300 lines
31 KiB
XML
300 lines
31 KiB
XML
<document id="88B2E0A5743B1194967DCF6121756354" ID-DOI="10.11646/zootaxa.4104.1.1" ID-GBIF-Dataset="9b3e7716-1099-48ca-803b-620c25d788f1" ID-ISSN="1175-5326" ID-Zenodo-Dep="265809" ID-ZooBank="D598E724-C9E4-4BBA-B25D-511300A47B1D" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1461050759050" checkinUser="plazi" docAuthor="Duellman, William E., Marion, Angela B. & Hedges, Blair" docDate="2016" docId="03EA87A5FFBD123DF3988E41367EF43A" docLanguage="en" docName="zootaxa.4104.pdf.imf" docOrigin="Zootaxa 4104 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Ecnomiohyla Faivovich, Haddad, Garcia, Frost, Campbell 2005" docType="treatment" docVersion="12" lastPageNumber="21" masterDocId="FFD3FFDDFFA91229F30F8D5E333EF35E" masterDocTitle="Phylogenetics, classification, and biogeography of the treefrogs (Amphibia: Anura: Arboranae)" masterLastPageNumber="109" masterPageNumber="1" pageNumber="21" updateTime="1698579563523" updateUser="plazi">
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<mods:mods id="81774702BB563E4E8A147DB75EB898CA" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="D5A2E3CFD82B3E2ED6D014508B7C53C8">
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<mods:title id="13A02AC38CDC16AEE4B027F5D6740848">Phylogenetics, classification, and biogeography of the treefrogs (Amphibia: Anura: Arboranae)</mods:title>
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</mods:titleInfo>
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<mods:name id="294E73911F83DA7FDA6D023B9EC06A82" type="personal">
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<mods:role id="5F5831F1FDA41DAF794966E696862811">
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<mods:roleTerm id="29E9DC1039F012B5BBBFDAAEB0BB32AE">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="D59BAD0ADE545BE682A1914C39ABAD1D">Duellman, William E.</mods:namePart>
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</mods:name>
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<mods:name id="4ADAC358918BD8F3C36E5EA913144B1C" type="personal">
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<mods:roleTerm id="9E3A17E5B2B2015E161A8484C5CB130E">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="DB3ADEFC180C6DE23FCACECEB06511F2">Marion, Angela B.</mods:namePart>
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</mods:name>
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<mods:name id="373DEF0CF003AA9E8E4FB80052BB67CC" type="personal">
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<mods:role id="19813451BFDE9CDA44A933F2A736BE3B">
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<mods:roleTerm id="4293144C2F641BF537AD7572CF752BF6">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="2A2D9963EA7E4FCDC3F063391A7EB5B2">Hedges, Blair</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="39C14A7F4D2081C7203C511C426991A5">text</mods:typeOfResource>
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<mods:titleInfo id="D6667FF3C21A31B97503BBA0E8616044">
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<mods:title id="C35FE786D70C3CCAEF01E92B7F581164">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="AA7C76637A135F421EC3149F175BAF36">
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<mods:date id="96D177439F9CD3A899F83B43F7388A9A">2016</mods:date>
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<mods:detail id="82879FD8070369CD53F1DC7ABBA9CE3B" type="volume">
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<mods:number id="F0219E117A365EA58EDEF61E81B8C800">4104</mods:number>
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</mods:detail>
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<mods:detail id="F6BC316C9E862D5CD4DF1635AC6F549B" type="issue">
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<mods:number id="4AD48E79D707A9876EF4C03713CD8A47">1</mods:number>
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<mods:start id="6C4F0AF37ADBC585C228A674C8A02744">1</mods:start>
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<mods:end id="266E627415133BD76506DCD36C3ECC90">109</mods:end>
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<mods:classification id="1843C7C4869ECF8E08A164C78F427F6B">journal article</mods:classification>
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<mods:identifier id="5C6CAAC0CB859A92F68B23E6C2922D93" type="DOI">10.11646/zootaxa.4104.1.1</mods:identifier>
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<mods:identifier id="ED813E409E6E209D8804526FDD287981" type="GBIF-Dataset">9b3e7716-1099-48ca-803b-620c25d788f1</mods:identifier>
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<mods:identifier id="5E8CDB9B0CC29CD1C82CF5FC50371281" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="705012703D447109296422DB427F1D9F" type="Zenodo-Dep">265809</mods:identifier>
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</mods:mods>
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<treatment id="03EA87A5FFBD123DF3988E41367EF43A" ID-DOI="http://doi.org/10.5281/zenodo.5458504" ID-GBIF-Taxon="119611966" ID-Zenodo-Dep="5458504" LSID="urn:lsid:plazi:treatment:03EA87A5FFBD123DF3988E41367EF43A" httpUri="http://treatment.plazi.org/id/03EA87A5FFBD123DF3988E41367EF43A" lastPageNumber="21" pageId="20" pageNumber="21">
