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<document id="5D6AA77ED1DB95997BB83397683BE138" ID-DOI="10.1051/acarologia/20152184" ID-ISSN="2107-7207" ID-Zenodo-Dep="5401498" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630334644620" checkinUser="marcus" docAuthor="Miko, L." docDate="2015" docId="03F887C4ED48826A8C6CD64D47F0F917" docLanguage="en" docName="Acarologia.2015.55.4.449-457.pdf" docOrigin="Acarologia 55 (4)" docSource="http://dx.doi.org/10.1051/acarologia/20152184" docStyle="DocumentStyle:8E0DCBB23C2CB3FD20935B7794B24F85.3:Acarologia.2010-2017.journal_article" docStyleId="8E0DCBB23C2CB3FD20935B7794B24F85" docStyleName="Acarologia.2010-2017.journal_article" docStyleVersion="3" docTitle="Dissorhina paleokrasica Miko 2015, n. sp." docType="treatment" docVersion="3" lastPageNumber="452" masterDocId="FFC1FFBCED4A826F8D7BD20B470AFFBC" masterDocTitle="Oribatid mite fossils from pre-Quaternary sediments in Slovenian caves III. Two new species of Dissorhina (Oppiidae) from the Pliocene" masterLastPageNumber="457" masterPageNumber="449" pageNumber="450" updateTime="1699232562429" updateUser="plazi" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-ND-4.0">
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<mods:title id="81CEC1892190CCE5AE9C31C96F0BC6A4">Oribatid mite fossils from pre-Quaternary sediments in Slovenian caves III. Two new species of Dissorhina (Oppiidae) from the Pliocene</mods:title>
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<mods:namePart id="1F79876B1B439116A2E1F2CF1B502788">Miko, L.</mods:namePart>
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<subSubSection id="C34B6559ED48826D8C6CD64D4588FBE2" box="[279,642,1094,1118]" pageId="2" pageNumber="450" type="nomenclature">
<paragraph id="8BEE36D2ED48826D8C6CD64D4588FBE2" blockId="2.[234,687,1094,1153]" box="[279,642,1094,1118]" pageId="2" pageNumber="450">
<heading id="D0A681BEED48826D8C6CD64D4588FBE2" bold="true" box="[279,642,1094,1118]" centered="true" fontSize="10" level="2" pageId="2" pageNumber="450" reason="2">
<emphasis id="B925EAC0ED48826D8C6CD64D4588FBE2" bold="true" box="[279,642,1094,1118]" pageId="2" pageNumber="450">
<taxonomicName id="4C514D51ED48826D8C6CD64D4533FBE2" authority="Miko, 2015" authorityName="Miko" authorityYear="2015" box="[279,569,1094,1118]" class="Arachnida" family="Oppiidae" genus="Dissorhina" kingdom="Animalia" order="Sarcoptiformes" pageId="2" pageNumber="450" phylum="Arthropoda" rank="species" species="paleokrasica" status="sp. nov.">
<emphasis id="B925EAC0ED48826D8C6CD64D4533FBE2" bold="true" box="[279,569,1094,1118]" italics="true" pageId="2" pageNumber="450">Dissorhina paleokrasica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21657BBED48826D8F44D64D4588FBE2" box="[575,642,1094,1118]" pageId="2" pageNumber="450" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
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<subSubSection id="C34B6559ED48826A8D91D66247F0F917" lastPageId="5" lastPageNumber="453" pageId="2" pageNumber="450" type="description">
<paragraph id="8BEE36D2ED48826D8D91D66245A5FB3D" blockId="2.[234,687,1094,1153]" box="[234,687,1129,1153]" pageId="2" pageNumber="450">
<emphasis id="B925EAC0ED48826D8D91D66245A5FB3D" bold="true" box="[234,687,1129,1153]" pageId="2" pageNumber="450">
(
<figureCitation id="136A2A57ED48826D8D8AD6624654FB3D" box="[241,350,1129,1153]" captionStart="FIGURE" captionStartId="3.[159,171,1243,1262]" captionTargetBox="[212,1343,346,1120]" captionTargetId="figure-6@3.[159,1444,302,1211]" captionTargetPageId="3" captionText="FIGURE 1: Dissorhina paleokrasica n. sp.: A dorsal view of holotype; B dorsal view of paratype; C detail of the prodorsum of paratype. Scale bar: A, B 100 µm, C 50 µm. Acronyms: bo bothridium, cos lamellar costula, ctr central tooth of rostrum, ibt interbothridial tubercle, lc lateral carina, ltr lateral tooth of rostrum, pbt postbothridial tubercle, pt I pedotectum I, pt II pedotectum II, ss sensillus. Setae are labeled by name." figureDoi="http://doi.org/10.5281/zenodo.5401500" httpUri="https://zenodo.org/record/5401500/files/figure.png" pageId="2" pageNumber="450">Figures 1</figureCitation>
A-C, 2A-E, 4A-G, and 5D-E)
</emphasis>
</paragraph>
<paragraph id="8BEE36D2ED48826D8DE4D6AE46F0FA33" blockId="2.[159,762,1189,1948]" pageId="2" pageNumber="450">
Diagnosis —
