155 lines
18 KiB
XML
155 lines
18 KiB
XML
<document id="8621E00C03377B69700CE4B0B849757D" ID-DOI="10.26107/RBZ-2020-0019" ID-ISSN="2345-7600" ID-Zenodo-Dep="4577217" ID-ZooBank="650E406A-6668-42DE-B6A6-637B154A10FE" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1614784910665" checkinUser="carolina" docAuthor="Nguyen, Tung T., Nguyen, Nam Q., Lam, Dang H. & Nguyen, Anh D." docDate="2020" docId="03EACC33FFCF05201387B38CB162FEA6" docLanguage="en" docName="RafflesBZool.68.220-236.pdf" docOrigin="Raffles Bulletin of Zoology 68" docStyle="DocumentStyle:AF09A0581917EACD12E7F0991C3E7C26.5:RafflesBZool.2019-.journal_article" docStyleId="AF09A0581917EACD12E7F0991C3E7C26" docStyleName="RafflesBZool.2019-.journal_article" docStyleVersion="5" docTitle="Metaphire Sims & Easton 1972" docType="treatment" docVersion="3" lastPageNumber="221" masterDocId="FFD3B44BFFCE0523104BB60CB065FFF0" masterDocTitle="Six new species of the genus Metaphire Sims & Easton, 1972 (Annelida: Oligochaeta: Megascolecidae) from southeastern Vietnam" masterLastPageNumber="236" masterPageNumber="220" pageNumber="221" updateTime="1698878656804" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="4E405C060AC556EF4EC77654EC3A3DBA">Six new species of the genus Metaphire Sims & Easton, 1972 (Annelida: Oligochaeta: Megascolecidae) from southeastern Vietnam</mods:title>
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<mods:roleTerm id="D10D9B879AEB2756A482A49DE4B6FD02">Author</mods:roleTerm>
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<mods:namePart id="EE33EB8FEB70F757E93A6F77F90CBE80">Nguyen, Tung T.</mods:namePart>
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<mods:namePart id="F0C5E07DD4FE8B1C1054E6DDE0BA71B7">Nguyen, Nam Q.</mods:namePart>
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<mods:namePart id="A4CC61E02DDEFCAE6CB862ECD57004B9">Lam, Dang H.</mods:namePart>
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<mods:name id="69A30006BBB1FB951755E2E5435A44F9" type="personal">
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<mods:namePart id="8047E4DB903A8DF3C25F8A0B1353BBEF">Nguyen, Anh D.</mods:namePart>
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<mods:typeOfResource id="4D3092362D9EA4CB9CA8A109764EFC5B">text</mods:typeOfResource>
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<mods:title id="E069ECE7F2154761673E6502303E8101">Raffles Bulletin of Zoology</mods:title>
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<mods:part id="A373C25B93D4FD0E34B71A339D363BC3">
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<mods:date id="E6C23B4E6D1BDC393E7A69CD31C95D59">2020</mods:date>
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<mods:number id="96A31519E263580956F67A5BD078F20A">2020-04-22</mods:number>
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<mods:detail id="E6EBA8A04AF3C309E6F70C4352791096" type="volume">
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<mods:number id="584903796D4901A94C23FFDD976F5F50">68</mods:number>
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<mods:start id="C2ADE25E5B22E6AB637A42C74073F1C6">220</mods:start>
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<mods:identifier id="2CBCFC15068845DB6E277D1323C74596" type="DOI">10.26107/RBZ-2020-0019</mods:identifier>
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<mods:identifier id="7993DEC2A875D3CA02BA767D738038CA" type="ISSN">2345-7600</mods:identifier>
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<treatment id="03EACC33FFCF05201387B38CB162FEA6" ID-DOI="http://doi.org/10.5281/zenodo.4577266" ID-Zenodo-Dep="4577266" LSID="urn:lsid:plazi:treatment:03EACC33FFCF05201387B38CB162FEA6" httpUri="http://treatment.plazi.org/id/03EACC33FFCF05201387B38CB162FEA6" lastPageId="3" lastPageNumber="221" pageId="1" pageNumber="221">
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<subSubSection id="C3592EAEFFCF05221387B38CB55DFA68" box="[972,1336,1408,1433]" pageId="1" pageNumber="221" type="nomenclature">
