treatments-xml/data/03/EA/87/03EA87A5FF891209F3988C823020F674.xml
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<document id="1730F4053FD1CCF2EA0D123A49922FB5" ID-DOI="10.11646/zootaxa.4104.1.1" ID-GBIF-Dataset="9b3e7716-1099-48ca-803b-620c25d788f1" ID-ISSN="1175-5326" ID-Zenodo-Dep="265809" ID-ZooBank="D598E724-C9E4-4BBA-B25D-511300A47B1D" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1461050759050" checkinUser="plazi" docAuthor="Duellman, William E., Marion, Angela B. &amp; Hedges, Blair" docDate="2016" docId="03EA87A5FF891209F3988C823020F674" docLanguage="en" docName="zootaxa.4104.pdf.imf" docOrigin="Zootaxa 4104 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Callimedusa Duellman, Marion &amp; Hedges, 2016, new genus" docType="treatment" docVersion="11" lastPageNumber="33" masterDocId="FFD3FFDDFFA91229F30F8D5E333EF35E" masterDocTitle="Phylogenetics, classification, and biogeography of the treefrogs (Amphibia: Anura: Arboranae)" masterLastPageNumber="109" masterPageNumber="1" pageNumber="33" updateTime="1698579563523" updateUser="plazi">
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<mods:title id="25BC10A58DFDCE46DFF5BB5ECBF83C78">Phylogenetics, classification, and biogeography of the treefrogs (Amphibia: Anura: Arboranae)</mods:title>
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<mods:namePart id="97DF170021DB086122CA254DDDB7214D">Duellman, William E.</mods:namePart>
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<mods:namePart id="40234D3E1642AF4124701C93AEAF84F5">Marion, Angela B.</mods:namePart>
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<mods:namePart id="EAF87DB2DD3F03E37A76671C3E4CFA1D">Hedges, Blair</mods:namePart>
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<mods:title id="83CB445EEDF18AAFF16327AE613B3B57">Zootaxa</mods:title>
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<mods:date id="085E48818A83A2585EB776815B04CAA6">2016</mods:date>
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<treatment id="03EA87A5FF891209F3988C823020F674" ID-DOI="http://doi.org/10.5281/zenodo.5458536" ID-GBIF-Taxon="119611986" ID-Zenodo-Dep="5458536" LSID="urn:lsid:plazi:treatment:03EA87A5FF891209F3988C823020F674" httpUri="http://treatment.plazi.org/id/03EA87A5FF891209F3988C823020F674" lastPageNumber="33" pageId="32" pageNumber="33">
<subSubSection id="C3596538FF891209F3988C823208F008" pageId="32" pageNumber="33" type="nomenclature">
<paragraph id="8BFC36B3FF891209F3988C823288F2A8" blockId="32.[151,438,476,502]" box="[151,438,476,502]" pageId="32" pageNumber="33">
<heading id="D0B481DFFF891209F3988C823288F2A8" bold="true" box="[151,438,476,502]" fontSize="11" level="1" pageId="32" pageNumber="33" reason="1">
<emphasis id="B937EAA1FF891209F3988C823288F2A8" bold="true" box="[151,438,476,502]" pageId="32" pageNumber="33">
<taxonomicName id="4C434D30FF891209F3988C823210F2A8" ID-CoL="3FFK" box="[151,302,476,502]" class="Amphibia" family="Phyllomedusidae" genus="Callimedusa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="32" pageNumber="33" phylum="Chordata" rank="genus" status="gen. nov.">
<emphasis id="B937EAA1FF891209F3988C823210F2A8" bold="true" box="[151,302,476,502]" italics="true" pageId="32" pageNumber="33">Callimedusa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20457DAFF891209F23A8C823288F2A8" box="[309,438,476,502]" pageId="32" pageNumber="33" rank="genus">new genus</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BFC36B3FF891209F3988F7C3064F166" blockId="32.[151,858,546,568]" box="[151,858,546,568]" pageId="32" pageNumber="33">
<emphasis id="B937EAA1FF891209F3988F7C3220F166" box="[151,286,546,568]" italics="true" pageId="32" pageNumber="33">
<taxonomicName id="4C434D30FF891209F3988F7C3224F166" box="[151,282,546,568]" class="Amphibia" family="Phyllomedusidae" genus="Callimedusa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="32" pageNumber="33" phylum="Chordata" rank="genus" status="gen. nov.">Callimedusa</taxonomicName>
.
