treatments-xml/data/03/EB/5A/03EB5A00FFF27D78ACE8D6A5FC149C4B.xml

<document id="E342091896CE048B78EAD6217711F094" ID-CLB-Dataset="21966" ID-DOI="10.5852/ejt.2016.195" ID-GBIF-Dataset="39071d6f-d900-4c81-ace1-dd03875018c5" ID-ISSN="2118-9773" ID-Zenodo-Dep="3837846" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1590071140602" checkinUser="valdenar" docAuthor="Likhitrakarn, Natdanai, Golovatch, Sergei I. &amp; Panha, Somsak" docDate="2016" docId="03EB5A00FFF27D78ACE8D6A5FC149C4B" docLanguage="en" docName="ejt-2016-195.pdf.imf" docOrigin="European Journal of Taxonomy 195" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Tylopus retusus Likhitrakarn &amp; Golovatch &amp; Panha 2016, sp. nov." docType="treatment" docUuid="40D74A7C-61D7-442B-B388-3868C47102C5" docUuidSource="ZooBank" docVersion="8" lastPageNumber="17" masterDocId="FFD22278FFFF7D68AE62D451FFD19824" masterDocTitle="The millipede genus Tylopus Jeekel, 1968 (Diplopoda, Polydesmida, Paradoxosomatidae), with a key and descriptions of eight new species from Indochina" masterLastPageNumber="47" masterPageNumber="1" pageNumber="14" updateTime="1698840626989" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0" zenodo-license-figures="CC-BY-3.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="F1F8DCFB56555F95A30D7EFF70B93594">The millipede genus Tylopus Jeekel, 1968 (Diplopoda, Polydesmida, Paradoxosomatidae), with a key and descriptions of eight new species from Indochina</mods:title>
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<mods:namePart id="F3B035199FF5AA2814F00EF2DB8DED0D">Likhitrakarn, Natdanai</mods:namePart>
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<mods:affiliation id="DB26D97E67C78C738A785F261EB650FA">Division of Plant Protection, Faculty of Agricultural Production, Maejo University, Chiang Mai 50290, Thailand. &amp; Email: kongerrrr @ hotmail. com &amp; urn: lsid: zoobank. org: author: 442 E 2 E 6 E- 1 CB 3 - 47 B 2 - 9896 - 1 A 922 CE 509 E 1</mods:affiliation>
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<mods:namePart id="FD5622D753C78FC1AC15EB241D96A339">Golovatch, Sergei I.</mods:namePart>
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<mods:affiliation id="5F65D563F5D965B80D0C2D676A9D5565">Institute for Problems of Ecology and Evolution, Russian Academy of Sciences, Leninsky pr. 33, Moscow 119071, Russia. &amp; Corresponding author: sgolovatch @ yandex. ru &amp; urn: lsid: zoobank. org: author: 71532 F 45 - BDD 5 - 415 D-BC 54 - 86256 E 5 D 5 D 4 A &amp; Corresponding author: sgolovatch @ yandex. ru &amp; Corresponding author: somsak. pan @ chula. ac. th</mods:affiliation>
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<mods:namePart id="1F5A3E2DE6DB8962F621B5FC038C1AA0">Panha, Somsak</mods:namePart>
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<mods:affiliation id="C19C493354471CB181CBC3B2BA8F9817">Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand. &amp; Corresponding author: somsak. pan @ chula. ac. th &amp; urn: lsid: zoobank. org: author: AC 935098 - D 901 - 4 F 35 - A 414 - 4 B 0 D 4 FE 44 E 79 &amp; Corresponding author: sgolovatch @ yandex. ru &amp; Corresponding author: somsak. pan @ chula. ac. th</mods:affiliation>
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<heading id="D0B55C7AFFF27D65ACE8D6A5FC7B9B2B" box="[650,938,756,783]" centered="true" fontSize="11" level="2" pageId="13" pageNumber="14" reason="2">
<taxonomicName id="4C429095FFF27D65ACE8D6A5FC999B2B" ID-CoL="7CZPY" authority="Likhitrakarn &amp; Golovatch &amp; Panha, 2016" authorityName="Likhitrakarn &amp; Golovatch &amp; Panha" authorityYear="2016" box="[650,840,756,783]" class="Diplopoda" family="Paradoxosomatidae" genus="Tylopus" kingdom="Animalia" order="Polydesmida" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="retusus" status="sp. nov.">
<emphasis id="B9363704FFF27D65ACE8D6A5FC999B2B" bold="true" box="[650,840,756,783]" italics="true" pageId="13" pageNumber="14">Tylopus retusus</emphasis>
