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<mods:title id="355FF8E25847B8E93A6FA6AFA05CCC6D">Trachyibana sarawakiensis gen. nov., sp. nov., a remarkable new genus and species from Malaysia (Acari: Uropodina: Trachyuropodidae)</mods:title>
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<subSubSection id="C359653C6066FFA453C7B66571614B60" pageId="5" pageNumber="277" type="nomenclature">
<paragraph id="8BFC36B76066FFA453C7B66573F44C21" blockId="5.[151,1004,800,826]" box="[151,1004,800,826]" pageId="5" pageNumber="277">
<heading id="D0B481DB6066FFA453C7B66573F44C21" bold="true" box="[151,1004,800,826]" fontSize="11" level="1" pageId="5" pageNumber="277" reason="1">
<emphasis id="B937EAA56066FFA453C7B66573F44C21" bold="true" box="[151,1004,800,826]" pageId="5" pageNumber="277">
Key to the genera of the family
<taxonomicName id="4C434D346066FFA45170B66573104C21" ID-CoL="HCL" ID-ENA="118557" box="[544,776,800,826]" class="Arachnida" family="Trachyuropodidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="5" pageNumber="277" phylum="Arthropoda" rank="family">Trachyuropodidae</taxonomicName>
(based on adults).
</emphasis>
</heading>
</paragraph>
<paragraph id="8BFC36B76066FFA453C7B62371614B60" blockId="5.[151,1436,870,1147]" pageId="5" pageNumber="277">
The key follows Hirschmanns (1979) generic classification, except that
<emphasis id="B937EAA56066FFA45098B623744F4C64" box="[968,1111,870,895]" italics="true" pageId="5" pageNumber="277">Foliatrachys</emphasis>
<bibRefCitation id="EFD24B466066FFA4570FB62375364C64" author="Hirschmann" box="[1119,1326,870,895]" pageId="5" pageNumber="277" refString="Hirschmann, W. (1979) Stadiensystematik der Parasitiformes Teil 1. Stadienfamilien und Stadiengattungen der Atrichopygidiina, erstellt im Vergleich zum Gangsystem Hirschmann 1979. Acarologie. Schriftenreihe fur Vergleichende Milbenkunde, 26, 57 - 70." type="journal article" year="1979">Hirschmann, 1979</bibRefCitation>
is known only from the immature stages.
<bibRefCitation id="EFD24B466066FFA45142B6CE72B84CB8" author="Karg" box="[530,672,907,931]" pageId="5" pageNumber="277" refString="Karg, W. (1989) Uropodina Kramer, Schildkrotenmilben. Acari (Acarina), Milben, Unterordnung Parasitiformes. Die Tierwelt Deutschlands, 67, 1 - 203." type="journal article" year="1989">Karg (1989)</bibRefCitation>
and
<bibRefCitation id="EFD24B466066FFA4518FB6CE73984CB8" author="Masan" box="[735,896,907,931]" pageId="5" pageNumber="277" refString="Masan, P. (2001) Roztoe kohorty Uropodina (Acarina, Mesostigmata) Slovenska (Mites of the cohort Uropodina (Acarina, Mesostigmata) in Slovakia. Annotationes Zoologicae et Botanicae, 22, 1 - 320." type="journal article" year="2001">Mašán (2001)</bibRefCitation>
included only two genera (
<emphasis id="B937EAA56066FFA4579AB6CF75704CB8" box="[1226,1384,906,931]" italics="true" pageId="5" pageNumber="277">Trachyuropda</emphasis>
and
<taxonomicName id="4C434D346066FFA453C7B6EB70FB4CDC" box="[151,227,942,967]" class="Arachnida" family="Trachyuropodidae" genus="Oplitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="5" pageNumber="277" phylum="Arthropoda" rank="genus">
<emphasis id="B937EAA56066FFA453C7B6EB70FB4CDC" box="[151,227,942,967]" italics="true" pageId="5" pageNumber="277">Oplitis</emphasis>
</taxonomicName>
) in this family. More recently
<bibRefCitation id="EFD24B466066FFA45110B6EB73274CDC" box="[576,831,942,967]" pageId="5" pageNumber="277" refString="Lindquist, E. E., Krantz, G. W. &amp; Walter, D. E. (2009) Order Mesostigmata. In: Krantz, G. W. &amp; Walter, D. E. (Eds.), A Manual of Acarology. Third edition. Texas Tech University Press, Lubbock, USA, pp. 124 - 232." type="book chapter">
Lindquist
<emphasis id="B937EAA56066FFA451E3B6EA72FC4CDC" box="[691,740,942,967]" italics="true" pageId="5" pageNumber="277">et al</emphasis>
. (2009)
</bibRefCitation>
and