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<paragraph id="8BFC7D25FFCF05221387B38CB55DFA68" blockId="1.[972,1336,1408,1433]" box="[972,1336,1408,1433]" pageId="1" pageNumber="221">
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<heading id="D0B4CA49FFCF05221387B38CB55DFA68" bold="true" box="[972,1336,1408,1433]" centered="true" fontSize="10" level="2" pageId="1" pageNumber="221" reason="2">
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<taxonomicName id="4C4306A6FFCF05221387B38CB55DFA68" authority="Sims & Easton, 1972" authorityName="Sims & Easton" authorityYear="1972" box="[972,1336,1408,1433]" class="Clitellata" family="Megascolecidae" genus="Metaphire" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="1" pageNumber="221" phylum="Annelida" rank="genus">
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<emphasis id="B937A137FFCF05221387B38CB55DFA68" bold="true" box="[972,1336,1408,1433]" pageId="1" pageNumber="221">
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<emphasis id="B937A137FFCF05221387B38CB45BFA68" bold="true" box="[972,1086,1408,1432]" italics="true" pageId="1" pageNumber="221">Metaphire</emphasis>
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<bibRefCitation id="EFD200D4FFCF0522140CB38CB55DFA68" author="Sims RW & Easton EG" box="[1095,1336,1408,1433]" pageId="1" pageNumber="221" pagination="169 - 268" refId="ref11775" refString="Sims RW & Easton EG (1972) A numerical revision of the earthworm genus Pheretima auct. (Megascolecidae: Oligochaeta) with the recognition of new genera and an appendix on the earthworms collected by the Royal Society North Borneo Expedition. Biological Journal of the Linnean Society, 4: 169 - 268." type="journal article" year="1972">Sims & Easton, 1972</bibRefCitation>
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3592EAEFFCF05221308B3B3B4DFFA25" box="[835,1210,1471,1493]" pageId="1" pageNumber="221" type="reference_group">
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<paragraph id="8BFC7D25FFCF05221308B3B3B4DFFA25" blockId="1.[835,1210,1471,1493]" box="[835,1210,1471,1493]" pageId="1" pageNumber="221">
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<treatmentCitationGroup id="AB535A0BFFCF05221308B3B3B4DFFA25" box="[835,1210,1471,1493]" pageId="1" pageNumber="221">
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<treatmentCitation id="0AE25B34FFCF05221308B3B3B4D3FA25" author="Sims RW & Easton EG" box="[835,1206,1471,1493]" page="215" pageId="1" pageNumber="221" year="1972">
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<taxonomicName id="4C4306A6FFCF05221308B3B3B4D3FA25" authority="Sims & Easton, 1972: 215" authorityName="Sims & Easton" authorityPageNumber="215" authorityYear="1972" box="[835,1206,1471,1493]" class="Clitellata" family="Megascolecidae" genus="Metaphire" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="1" pageNumber="221" phylum="Annelida" rank="genus">
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<emphasis id="B937A137FFCF05221308B3B3B3C2FA24" box="[835,935,1471,1492]" italics="true" pageId="1" pageNumber="221">Metaphire</emphasis>
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<bibRefCitation id="EFD200D4FFCF052213E4B3B3B4D3FA25" author="Sims RW & Easton EG" box="[943,1206,1471,1493]" pageId="1" pageNumber="221" pagination="169 - 268" refId="ref11775" refString="Sims RW & Easton EG (1972) A numerical revision of the earthworm genus Pheretima auct. (Megascolecidae: Oligochaeta) with the recognition of new genera and an appendix on the earthworms collected by the Royal Society North Borneo Expedition. Biological Journal of the Linnean Society, 4: 169 - 268." type="journal article" year="1972">Sims & Easton, 1972: 215</bibRefCitation>
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</taxonomicName>
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</treatmentCitation>
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.