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<figureCitation id="1379F793FFF27D65ACA1D76AFC979B71" box="[707,838,827,853]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="14.[189,232,1895,1921]" captionStartId-1="15.[189,232,1933,1959]" captionTargetBox-0="[189,1398,601,1850]" captionTargetBox-1="[241,1336,435,1859]" captionTargetId-0="figure@14.[189,1398,601,1850]" captionTargetId-1="figure@15.[238,1340,431,1892]" captionTargetPageId-0="14" captionTargetPageId-1="15" captionText-0="Fig. 10. Tylopus retusus sp. nov., holotype, Ƌ. A. Habitus, live coloration. B–C. Anterior part of body, dorsal and lateral views, respectively. D–E. Segments 10 and 11, dorsal and lateral views, respectively. F–H. Posterior part of body, dorsal, ventral and lateral views, respectively. I–J. Sternal cones between coxae 4, caudal and sublateral views, respectively." captionText-1="Fig. 11. Tylopus retusus sp. nov., holotype, Ƌ, left gonopod. A–D. Mesal, lateral, subcaudal and suboral views, respectively. E. Sterna of segment 10. F. Leg 6. G. Leg 10. Scale bars = 0.2 mm. Abbreviations are explained in the text." figureDoi-0="http://doi.org/10.5281/zenodo.3837893" figureDoi-1="http://doi.org/10.5281/zenodo.3837895" httpUri-0="https://zenodo.org/record/3837893/files/figure.png" httpUri-1="https://zenodo.org/record/3837895/files/figure.png" pageId="13" pageNumber="14">
Figs 10 
<emphasis id="B9363704FFF27D65AD78D76AFCF89B71" bold="true" box="[794,809,827,853]" pageId="13" pageNumber="14">–</emphasis>
11
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, 
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<heading id="D0B55C7AFFF27D65AEDFD72DFEE79BB2" bold="true" box="[189,310,892,918]" fontSize="11" level="3" pageId="13" pageNumber="14" reason="3">
<emphasis id="B9363704FFF27D65AEDFD72DFEE79BB2" bold="true" box="[189,310,892,918]" pageId="13" pageNumber="14">Diagnosis</emphasis>
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Distinguished by its relatively small body with four transverse rows of setae forming no regular pattern of setation; in gonopod structure it is similar to 
<taxonomicName id="4C429095FFF27D65AC84D79EFB239BCC" authority="Golovatch &amp; Enghoff, 1993" authorityName="Golovatch &amp; Enghoff" authorityYear="1993" box="[742,1266,974,1000]" class="Diplopoda" family="Paradoxosomatidae" genus="Tylopus" kingdom="Animalia" order="Polydesmida" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="haplorugosus">
<emphasis id="B9363704FFF27D65AC84D79EFC739BCC" box="[742,930,974,1000]" italics="true" pageId="13" pageNumber="14">T. haplorugosus</emphasis>
<bibRefCitation id="EFD396E7FFF27D65ADCBD79FFB239BCC" author="Golovatch S. I. &amp; Enghoff H." box="[937,1266,974,1000]" pageId="13" pageNumber="14" pagination="85 - 125" refId="ref19643" refString="Golovatch S. I. &amp; Enghoff H. 1993. Review of the millipede genus Tylopus, with descriptions of new species from Thailand (Diplopoda, Polydesmida, Paradoxosomatidae). Steenstrupia 19 (3): 85 - 125." type="journal article" year="1993">Golovatch &amp; Enghoff, 1993</bibRefCitation>
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, but differs in process 
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being rather short, slightly flattened, blunt, coupled with the solenophore being strongly coiled.
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<emphasis id="B9363704FFF27D65AEDFD007FE959C54" bold="true" box="[189,324,1110,1136]" pageId="13" pageNumber="14">Etymology</emphasis>
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To emphasize the blunt tip of process 
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; adjective.
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<heading id="D0B55C7AFFF27D65AEDFD097FE7B9CC4" bold="true" box="[189,426,1222,1248]" fontSize="11" level="3" pageId="13" pageNumber="14" reason="3">
<emphasis id="B9363704FFF27D65AEDFD097FE7B9CC4" bold="true" box="[189,426,1222,1248]" pageId="13" pageNumber="14">Material examined</emphasis>
</heading>
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<collectingCountry id="F355AB86FFF27D65AEDFD149FEDF9D16" box="[189,270,1304,1330]" name="Laos" pageId="13" pageNumber="14">LAOS</collectingCountry>
: Ƌ, 
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, 
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, 
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, 
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, 
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, ca 
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a.s.l.
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, mango garden, 
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, leg. 
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&amp; 
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. 
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(
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).