<bibRefCitation id="EFD24B466066FFA45027B6EB74754CDC" box="[887,1133,942,967]" pageId="5" pageNumber="277" refString="Beaulieu, F., Dowling, A. P. G., Klompen, H., de Moraes, G. J. &amp; Walter, D. E. (2011) Superorder Parasitiformes Reuter, 1909. In: Zhang, Z. - Q. (Ed.), Animal biodiversity: an outline of higher-level classification and survey of taxonomic richness. Zootaxa, 3148, 123 - 128." type="journal article">
Beaulieu
<emphasis id="B937EAA56066FFA450B3B6EA740C4CDC" box="[995,1044,942,967]" italics="true" pageId="5" pageNumber="277">et al</emphasis>
. (2011)
</bibRefCitation>
separated
<taxonomicName id="4C434D346066FFA457B7B6EB752B4CDC" box="[1255,1331,942,967]" class="Arachnida" family="Trachyuropodidae" genus="Oplitis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="5" pageNumber="277" phylum="Arthropoda" rank="genus">
<emphasis id="B937EAA56066FFA457B7B6EB752B4CDC" box="[1255,1331,942,967]" italics="true" pageId="5" pageNumber="277">Oplitis</emphasis>
</taxonomicName>
from the
<taxonomicName id="4C434D346066FFA453C7B697717B4CF0" box="[151,355,978,1003]" class="Arachnida" family="Trachyuropodidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="5" pageNumber="277" phylum="Arthropoda" rank="family">Trachyuropodidae</taxonomicName>
into the family
<taxonomicName id="4C434D346066FFA45172B69772954CF0" box="[546,653,978,1003]" class="Arachnida" family="Oplitidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="5" pageNumber="277" phylum="Arthropoda" rank="family">Oplitidae</taxonomicName>
, and that decision is accepted here. The classification of the family
<taxonomicName id="4C434D346066FFA453C7B6B3717B4B14" box="[151,355,1014,1039]" class="Arachnida" family="Trachyuropodidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="5" pageNumber="277" phylum="Arthropoda" rank="family">Trachyuropodidae</taxonomicName>
is more complex than was suggested by the European studies, and several endemic genera exist in tropical areas. The present paper therefore follows Hirschmanns (1979) generic classification, which gives a more detailed subdivision of the family
<taxonomicName id="4C434D346066FFA4510FB17B73364B4C" box="[607,814,1086,1111]" class="Arachnida" family="Trachyuropodidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="5" pageNumber="277" phylum="Arthropoda" rank="family">Trachyuropodidae</taxonomicName>
, despite the fact that it was not used by
<bibRefCitation id="EFD24B466066FFA457ABB17B716D4B60" author="Wisniewski" pageId="5" pageNumber="277" refString="Wisniewski, J. &amp; Hirschmann, W. (1993) Gangsystematik der Parasitiformes Teil 548. Katalog der Ganggattungen, Untergattungen, Gruppen und Arten der Uropodiden der Erde. Acarologie. Schriftenreihe fur Vergleichende Milbenkunde, 40, 1 - 220." type="journal article" year="1993">Wiśniewski &amp; Hirschmann (1993)</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C359653C6066FFA753C7B1EC75844FD1" lastPageId="6" lastPageNumber="278" pageId="5" pageNumber="277" type="key">
<keyStep id="30B72E126066FFA453C7B1EC75834BC6" pageId="5" pageNumber="277">
<paragraph id="8BFC36B76066FFA453C7B1EC75834BA4" blockId="5.[151,1436,1193,2037]" box="[151,1435,1193,1215]" pageId="5" pageNumber="277">
<keyLead id="30B295826066FFA453C7B1EC75834BA4" box="[151,1435,1193,1215]" pageId="5" pageNumber="277">1. Opisthogastric region of ventral idiosoma with a pair of deep furrows............................................ 2</keyLead>
</paragraph>
<paragraph id="8BFC36B76066FFA453C7B18275834BC6" blockId="5.[151,1436,1193,2037]" box="[151,1435,1223,1245]" pageId="5" pageNumber="277">
<keyLead id="30B295826066FFA453C7B18275834BC6" box="[151,1435,1223,1245]" pageId="5" pageNumber="277">- Opisthogastric region of ventral idiosoma without deep furrows................................................. 3</keyLead>
</paragraph>
</keyStep>
<keyStep id="30B72E126066FFA453C7B1A175844A03" pageId="5" pageNumber="277">
<paragraph id="8BFC36B76066FFA453C7B1A175844BE1" blockId="5.[151,1436,1193,2037]" box="[151,1436,1252,1274]" pageId="5" pageNumber="277">
<keyLead id="30B295826066FFA453C7B1A175844BE1" box="[151,1436,1252,1274]" pageId="5" pageNumber="277">
