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<document id="753119251FA10E2E79F2F555FDDF6765" ID-CLB-Dataset="21966" ID-DOI="10.5852/ejt.2016.195" ID-GBIF-Dataset="39071d6f-d900-4c81-ace1-dd03875018c5" ID-ISSN="2118-9773" ID-Zenodo-Dep="3837846" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1590071140602" checkinUser="valdenar" docAuthor="Likhitrakarn, Natdanai, Golovatch, Sergei I. &amp; Panha, Somsak" docDate="2016" docId="03EB5A00FFF97D61AC35D55AFCA79CB6" docLanguage="en" docName="ejt-2016-195.pdf.imf" docOrigin="European Journal of Taxonomy 195" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Tylopus hongkhraiensis Likhitrakarn &amp; Golovatch &amp; Panha 2016, sp. nov." docType="treatment" docUuid="E217CF1E-83D8-4EFD-875B-7C3845D01580" docUuidSource="ZooBank" docVersion="8" lastPageNumber="10" masterDocId="FFD22278FFFF7D68AE62D451FFD19824" masterDocTitle="The millipede genus Tylopus Jeekel, 1968 (Diplopoda, Polydesmida, Paradoxosomatidae), with a key and descriptions of eight new species from Indochina" masterLastPageNumber="47" masterPageNumber="1" pageNumber="7" updateTime="1698840626989" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0" zenodo-license-figures="CC-BY-3.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="B1D0727BC38899C73191CAD6A90BDA7C">The millipede genus Tylopus Jeekel, 1968 (Diplopoda, Polydesmida, Paradoxosomatidae), with a key and descriptions of eight new species from Indochina</mods:title>
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<mods:namePart id="9D5F22FE8370E962CC95EB483D8D14F5">Likhitrakarn, Natdanai</mods:namePart>
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<mods:affiliation id="9D636550F57829F4D2C280666EBE093C">Division of Plant Protection, Faculty of Agricultural Production, Maejo University, Chiang Mai 50290, Thailand. &amp; Email: kongerrrr @ hotmail. com &amp; urn: lsid: zoobank. org: author: 442 E 2 E 6 E- 1 CB 3 - 47 B 2 - 9896 - 1 A 922 CE 509 E 1</mods:affiliation>
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<mods:namePart id="76F0EFC7F954174C9EE4D1F07CC6D7F9">Golovatch, Sergei I.</mods:namePart>
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<mods:affiliation id="A72BE6F825A268F5E29DA60517DEFB9C">Institute for Problems of Ecology and Evolution, Russian Academy of Sciences, Leninsky pr. 33, Moscow 119071, Russia. &amp; Corresponding author: sgolovatch @ yandex. ru &amp; urn: lsid: zoobank. org: author: 71532 F 45 - BDD 5 - 415 D-BC 54 - 86256 E 5 D 5 D 4 A &amp; Corresponding author: sgolovatch @ yandex. ru &amp; Corresponding author: somsak. pan @ chula. ac. th</mods:affiliation>
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<mods:namePart id="35E53AE1B3812A6571ABCC671C348CBB">Panha, Somsak</mods:namePart>
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<mods:affiliation id="AEF812E940425247ABA8E06C85658E56">Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand. &amp; Corresponding author: somsak. pan @ chula. ac. th &amp; urn: lsid: zoobank. org: author: AC 935098 - D 901 - 4 F 35 - A 414 - 4 B 0 D 4 FE 44 E 79 &amp; Corresponding author: sgolovatch @ yandex. ru &amp; Corresponding author: somsak. pan @ chula. ac. th</mods:affiliation>
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<emphasis id="B9363704FFF97D6EAC35D55AFCA99902" bold="true" box="[599,888,267,294]" italics="true" pageId="6" pageNumber="7">Tylopus hongkhraiensis</emphasis>
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<paragraph id="8BFDEB16FFF97D6EACB3D503FCB39948" blockId="6.[369,1218,267,364]" box="[721,866,338,364]" pageId="6" pageNumber="7">
<figureCitation id="1379F793FFF97D6EACB3D503FCEB9948" box="[721,826,338,364]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStartId-0="6.[189,232,1858,1884]" captionStartId-1="7.[189,232,1964,1990]" captionStartId-2="8.[189,232,1930,1956]" captionTargetBox-0="[189,1398,567,1816]" captionTargetBox-1="[339,1250,1163,1911]" captionTargetBox-2="[189,1398,682,1888]" captionTargetId-0="figure@6.[189,1398,567,1816]" captionTargetId-1="figure@7.[317,1262,1158,1917]" captionTargetId-2="figure@8.[189,1398,682,1888]" captionTargetPageId-0="6" captionTargetPageId-1="7" captionTargetPageId-2="8" captionText-0="Fig. 4. Tylopus hongkhraiensis sp. nov., holotype, Ƌ (B–J), paratypes, Ƌ, ♀ (A). A. Habitus, live coloration. B–C. Anterior part of body, dorsal and lateral views, respectively. D–E. Segments 10 and 11, dorsal and lateral views, respectively. F–H. Posterior part of body, dorsal, ventral and lateral views, respectively I–J. Sternal cones between coxae 4, subcaudal and sublateral views, respectively. Abbreviations are explained in the text." captionText-1="Fig. 5. Tylopus hongkhraiensis sp. nov., holotype, Ƌ, right gonopod. A–C. Lateral, mesal and suboral views, respectively. Scale bar = 0.2 mm. Abbreviations are explained in the text." captionText-2="Fig. 6. Tylopus hongkhraiensis sp. nov., holotype, Ƌ, left gonopod. A–B. Mesal and lateral views, respectively. C–F. Distal part, sublateral, submesal, suboral and subcaudal views, respectively. Scale bars = 0.1 mm." figureDoi-0="http://doi.org/10.5281/zenodo.3837879" figureDoi-1="http://doi.org/10.5281/zenodo.3837881" figureDoi-2="http://doi.org/10.5281/zenodo.3837885" httpUri-0="https://zenodo.org/record/3837879/files/figure.png" httpUri-1="https://zenodo.org/record/3837881/files/figure.png" httpUri-2="https://zenodo.org/record/3837885/files/figure.png" pageId="6" pageNumber="7">Figs 4–6</figureCitation>
