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<mods:title id="0DA3DCAB1F5A7FD87C5744419FE9068E">Terrestrial isopods from the Oued Laou basin, north-eastern Morocco (Crustacea: Oniscidea), with descriptions of two new genera and seven new species</mods:title>
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<mods:namePart id="CD3645981F2C925CB9AB83FE0A814C71">Taiti, Stefano</mods:namePart>
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<treatment id="03F887936B61645839C4FDAEFD27FB07" ID-DOI="http://doi.org/10.5281/zenodo.4337174" ID-GBIF-Taxon="167307718" ID-Zenodo-Dep="4337174" LSID="urn:lsid:plazi:treatment:03F887936B61645839C4FDAEFD27FB07" httpUri="http://treatment.plazi.org/id/03F887936B61645839C4FDAEFD27FB07" lastPageNumber="2129" pageId="62" pageNumber="2129">
<subSubSection id="C34B650E6B61645839C4FDAEFCFCFDCA" box="[493,890,535,559]" pageId="62" pageNumber="2129" type="nomenclature">
<paragraph id="8BEE36856B61645839C4FDAEFCFCFDCA" blockId="62.[493,890,503,559]" box="[493,890,535,559]" pageId="62" pageNumber="2129">
<heading id="D0A681E96B61645839C4FDAEFCFCFDCA" box="[493,890,535,559]" centered="true" fontSize="10" level="2" pageId="62" pageNumber="2129" reason="2">
<taxonomicName id="4C514D066B61645839C4FDAEFCFCFDCA" ID-CoL="GRBY" authority="Dollfus, 1887" authorityName="Dollfus" authorityYear="1887" box="[493,890,535,559]" class="Malacostraca" family="Armadillidiidae" genus="Armadillidium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Isopoda" pageId="62" pageNumber="2129" phylum="Arthropoda" rank="species" species="album">
<emphasis id="B925EA976B61645839C4FDAEFD5EFDCA" bold="true" box="[493,728,535,559]" italics="true" pageId="62" pageNumber="2129">Armadillidium album</emphasis>
<bibRefCitation id="EFC04B746B6164583AC8FDAEFCFCFDCA" author="Dollfus A" box="[737,890,535,559]" pageId="62" pageNumber="2129" pagination="89 - 95" refId="ref20523" refString="Dollfus A. 1887. Diagnoses d' especes nouvelles et catalogue des especes francaises de la tribu des Armadilliens (Crustaces, Isopodes terrestres). Bull Soc Etud Sci Paris. 9: 89 - 95." type="journal article" year="1887">Dollfus, 1887</bibRefCitation>
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<subSubSection id="C34B650E6B61645838FDFDEEFC9EFD8A" box="[212,792,599,623]" pageId="62" pageNumber="2129" type="nomenclature">
<paragraph id="8BEE36856B61645838FDFDEEFC9EFD8A" blockId="62.[212,792,599,623]" box="[212,792,599,623]" pageId="62" pageNumber="2129">
<heading id="D0A681E96B61645838FDFDEEFC9EFD8A" box="[212,792,599,623]" fontSize="10" level="3" pageId="62" pageNumber="2129" reason="5">
<taxonomicName id="4C514D066B61645838FDFDEEFE39FD8A" ID-CoL="GRBY" authorityName="Dollfus" authorityYear="1887" box="[212,447,599,623]" class="Malacostraca" family="Armadillidiidae" genus="Armadillidium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Isopoda" pageId="62" pageNumber="2129" phylum="Arthropoda" rank="species" species="album">
<emphasis id="B925EA976B61645838FDFDEEFE39FD8A" box="[212,447,599,623]" italics="true" pageId="62" pageNumber="2129">Armadillidium album</emphasis>
</taxonomicName>
;
<bibRefCitation id="EFC04B746B61645839F9FDEEFD48FD8A" author="Colombini I &amp; Chaouti A &amp; Fallaci M &amp; Gagnarli E &amp; Scapini F &amp; Bayed A &amp; Chelazzi L" box="[464,718,599,623]" pageId="62" pageNumber="2129" pagination="81 - 91" refId="ref20395" refString="Colombini I, Chaouti A, Fallaci M, Gagnarli E, Scapini F, Bayed A, Chelazzi L. 2008. An assessment of sandy beach macroinvertebrates inhabiting the coastal fringe of the Oued Laou river catchment area (Northern Morocco). In: Bayed A, Ater M, editors. Du bassin versant vers la mer: analyse multidisciplinaire pour une gestion durable. Trav Inst Sci Rabat (Ser Gen). 5: 81 - 91." type="journal article" year="2008">Colombini et al., 2008</bibRefCitation>