2. Idiosoma subtriangular or pentagonal, anterior area of marginal shield broad.................
<taxonomicName id="4C434D346066FFA457C7B1A075844BE1" authority="Berlese, 1903" authorityName="Berlese" authorityYear="1903" box="[1175,1436,1252,1274]" class="Arachnida" family="Trachyuropodidae" genus="Leonardiella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="5" pageNumber="277" phylum="Arthropoda" rank="genus">
<emphasis id="B937EAA56066FFA457C7B1A0750A4BE1" box="[1175,1298,1253,1274]" italics="true" pageId="5" pageNumber="277">Leonardiella</emphasis>
<bibRefCitation id="EFD24B466066FFA45648B1A175844BE1" author="Berlese" box="[1304,1436,1252,1274]" pageId="5" pageNumber="277" refString="Berlese, A. (1903) Acari Mirmecofili. Redia, 1, 299 - 453." type="journal article" year="1903">Berlese, 1903</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BFC36B76066FFA453C7B04775844A03" blockId="5.[151,1436,1193,2037]" box="[151,1436,1281,1304]" pageId="5" pageNumber="277">
<keyLead id="30B295826066FFA453C7B04775844A03" box="[151,1436,1281,1304]" pageId="5" pageNumber="277">
- Idiosoma oval, lemon-shaped, anterior area of marginal shield narrow..........................
<taxonomicName id="4C434D346066FFA45792B04775234A0C" box="[1218,1339,1282,1303]" class="Arachnida" family="Trachyuropodidae" genus="Trachyibana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="5" pageNumber="277" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B937EAA56066FFA45792B04775234A0C" box="[1218,1339,1282,1303]" italics="true" pageId="5" pageNumber="277">Trachyibana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20457DE6066FFA45612B04775844A03" box="[1346,1436,1282,1304]" pageId="5" pageNumber="277" rank="genus">
<emphasis id="B937EAA56066FFA45612B04775844A03" bold="true" box="[1346,1436,1282,1304]" pageId="5" pageNumber="277">gen. nov.</emphasis>
</taxonomicNameLabel>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30B72E126066FFA453C7B05A75834A49" pageId="5" pageNumber="277">
<paragraph id="8BFC36B76066FFA453C7B05A75834A2E" blockId="5.[151,1436,1193,2037]" box="[151,1435,1311,1333]" pageId="5" pageNumber="277">
<keyLead id="30B295826066FFA453C7B05A75834A2E" box="[151,1435,1311,1333]" pageId="5" pageNumber="277">3. Dorsal shield with strongly sclerotised lines, rings and other structures........................................... 5</keyLead>
</paragraph>
<paragraph id="8BFC36B76066FFA453C7B07875834A49" blockId="5.[151,1436,1193,2037]" box="[151,1435,1340,1362]" pageId="5" pageNumber="277">
<keyLead id="30B295826066FFA453C7B07875834A49" box="[151,1435,1340,1362]" pageId="5" pageNumber="277">- Dorsal shield without sclerotised lines, rings and other structures................................................ 4</keyLead>
</paragraph>
</keyStep>
<keyStep id="30B72E126066FFA453C7B01C75844A96" pageId="5" pageNumber="277">
<paragraph id="8BFC36B76066FFA453C7B01C75844A74" blockId="5.[151,1436,1193,2037]" box="[151,1436,1369,1391]" pageId="5" pageNumber="277">
<keyLead id="30B295826066FFA453C7B01C75844A74" box="[151,1436,1369,1391]" pageId="5" pageNumber="277">