</emphasis>
<typeStatus id="54F88811FF891209F22C8F7D326BF166" box="[291,341,547,568]" pageId="32" pageNumber="33">Type</typeStatus>
species:
<taxonomicName id="4C434D30FF891209F2BD8F7C3068F166" authority="Duellman, 1973" authorityName="Duellman" authorityYear="1973" box="[434,854,546,568]" class="Amphibia" family="Hylidae" genus="Phyllomedusa" kingdom="Animalia" order="Anura" pageId="32" pageNumber="33" phylum="Chordata" rank="species" species="perinesos">
<emphasis id="B937EAA1FF891209F2BD8F7C3197F166" box="[434,681,546,568]" italics="true" pageId="32" pageNumber="33">Phyllomedusa perinesos</emphasis>
Duellman, 1973
</taxonomicName>
.
</paragraph>
<paragraph id="8BFC36B3FF891209F3988F3B30A7F1B4" blockId="32.[151,1436,613,1322]" pageId="32" pageNumber="33">
<emphasis id="B937EAA1FF891209F3988F3B3228F120" bold="true" box="[151,278,613,638]" pageId="32" pageNumber="33">Definition.</emphasis>
Small to medium-sized species (SVL of
<quantity id="4CBB9B56FF891209F0148F38304DF123" box="[795,883,614,638]" metricMagnitude="-2" metricUnit="m" metricValue="4.4" pageId="32" pageNumber="33" unit="mm" value="44.0">44 mm</quantity>
in
<taxonomicName id="4C434D30FF891209F0A98F3B37FAF120" box="[934,1220,613,638]" class="Amphibia" family="Phyllomedusidae" genus="Callimedusa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="32" pageNumber="33" phylum="Chordata" rank="species" species="atelopoides">
<emphasis id="B937EAA1FF891209F0A98F3B37FAF120" box="[934,1220,613,638]" italics="true" pageId="32" pageNumber="33">Callimedusa atelopoides</emphasis>
</taxonomicName>
to
<quantity id="4CBB9B56FF891209F7F88F3B3670F123" box="[1271,1358,613,638]" metricMagnitude="-2" metricUnit="m" metricValue="6.2" pageId="32" pageNumber="33" unit="mm" value="62.0">62 mm</quantity>
in
<taxonomicName id="4C434D30FF891209F68D8F383226F1FF" class="Amphibia" family="Phyllomedusidae" genus="Callimedusa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="32" pageNumber="33" phylum="Chordata" rank="species" species="tomopterna">
<emphasis id="B937EAA1FF891209F68D8F383226F1FF" italics="true" pageId="32" pageNumber="33">C. tomopterna</emphasis>
</taxonomicName>
); Toe I slightly longer than Toe II, not opposable; vomerine teeth present (except in
<taxonomicName id="4C434D30FF891209F7E88FD436B5F1FC" box="[1255,1419,649,674]" class="Amphibia" family="Phyllomedusidae" genus="Callimedusa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="32" pageNumber="33" phylum="Chordata" rank="species" species="atelopoides">
<emphasis id="B937EAA1FF891209F7E88FD436B5F1FC" box="[1255,1419,649,674]" italics="true" pageId="32" pageNumber="33">C. atelopoides</emphasis>
</taxonomicName>
); flash colors on flanks and hidden surfaces of thighs purple or black; palpebral membrane reticulated or not; tadpoles having moderately small oral discs directed anteroventrally.