<taxonomicName id="4C514D51ED48826D8C29D6AE46C2FB01" authorityName="Hull" authorityYear="1916" box="[338,456,1189,1213]" class="Arachnida" family="Oppiidae" genus="Dissorhina" kingdom="Animalia" order="Sarcoptiformes" pageId="2" pageNumber="450" phylum="Arthropoda" rank="genus">
<emphasis id="B925EAC0ED48826D8C29D6AE46C2FB01" box="[338,456,1189,1213]" italics="true" pageId="2" pageNumber="450">Dissorhina</emphasis>
</taxonomicName>
(see
<bibRefCitation id="EFC04B23ED48826D8F61D6AE45A0FB01" author="Miko L." box="[538,682,1189,1213]" pageId="2" pageNumber="450" pagination="263 - 296" refId="ref3854" refString="Miko L. 2006 - Oppiidae Grandjean, 1951 - In: Weigmann G. Hornmilben (Oribatida). Keltern: Goecke &amp; Evers. p 263 - 296" type="book chapter" year="2006">Miko, 2006</bibRefCitation>
) with broadly pentagonal rostrum, fine and rather straight lamellar costula. Interbothridial tubercles small and not fused with anterior border of notogaster. Notogaster in axial part protruding anteriad and notogastral as well as interlamellar setae with about 3-5 bristles unilaterally.
</paragraph>
<paragraph id="8BEE36D2ED48826D8DBED7AD47E8F9D1" blockId="2.[159,762,1189,1948]" pageId="2" pageNumber="450">
Measurements — Available specimens partly crushed, not allowing precise measurements. Body length of
<typeStatus id="54EA8870ED48826D8C6BD7E74672F9B8" box="[272,376,1516,1540]" pageId="2" pageNumber="450" type="holotype">holotype</typeStatus>
about 300 µm, length of prodorsum about 100 µm, maximum notogaster width about 150 µm (actual values for damaged specimen).
</paragraph>
<paragraph id="8BEE36D2ED48826D8DBED48F454FF8F7" blockId="2.[159,762,1189,1948]" pageId="2" pageNumber="450">General characters — Original colour difficult to estimate, available fossil material pale, yellowish brown, partly covered with debris and dirt, making observation of finer structure difficult. Cuticle smooth as far as possible to observe, prodorsum laterally with granular microstructure.</paragraph>
<paragraph id="8BEE36D2ED48826D8DBED56A44F1FA68" blockId="2.[159,762,1189,1948]" lastBlockId="2.[826,1429,311,1492]" pageId="2" pageNumber="450">
Prodorsum — Prodorsum relatively narrow, delicate. Rostrum with two incisions as usual for
<taxonomicName id="4C514D51ED48826D8FB1D58E4483FEF3" authorityName="Hull" authorityYear="1916" box="[714,905,311,1948]" class="Arachnida" family="Oppiidae" genus="Dissorhina" kingdom="Animalia" order="Sarcoptiformes" pageId="2" pageNumber="450" phylum="Arthropoda" rank="genus">
<emphasis id="B925EAC0ED48826D8FB1D58E4483FEF3" box="[714,905,311,1948]" italics="true" pageId="2" pageNumber="450">Dissorhina</emphasis>
</taxonomicName>
, central tooth broadly pentagonal, blunt anteriorly, bearing a pair of rostral setae, inserted close to each other. Lateral teeth of rostrum short, broad, with blunt, rounded tips. Pedotectum I weakly developed as triangular projection with rounded tip, only slightly projecting over prodorsal outline. Another blunt projection visible on the place of pedotectum II. Only proximal part of lamellar costulae developed (
<emphasis id="B925EAC0ED48826D8977D05B4321FDDB" box="[1036,1067,592,615]" italics="true" pageId="2" pageNumber="450">i.e.</emphasis>
reaching from bothridium to prodorsal groove and not beyond), costulae finely developed, oblique, straight or only slightly bent. Bothridia relatively small, rounded, with no or only indistinct posterior lobe. Postbothridial tubercle separated from bothridia, developed as small round tubercle. Interbothridial tubercles rather small and fine, not fused with anterior border of notogaster, their mutual distance same or smaller than distance between them and bothridia. Longitudinal, laterally rounded and flat &quot;ear-like&quot; carina developed in lateral part of prodorsum, above insertion of leg II. Sensillus moderately long (65 70 µm), lanceiform, smooth, stalk about one half of the length or slightly longer, lanceiform head about 2,5 times broader than stalk. Prodorsal setae of different lengths and forms. Rostral setae rather fine, slightly bent or straight, smooth or with one bristle, about 27 µm long. Lamellar setae shortest, fine, straight, about 15 µm long. Interlamellar setae stronger, straight, slightly over 30 µm long with about 5-6 small bristles unilaterally, better visible in lateral view. Exobothridial setae not observed, their insertions nevertheless positioned on small tubercles. Another small tubercle may be present in close vicinity to them.