, p. 86
</heading>
</paragraph>
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<subSubSection id="C34B650E6B61645838FDFD7DFDE9FC7D" pageId="62" pageNumber="2129" type="materials_examined">
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<emphasis id="B925EA976B61645838FDFD7DFE21FD39" box="[212,423,708,732]" italics="true" pageId="62" pageNumber="2129">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="8BEE36856B61645838FDFD52FB12FCE6" blockId="62.[212,1172,708,772]" box="[212,1172,747,772]" pageId="62" pageNumber="2129">
<materialsCitation id="3B393CD86B61645838FDFD52FB16FCE6" ID-GBIF-Occurrence="2848654407" box="[212,1168,747,772]" collectingDate="2004-05-01" collectionCode="MZUF" collectorName="St. &amp; S. Taiti &amp; C. Rossano" pageId="62" pageNumber="2129" specimenCode="MZUF 9569" specimenCount="3" specimenCount-female="1" specimenCount-male="2">
<specimenCount id="9D57FD0C6B61645838FDFD52FE96FCE6" box="[212,272,747,771]" pageId="62" pageNumber="2129" type="male">2 ♂♂</specimenCount>
,
<specimenCount id="9D57FD0C6B6164583937FD52FEC0FCE6" box="[286,326,747,771]" pageId="62" pageNumber="2129" type="female">1 ♀</specimenCount>
,
<collectorName id="26A453536B616458397DFD52FEF4FCE6" box="[340,370,747,771]" pageId="62" pageNumber="2129">St.</collectorName>
4, under logs, leg.
<collectorName id="26A453536B6164583A6CFD52FD11FCE6" box="[581,663,747,771]" pageId="62" pageNumber="2129">S. Taiti</collectorName>
and
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,
<date id="FFEF10456B6164583B49FD52FC62FCE6" box="[864,996,747,772]" pageId="62" pageNumber="2129" value="2004-05-01">
<collectingDate id="EFABE9AD6B6164583B49FD52FC62FCE6" box="[864,996,747,772]" pageId="62" pageNumber="2129" value="2004-05-01">1 May 2004</collectingDate>
</date>
(
<specimenCode id="DBF79EFE6B6164583BDAFD52FB0FFCE6" box="[1011,1161,747,771]" collectionCode="MZUF" country="Italy" httpUri="http://grbio.org/cool/mvk4-k796" name="Museo Zoologico La Specola, Universita di Firenze" pageId="62" pageNumber="2129">MZUF 9569</specimenCode>
)
</materialsCitation>
.
</paragraph>
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<emphasis id="B925EA976B61645838FDFCE0FE31FC94" box="[212,439,857,881]" italics="true" pageId="62" pageNumber="2129">Previous Rif records</emphasis>
</heading>
</paragraph>
<paragraph id="8BEE36856B61645838FDFC39FDE9FC7D" blockId="62.[212,623,857,920]" box="[212,623,896,920]" pageId="62" pageNumber="2129">
Oued Laou (
<bibRefCitation id="EFC04B746B616458394FFC39FDD8FC7D" author="Colombini I &amp; Chaouti A &amp; Fallaci M &amp; Gagnarli E &amp; Scapini F &amp; Bayed A &amp; Chelazzi L" box="[358,606,896,920]" pageId="62" pageNumber="2129" pagination="81 - 91" refId="ref20395" refString="Colombini I, Chaouti A, Fallaci M, Gagnarli E, Scapini F, Bayed A, Chelazzi L. 2008. An assessment of sandy beach macroinvertebrates inhabiting the coastal fringe of the Oued Laou river catchment area (Northern Morocco). In: Bayed A, Ater M, editors. Du bassin versant vers la mer: analyse multidisciplinaire pour une gestion durable. Trav Inst Sci Rabat (Ser Gen). 5: 81 - 91." type="journal article" year="2008">Colombini et al. 2008</bibRefCitation>
).