4. Dorsal shield covered by oval pits..............................................
<emphasis id="B937EAA56066FFA45709B01F74FB4A74" box="[1113,1251,1370,1391]" italics="true" pageId="5" pageNumber="277">Magnatrachys</emphasis>
<bibRefCitation id="EFD24B466066FFA457BBB01C75844A74" author="Hirschmann" box="[1259,1436,1369,1391]" pageId="5" pageNumber="277" refString="Hirschmann, W. (1979) Stadiensystematik der Parasitiformes Teil 1. Stadienfamilien und Stadiengattungen der Atrichopygidiina, erstellt im Vergleich zum Gangsystem Hirschmann 1979. Acarologie. Schriftenreihe fur Vergleichende Milbenkunde, 26, 57 - 70." type="journal article" year="1979">Hirschmann, 1979</bibRefCitation>
</keyLead>
</paragraph>
<paragraph id="8BFC36B76066FFA453C7B03275844A96" blockId="5.[151,1436,1193,2037]" box="[151,1436,1399,1421]" pageId="5" pageNumber="277">
<keyLead id="30B295826066FFA453C7B03275844A96" box="[151,1436,1399,1421]" pageId="5" pageNumber="277">
- Dorsal shield covered by reticulate sculptural pattern.............................
<emphasis id="B937EAA56066FFA45714B03274FD4A97" box="[1092,1253,1399,1420]" italics="true" pageId="5" pageNumber="277">Lindquisttrachys</emphasis>
<bibRefCitation id="EFD24B466066FFA457BBB03275844A96" author="Hirschmann" box="[1259,1436,1399,1421]" pageId="5" pageNumber="277" refString="Hirschmann, W. (1979) Stadiensystematik der Parasitiformes Teil 1. Stadienfamilien und Stadiengattungen der Atrichopygidiina, erstellt im Vergleich zum Gangsystem Hirschmann 1979. Acarologie. Schriftenreihe fur Vergleichende Milbenkunde, 26, 57 - 70." type="journal article" year="1979">Hirschmann, 1979</bibRefCitation>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30B72E126066FFA453C7B0D175844ADC" pageId="5" pageNumber="277">
<paragraph id="8BFC36B76066FFA453C7B0D175834AB1" blockId="5.[151,1436,1193,2037]" box="[151,1435,1428,1450]" pageId="5" pageNumber="277">
<keyLead id="30B295826066FFA453C7B0D175834AB1" box="[151,1435,1428,1450]" pageId="5" pageNumber="277">5. Dorsal shield with line-like strongly sclerotised structures..................................................... 6</keyLead>
</paragraph>
<paragraph id="8BFC36B76066FFA453C7B0F775844ADC" blockId="5.[151,1436,1193,2037]" box="[151,1436,1457,1479]" pageId="5" pageNumber="277">
<keyLead id="30B295826066FFA453C7B0F775844ADC" box="[151,1436,1457,1479]" pageId="5" pageNumber="277">- Dorsal shield with circular, semi-circular or subtriangular strongly sclerotised structures........................... 10</keyLead>
</paragraph>
</keyStep>
<keyStep id="30B72E126066FFA453C7B08A75834919" pageId="5" pageNumber="277">
<paragraph id="8BFC36B76066FFA453C7B08A75834AFF" blockId="5.[151,1436,1193,2037]" box="[151,1435,1487,1509]" pageId="5" pageNumber="277">
<keyLead id="30B295826066FFA453C7B08A75834AFF" box="[151,1435,1487,1509]" pageId="5" pageNumber="277">6. Strongly sclerotised line present, parallel with the axis of body.................................................. 7</keyLead>
</paragraph>
<paragraph id="8BFC36B76066FFA453C7B0A875834919" blockId="5.[151,1436,1193,2037]" box="[151,1435,1516,1538]" pageId="5" pageNumber="277">
<keyLead id="30B295826066FFA453C7B0A875834919" box="[151,1435,1516,1538]" pageId="5" pageNumber="277">- Strongly sclerotised line not parallel with the axis of body.................................................... 8</keyLead>
</paragraph>
</keyStep>
<keyStep id="30B72E126066FFA453C7B34F75844926" pageId="5" pageNumber="277">
<paragraph id="8BFC36B76066FFA453C7B34F75844904" blockId="5.[151,1436,1193,2037]" box="[151,1436,1545,1567]" pageId="5" pageNumber="277">
<keyLead id="30B295826066FFA453C7B34F75844904" box="[151,1436,1545,1567]" pageId="5" pageNumber="277">