</paragraph>
<paragraph id="8BEE36D2ED48826D8E1BD7FA424CF8DB" blockId="2.[826,1429,1521,1896]" pageId="2" pageNumber="450">
Notogaster — Broadly oval in shape, with anterior border protruding anteriad, darker in transmitted light. Notogastral crista present, short, bearing short setae
<emphasis id="B925EAC0ED48826D8EBAD45144D0F9CE" box="[961,986,1626,1650]" italics="true" pageId="2" pageNumber="450">c1</emphasis>
. Other notogastral setae longer, fine, attenuated towards the end, adhering to the surface. Setae
<emphasis id="B925EAC0ED48826D8EFAD4AB4496F904" box="[897,924,1696,1720]" italics="true" pageId="2" pageNumber="450">h1</emphasis>
inserted closely to each other, distally divergent. Not all setae observable on available material, but at least setae
<emphasis id="B925EAC0ED48826D8928D4ED436CF942" box="[1107,1126,1766,1790]" italics="true" pageId="2" pageNumber="450">la</emphasis>
,
<emphasis id="B925EAC0ED48826D890DD4ED4381F942" box="[1142,1163,1766,1790]" italics="true" pageId="2" pageNumber="450">lp</emphasis>
,
<emphasis id="B925EAC0ED48826D89E2D4ED43BEF942" box="[1177,1204,1766,1790]" italics="true" pageId="2" pageNumber="450">h2</emphasis>
and
<emphasis id="B925EAC0ED48826D898AD4ED4201F942" box="[1265,1291,1766,1790]" italics="true" pageId="2" pageNumber="450">h3</emphasis>
with about 3-4 fine bristles unilaterally. Setae
<emphasis id="B925EAC0ED48826D89BFD50243D5F89D" box="[1220,1247,1801,1825]" italics="true" pageId="2" pageNumber="450">h1</emphasis>
and
<emphasis id="B925EAC0ED48826D886DD501423BF89E" box="[1302,1329,1802,1826]" italics="true" pageId="2" pageNumber="450">p1</emphasis>
smooth, without bristles. Approximate length of setae
<emphasis id="B925EAC0ED48826D8824D5264278F8F9" box="[1375,1394,1837,1861]" italics="true" pageId="2" pageNumber="450">la</emphasis>
21 µm,
<emphasis id="B925EAC0ED48826D8E15D55B4489F8D4" box="[878,899,1872,1896]" italics="true" pageId="2" pageNumber="450">lp</emphasis>
29 µm,
<emphasis id="B925EAC0ED48826D8EA5D55B44F3F8D4" box="[990,1017,1872,1896]" italics="true" pageId="2" pageNumber="450">h1</emphasis>
and
<emphasis id="B925EAC0ED48826D8948D55B4347F8D4" box="[1075,1101,1872,1896]" italics="true" pageId="2" pageNumber="450">h2</emphasis>
32 µm and
<emphasis id="B925EAC0ED48826D89ADD55B43FBF8D4" box="[1238,1265,1872,1896]" italics="true" pageId="2" pageNumber="450">p1</emphasis>
19 µm.
