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<document id="57F9EDA85CFB424457218F96D3F21B10" ID-DOI="10.5281/zenodo.5042023" ID-ISSN="1942-1354" ID-Zenodo-Dep="5042023" ID-ZooBank="1738B3CE-22AC-409B-9B04-DAD91322B278" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1624979611302" checkinUser="carolina" docAuthor="Phillips-Rodríguez, Eugenie, Brown, John W., Hallwachs, Winnie &amp; Janzen, Daniel H." docDate="2021" docId="03EB577BFFADF076FF67FD54FEF6FC04" docLanguage="en" docName="InsectaMundi.0868.1-96.pdf" docOrigin="Insecta Mundi 2021 (868)" docStyle="DocumentStyle:F9976E9B7F8A094D890004515A7A0592.3:InsectaMundi.2020-.journal_article" docStyleId="F9976E9B7F8A094D890004515A7A0592" docStyleName="InsectaMundi.2020-.journal_article" docStyleVersion="3" docTitle="Chlamydastis gladysrojasae Phillips and Brown 2021, new species" docType="treatment" docVersion="3" lastPageNumber="27" masterDocId="FFD22F03FFB6F06AFFA7FFA9FFE1FFE2" masterDocTitle="Chlamydastis Meyrick of Costa Rica: barcodes, biology, and descriptions of 36 new species (Lepidoptera: Depressariidae)" masterLastPageNumber="96" masterPageNumber="1" pageNumber="26" updateTime="1698978567902" updateUser="plazi" zenodo-license-document="CC-BY-NC-3.0">
<mods:mods id="32476E9B2D5D43EEF622096D6ABE26E9" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="8BC9E865AE0E391BF8C214F075C1E86D">Chlamydastis Meyrick of Costa Rica: barcodes, biology, and descriptions of 36 new species (Lepidoptera: Depressariidae)</mods:title>
</mods:titleInfo>
<mods:name id="3028C8CD9E0B1D6657E86798893B7BC0" type="personal">
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<mods:namePart id="5761D25D97D78F853D4D3612E6BBDC89">Phillips-Rodríguez, Eugenie</mods:namePart>
<mods:affiliation id="F552E1D395D51C92A7D6656E1B4BC177">Dry Forest Conservation Fund, Sector Santa Rosa, Área de Conservación Guanacaste, phillipsrodriguez @ gmail. com urn: lsid: zoobank. org: author: E 798045 C-A 872 - 4 F 6 A- 82 F 4 - 5 AC 78 A 04 DDA 3</mods:affiliation>
</mods:name>
<mods:name id="D4F03C2E769B78A02E0935DBED7C9AF3" type="personal">
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<mods:namePart id="DA0D913C9ABE6C0B824BA32DF5B3B7E8">Brown, John W.</mods:namePart>
<mods:affiliation id="D80E57FDF600C7FD324D0A6ED0D1E02D">National Museum of Natural History, Smithsonian Institution Washington, DC, 20013 - 7012, USA tortricidae. jwb @ gmail. com urn: lsid: zoobank. org: author: 3 C 52 FC 4 E-E 988 - 4 AD 6 - B 0 D 1 - 9 A 5 CA 74 CB 24 C</mods:affiliation>
</mods:name>
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<mods:namePart id="E832D1BB69FA347111B5EE95FF5D96DD">Hallwachs, Winnie</mods:namePart>
<mods:affiliation id="1781CC52E8B745C9A1D5C9A8E54B502F">Department of Biology, University of Pennsylvania Philadelphia, PA 19104, USA</mods:affiliation>
</mods:name>
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<mods:namePart id="5F2E8A7773B161DF9391782C7CF3F203">Janzen, Daniel H.</mods:namePart>
<mods:affiliation id="B496AD41C9D5ED6F83ABAE4FF92FDA90">Department of Biology, University of Pennsylvania Philadelphia, PA 19104, USA</mods:affiliation>
</mods:name>
<mods:typeOfResource id="A78C2074C15F599008C5474AEEBCC078">text</mods:typeOfResource>
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<mods:title id="EF0D8FA83F2641507EB5908849D02973">Insecta Mundi</mods:title>
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<mods:part id="200625AEA6CC045DD8C2710DEDE397BB">
<mods:date id="048385F2E75612E14A8B27280BD1DD0F">2021</mods:date>
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<mods:number id="8444DDA723AA415CCDDE70C2F7210128">2021-05-28</mods:number>
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<mods:number id="61182129591A8CADC2B6C5CAC32C529C">2021</mods:number>
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<mods:number id="730736301B3BD96BC390594107AD9CCD">868</mods:number>
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<mods:start id="0CBD7311872ED0DBACFCDE926B504F19">1</mods:start>
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<mods:identifier id="F4AC241C8971625DFA1A7F0EFCA81392" type="DOI">10.5281/zenodo.5042023</mods:identifier>
<mods:identifier id="07D2700FD2019E21D285ECDE6DE3C1EB" type="ISSN">1942-1354</mods:identifier>
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<treatment id="03EB577BFFADF076FF67FD54FEF6FC04" LSID="urn:lsid:plazi:treatment:03EB577BFFADF076FF67FD54FEF6FC04" httpUri="http://treatment.plazi.org/id/03EB577BFFADF076FF67FD54FEF6FC04" lastPageId="28" lastPageNumber="27" pageId="27" pageNumber="26">
<subSubSection id="C358B5E6FFADF071FF67FD54FC5FFCF8" box="[192,958,765,794]" pageId="27" pageNumber="26" type="nomenclature">
