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<document id="A05AAC6D5773DB92637D32A07CEFF4B0" ID-CLB-Dataset="3889" ID-DOI="10.11646/zootaxa.5039.3.5" ID-GBIF-Dataset="f0ac1e8c-b600-4930-94f1-3992327986d4" ID-ISSN="1175-5326" ID-Zenodo-Dep="5511499" ID-ZooBank="A3F75F9C-0E13-409B-B9C5-C4307193242B" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1631776023332" checkinUser="plazi" docAuthor="Zhang, Aihuan" docDate="2021" docId="03F33310DA38FF875DB5FDF3FC2AF8CD" docLanguage="en" docName="zootaxa.5039.3.5.pdf" docOrigin="Zootaxa 5039 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Gibberifera tetracornata Zhang 2021, sp. nov." docType="treatment" docVersion="7" lastPageNumber="411" masterDocId="FFCA4B68DA3AFF855D22FFD6FFAEFF82" masterDocTitle="Review of Gibberifera Obraztsov (Lepidoptera: Tortricidae) from China, with description of a new species" masterLastPageNumber="416" masterPageNumber="409" pageNumber="411" updateTime="1699449204683" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="92911DF82052E3A0789E650057769B66" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="DD9F2568B8F2F821EE5A927FF5E6416F">
<mods:title id="1D49D93DC93338EBDAF6ED6072A9FCBB">Review of Gibberifera Obraztsov (Lepidoptera: Tortricidae) from China, with description of a new species</mods:title>
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<mods:namePart id="ABF6224AFFFAA95A8652307E4DFC38F7">Zhang, Aihuan</mods:namePart>
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<mods:title id="CCDEB62ED0FBDF21BCD2ECAB0AE14101">Zootaxa</mods:title>
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<mods:part id="A072AAC33074A5C5BE0B0A6F07FA95E5">
<mods:date id="34D1E744B81713C2976ED56A42320F1A">2021</mods:date>
<mods:detail id="BAA3DD20DB35C82C98D2F0395824F061" type="pubDate">
<mods:number id="E405A675B8CED9F8992BCC6F26C03590">2021-09-16</mods:number>
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<mods:number id="1B6121642EF769F3D67EEC34600100E6">5039</mods:number>
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<mods:detail id="2013EACAD3ED87ADDACAE3BE3D4C163E" type="issue">
<mods:number id="D95FCF143AC339748DAC59BC9D0B579A">3</mods:number>
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<mods:classification id="F714C4839DEDE9B8A4A073A2236F959E">journal article</mods:classification>
<mods:identifier id="414EB962CFF7198F382FF0636DF7440B" type="CLB-Dataset">3889</mods:identifier>
<mods:identifier id="9430B128D54CD2B681C99BFCE7574026" type="DOI">10.11646/zootaxa.5039.3.5</mods:identifier>
<mods:identifier id="C834FAD9CCB066840954560B6DAA6AEF" type="GBIF-Dataset">f0ac1e8c-b600-4930-94f1-3992327986d4</mods:identifier>
<mods:identifier id="10BFD6D89B3E6D1B99B5290D670785F2" type="ISSN">1175-5326</mods:identifier>
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<treatment id="03F33310DA38FF875DB5FDF3FC2AF8CD" ID-DOI="http://doi.org/10.5281/zenodo.5511501" ID-GBIF-Taxon="189334342" ID-Zenodo-Dep="5511501" LSID="urn:lsid:plazi:treatment:03F33310DA38FF875DB5FDF3FC2AF8CD" httpUri="http://treatment.plazi.org/id/03F33310DA38FF875DB5FDF3FC2AF8CD" lastPageNumber="411" pageId="2" pageNumber="411">
<subSubSection id="C340D18DDA38FF875DB5FDF3FC69FDE6" pageId="2" pageNumber="411" type="nomenclature">
<paragraph id="8BE58206DA38FF875DB5FDF3FD2AFDBD" blockId="2.[151,967,549,647]" box="[151,644,549,576]" pageId="2" pageNumber="411">
<heading id="D0AD356ADA38FF875DB5FDF3FD2AFDBD" bold="true" box="[151,644,549,576]" fontSize="11" level="1" pageId="2" pageNumber="411" reason="1">
<emphasis id="B92E5E14DA38FF875DB5FDF3FD2AFDBD" bold="true" box="[151,644,549,576]" pageId="2" pageNumber="411">
<taxonomicName id="4C5AF985DA38FF875DB5FDF3FDB7FDBD" authority="Zhang" authorityName="Zhang" authorityYear="2021" box="[151,537,549,576]" class="Insecta" family="Tortricidae" genus="Gibberifera" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="411" phylum="Arthropoda" rank="species" species="tetracornata" status="sp. nov.">