</paragraph>
<paragraph id="8BEE36D2ED48826C8E1BD58F46CBF8A0" blockId="2.[864,1428,1924,1948]" lastBlockId="3.[159,762,1410,1820]" lastPageId="3" lastPageNumber="451" pageId="2" pageNumber="450">
Ventral characters — Infracapitulum as usual in
<taxonomicName id="4C514D51ED49826C8DE4D789461FFA26" authorityName="Hull" authorityYear="1916" box="[159,277,1410,1434]" class="Arachnida" family="Oppiidae" genus="Dissorhina" kingdom="Animalia" order="Sarcoptiformes" pageId="3" pageNumber="451" phylum="Arthropoda" rank="genus">
<emphasis id="B925EAC0ED49826C8DE4D789461FFA26" box="[159,277,1410,1434]" italics="true" pageId="3" pageNumber="451">Dissorhina</emphasis>
</taxonomicName>
, setae of infracapitulum and palps fine and smooth. Epimeral setation 3-1-3-3, epimeral setae straight, smooth, about 11 12 µm long,
<emphasis id="B925EAC0ED49826C8F9AD7C345F0FA5C" box="[737,762,1480,1504]" italics="true" pageId="3" pageNumber="451">1c</emphasis>
and
<emphasis id="B925EAC0ED49826C8DAFD7E047E6F9BF" box="[212,236,1515,1539]" italics="true" pageId="3" pageNumber="451">3c</emphasis>
and
<emphasis id="B925EAC0ED49826C8C52D7E0465CF9BF" box="[297,342,1515,1539]" italics="true" pageId="3" pageNumber="451">4b-c</emphasis>
being the longest. Trochanteral setae I and II very long, bent, with few short bristles. Epimeral area with several lighter maculae in central part. Discidium developed as small, blunt tubercle. Anogenital formula as usual (
<emphasis id="B925EAC0ED49826C8F1CD4734588F933" box="[615,642,1656,1679]" italics="true" pageId="3" pageNumber="451">
<quantity id="4CA99B37ED49826C8F1CD4734588F933" box="[615,642,1656,1679]" metricMagnitude="-3" metricUnit="kg" metricValue="5.0" pageId="3" pageNumber="451" unit="g" value="5.0">5g</quantity>
</emphasis>
,
<emphasis id="B925EAC0ED49826C8FE8D47345B0F92C" box="[659,698,1656,1680]" italics="true" pageId="3" pageNumber="451">1ag</emphasis>
,
<emphasis id="B925EAC0ED49826C8FB0D47345F9F92C" box="[715,755,1656,1680]" italics="true" pageId="3" pageNumber="451">2an</emphasis>
,
<emphasis id="B925EAC0ED49826C8DE4D49047CCF90F" box="[159,198,1691,1715]" italics="true" pageId="3" pageNumber="451">3ad</emphasis>
), setae smooth and narrow, mostly a bit longer than epimeral ones (
<emphasis id="B925EAC0ED49826C8CEDD4B546BBF96A" box="[406,433,1726,1750]" italics="true" pageId="3" pageNumber="451">g1</emphasis>
8 µm,
<emphasis id="B925EAC0ED49826C8F73D4B54528F96A" box="[520,546,1726,1750]" italics="true" pageId="3" pageNumber="451">g6</emphasis>
13 µm,
<emphasis id="B925EAC0ED49826C8FFDD4B54595F969" box="[646,671,1726,1749]" italics="true" pageId="3" pageNumber="451">ag</emphasis>
14 µm,
<emphasis id="B925EAC0ED49826C8DE4D4EA47CDF945" box="[159,199,1761,1785]" italics="true" pageId="3" pageNumber="451">an2</emphasis>
15 µm). Adanal setae longer,
<emphasis id="B925EAC0ED49826C8F56D4EA455EF945" box="[557,596,1761,1785]" italics="true" pageId="3" pageNumber="451">ad1</emphasis>
posterior,
<emphasis id="B925EAC0ED49826C8FA8D4EA45F0F945" box="[723,762,1761,1785]" italics="true" pageId="3" pageNumber="451">ad3</emphasis>
anterior to anal opening.