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</emphasis>
</paragraph>
<paragraph id="8BFDEB16FFF27D65AEDFD1F1FCD19DF9" blockId="13.[189,1399,1405,1501]" pageId="13" pageNumber="14">
<collectingCountry id="F355AB86FFF27D65AEDFD1F1FEDF9D9E" box="[189,270,1440,1466]" name="Laos" pageId="13" pageNumber="14">LAOS</collectingCountry>
: 
<specimenCount id="9D44209FFFF27D65AF79D1F1FE989D9E" box="[283,329,1440,1466]" pageId="13" pageNumber="14" type="female">1 ♀</specimenCount>
, Khoua, 
<collectingRegion id="498625F4FFF27D65AFDAD1F1FDE29D9E" box="[440,563,1440,1466]" country="Laos" name="Phongsali" pageId="13" pageNumber="14">Phongsaly</collectingRegion>
, ca 
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a.s.l., 
<geoCoordinate id="EE768DD1FFF27D65AD62D1CEFC449D9E" box="[768,917,1439,1466]" degrees="21" direction="north" minutes="11" orientation="latitude" pageId="13" pageNumber="14" precision="15" seconds="55" value="21.19861">21°11'55&quot; N</geoCoordinate>
, 
<geoCoordinate id="EE768DD1FFF27D65ADC1D1CEFB959D9E" box="[931,1092,1439,1466]" degrees="102" direction="east" minutes="06" orientation="longitude" pageId="13" pageNumber="14" precision="15" seconds="40" value="102.11111">102°06'40&quot; E</geoCoordinate>
, forest near road, 
<date id="FFFCCDD6FFF27D65AB7CD1CEFF2A9DF9" pageId="13" pageNumber="14" value="2014-10-15">15 Oct. 2014</date>
, leg. C. Sutcharit &amp; 
<collectionCode id="ED5373D3FFF27D65AF88D192FE2F9DF9" box="[490,510,1475,1501]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="13" pageNumber="14">R</collectionCode>
. Srisonchai (
<collectionCode id="ED5373D3FFF27D65ACF8D192FD259DF9" box="[666,756,1475,1501]" country="0" httpUri="http://grbio.org/cool/da58-p0xp" name="Cameroon University, Museum of Zoology" pageId="13" pageNumber="14">CUMZ</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="C358B89DFFF27D78AEDFD255FC2B9BDB" lastPageId="16" lastPageNumber="17" pageId="13" pageNumber="14" type="description">
<paragraph id="8BFDEB16FFF27D65AEDFD255FE9F9E3A" blockId="13.[189,334,1540,1566]" box="[189,334,1540,1566]" pageId="13" pageNumber="14">
<heading id="D0B55C7AFFF27D65AEDFD255FE9F9E3A" bold="true" box="[189,334,1540,1566]" fontSize="11" level="3" pageId="13" pageNumber="14" reason="3">
<emphasis id="B9363704FFF27D65AEDFD255FE9F9E3A" bold="true" box="[189,334,1540,1566]" pageId="13" pageNumber="14">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8BFDEB16FFF27D65AEDFD262FC859EFE" blockId="13.[189,1399,1587,1754]" pageId="13" pageNumber="14">
MEASUREMENTS AND COLOUR. Length 16.3 (Ƌ) or 
<quantity id="4CBA46F3FFF27D65AC86D262FC9B9E69" box="[740,842,1587,1613]" metricMagnitude="-2" metricUnit="m" metricValue="2.04" pageId="13" pageNumber="14" unit="mm" value="20.4">20.4 mm</quantity>
(
<specimenCount id="9D44209FFFF27D65AD39D262FCBF9E69" box="[859,878,1587,1613]" pageId="13" pageNumber="14" type="female">♀</specimenCount>
), width of midbody pro- and metazonae 1.12 and 
<quantity id="4CBA46F3FFF27D65AE8CD207FE879E54" box="[238,342,1622,1648]" metricMagnitude="-3" metricUnit="m" metricValue="2.07" pageId="13" pageNumber="14" unit="mm" value="2.07">2.07 mm</quantity>
(Ƌ) or 2.7 and 
<quantity id="4CBA46F3FFF27D65AC6AD207FDB09E54" box="[520,609,1622,1648]" metricMagnitude="-3" metricUnit="m" metricValue="3.7" pageId="13" pageNumber="14" unit="mm" value="3.7">3.7 mm</quantity>
(
<specimenCount id="9D44209FFFF27D65AC11D207FD579E54" box="[627,646,1622,1648]" pageId="13" pageNumber="14" type="female">♀</specimenCount>
), respectively. Live coloration light brown (
<figureCitation id="1379F793FFF27D65AAE8D207FB279E54" box="[1162,1270,1622,1648]" captionStart="Fig" captionStartId="14.[189,232,1895,1921]" captionTargetBox="[189,1398,601,1850]" captionTargetId="figure@14.[189,1398,601,1850]" captionTargetPageId="14" captionText="Fig. 10. Tylopus retusus sp. nov., holotype, Ƌ. A. Habitus, live coloration. B–C. Anterior part of body, dorsal and lateral views, respectively. D–E. Segments 10 and 11, dorsal and lateral views, respectively. F–H. Posterior part of body, dorsal, ventral and lateral views, respectively. I–J. Sternal cones between coxae 4, caudal and sublateral views, respectively." figureDoi="http://doi.org/10.5281/zenodo.3837893" httpUri="https://zenodo.org/record/3837893/files/figure.png" pageId="13" pageNumber="14">Fig. 10A</figureCitation>
); prozonae dark brown; paraterga, antennae, head and legs light brown to yellow-brown. Coloration of alcohol material after one year of preservation faded to light brown to yellow-brown; paraterga, antennae, head, legs and venter light brown to light yellow (
<figureCitation id="1379F793FFF27D65ACA2D2EEFCC59EFE" box="[704,788,1727,1754]" captionStart="Fig" captionStartId="14.[189,232,1895,1921]" captionTargetBox="[189,1398,601,1850]" captionTargetId="figure@14.[189,1398,601,1850]" captionTargetPageId="14" captionText="Fig. 10. Tylopus retusus sp. nov., holotype, Ƌ. A. Habitus, live coloration. B–C. Anterior part of body, dorsal and lateral views, respectively. D–E. Segments 10 and 11, dorsal and lateral views, respectively. F–H. Posterior part of body, dorsal, ventral and lateral views, respectively. I–J. Sternal cones between coxae 4, caudal and sublateral views, respectively." figureDoi="http://doi.org/10.5281/zenodo.3837893" httpUri="https://zenodo.org/record/3837893/files/figure.png" pageId="13" pageNumber="14">Fig. 10</figureCitation>
B–J).