</paragraph>
<paragraph id="8BFC36B3FF891209F3C88FAB3208F008" blockId="32.[151,1436,613,1322]" pageId="32" pageNumber="33">
<emphasis id="B937EAA1FF891209F3C88FAB3210F050" bold="true" box="[199,302,757,782]" pageId="32" pageNumber="33">Content.</emphasis>
Six species:
<taxonomicName id="4C434D30FF891209F2C78FAB375EF050" authority="Duellman, Cadle &amp; Cannatella" authorityName="Duellman, Cadle &amp; Cannatella" box="[456,1120,757,782]" class="Amphibia" family="Phyllomedusidae" genus="Callimedusa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="32" pageNumber="33" phylum="Chordata" rank="species" species="atelopoides">
<emphasis id="B937EAA1FF891209F2C78FAB31DEF050" box="[456,736,757,782]" italics="true" pageId="32" pageNumber="33">Callimedusa atelopoides</emphasis>
(Duellman, Cadle &amp; Cannatella)
</taxonomicName>
,
<taxonomicName id="4C434D30FF891209F7628FAB36A9F050" authority="Duellman &amp; Toft" authorityName="Duellman &amp; Toft" box="[1133,1431,757,782]" class="Amphibia" family="Phyllomedusidae" genus="Callimedusa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="32" pageNumber="33" phylum="Chordata" rank="species" species="baltea">
<emphasis id="B937EAA1FF891209F7628FAB378DF050" box="[1133,1203,757,782]" italics="true" pageId="32" pageNumber="33">baltea</emphasis>
(Duellman &amp; Toft)
</taxonomicName>
,
<taxonomicName id="4C434D30FF891209F3988E47329BF06C" authority="Cannatella" authorityName="Cannatella" box="[151,421,793,818]" class="Amphibia" family="Phyllomedusidae" genus="Callimedusa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="32" pageNumber="33" phylum="Chordata" rank="species" species="duellmani">
<emphasis id="B937EAA1FF891209F3988E473239F06C" box="[151,263,793,818]" italics="true" pageId="32" pageNumber="33">duellmani</emphasis>
(Cannatella)
</taxonomicName>
,
<taxonomicName id="4C434D30FF891209F2B48E4431DFF06C" authority="Cannatella" authorityName="Cannatella" box="[443,737,793,818]" class="Amphibia" family="Phyllomedusidae" genus="Callimedusa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="32" pageNumber="33" phylum="Chordata" rank="species" species="ecuatoriana">
<emphasis id="B937EAA1FF891209F2B48E44317CF06F" box="[443,578,794,817]" italics="true" pageId="32" pageNumber="33">ecuatoriana</emphasis>
(Cannatella)
</taxonomicName>
,
<taxonomicName id="4C434D30FF891209F1F98E4430C9F06C" authority="Duellman" authorityName="Duellman" box="[758,1015,793,818]" class="Amphibia" family="Hylidae" genus="Phyllomedusa" kingdom="Animalia" order="Anura" pageId="32" pageNumber="33" phylum="Chordata" rank="species" species="perinesos">
<emphasis id="B937EAA1FF891209F1F98E44305FF06F" box="[758,865,794,817]" italics="true" pageId="32" pageNumber="33">perinesos</emphasis>
(Duellman)
</taxonomicName>
, and
<taxonomicName id="4C434D30FF891209F7498E443614F06C" authority="Cope" authorityName="Cope" box="[1094,1322,794,818]" class="Amphibia" family="Phyllomedusidae" genus="Callimedusa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="32" pageNumber="33" phylum="Chordata" rank="species" species="tomopterna">
<emphasis id="B937EAA1FF891209F7498E4437F9F06F" box="[1094,1223,794,817]" italics="true" pageId="32" pageNumber="33">tomopterna</emphasis>
(Cope)
</taxonomicName>
; all new combinations.