</paragraph>
<caption id="DF2E665AED49826C8DE4D6D045C7FAFD" ID-DOI="http://doi.org/10.5281/zenodo.5401500" ID-Zenodo-Dep="5401500" httpUri="https://zenodo.org/record/5401500/files/figure.png" pageId="3" pageNumber="451" startId="3.[159,171,1243,1262]" targetBox="[212,1343,346,1120]" targetPageId="3">
<paragraph id="8BEE36D2ED49826C8DE4D6D045C7FAFD" blockId="3.[159,1429,1242,1345]" pageId="3" pageNumber="451">
FIGURE 1:
<taxonomicName id="4C514D51ED49826C8C77D6D146D3FB51" authority="Miko, 2015" authorityName="Miko" authorityYear="2015" box="[268,473,1242,1261]" class="Arachnida" family="Oppiidae" genus="Dissorhina" kingdom="Animalia" order="Sarcoptiformes" pageId="3" pageNumber="451" phylum="Arthropoda" rank="species" species="paleokrasica" status="sp. nov.">
<emphasis id="B925EAC0ED49826C8C77D6D146D3FB51" box="[268,473,1242,1261]" italics="true" pageId="3" pageNumber="451">Dissorhina paleokrasica</emphasis>
</taxonomicName>
<emphasis id="B925EAC0ED49826C8C9AD6D14516FB51" bold="true" box="[481,540,1242,1261]" pageId="3" pageNumber="451">
<taxonomicNameLabel id="A21657BBED49826C8C9AD6D14516FB51" box="[481,540,1242,1261]" pageId="3" pageNumber="451" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
: A dorsal view of holotype; B dorsal view of paratype; C detail of the prodorsum of paratype. Scale bar: A, B 100 µm, C 50 µm. Acronyms: bo bothridium, cos lamellar costula, ctr central tooth of rostrum, ibt interbothridial tubercle, lc lateral carina, ltr lateral tooth of rostrum, pbt postbothridial tubercle, pt I pedotectum I, pt II pedotectum II, ss sensillus. Setae are labeled by name.
</paragraph>
</caption>
<paragraph id="8BEE36D2ED49826C8DBED53443ACF945" blockId="3.[159,762,1854,1948]" lastBlockId="3.[826,1429,1410,1785]" pageId="3" pageNumber="451">
Legs — One of the two individuals (
<typeStatus id="54EA8870ED49826C8FFCD53545FBF8EA" box="[647,753,1854,1878]" pageId="3" pageNumber="451" type="holotype">holotype</typeStatus>
) was relatively well preserved, with almost all legs present, even if incomplete. Due to the risk of further destruction, only leg I was investigated. As far as possible to observe, leg setation formula corresponds to those of
<taxonomicName id="4C514D51ED49826C896FD7C34380FA5C" authorityName="Hull" authorityYear="1916" box="[1044,1162,1480,1504]" class="Arachnida" family="Oppiidae" genus="Dissorhina" kingdom="Animalia" order="Sarcoptiformes" pageId="3" pageNumber="451" phylum="Arthropoda" rank="genus">
<emphasis id="B925EAC0ED49826C896FD7C34380FA5C" box="[1044,1162,1480,1504]" italics="true" pageId="3" pageNumber="451">Dissorhina</emphasis>
</taxonomicName>
as usual
<emphasis id="B925EAC0ED49826C898FD7C24219FA5C" box="[1268,1299,1481,1504]" italics="true" pageId="3" pageNumber="451">i.e.</emphasis>
setation of leg I: 1-5-2(1)-4(2)-20(2). Setae moderately long, at least some of them with several short bristles. Tarsal solenidia
<emphasis id="B925EAC0ED49826C8ED7D43A44B7F9F5" box="[940,957,1585,1609]" italics="true" pageId="3" pageNumber="451">ω</emphasis>
<subScript id="17D53497ED49826C8EC5D43044CDF9F0" attach="left" box="[958,967,1595,1612]" fontSize="7" pageId="3" pageNumber="451">1</subScript>
and
<emphasis id="B925EAC0ED49826C897ED43A431CF9F5" box="[1029,1046,1585,1609]" italics="true" pageId="3" pageNumber="451">ω</emphasis>
<subScript id="17D53497ED49826C896CD430432AF9F0" attach="left" box="[1047,1056,1595,1612]" fontSize="7" pageId="3" pageNumber="451">2</subScript>
of about same length,
<emphasis id="B925EAC0ED49826C8849D43A4249F9F5" box="[1330,1347,1585,1609]" italics="true" pageId="3" pageNumber="451">ω</emphasis>
<subScript id="17D53497ED49826C883FD4304247F9F0" attach="left" box="[1348,1357,1595,1612]" fontSize="7" pageId="3" pageNumber="451">1</subScript>
blunt and bent towards the tarsal surface. Tibial solenidion
<emphasis id="B925EAC0ED49826C8E00D4734486F92C" box="[891,908,1656,1680]" italics="true" pageId="3" pageNumber="451"></emphasis>
<subScript id="17D53497ED49826C8EF6D48A449CF92E" attach="left" box="[909,918,1665,1682]" fontSize="7" pageId="3" pageNumber="451">1</subScript>
setiform, longer than the length of tibia. Solenidion
<emphasis id="B925EAC0ED49826C8EBAD49044D8F90F" box="[961,978,1691,1715]" italics="true" pageId="3" pageNumber="451"></emphasis>
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much shorter, blunt. Genual solenidion
<emphasis id="B925EAC0ED49826C8E02D4B54482F96A" box="[889,904,1726,1750]" italics="true" pageId="3" pageNumber="451">σ</emphasis>
long, setiform, more than two times longer than the length of the segment.