7. Strongly sclerotised line divided into two parts....................................
<emphasis id="B937EAA56066FFA45732B34F74C34904" box="[1122,1243,1546,1567]" italics="true" pageId="5" pageNumber="277">Castritrachy</emphasis>
s
<bibRefCitation id="EFD24B466066FFA457BBB34C75844904" author="Hirschmann" box="[1259,1436,1545,1567]" pageId="5" pageNumber="277" refString="Hirschmann, W. (1979) Stadiensystematik der Parasitiformes Teil 1. Stadienfamilien und Stadiengattungen der Atrichopygidiina, erstellt im Vergleich zum Gangsystem Hirschmann 1979. Acarologie. Schriftenreihe fur Vergleichende Milbenkunde, 26, 57 - 70." type="journal article" year="1979">Hirschmann, 1979</bibRefCitation>
</keyLead>
</paragraph>
<paragraph id="8BFC36B76066FFA453C7B36275844926" blockId="5.[151,1436,1193,2037]" box="[151,1436,1575,1597]" pageId="5" pageNumber="277">
<keyLead id="30B295826066FFA453C7B36275844926" box="[151,1436,1575,1597]" pageId="5" pageNumber="277">
- Strongly sclerotised line divided into three parts..................................
<emphasis id="B937EAA56066FFA45702B36274FC4927" box="[1106,1252,1575,1596]" italics="true" pageId="5" pageNumber="277">Trogulotrachys</emphasis>
<bibRefCitation id="EFD24B466066FFA457BBB36275844926" author="Hirschmann" box="[1259,1436,1575,1597]" pageId="5" pageNumber="277" refString="Hirschmann, W. (1979) Stadiensystematik der Parasitiformes Teil 1. Stadienfamilien und Stadiengattungen der Atrichopygidiina, erstellt im Vergleich zum Gangsystem Hirschmann 1979. Acarologie. Schriftenreihe fur Vergleichende Milbenkunde, 26, 57 - 70." type="journal article" year="1979">Hirschmann, 1979</bibRefCitation>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30B72E126066FFA453C7B3017583496C" pageId="5" pageNumber="277">
<paragraph id="8BFC36B76066FFA453C7B30175844941" blockId="5.[151,1436,1193,2037]" box="[151,1436,1604,1626]" pageId="5" pageNumber="277">
<keyLead id="30B295826066FFA453C7B30175844941" box="[151,1436,1604,1626]" pageId="5" pageNumber="277">
8. Dorsal shield with deep horizontal furrow covered by two sclerotised lines.............
<emphasis id="B937EAA56066FFA45702B30074FC4941" box="[1106,1252,1605,1626]" italics="true" pageId="5" pageNumber="277">Bostocktrachys</emphasis>
<bibRefCitation id="EFD24B466066FFA457BBB30175844941" author="Hirschmann" box="[1259,1436,1604,1626]" pageId="5" pageNumber="277" refString="Hirschmann, W. (1979) Stadiensystematik der Parasitiformes Teil 1. Stadienfamilien und Stadiengattungen der Atrichopygidiina, erstellt im Vergleich zum Gangsystem Hirschmann 1979. Acarologie. Schriftenreihe fur Vergleichende Milbenkunde, 26, 57 - 70." type="journal article" year="1979">Hirschmann, 1979</bibRefCitation>
</keyLead>
</paragraph>
<paragraph id="8BFC36B76066FFA453C7B3277583496C" blockId="5.[151,1436,1193,2037]" box="[151,1435,1633,1655]" pageId="5" pageNumber="277">
<keyLead id="30B295826066FFA453C7B3277583496C" box="[151,1435,1633,1655]" pageId="5" pageNumber="277">- Deep horizontal furrow absent........................................................................... 9</keyLead>
</paragraph>
</keyStep>
<keyStep id="30B72E126066FFA453C7B33A758449A9" pageId="5" pageNumber="277">
<paragraph id="8BFC36B76066FFA453C7B33A7584498E" blockId="5.[151,1436,1193,2037]" box="[151,1436,1663,1685]" pageId="5" pageNumber="277">
<keyLead id="30B295826066FFA453C7B33A7584498E" box="[151,1436,1663,1685]" pageId="5" pageNumber="277">