</paragraph>
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<subSubSection id="C34B650E6B61645838FDFC54FB0BFBC8" box="[212,1165,1005,1069]" pageId="62" pageNumber="2129" type="distribution">
<paragraph id="8BEE36856B61645838FDFC54FEDEFBE0" blockId="62.[212,1165,1005,1069]" box="[212,344,1005,1029]" pageId="62" pageNumber="2129">
<heading id="D0A681E96B61645838FDFC54FEDEFBE0" box="[212,344,1005,1029]" fontSize="10" level="3" pageId="62" pageNumber="2129" reason="5">
<emphasis id="B925EA976B61645838FDFC54FEDEFBE0" box="[212,344,1005,1029]" italics="true" pageId="62" pageNumber="2129">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="8BEE36856B61645838FDFBACFB0BFBC8" blockId="62.[212,1165,1005,1069]" box="[212,1165,1045,1069]" pageId="62" pageNumber="2129">
Atlantic coast of Europe, western and central Mediterranean as far East as
<collectingCountry id="F34676156B6164583C1EFBACFB0FFBC8" box="[1079,1161,1045,1069]" name="Greece" pageId="62" pageNumber="2129">Greece</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C34B650E6B61645838FDFB3BFD27FB07" pageId="62" pageNumber="2129" type="discussion">
<paragraph id="8BEE36856B61645838FDFB3BFEB0FB7F" blockId="62.[212,1172,1154,1250]" box="[212,310,1154,1178]" pageId="62" pageNumber="2129">
<heading id="D0A681E96B61645838FDFB3BFEB0FB7F" box="[212,310,1154,1178]" fontSize="10" level="3" pageId="62" pageNumber="2129" reason="5">
<emphasis id="B925EA976B61645838FDFB3BFEB0FB7F" box="[212,310,1154,1178]" italics="true" pageId="62" pageNumber="2129">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="8BEE36856B61645838FDFB13FD27FB07" blockId="62.[212,1172,1154,1250]" pageId="62" pageNumber="2129">
For diagnostic characters of
<taxonomicName id="4C514D066B6164583A03FB13FD15FB27" authorityName="Dollfus" authorityYear="1887" box="[554,659,1194,1218]" class="Malacostraca" family="Armadillidiidae" genus="Armadillidium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Isopoda" pageId="62" pageNumber="2129" phylum="Arthropoda" rank="species" species="album">
<emphasis id="B925EA976B6164583A03FB13FD15FB27" box="[554,659,1194,1218]" italics="true" pageId="62" pageNumber="2129">A. album</emphasis>
</taxonomicName>
see
<bibRefCitation id="EFC04B746B6164583AE4FB13FCE1FB27" author="Vandel A" box="[717,871,1194,1218]" pageId="62" pageNumber="2129" pagination="417 - 931" refId="ref22666" refString="Vandel A. 1962. Isopodes terrestres (Deuxieme partie). Faune Fr. 66: 417 - 931." type="journal article" year="1962">Vandel (1962</bibRefCitation>
, p. 841, Figure 403) and
<bibRefCitation id="EFC04B746B61645838FDFB73FE1EFB07" author="Schmalfuss H" box="[212,408,1226,1250]" pageId="62" pageNumber="2129" pagination="153 - 201" refId="ref21590" refString="Schmalfuss H. 2008. The terrestrial isopods (Isopoda: Oniscidea) of Greece. 25 th contribution: the genus Armadillidium (Armadillidiidae) in the provinces Macedonia and Thrace. Stuttg Beitr Naturk, Ser A (N Ser). 1: 153 - 201." type="journal article" year="2008">Schmalfuss (2008</bibRefCitation>
, p. 154,