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</treatmentCitationGroup>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3592EAEFFCF05221308B3F6B3DDF9C3" pageId="1" pageNumber="221" type="type_taxon">
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<paragraph id="8BFC7D25FFCF05221308B3F6B3DDF9C3" blockId="1.[835,1473,1530,1587]" pageId="1" pageNumber="221">
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<emphasis id="B937A137FFCF05221308B3F6B389F9E2" bold="true" box="[835,1004,1530,1554]" pageId="1" pageNumber="221">
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<typeStatus id="54F8C387FFCF05221308B3F6B3E4F9E2" box="[835,897,1530,1554]" pageId="1" pageNumber="221">Type</typeStatus>
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species.
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</emphasis>
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<taxonomicName id="4C4306A6FFCF052213B3B3F6B5F0F9E2" authority="Kinberg, 1867" authorityName="Kinberg" authorityYear="1867" box="[1016,1429,1530,1554]" class="Aves" family="Trochilidae" genus="Rhodopis" kingdom="Animalia" order="Apodiformes" pageId="1" pageNumber="221" phylum="Chordata" rank="species" species="javanica">
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<emphasis id="B937A137FFCF052213B3B3F6B4BDF9E2" box="[1016,1240,1530,1554]" italics="true" pageId="1" pageNumber="221">Rhodopis javanica</emphasis>
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<bibRefCitation id="EFD200D4FFCF052214AFB3F6B5F0F9E2" author="Kinberg JGH" box="[1252,1429,1530,1554]" pageId="1" pageNumber="221" pagination="97 - 103" refId="ref10978" refString="Kinberg JGH (1867) Annulata Nova. Ofversigt af Kongl. Vetenskaps-Akademiens Forhandlingar, 23: 97 - 103." type="journal article" year="1867">Kinberg, 1867</bibRefCitation>
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</taxonomicName>
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, by monotypy.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3592EAEFFCF05221308B056B41EF942" pageId="1" pageNumber="221" type="distribution">
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<paragraph id="8BFC7D25FFCF05221308B056B41EF942" blockId="1.[834,1474,1626,1714]" pageId="1" pageNumber="221">
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<emphasis id="B937A137FFCF05221308B056B3B3F982" bold="true" box="[835,982,1626,1650]" pageId="1" pageNumber="221">Distribution.</emphasis>
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The genus is widely distributed in the Oriental region, from
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<collectingCountry id="F3543DB5FFCF05221383B077B461F962" box="[968,1028,1659,1682]" name="Japan" pageId="1" pageNumber="221">Japan</collectingCountry>
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through to the Indo-Australian archipelago (
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<bibRefCitation id="EFD200D4FFCF05221302B096B40AF942" author="Sims RW & Easton EG" box="[841,1135,1690,1714]" pageId="1" pageNumber="221" pagination="169 - 268" refId="ref11775" refString="Sims RW & Easton EG (1972) A numerical revision of the earthworm genus Pheretima auct. (Megascolecidae: Oligochaeta) with the recognition of new genera and an appendix on the earthworms collected by the Royal Society North Borneo Expedition. Biological Journal of the Linnean Society, 4: 169 - 268." type="journal article" year="1972">Sims & Easton, 1972: 215</bibRefCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="C3592EAEFFCF05201308B0D6B162FEA6" lastPageId="3" lastPageNumber="223" pageId="1" pageNumber="221" type="discussion">
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<paragraph id="8BFC7D25FFCF05221308B0D6B3E6F842" blockId="1.[835,1474,1754,1970]" pageId="1" pageNumber="221">
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<emphasis id="B937A137FFCF05221308B0D6B3D7F902" bold="true" box="[835,946,1754,1778]" pageId="1" pageNumber="221">Remarks.</emphasis>
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According to
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<bibRefCitation id="EFD200D4FFCF05221418B0D6B529F902" author="Sims RW & Easton EG" box="[1107,1356,1754,1778]" pageId="1" pageNumber="221" pagination="169 - 268" refId="ref11775" refString="Sims RW & Easton EG (1972) A numerical revision of the earthworm genus Pheretima auct. (Megascolecidae: Oligochaeta) with the recognition of new genera and an appendix on the earthworms collected by the Royal Society North Borneo Expedition. Biological Journal of the Linnean Society, 4: 169 - 268." type="journal article" year="1972">Sims & Easton (1972)</bibRefCitation>