9. Numerous, narrow sclerotised horizontal lines present..................................
<taxonomicName id="4C434D346066FFA457C4B33A7584498E" authority="Berlese, 1903" authorityName="Berlese" authorityYear="1903" box="[1172,1436,1663,1685]" class="Arachnida" family="Trachyuropodidae" genus="Urotrachytes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="5" pageNumber="277" phylum="Arthropoda" rank="genus">
<emphasis id="B937EAA56066FFA457C4B33A750A498F" box="[1172,1298,1663,1684]" italics="true" pageId="5" pageNumber="277">Urotrachytes</emphasis>
<bibRefCitation id="EFD24B466066FFA45648B33A7584498E" author="Berlese" box="[1304,1436,1663,1685]" pageId="5" pageNumber="277" refString="Berlese, A. (1903) Acari Mirmecofili. Redia, 1, 299 - 453." type="journal article" year="1903">Berlese, 1903</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BFC36B76066FFA453C7B3D8758449A9" blockId="5.[151,1436,1193,2037]" box="[151,1436,1692,1714]" pageId="5" pageNumber="277">
<keyLead id="30B295826066FFA453C7B3D8758449A9" box="[151,1436,1692,1714]" pageId="5" pageNumber="277">
- Numerous, narrow sclerotised horizontal lines absent..................................
<taxonomicName id="4C434D346066FFA4572DB3D8758449A9" authority="Berlese, 1903" authorityName="Berlese" authorityYear="1903" box="[1149,1436,1692,1714]" class="Arachnida" family="Trachyuropodidae" genus="Cephalojanetia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="5" pageNumber="277" phylum="Arthropoda" rank="genus">
<emphasis id="B937EAA56066FFA4572DB3D8750949A9" box="[1149,1297,1693,1714]" italics="true" pageId="5" pageNumber="277">Cephalojanetia</emphasis>
<bibRefCitation id="EFD24B466066FFA45648B3D9758449A9" author="Berlese" box="[1304,1436,1692,1714]" pageId="5" pageNumber="277" refString="Berlese, A. (1903) Acari Mirmecofili. Redia, 1, 299 - 453." type="journal article" year="1903">Berlese, 1903</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30B72E126066FFA453C7B3FC758349F6" pageId="5" pageNumber="277">
<paragraph id="8BFC36B76066FFA453C7B3FC758349D4" blockId="5.[151,1436,1193,2037]" box="[151,1435,1721,1743]" pageId="5" pageNumber="277">
<keyLead id="30B295826066FFA453C7B3FC758349D4" box="[151,1435,1721,1743]" pageId="5" pageNumber="277">10. Dorsal shield with one pair of strongly sclerotised structures................................................. 13</keyLead>
</paragraph>
<paragraph id="8BFC36B76066FFA453C7B392758349F6" blockId="5.[151,1436,1193,2037]" box="[151,1435,1751,1773]" pageId="5" pageNumber="277">
<keyLead id="30B295826066FFA453C7B392758349F6" box="[151,1435,1751,1773]" pageId="5" pageNumber="277">- Dorsal shield with more than one pair of strongly sclerotised structures......................................... 11</keyLead>
</paragraph>
</keyStep>
<keyStep id="30B72E126066FFA453C7B3B17583483C" pageId="5" pageNumber="277">
<paragraph id="8BFC36B76066FFA453C7B3B175844811" blockId="5.[151,1436,1193,2037]" box="[151,1436,1780,1802]" pageId="5" pageNumber="277">
<keyLead id="30B295826066FFA453C7B3B175844811" box="[151,1436,1780,1802]" pageId="5" pageNumber="277">