</paragraph>
<paragraph id="8BFDEB16FFF27D65AEDFD350FC439FEF" blockId="13.[189,1399,1793,1995]" pageId="13" pageNumber="14">
HEAD. Clypeolabral region and vertex sparsely setose, epicranial suture distinct. Antennae (
<figureCitation id="1379F793FFF27D65AB61D350FF009F1B" captionStart="Fig" captionStartId="14.[189,232,1895,1921]" captionTargetBox="[189,1398,601,1850]" captionTargetId="figure@14.[189,1398,601,1850]" captionTargetPageId="14" captionText="Fig. 10. Tylopus retusus sp. nov., holotype, Ƌ. A. Habitus, live coloration. B–C. Anterior part of body, dorsal and lateral views, respectively. D–E. Segments 10 and 11, dorsal and lateral views, respectively. F–H. Posterior part of body, dorsal, ventral and lateral views, respectively. I–J. Sternal cones between coxae 4, caudal and sublateral views, respectively." figureDoi="http://doi.org/10.5281/zenodo.3837893" httpUri="https://zenodo.org/record/3837893/files/figure.png" pageId="13" pageNumber="14">Fig. 10A, B</figureCitation>
) rather short and stout, antennomeres 2–6 subequal in length, reaching body segment 3 (Ƌ, 
<specimenCount id="9D44209FFFF27D65AB6DD374FAF29F1B" box="[1295,1315,1829,1855]" pageId="13" pageNumber="14" type="female">♀</specimenCount>
) when stretched dorsally. In width, head &lt;segment 3 &lt;4 &lt;collum &lt;segments 5–15; thereafter body gently and gradually tapering. Collum with three transverse rows of setae: 6+6 anterior, 4+4 intermediate and 6+6 posterior; a small lateral incision at about midway; caudal corner very broadly rounded, paraterga declined ventrad, slightly produced behind rear tergal margin.
</paragraph>
<paragraph id="8BFDEB16FFF17D78AEDFD55DFD499A47" blockId="14.[189,1399,267,540]" lastBlockId="16.[189,1399,268,612]" lastPageId="16" lastPageNumber="17" pageId="14" pageNumber="15">
BODY. Tegument smooth and shining, prozonae finely shagreened, metaterga leathery, finely rugulose (
<figureCitation id="1379F793FFF17D66AEA4D57FFE5B996D" box="[198,394,302,329]" captionStart="Fig" captionStartId="14.[189,232,1895,1921]" captionTargetBox="[189,1398,601,1850]" captionTargetId="figure@14.[189,1398,601,1850]" captionTargetPageId="14" captionText="Fig. 10. Tylopus retusus sp. nov., holotype, Ƌ. A. Habitus, live coloration. B–C. Anterior part of body, dorsal and lateral views, respectively. D–E. Segments 10 and 11, dorsal and lateral views, respectively. F–H. Posterior part of body, dorsal, ventral and lateral views, respectively. I–J. Sternal cones between coxae 4, caudal and sublateral views, respectively." figureDoi="http://doi.org/10.5281/zenodo.3837893" httpUri="https://zenodo.org/record/3837893/files/figure.png" pageId="14" pageNumber="15">Fig. 10B, D, F</figureCitation>
); surface below paraterga roughly microgranulate (
<figureCitation id="1379F793FFF17D66AA72D57FFB07996D" box="[1040,1238,302,329]" captionStart="Fig" captionStartId="14.[189,232,1895,1921]" captionTargetBox="[189,1398,601,1850]" captionTargetId="figure@14.[189,1398,601,1850]" captionTargetPageId="14" captionText="Fig. 10. Tylopus retusus sp. nov., holotype, Ƌ. A. Habitus, live coloration. B–C. Anterior part of body, dorsal and lateral views, respectively. D–E. Segments 10 and 11, dorsal and lateral views, respectively. F–H. Posterior part of body, dorsal, ventral and lateral views, respectively. I–J. Sternal cones between coxae 4, caudal and sublateral views, respectively." figureDoi="http://doi.org/10.5281/zenodo.3837893" httpUri="https://zenodo.org/record/3837893/files/figure.png" pageId="14" pageNumber="15">Fig. 10C, E, H</figureCitation>
). Postcollum metaterga with four transverse rows of setae forming no regulated pattern of setation, anterior row with the most dense setae growing increasingly dense towards segment 19 (
<figureCitation id="1379F793FFF17D66AA7AD524FB0199AB" box="[1048,1232,373,399]" captionStart="Fig" captionStartId="14.[189,232,1895,1921]" captionTargetBox="[189,1398,601,1850]" captionTargetId="figure@14.[189,1398,601,1850]" captionTargetPageId="14" captionText="Fig. 10. Tylopus retusus sp. nov., holotype, Ƌ. A. Habitus, live coloration. B–C. Anterior part of body, dorsal and lateral views, respectively. D–E. Segments 10 and 11, dorsal and lateral views, respectively. F–H. Posterior part of body, dorsal, ventral and lateral views, respectively. I–J. Sternal cones between coxae 4, caudal and sublateral views, respectively." figureDoi="http://doi.org/10.5281/zenodo.3837893" httpUri="https://zenodo.org/record/3837893/files/figure.png" pageId="14" pageNumber="15">Fig. 10B, D, F</figureCitation>
). Tergal setae long and slender, about 2/3 as long as metaterga. Axial line well visible on metazonae, traceable also on prozonae. Paraterga strongly developed (