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<subSubSection id="C3596538FFBD123DF3988E41375FF067" box="[151,1121,799,825]" pageId="20" pageNumber="21" type="nomenclature">
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<paragraph id="8BFC36B3FFBD123DF3988E41375FF067" blockId="20.[151,1121,799,825]" box="[151,1121,799,825]" pageId="20" pageNumber="21">
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<heading id="D0B481DFFFBD123DF3988E41375FF067" bold="true" box="[151,1121,799,825]" fontSize="11" level="1" pageId="20" pageNumber="21" reason="1">
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<taxonomicName id="4C434D30FFBD123DF3988E41375FF067" ID-CoL="49TJ" authority="Faivovich, Haddad, Garcia, Frost, Campbell" authorityName="Faivovich, Haddad, Garcia, Frost, Campbell" authorityYear="2005" box="[151,1121,799,825]" class="Amphibia" family="Hylidae" genus="Ecnomiohyla" kingdom="Animalia" order="Anura" pageId="20" pageNumber="21" phylum="Chordata" rank="genus">
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<emphasis id="B937EAA1FFBD123DF3988E41375FF067" bold="true" box="[151,1121,799,825]" pageId="20" pageNumber="21">
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<emphasis id="B937EAA1FFBD123DF3988E413206F067" bold="true" box="[151,312,799,825]" italics="true" pageId="20" pageNumber="21">Ecnomiohyla</emphasis>
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<bibRefCitation id="EFD24B42FFBD123DF2308E41375FF067" author="Faivovich" box="[319,1121,799,825]" pageId="20" pageNumber="21" refString="Faivovich, J., Haddad, C. F. B., Garcia, P. C. A., Frost, D. R., Campbell, J. A. & Wheeler, W. C. (2005) Systematic review of the frog family Hylidae, with special reference to Hylinae: phylogenetic analysis and taxonomic revision. Bulletin of the American Museum of Natural History, 294, 1 - 240. http: // dx. doi. org / 10.1206 / 0003 - 0090 (2005) 294 [0001: SROTFF] 2.0. CO; 2" type="journal article" year="2005">Faivovich, Haddad, Garcia, Frost, Campbell, and Wheeler, 2005</bibRefCitation>
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3596538FFBD123DF3988E38376DF782" pageId="20" pageNumber="21" type="reference_group">
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<paragraph id="8BFC36B3FFBD123DF3988E3832C9F0C5" blockId="20.[151,1436,870,923]" pageId="20" pageNumber="21">
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<treatmentCitationGroup id="AB53119DFFBD123DF3988E3832C9F0C5" pageId="20" pageNumber="21">
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<taxonomicName id="4C434D30FFBD123DF3988E3830FFF022" ID-CoL="49TJ" authority="Faivovich, Haddad, Garcia, Frost, Campbell" authorityName="Faivovich, Haddad, Garcia, Frost, Campbell" authorityYear="2005" box="[151,961,870,892]" class="Amphibia" family="Hylidae" genus="Ecnomiohyla" kingdom="Animalia" order="Anura" pageId="20" pageNumber="21" phylum="Chordata" rank="genus">
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<emphasis id="B937EAA1FFBD123DF3988E383223F022" box="[151,285,870,892]" italics="true" pageId="20" pageNumber="21">Ecnomiohyla</emphasis>
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<treatmentCitation id="0AE210A2FFBD123DF22B8E3830FFF022" ID-DOI="http://doi.org/10.5281/zenodo.5457768" ID-Zenodo-Dep="5457768" author="Faivovich" box="[292,961,870,892]" httpUri="http://treatment.plazi.org/id/03D887A5FF8C8934FF37FB4ECD6AFC90" page="100" pageId="20" pageNumber="21" year="2005">
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<bibRefCitation id="EFD24B42FFBD123DF22B8E3830FFF022" author="Faivovich" box="[292,961,870,892]" pageId="20" pageNumber="21" refString="Faivovich, J., Haddad, C. F. B., Garcia, P. C. A., Frost, D. R., Campbell, J. A. & Wheeler, W. C. (2005) Systematic review of the frog family Hylidae, with special reference to Hylinae: phylogenetic analysis and taxonomic revision. Bulletin of the American Museum of Natural History, 294, 1 - 240. http: // dx. doi. org / 10.1206 / 0003 - 0090 (2005) 294 [0001: SROTFF] 2.0. CO; 2" type="journal article" year="2005">Faivovich, Haddad, Garcia, Frost, Campbell, and Wheeler, 2005</bibRefCitation>
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</treatmentCitation>
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</taxonomicName>
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:100.
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<typeStatus id="54F88811FFBD123DF0F88E393717F022" box="[1015,1065,871,892]" pageId="20" pageNumber="21">Type</typeStatus>
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species:
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<taxonomicName id="4C434D30FFBD123DF7858E3833C2F0C5" ID-CoL="3P3PM" authority="Cope, 1886" authorityName="Cope" authorityYear="1886" class="Amphibia" family="Hylidae" genus="Hypsiboas" kingdom="Animalia" order="Anura" pageId="20" pageNumber="21" phylum="Chordata" rank="species" species="miliarius">
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<emphasis id="B937EAA1FFBD123DF7858E383666F022" box="[1162,1368,870,892]" italics="true" pageId="20" pageNumber="21">Hypsiboas miliarius</emphasis>
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Cope, 1886
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</taxonomicName>
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, by original designation.