<paragraph id="8BFDE66DFFADF071FF67FD54FC5FFCF8" blockId="27.[192,958,765,828]" box="[192,958,765,794]" pageId="27" pageNumber="26">
<heading id="D0B55101FFADF071FF67FD54FC5FFCF8" box="[192,958,765,794]" fontSize="12" level="2" pageId="27" pageNumber="26" reason="2">
<emphasis id="B9363A7FFFADF071FF67FD54FC5FFCF8" bold="true" box="[192,958,765,794]" pageId="27" pageNumber="26">
<taxonomicName id="4C429DEEFFADF071FF67FD54FCFDFCF8" authority="Phillips and Brown" authorityName="Phillips and Brown" authorityYear="2021" box="[192,796,765,794]" class="Insecta" family="Oecophoridae" genus="Chlamydastis" kingdom="Animalia" order="Lepidoptera" pageId="27" pageNumber="26" phylum="Arthropoda" rank="species" species="gladysrojasae" status="sp. nov.">
<emphasis id="B9363A7FFFADF071FF67FD54FDFFFCF8" bold="true" box="[192,542,765,794]" italics="true" pageId="27" pageNumber="26">Chlamydastis gladysrojasae</emphasis>
Phillips and Brown
</taxonomicName>
,
<taxonomicNameLabel id="A2058704FFADF071FC8EFD57FC5FFCF8" box="[809,958,765,794]" pageId="27" pageNumber="26" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C358B5E6FFADF071FF67FC8BFE71FCDE" box="[192,400,802,828]" pageId="27" pageNumber="26" type="description">
<paragraph id="8BFDE66DFFADF071FF67FC8BFE71FCDE" blockId="27.[192,958,765,828]" box="[192,400,802,828]" pageId="27" pageNumber="26">
<figureCitation id="1379FAE8FFADF071FF67FC8BFED7FCDE" box="[192,310,802,828]" captionStart="Figures 1622" captionStartId="22.[192,279,1765,1791]" captionTargetBox="[208,1438,271,1674]" captionTargetId="figure-14@22.[192,1441,248,1694]" captionTargetPageId="22" captionText="Figures 1622. Adults of Chlamydastis from Costa Rica Mendoron and Phytoptera Species Groups. 16) C. carlosviquezi, holotype, male, 09-SRNP-107366. 17) C. christerhanssoni, holotype, male, INBIOCRI000844209. 18) C. phytoptera, male, 12-SRNP-105632. 19) C. christhompsoni, male, holotype, 09-SRNP-20126. 20) C. paulhansoni, male, holotype, INB0004269422.21) C. elenaulateae, female, paratype, 03-SRNP-10412.22) C. gladysrojasae, female, paratype, 06-SRNP-109148." figureDoi="http://doi.org/10.5281/zenodo.5042029" httpUri="https://zenodo.org/record/5042029/files/figure.png" pageId="27" pageNumber="26">Figures 22</figureCitation>
,
<figureCitation id="1379FAE8FFADF071FEE7FC8BFEBCFCDE" box="[320,349,802,828]" captionStart="Figures 6569" captionStartId="51.[192,279,1799,1825]" captionTargetBox="[253,1376,265,1740]" captionTargetId="figure-69@51.[192,1440,248,1755]" captionTargetPageId="51" captionText="Figures 6569. Male genitalia (phallus detached) of Chlamydastis from Costa Rica Phytoptera Species Group. 65) C. phytoptera, 10-SRNP-101654 (65a = last abdominal segment). 66) C. christhompsoni, paratype, INBI- OCRI000543326 (66a = last abdominal segment). 67) C. paulhansoni, paratype, INBIOCRI0004269421. 68) C. elenaulateae, paratype, 10-SRNP-1458. 69) C. gladysrojasae, paratype, 09-SRNP-101325." figureDoi="http://doi.org/10.5281/zenodo.5042051" httpUri="https://zenodo.org/record/5042051/files/figure.png" pageId="27" pageNumber="26">69</figureCitation>
,
<figureCitation id="1379FAE8FFADF071FECFFC8BFE71FCDE" box="[360,400,802,828]" captionStart="Figures 103110" captionStartId="58.[192,279,1799,1825]" captionTargetBox="[297,1376,232,1751]" captionTargetId="figure-70@58.[237,1394,208,1769]" captionTargetPageId="58" captionText="Figures 103110. Female genitalia of Chlamydastis. 103) C. phytoptera, INBIOCRI000650708. 104) C. christhompsoni, paratype, INBIOCRI002553979. 105) C. elenaulateae, paratype, 03-SRNP-10412. 106) C. gladysrojasae, paratype, 08-SRNP-102184. 107) C. powelli, paratype, 06-SRNP-3185. 108) C. isidrochaconi, paratype, INB00151483. 109) C. jimlewisi, paratype, INB0073276. 110) C. jimmilleri, paratype, INB003326762." figureDoi="http://doi.org/10.5281/zenodo.5042071" httpUri="https://zenodo.org/record/5042071/files/figure.png" pageId="27" pageNumber="26">106</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C358B5E6FFADF071FF67FCE5FDBEFC50" pageId="27" pageNumber="26" type="materials_examined">
<paragraph id="8BFDE66DFFADF071FF67FCE5FB67FC6A" blockId="27.[192,1441,844,905]" pageId="27" pageNumber="26">
<materialsCitation id="3B2AEC30FFADF071FF67FCE5FB67FC6A" accessionNumber="HQ556226" collectingDate="2009-03-13" collectionCode="USNM" collectorName="F. Quesada &amp; S. Rios &amp; GenBank" country="Costa Rica" county="Male" elevation="491" location="Manta Hugo" municipality="Sector Rincon Rain Forest" pageId="27" pageNumber="26" specimenCount="1" stateProvince="Guanacaste" typeStatus="holotype">
<emphasis id="B9363A7FFFADF071FF67FCE5FED5FC84" bold="true" box="[192,308,844,870]" pageId="27" pageNumber="26">
<typeStatus id="54F958CFFFADF071FF67FCE5FECEFC84" box="[192,303,844,870]" pageId="27" pageNumber="26" type="holotype">Holotype</typeStatus>
.