<figureCitation id="1379F793FFF17D66ACFED5EAFD2199F1" box="[668,752,443,469]" captionStart="Fig" captionStartId="14.[189,232,1895,1921]" captionTargetBox="[189,1398,601,1850]" captionTargetId="figure@14.[189,1398,601,1850]" captionTargetPageId="14" captionText="Fig. 10. Tylopus retusus sp. nov., holotype, Ƌ. A. Habitus, live coloration. B–C. Anterior part of body, dorsal and lateral views, respectively. D–E. Segments 10 and 11, dorsal and lateral views, respectively. F–H. Posterior part of body, dorsal, ventral and lateral views, respectively. I–J. Sternal cones between coxae 4, caudal and sublateral views, respectively." figureDoi="http://doi.org/10.5281/zenodo.3837893" httpUri="https://zenodo.org/record/3837893/files/figure.png" pageId="14" pageNumber="15">Fig. 10</figureCitation>
B–H), especially so in Ƌ, set rather high (at upper 1/3 of body height), slightly upturned, but lying below dorsum; anterior edge broadly rounded and narrowly bordered, fused to callus; caudal corner very narrowly rounded, starting with segment 16 extending increasingly behind rear tergal margin (
<figureCitation id="1379F793FFF07D67ACFAD55AFCF29902" box="[664,803,267,294]" captionStart="Fig" captionStartId="14.[189,232,1895,1921]" captionTargetBox="[189,1398,601,1850]" captionTargetId="figure@14.[189,1398,601,1850]" captionTargetPageId="14" captionText="Fig. 10. Tylopus retusus sp. nov., holotype, Ƌ. A. Habitus, live coloration. B–C. Anterior part of body, dorsal and lateral views, respectively. D–E. Segments 10 and 11, dorsal and lateral views, respectively. F–H. Posterior part of body, dorsal, ventral and lateral views, respectively. I–J. Sternal cones between coxae 4, caudal and sublateral views, respectively." figureDoi="http://doi.org/10.5281/zenodo.3837893" httpUri="https://zenodo.org/record/3837893/files/figure.png" pageId="15" pageNumber="16">Fig. 10F, H</figureCitation>
). Paraterga 2 and 3 with two evident incisions at lateral edge (
<figureCitation id="1379F793FFF07D67AF34D57FFE13996D" box="[342,450,302,329]" captionStart="Fig" captionStartId="14.[189,232,1895,1921]" captionTargetBox="[189,1398,601,1850]" captionTargetId="figure@14.[189,1398,601,1850]" captionTargetPageId="14" captionText="Fig. 10. Tylopus retusus sp. nov., holotype, Ƌ. A. Habitus, live coloration. B–C. Anterior part of body, dorsal and lateral views, respectively. D–E. Segments 10 and 11, dorsal and lateral views, respectively. F–H. Posterior part of body, dorsal, ventral and lateral views, respectively. I–J. Sternal cones between coxae 4, caudal and sublateral views, respectively." figureDoi="http://doi.org/10.5281/zenodo.3837893" httpUri="https://zenodo.org/record/3837893/files/figure.png" pageId="15" pageNumber="16">Fig. 10B</figureCitation>
). Following poreless segments with two similar incisions; pore-boring segments with one, often setigerous incision lying in front of pore (
<figureCitation id="1379F793FFF07D67AD01D503FC229948" box="[867,1011,338,364]" captionStart="Fig" captionStartId="14.[189,232,1895,1921]" captionTargetBox="[189,1398,601,1850]" captionTargetId="figure@14.[189,1398,601,1850]" captionTargetPageId="14" captionText="Fig. 10. Tylopus retusus sp. nov., holotype, Ƌ. A. Habitus, live coloration. B–C. Anterior part of body, dorsal and lateral views, respectively. D–E. Segments 10 and 11, dorsal and lateral views, respectively. F–H. Posterior part of body, dorsal, ventral and lateral views, respectively. I–J. Sternal cones between coxae 4, caudal and sublateral views, respectively." figureDoi="http://doi.org/10.5281/zenodo.3837893" httpUri="https://zenodo.org/record/3837893/files/figure.png" pageId="15" pageNumber="16">Fig. 10D, E</figureCitation>
). Calluses of paraterga delimited by a sulcus both dorsally and ventrally. Ozopores evident, lateral, lying in an ovoid groove at about 1/3 of metatergal length in front of posterior edge of metaterga. Transverse sulcus usually distinct (
<figureCitation id="1379F793FFEF7D78AB64D57EFF249949" captionStart="Fig" captionStartId="14.[189,232,1895,1921]" captionTargetBox="[189,1398,601,1850]" captionTargetId="figure@14.[189,1398,601,1850]" captionTargetPageId="14" captionText="Fig. 10. Tylopus retusus sp. nov., holotype, Ƌ. A. Habitus, live coloration. B–C. Anterior part of body, dorsal and lateral views, respectively. D–E. Segments 10 and 11, dorsal and lateral views, respectively. F–H. Posterior part of body, dorsal, ventral and lateral views, respectively. I–J. Sternal cones between coxae 4, caudal and sublateral views, respectively." figureDoi="http://doi.org/10.5281/zenodo.3837893" httpUri="https://zenodo.org/record/3837893/files/figure.png" pageId="16" pageNumber="17">Fig. 10B, D, F</figureCitation>