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</treatmentCitationGroup>
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</paragraph>
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<paragraph id="8BFC36B3FFBD123DF3988E993133F776" blockId="20.[151,1436,967,1892]" pageId="20" pageNumber="21">
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<emphasis id="B937EAA1FFBD123DF3988E993228F0BE" bold="true" box="[151,278,967,992]" pageId="20" pageNumber="21">Definition.</emphasis>
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Large treefrogs (SVL in males to
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<quantity id="4CBB9B56FFBD123DF1B78E963024F0BE" box="[696,794,968,992]" metricMagnitude="-1" metricUnit="m" metricValue="1.1" pageId="20" pageNumber="21" unit="mm" value="110.0">110 mm</quantity>
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) with dermal fringes on the outer edges of the limbs, extensive webbing on the hands and feet, and an enlarged prepollex (
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<figureCitation id="13782A36FFBD123DF0A08EB530CAF75A" box="[943,1012,1003,1028]" captionStart="FIGURE 7. A" captionStartId="20.[151,250,671,693]" captionTargetBox="[151,1436,195,649]" captionTargetId="figure@20.[151,1436,195,649]" captionTargetPageId="20" captionText="FIGURE 7. A. Rheohyla miotympanum, KU 100965. Salto Cola de Caballo, 710 m, Nuevo León, Mexico. B. Ecnomiohyla miliaria, KU 101610, Finca Santa Clara, 1200 m, Chiriquí, Panama. Both by W. E. Duellman." httpUri="https://zenodo.org/record/265819/files/figure.png" pageId="20" pageNumber="21">Fig. 7</figureCitation>
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B). Tadpoles with a LTRF of 2/3 and developing in water in tree holes.
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</paragraph>
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<paragraph id="8BFC36B3FFBD123DF3C8896D376DF782" blockId="20.[151,1436,967,1892]" pageId="20" pageNumber="21">
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<emphasis id="B937EAA1FFBD123DF3C8896D3210F712" bold="true" box="[199,302,1075,1100]" pageId="20" pageNumber="21">Content.</emphasis>
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Twelve species:
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<taxonomicName id="4C434D30FFBD123DF2E1896D362FF715" ID-CoL="38NH6" authority="Batista, Hertz, Mebert, Kohler, Lotzkat, Ponce" authorityName="Batista, Hertz, Mebert, Kohler, Lotzkat, Ponce" box="[494,1297,1075,1100]" class="Amphibia" family="Hylidae" genus="Ecnomiohyla" kingdom="Animalia" order="Anura" pageId="20" pageNumber="21" phylum="Chordata" rank="species" species="bailarina">
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<emphasis id="B937EAA1FFBD123DF2E1896D31C1F712" box="[494,767,1075,1100]" italics="true" pageId="20" pageNumber="21">Ecnomiohyla bailarina*</emphasis>
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Batista, Hertz, Mebert, Köhler, Lotzkat, Ponce
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</taxonomicName>
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, and Vesely,
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<emphasis id="B937EAA1FFBD123DF3988909323BF72E" box="[151,261,1111,1136]" italics="true" pageId="20" pageNumber="21">
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<taxonomicName id="4C434D30FFBD123DF398890933C4F72E" ID-CoL="38NH7" authority="Duellman" authorityName="Duellman" box="[151,250,1111,1136]" class="Amphibia" family="Hylidae" genus="Ecnomiohyla" kingdom="Animalia" order="Anura" pageId="20" pageNumber="21" phylum="Chordata" rank="species" species="echinata">echinata</taxonomicName>
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*
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</emphasis>
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(Duellman),
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<emphasis id="B937EAA1FFBD123DF2A689093169F72E" box="[425,599,1111,1136]" italics="true" pageId="20" pageNumber="21">
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<taxonomicName id="4C434D30FFBD123DF2A689093173F72E" ID-CoL="38NH8" authority="Taylor" authorityName="Taylor" box="[425,589,1111,1136]" class="Amphibia" family="Hylidae" genus="Ecnomiohyla" kingdom="Animalia" order="Anura" pageId="20" pageNumber="21" phylum="Chordata" rank="species" species="fimbrimembra">fimbrimembra</taxonomicName>
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*
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</emphasis>
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(Taylor),
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<taxonomicName id="4C434D30FFBD123DF1DA890930B0F72E" ID-CoL="38NH9" authority="Cope" authorityName="Cope" box="[725,910,1111,1136]" class="Amphibia" family="Hylidae" genus="Ecnomiohyla" kingdom="Animalia" order="Anura" pageId="20" pageNumber="21" phylum="Chordata" rank="species" species="miliaria">