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, the genus
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<taxonomicName id="4C4306A6FFCF05221308B0F6B3D7F8E2" authorityName="Sims & Easton" authorityYear="1972" box="[835,946,1786,1810]" class="Clitellata" family="Megascolecidae" genus="Metaphire" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="1" pageNumber="221" phylum="Annelida" rank="genus">
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<emphasis id="B937A137FFCF05221308B0F6B3D7F8E2" box="[835,946,1786,1810]" italics="true" pageId="1" pageNumber="221">Metaphire</emphasis>
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</taxonomicName>
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is distinguished from
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<taxonomicName id="4C4306A6FFCF052214E7B0F6B5DBF8E2" authority="Kinberg, 1867" authorityName="Kinberg" authorityYear="1867" box="[1196,1470,1786,1810]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Opisthopora" pageId="1" pageNumber="221" phylum="Annelida" rank="genus">
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<emphasis id="B937A137FFCF052214E7B0F6B570F8E2" box="[1196,1301,1786,1810]" italics="true" pageId="1" pageNumber="221">Amynthas</emphasis>
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<bibRefCitation id="EFD200D4FFCF05221554B0F6B5DBF8E2" author="Kinberg JGH" box="[1311,1470,1786,1810]" pageId="1" pageNumber="221" pagination="97 - 103" refId="ref10978" refString="Kinberg JGH (1867) Annulata Nova. Ofversigt af Kongl. Vetenskaps-Akademiens Forhandlingar, 23: 97 - 103." type="journal article" year="1867">Kinberg, 1867</bibRefCitation>
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</taxonomicName>
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, mainly by the presence of copulatory pouches, from
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<taxonomicName id="4C4306A6FFCF05221308B136B430F8A2" authority="Kinberg, 1867" authorityName="Kinberg" authorityYear="1867" box="[835,1109,1850,1874]" class="Clitellata" family="Megascolecidae" genus="Pheretima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="1" pageNumber="221" phylum="Annelida" rank="genus">
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<emphasis id="B937A137FFCF05221308B136B3D7F8A2" box="[835,946,1850,1874]" italics="true" pageId="1" pageNumber="221">Pheretima</emphasis>
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<bibRefCitation id="EFD200D4FFCF052213F2B136B430F8A2" author="Kinberg JGH" box="[953,1109,1850,1874]" pageId="1" pageNumber="221" pagination="97 - 103" refId="ref10978" refString="Kinberg JGH (1867) Annulata Nova. Ofversigt af Kongl. Vetenskaps-Akademiens Forhandlingar, 23: 97 - 103." type="journal article" year="1867">Kinberg, 1867</bibRefCitation>
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</taxonomicName>
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, by the absence of micronephridia on the spermathecal ducts, and from
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<taxonomicName id="4C4306A6FFCF052214A4B156B3B6F862" authority="Sims & Easton, 1972" authorityName="Sims & Easton" authorityYear="1972" class="Clitellata" family="Megascolecidae" genus="Pithemera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="1" pageNumber="221" phylum="Annelida" rank="genus">
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<emphasis id="B937A137FFCF052214A4B156B504F882" box="[1263,1377,1882,1906]" italics="true" pageId="1" pageNumber="221">Pithemera</emphasis>
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<bibRefCitation id="EFD200D4FFCF05221527B156B3B6F862" author="Sims RW & Easton EG" pageId="1" pageNumber="221" pagination="169 - 268" refId="ref11775" refString="Sims RW & Easton EG (1972) A numerical revision of the earthworm genus Pheretima auct. (Megascolecidae: Oligochaeta) with the recognition of new genera and an appendix on the earthworms collected by the Royal Society North Borneo Expedition. Biological Journal of the Linnean Society, 4: 169 - 268." type="journal article" year="1972">Sims & Easton, 1972</bibRefCitation>
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</taxonomicName>
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, by the presence of intestinal caeca at or near xxvii.