<emphasis id="B92E5E14DA38FF875DB5FDF3FE11FDC2" bold="true" box="[151,447,549,576]" italics="true" pageId="2" pageNumber="411">Gibberifera tetracornata</emphasis>
Zhang
</taxonomicName>
,
<taxonomicNameLabel id="A21DE36FDA38FF875F04FDF3FD2AFDBD" box="[550,644,549,575]" pageId="2" pageNumber="411" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BE58206DA38FF875DB5FD9FFC69FDE6" blockId="2.[151,967,549,647]" box="[151,967,585,612]" pageId="2" pageNumber="411">
<uri id="FFCB8E04DA38FF875DB5FD9FFC69FDE6" box="[151,967,585,612]" pageId="2" pageNumber="411">
urn:lsid:zoobank.org:pub:
<uuid id="FFFCB8D3DA38FF875C96FD9FFC69FDE6" box="[436,967,585,612]" pageId="2" pageNumber="411">A3F75F9C-0E13-409B-B9C5-C4307193242B</uuid>
</uri>
</paragraph>
</subSubSection>
<subSubSection id="C340D18DDA38FF875DB5FDBBFEBCFD05" box="[151,274,621,647]" pageId="2" pageNumber="411" type="description">
<paragraph id="8BE58206DA38FF875DB5FDBBFEBCFD05" blockId="2.[151,967,549,647]" box="[151,274,621,647]" pageId="2" pageNumber="411">
(
<figureCitation id="13619E83DA38FF875DBDFDBBFEA7FD05" box="[159,265,621,647]" captionStart="FIGURES 1–3" captionStartId="3.[151,264,1294,1318]" captionTargetBox="[159,1420,186,1260]" captionTargetId="figure-15@3.[151,1436,179,1268]" captionTargetPageId="3" captionText="FIGURES 1–3. Gibberifera tetracornata Zhang, sp. nov. 1. Adult, holotype; 2. Male genitalia (ZAH19003, holotype); 3. Female genitalia (ZAH19030, paratype) (Scale bars: 1 = 2 mm; 2, 3 = 0.5 mm)" figureDoi="http://doi.org/10.5281/zenodo.5511503" httpUri="https://zenodo.org/record/5511503/files/figure.png" pageId="2" pageNumber="411">Figs. 1–3</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C340D18DDA38FF875DB5FD63FDBCFBD9" pageId="2" pageNumber="411" type="diagnosis">
<paragraph id="8BE58206DA38FF875DB5FD63FC74FC49" blockId="2.[151,1437,693,1871]" pageId="2" pageNumber="411">
<emphasis id="B92E5E14DA38FF875DB5FD63FEBCFD4D" bold="true" box="[151,274,693,719]" pageId="2" pageNumber="411">Diagnosis.</emphasis>
This species is similar to
<taxonomicName id="4C5AF985DA38FF875F16FD60FCE9FD52" authority="Kuznetzov" authorityName="Kuznetzov" box="[564,839,693,720]" class="Insecta" family="Tortricidae" genus="Gibberifera" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="411" phylum="Arthropoda" rank="species" species="monticola">
<emphasis id="B92E5E14DA38FF875F16FD60FD68FD4D" box="[564,710,693,720]" italics="true" pageId="2" pageNumber="411">G. monticola</emphasis>
Kuznetzov
</taxonomicName>
, but can be distinguished by darker grey forewing; in the male genitalia by uncus having straight end and vesica with four (arranged in two pairs) non-deciduous spinous cornuti; in the female genitalia by the sterigma which is represented by a wide oblong sclerite, and having the anterior and posterior edge each with a prominence, the four angles are blunt, ductus bursae is longer than the corpus bursae, the corpus bursae is pear-shaped. In
<taxonomicName id="4C5AF985DA38FF875FA7FC90FCB9FCDD" box="[645,791,837,864]" class="Insecta" family="Tortricidae" genus="Gibberifera" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="411" phylum="Arthropoda" rank="species" species="monticola">
<emphasis id="B92E5E14DA38FF875FA7FC90FCB9FCDD" box="[645,791,837,864]" italics="true" pageId="2" pageNumber="411">G. monticola</emphasis>
</taxonomicName>
the forewing is light grey; in the male genitalia the uncus is blunt and round apically and the vesica has two or more (not arranged in pairs) non-deciduous cornuti apically; in the female genitalia the median part and two posterior angles of the sterigma are protuberant, ductus bursae is shorter than the corpus bursae, and the corpus bursae is large and globular.