</emphasis>
<collectingCounty id="629C9EE1FFADF071FE9BFCE5FE97FC84" box="[316,374,844,870]" pageId="27" pageNumber="26">Male</collectingCounty>
,
<collectingCountry id="F355A6FDFFADF071FE25FCE5FE1EFC84" box="[386,511,844,870]" name="Costa Rica" pageId="27" pageNumber="26">Costa Rica</collectingCountry>
,
<collectingRegion id="4986288FFFADF071FDADFCE5FD6CFC84" box="[522,653,844,870]" country="Costa Rica" name="Guanacaste" pageId="27" pageNumber="26">Guanacaste</collectingRegion>
, ACG,
<collectingMunicipality id="6B997C17FFADF071FD46FCE5FBECFC84" box="[737,1037,844,870]" pageId="27" pageNumber="26">Sector Rincón Rain Forest</collectingMunicipality>
,
<location id="8E9DB0B6FFADF071FBBEFCE5FB4BFC84" LSID="urn:lsid:plazi:treatment:03EB577BFFADF076FF67FD54FEF6FC04:8E9DB0B6FFADF071FBBEFCE5FB4BFC84" box="[1049,1194,844,870]" country="Costa Rica" county="Male" municipality="Sector Rincon Rain Forest" name="Manta Hugo" pageId="27" pageNumber="26" stateProvince="Guanacaste">Manta Hugo</location>
,
<quantity id="4CBA4B88FFADF071FB11FCE5FB1EFC84" box="[1206,1279,844,870]" metricMagnitude="2" metricUnit="m" metricValue="4.91" pageId="27" pageNumber="26" unit="m" value="491.0">
<elevation id="006F015EFFADF071FB11FCE5FB1EFC84" box="[1206,1279,844,870]" metricMagnitude="2" metricUnit="m" metricValue="4.91" pageId="27" pageNumber="26" unit="m" value="491.0">491 m</elevation>
</quantity>
,
<date id="FFFCC0ADFFADF071FAACFCE5FA60FC84" box="[1291,1409,844,870]" pageId="27" pageNumber="26" value="2009-03-13">
<collectingDate id="EFB83945FFADF071FAACFCE5FA60FC84" box="[1291,1409,844,870]" pageId="27" pageNumber="26" value="2009-03-13">13.iii.2009</collectingDate>
</date>
,
<collectorName id="26B783BBFFADF071FA29FCE5FEC5FC6A" pageId="27" pageNumber="26">F. Quesada</collectorName>
and
<collectorName id="26B783BBFFADF071FEFDFCC7FE49FC6A" box="[346,424,878,904]" pageId="27" pageNumber="26">S. Rios</collectorName>
, 10-SRNP-106674,
<collectorName id="26B783BBFFADF071FD27FCC7FD0BFC6A" box="[640,746,878,904]" pageId="27" pageNumber="26">GenBank</collectorName>
accession code HQ556226 (
<collectionCode id="ED537EA8FFADF071FB83FCC7FB9BFC6A" box="[1060,1146,878,904]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="27" pageNumber="26" type="Museum">USNM</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="8BFDE66DFFADF071FF67FC31FDBEFC50" blockId="27.[192,607,920,947]" box="[192,607,920,947]" pageId="27" pageNumber="26">
<emphasis id="B9363A7FFFADF071FF67FC31FED3FC50" bold="true" box="[192,306,920,946]" pageId="27" pageNumber="26">
<typeStatus id="54F958CFFFADF071FF67FC31FED3FC50" box="[192,306,920,946]" pageId="27" pageNumber="26" type="paratype">Paratypes</typeStatus>
</emphasis>
(
<specimenCount id="9D442DE4FFADF071FEE6FC31FE84FC50" box="[321,357,920,946]" pageId="27" pageNumber="26" type="male">2♂</specimenCount>
,
<specimenCount id="9D442DE4FFADF071FED5FC31FE75FC50" box="[370,404,920,946]" pageId="27" pageNumber="26" type="female">3♀</specimenCount>
). See Appendix 1.
</paragraph>
</subSubSection>
<subSubSection id="C358B5E6FFADF071FF67FC6BFA72FBC3" pageId="27" pageNumber="26" type="diagnosis">
<paragraph id="8BFDE66DFFADF071FF67FC6BFA72FBC3" blockId="27.[192,1442,962,1057]" pageId="27" pageNumber="26">
<emphasis id="B9363A7FFFADF071FF67FC6BFEDDFC3E" bold="true" box="[192,316,962,988]" pageId="27" pageNumber="26">Diagnosis.</emphasis>
<taxonomicName id="4C429DEEFFADF071FEE4FC6BFD8EFC3E" authorityName="Phillips and Brown" authorityYear="2021" box="[323,623,962,988]" class="Insecta" family="Oecophoridae" genus="Chlamydastis" kingdom="Animalia" order="Lepidoptera" pageId="27" pageNumber="26" phylum="Arthropoda" rank="species" species="gladysrojasae">
<emphasis id="B9363A7FFFADF071FEE4FC6BFD8EFC3E" box="[323,623,962,988]" italics="true" pageId="27" pageNumber="26">Chlamydastis gladysrojasae</emphasis>
</taxonomicName>
is most similar to
<taxonomicName id="4C429DEEFFADF071FCE3FC6AFC0BFC3E" authorityName="Phillips and Brown" authorityYear="2021" box="[836,1002,962,989]" class="Insecta" family="Oecophoridae" genus="Chlamydastis" kingdom="Animalia" order="Lepidoptera" pageId="27" pageNumber="26" phylum="Arthropoda" rank="species" species="paulhansoni">
<emphasis id="B9363A7FFFADF071FCE3FC6AFC0BFC3E" box="[836,1002,962,989]" italics="true" pageId="27" pageNumber="26">C. paulhansoni</emphasis>
</taxonomicName>
and
<taxonomicName id="4C429DEEFFADF071FB84FC6AFB2BFC3E" authorityName="Phillips and Brown" authorityYear="2021" box="[1059,1226,962,989]" class="Insecta" family="Oecophoridae" genus="Chlamydastis" kingdom="Animalia" order="Lepidoptera" pageId="27" pageNumber="26" phylum="Arthropoda" rank="species" species="elenaulateae">