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<emphasis id="B937EAA1FFBD123DF1DA89093011F72E" box="[725,815,1111,1136]" italics="true" pageId="20" pageNumber="21">miliaria</emphasis>
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(Cope)
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</taxonomicName>
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,
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<taxonomicName id="4C434D30FFBD123DF0AF890730D1F72E" ID-CoL="6DWMQ" box="[928,1007,1113,1136]" class="Amphibia" family="Hylidae" genus="Ecnomiohyla" kingdom="Animalia" order="Anura" pageId="20" pageNumber="21" phylum="Chordata" rank="species" species="minera">
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<emphasis id="B937EAA1FFBD123DF0AF890730D1F72E" box="[928,1007,1113,1136]" italics="true" pageId="20" pageNumber="21">minera</emphasis>
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</taxonomicName>
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(Wilson, McCranie, and Williams),
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<emphasis id="B937EAA1FFBD123DF39889253263F7CA" box="[151,349,1147,1172]" italics="true" pageId="20" pageNumber="21">
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<taxonomicName id="4C434D30FFBD123DF3988925326CF7CA" ID-CoL="38NHC" authority="Dunn" authorityName="Dunn" box="[151,338,1147,1172]" class="Amphibia" family="Hylidae" genus="Ecnomiohyla" kingdom="Animalia" order="Anura" pageId="20" pageNumber="21" phylum="Chordata" rank="species" species="phantasmagoria">phantasmagoria</taxonomicName>
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*
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</emphasis>
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(Dunn),
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<taxonomicName id="4C434D30FFBD123DF2DA89253754F7CA" ID-CoL="38NHD" authority="Mendelson, Savage, Griffith, Ross, Kubicki" authorityName="Mendelson, Savage, Griffith, Ross, Kubicki" box="[469,1130,1147,1172]" class="Amphibia" family="Hylidae" genus="Ecnomiohyla" kingdom="Animalia" order="Anura" pageId="20" pageNumber="21" phylum="Chordata" rank="species" species="rabborum">
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<emphasis id="B937EAA1FFBD123DF2DA8925317BF7CA" box="[469,581,1147,1172]" italics="true" pageId="20" pageNumber="21">rabborum</emphasis>
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Mendelson, Savage, Griffith, Ross, Kubicki
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</taxonomicName>
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, and Gagliardo,
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<emphasis id="B937EAA1FFBD123DF631892536A2F7CA" box="[1342,1436,1147,1172]" italics="true" pageId="20" pageNumber="21">salvaje*</emphasis>
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(Wilson, McCranie, and Williams),
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<emphasis id="B937EAA1FFBD123DF12889C13151F7E6" box="[551,623,1183,1208]" italics="true" pageId="20" pageNumber="21">
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<taxonomicName id="4C434D30FFBD123DF12889C1315BF7E6" ID-CoL="38NHF" authority="Savage" authorityName="Savage" box="[551,613,1183,1208]" class="Amphibia" family="Hylidae" genus="Ecnomiohyla" kingdom="Animalia" order="Anura" pageId="20" pageNumber="21" phylum="Chordata" rank="species" species="sukia">sukia</taxonomicName>
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*
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</emphasis>
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Savage and Kubicki,
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<emphasis id="B937EAA1FFBD123DF06A89C130DEF7E6" box="[869,992,1183,1208]" italics="true" pageId="20" pageNumber="21">
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<taxonomicName id="4C434D30FFBD123DF06A89C130EBF7E6" ID-CoL="38NHG" authority="Duellman" authorityName="Duellman" box="[869,981,1183,1208]" class="Amphibia" family="Hylidae" genus="Ecnomiohyla" kingdom="Animalia" order="Anura" pageId="20" pageNumber="21" phylum="Chordata" rank="species" species="thysanota">thysanota</taxonomicName>
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*
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</emphasis>
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(Duellman),
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<emphasis id="B937EAA1FFBD123DF77A89C137CBF7E6" box="[1141,1269,1183,1208]" italics="true" pageId="20" pageNumber="21">valancifer*</emphasis>
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(Firschein and Smith), and
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<emphasis id="B937EAA1FFBD123DF22F899A328AF785" box="[288,436,1220,1243]" italics="true" pageId="20" pageNumber="21">