), slightly incomplete on segment 19, complete on segments 5–18, narrow, undulate, shallow, not reaching bases of paraterga, beaded at bottom. Stricture between pro- and metazonae evident, wide and deep, beaded at bottom down to base of paraterga (
<figureCitation id="1379F793FFEF7D78AD2AD5C8FC269990" box="[840,1015,409,436]" captionStart="Fig" captionStartId="14.[189,232,1895,1921]" captionTargetBox="[189,1398,601,1850]" captionTargetId="figure@14.[189,1398,601,1850]" captionTargetPageId="14" captionText="Fig. 10. Tylopus retusus sp. nov., holotype, Ƌ. A. Habitus, live coloration. B–C. Anterior part of body, dorsal and lateral views, respectively. D–E. Segments 10 and 11, dorsal and lateral views, respectively. F–H. Posterior part of body, dorsal, ventral and lateral views, respectively. I–J. Sternal cones between coxae 4, caudal and sublateral views, respectively." figureDoi="http://doi.org/10.5281/zenodo.3837893" httpUri="https://zenodo.org/record/3837893/files/figure.png" pageId="16" pageNumber="17">
Fig. 10B, D 
<emphasis id="B9363704FFEF7D78ADB6D5C8FC329997" bold="true" box="[980,995,409,435]" pageId="16" pageNumber="17">–</emphasis>
F
</figureCitation>
). Pleurosternal carinae complete crests with a sharp caudal tooth on segments 2–7 (Ƌ) or 2–4 (
<specimenCount id="9D44209FFFEF7D78ADD7D5EDFC1999F2" box="[949,968,444,470]" pageId="16" pageNumber="17" type="female">♀</specimenCount>
), thereafter increasingly reduced to a small tooth until segment 15 (Ƌ) or 13 (
<specimenCount id="9D44209FFFEF7D78ACD8D5B1FD1C99DE" box="[698,717,480,506]" pageId="16" pageNumber="17" type="female">♀</specimenCount>
), missing more caudally. Epiproct (
<figureCitation id="1379F793FFEF7D78AA1FD58EFB0399DE" box="[1149,1234,479,506]" captionStart="Fig" captionStartId="14.[189,232,1895,1921]" captionTargetBox="[189,1398,601,1850]" captionTargetId="figure@14.[189,1398,601,1850]" captionTargetPageId="14" captionText="Fig. 10. Tylopus retusus sp. nov., holotype, Ƌ. A. Habitus, live coloration. B–C. Anterior part of body, dorsal and lateral views, respectively. D–E. Segments 10 and 11, dorsal and lateral views, respectively. F–H. Posterior part of body, dorsal, ventral and lateral views, respectively. I–J. Sternal cones between coxae 4, caudal and sublateral views, respectively." figureDoi="http://doi.org/10.5281/zenodo.3837893" httpUri="https://zenodo.org/record/3837893/files/figure.png" pageId="16" pageNumber="17">Fig. 10</figureCitation>
F–H) conical, flattened dorsoventrally, with two evident, large, rounded apical papillae; tip subtruncate; lateral preapical papillae small, lying close to tip. Hypoproct (
<figureCitation id="1379F793FFEF7D78AD7AD677FC539A64" box="[792,898,550,576]" captionStart="Fig" captionStartId="14.[189,232,1895,1921]" captionTargetBox="[189,1398,601,1850]" captionTargetId="figure@14.[189,1398,601,1850]" captionTargetPageId="14" captionText="Fig. 10. Tylopus retusus sp. nov., holotype, Ƌ. A. Habitus, live coloration. B–C. Anterior part of body, dorsal and lateral views, respectively. D–E. Segments 10 and 11, dorsal and lateral views, respectively. F–H. Posterior part of body, dorsal, ventral and lateral views, respectively. I–J. Sternal cones between coxae 4, caudal and sublateral views, respectively." figureDoi="http://doi.org/10.5281/zenodo.3837893" httpUri="https://zenodo.org/record/3837893/files/figure.png" pageId="16" pageNumber="17">Fig. 10G</figureCitation>
) roundly subtrapeziform, setigerous knobs at caudal edge small and well-separated.
</paragraph>
<caption id="DF3DBB9EFFF17D66AEDFD336FCDD9FCF" ID-DOI="http://doi.org/10.5281/zenodo.3837893" ID-Zenodo-Dep="3837893" httpUri="https://zenodo.org/record/3837893/files/figure.png" pageId="14" pageNumber="15" startId="14.[189,232,1895,1921]" targetBox="[189,1398,601,1850]" targetPageId="14">
<paragraph id="8BFDEB16FFF17D66AEDFD336FCDD9FCF" blockId="14.[189,1399,1895,2028]" pageId="14" pageNumber="15">
<emphasis id="B9363704FFF17D66AEDFD336FECB9FA6" bold="true" box="[189,282,1895,1922]" pageId="14" pageNumber="15">Fig. 10.</emphasis>
<taxonomicName id="4C429095FFF17D66AF40D339FE089FA6" authority="Likhitrakarn &amp; Golovatch &amp; Panha, 2016" authorityName="Likhitrakarn &amp; Golovatch &amp; Panha" authorityYear="2016" box="[290,473,1896,1922]" class="Diplopoda" family="Paradoxosomatidae" genus="Tylopus" kingdom="Animalia" order="Polydesmida" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="retusus" status="sp. nov.">
<emphasis id="B9363704FFF17D66AF40D339FE089FA6" box="[290,473,1896,1922]" italics="true" pageId="14" pageNumber="15">Tylopus retusus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2058A7FFFF17D66AF82D339FDEF9FA6" box="[480,574,1896,1922]" pageId="14" pageNumber="15" rank="species">sp. nov.</taxonomicNameLabel>
, holotype, Ƌ. 