<emphasis id="B9363A7FFFADF071FB84FC6AFB2BFC3E" box="[1059,1226,962,989]" italics="true" pageId="27" pageNumber="26">C. elenaulateae</emphasis>
</taxonomicName>
, from which it can be distinguished by its shorter uncus with a rounded apex, its shorter gnathos, and its more rounded sacculus. Superficially, the vertex of the head and the adjacent collar of scales are much darker brown in
<taxonomicName id="4C429DEEFFADF071FB7AFBAEFA6CFBC3" authorityName="Phillips and Brown" authorityYear="2021" box="[1245,1421,1031,1057]" class="Insecta" family="Oecophoridae" genus="Chlamydastis" kingdom="Animalia" order="Lepidoptera" pageId="27" pageNumber="26" phylum="Arthropoda" rank="species" species="gladysrojasae">
<emphasis id="B9363A7FFFADF071FB7AFBAEFA6CFBC3" box="[1245,1421,1031,1057]" italics="true" pageId="27" pageNumber="26">C. gladysrojasae</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C358B5E6FFADF071FF67FB98FB6AF9B6" pageId="27" pageNumber="26" type="description">
<paragraph id="8BFDE66DFFADF071FF67FB98FCB8FABE" blockId="27.[191,1442,1073,1544]" pageId="27" pageNumber="26">
<emphasis id="B9363A7FFFADF071FF67FB98FEB3FBA9" bold="true" box="[192,338,1073,1099]" pageId="27" pageNumber="26">Description.</emphasis>
MALE (
<figureCitation id="1379FAE8FFADF071FE10FB98FDEBFBA9" box="[439,522,1073,1099]" captionStart="Figures 1622" captionStartId="22.[192,279,1765,1791]" captionTargetBox="[208,1438,271,1674]" captionTargetId="figure-14@22.[192,1441,248,1694]" captionTargetPageId="22" captionText="Figures 1622. Adults of Chlamydastis from Costa Rica Mendoron and Phytoptera Species Groups. 16) C. carlosviquezi, holotype, male, 09-SRNP-107366. 17) C. christerhanssoni, holotype, male, INBIOCRI000844209. 18) C. phytoptera, male, 12-SRNP-105632. 19) C. christhompsoni, male, holotype, 09-SRNP-20126. 20) C. paulhansoni, male, holotype, INB0004269422.21) C. elenaulateae, female, paratype, 03-SRNP-10412.22) C. gladysrojasae, female, paratype, 06-SRNP-109148." figureDoi="http://doi.org/10.5281/zenodo.5042029" httpUri="https://zenodo.org/record/5042029/files/figure.png" pageId="27" pageNumber="26">Fig. 22</figureCitation>
).
<emphasis id="B9363A7FFFADF071FDB8FB98FDBBFBA9" box="[543,602,1073,1099]" italics="true" pageId="27" pageNumber="26">Head</emphasis>
. Frons whitish, vertex blackish, collar blackish; labial palpus with all segments light brown, second segment with a lighter ring at anterior end; antenna with sensory setae ca. 1.5 times width of flagellomere.
<emphasis id="B9363A7FFFADF071FE10FBDCFDE4FB6D" box="[439,517,1141,1167]" italics="true" pageId="27" pageNumber="26">Thorax</emphasis>
. Base of tegula and anterior margin of prothorax brown, mesothorax and metathorax gray with faint pale purplish tint. FW length 9.511.0 mm; FW brownish with faint olive green overscaling and small green irregular blotches; costal 0.5 darker with faint black lines along veins; lower 0.5 paler, without lines along veins.
<emphasis id="B9363A7FFFADF071FE2AFB75FE14FB14" box="[397,501,1244,1270]" italics="true" pageId="27" pageNumber="26">Abdomen</emphasis>
. Dorsum grayish, venter darker with paired whitish spots. Genitalia (
<figureCitation id="1379FAE8FFADF071FAAAFB75FABFFB15" box="[1293,1374,1244,1271]" captionStart="Figures 6569" captionStartId="51.[192,279,1799,1825]" captionTargetBox="[253,1376,265,1740]" captionTargetId="figure-69@51.[192,1440,248,1755]" captionTargetPageId="51" captionText="Figures 6569. Male genitalia (phallus detached) of Chlamydastis from Costa Rica Phytoptera Species Group. 65) C. phytoptera, 10-SRNP-101654 (65a = last abdominal segment). 66) C. christhompsoni, paratype, INBI- OCRI000543326 (66a = last abdominal segment). 67) C. paulhansoni, paratype, INBIOCRI0004269421. 68) C. elenaulateae, paratype, 10-SRNP-1458. 69) C. gladysrojasae, paratype, 09-SRNP-101325." figureDoi="http://doi.org/10.5281/zenodo.5042051" httpUri="https://zenodo.org/record/5042051/files/figure.png" pageId="27" pageNumber="26">Fig. 69</figureCitation>
) with uncus slender, ca. 0.5 length of valva, with rounded apex; gnathos short; valva subrectangular with rounded apex, strongly veined, with dense cluster of specialized setae near mid-costa; sacculus a short, free, rounded lobe; phallus curved throughout, with round-tipped sclerite in vesica.
</paragraph>
<paragraph id="8BFDE66DFFADF071FEA7FACCFE05F9E5" blockId="27.[191,1442,1073,1544]" pageId="27" pageNumber="26">
FEMALE.