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<taxonomicName id="4C434D30FFBD123DF22F899A3297F785" ID-CoL="38NHK" authority="Batista" authorityName="Batista" box="[288,425,1220,1243]" class="Amphibia" family="Hylidae" genus="Ecnomiohyla" kingdom="Animalia" order="Anura" pageId="20" pageNumber="21" phylum="Chordata" rank="species" species="veraguensis">veraguensis</taxonomicName>
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*
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</emphasis>
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Batista, Hertz, Mebert, Köhler, Lotzkat, Ponce, and Vesely.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3596538FFBD123DF3C889B93655F65E" box="[199,1387,1255,1280]" pageId="20" pageNumber="21" type="distribution">
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<paragraph id="8BFC36B3FFBD123DF3C889B93655F65E" blockId="20.[151,1436,967,1892]" box="[199,1387,1255,1280]" pageId="20" pageNumber="21">
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<emphasis id="B937EAA1FFBD123DF3C889B9325EF65E" bold="true" box="[199,352,1255,1280]" pageId="20" pageNumber="21">Distribution.</emphasis>
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Southern
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<collectingCountry id="F3547623FFBD123DF2DC89B93116F65E" box="[467,552,1255,1280]" name="Mexico" pageId="20" pageNumber="21">Mexico</collectingCountry>
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through Central
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<collectingCountry id="F3547623FFBD123DF1E889B93076F65E" box="[743,840,1255,1280]" name="United States of America" pageId="20" pageNumber="21">America</collectingCountry>
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to western
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<collectingCountry id="F3547623FFBD123DF0C689B93709F65E" box="[969,1079,1255,1280]" name="Colombia" pageId="20" pageNumber="21">Colombia</collectingCountry>
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and northwestern
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<collectingCountry id="F3547623FFBD123DF60789B93656F65E" box="[1288,1384,1255,1280]" name="Ecuador" pageId="20" pageNumber="21">Ecuador</collectingCountry>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3596538FFBD123DF3C888553111F616" pageId="20" pageNumber="21" type="etymology">
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<paragraph id="8BFC36B3FFBD123DF3C888553111F616" blockId="20.[151,1436,967,1892]" pageId="20" pageNumber="21">
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<emphasis id="B937EAA1FFBD123DF3C888553270F67A" bold="true" box="[199,334,1291,1316]" pageId="20" pageNumber="21">Etymology.</emphasis>
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According to Faivovich
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<emphasis id="B937EAA1FFBD123DF17288533185F67A" box="[637,699,1291,1316]" italics="true" pageId="20" pageNumber="21">et al.</emphasis>
|
|
(2005:100), “From the Greek,
|
|
<emphasis id="B937EAA1FFBD123DF73A88533798F67A" box="[1077,1190,1293,1316]" italics="true" pageId="20" pageNumber="21">ecnomios,</emphasis>
|
|
meaning marvelous, unusual …” The gender is feminine.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="C3596538FFBD123DF3C8880D367EF43A" pageId="20" pageNumber="21" type="discussion">
|
|
<paragraph id="8BFC36B3FFBD123DF3C8880D32B8F6A2" blockId="20.[151,1436,967,1892]" pageId="20" pageNumber="21">
|
|
<emphasis id="B937EAA1FFBD123DF3C8880D320AF632" bold="true" box="[199,308,1363,1388]" pageId="20" pageNumber="21">Remarks</emphasis>
|
|
.
|
|
<bibRefCitation id="EFD24B42FFBD123DF24C880D3168F632" box="[323,598,1363,1388]" pageId="20" pageNumber="21" refString="Mendelson, J. R., III, Savage, J. M., Griffith, E., Ross, H., Kubicki, B. & Gagliardo, R. (2008) Spectacular new gliding species of Ecnomiohyla (Anura: Hylidae) from central Panama. Journal of Herpetology, 42, 750 - 759. http: // dx. doi. org / 10.1670 / 08 - 025 R 1.1" type="journal article">
|
|
Mendelson
|
|
<emphasis id="B937EAA1FFBD123DF2C7880B313CF632" box="[456,514,1363,1388]" italics="true" pageId="20" pageNumber="21">et al.</emphasis>
|
|
(2008)
|
|
</bibRefCitation>
|
|
,
|
|
<bibRefCitation id="EFD24B42FFBD123DF16C880A30A3F632" author="Savage" box="[611,925,1363,1388]" pageId="20" pageNumber="21" refString="Savage, J. M. & Kubicki, B. (2010) A new species of fringe-limbed frog, genus Ecnomiohyla (Anura: Hylidae) from the Atlantic slope of Costa Rica, Central America. Zootaxa, 2719, 21 - 34." type="journal article" year="2010">Savage and Kubicki (2010)</bibRefCitation>
|
|
, and
|
|