</paragraph>
<paragraph id="8BE58206DA38FF875DE5FC03FDBCFBD9" blockId="2.[151,1437,693,1871]" pageId="2" pageNumber="411">
This species is also similar to
<taxonomicName id="4C5AF985DA38FF875F33FC00FD7CFC6D" authority="Zhang" authorityName="Zhang &amp; Li" authorityYear="2004" box="[529,722,981,1008]" class="Insecta" family="Tortricidae" genus="Gibberifera" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="411" phylum="Arthropoda" rank="species" species="clavata">
<emphasis id="B92E5E14DA38FF875F33FC00FD2AFC6D" box="[529,644,981,1008]" italics="true" pageId="2" pageNumber="411">G. clavata</emphasis>
Zhang
</taxonomicName>
in appearance, but can be separated by the uncus having straight end, the saccular angle is obtuse and the vesica has four (arranged in two pairs, one pair longer than the other) nondeciduous spinous cornuti. In the latter species the uncus is blunt and round apically, the saccular angle is blunt and non-deciduous cornuti are absent.
</paragraph>
</subSubSection>
<subSubSection id="C340D18DDA38FF875DE5FBB3FD1EF885" pageId="2" pageNumber="411" type="description">
<paragraph id="8BE58206DA38FF875DE5FBB3FAD5F919" blockId="2.[151,1437,693,1871]" pageId="2" pageNumber="411">
<emphasis id="B92E5E14DA38FF875DE5FBB3FEF7FBFD" bold="true" box="[199,345,1125,1151]" pageId="2" pageNumber="411">Description.</emphasis>
Adult (
<figureCitation id="13619E83DA38FF875C93FBB3FE5AFBFD" box="[433,500,1125,1151]" captionStart="FIGURES 1–3" captionStartId="3.[151,264,1294,1318]" captionTargetBox="[159,1420,186,1260]" captionTargetId="figure-15@3.[151,1436,179,1268]" captionTargetPageId="3" captionText="FIGURES 1–3. Gibberifera tetracornata Zhang, sp. nov. 1. Adult, holotype; 2. Male genitalia (ZAH19003, holotype); 3. Female genitalia (ZAH19030, paratype) (Scale bars: 1 = 2 mm; 2, 3 = 0.5 mm)" figureDoi="http://doi.org/10.5281/zenodo.5511503" httpUri="https://zenodo.org/record/5511503/files/figure.png" pageId="2" pageNumber="411">Fig. 1</figureCitation>
).
<emphasis id="B92E5E14DA38FF875F2FFBB3FDE4FBFD" box="[525,586,1125,1151]" italics="true" pageId="2" pageNumber="411">Head</emphasis>
: Vertex greyish brown; frons white. Antenna brown. Labial palpus greyish brown, inner side white; terminal segment slender, drooping.
<emphasis id="B92E5E14DA38FF875E6EFB5FFC08FB21" box="[844,934,1161,1187]" italics="true" pageId="2" pageNumber="411">Thorax:</emphasis>
Thorax and tegula greyish brown. Forewing length 7.5–8.0 mm. Forewing ground color dark grey, termen brown; costa with seven pairs of white strigulae from basal 1/4 to apex; basal patch brown, extending from costal 1/4 to 2/5 of dorsum, occupying about basal 1/4; median fascia dark grey, originating from between costal 1/4 and half, reaching 3/4 of dorsum, outer side darker; ocelloid patch large oval, slivery grey; a small triangular dark spot on dorsum between median fascia and tornus; cilia brown; underside of forewing dark grey. Hindwing and cilia grey; underside of hindwing grey. Foreleg brown, midleg and hindleg grey, with brown scales on tarsi.