<emphasis id="B9363A7FFFADF071FED1FACDFE50FA9C" box="[374,433,1380,1406]" italics="true" pageId="27" pageNumber="26">Head</emphasis>
and
<emphasis id="B9363A7FFFADF071FE43FACDFDD3FA9C" box="[484,562,1380,1406]" italics="true" pageId="27" pageNumber="26">Thorax</emphasis>
. Essentially as described for male, except sensory setae of antenna short, sparse; FW length
<quantity id="4CBA4B88FFADF071FEE5FA2FFE09FA43" box="[322,488,1414,1441]" metricMagnitude="-2" metricUnit="m" metricValue="1.3199999999999998" metricValueMax="1.35" metricValueMin="1.29" pageId="27" pageNumber="26" unit="mm" value="13.2" valueMax="13.5" valueMin="12.9">12.913.5 mm</quantity>
.
<emphasis id="B9363A7FFFADF071FE51FA2EFD81FA43" box="[502,608,1415,1441]" italics="true" pageId="27" pageNumber="26">Abdomen</emphasis>
. Genitalia (
<figureCitation id="1379FAE8FFADF071FD40FA2EFCA4FA40" box="[743,837,1415,1442]" captionStart="Figures 103110" captionStartId="58.[192,279,1799,1825]" captionTargetBox="[297,1376,232,1751]" captionTargetId="figure-70@58.[237,1394,208,1769]" captionTargetPageId="58" captionText="Figures 103110. Female genitalia of Chlamydastis. 103) C. phytoptera, INBIOCRI000650708. 104) C. christhompsoni, paratype, INBIOCRI002553979. 105) C. elenaulateae, paratype, 03-SRNP-10412. 106) C. gladysrojasae, paratype, 08-SRNP-102184. 107) C. powelli, paratype, 06-SRNP-3185. 108) C. isidrochaconi, paratype, INB00151483. 109) C. jimlewisi, paratype, INB0073276. 110) C. jimmilleri, paratype, INB003326762." figureDoi="http://doi.org/10.5281/zenodo.5042071" httpUri="https://zenodo.org/record/5042071/files/figure.png" pageId="27" pageNumber="26">Fig. 106</figureCitation>
) with papillae anales slightly diverging and rounded posteriorly, each papilla with a distinct broad line of sclerotization; ductus bursae slender, weakly broadened anteriorly to ill-defined junction with corpus bursae; corpus bursae pear-shaped, signum subrectangular, gradually narrowed at each end.
</paragraph>
<paragraph id="8BFDE66DFFADF071FF67F9BEFB6AF9B6" blockId="27.[192,1440,1559,1620]" pageId="27" pageNumber="26">
<emphasis id="B9363A7FFFADF071FF67F9BEFE92F9D0" bold="true" box="[192,371,1559,1586]" pageId="27" pageNumber="26">DNA barcodes.</emphasis>
The 13 barcodes of
<taxonomicName id="4C429DEEFFADF071FDF2F9B1FCE7F9D3" authorityName="Phillips and Brown" authorityYear="2021" box="[597,774,1559,1586]" class="Insecta" family="Oecophoridae" genus="Chlamydastis" kingdom="Animalia" order="Lepidoptera" pageId="27" pageNumber="26" phylum="Arthropoda" rank="species" species="gladysrojasae">
<emphasis id="B9363A7FFFADF071FDF2F9B1FCE7F9D3" box="[597,774,1559,1586]" italics="true" pageId="27" pageNumber="26">C. gladysrojasae</emphasis>
</taxonomicName>
form a BIN (BOLD:AAD8085) with an average distance of 0.32% among barcodes, and a distance of 7.22% to its nearest neighbor,
<taxonomicName id="4C429DEEFFADF071FC79F993FB64F9B6" authorityName="Phillips and Brown" authorityYear="2021" box="[990,1157,1594,1620]" class="Insecta" family="Oecophoridae" genus="Chlamydastis" kingdom="Animalia" order="Lepidoptera" pageId="27" pageNumber="26" phylum="Arthropoda" rank="species" species="elenaulateae">
<emphasis id="B9363A7FFFADF071FC79F993FB64F9B6" box="[990,1157,1594,1620]" italics="true" pageId="27" pageNumber="26">C. elenaulateae</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C358B5E6FFADF071FF67F9CAFB79F99C" box="[192,1176,1635,1662]" pageId="27" pageNumber="26" type="distribution">
<paragraph id="8BFDE66DFFADF071FF67F9CAFB79F99C" blockId="27.[192,1176,1635,1662]" box="[192,1176,1635,1662]" pageId="27" pageNumber="26">
<emphasis id="B9363A7FFFADF071FF67F9CAFEBBF99F" bold="true" box="[192,346,1635,1661]" pageId="27" pageNumber="26">Distribution.</emphasis>
<taxonomicName id="4C429DEEFFADF071FEC7F9CDFD6DF99C" authorityName="Phillips and Brown" authorityYear="2021" box="[352,652,1636,1662]" class="Insecta" family="Oecophoridae" genus="Chlamydastis" kingdom="Animalia" order="Lepidoptera" pageId="27" pageNumber="26" phylum="Arthropoda" rank="species" species="gladysrojasae">
<emphasis id="B9363A7FFFADF071FEC7F9CDFD6DF99C" box="[352,652,1636,1662]" italics="true" pageId="27" pageNumber="26">Chlamydastis gladysrojasae</emphasis>
</taxonomicName>
has been collected from
<quantity id="4CBA4B88FFADF071FC05F9CDFBD5F99C" box="[930,1076,1636,1662]" metricMagnitude="2" metricUnit="m" metricValue="5.505" metricValueMax="6.1" metricValueMin="4.91" pageId="27" pageNumber="26" unit="m" value="550.5" valueMax="610.0" valueMin="491.0">491 to 610 m</quantity>
in ACG.