</paragraph>
</subSubSection>
<subSubSection id="C3596538FF891209F3C88E3F3177F0C0" pageId="32" pageNumber="33" type="distribution">
<paragraph id="8BFC36B3FF891209F3C88E3F3177F0C0" blockId="32.[151,1436,613,1322]" pageId="32" pageNumber="33">
<emphasis id="B937EAA1FF891209F3C88E3F325EF024" bold="true" box="[199,352,865,890]" pageId="32" pageNumber="33">Distribution.</emphasis>
Amazonian slopes of the Andes from
<collectingCountry id="F3547623FF891209F0398E3F30ADF024" box="[822,915,865,890]" name="Ecuador" pageId="32" pageNumber="33">Ecuador</collectingCountry>
to central
<collectingCountry id="F3547623FF891209F7158E3C376FF027" box="[1050,1105,866,889]" name="Peru" pageId="32" pageNumber="33">Peru</collectingCountry>
; upper Amazon Basin from
<collectingCountry id="F3547623FF891209F3988EDB323BF0C0" box="[151,261,901,926]" name="Colombia" pageId="32" pageNumber="33">Colombia</collectingCountry>
to
<collectingCountry id="F3547623FF891209F2268EDB3241F0C0" box="[297,383,901,926]" name="Bolivia" pageId="32" pageNumber="33">Bolivia</collectingCountry>
; Guianan Region.
</paragraph>
</subSubSection>
<subSubSection id="C3596538FF891209F3C88EF7361FF0B8" pageId="32" pageNumber="33" type="etymology">
<paragraph id="8BFC36B3FF891209F3C88EF7361FF0B8" blockId="32.[151,1436,613,1322]" pageId="32" pageNumber="33">
<emphasis id="B937EAA1FF891209F3C88EF73270F09C" bold="true" box="[199,334,937,962]" pageId="32" pageNumber="33">Etymology.</emphasis>
The generic name is derived from the Greek
<emphasis id="B937EAA1FF891209F0488EF730BCF09C" box="[839,898,937,962]" italics="true" pageId="32" pageNumber="33">kalos</emphasis>
meaning beautiful and the Greek
<emphasis id="B937EAA1FF891209F7F38EF73655F09C" box="[1276,1387,937,962]" italics="true" pageId="32" pageNumber="33">Medousa.</emphasis>
The name alludes to the beautiful coloration of the flanks of members of this genus. The gender is feminine.
</paragraph>
</subSubSection>
<subSubSection id="C3596538FF891209F3C88EAF3020F674" pageId="32" pageNumber="33" type="discussion">
<paragraph id="8BFC36B3FF891209F3C88EAF3020F674" blockId="32.[151,1436,613,1322]" pageId="32" pageNumber="33">
<emphasis id="B937EAA1FF891209F3C88EAF3205F754" bold="true" box="[199,315,1009,1034]" pageId="32" pageNumber="33">Remarks.</emphasis>
The monophyly of this genus is rather well supported (78%), but that of the “
<taxonomicName id="4C434D30FF891209F7F28EAF323DF773" class="Amphibia" family="Hylidae" genus="Phyllomedusa" kingdom="Animalia" order="Anura" pageId="32" pageNumber="33" phylum="Chordata" rank="species" species="perinesos">
<emphasis id="B937EAA1FF891209F7F28EAF323DF773" italics="true" pageId="32" pageNumber="33">Phyllomedusa perinesos</emphasis>
</taxonomicName>
Group” of
<bibRefCitation id="EFD24B42FF891209F28A894B3171F770" author="Cannatella" box="[389,591,1045,1070]" pageId="32" pageNumber="33" refString="Cannatella, D. C. (1982) Leaf-frogs of the Phyllomedusa perinesos Group (Anura: Hylidae). Copeia, 1982, 501 - 513. http: // dx. doi. org / 10.2307 / 1444649" type="journal article" year="1982">Cannatella (1982)</bibRefCitation>
within
<taxonomicName id="4C434D30FF891209F1A9894B300AF770" box="[678,820,1045,1070]" class="Amphibia" family="Phyllomedusidae" genus="Callimedusa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="32" pageNumber="33" phylum="Chordata" rank="genus" status="gen. nov.">