</paragraph>
<paragraph id="8BEE36D2ED49826C8E1BD510429FF820" blockId="3.[826,1429,1819,1949]" pageId="3" pageNumber="451">
Material examined — Two individuals were available from the clastic sediments of Trhlovca cave in Karst,
<collectingCountry id="F3467642ED49826C8E99D56A4340F8C5" box="[994,1098,1889,1913]" name="Slovenia" pageId="3" pageNumber="451">Slovenia</collectingCountry>
. Individual from sample T5 from Pliocene red clay sediments (about 3,8 millions
</paragraph>
<paragraph id="8BEE36D2ED4E826B8DE4D2F147E3FEB1" blockId="4.[159,233,250,269]" box="[159,233,250,269]" pageId="4" pageNumber="452">Miko L.</paragraph>
<caption id="DF2E665AED4E826B8DE4D6D04299FAB6" ID-DOI="http://doi.org/10.5281/zenodo.5401502" ID-Zenodo-Dep="5401502" httpUri="https://zenodo.org/record/5401502/files/figure.png" pageId="4" pageNumber="452" startId="4.[159,171,1243,1262]" targetBox="[159,1444,302,1211]" targetPageId="4">
<paragraph id="8BEE36D2ED4E826B8DE4D6D04299FAB6" blockId="4.[159,1429,1242,1290]" pageId="4" pageNumber="452">
FIGURE 2:
<taxonomicName id="4C514D51ED4E826B8C7FD6D146C4FB51" authority="Miko, 2015" authorityName="Miko" authorityYear="2015" box="[260,462,1242,1261]" class="Arachnida" family="Oppiidae" genus="Dissorhina" kingdom="Animalia" order="Sarcoptiformes" pageId="4" pageNumber="452" phylum="Arthropoda" rank="species" species="paleokrasica" status="sp. nov.">
<emphasis id="B925EAC0ED4E826B8C7FD6D146C4FB51" box="[260,462,1242,1261]" italics="true" pageId="4" pageNumber="452">Dissorhina paleokrasica</emphasis>
</taxonomicName>
<emphasis id="B925EAC0ED4E826B8CA8D6D1450DFB51" bold="true" box="[467,519,1242,1261]" pageId="4" pageNumber="452">
<taxonomicNameLabel id="A21657BBED4E826B8CA8D6D1450DFB51" box="[467,519,1242,1261]" pageId="4" pageNumber="452" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
: A ventral view; B sensillus; C prodorsal setae; D notogastral setae; E leg I. Scale bar: A 100 µm, BE 50 µm. Acronyms: tr III trochanter of leg III, tr IV trochanter of leg IV. Setae and lyrifissurae are labelled by name.
</paragraph>
</caption>
<paragraph id="8BEE36D2ED4E826B8DE4D74145A0F8F1" blockId="4.[159,762,1354,1949]" pageId="4" pageNumber="452">
<materialsCitation id="3B393C8FED4E826B8DE4D74145ACF8F1" collectionCode="T" country="Romania" county="Emil Racovita Institute of Speleology" location="Pliocene" municipality="Individuals" pageId="4" pageNumber="452" specimenCode="T2" specimenCount="2" stateProvince="Cluj" typeStatus="holotype">
of years old; see
<bibRefCitation id="EFC04B23ED4E826B8C27D7414545FADF" author="Moldovan O. T. &amp; Mihevc A. &amp; Miko L. &amp; Constantin S. &amp; Meleg I. N. &amp; Petculescu A. &amp; Bosak P." box="[348,591,1354,1379]" pageId="4" pageNumber="452" pagination="1825 - 1837" refId="ref4027" refString="Moldovan O. T., Mihevc A., Miko L., Constantin S., Meleg I. N., Petculescu A., Bosak P. 2011 - Invertebrate fossils from cave sediments: a new proxy for pre-Quaternary paleoenvironments - Biogeosciences, 8, 1825 - 1837. Available from: http: // dx. doi. org / 10.5194 / bg- 8 - 1825 - 2011 doi: 10.5194 / bg- 8 - 1825 - 2011" type="journal article" year="2011">
Moldovan
<emphasis id="B925EAC0ED4E826B8CA0D7404504FADE" box="[475,526,1354,1378]" italics="true" pageId="4" pageNumber="452">et al.</emphasis>
, 2011
</bibRefCitation>
for further details) was much better preserved, and is designed as
<typeStatus id="54EA8870ED4E826B8DC4D79A4620FA15" box="[191,298,1425,1449]" pageId="4" pageNumber="452" type="holotype">holotype</typeStatus>
. Individual (
<typeStatus id="54EA8870ED4E826B8CBED79A453AFA15" box="[453,560,1425,1449]" pageId="4" pageNumber="452" type="paratype">paratype</typeStatus>
) from sample
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from
<location id="8E8E6009ED4E826B8D9BD7BF4649FA70" LSID="urn:lsid:plazi:treatment:03F887C4ED48826A8C6CD64D47F0F917:8E8E6009ED4E826B8D9BD7BF4649FA70" box="[224,323,1460,1484]" country="Romania" county="Emil Racovita Institute of Speleology" municipality="Individuals" name="Pliocene" pageId="4" pageNumber="452" stateProvince="Cluj">Pliocene</location>
beige clay sediments (about 2,7 million years old) was damaged, in poor condition, without legs and practically without observable setation.