</paragraph>
</subSubSection>
<subSubSection id="C358B5E6FFADF071FF67F927FCC1F94A" box="[192,800,1678,1704]" pageId="27" pageNumber="26" type="biology_ecology">
<paragraph id="8BFDE66DFFADF071FF67F927FCC1F94A" blockId="27.[192,800,1678,1704]" box="[192,800,1678,1704]" pageId="27" pageNumber="26">
<emphasis id="B9363A7FFFADF071FF67F927FEC1F94A" bold="true" box="[192,288,1678,1704]" pageId="27" pageNumber="26">Biology.</emphasis>
The immatures and host plants are unknown.
</paragraph>
</subSubSection>
<subSubSection id="C358B5E6FFADF076FF67F911FEF6FC04" lastPageId="28" lastPageNumber="27" pageId="27" pageNumber="26" type="etymology">
<paragraph id="8BFDE66DFFADF071FF67F911FCE0F916" blockId="27.[192,1440,1720,1780]" pageId="27" pageNumber="26">
<emphasis id="B9363A7FFFADF071FF67F911FEA5F930" bold="true" box="[192,324,1720,1746]" pageId="27" pageNumber="26">Etymology.</emphasis>
<taxonomicName id="4C429DEEFFADF071FEECF911FD97F930" authorityName="Phillips and Brown" authorityYear="2021" box="[331,630,1720,1746]" class="Insecta" family="Oecophoridae" genus="Chlamydastis" kingdom="Animalia" order="Lepidoptera" pageId="27" pageNumber="26" phylum="Arthropoda" rank="species" species="gladysrojasae">
<emphasis id="B9363A7FFFADF071FEECF911FD97F930" box="[331,630,1720,1746]" italics="true" pageId="27" pageNumber="26">Chlamydastis gladysrojasae</emphasis>
</taxonomicName>
is named in honor of Gladys Rojas in recognition of her logistic support of the national biodiversity inventory of
<collectingCountry id="F355A6FDFFADF071FD25F973FD1CF916" box="[642,765,1754,1780]" name="Costa Rica" pageId="27" pageNumber="26">Costa Rica</collectingCountry>
.
</paragraph>
<paragraph id="8BFDE66DFFADF071FD3EF8B7FC26F8D9" blockId="27.[665,967,1822,1851]" box="[665,967,1822,1851]" pageId="27" pageNumber="26">
<emphasis id="B9363A7FFFADF071FD3EF8B7FC26F8D9" bold="true" box="[665,967,1822,1851]" pageId="27" pageNumber="26">Tryphon Species Group</emphasis>
</paragraph>
<paragraph id="8BFDE66DFFADF076FF67F8E3FC5EFE5D" blockId="27.[192,1441,1866,1927]" lastBlockId="28.[191,1442,214,998]" lastPageId="28" lastPageNumber="27" pageId="27" pageNumber="26">
The Tryphon Species Group is composed of six species:
<taxonomicName id="4C429DEEFFADF071FC80F8E3FC73F886" authorityName="Phillips and Brown" authorityYear="2021" box="[807,914,1866,1892]" class="Insecta" family="Oecophoridae" genus="Chlamydastis" kingdom="Animalia" order="Lepidoptera" pageId="27" pageNumber="26" phylum="Arthropoda" rank="species" species="powelli">
<emphasis id="B9363A7FFFADF071FC80F8E3FC73F886" box="[807,914,1866,1892]" italics="true" pageId="27" pageNumber="26">C. powelli</emphasis>
</taxonomicName>
,
<taxonomicName id="4C429DEEFFADF071FC38F8E3FBABF886" authorityName="Phillips and Brown" authorityYear="2021" box="[927,1098,1866,1892]" class="Insecta" family="Oecophoridae" genus="Chlamydastis" kingdom="Animalia" order="Lepidoptera" pageId="27" pageNumber="26" phylum="Arthropoda" rank="species" species="gracewoodae">
<emphasis id="B9363A7FFFADF071FC38F8E3FBABF886" box="[927,1098,1866,1892]" italics="true" pageId="27" pageNumber="26">C. gracewoodae</emphasis>
</taxonomicName>
,
<taxonomicName id="4C429DEEFFADF071FBF2F8E3FB07F886" authorityName="Phillips and Brown" authorityYear="2021" box="[1109,1254,1866,1892]" class="Insecta" family="Oecophoridae" genus="Chlamydastis" kingdom="Animalia" order="Lepidoptera" pageId="27" pageNumber="26" phylum="Arthropoda" rank="species" species="juanmatai">
<emphasis id="B9363A7FFFADF071FBF2F8E3FB07F886" box="[1109,1254,1866,1892]" italics="true" pageId="27" pageNumber="26">C. juanmatai</emphasis>
</taxonomicName>
,
<taxonomicName id="4C429DEEFFADF071FB56F8E3FA40F886" authorityName="Phillips and Brown" authorityYear="2021" box="[1265,1441,1866,1892]" class="Insecta" family="Oecophoridae" genus="Chlamydastis" kingdom="Animalia" order="Lepidoptera" pageId="27" pageNumber="26" phylum="Arthropoda" rank="species" species="isidrochaconi">
<emphasis id="B9363A7FFFADF071FB56F8E3FA40F886" box="[1265,1441,1866,1892]" italics="true" pageId="27" pageNumber="26">C. isidrochaconi</emphasis>
</taxonomicName>
(all from
<collectingCountry id="F355A6FDFFADF071FE80F8C5FE42F865" box="[295,419,1900,1927]" name="Costa Rica" pageId="27" pageNumber="26">Costa Rica</collectingCountry>
),
<taxonomicName id="4C429DEEFFADF071FE14F8C5FD26F864" authority="(Busck, 1920)" baseAuthorityName="Busck" baseAuthorityYear="1920" box="[435,711,1900,1926]" class="Insecta" family="Oecophoridae" genus="Chlamydastis" kingdom="Animalia" order="Lepidoptera" pageId="27" pageNumber="26" phylum="Arthropoda" rank="species" species="tryphon">
<emphasis id="B9363A7FFFADF071FE14F8C5FDC9F864" box="[435,552,1900,1926]" italics="true" pageId="27" pageNumber="26">C. tryphon</emphasis>