<bibRefCitation id="EFD24B42FFBD123DF0D4880D3780F632" box="[987,1214,1363,1388]" pageId="20" pageNumber="21" refString="Batista, A., Hertz, A., Mebert, K., Kohler, G., Lotzkat, S., Ponce, M. & Vesely, M. (2014) Two new fringe-limbed frogs of the genus Ecnomiohyla (Anura: Hylidae) from Panama. Zootaxa, 3826 (3), 449 - 474. http: // dx. doi. org / 10.11646 / zootaxa. 3826.3.2" type="journal article">
|
|
Batista
|
|
<emphasis id="B937EAA1FFBD123DF73E880B3755F632" box="[1073,1131,1363,1388]" italics="true" pageId="20" pageNumber="21">et al.</emphasis>
|
|
(2014)
|
|
</bibRefCitation>
|
|
have expanded our knowledge of this genus, which still contains species known only from their
|
|
<typeStatus id="54F88811FFBD123DF0E188293765F6CE" box="[1006,1115,1399,1424]" pageId="20" pageNumber="21" type="holotype">holotypes</typeStatus>
|
|
(e.g.,
|
|
<taxonomicName id="4C434D30FFBD123DF7AE882936A2F6CE" box="[1185,1436,1399,1424]" class="Amphibia" family="Hylidae" genus="Ecnomiohyla" kingdom="Animalia" order="Anura" pageId="20" pageNumber="21" phylum="Chordata" rank="species" species="echinata">
|
|
<emphasis id="B937EAA1FFBD123DF7AE882936A2F6CE" box="[1185,1436,1399,1424]" italics="true" pageId="20" pageNumber="21">Ecnomiohyla echinata</emphasis>
|
|
</taxonomicName>
|
|
and
|
|
<taxonomicName id="4C434D30FFBD123DF3C688C33266F6EA" box="[201,344,1435,1460]" class="Amphibia" family="Hylidae" genus="Ecnomiohyla" kingdom="Animalia" order="Anura" pageId="20" pageNumber="21" phylum="Chordata" rank="species" species="thysanota">
|
|
<emphasis id="B937EAA1FFBD123DF3C688C33266F6EA" box="[201,344,1435,1460]" italics="true" pageId="20" pageNumber="21">E. thysanota</emphasis>
|
|
</taxonomicName>
|
|
). For more than half of a century,
|
|
<taxonomicName id="4C434D30FFBD123DF1E288C330F9F6EA" box="[749,967,1435,1460]" class="Amphibia" family="Hylidae" genus="Ecnomiohyla" kingdom="Animalia" order="Anura" pageId="20" pageNumber="21" phylum="Chordata" rank="species" species="phantasmagoria">
|
|
<emphasis id="B937EAA1FFBD123DF1E288C330F9F6EA" box="[749,967,1435,1460]" italics="true" pageId="20" pageNumber="21">E. phantasmagoria</emphasis>
|
|
</taxonomicName>
|
|
has been known only from the
|
|
<typeStatus id="54F88811FFBD123DF63588C536A2F6EA" box="[1338,1436,1435,1460]" pageId="20" pageNumber="21" type="holotype">holotype</typeStatus>
|
|
from the Río Cauca in
|
|
<collectingCountry id="F3547623FFBD123DF29688E13134F686" box="[409,522,1471,1496]" name="Colombia" pageId="20" pageNumber="21">Colombia</collectingCountry>
|
|
, but recently was discovered in the Provincia de Esmeraldas in
|
|
<collectingCountry id="F3547623FFBD123DF7D688E13608F686" box="[1241,1334,1471,1496]" name="Ecuador" pageId="20" pageNumber="21">Ecuador</collectingCountry>
|
|
(Ortega- Andrade
|
|
<emphasis id="B937EAA1FFBD123DF3F188BB3208F6A2" box="[254,310,1507,1532]" italics="true" pageId="20" pageNumber="21">et al.</emphasis>
|
|
2010).
|
|
</paragraph>
|
|
<paragraph id="8BFC36B3FFBD123DF3C88B563022F5EE" blockId="20.[151,1436,967,1892]" pageId="20" pageNumber="21">
|
|
Our tree (
|
|
<figureCitation id="13782A36FFBD123DF2368B5932BEF57E" box="[313,384,1543,1568]" captionStart="FIGURE 4" captionStartId="10.[151,250,1915,1937]" captionTargetBox="[205,1424,1381,1881]" captionTargetId="figure@10.[174,1472,1381,1884]" captionTargetPageId="10" captionText="FIGURE 4. Maximum likelihood phylogeny of arboranan frogs based on DNA sequence data from 19 genes (16,128 aligned sites). The tree is rooted with Ceuthomantis smaragdinus, Dendrobates auratus, Haddadus binotatus, and Rhinoderma darwinii (not shown). Bootstrap support values are indicated at nodes." httpUri="https://zenodo.org/record/265813/files/figure.png" pageId="20" pageNumber="21">Fig. 4</figureCitation>
|
|
) shows
|
|
<taxonomicName id="4C434D30FFBD123DF2EC8B5931CFF57E" box="[483,753,1543,1568]" class="Amphibia" family="Hylidae" genus="Ecnomiohyla" kingdom="Animalia" order="Anura" pageId="20" pageNumber="21" phylum="Chordata" rank="species" species="rabborum">
|
|
<emphasis id="B937EAA1FFBD123DF2EC8B5931CFF57E" box="[483,753,1543,1568]" italics="true" pageId="20" pageNumber="21">Ecnomiohyla rabborum</emphasis>
|
|
</taxonomicName>
|
|
as the sister species of
|
|
<taxonomicName id="4C434D30FFBD123DF7038B5737B6F57E" box="[1036,1160,1543,1568]" class="Amphibia" family="Hylidae" genus="Ecnomiohyla" kingdom="Animalia" order="Anura" pageId="20" pageNumber="21" phylum="Chordata" rank="species" species="malaria">
|
|
<emphasis id="B937EAA1FFBD123DF7038B5737B6F57E" box="[1036,1160,1543,1568]" italics="true" pageId="20" pageNumber="21">E. malaria</emphasis>
|
|
</taxonomicName>
|
|
+
|
|
<emphasis id="B937EAA1FFBD123DF7A38B573618F57E" box="[1196,1318,1545,1568]" italics="true" pageId="20" pageNumber="21">
|
|
<taxonomicName id="4C434D30FFBD123DF7A38B57361CF57E" box="[1196,1314,1545,1568]" class="Amphibia" family="Hylidae" genus="Ecnomiohyla" kingdom="Animalia" order="Anura" pageId="20" pageNumber="21" phylum="Chordata" rank="species" species="minera">E. minera</taxonomicName>
|
|
.