<location id="8E8E6009ED4E826B8D8DD4164673F989" LSID="urn:lsid:plazi:treatment:03F887C4ED48826A8C6CD64D47F0F917:8E8E6009ED4E826B8D8DD4164673F989" box="[246,377,1565,1589]" country="Romania" county="Emil Racovita Institute of Speleology" municipality="Individuals" name="Notogaster" pageId="4" pageNumber="452" stateProvince="Cluj">Notogaster</location>
was largely damaged and dirty, not allowing more detailed study.
<location id="8E8E6009ED4E826B8F47D44B459CF9E4" LSID="urn:lsid:plazi:treatment:03F887C4ED48826A8C6CD64D47F0F917:8E8E6009ED4E826B8F47D44B459CF9E4" box="[572,662,1600,1624]" country="Romania" county="Emil Racovita Institute of Speleology" municipality="Individuals" name="Specific" pageId="4" pageNumber="452" stateProvince="Cluj">Specific</location>
identity of the two individuals is judged on the basis of the same development of the rostrum, bothridia and lamellar costula, interbothridial and postbohridial tubercles and lateral part of prodorsum, and due to its occurrence in the same locality.
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are preserved in the collection of
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in
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,
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</materialsCitation>
.
</paragraph>
<paragraph id="8BEE36D2ED4E826B8DBED56A4326FA15" blockId="4.[159,762,1354,1949]" lastBlockId="4.[826,1429,1354,1449]" pageId="4" pageNumber="452">
Derivatio nominis — The name &quot;
<taxonomicName id="4C514D51ED4E826B8F27D56A45FBF8C5" authorityName="Miko" authorityYear="2015" box="[604,753,1889,1913]" class="Arachnida" family="Oppiidae" genus="Dissorhina" kingdom="Animalia" order="Sarcoptiformes" pageId="4" pageNumber="452" phylum="Arthropoda" rank="species" species="paleokrasica">paleokrasica</taxonomicName>
&quot; is derived from &quot;Kras&quot;, the name of the Slovenian geomorphological unit where is the
<typeStatus id="54EA8870ED4E826B8980D7404225FADF" box="[1275,1327,1355,1379]" pageId="4" pageNumber="452">type</typeStatus>
locality, Trhlovca cave. Prefix paleo- refers to the fossil character of the material.