11. Dorsal shield with two pairs of subtriangular strongly sclerotised structures............
<emphasis id="B937EAA56066FFA45715B3B074FD4811" box="[1093,1253,1781,1802]" italics="true" pageId="5" pageNumber="277">Excavatatrachys</emphasis>
<bibRefCitation id="EFD24B466066FFA457BBB3B175844811" author="Hirschmann" box="[1259,1436,1780,1802]" pageId="5" pageNumber="277" refString="Hirschmann, W. (1979) Stadiensystematik der Parasitiformes Teil 1. Stadienfamilien und Stadiengattungen der Atrichopygidiina, erstellt im Vergleich zum Gangsystem Hirschmann 1979. Acarologie. Schriftenreihe fur Vergleichende Milbenkunde, 26, 57 - 70." type="journal article" year="1979">Hirschmann, 1979</bibRefCitation>
</keyLead>
</paragraph>
<paragraph id="8BFC36B76066FFA453C7B2577583483C" blockId="5.[151,1436,1193,2037]" box="[151,1435,1809,1831]" pageId="5" pageNumber="277">
<keyLead id="30B295826066FFA453C7B2577583483C" box="[151,1435,1809,1831]" pageId="5" pageNumber="277">- Dorsal shield with more than two pairs of semicircular strongly sclerotised structures.............................. 12</keyLead>
</paragraph>
</keyStep>
<keyStep id="30B72E126066FFA453C7B26A75844864" pageId="5" pageNumber="277">
<paragraph id="8BFC36B76066FFA453C7B26A7584485E" blockId="5.[151,1436,1193,2037]" box="[151,1436,1839,1861]" pageId="5" pageNumber="277">
<keyLead id="30B295826066FFA453C7B26A7584485E" box="[151,1436,1839,1861]" pageId="5" pageNumber="277">
12. Dorsal shield with three pairs of semicircular strongly sclerotised structures...................
<taxonomicName id="4C434D346066FFA457FAB26A7584485E" authority="Berlese, 1903" authorityName="Berlese" authorityYear="1903" box="[1194,1436,1839,1861]" class="Arachnida" family="Trachyuropodidae" genus="Urojanetia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="5" pageNumber="277" phylum="Arthropoda" rank="genus">
<emphasis id="B937EAA56066FFA457FAB26A750A485F" box="[1194,1298,1839,1860]" italics="true" pageId="5" pageNumber="277">Urojanetia</emphasis>
<bibRefCitation id="EFD24B466066FFA45648B26A7584485E" author="Berlese" box="[1304,1436,1839,1861]" pageId="5" pageNumber="277" refString="Berlese, A. (1903) Acari Mirmecofili. Redia, 1, 299 - 453." type="journal article" year="1903">Berlese, 1903</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BFC36B76066FFA453C7B20875844864" blockId="5.[151,1436,1193,2037]" pageId="5" pageNumber="277">
<keyLead id="30B295826066FFA453C7B20875844864" pageId="5" pageNumber="277">
- Dorsal shield with numerous (more than three pairs) semicircular strongly sclerotised structures...................................................................................................
<taxonomicName id="4C434D346066FFA45737B22F75844864" authority="Willmann, 1951" authorityName="Willmann" authorityYear="1951" box="[1127,1436,1897,1919]" class="Arachnida" family="Trachyuropodidae" genus="Cephalojanetia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="5" pageNumber="277" phylum="Arthropoda" rank="genus">
<emphasis id="B937EAA56066FFA45737B22F74E34864" box="[1127,1275,1898,1919]" italics="true" pageId="5" pageNumber="277">Cephalojanetia</emphasis>
<bibRefCitation id="EFD24B466066FFA45651B22C75844864" author="Willmann" box="[1281,1436,1897,1919]" pageId="5" pageNumber="277" refString="Willmann, C. (1951) Untersuchungen uber die terrestrische Milbenfauna im pannonischen Klimagebiet Osterreichs. Sitzungsberichte Osterreichische Akademie der Wissenschaften, 160, 91 - 176." type="journal article" year="1951">Willmann, 1951</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30B72E126066FFA453C7B2C2758348A1" pageId="5" pageNumber="277">
<paragraph id="8BFC36B76066FFA453C7B2C275844886" blockId="5.[151,1436,1193,2037]" box="[151,1436,1927,1949]" pageId="5" pageNumber="277">
<keyLead id="30B295826066FFA453C7B2C275844886" box="[151,1436,1927,1949]" pageId="5" pageNumber="277">