|
|
</emphasis>
|
|
The most extensive molecular phylogenetic tree, based only on the 16S rRNA mitochondrial gene, of
|
|
<taxonomicName id="4C434D30FFBD123DF7AC8B753609F51A" box="[1187,1335,1579,1604]" class="Amphibia" family="Hylidae" genus="Ecnomiohyla" kingdom="Animalia" order="Anura" pageId="20" pageNumber="21" phylum="Chordata" rank="genus">
|
|
<emphasis id="B937EAA1FFBD123DF7AC8B753609F51A" box="[1187,1335,1579,1604]" italics="true" pageId="20" pageNumber="21">Ecnomiohyla</emphasis>
|
|
</taxonomicName>
|
|
contains six species (
|
|
<bibRefCitation id="EFD24B42FFBD123DF2268B113135F536" author="Batista" box="[297,523,1615,1640]" pageId="20" pageNumber="21" refString="Batista, A., Hertz, A., Mebert, K., Kohler, G., Lotzkat, S., Ponce, M. & Vesely, M. (2014) Two new fringe-limbed frogs of the genus Ecnomiohyla (Anura: Hylidae) from Panama. Zootaxa, 3826 (3), 449 - 474. http: // dx. doi. org / 10.11646 / zootaxa. 3826.3.2" type="journal article" year="2014">
|
|
Batista
|
|
<emphasis id="B937EAA1FFBD123DF28B8B0F32FCF536" box="[388,450,1615,1640]" italics="true" pageId="20" pageNumber="21">et al.</emphasis>
|
|
2014
|
|
</bibRefCitation>
|
|
). In their maximum likelihood consensus tree, two well-supported clades are evident. One contains
|
|
<taxonomicName id="4C434D30FFBD123DF29E8B2B316EF5D2" box="[401,592,1651,1676]" class="Amphibia" family="Hylidae" genus="Ecnomiohyla" kingdom="Animalia" order="Anura" pageId="20" pageNumber="21" phylum="Chordata" rank="species" species="fimbrimembra">
|
|
<emphasis id="B937EAA1FFBD123DF29E8B2B316EF5D2" box="[401,592,1651,1676]" italics="true" pageId="20" pageNumber="21">E. fimbrimembra</emphasis>
|
|
</taxonomicName>
|
|
as the sister species of
|
|
<taxonomicName id="4C434D30FFBD123DF0598B2B30DBF5D2" box="[854,997,1651,1676]" class="Amphibia" family="Hylidae" genus="Ecnomiohyla" kingdom="Animalia" order="Anura" pageId="20" pageNumber="21" phylum="Chordata" rank="species" species="rabborum">
|
|
<emphasis id="B937EAA1FFBD123DF0598B2B30DBF5D2" box="[854,997,1651,1676]" italics="true" pageId="20" pageNumber="21">E. rabborum</emphasis>
|
|
</taxonomicName>
|
|
+
|
|
<taxonomicName id="4C434D30FFBD123DF70D8B2B37B7F5D2" box="[1026,1161,1651,1676]" class="Amphibia" family="Hylidae" genus="Ecnomiohyla" kingdom="Animalia" order="Anura" pageId="20" pageNumber="21" phylum="Chordata" rank="species" species="bailarina">
|
|
<emphasis id="B937EAA1FFBD123DF70D8B2B37B7F5D2" box="[1026,1161,1651,1676]" italics="true" pageId="20" pageNumber="21">E. bailarina</emphasis>
|
|
</taxonomicName>
|
|
; the second clade has
|
|
<taxonomicName id="4C434D30FFBD123DF68B8B2B33CFF5EE" class="Amphibia" family="Hylidae" genus="Ecnomiohyla" kingdom="Animalia" order="Anura" pageId="20" pageNumber="21" phylum="Chordata" rank="species" species="miliaria">
|
|
<emphasis id="B937EAA1FFBD123DF68B8B2B33CFF5EE" italics="true" pageId="20" pageNumber="21">E. miliaria</emphasis>
|
|
</taxonomicName>
|
|
as the sister species of
|
|
<taxonomicName id="4C434D30FFBD123DF2F78BC7316DF5EE" box="[504,595,1687,1712]" class="Amphibia" family="Hylidae" genus="Ecnomiohyla" kingdom="Animalia" order="Anura" pageId="20" pageNumber="21" phylum="Chordata" rank="species" species="sukia">
|
|
<emphasis id="B937EAA1FFBD123DF2F78BC7316DF5EE" box="[504,595,1687,1712]" italics="true" pageId="20" pageNumber="21">E. sukia</emphasis>
|
|
</taxonomicName>
|
|
+
|
|
<emphasis id="B937EAA1FFBD123DF17F8BC73022F5EE" box="[624,796,1689,1712]" italics="true" pageId="20" pageNumber="21">
|
|
<taxonomicName id="4C434D30FFBD123DF17F8BC73026F5EE" box="[624,792,1689,1712]" class="Amphibia" family="Hylidae" genus="Ecnomiohyla" kingdom="Animalia" order="Anura" pageId="20" pageNumber="21" phylum="Chordata" rank="species" species="veraguensis">E. veraguensis</taxonomicName>
|
|
.