</paragraph>
<paragraph id="8BEE36D2ED4E826B8E1BD7C14387F96B" blockId="4.[826,1429,1482,1751]" pageId="4" pageNumber="452">
Remarks —
<taxonomicName id="4C514D51ED4E826B8E8FD7C143FEFA5E" authority="Miko, 2015" authorityName="Miko" authorityYear="2015" box="[1012,1268,1482,1506]" class="Arachnida" family="Oppiidae" genus="Dissorhina" kingdom="Animalia" order="Sarcoptiformes" pageId="4" pageNumber="452" phylum="Arthropoda" rank="species" species="paleokrasica" status="sp. nov.">
<emphasis id="B925EAC0ED4E826B8E8FD7C143FEFA5E" box="[1012,1268,1482,1506]" italics="true" pageId="4" pageNumber="452">Dissorhina paleokrasica</emphasis>
</taxonomicName>
<emphasis id="B925EAC0ED4E826B8986D7C1424DFA5E" bold="true" box="[1277,1351,1482,1506]" pageId="4" pageNumber="452">
<taxonomicNameLabel id="A21657BBED4E826B8986D7C1424DFA5E" box="[1277,1351,1482,1506]" pageId="4" pageNumber="452" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
bears all important characters of the genus, and is close to the widely distributed and common species
<taxonomicName id="4C514D51ED4E826B8802D41A4475F9F7" baseAuthorityName="Oudemans" baseAuthorityYear="1900" class="Arachnida" family="Oppiidae" genus="Dissorhina" kingdom="Animalia" order="Sarcoptiformes" pageId="4" pageNumber="452" phylum="Arthropoda" rank="species" species="ornata">
<emphasis id="B925EAC0ED4E826B8802D41A4475F9F7" italics="true" pageId="4" pageNumber="452">D. ornata</emphasis>
</taxonomicName>
. Our own studies showed that several characters of
<taxonomicName id="4C514D51ED4E826B8EDCD45C431AF9D2" baseAuthorityName="Oudemans" baseAuthorityYear="1900" box="[935,1040,1623,1646]" class="Arachnida" family="Oppiidae" genus="Dissorhina" kingdom="Animalia" order="Sarcoptiformes" pageId="4" pageNumber="452" phylum="Arthropoda" rank="species" species="ornata">
<emphasis id="B925EAC0ED4E826B8EDCD45C431AF9D2" box="[935,1040,1623,1646]" italics="true" pageId="4" pageNumber="452">D. ornata</emphasis>
</taxonomicName>
may be rather variable (see also figures in
<bibRefCitation id="EFC04B23ED4E826B8ECFD4724336F92D" author="Miko L." box="[948,1084,1657,1681]" pageId="4" pageNumber="452" pagination="263 - 296" refId="ref3854" refString="Miko L. 2006 - Oppiidae Grandjean, 1951 - In: Weigmann G. Hornmilben (Oribatida). Keltern: Goecke &amp; Evers. p 263 - 296" type="book chapter" year="2006">Miko, 2006</bibRefCitation>
), so differences of the newly described fossil species needed to be carefully analysed against that variability.
</paragraph>
<paragraph id="8BEE36D2ED4E826A8E1BD4F247F0F917" blockId="4.[826,1429,1784,1948]" lastBlockId="5.[159,762,1438,1707]" lastPageId="5" lastPageNumber="453" pageId="4" pageNumber="452">
However, the newly described species can be distinguished from all other
<taxonomicName id="4C514D51ED4E826B89E2D5104266F88F" box="[1177,1388,1819,1843]" class="Arachnida" family="Oppiidae" genus="Dissorhina" kingdom="Animalia" order="Sarcoptiformes" pageId="4" pageNumber="452" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B925EAC0ED4E826B89E2D5104205F88F" box="[1177,1295,1819,1843]" italics="true" pageId="4" pageNumber="452">Dissorhina</emphasis>
species
</taxonomicName>
by a combination of some specific characters, namely: (a) interbothridial tubercles are fine, weakly developed and not fused with anterior margin of notogaster (developed as separate tubercles); (b) anterior part of notogaster between setae
<emphasis id="B925EAC0ED4F826A8F11D7CA4589FA65" box="[618,643,1473,1497]" italics="true" pageId="5" pageNumber="453">c2</emphasis>
is arched and protruding anteriad; (c) central tooth of rostrum is broad, bluntly pentagonal or almost quadrangular, not trapezoid as in
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<emphasis id="B925EAC0ED4F826A8C96D4204559F9FE" box="[493,595,1579,1602]" italics="true" pageId="5" pageNumber="453">D. ornata</emphasis>
</taxonomicName>
or some other species; (d) notogastral setae except
<emphasis id="B925EAC0ED4F826A8F26D446457FF9D9" box="[605,629,1613,1637]" italics="true" pageId="5" pageNumber="453">c2</emphasis>
and
<emphasis id="B925EAC0ED4F826A8FCCD44645F3F9D9" box="[695,761,1613,1637]" italics="true" pageId="5" pageNumber="453">p1p3</emphasis>
are quite long, and at least
<emphasis id="B925EAC0ED4F826A8C9FD47B46FDF934" box="[484,503,1648,1672]" italics="true" pageId="5" pageNumber="453">la</emphasis>
,
<emphasis id="B925EAC0ED4F826A8F7CD47B4516F934" box="[519,540,1648,1672]" italics="true" pageId="5" pageNumber="453">lp</emphasis>
and
<emphasis id="B925EAC0ED4F826A8F22D47B457EF934" box="[601,628,1648,1672]" italics="true" pageId="5" pageNumber="453">h2</emphasis>
with small bristles.
</paragraph>
</subSubSection>
</treatment>
</document>