<emphasis id="B9363704FFF17D66AC84D336FD2A9FA5" bold="true" box="[742,763,1895,1921]" pageId="14" pageNumber="15">A</emphasis>
. Habitus, live coloration. 
<emphasis id="B9363704FFF17D66AA52D336FBB69FA5" bold="true" box="[1072,1127,1895,1921]" pageId="14" pageNumber="15">B–C</emphasis>
. Anterior part of body, dorsal and lateral views, respectively. 
<emphasis id="B9363704FFF17D66AC1FD3DAFD659F81" bold="true" box="[637,692,1931,1957]" pageId="14" pageNumber="15">D–E</emphasis>
. Segments 10 and 11, dorsal and lateral views, respectively. 
<emphasis id="B9363704FFF17D66AEDFD3FFFF259FEC" bold="true" box="[189,244,1966,1992]" pageId="14" pageNumber="15">F–H</emphasis>
. Posterior part of body, dorsal, ventral and lateral views, respectively. 
<emphasis id="B9363704FFF17D66AA55D3FFFBB19FEC" bold="true" box="[1079,1120,1966,1992]" pageId="14" pageNumber="15">I–J</emphasis>
. Sternal cones between coxae 4, caudal and sublateral views, respectively.
</paragraph>
</caption>
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<paragraph id="8BFDEB16FFF07D67AEDFD3DCFE0E9FCA" blockId="15.[189,1398,1933,2030]" pageId="15" pageNumber="16">
<emphasis id="B9363704FFF07D67AEDFD3DCFEC69F83" bold="true" box="[189,279,1933,1959]" pageId="15" pageNumber="16">Fig. 11.</emphasis>
<taxonomicName id="4C429095FFF07D67AF7CD3DCFE029F83" authority="Likhitrakarn &amp; Golovatch &amp; Panha, 2016" authorityName="Likhitrakarn &amp; Golovatch &amp; Panha" authorityYear="2016" box="[286,467,1933,1959]" class="Diplopoda" family="Paradoxosomatidae" genus="Tylopus" kingdom="Animalia" order="Polydesmida" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="retusus" status="sp. nov.">
<emphasis id="B9363704FFF07D67AF7CD3DCFE029F83" box="[286,467,1933,1959]" italics="true" pageId="15" pageNumber="16">Tylopus retusus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2058A7FFFF07D67AFB8D3DCFDE69F83" box="[474,567,1933,1959]" pageId="15" pageNumber="16" rank="species">sp. nov.</taxonomicNameLabel>
, holotype, Ƌ, left gonopod. 
<emphasis id="B9363704FFF07D67AD1CD3DCFC669F83" bold="true" box="[894,951,1933,1959]" pageId="15" pageNumber="16">A–D</emphasis>
. Mesal, lateral, subcaudal and suboral views, respectively. 
<emphasis id="B9363704FFF07D67AFC9D3E1FE6E9FEE" bold="true" box="[427,447,1968,1994]" pageId="15" pageNumber="16">E</emphasis>
. Sterna of segment 10. 
<emphasis id="B9363704FFF07D67ACB4D3E1FD399FEE" bold="true" box="[726,744,1968,1994]" pageId="15" pageNumber="16">F</emphasis>
. Leg 6. 
<emphasis id="B9363704FFF07D67AD28D3E1FCB09FEE" bold="true" box="[842,865,1968,1994]" pageId="15" pageNumber="16">G</emphasis>
. Leg 10. Scale bars = 0.2 mm. Abbreviations are explained in the text.