<emphasis id="B937EAA1FF891209F1A9894B300AF770" box="[678,820,1045,1070]" italics="true" pageId="32" pageNumber="33">Callimedusa</emphasis>
</taxonomicName>
is only moderately supported (66%). The four species having allopatric ranges on the Amazon slopes of the Andes in
<collectingCountry id="F3547623FF891209F0BD89673731F70C" box="[946,1039,1081,1106]" name="Ecuador" pageId="32" pageNumber="33">Ecuador</collectingCountry>
and
<collectingCountry id="F3547623FF891209F759896437B7F70F" box="[1110,1161,1082,1105]" name="Peru" pageId="32" pageNumber="33">Peru</collectingCountry>
(
<emphasis id="B937EAA1FF891209F7AE8964321BF72B" italics="true" pageId="32" pageNumber="33">
<taxonomicName id="4C434D30FF891209F7AE8964362FF70C" box="[1185,1297,1081,1106]" class="Amphibia" family="Phyllomedusidae" genus="Callimedusa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="32" pageNumber="33" phylum="Chordata" rank="species" species="baltea">C. baltea</taxonomicName>
,
<taxonomicName id="4C434D30FF891209F62A896736A6F70C" box="[1317,1432,1081,1106]" class="Amphibia" family="Phyllomedusidae" genus="Callimedusa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="32" pageNumber="33" phylum="Chordata" rank="species" species="duellmani">duellmani</taxonomicName>
,
<taxonomicName id="4C434D30FF891209F3988900321FF72B" box="[151,289,1118,1141]" class="Amphibia" family="Phyllomedusidae" genus="Callimedusa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="32" pageNumber="33" phylum="Chordata" rank="species" species="ecuatoriana">ecuatoriana</taxonomicName>
,
</emphasis>
and
<taxonomicName id="4C434D30FF891209F255890032FBF72B" box="[346,453,1118,1141]" class="Amphibia" family="Hylidae" genus="Phyllomedusa" kingdom="Animalia" order="Anura" pageId="32" pageNumber="33" phylum="Chordata" rank="species" species="perinesos">
<emphasis id="B937EAA1FF891209F255890032FBF72B" box="[346,453,1118,1141]" italics="true" pageId="32" pageNumber="33">perinesos</emphasis>
</taxonomicName>
) closely resemble one another (
<figureCitation id="13782A36FF891209F026890330BCF728" box="[809,898,1117,1142]" captionStart="FIGURE 14" captionStartId="33.[151,250,1707,1729]" captionTargetBox="[153,1436,776,1683]" captionTargetId="figure@33.[151,1436,776,1686]" captionTargetPageId="33" captionText="FIGURE 14. Species of Callimedusa A. C. perinesos, KU 164450, Río Salado, 1 km upstream from Río Coca, 1410 m, Napo, Ecuador. B. C. duellmani, KU 181813, 8 km north-northeast Balzapata, 1860 m, Amazonas, Peru. C. C. tomopterna, KU 220332, Río Sucusari, Loreto, 210 m, Peru. D. C. atelopoides, KU 204764, Cusco Amazónico, 15 km east Puerto Maldonado, 200 m, Madre de Dios, Peru. All by W. E. Duellman." httpUri="https://zenodo.org/record/265826/files/figure.png" pageId="32" pageNumber="33">Figs. 14</figureCitation>
A and B). In our analysis, the sister species is
<taxonomicName id="4C434D30FF891209F68D89003226F7C4" class="Amphibia" family="Phyllomedusidae" genus="Callimedusa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="32" pageNumber="33" phylum="Chordata" rank="species" species="atelopoides">
<emphasis id="B937EAA1FF891209F68D89003226F7C4" italics="true" pageId="32" pageNumber="33">C. atelopoides</emphasis>
</taxonomicName>
(