<emphasis id="B92E5E14DA38FF875F47FAB7FD61FAF9" box="[613,719,1377,1403]" italics="true" pageId="2" pageNumber="411">Abdomen</emphasis>
: Male genitalia (
<figureCitation id="13619E83DA38FF875EADFAB7FC7AFAF9" box="[911,980,1377,1403]" captionStart="FIGURES 1–3" captionStartId="3.[151,264,1294,1318]" captionTargetBox="[159,1420,186,1260]" captionTargetId="figure-15@3.[151,1436,179,1268]" captionTargetPageId="3" captionText="FIGURES 1–3. Gibberifera tetracornata Zhang, sp. nov. 1. Adult, holotype; 2. Male genitalia (ZAH19003, holotype); 3. Female genitalia (ZAH19030, paratype) (Scale bars: 1 = 2 mm; 2, 3 = 0.5 mm)" figureDoi="http://doi.org/10.5281/zenodo.5511503" httpUri="https://zenodo.org/record/5511503/files/figure.png" pageId="2" pageNumber="411">Fig. 2</figureCitation>
). Uncus elongate club-shaped with truncate apex. Socii broad triangular, hairy. Valva broad at base, with neck as wide as 2/3 of base; saccular angle obtuse; cucullus oblong, bending inward, setose, with a slender process on the ventral angle bearing a short stout spine at apex. Phallus broad and short; vesica with numerous deciduous cornuti sockets and four (arranged in two pairs, one pair longer than the other) non-deciduous, spinous cornuti. Female genitalia (
<figureCitation id="13619E83DA38FF875ED5FA27FB97F989" box="[1015,1081,1521,1547]" captionStart="FIGURES 1–3" captionStartId="3.[151,264,1294,1318]" captionTargetBox="[159,1420,186,1260]" captionTargetId="figure-15@3.[151,1436,179,1268]" captionTargetPageId="3" captionText="FIGURES 1–3. Gibberifera tetracornata Zhang, sp. nov. 1. Adult, holotype; 2. Male genitalia (ZAH19003, holotype); 3. Female genitalia (ZAH19030, paratype) (Scale bars: 1 = 2 mm; 2, 3 = 0.5 mm)" figureDoi="http://doi.org/10.5281/zenodo.5511503" httpUri="https://zenodo.org/record/5511503/files/figure.png" pageId="2" pageNumber="411">Fig. 3</figureCitation>
) with papilla analis narrow and long (dorsal part lost), with some little prominences setose. Apophysis posterioris as long as apophysis anterioris. Sterigma a wide oblong plate, with a prominence on the anterior edge and one on the posterior edge, four blunt angles. Ductus bursae longer than corpus bursae, colliculum a moderately wide sclerotized ring near ostium bursae. Ductus seminalis originating from colliculum. Corpus bursae pear-shaped, with two horned signa, equal in size.
</paragraph>
<paragraph id="8BE58206DA38FF875DE5F973FD1EF885" blockId="2.[151,1437,693,1871]" pageId="2" pageNumber="411">
<materialsCitation id="3B32885BDA38FF875DE5F973FB1BF961" ID-GBIF-Occurrence="3392778306" accessionNumber="ZAH19003" collectingDate="2017-07-19" collectorName="Qi M. J. &amp; Yang X. F." country="China" elevation="2577" latitude="32.97" location="Wanglang National Natural Reserve" longLatPrecision="725" longitude="104.1" municipality="Muyangchang" pageId="2" pageNumber="411" specimenCount="1" specimenCount-male="1" stateProvince="Sichuan Province" typeStatus="holotype">
<typeStatus id="54E13CA4DA38FF875DE5F973FE81F93D" box="[199,303,1701,1727]" pageId="2" pageNumber="411" type="holotype">Holotype</typeStatus>
<specimenCount id="9D5C498FDA38FF875C19F970FEFCF942" box="[315,338,1702,1728]" pageId="2" pageNumber="411" type="male">♂</specimenCount>
,
<emphasis id="B92E5E14DA38FF875C40F973FE66F93D" bold="true" box="[354,456,1701,1727]" pageId="2" pageNumber="411">
<collectingCountry id="F34DC296DA38FF875C40F973FE6AF93D" box="[354,452,1701,1727]" name="China" pageId="2" pageNumber="411">CHINA</collectingCountry>
:
</emphasis>
<collectingMunicipality id="6B81187CDA38FF875CF6F973FDDAF93D" box="[468,628,1701,1727]" pageId="2" pageNumber="411">Muyangchang</collectingMunicipality>
(
<geoCoordinate id="EE6EE4C1DA38FF875FA5F973FD5DF93D" box="[647,755,1701,1727]" degrees="32.97" direction="north" orientation="latitude" pageId="2" pageNumber="411" precision="555" value="32.97">32.97 ºN</geoCoordinate>
,
<geoCoordinate id="EE6EE4C1DA38FF875E20F973FCD9F93D" box="[770,887,1701,1727]" degrees="104.10" direction="east" orientation="longitude" pageId="2" pageNumber="411" precision="555" value="104.1">104.10 ºE</geoCoordinate>
),
<location id="8E85D4DDDA38FF875EADF973FA9DF942" LSID="urn:lsid:plazi:treatment:03F33310DA38FF875DB5FDF3FC2AF8CD:8E85D4DDDA38FF875EADF973FA9DF942" box="[911,1331,1701,1728]" country="China" latitude="32.97" longLatPrecision="725" longitude="104.1" municipality="Muyangchang" name="Wanglang National Natural Reserve" pageId="2" pageNumber="411" stateProvince="Sichuan Province">Wanglang National Natural Reserve</location>
,
<collectingRegion id="499E4CE4DA38FF875860F973FF53F961" country="China" name="Sichuan" pageId="2" pageNumber="411">Sichuan Province</collectingRegion>
, alt.