, 
<figureCitation id="1379F793FFF97D6EAD27D503FCB39948" box="[837,866,338,364]" captionStart="Fig" captionStartId="38.[189,232,1821,1847]" captionTargetBox="[266,1310,1040,1771]" captionTargetId="figure@38.[266,1310,1040,1771]" captionTargetPageId="38" captionText="Fig. 26. Distribution of the new eight species of Tylopus. Filled triangle. T. punctus sp. nov. Filled circle. T. acuminatus sp. nov. Crossed circle. T. acuminatus sp. nov., T. dorsalis sp. nov. Crossed square. T. acuminatus sp. nov. T. retusus sp. nov. Open circle. T. retusus sp. nov. Open triangle. T. baenzigeri Golovatch &amp; Enghoff, 1993. Filled square. T. hongkhraiensis sp. nov. Open diamond. T. thunghaihin sp. nov. Inverted filled triangle. T. flavolineatus sp. nov. Open square. T. moniliformis sp. nov." figureDoi="http://doi.org/10.5281/zenodo.3837927" httpUri="https://zenodo.org/record/3837927/files/figure.png" pageId="6" pageNumber="7">26</figureCitation>
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<heading id="D0B55C7AFFF97D6EAEDFD5C2FEE79989" bold="true" box="[189,310,403,429]" fontSize="11" level="3" pageId="6" pageNumber="7" reason="3">
<emphasis id="B9363704FFF97D6EAEDFD5C2FEE79989" bold="true" box="[189,310,403,429]" pageId="6" pageNumber="7">Diagnosis</emphasis>
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Differs from all known congeners in the almost missing paraterga, coupled with the gonopod being especially similar to that of 
<taxonomicName id="4C429095FFF97D6EAC7AD5B7FCA199DB" authority="Golovatch, 1984" authorityName="Golovatch" authorityYear="1984" box="[536,880,485,511]" class="Diplopoda" family="Paradoxosomatidae" genus="Tylopus" kingdom="Animalia" order="Polydesmida" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="magicus">
<emphasis id="B9363704FFF97D6EAC7AD5B7FD4A99DB" box="[536,667,486,511]" italics="true" pageId="6" pageNumber="7">T. magicus</emphasis>
<bibRefCitation id="EFD396E7FFF97D6EACC4D5B4FCA199DB" author="Golovatch S. I." box="[678,880,485,511]" pageId="6" pageNumber="7" pagination="53 - 77" refId="ref19517" refString="Golovatch S. I. 1984. Contributions to the millipede fauna of Vietnam (Diplopoda) II. Acta Zoologica Hungarica 30: 53 - 77." type="journal article" year="1984">Golovatch, 1984</bibRefCitation>
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, but differs in process 
<emphasis id="B9363704FFF97D6EAAEDD5B4FB4E99DB" bold="true" box="[1167,1183,485,511]" pageId="6" pageNumber="7">h</emphasis>
being strong and
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<paragraph id="8BFDEB16FFF97D6EAEDFD313FD5D9FCE" blockId="6.[189,1399,1858,2026]" pageId="6" pageNumber="7">
<emphasis id="B9363704FFF97D6EAEDFD313FEDE9F78" bold="true" box="[189,271,1858,1884]" pageId="6" pageNumber="7">Fig. 4.</emphasis>
<taxonomicName id="4C429095FFF97D6EAF79D312FDE49F79" authority="Likhitrakarn &amp; Golovatch &amp; Panha, 2016" authorityName="Likhitrakarn &amp; Golovatch &amp; Panha" authorityYear="2016" box="[283,565,1859,1885]" class="Diplopoda" family="Paradoxosomatidae" genus="Tylopus" kingdom="Animalia" order="Polydesmida" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="hongkhraiensis" status="sp. nov.">
<emphasis id="B9363704FFF97D6EAF79D312FDE49F79" box="[283,565,1859,1885]" italics="true" pageId="6" pageNumber="7">Tylopus hongkhraiensis</emphasis>
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, holotype, Ƌ (
<emphasis id="B9363704FFF97D6EAD38D312FC5A9F79" bold="true" box="[858,907,1859,1885]" pageId="6" pageNumber="7">B–J</emphasis>
), paratypes, Ƌ, ♀ (
<emphasis id="B9363704FFF97D6EAAE3D313FB479F78" bold="true" box="[1153,1174,1858,1884]" pageId="6" pageNumber="7">A</emphasis>
). 
<emphasis id="B9363704FFF97D6EAAD1D313FB199F78" bold="true" box="[1203,1224,1858,1884]" pageId="6" pageNumber="7">A</emphasis>
. Habitus, live coloration. 
<emphasis id="B9363704FFF97D6EAF2AD337FEAE9FA4" bold="true" box="[328,383,1894,1920]" pageId="6" pageNumber="7">B–C</emphasis>
. Anterior part of body, dorsal and lateral views, respectively. 
<emphasis id="B9363704FFF97D6EAAECD337FB149FA4" bold="true" box="[1166,1221,1894,1920]" pageId="6" pageNumber="7">D–E</emphasis>
. Segments 10 and 11, dorsal and lateral views, respectively. 
<emphasis id="B9363704FFF97D6EAC8AD3D8FCCE9F87" bold="true" box="[744,799,1929,1955]" pageId="6" pageNumber="7">F–H</emphasis>
. Posterior part of body, dorsal, ventral and lateral views, respectively 
<emphasis id="B9363704FFF97D6EAFCBD3FDFE039FE2" bold="true" box="[425,466,1964,1990]" pageId="6" pageNumber="7">I–J</emphasis>
. Sternal cones between coxae 4, subcaudal and sublateral views, respectively. Abbreviations are explained in the text.
</paragraph>
</caption>
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pointed, while process 
<emphasis id="B9363704FFF87D6FAFA9D55DFE099902" bold="true" box="[459,472,268,294]" pageId="7" pageNumber="8">z</emphasis>
long, slender and pointed.