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</paragraph>
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<paragraph id="8BFC7D25FFCF05201308B1D6B162FEA6" blockId="1.[835,1475,2010,2066]" lastBlockId="3.[136,777,189,342]" lastPageId="3" lastPageNumber="223" pageId="1" pageNumber="221">
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Morphologically,
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<taxonomicName id="4C4306A6FFCF0522145DB1D6B4E9F802" authorityName="Sims & Easton" authorityYear="1972" box="[1046,1164,2010,2034]" class="Clitellata" family="Megascolecidae" genus="Metaphire" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="1" pageNumber="221" phylum="Annelida" rank="genus">
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<emphasis id="B937A137FFCF0522145DB1D6B4E9F802" box="[1046,1164,2010,2034]" italics="true" pageId="1" pageNumber="221">Metaphire</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="4C4306A6FFCF05221480B1D6B55FF802" authorityName="Kinberg" authorityYear="1867" box="[1227,1338,2010,2034]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Opisthopora" pageId="1" pageNumber="221" phylum="Annelida" rank="genus">
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<emphasis id="B937A137FFCF05221480B1D6B55FF802" box="[1227,1338,2010,2034]" italics="true" pageId="1" pageNumber="221">Amynthas</emphasis>
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</taxonomicName>
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are clearly separated by the presence or absence of copulatory pouches. However, recent molecular studies show the polyphyletic relationship between
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<taxonomicName id="4C4306A6FFCD05201124B6D1B1BBFF05" authorityName="Sims & Easton" authorityYear="1972" box="[367,478,221,245]" class="Clitellata" family="Megascolecidae" genus="Metaphire" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="3" pageNumber="223" phylum="Annelida" rank="genus">
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<emphasis id="B937A137FFCD05201124B6D1B1BBFF05" box="[367,478,221,245]" italics="true" pageId="3" pageNumber="223">Metaphire</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="4C4306A6FFCD0520125FB6D1B218FF05" authorityName="Kinberg" authorityYear="1867" box="[532,637,221,245]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Opisthopora" pageId="3" pageNumber="223" phylum="Annelida" rank="genus">
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<emphasis id="B937A137FFCD0520125FB6D1B218FF05" box="[532,637,221,245]" italics="true" pageId="3" pageNumber="223">Amynthas</emphasis>
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</taxonomicName>
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(
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<bibRefCitation id="EFD200D4FFCD052012C0B6D1B0A7FEE5" author="Zhao Q & Cluzeau D & Jiang J & Petit EJ & Briard C & Sun J & Prinzing A & Qiu J" pageId="3" pageNumber="223" pagination="138" refId="ref12030" refString="Zhao Q, Cluzeau D, Jiang J, Petit EJ, Briard C, Sun J, Prinzing A & Qiu J (2015) Molecular phylogeny of pheretimoid earthworms (Haplotaxina: Megascolecidae) based on mitochondrial DNA in Hainan Island, China. Molecular Biology, 4 (4): 138." type="journal article" year="2015">Zhao et al., 2015</bibRefCitation>
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;
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<bibRefCitation id="EFD200D4FFCD05201085B6F1B1B0FEE5" author="Jeratthitikul E & Bantaowong U & Panha S" box="[206,469,253,277]" pageId="3" pageNumber="223" pagination="39 - 47" refId="ref10934" refString="Jeratthitikul E, Bantaowong U & Panha S (2017) DNA barcoding of the Thai species of terrestrial earthworms in the genera Amynthas and Metaphire (Haplotaxida: Megascolecidae). European Journal of Soil Biology, 81: 39 - 47." type="journal article" year="2017">Jeratthitikul et al., 2017</bibRefCitation>
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). Thus, more data is needed for better understanding of the relationship between these two genera.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |