]" class="Arachnida" family="Gnaphosidae" genus="Nomisia" kingdom="Animalia" order="Araneae" pageId="26" pageNumber="94" phylum="Arthropoda" rank="species" species="ripariensis">
<emphasis id="B924EA810365EF7B1F627A62FE37686B" box="[342,502,1297,1322]" italics="true" pageId="26" pageNumber="94">N. ripariensis</emphasis>
</taxonomicName>
, see
<bibRefCitation id="EFC14B620365EF7B1C057A63FD1E686A" author="Chatzaki M." box="[561,735,1296,1323]" pageId="26" pageNumber="94" pagination="1 - 22" refId="ref17040" refString="Chatzaki M. 2010. A revision of the genus Nomisia in Greece and neighboring regions with the description of two new species. Zootaxa 2501 (1): 1 - 22. https: // doi. org / 10.11646 / zootaxa. 2501.1.1" type="journal article" year="2010">Chatzaki 2010</bibRefCitation>
: figs 4748, and small and vertical embolus connected to a long, transverse terminal apophysis in
<taxonomicName id="4C504D100365EF7B1CA27A46FD31680E" authorityName="Chatzaki" authorityYear="2010" box="[662,752,1333,1359]" class="Arachnida" family="Gnaphosidae" genus="Echemus" kingdom="Animalia" order="Araneae" pageId="26" pageNumber="94" phylum="Arthropoda" rank="species" species="levyi">
<emphasis id="B924EA810365EF7B1CA27A46FD31680E" box="[662,752,1333,1359]" italics="true" pageId="26" pageNumber="94">N. levyi</emphasis>
</taxonomicName>
, see
<bibRefCitation id="EFC14B620365EF7B1D1F7A46FC1B680E" author="Chatzaki M." box="[811,986,1333,1359]" pageId="26" pageNumber="94" pagination="1 - 22" refId="ref17040" refString="Chatzaki M. 2010. A revision of the genus Nomisia in Greece and neighboring regions with the description of two new species. Zootaxa 2501 (1): 1 - 22. https: // doi. org / 10.11646 / zootaxa. 2501.1.1" type="journal article" year="2010">Chatzaki 2010</bibRefCitation>
: figs 2829).
</paragraph>
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<emphasis id="B924EA810365EF7B1E897A04FE8568D0" bold="true" box="[189,324,1399,1425]" pageId="26" pageNumber="94">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="8BEF36930365EF7B1E897AD5FD2D68A5" blockId="26.[189,1398,1446,1509]" pageId="26" pageNumber="94">The specific epithet is a noun in the genitive case and refers to Ameretat, the Avestan language name of the Zoroastrian divinity concept of immortality.</paragraph>
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<emphasis id="B924EA810365EF7B1E89797EFEAD6B67" bold="true" box="[189,364,1548,1575]" pageId="26" pageNumber="94">Type material</emphasis>
</heading>
</paragraph>
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<typeStatus id="54EB88310365EF7B1EDE794FFE9D6B17" box="[234,348,1596,1622]" pageId="26" pageNumber="94" type="holotype">Holotype</typeStatus>
</emphasis>
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;
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,
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;
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;
<collectorName id="26A553450365EF7B1B087912FEDE6BDF" pageId="26" pageNumber="94">Y.M. Marusik</collectorName>
leg.;
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.
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</paragraph>
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<emphasis id="B924EA810365EF7B1E8979B5FE8F6BA1" bold="true" box="[189,334,1734,1760]" pageId="26" pageNumber="94">Description</emphasis>
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</paragraph>
<paragraph id="8BEF36930365EF7B1EDE7985FEEB6A51" blockId="26.[187,1399,1782,2025]" box="[234,298,1782,1808]" pageId="26" pageNumber="94">
<emphasis id="B924EA810365EF7B1EDE7985FEEB6A51" bold="true" box="[234,298,1782,1808]" pageId="26" pageNumber="94">Male</emphasis>
</paragraph>
<paragraph id="8BEF36930365EF7D1E897869FE076C2D" blockId="26.[187,1399,1782,2025]" lastBlockId="28.[188,1400,265,365]" lastPageId="28" lastPageNumber="96" pageId="26" pageNumber="94">
Habitus as in
<figureCitation id="136B2A160365EF7B1F627869FE206A75" box="[342,481,1818,1844]" captionStart="Fig" captionStartId="27.[189,232,1892,1918]" captionTargetBox="[230,1367,266,1854]" captionTargetId="figure-14@27.[220,1367,265,1854]" captionTargetPageId="27" captionText="Fig. 17. Nomisia ameretatae sp. nov., holotype, ♂ (MMUE). AB. Habitus, dorsal and ventral views. CD. Bulb, dorsal and retrolateral views. EG. Palp, ventral, retroventral and retrolateral views. H. Chelicera, posterior view. IK. Tibial apophysis, dorsal, retrolateral and ventral views. Scale bars: 0.2 mm, unless stated otherwise." figureDoi="http://doi.org/10.5281/zenodo.4906057" httpUri="https://zenodo.org/record/4906057/files/figure.png" pageId="26" pageNumber="94">Fig. 17AB</figureCitation>
. Total length 7.30. Carapace 3.38 long, 2.45 wide. Sternum 1.74 long, 1.60 wide. Eye sizes and interdistance of PMEs: AME = 0.12, ALE = 0.13, PME = 0.15, PLE = 0.13, PMEPME = 0.08. Carapace, sternum, labium, chelicerae and maxillae dark reddish brown; carapace with numerous dark lines radiating from fovea to submarginal area. Chelicera (
<figureCitation id="136B2A160365EF7B1DB978F5FC356AE1" box="[909,1012,1926,1952]" captionStart="Fig" captionStartId="27.[189,232,1892,1918]" captionTargetBox="[230,1367,266,1854]" captionTargetId="figure-14@27.[220,1367,265,1854]" captionTargetPageId="27" captionText="Fig. 17. Nomisia ameretatae sp. nov., holotype, ♂ (MMUE). AB. Habitus, dorsal and ventral views. CD. Bulb, dorsal and retrolateral views. EG. Palp, ventral, retroventral and retrolateral views. H. Chelicera, posterior view. IK. Tibial apophysis, dorsal, retrolateral and ventral views. Scale bars: 0.2 mm, unless stated otherwise." figureDoi="http://doi.org/10.5281/zenodo.4906057" httpUri="https://zenodo.org/record/4906057/files/figure.png" pageId="26" pageNumber="94">Fig. 17H</figureCitation>
) with serrated retromarginal keel. Legs light brown, with annulations. Abdomen light gray, dorsally with distinct darker pattern medially and marginally, ventrally with two light brown tracheal marks. Anterior spinnerets dark reddish brown, slightly lighter in the apex. Posterior spinnerets slightly lighter than abdomen. Measurements of legs: I: 13.78 (3.48, 1.87, 3.16, 3.25, 2.02), II: missing, III: 8.32 (2.22, 1.17, 1.45, 2.10, 1.42), IV: 11.74 (3.15, 1.44, 2.50, 3.24, 1.41).
</paragraph>
<caption id="DF2F661B0364EF7A1E897817FDFC6AA9" ID-DOI="http://doi.org/10.5281/zenodo.4906057" ID-Zenodo-Dep="4906057" httpUri="https://zenodo.org/record/4906057/files/figure.png" pageId="27" pageNumber="95" startId="27.[189,232,1892,1918]" targetBox="[230,1367,266,1854]" targetPageId="27">
<paragraph id="8BEF36930364EF7A1E897817FDFC6AA9" blockId="27.[189,1399,1892,2024]" pageId="27" pageNumber="95">
<emphasis id="B924EA810364EF7A1E897817FEDA6A3F" bold="true" box="[189,283,1892,1918]" pageId="27" pageNumber="95">Fig. 17.</emphasis>
<taxonomicName id="4C504D100364EF7A1F117816FDD36A3F" authority="Zamani &amp; Chatzaki &amp; Esyunin &amp; Marusik, 2021" authorityName="Zamani &amp; Chatzaki &amp; Esyunin &amp; Marusik" authorityYear="2021" box="[293,530,1893,1918]" class="Arachnida" family="Gnaphosidae" genus="Nomisia" kingdom="Animalia" order="Araneae" pageId="27" pageNumber="95" phylum="Arthropoda" rank="species" species="ameretatae" status="sp. nov.">
<emphasis id="B924EA810364EF7A1F117816FDD36A3F" box="[293,530,1893,1918]" italics="true" pageId="27" pageNumber="95">Nomisia ameretatae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21757FA0364EF7A1C2F7816FDB96A3E" box="[539,632,1893,1919]" pageId="27" pageNumber="95" rank="species">sp. nov.</taxonomicNameLabel>
, holotype, ♂ (MMUE).
<emphasis id="B924EA810364EF7A1DAE7817FC136A3F" bold="true" box="[922,978,1892,1918]" pageId="27" pageNumber="95">AB</emphasis>
. Habitus, dorsal and ventral views.
<emphasis id="B924EA810364EF7A1E8A78F4FF366AE0" bold="true" box="[190,247,1927,1953]" pageId="27" pageNumber="95">CD</emphasis>
. Bulb, dorsal and retrolateral views.
<emphasis id="B924EA810364EF7A1CF878F4FCC66AE0" bold="true" box="[716,775,1927,1953]" pageId="27" pageNumber="95">EG</emphasis>
. Palp, ventral, retroventral and retrolateral views.
<emphasis id="B924EA810364EF7A1E8978D8FF156A84" bold="true" box="[189,212,1963,1989]" pageId="27" pageNumber="95">H</emphasis>
. Chelicera, posterior view.
<emphasis id="B924EA810364EF7A1C2A78D8FD8F6A84" bold="true" box="[542,590,1963,1989]" pageId="27" pageNumber="95">IK</emphasis>
. Tibial apophysis, dorsal, retrolateral and ventral views. Scale bars: 0.2 mm, unless stated otherwise.
</paragraph>
</caption>
<paragraph id="8BEF36930363EF7D1E897EE6FB3B6F00" blockId="28.[189,1399,405,577]" pageId="28" pageNumber="96">
Palp as in
<figureCitation id="136B2A160363EF7D1F0C7EE6FDC56CEE" box="[312,516,405,431]" captionStart="Fig" captionStartId="27.[189,232,1892,1918]" captionTargetBox="[230,1367,266,1854]" captionTargetId="figure-14@27.[220,1367,265,1854]" captionTargetPageId="27" captionText="Fig. 17. Nomisia ameretatae sp. nov., holotype, ♂ (MMUE). AB. Habitus, dorsal and ventral views. CD. Bulb, dorsal and retrolateral views. EG. Palp, ventral, retroventral and retrolateral views. H. Chelicera, posterior view. IK. Tibial apophysis, dorsal, retrolateral and ventral views. Scale bars: 0.2 mm, unless stated otherwise." figureDoi="http://doi.org/10.5281/zenodo.4906057" httpUri="https://zenodo.org/record/4906057/files/figure.png" pageId="28" pageNumber="96">Fig. 17CG, IK</figureCitation>
. Tibia with small, membranous retroventral apophysis and well sclerotized retrolateral one; retrolateral apophysis shorter than tibia, broader in central part than at the base, tip rounded, central part with conical process; cymbium 2.3 times as long as wide; tegulum with massive protrusion directed prolaterally; terminal apophysis subtriangular, well sclerotized, parallel to embolus, arising behind tegulum; embolus pike-like, oblique, directed at an almost 2 oclock position.
</paragraph>
<paragraph id="8BEF36930363EF7D1EDE7D19FE846FC5" blockId="28.[189,325,618,680]" box="[234,325,618,644]" pageId="28" pageNumber="96">
<emphasis id="B924EA810363EF7D1EDE7D19FE846FC5" bold="true" box="[234,325,618,644]" pageId="28" pageNumber="96">Female</emphasis>
</paragraph>
<paragraph id="8BEF36930363EF7D1E897DFDFEF86FE9" blockId="28.[189,325,618,680]" box="[189,313,654,680]" pageId="28" pageNumber="96">Unknown.</paragraph>
</subSubSection>
<subSubSection id="C34A65180363EF7D1E897DA2FC286E5A" box="[189,1001,721,795]" pageId="28" pageNumber="96" type="distribution">
<paragraph id="8BEF36930363EF7D1E897DA2FE976FAA" blockId="28.[189,342,721,747]" box="[189,342,721,747]" pageId="28" pageNumber="96">
<heading id="D0A781FF0363EF7D1E897DA2FE976FAA" bold="true" box="[189,342,721,747]" fontSize="11" level="3" pageId="28" pageNumber="96" reason="3">
<emphasis id="B924EA810363EF7D1E897DA2FE976FAA" bold="true" box="[189,342,721,747]" pageId="28" pageNumber="96">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="8BEF36930363EF7D1E897C72FC286E5A" blockId="28.[189,1001,769,795]" box="[189,1001,769,795]" pageId="28" pageNumber="96">
Known only from the
<typeStatus id="54EB88310363EF7D1FF67C72FE346E5A" box="[450,501,769,795]" pageId="28" pageNumber="96">type</typeStatus>
locality in
<collectingRegion id="4994F8710363EF7D1C4C7C72FCFD6E5A" box="[632,828,769,795]" country="Iran" name="Tehran" pageId="28" pageNumber="96">Tehran Province</collectingRegion>
, northern
<collectingCountry id="F34776030363EF7D1D867C72FC246E5A" box="[946,997,769,795]" name="Iran" pageId="28" pageNumber="96">Iran</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>