(
<bibRefCitation id="EFD39B9CFFADF071FD91F8C5FD5EF864" author="Busck A." box="[566,703,1900,1926]" pageId="27" pageNumber="26" pagination="83 - 95" refId="ref37386" refString="Busck A. 1920. Descriptions of new Central American microlepidoptera. Insecutor Inscitiae Menstruus 8: 83 - 95." type="journal article" year="1920">Busck, 1920</bibRefCitation>
)
</taxonomicName>
from
<collectingCountry id="F355A6FDFFADF071FCAEF8C5FC64F864" box="[777,901,1900,1926]" name="Guatemala" pageId="27" pageNumber="26">Guatemala</collectingCountry>
, and
<taxonomicName id="4C429DEEFFADF071FC1AF8C5FB53F864" authority="(Busck, 1911)" baseAuthorityName="Busck" baseAuthorityYear="1911" box="[957,1202,1900,1926]" class="Insecta" family="Oecophoridae" genus="Stenoma" kingdom="Animalia" order="Lepidoptera" pageId="27" pageNumber="26" phylum="Arthropoda" rank="species" species="lactis">
<emphasis id="B9363A7FFFADF071FC1AF8C5FBF5F864" box="[957,1044,1900,1926]" italics="true" pageId="27" pageNumber="26">C. lactis</emphasis>
(Busck, 1911)
</taxonomicName>
from
<collectingCountry id="F355A6FDFFADF071FB53F8C5FA7DF864" box="[1268,1436,1900,1926]" name="French Guiana" pageId="27" pageNumber="26">French Guiana</collectingCountry>
, the last illustrated by
<bibRefCitation id="EFD39B9CFFAAF076FE0EFF7FFD99FF13" author="Clarke JFG" box="[425,632,214,241]" pageId="28" pageNumber="27" refId="ref37407" refString="Clarke JFG. 1955. Catalogue of the type specimens of microlepidoptera in the British Museum (Natural History) described by Edward Meyrick, volume 2. Jarrold and Sons Ltd.; Norwich. 531 p." type="book" year="1955">Clarke (1955: 175)</bibRefCitation>
. The forewing pattern is sexually dimorphic: males typically have a broad, dark media fascia originating near the middle of the costa and extending to the hind margin, leaving a uniform pale area in the basal half of the forewing (the distal portion beyond the fascia may be pale or concolorous with the fascia), and the costa has an extremely shallow, broad concavity near the middle; females typically have a more uniform ground color (light or dark) with a dark semicircular blotch near the middle of the hind margin. Males of all species have a patch of long, fine, pale scales along the surface of the antenna opposite of the sensory setae, extending from the pedicel to ca. 0.5 the length of the antenna.
</paragraph>
<paragraph id="8BFDE66DFFAAF076FEA7FE6EFB13FD16" blockId="28.[191,1442,214,998]" pageId="28" pageNumber="27">
The six species share very similar male genitalia with a long, slender, slightly curved uncus; a well-developed gnathos with a flattened distal process at the junction of the lateral arms; a valva with a large, distally attenuate sacculus that projects outward from the vinculum and bears a dense patch of scales (hairpencil); and the dorsoposterior portion of the valva with a broad rounded-triangular lobe near the middle of the outer margin bearing a second dense patch of scales (hairpencil), and its upper, apical portion with a rounded or pointed apex. The specialized arrowhead-tipped setae of the male originate as a dense bundle slightly basal of the middle of the costa of the valva.
<taxonomicName id="4C429DEEFFAAF076FEF5FD3CFDDAFD4D" authorityName="Phillips and Brown" authorityYear="2021" box="[338,571,661,687]" class="Insecta" family="Oecophoridae" genus="Chlamydastis" kingdom="Animalia" order="Lepidoptera" pageId="28" pageNumber="27" phylum="Arthropoda" rank="species" species="powelli">
<emphasis id="B9363A7FFFAAF076FEF5FD3CFDDAFD4D" box="[338,571,661,687]" italics="true" pageId="28" pageNumber="27">Chlamydastis powelli</emphasis>
</taxonomicName>
,
<taxonomicName id="4C429DEEFFAAF076FDEDFD3FFD16FD4D" authorityName="Phillips and Brown" authorityYear="2021" box="[586,759,661,688]" class="Insecta" family="Oecophoridae" genus="Chlamydastis" kingdom="Animalia" order="Lepidoptera" pageId="28" pageNumber="27" phylum="Arthropoda" rank="species" species="gracewoodae">
<emphasis id="B9363A7FFFAAF076FDEDFD3FFD16FD4D" box="[586,759,661,688]" italics="true" pageId="28" pageNumber="27">C. gracewoodae</emphasis>
</taxonomicName>
, and
<taxonomicName id="4C429DEEFFAAF076FC9EFD3FFC2DFD4D" authorityName="Phillips and Brown" authorityYear="2021" box="[825,972,662,688]" class="Insecta" family="Oecophoridae" genus="Chlamydastis" kingdom="Animalia" order="Lepidoptera" pageId="28" pageNumber="27" phylum="Arthropoda" rank="species" species="juanmatai">
<emphasis id="B9363A7FFFAAF076FC9EFD3FFC2DFD4D" box="[825,972,662,688]" italics="true" pageId="28" pageNumber="27">C. juanmatai</emphasis>
</taxonomicName>
share a dorsal, semi-membranous, hump near the middle of the phallus.