<figureCitation id="136A2A006B61645839D7FB73FD17FB07" box="[510,657,1226,1250]" captionStart-0="Figure 1" captionStart-1="Figure 2" captionStart-10="Figure 11" captionStart-11="Figure 12" captionStart-2="Figure 3" captionStart-3="Figure 4" captionStart-4="Figure 5" captionStart-5="Figure 6" captionStart-6="Figure 7" captionStart-7="Figure 8" captionStart-8="Figure 9" captionStart-9="Figure 10" captionStartId-0="4.[212,279,1518,1540]" captionStartId-1="5.[144,211,1517,1539]" captionStartId-10="18.[212,279,1512,1534]" captionStartId-11="20.[212,279,1520,1542]" captionStartId-2="7.[144,211,1189,1211]" captionStartId-3="8.[212,279,1509,1531]" captionStartId-4="9.[144,211,1182,1204]" captionStartId-5="11.[144,211,1511,1533]" captionStartId-6="12.[212,279,1524,1546]" captionStartId-7="13.[144,211,1525,1547]" captionStartId-8="16.[212,279,1513,1535]" captionStartId-9="17.[144,211,1379,1401]" captionTargetBox-0="[222,1164,178,1486]" captionTargetBox-1="[144,1104,168,1485]" captionTargetBox-10="[212,1164,178,1475]" captionTargetBox-11="[213,1167,168,1488]" captionTargetBox-2="[144,1104,168,1157]" captionTargetBox-3="[222,1163,174,1458]" captionTargetBox-4="[154,1086,178,1130]" captionTargetBox-5="[162,1096,168,1478]" captionTargetBox-6="[222,1164,178,1468]" captionTargetBox-7="[154,1096,178,1479]" captionTargetBox-8="[222,1164,168,1458]" captionTargetBox-9="[158,1096,168,1347]" captionTargetId-0="figure@4.[212,1172,168,1486]" captionTargetId-1="figure@5.[144,1104,168,1485]" captionTargetId-10="figure@18.[212,1172,168,1480]" captionTargetId-11="figure@20.[212,1172,168,1488]" captionTargetId-2="figure@7.[144,1104,168,1157]" captionTargetId-3="figure@8.[212,1172,168,1477]" captionTargetId-4="figure@9.[144,1104,168,1150]" captionTargetId-5="figure@11.[144,1104,168,1479]" captionTargetId-6="figure@12.[212,1172,168,1492]" captionTargetId-7="figure@13.[144,1104,168,1493]" captionTargetId-8="figure@16.[212,1172,168,1481]" captionTargetId-9="figure@17.[144,1104,168,1347]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionTargetPageId-10="18" captionTargetPageId-11="20" captionTargetPageId-2="7" captionTargetPageId-3="8" captionTargetPageId-4="9" captionTargetPageId-5="11" captionTargetPageId-6="12" captionTargetPageId-7="13" captionTargetPageId-8="16" captionTargetPageId-9="17" captionText-0="Figure 1. Trichoniscus microphthalmus sp. nov. from St. 8, paratype ♀: (A) adult specimen, dorsal view; (B) dorsal scale-seta; (C) cephalon, frontal view; (D) pleonite 5, telson and left uropod; (E) antennule; (F) antenna; (G) right mandible; (H) left mandible." captionText-1="Figure 2. Trichoniscus microphthalmus sp. nov. from St. 8, paratype ♀: (A) maxillule; (B) maxilla; (C) maxilliped. Paratype ♂: (D) pereopod 1; (E) pereopod 7; (F) pleopod 1; (G) pleopod 2." captionText-10="Figure 11. Paractenoscia cavernicola sp. nov. from St. 8, paratype ♂: (A) antenna; (B) pereopod 1; (C) pereopod 7; (D) genital papilla and pleopod 1; (E) pleopod 2; (F) pleopod 3 exopod; (G) pleopod 4 exopod; (H) pleopod 5 exopod." captionText-11="Figure 12. Bathytropa rifensis sp. nov. from St. 15, paratype ♀: (A) adult specimen, dorsal view; (B) dorsal scale-seta; (C) scale-seta on posterior margins of segments; (D) cephalon, dorsal view; (E) cephalon, frontal view; (F) pleonite 5, telson and right uropod; (G) antennule; (H) antenna." captionText-2="Figure 3. Graeconiscus thermophilus from St. 8, ♂: (A) adult specimen, dorsal view; (B) pereonites 6, 7 and pleon, ventral view; (C) pereopod 7." captionText-3="Figure 4. Graeconiscus thermophilus from St. 8, ♀: (A) pleonite 5, telson and uropods; (B) antennule; (C) antenna; (D) right mandible; (E) left mandible; (F) maxillule; (G) maxilla; (H) maxilliped." captionText-4="Figure 5. Graeconiscus thermophilus from St. 8, ♂: (A) pereopod 1; (B) pereopod 7; (C) pleopod 1; (D) pleopod 2." captionText-5="Figure 6. Ctenoscia minima from St. 14, ♀: (A) adult specimen, dorsal view; (B) dorsal scaleseta; (C) co-ordinates of noduli laterals; (D) cephalon, dorsal view; (E) cephalon, frontal view; (F) left side of pereon showing disposition of noduli laterales; (G) pereonite 1, left side; (H) pleonites 4, 5, telson and uropods; (I) antennule." captionText-6="Figure 7. Ctenoscia minima from St. 14, ♀: (A) antenna; (B) left mandible; (C) right mandible; (D) maxillule; (E) maxilla; (F) maxilliped." captionText-7="Figure 8. Ctenoscia minima from St. 14, ♂: (A) pereopod 1; (B) pereopod 7; (C) pleopod 1 (specimen 4.5 mm long); (D) genital papilla and pleopod 1 (specimen 4.0 mm long); (E) pleopod 2; (F) pleopod 3 exopod; (G) pleopod 4 exopod; (H) pleopod 5 exopod." captionText-8="Figure 9. Paractenoscia cavernicola sp. nov. from St. 8, paratype ♀: (A) adult specimen, dorsal view; (B) dorsal scale-seta; (C) co-ordinates of noduli laterals; (D) cephalon, dorsal view; (E) cephalon, frontal view; (F) right side of pereon showing disposition of noduli laterales; (G) pleonites 35, telson and uropods; (H) antennule; (I) antenna." captionText-9="Figure 10. Paractenoscia cavernicola sp. nov. from St. 8, paratype ♀: (A) left mandible; (B) right mandible; (C) maxillule; (D) maxilla; (E) maxilliped." figureDoi-0="http://doi.org/10.5281/zenodo.3999871" figureDoi-1="http://doi.org/10.5281/zenodo.3999873" figureDoi-10="http://doi.org/10.5281/zenodo.3999893" figureDoi-11="http://doi.org/10.5281/zenodo.3999895" figureDoi-2="http://doi.org/10.5281/zenodo.3999875" figureDoi-3="http://doi.org/10.5281/zenodo.3999879" figureDoi-4="http://doi.org/10.5281/zenodo.3999881" figureDoi-5="http://doi.org/10.5281/zenodo.3999883" figureDoi-6="http://doi.org/10.5281/zenodo.3999885" figureDoi-7="http://doi.org/10.5281/zenodo.3999887" figureDoi-8="http://doi.org/10.5281/zenodo.3999889" figureDoi-9="http://doi.org/10.5281/zenodo.3999891" httpUri-0="https://zenodo.org/record/3999871/files/figure.png" httpUri-1="https://zenodo.org/record/3999873/files/figure.png" httpUri-10="https://zenodo.org/record/3999893/files/figure.png" httpUri-11="https://zenodo.org/record/3999895/files/figure.png" httpUri-2="https://zenodo.org/record/3999875/files/figure.png" httpUri-3="https://zenodo.org/record/3999879/files/figure.png" httpUri-4="https://zenodo.org/record/3999881/files/figure.png" httpUri-5="https://zenodo.org/record/3999883/files/figure.png" httpUri-6="https://zenodo.org/record/3999885/files/figure.png" httpUri-7="https://zenodo.org/record/3999887/files/figure.png" httpUri-8="https://zenodo.org/record/3999889/files/figure.png" httpUri-9="https://zenodo.org/record/3999891/files/figure.png" pageId="62" pageNumber="2129">Figures 112</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>