</paragraph>
</subSubSection>
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<paragraph id="8BFDEB16FFF87D6FAEDFD51CFE959943" blockId="7.[189,324,333,359]" box="[189,324,333,359]" pageId="7" pageNumber="8">
<heading id="D0B55C7AFFF87D6FAEDFD51CFE959943" bold="true" box="[189,324,333,359]" fontSize="11" level="3" pageId="7" pageNumber="8" reason="3">
<emphasis id="B9363704FFF87D6FAEDFD51CFE959943" bold="true" box="[189,324,333,359]" pageId="7" pageNumber="8">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="8BFDEB16FFF87D6FAEDFD52AFC4F99B1" blockId="7.[189,926,379,406]" box="[189,926,379,406]" pageId="7" pageNumber="8">
The species is named to emphasize the 
<typeStatus id="54F955B4FFF87D6FACE9D52DFD6F99B2" box="[651,702,380,406]" pageId="7" pageNumber="8">type</typeStatus>
locality; adjective.
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<subSubSection id="C358B89DFFF87D6FAEDFD5ECFDC49AF7" pageId="7" pageNumber="8" type="materials_examined">
<paragraph id="8BFDEB16FFF87D6FAEDFD5ECFE7B99F3" blockId="7.[189,426,445,471]" box="[189,426,445,471]" pageId="7" pageNumber="8">
<heading id="D0B55C7AFFF87D6FAEDFD5ECFE7B99F3" bold="true" box="[189,426,445,471]" fontSize="11" level="3" pageId="7" pageNumber="8" reason="3">
<emphasis id="B9363704FFF87D6FAEDFD5ECFE7B99F3" bold="true" box="[189,426,445,471]" pageId="7" pageNumber="8">Material examined</emphasis>
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<typeStatus id="54F955B4FFF87D6FAE88D5BDFE8D9A22" box="[234,348,492,518]" pageId="7" pageNumber="8" type="holotype">Holotype</typeStatus>
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: Ƌ, 
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Huai 
<collectingMunicipality id="6B99716CFFF87D6FAF81D65EFCAD9A0C" box="[483,892,526,553]" pageId="7" pageNumber="8">
Hong 
<location id="8E9DBDCDFFF87D6FAC57D65FFCAD9A0C" LSID="urn:lsid:plazi:treatment:03EB5A00FFF97D61AC35D55AFCA79CB6:8E9DBDCDFFF87D6FAC57D65FFCAD9A0C" box="[565,892,526,552]" country="Thailand" county="Huai Hong Khrai Royal Development" latitude="18.879168" longLatPrecision="21" longitude="99.211105" municipality="Hong Khrai Royal Development Study" name="Khrai Royal Development Study Centre" pageId="7" pageNumber="8" stateProvince="Chiang Mai">Khrai Royal Development</location>
</collectingMunicipality>
</collectingCounty>
Study Centre, 
<location id="8E9DBDCDFFF87D6FAA24D65FFB179A0C" LSID="urn:lsid:plazi:treatment:03EB5A00FFF97D61AC35D55AFCA79CB6:8E9DBDCDFFF87D6FAA24D65FFB179A0C" box="[1094,1222,526,552]" country="Thailand" county="Huai Hong Khrai Royal Development" latitude="18.879168" longLatPrecision="21" longitude="99.211105" municipality="Hong Khrai Royal Development Study" name="Doi Saket" pageId="7" pageNumber="8" stateProvince="Chiang Mai">Doi Saket</location>
, 
<collectingRegion id="498625F4FFF87D6FAAB9D65FFAA39A0C" box="[1243,1394,526,552]" country="Thailand" name="Chiang Mai" pageId="7" pageNumber="8">Chiang Mai</collectingRegion>
, 
<geoCoordinate id="EE768DD1FFF87D6FAEDFD660FE809A68" box="[189,337,561,588]" degrees="18" direction="north" minutes="52" orientation="latitude" pageId="7" pageNumber="8" precision="15" seconds="45" value="18.879168">18°52'45&quot; N</geoCoordinate>
, 
<geoCoordinate id="EE768DD1FFF87D6FAF3FD663FE3C9A68" box="[349,493,562,588]" degrees="99" direction="east" minutes="12" orientation="longitude" pageId="7" pageNumber="8" precision="15" seconds="40" value="99.211105">99°12'40&quot; E</geoCoordinate>
, ca 
<elevation id="006F0C25FFF87D6FAC79D663FD749A68" box="[539,677,562,588]" metricMagnitude="2" metricUnit="m" metricValue="4.2" pageId="7" pageNumber="8" unit="m" value="420.0">
<quantity id="4CBA46F3FFF87D6FAC79D663FDB79A68" box="[539,614,562,588]" metricMagnitude="2" metricUnit="m" metricValue="4.2" pageId="7" pageNumber="8" unit="m" value="420.0">420 m</quantity>
a.s.l.
</elevation>
, 
<date id="FFFCCDD6FFF87D6FACD0D663FC979A68" box="[690,838,561,588]" pageId="7" pageNumber="8" value="2015-07-29">
<collectingDate id="EFB8343EFFF87D6FACD0D663FC979A68" box="[690,838,561,588]" pageId="7" pageNumber="8" value="2015-07-29">29 Jul. 2015</collectingDate>
</date>
, leg. 
<collectorName id="26B78EC0FFF87D6FADE6D663FBEB9A68" box="[900,1082,562,588]" pageId="7" pageNumber="8">N. Likhitrakarn</collectorName>
(
<collectionCode id="ED5373D3FFF87D6FAA28D663FB779A68" box="[1098,1190,562,588]" country="0" httpUri="http://grbio.org/cool/da58-p0xp" name="Cameroon University, Museum of Zoology" pageId="7" pageNumber="8">CUMZ</collectionCode>
).