<taxonomicName id="4C429DEEFFAAF076FDB9FD11FCF1FD30" baseAuthorityName="Busck" baseAuthorityYear="1920" box="[542,784,696,722]" class="Insecta" family="Oecophoridae" genus="Chlamydastis" kingdom="Animalia" order="Lepidoptera" pageId="28" pageNumber="27" phylum="Arthropoda" rank="species" species="tryphon">
<emphasis id="B9363A7FFFAAF076FDB9FD11FCF1FD30" box="[542,784,696,722]" italics="true" pageId="28" pageNumber="27">Chlamydastis tryphon</emphasis>
</taxonomicName>
can be distinguished from the four Costa Rican species by its more pointed valva and its short, stout phallus with a small, ventrally directed, sclerotized tip.
</paragraph>
<paragraph id="8BFDE66DFFAAF076FEA7FD54FB17FCBE" blockId="28.[191,1442,214,998]" pageId="28" pageNumber="27">
The male genitalia of
<taxonomicName id="4C429DEEFFAAF076FE5DFD54FD87FCF5" authorityName="Phillips and Brown" authorityYear="2021" box="[506,614,765,791]" class="Insecta" family="Oecophoridae" genus="Chlamydastis" kingdom="Animalia" order="Lepidoptera" pageId="28" pageNumber="27" phylum="Arthropoda" rank="species" species="powelli">
<emphasis id="B9363A7FFFAAF076FE5DFD54FD87FCF5" box="[506,614,765,791]" italics="true" pageId="28" pageNumber="27">C. powelli</emphasis>
</taxonomicName>
,
<taxonomicName id="4C429DEEFFAAF076FDD2FD54FCC2FCF5" authorityName="Phillips and Brown" authorityYear="2021" box="[629,803,765,791]" class="Insecta" family="Oecophoridae" genus="Chlamydastis" kingdom="Animalia" order="Lepidoptera" pageId="28" pageNumber="27" phylum="Arthropoda" rank="species" species="gracewoodae">
<emphasis id="B9363A7FFFAAF076FDD2FD54FCC2FCF5" box="[629,803,765,791]" italics="true" pageId="28" pageNumber="27">C. gracewoodae</emphasis>
</taxonomicName>
, and
<taxonomicName id="4C429DEEFFAAF076FCC1FD54FC18FCF5" authorityName="Phillips and Brown" authorityYear="2021" box="[870,1017,765,791]" class="Insecta" family="Oecophoridae" genus="Chlamydastis" kingdom="Animalia" order="Lepidoptera" pageId="28" pageNumber="27" phylum="Arthropoda" rank="species" species="juanmatai">
<emphasis id="B9363A7FFFAAF076FCC1FD54FC18FCF5" box="[870,1017,765,791]" italics="true" pageId="28" pageNumber="27">C. juanmatai</emphasis>
</taxonomicName>
are nearly identical, differing only in subtle and/or qualitative features. Of the three, females are known for
<taxonomicName id="4C429DEEFFAAF076FC7AFC89FBAAFCDB" authorityName="Phillips and Brown" authorityYear="2021" box="[989,1099,799,826]" class="Insecta" family="Oecophoridae" genus="Chlamydastis" kingdom="Animalia" order="Lepidoptera" pageId="28" pageNumber="27" phylum="Arthropoda" rank="species" species="powelli">
<emphasis id="B9363A7FFFAAF076FC7AFC89FBAAFCDB" box="[989,1099,799,826]" italics="true" pageId="28" pageNumber="27">C. powelli</emphasis>
</taxonomicName>
and
<taxonomicName id="4C429DEEFFAAF076FB23FC89FACEFCDB" authorityName="Phillips and Brown" authorityYear="2021" box="[1156,1327,799,826]" class="Insecta" family="Oecophoridae" genus="Chlamydastis" kingdom="Animalia" order="Lepidoptera" pageId="28" pageNumber="27" phylum="Arthropoda" rank="species" species="gracewoodae">
<emphasis id="B9363A7FFFAAF076FB23FC89FACEFCDB" box="[1156,1327,799,826]" italics="true" pageId="28" pageNumber="27">C. gracewoodae</emphasis>
</taxonomicName>
, and they likewise are nearly identical in appearance. However, barcodes convincingly separate the species.
</paragraph>
<paragraph id="8BFDE66DFFAAF076FEA7FCCCFEF6FC04" blockId="28.[191,1442,214,998]" pageId="28" pageNumber="27">
In the female genitalia the length of the anterior apophysis may be useful for separating
<taxonomicName id="4C429DEEFFAAF076FB44FCCCFAB1FC9C" authorityName="Phillips and Brown" authorityYear="2021" box="[1251,1360,868,895]" class="Insecta" family="Oecophoridae" genus="Chlamydastis" kingdom="Animalia" order="Lepidoptera" pageId="28" pageNumber="27" phylum="Arthropoda" rank="species" species="powelli">
<emphasis id="B9363A7FFFAAF076FB44FCCCFAB1FC9C" box="[1251,1360,868,895]" italics="true" pageId="28" pageNumber="27">C. powelli</emphasis>
</taxonomicName>
and
<taxonomicName id="4C429DEEFFAAF076FA2EFCCCFEACFC43" authorityName="Phillips and Brown" authorityYear="2021" class="Insecta" family="Oecophoridae" genus="Chlamydastis" kingdom="Animalia" order="Lepidoptera" pageId="28" pageNumber="27" phylum="Arthropoda" rank="species" species="gracewoodae">
<emphasis id="B9363A7FFFAAF076FA2EFCCCFEACFC43" italics="true" pageId="28" pageNumber="27">C. gracewoodae</emphasis>
</taxonomicName>
. Females have a long, slender ductus bursae that gradually broadens into the corpus bursae. The signum is a U-shaped sclerite at the anterior end of the corpus bursae, spiny at each distal 0.33, unspined in the middle.
</paragraph>
</subSubSection>
</treatment>
</document>