|
|
</emphasis>
|
|
</paragraph>
|
|
<paragraph id="8BFC36B3FFBD123DF3C88BE5367EF43A" blockId="20.[151,1436,967,1892]" pageId="20" pageNumber="21">
|
|
<bibRefCitation id="EFD24B42FFBD123DF3C88BE532E5F58A" box="[199,475,1723,1748]" pageId="20" pageNumber="21" refString="Mendelson, J. R., III, Savage, J. M., Griffith, E., Ross, H., Kubicki, B. & Gagliardo, R. (2008) Spectacular new gliding species of Ecnomiohyla (Anura: Hylidae) from central Panama. Journal of Herpetology, 42, 750 - 759. http: // dx. doi. org / 10.1670 / 08 - 025 R 1.1" type="journal article">
|
|
Mendelson
|
|
<emphasis id="B937EAA1FFBD123DF2428BE332B6F58A" box="[333,392,1723,1748]" italics="true" pageId="20" pageNumber="21">et al.</emphasis>
|
|
(2008)
|
|
</bibRefCitation>
|
|
emphasized that the Amazonian “
|
|
<taxonomicName id="4C434D30FFBD123DF0668BE53712F58A" authority="Boulenger" authorityName="Boulenger" box="[873,1068,1723,1748]" class="Amphibia" family="Hylidae" genus="Hyla" kingdom="Animalia" order="Anura" pageId="20" pageNumber="21" phylum="Chordata" rank="species" species="tuberculosa">
|
|
<emphasis id="B937EAA1FFBD123DF0668BE53712F58A" box="[873,1068,1723,1748]" italics="true" pageId="20" pageNumber="21">Hyla tuberculosa</emphasis>
|
|
</taxonomicName>
|
|
” Boulenger is not a member of
|
|
<taxonomicName id="4C434D30FFBD123DF3988B813215F5A6" box="[151,299,1759,1784]" class="Amphibia" family="Hylidae" genus="Ecnomiohyla" kingdom="Animalia" order="Anura" pageId="20" pageNumber="21" phylum="Chordata" rank="genus">
|
|
<emphasis id="B937EAA1FFBD123DF3988B813215F5A6" box="[151,299,1759,1784]" italics="true" pageId="20" pageNumber="21">Ecnomiohyla</emphasis>
|
|
</taxonomicName>
|
|
and should be designated
|
|
<emphasis id="B937EAA1FFBD123DF1648BBF302CF5A6" box="[619,786,1759,1784]" italics="true" pageId="20" pageNumber="21">incertae sedis.</emphasis>
|
|
<bibRefCitation id="EFD24B42FFBD123DF0128BBE3761F5A6" author="Savage" box="[797,1119,1759,1784]" pageId="20" pageNumber="21" refString="Savage, J. M. & Kubicki, B. (2010) A new species of fringe-limbed frog, genus Ecnomiohyla (Anura: Hylidae) from the Atlantic slope of Costa Rica, Central America. Zootaxa, 2719, 21 - 34." type="journal article" year="2010">Savage and Kubicki (2010)</bibRefCitation>
|
|
regarded the placement of
|
|
<taxonomicName id="4C434D30FFBD123DF3988A5D32D2F442" authority="in Economiohyla" authorityName="in Economiohyla" box="[151,492,1795,1820]" class="Amphibia" family="Hylidae" genus="Hyla" kingdom="Animalia" order="Anura" pageId="20" pageNumber="21" phylum="Chordata" rank="species" species="tuberculosa">
|
|
<emphasis id="B937EAA1FFBD123DF3988A5D3224F442" box="[151,282,1795,1820]" italics="true" pageId="20" pageNumber="21">tuberculosa</emphasis>
|
|
in
|
|
<emphasis id="B937EAA1FFBD123DF2458A5D32D2F442" box="[330,492,1795,1820]" italics="true" pageId="20" pageNumber="21">Economiohyla</emphasis>
|
|
</taxonomicName>
|
|
as problematic because it lacked the synapomorphic morphological characters, principally an enlarged prepollex with keratinous spines, of the genus. We await molecular data for this species but herein we consider “
|
|
<taxonomicName id="4C434D30FFBD123DF2718A153100F43A" box="[382,574,1867,1892]" class="Amphibia" family="Hylidae" genus="Hyla" kingdom="Animalia" order="Anura" pageId="20" pageNumber="21" phylum="Chordata" rank="species" species="tuberculosa">
|
|
<emphasis id="B937EAA1FFBD123DF2718A153100F43A" box="[382,574,1867,1892]" italics="true" pageId="20" pageNumber="21">Hyla tuberculosa</emphasis>
|
|
</taxonomicName>
|
|
” to be a member of the South American catch-all genus,
|
|
<emphasis id="B937EAA1FFBD123DF7CD8A15367EF43A" box="[1218,1344,1867,1892]" italics="true" pageId="20" pageNumber="21">
|
|
<taxonomicName id="4C434D30FFBD123DF7CD8A153602F43A" box="[1218,1340,1867,1892]" class="Amphibia" family="Hylidae" genus="Hypsiboas" kingdom="Animalia" order="Anura" pageId="20" pageNumber="21" phylum="Chordata" rank="genus">Hypsiboas</taxonomicName>
|
|
.
|
|
</emphasis>
|
|
</paragraph>
|
|
</subSubSection>
|
|
</treatment>
|
|
</document> |