</paragraph>
</materialsCitation>
<paragraph id="8BFDEB16FFF87D6FAE88D622FEB69AA9" blockId="7.[189,1398,627,724]" box="[234,359,627,653]" pageId="7" pageNumber="8">
<emphasis id="B9363704FFF87D6FAE88D622FEB69AA9" bold="true" box="[234,359,627,653]" pageId="7" pageNumber="8">
<typeStatus id="54F955B4FFF87D6FAE88D622FEB69AA9" box="[234,359,627,653]" pageId="7" pageNumber="8" type="paratype">Paratypes</typeStatus>
</emphasis>
</paragraph>
<paragraph id="8BFDEB16FFF87D6FAEDFD6C6FDC49AF7" blockId="7.[189,1398,627,724]" pageId="7" pageNumber="8">
<collectingCountry id="F355AB86FFF87D6FAEDFD6C6FE889A95" box="[189,345,663,689]" name="Thailand" pageId="7" pageNumber="8">THAILAND</collectingCountry>
: 4 ƋƋ, 
<specimenCount id="9D44209FFFF87D6FAFD1D6C7FE249A95" box="[435,501,662,689]" pageId="7" pageNumber="8" type="female">4 ♀♀</specimenCount>
(
<collectionCode id="ED5373D3FFF87D6FAC66D6C7FD8E9A94" box="[516,607,662,688]" country="0" httpUri="http://grbio.org/cool/da58-p0xp" name="Cameroon University, Museum of Zoology" pageId="7" pageNumber="8">CUMZ</collectionCode>
); 1 Ƌ, 
<specimenCount id="9D44209FFFF87D6FACCED6C7FD099A95" box="[684,728,662,689]" pageId="7" pageNumber="8" type="female">1 ♀</specimenCount>
(
<collectionCode id="ED5373D3FFF87D6FAC84D6C7FC929A94" box="[742,835,662,688]" country="China" httpUri="http://grbio.org/cool/49rk-3jpq" name="Zoological Museum, University of Amoy" pageId="7" pageNumber="8">ZMUM</collectionCode>
ρ3060); 1 Ƌ, 
<specimenCount id="9D44209FFFF87D6FAD84D6C7FBC09A95" box="[998,1041,662,689]" pageId="7" pageNumber="8" type="female">1 ♀</specimenCount>
(
<collectionCode id="ED5373D3FFF87D6FAA42D6C7FBAA9A94" box="[1056,1147,662,688]" country="Denmark" httpUri="http://grbio.org/cool/mci8-ehqk" name="Zoological Museum, University of Copenhagen" pageId="7" pageNumber="8">ZMUC</collectionCode>
); 1 Ƌ, 
<specimenCount id="9D44209FFFF87D6FAAABD6C7FB249A95" box="[1225,1269,662,689]" pageId="7" pageNumber="8" type="female">1 ♀</specimenCount>
(
<collectionCode id="ED5373D3FFF87D6FAB61D6C7FAB89A94" box="[1283,1385,662,688]" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="7" pageNumber="8" type="Museum">NHMW</collectionCode>
), all same data as for 
<typeStatus id="54F955B4FFF87D6FAFC5D6E8FDC39AF7" box="[423,530,697,723]" pageId="7" pageNumber="8" type="holotype">holotype</typeStatus>
.
</paragraph>
</subSubSection>
<subSubSection id="C358B89DFFF87D61AEDFD6AAFDEF9BFF" lastPageId="9" lastPageNumber="10" pageId="7" pageNumber="8" type="description">
<paragraph id="8BFDEB16FFF87D6FAEDFD6AAFE9F9B31" blockId="7.[189,334,763,789]" box="[189,334,763,789]" pageId="7" pageNumber="8">
<heading id="D0B55C7AFFF87D6FAEDFD6AAFE9F9B31" bold="true" box="[189,334,763,789]" fontSize="11" level="3" pageId="7" pageNumber="8" reason="3">
<emphasis id="B9363704FFF87D6FAEDFD6AAFE9F9B31" bold="true" box="[189,334,763,789]" pageId="7" pageNumber="8">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8BFDEB16FFF87D6FAEDFD77BFAB09BF4" blockId="7.[189,1399,809,977]" pageId="7" pageNumber="8">
MEASUREMENTS AND COLOUR. Length 11.2–12.8 (Ƌ) or 
<quantity id="4CBA46F3FFF87D6FAD23D778FC3A9B60" box="[833,1003,809,836]" metricMagnitude="-2" metricUnit="m" metricValue="1.205" metricValueMax="1.26" metricValueMin="1.15" pageId="7" pageNumber="8" unit="mm" value="12.05" valueMax="12.6" valueMin="11.5">11.5–12.6 mm</quantity>
(
<specimenCount id="9D44209FFFF87D6FAA60D77BFBC49B60" box="[1026,1045,810,836]" pageId="7" pageNumber="8" type="female">♀</specimenCount>
), width of midbody pro- and metazonae 0.85–0.96 and 
<quantity id="4CBA46F3FFF87D6FAF8BD71CFD539B43" box="[489,642,845,871]" metricMagnitude="-3" metricUnit="m" metricValue="1.12" metricValueMax="1.2" metricValueMin="1.04" pageId="7" pageNumber="8" unit="mm" value="1.12" valueMax="1.2" valueMin="1.04">1.04–1.2 mm</quantity>
(Ƌ) or 0.85–1.05 and 
<quantity id="4CBA46F3FFF87D6FAD1DD71DFBF79B43" box="[895,1062,844,871]" metricMagnitude="-3" metricUnit="m" metricValue="1.185" metricValueMax="1.32" metricValueMin="1.05" pageId="7" pageNumber="8" unit="mm" value="1.185" valueMax="1.32" valueMin="1.05">1.05–1.32 mm</quantity>
(
<specimenCount id="9D44209FFFF87D6FAA5AD71CFB9A9B43" box="[1080,1099,845,871]" pageId="7" pageNumber="8" type="female">♀</specimenCount>
), respectively. Coloration of live animals dark brown (
<figureCitation id="1379F793FFF87D6FAC67D721FDB29BAE" box="[517,611,880,906]" captionStart="Fig" captionStartId="6.[189,232,1858,1884]" captionTargetBox="[189,1398,567,1816]" captionTargetId="figure@6.[189,1398,567,1816]" captionTargetPageId="6" captionText="Fig. 4. Tylopus hongkhraiensis sp. nov., holotype, Ƌ (B–J), paratypes, Ƌ, ♀ (A). A. Habitus, live coloration. B–C. Anterior part of body, dorsal and lateral views, respectively. D–E. Segments 10 and 11, dorsal and lateral views, respectively. F–H. Posterior part of body, dorsal, ventral and lateral views, respectively I–J. Sternal cones between coxae 4, subcaudal and sublateral views, respectively. Abbreviations are explained in the text." figureDoi="http://doi.org/10.5281/zenodo.3837879" httpUri="https://zenodo.org/record/3837879/files/figure.png" pageId="7" pageNumber="8">Fig. 4A</figureCitation>
); legs light brown, venter and a few basal podomeres light brown to yellow-brown. Coloration in alcohol after three months of preservation faded to light brown; antennae and epiproct light brown to pallid, venter and a few basal podomeres light brown to pallid (
<figureCitation id="1379F793FFF87D6FAABCD7E7FAF29BF4" box="[1246,1315,950,976]" captionStart="Fig" captionStartId="6.[189,232,1858,1884]" captionTargetBox="[189,1398,567,1816]" captionTargetId="figure@6.[189,1398,567,1816]" captionTargetPageId="6" captionText="Fig. 4. Tylopus hongkhraiensis sp. nov., holotype, Ƌ (B–J), paratypes, Ƌ, ♀ (A). A. Habitus, live coloration. B–C. Anterior part of body, dorsal and lateral views, respectively. D–E. Segments 10 and 11, dorsal and lateral views, respectively. F–H. Posterior part of body, dorsal, ventral and lateral views, respectively I–J. Sternal cones between coxae 4, subcaudal and sublateral views, respectively. Abbreviations are explained in the text." figureDoi="http://doi.org/10.5281/zenodo.3837879" httpUri="https://zenodo.org/record/3837879/files/figure.png" pageId="7" pageNumber="8">Fig. 4</figureCitation>
B–J).