<figureCitation id="13782A36FF891209F23D89DF32B1F7C4" box="[306,399,1153,1178]" captionStart="FIGURE 14" captionStartId="33.[151,250,1707,1729]" captionTargetBox="[153,1436,776,1683]" captionTargetId="figure@33.[151,1436,776,1686]" captionTargetPageId="33" captionText="FIGURE 14. Species of Callimedusa A. C. perinesos, KU 164450, Río Salado, 1 km upstream from Río Coca, 1410 m, Napo, Ecuador. B. C. duellmani, KU 181813, 8 km north-northeast Balzapata, 1860 m, Amazonas, Peru. C. C. tomopterna, KU 220332, Río Sucusari, Loreto, 210 m, Peru. D. C. atelopoides, KU 204764, Cusco Amazónico, 15 km east Puerto Maldonado, 200 m, Madre de Dios, Peru. All by W. E. Duellman." httpUri="https://zenodo.org/record/265826/files/figure.png" pageId="32" pageNumber="33">Fig. 14</figureCitation>
D), an inhabitant of the upper Amazon Basin. This species differs from all other phyllomedusids by being small, brown, terrestrial; furthermore, it is unlike other
<taxonomicName id="4C434D30FF891209F72989FB378AF7E0" box="[1062,1204,1189,1214]" class="Amphibia" family="Phyllomedusidae" genus="Callimedusa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="32" pageNumber="33" phylum="Chordata" rank="genus" status="gen. nov.">
<emphasis id="B937EAA1FF891209F72989FB378AF7E0" box="[1062,1204,1189,1214]" italics="true" pageId="32" pageNumber="33">Callimedusa</emphasis>
</taxonomicName>
in lacking vomerine teeth and clavicles in the pectoral girdle (
<bibRefCitation id="EFD24B42FF891209F17C89973055F7BC" author="Duellman" box="[627,875,1225,1250]" pageId="32" pageNumber="33" refString="Duellman, W. E., Cadle, J. E. &amp; Cannatella, D. C. (1988) A new species of terrestrial Phyllomedusa (Anura: Hylidae) from southern Peru. Herpetologica, 44, 91 - 95." type="journal article" year="1988">
Duellman
<emphasis id="B937EAA1FF891209F1E389943018F7BC" box="[748,806,1225,1250]" italics="true" pageId="32" pageNumber="33">et al.</emphasis>
1988
</bibRefCitation>
). The analysis places
<taxonomicName id="4C434D30FF891209F76789943635F7BF" box="[1128,1291,1226,1249]" class="Amphibia" family="Phyllomedusidae" genus="Callimedusa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="32" pageNumber="33" phylum="Chordata" rank="species" species="tomopterna">
<emphasis id="B937EAA1FF891209F76789943635F7BF" box="[1128,1291,1226,1249]" italics="true" pageId="32" pageNumber="33">C. tomopterna</emphasis>
</taxonomicName>
as the sister species to the remaining members of the genus (
<figureCitation id="13782A36FF891209F1B189B3302FF658" box="[702,785,1261,1286]" captionStart="FIGURE 14" captionStartId="33.[151,250,1707,1729]" captionTargetBox="[153,1436,776,1683]" captionTargetId="figure@33.[151,1436,776,1686]" captionTargetPageId="33" captionText="FIGURE 14. Species of Callimedusa A. C. perinesos, KU 164450, Río Salado, 1 km upstream from Río Coca, 1410 m, Napo, Ecuador. B. C. duellmani, KU 181813, 8 km north-northeast Balzapata, 1860 m, Amazonas, Peru. C. C. tomopterna, KU 220332, Río Sucusari, Loreto, 210 m, Peru. D. C. atelopoides, KU 204764, Cusco Amazónico, 15 km east Puerto Maldonado, 200 m, Madre de Dios, Peru. All by W. E. Duellman." httpUri="https://zenodo.org/record/265826/files/figure.png" pageId="32" pageNumber="33">Fig. 14</figureCitation>
C). This colorful frog that ranges throughout much of the Amazon Basin is widely allopatric to the Andean species.
</paragraph>
</subSubSection>
</treatment>
</document>