</paragraph>
</caption>
<paragraph id="8BFDEB16FFEF7D78AEDFD6DAFE1D9B71" blockId="16.[189,1399,650,853]" pageId="16" pageNumber="17">
STERNA. Densely setose, with a small cone caudally near each coxa, rear cones being a little larger than fore ones (
<figureCitation id="1379F793FFEF7D78AF5FD6FFFE779AEC" box="[317,422,686,712]" captionStart="Fig" captionStartId="15.[189,232,1933,1959]" captionTargetBox="[241,1336,435,1859]" captionTargetId="figure@15.[238,1340,431,1892]" captionTargetPageId="15" captionText="Fig. 11. Tylopus retusus sp. nov., holotype, Ƌ, left gonopod. A–D. Mesal, lateral, subcaudal and suboral views, respectively. E. Sterna of segment 10. F. Leg 6. G. Leg 10. Scale bars = 0.2 mm. Abbreviations are explained in the text." figureDoi="http://doi.org/10.5281/zenodo.3837895" httpUri="https://zenodo.org/record/3837895/files/figure.png" pageId="16" pageNumber="17">Fig. 11E</figureCitation>
); two small, rough, fully separated, setose cones between Ƌ coxae 4 (
<figureCitation id="1379F793FFEF7D78AA88D6FFFAB99AEC" box="[1258,1384,686,712]" captionStart="Fig" captionStartId="14.[189,232,1895,1921]" captionTargetBox="[189,1398,601,1850]" captionTargetId="figure@14.[189,1398,601,1850]" captionTargetPageId="14" captionText="Fig. 10. Tylopus retusus sp. nov., holotype, Ƌ. A. Habitus, live coloration. B–C. Anterior part of body, dorsal and lateral views, respectively. D–E. Segments 10 and 11, dorsal and lateral views, respectively. F–H. Posterior part of body, dorsal, ventral and lateral views, respectively. I–J. Sternal cones between coxae 4, caudal and sublateral views, respectively." figureDoi="http://doi.org/10.5281/zenodo.3837893" httpUri="https://zenodo.org/record/3837893/files/figure.png" pageId="16" pageNumber="17">Fig. 10I, J</figureCitation>
). Legs rather long and slender, midbody ones ca 1.2–1.3 (Ƌ) or 0.9–1.0 (
<specimenCount id="9D44209FFFEF7D78AA4DD680FB939ACF" box="[1071,1090,721,747]" pageId="16" pageNumber="17" type="female">♀</specimenCount>
) as long as body height; all male legs until segment 16 with tarsal brushes, prefemora not swollen; prefemora, femora and tibiae with ventral microgranulations; femora, tibiae and tarsi each with an evident adenostyle (tubercle) medially (
<figureCitation id="1379F793FFEF7D78AF56D76BFE6E9B71" box="[308,447,826,853]" captionStart="Fig" captionStartId="15.[189,232,1933,1959]" captionTargetBox="[241,1336,435,1859]" captionTargetId="figure@15.[238,1340,431,1892]" captionTargetPageId="15" captionText="Fig. 11. Tylopus retusus sp. nov., holotype, Ƌ, left gonopod. A–D. Mesal, lateral, subcaudal and suboral views, respectively. E. Sterna of segment 10. F. Leg 6. G. Leg 10. Scale bars = 0.2 mm. Abbreviations are explained in the text." figureDoi="http://doi.org/10.5281/zenodo.3837895" httpUri="https://zenodo.org/record/3837895/files/figure.png" pageId="16" pageNumber="17">
Fig. 11F 
<emphasis id="B9363704FFEF7D78AFF5D76AFE779B71" bold="true" box="[407,422,827,853]" pageId="16" pageNumber="17">–</emphasis>
G
</figureCitation>
).
</paragraph>
<paragraph id="8BFDEB16FFEF7D78AEDFD72DFC2B9BDB" blockId="16.[189,1399,892,1024]" pageId="16" pageNumber="17">
GONOPOD. Rather simple (
<figureCitation id="1379F793FFEF7D78AF98D72DFD809BB2" box="[506,593,892,918]" captionStart="Fig" captionStartId="15.[189,232,1933,1959]" captionTargetBox="[241,1336,435,1859]" captionTargetId="figure@15.[238,1340,431,1892]" captionTargetPageId="15" captionText="Fig. 11. Tylopus retusus sp. nov., holotype, Ƌ, left gonopod. A–D. Mesal, lateral, subcaudal and suboral views, respectively. E. Sterna of segment 10. F. Leg 6. G. Leg 10. Scale bars = 0.2 mm. Abbreviations are explained in the text." figureDoi="http://doi.org/10.5281/zenodo.3837895" httpUri="https://zenodo.org/record/3837895/files/figure.png" pageId="16" pageNumber="17">Fig. 11</figureCitation>
A–D); coxa a little curved caudad, sparsely setose distoventrally. Femorite slightly curved and stout, with an evident mesal groove and a clear distolateral sulcus demarcating a postfemoral part; lobe 
<emphasis id="B9363704FFEF7D78AC1AD792FD519BF9" bold="true" box="[632,640,963,989]" pageId="16" pageNumber="17">l</emphasis>
evident, rounded, with process 
<emphasis id="B9363704FFEF7D78AD99D792FBDA9BF9" bold="true" box="[1019,1035,963,989]" pageId="16" pageNumber="17">h</emphasis>
rather short, slightly flattened, blunt; solenophore (
<emphasis id="B9363704FFEF7D78AFCBD7B7FE049C24" bold="true" box="[425,469,998,1024]" pageId="16" pageNumber="17">sph</emphasis>
) strongly coiled, devoid of processes, pointed.
</paragraph>
</subSubSection>
<subSubSection id="C358B89DFFEF7D78AEDFD076FC149C4B" box="[189,965,1063,1136]" pageId="16" pageNumber="17" type="discussion">
<paragraph id="8BFDEB16FFEF7D78AEDFD076FEF59C65" blockId="16.[189,292,1063,1089]" box="[189,292,1063,1089]" pageId="16" pageNumber="17">
<heading id="D0B55C7AFFEF7D78AEDFD076FEF59C65" bold="true" box="[189,292,1063,1089]" fontSize="11" level="3" pageId="16" pageNumber="17" reason="3">
<emphasis id="B9363704FFEF7D78AEDFD076FEF59C65" bold="true" box="[189,292,1063,1089]" pageId="16" pageNumber="17">Remark</emphasis>
</heading>
</paragraph>
<paragraph id="8BFDEB16FFEF7D78AEDFD004FC149C4B" blockId="16.[189,965,1109,1136]" box="[189,965,1109,1136]" pageId="16" pageNumber="17">This species was found in a mango plantation in Muang Xay City.</paragraph>
</subSubSection>
</treatment>
</document>