</paragraph>
<paragraph id="8BFDEB16FFF87D60AEDFD7A9FDFB9948" blockId="7.[189,1399,1016,1113]" lastBlockId="8.[189,1399,267,364]" lastPageId="8" lastPageNumber="9" pageId="7" pageNumber="8">
HEAD. Clypeolabral region and vertex sparsely setose, epicranial suture distinct. Antennae rather short (
<figureCitation id="1379F793FFF87D6FAEA4D04AFEF59C11" box="[198,292,1051,1077]" captionStart="Fig" captionStartId="6.[189,232,1858,1884]" captionTargetBox="[189,1398,567,1816]" captionTargetId="figure@6.[189,1398,567,1816]" captionTargetPageId="6" captionText="Fig. 4. Tylopus hongkhraiensis sp. nov., holotype, Ƌ (B–J), paratypes, Ƌ, ♀ (A). A. Habitus, live coloration. B–C. Anterior part of body, dorsal and lateral views, respectively. D–E. Segments 10 and 11, dorsal and lateral views, respectively. F–H. Posterior part of body, dorsal, ventral and lateral views, respectively I–J. Sternal cones between coxae 4, subcaudal and sublateral views, respectively. Abbreviations are explained in the text." figureDoi="http://doi.org/10.5281/zenodo.3837879" httpUri="https://zenodo.org/record/3837879/files/figure.png" pageId="7" pageNumber="8">Fig. 4A</figureCitation>
), surpassing body segment 2 (Ƌ, 
<specimenCount id="9D44209FFFF87D6FACCBD04AFD6C9C11" box="[681,701,1051,1077]" pageId="7" pageNumber="8" type="female">♀</specimenCount>
) when stretched dorsally. In width, segment 3 &lt;2 &lt;collum &lt;segment 4 &lt;5 &lt;6 &lt;7 &lt;head &lt;8–17 (Ƌ) or segment 2 &lt;3 &lt;collum &lt;segment 4 &lt;5 = 6 &lt;7 &lt;head &lt;8–17 (
<specimenCount id="9D44209FFFF77D60AF6FD55AFEF09901" box="[269,289,267,293]" pageId="8" pageNumber="9" type="female">♀</specimenCount>
); thereafter body gently and gradually tapering. Collum with three transverse rows of setae: 3+3 anterior, 2+2 intermediate and 3+3 posterior; caudal corner very narrowly rounded, not surpassing rear tergal margin (
<figureCitation id="1379F793FFF77D60AFFCD503FDCB9948" box="[414,538,338,364]" captionStart="Fig" captionStartId="6.[189,232,1858,1884]" captionTargetBox="[189,1398,567,1816]" captionTargetId="figure@6.[189,1398,567,1816]" captionTargetPageId="6" captionText="Fig. 4. Tylopus hongkhraiensis sp. nov., holotype, Ƌ (B–J), paratypes, Ƌ, ♀ (A). A. Habitus, live coloration. B–C. Anterior part of body, dorsal and lateral views, respectively. D–E. Segments 10 and 11, dorsal and lateral views, respectively. F–H. Posterior part of body, dorsal, ventral and lateral views, respectively I–J. Sternal cones between coxae 4, subcaudal and sublateral views, respectively. Abbreviations are explained in the text." figureDoi="http://doi.org/10.5281/zenodo.3837879" httpUri="https://zenodo.org/record/3837879/files/figure.png" pageId="8" pageNumber="9">
Fig. 4B 
<emphasis id="B9363704FFF77D60AF95D503FDD79948" bold="true" box="[503,518,338,364]" pageId="8" pageNumber="9">–</emphasis>
C
</figureCitation>
).
</paragraph>
<caption id="DF3DBB9EFFF87D6FAEDFD3FDFBBD9FCE" ID-DOI="http://doi.org/10.5281/zenodo.3837881" ID-Zenodo-Dep="3837881" httpUri="https://zenodo.org/record/3837881/files/figure.png" pageId="7" pageNumber="8" startId="7.[189,232,1964,1990]" targetBox="[339,1250,1163,1911]" targetPageId="7">
<paragraph id="8BFDEB16FFF87D6FAEDFD3FDFBBD9FCE" blockId="7.[189,1398,1964,2026]" pageId="7" pageNumber="8">
<emphasis id="B9363704FFF87D6FAEDFD3FDFEDD9FE3" bold="true" box="[189,268,1964,1991]" pageId="7" pageNumber="8">Fig. 5.</emphasis>
<taxonomicName id="4C429095FFF87D6FAF77D3FDFDFA9FE2" authority="Likhitrakarn &amp; Golovatch &amp; Panha, 2016" authorityName="Likhitrakarn &amp; Golovatch &amp; Panha" authorityYear="2016" box="[277,555,1964,1990]" class="Diplopoda" family="Paradoxosomatidae" genus="Tylopus" kingdom="Animalia" order="Polydesmida" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="hongkhraiensis" status="sp. nov.">
<emphasis id="B9363704FFF87D6FAF77D3FDFDFA9FE2" box="[277,555,1964,1990]" italics="true" pageId="7" pageNumber="8">Tylopus hongkhraiensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2058A7FFFF87D6FAC56D3FCFD429FE3" box="[564,659,1965,1991]" pageId="7" pageNumber="8" rank="species">sp. nov.</taxonomicNameLabel>
, holotype, Ƌ, right gonopod. 