<quantity id="4CA22FE3DA38FF875C16F91FFE22F966" box="[308,396,1737,1764]" metricMagnitude="3" metricUnit="m" metricValue="2.577" pageId="2" pageNumber="411" unit="m" value="2577.0">
<elevation id="00776535DA38FF875C16F91FFE22F966" box="[308,396,1737,1764]" metricMagnitude="3" metricUnit="m" metricValue="2.577" pageId="2" pageNumber="411" unit="m" value="2577.0">2577 m</elevation>
</quantity>
,
<date id="FFE4A4C6DA38FF875CBAF91FFD83F961" box="[408,557,1737,1763]" pageId="2" pageNumber="411" value="2017-07-19">
<collectingDate id="EFA05D2EDA38FF875CBAF91FFD83F961" box="[408,557,1737,1763]" pageId="2" pageNumber="411" value="2017-07-19">19 July 2017</collectingDate>
</date>
, leg.
<collectorName id="26AFE7D0DA38FF875F48F91FFD11F966" box="[618,703,1737,1764]" pageId="2" pageNumber="411">Qi M.J.</collectorName>
&amp;
<collectorName id="26AFE7D0DA38FF875FC1F91CFCF9F966" box="[739,855,1738,1764]" pageId="2" pageNumber="411">Yang X.F.</collectorName>
, genitalia slide no. ZAH19003
</materialsCitation>
.
<materialsCitation id="3B32885BDA38FF8759E2F91CFD02F885" ID-GBIF-Occurrence="3392778327" accessionNumber="ZAH19030" collectingDate="2017-07-19" collectorName="Qi M. J. &amp; Yang X. F." country="China" elevation="2577" latitude="32.97" location="Wanglang National Natural Reserve" longLatPrecision="725" longitude="104.1" municipality="Muyangchang" pageId="2" pageNumber="411" specimenCount="1" specimenCount-female="1" stateProvince="Sichuan" typeStatus="paratype">
<typeStatus id="54E13CA4DA38FF8759E2F91CFA88F966" box="[1216,1318,1738,1764]" pageId="2" pageNumber="411" type="paratype">Paratype</typeStatus>
.
<specimenCount id="9D5C498FDA38FF875813F91FFAFAF961" box="[1329,1364,1737,1763]" pageId="2" pageNumber="411" type="female">1♀</specimenCount>
, same data as holotype, genitalia slide no. ZAH19030
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C340D18DDA38FF875DE5F8C7FC2AF8CD" pageId="2" pageNumber="411" type="etymology">
<paragraph id="8BE58206DA38FF875DE5F8C7FC2AF8CD" blockId="2.[151,1437,693,1871]" pageId="2" pageNumber="411">
<emphasis id="B92E5E14DA38FF875DE5F8C7FEE0F8A9" bold="true" box="[199,334,1809,1835]" pageId="2" pageNumber="411">Etymology.</emphasis>
The specific name is derived from the Latin
<emphasis id="B92E5E14DA38FF875E60F8C4FCD9F8A9" box="[834,887,1810,1835]" italics="true" pageId="2" pageNumber="411">tetra</emphasis>
- (= four) and -
<emphasis id="B92E5E14DA38FF87593FF8C4FBD1F8A9" box="[1053,1151,1810,1835]" italics="true" pageId="2" pageNumber="411">cornatus</emphasis>
, referring to the vesica in the male genitalia with four apical non-deciduous, spinous cornuti.
</paragraph>
</subSubSection>
</treatment>
</document>
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