<emphasis id="B9363704FFF87D6FAD96D3FDFBFC9FE2" bold="true" box="[1012,1069,1964,1990]" pageId="7" pageNumber="8">A–C</emphasis>
. Lateral, mesal and suboral views, respectively. Scale bar = 0.2 mm. Abbreviations are explained in the text.
</paragraph>
</caption>
<paragraph id="8BFDEB16FFF77D61AEDFD5C5FC889AA1" blockId="8.[189,1399,403,606]" lastBlockId="9.[189,1399,267,646]" lastPageId="9" lastPageNumber="10" pageId="8" pageNumber="9">
BODY. Tegument smooth and shining, prozonae very finely shagreened, metaterga nearly smooth, faintly rugulose and leathery (
<figureCitation id="1379F793FFF77D60AFB6D5E7FDCD99F4" box="[468,540,438,464]" captionStart="Fig" captionStartId="6.[189,232,1858,1884]" captionTargetBox="[189,1398,567,1816]" captionTargetId="figure@6.[189,1398,567,1816]" captionTargetPageId="6" captionText="Fig. 4. Tylopus hongkhraiensis sp. nov., holotype, Ƌ (B–J), paratypes, Ƌ, ♀ (A). A. Habitus, live coloration. B–C. Anterior part of body, dorsal and lateral views, respectively. D–E. Segments 10 and 11, dorsal and lateral views, respectively. F–H. Posterior part of body, dorsal, ventral and lateral views, respectively I–J. Sternal cones between coxae 4, subcaudal and sublateral views, respectively. Abbreviations are explained in the text." figureDoi="http://doi.org/10.5281/zenodo.3837879" httpUri="https://zenodo.org/record/3837879/files/figure.png" pageId="8" pageNumber="9">Fig. 4</figureCitation>
B–F, H). Postcollum metaterga with two transverse rows of setae: 2+2 anterior and 2+2 posterior, traceable at least as insertion points when setae broken off. Tergal setae simple, slender, rather short, about 1/5 of metatergal length. Axial line visible, traceable only on metazonae. Postcollum paraterga virtually missing, in pore-bearing segments lateral bulges with ozopores set at about half of midbody height. Ozopores (
<emphasis id="B9363704FFF77D60ACFCD612FD149A79" bold="true" box="[670,709,579,605]" pageId="8" pageNumber="9">Op</emphasis>
) evident (
<figureCitation id="1379F793FFF77D60AD59D612FC449A79" box="[827,917,579,605]" captionStart="Fig" captionStartId="6.[189,232,1858,1884]" captionTargetBox="[189,1398,567,1816]" captionTargetId="figure@6.[189,1398,567,1816]" captionTargetPageId="6" captionText="Fig. 4. Tylopus hongkhraiensis sp. nov., holotype, Ƌ (B–J), paratypes, Ƌ, ♀ (A). A. Habitus, live coloration. B–C. Anterior part of body, dorsal and lateral views, respectively. D–E. Segments 10 and 11, dorsal and lateral views, respectively. F–H. Posterior part of body, dorsal, ventral and lateral views, respectively I–J. Sternal cones between coxae 4, subcaudal and sublateral views, respectively. Abbreviations are explained in the text." figureDoi="http://doi.org/10.5281/zenodo.3837879" httpUri="https://zenodo.org/record/3837879/files/figure.png" pageId="8" pageNumber="9">Fig. 4E</figureCitation>
), lateral, located at about 1/3 of metatergal length in front of posterior edge of metaterga. Transverse sulcus complete on segments 6–16, incomplete on segments 7 and 18 (Ƌ, 
<specimenCount id="9D44209FFFF67D61AC63D57EFDC4996D" box="[513,533,303,329]" pageId="9" pageNumber="10" type="female">♀</specimenCount>
), very narrow, shallow, not reaching bases of paraterga (
<figureCitation id="1379F793FFF67D61AAA5D57FFADF996D" box="[1223,1294,302,329]" captionStart="Fig" captionStartId="6.[189,232,1858,1884]" captionTargetBox="[189,1398,567,1816]" captionTargetId="figure@6.[189,1398,567,1816]" captionTargetPageId="6" captionText="Fig. 4. Tylopus hongkhraiensis sp. nov., holotype, Ƌ (B–J), paratypes, Ƌ, ♀ (A). A. Habitus, live coloration. B–C. Anterior part of body, dorsal and lateral views, respectively. D–E. Segments 10 and 11, dorsal and lateral views, respectively. F–H. Posterior part of body, dorsal, ventral and lateral views, respectively I–J. Sternal cones between coxae 4, subcaudal and sublateral views, respectively. Abbreviations are explained in the text." figureDoi="http://doi.org/10.5281/zenodo.3837879" httpUri="https://zenodo.org/record/3837879/files/figure.png" pageId="9" pageNumber="10">Fig. 4</figureCitation>
B–F, H). Stricture between pro- and metazonae wide, deep, evidently ribbed at bottom down to base of paraterga (
<figureCitation id="1379F793FFF67D61AEA4D524FEDF99AB" box="[198,270,373,399]" captionStart="Fig" captionStartId="6.[189,232,1858,1884]" captionTargetBox="[189,1398,567,1816]" captionTargetId="figure@6.[189,1398,567,1816]" captionTargetPageId="6" captionText="Fig. 4. Tylopus hongkhraiensis sp. nov., holotype, Ƌ (B–J), paratypes, Ƌ, ♀ (A). A. Habitus, live coloration. B–C. Anterior part of body, dorsal and lateral views, respectively. D–E. Segments 10 and 11, dorsal and lateral views, respectively. F–H. Posterior part of body, dorsal, ventral and lateral views, respectively I–J. Sternal cones between coxae 4, subcaudal and sublateral views, respectively. Abbreviations are explained in the text." figureDoi="http://doi.org/10.5281/zenodo.3837879" httpUri="https://zenodo.org/record/3837879/files/figure.png" pageId="9" pageNumber="10">Fig. 4</figureCitation>
B–F, H). Pleurosternal carinae complete crests with a very sharp median tooth on segment 2, increasingly reduced to a rounded caudal crest bulged on segments 2 and 3 (Ƌ, 
<specimenCount id="9D44209FFFF67D61AA0BD5C8FBAC9997" box="[1129,1149,409,435]" pageId="9" pageNumber="10" type="female">♀</specimenCount>
), retained only as an evident, sharp, caudal tooth on segments 5 and 6, thereafter increasingly reduced in size until segment 17, remaining as a small caudal tooth on segment 18 (Ƌ) (
<figureCitation id="1379F793FFF67D61AD1DD58FFBF799DD" box="[895,1062,478,505]" captionStart="Fig" captionStartId="6.[189,232,1858,1884]" captionTargetBox="[189,1398,567,1816]" captionTargetId="figure@6.[189,1398,567,1816]" captionTargetPageId="6" captionText="Fig. 4. Tylopus hongkhraiensis sp. nov., holotype, Ƌ (B–J), paratypes, Ƌ, ♀ (A). A. Habitus, live coloration. B–C. Anterior part of body, dorsal and lateral views, respectively. D–E. Segments 10 and 11, dorsal and lateral views, respectively. F–H. Posterior part of body, dorsal, ventral and lateral views, respectively I–J. Sternal cones between coxae 4, subcaudal and sublateral views, respectively. Abbreviations are explained in the text." figureDoi="http://doi.org/10.5281/zenodo.3837879" httpUri="https://zenodo.org/record/3837879/files/figure.png" pageId="9" pageNumber="10">Fig. 4C, E, H</figureCitation>
), or increasingly reduced to caudally roughly granulated bulges on segments 5 and 6, thereafter missing (
<specimenCount id="9D44209FFFF67D61AA2DD653FBB29A38" box="[1103,1123,514,540]" pageId="9" pageNumber="10" type="female">♀</specimenCount>
). Epiproct (
<figureCitation id="1379F793FFF67D61AA90D653FAE99A38" box="[1266,1336,514,540]" captionStart="Fig" captionStartId="6.[189,232,1858,1884]" captionTargetBox="[189,1398,567,1816]" captionTargetId="figure@6.[189,1398,567,1816]" captionTargetPageId="6" captionText="Fig. 4. Tylopus hongkhraiensis sp. nov., holotype, Ƌ (B–J), paratypes, Ƌ, ♀ (A). A. Habitus, live coloration. B–C. Anterior part of body, dorsal and lateral views, respectively. D–E. Segments 10 and 11, dorsal and lateral views, respectively. F–H. Posterior part of body, dorsal, ventral and lateral views, respectively I–J. Sternal cones between coxae 4, subcaudal and sublateral views, respectively. Abbreviations are explained in the text." figureDoi="http://doi.org/10.5281/zenodo.3837879" httpUri="https://zenodo.org/record/3837879/files/figure.png" pageId="9" pageNumber="10">Fig. 4</figureCitation>
F–H) conical, flattened dorsoventrally, with two small, but evident, rounded apical papillae; tip subtruncate; lateral pre-apical papillae very small, lying close to tip. Hypoproct roundly subtrapeziform, setigerous knobs at caudal edge small and well-separated (
<figureCitation id="1379F793FFF67D61AC8FD63AFC9D9AA1" box="[749,844,619,645]" captionStart="Fig" captionStartId="6.[189,232,1858,1884]" captionTargetBox="[189,1398,567,1816]" captionTargetId="figure@6.[189,1398,567,1816]" captionTargetPageId="6" captionText="Fig. 4. Tylopus hongkhraiensis sp. nov., holotype, Ƌ (B–J), paratypes, Ƌ, ♀ (A). A. Habitus, live coloration. B–C. Anterior part of body, dorsal and lateral views, respectively. D–E. Segments 10 and 11, dorsal and lateral views, respectively. F–H. Posterior part of body, dorsal, ventral and lateral views, respectively I–J. Sternal cones between coxae 4, subcaudal and sublateral views, respectively. Abbreviations are explained in the text." figureDoi="http://doi.org/10.5281/zenodo.3837879" httpUri="https://zenodo.org/record/3837879/files/figure.png" pageId="9" pageNumber="10">Fig. 4G</figureCitation>
).
</paragraph>
<caption id="DF3DBB9EFFF77D60AEDFD3DBFEBF9FCF" ID-DOI="http://doi.org/10.5281/zenodo.3837885" ID-Zenodo-Dep="3837885" httpUri="https://zenodo.org/record/3837885/files/figure.png" pageId="8" pageNumber="9" startId="8.[189,232,1930,1956]" targetBox="[189,1398,682,1888]" targetPageId="8">
<paragraph id="8BFDEB16FFF77D60AEDFD3DBFEBF9FCF" blockId="8.[189,1399,1930,2027]" pageId="8" pageNumber="9">
<emphasis id="B9363704FFF77D60AEDFD3DBFEDE9F80" bold="true" box="[189,271,1930,1956]" pageId="8" pageNumber="9">Fig. 6.</emphasis>
<taxonomicName id="4C429095FFF77D60AF79D3DBFDE59F80" authority="Likhitrakarn &amp; Golovatch &amp; Panha, 2016" authorityName="Likhitrakarn &amp; Golovatch &amp; Panha" authorityYear="2016" box="[283,564,1930,1956]" class="Diplopoda" family="Paradoxosomatidae" genus="Tylopus" kingdom="Animalia" order="Polydesmida" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="hongkhraiensis" status="sp. nov.">
<emphasis id="B9363704FFF77D60AF79D3DBFDE59F80" box="[283,564,1930,1956]" italics="true" pageId="8" pageNumber="9">Tylopus hongkhraiensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2058A7FFFF77D60AC22D3DAFD739F81" box="[576,674,1931,1957]" pageId="8" pageNumber="9" rank="species">sp. nov.</taxonomicNameLabel>
, holotype, Ƌ, left gonopod. 
<emphasis id="B9363704FFF77D60AA63D3DBFBE99F80" bold="true" box="[1025,1080,1930,1956]" pageId="8" pageNumber="9">A–B</emphasis>
. Mesal and lateral views, respectively. 
<emphasis id="B9363704FFF77D60AF38D3FCFE419FE3" bold="true" box="[346,400,1965,1991]" pageId="8" pageNumber="9">C–F</emphasis>
. Distal part, sublateral, submesal, suboral and subcaudal views, respectively. Scale bars = 0.1 mm.
</paragraph>
</caption>
<paragraph id="8BFDEB16FFF67D61AEDFD6FCFC6D9B14" blockId="9.[189,1399,685,817]" pageId="9" pageNumber="10">
STERNA. Sparsely setose, without modifications; a small, rounded, densely setose sternal cone between Ƌ coxae 4 (
<figureCitation id="1379F793FFF67D61AF2BD681FE699ACE" box="[329,440,720,746]" captionStart="Fig" captionStartId="6.[189,232,1858,1884]" captionTargetBox="[189,1398,567,1816]" captionTargetId="figure@6.[189,1398,567,1816]" captionTargetPageId="6" captionText="Fig. 4. Tylopus hongkhraiensis sp. nov., holotype, Ƌ (B–J), paratypes, Ƌ, ♀ (A). A. Habitus, live coloration. B–C. Anterior part of body, dorsal and lateral views, respectively. D–E. Segments 10 and 11, dorsal and lateral views, respectively. F–H. Posterior part of body, dorsal, ventral and lateral views, respectively I–J. Sternal cones between coxae 4, subcaudal and sublateral views, respectively. Abbreviations are explained in the text." figureDoi="http://doi.org/10.5281/zenodo.3837879" httpUri="https://zenodo.org/record/3837879/files/figure.png" pageId="9" pageNumber="10">Fig. 4I, J</figureCitation>
). A paramedian pair of small tubercles in front of gonopod aperture. Legs rather rather short, slightly incrassate in Ƌ, midbody ones ca 1.1–1.3 (Ƌ) or 0.8–0.9 times (
<specimenCount id="9D44209FFFF67D61AAFED6A2FB7E9B29" box="[1180,1199,755,781]" pageId="9" pageNumber="10" type="female">♀</specimenCount>
) as long as body height, prefemora without modifications, Ƌ tarsal brushes absent.
</paragraph>
<paragraph id="8BFDEB16FFF67D61AEDFD709FDEF9BFF" blockId="9.[189,1399,856,988]" pageId="9" pageNumber="10">
GONOPOD. Simple (
<figureCitation id="1379F793FFF67D61AFFAD709FE329B57" box="[408,483,856,883]" captionStart="Fig" captionStartId="7.[189,232,1964,1990]" captionTargetBox="[339,1250,1163,1911]" captionTargetId="figure@7.[317,1262,1158,1917]" captionTargetPageId="7" captionText="Fig. 5. Tylopus hongkhraiensis sp. nov., holotype, Ƌ, right gonopod. A–C. Lateral, mesal and suboral views, respectively. Scale bar = 0.2 mm. Abbreviations are explained in the text." figureDoi="http://doi.org/10.5281/zenodo.3837881" httpUri="https://zenodo.org/record/3837881/files/figure.png" pageId="9" pageNumber="10">Figs 5</figureCitation>
, 
<figureCitation id="1379F793FFF67D61AF92D709FDD09B57" box="[496,513,856,883]" captionStart="Fig" captionStartId="8.[189,232,1930,1956]" captionTargetBox="[189,1398,682,1888]" captionTargetId="figure@8.[189,1398,682,1888]" captionTargetPageId="8" captionText="Fig. 6. Tylopus hongkhraiensis sp. nov., holotype, Ƌ, left gonopod. A–B. Mesal and lateral views, respectively. C–F. Distal part, sublateral, submesal, suboral and subcaudal views, respectively. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3837885" httpUri="https://zenodo.org/record/3837885/files/figure.png" pageId="9" pageNumber="10">6</figureCitation>
); fermorite faintly curved, stout, slightly expanded distad, showing a strong mesal groove; process 
<emphasis id="B9363704FFF67D61AFAAD72AFE099BB1" bold="true" box="[456,472,891,917]" pageId="9" pageNumber="10">h</emphasis>
prominent, slightly curved, pointed; process 
<emphasis id="B9363704FFF67D61AD88D72AFC269BB1" bold="true" box="[1002,1015,891,917]" pageId="9" pageNumber="10">z</emphasis>
long and rather slender, evident, likewise curved and pointed; solenophore (
<emphasis id="B9363704FFF67D61ACD0D7CFFD0F9B9C" bold="true" box="[690,734,926,952]" pageId="9" pageNumber="10">sph</emphasis>
) expanded distally and flattened, typically coiled, clearly bifid, with a pair of small spines.
</paragraph>
</subSubSection>
<subSubSection id="C358B89DFFF67D61AEDFD052FCA79CB6" pageId="9" pageNumber="10" type="discussion">
<paragraph id="8BFDEB16FFF67D61AEDFD052FEFE9C39" blockId="9.[189,303,1027,1053]" box="[189,303,1027,1053]" pageId="9" pageNumber="10">
<heading id="D0B55C7AFFF67D61AEDFD052FEFE9C39" bold="true" box="[189,303,1027,1053]" fontSize="11" level="3" pageId="9" pageNumber="10" reason="3">
<emphasis id="B9363704FFF67D61AEDFD052FEFE9C39" bold="true" box="[189,303,1027,1053]" pageId="9" pageNumber="10">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="8BFDEB16FFF67D61AEDFD060FCA79CB6" blockId="9.[189,1399,1073,1170]" pageId="9" pageNumber="10">
The millipedes were found mating on bamboo logs on the soil surface during rain. The male was observed walking along the female’s back and stimulating her with rhythmic pulses of his legs. The tips of the solenophore and solenomere in 
<figureCitation id="1379F793FFF67D61AC1FD029FD139CB6" box="[637,706,1144,1170]" captionStart="Fig" captionStartId="8.[189,232,1930,1956]" captionTargetBox="[189,1398,682,1888]" captionTargetId="figure@8.[189,1398,682,1888]" captionTargetPageId="8" captionText="Fig. 6. Tylopus hongkhraiensis sp. nov., holotype, Ƌ, left gonopod. A–B. Mesal and lateral views, respectively. C–F. Distal part, sublateral, submesal, suboral and subcaudal views, respectively. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3837885" httpUri="https://zenodo.org/record/3837885/files/figure.png" pageId="9" pageNumber="10">Fig. 6</figureCitation>
are broken off.
</paragraph>
</subSubSection>
</treatment>
</document>