13. Dorsal strongly sclerotised structure circular (eye-like)............................
<emphasis id="B937EAA56066FFA45704B2C274FD4887" box="[1108,1253,1927,1948]" italics="true" pageId="5" pageNumber="277">Origmatrachys</emphasis>
<bibRefCitation id="EFD24B466066FFA457BBB2C275844886" author="Hirschmann" box="[1259,1436,1927,1949]" pageId="5" pageNumber="277" refString="Hirschmann, W. (1979) Stadiensystematik der Parasitiformes Teil 1. Stadienfamilien und Stadiengattungen der Atrichopygidiina, erstellt im Vergleich zum Gangsystem Hirschmann 1979. Acarologie. Schriftenreihe fur Vergleichende Milbenkunde, 26, 57 - 70." type="journal article" year="1979">Hirschmann, 1979</bibRefCitation>
</keyLead>
</paragraph>
<paragraph id="8BFC36B76066FFA453C7B2E0758348A1" blockId="5.[151,1436,1193,2037]" box="[151,1435,1956,1978]" pageId="5" pageNumber="277">
<keyLead id="30B295826066FFA453C7B2E0758348A1" box="[151,1435,1956,1978]" pageId="5" pageNumber="277">- Dorsal strongly sclerotised structure not circular........................................................... 14</keyLead>
</paragraph>
</keyStep>
<keyStep id="30B72E126066FFA453C7B287758348EE" pageId="5" pageNumber="277">
<paragraph id="8BFC36B76066FFA453C7B287758448C3" blockId="5.[151,1436,1193,2037]" box="[151,1436,1986,2008]" pageId="5" pageNumber="277">
<keyLead id="30B295826066FFA453C7B287758448C3" box="[151,1436,1986,2008]" pageId="5" pageNumber="277">
14. Dorsal strongly sclerotised structure U-shaped........................................
<taxonomicName id="4C434D346066FFA4572EB287758448C3" authority="Berlese, 1888" authorityName="Berlese" authorityYear="1888" box="[1150,1436,1986,2008]" class="Arachnida" family="Trachyuropodidae" genus="Trachyuropoda" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="5" pageNumber="277" phylum="Arthropoda" rank="genus">
<emphasis id="B937EAA56066FFA4572EB287750948CC" box="[1150,1297,1986,2007]" italics="true" pageId="5" pageNumber="277">Trachyuropoda</emphasis>
<bibRefCitation id="EFD24B466066FFA45648B287758448C3" author="Berlese" box="[1304,1436,1986,2008]" pageId="5" pageNumber="277" refString="Berlese, A. (1888) Acari Austro-Americani quos collegit Aloysius Balzan. Manipulus primus. Species novas circiter quinquaginta complectens. Bollettino della Societa Entomologica Italiana, 20, 171 - 222. + Plates V - XIII." type="journal article" year="1888">Berlese, 1888</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BFC36B76066FFA453C7B29A758348EE" blockId="5.[151,1436,1193,2037]" box="[151,1435,2015,2037]" pageId="5" pageNumber="277">
<keyLead id="30B295826066FFA453C7B29A758348EE" box="[151,1435,2015,2037]" pageId="5" pageNumber="277">- Dorsal strongly sclerotised structure C-shaped............................................................ 15</keyLead>
</paragraph>
</keyStep>
<keyStep id="30B72E126065FFA753C7B5D275844FD1" pageId="6" pageNumber="278">
<paragraph id="8BFC36B76065FFA753C7B5D275844FB6" blockId="6.[151,1436,151,202]" box="[151,1436,151,173]" pageId="6" pageNumber="278">
<keyLead id="30B295826065FFA753C7B5D275844FB6" box="[151,1436,151,173]" pageId="6" pageNumber="278">
15. C-shaped dorsal strongly sclerotised structure large..............................
<emphasis id="B937EAA56065FFA7571BB5D274FC4FB7" box="[1099,1252,151,172]" italics="true" pageId="6" pageNumber="278">Arculatatrachys</emphasis>
<bibRefCitation id="EFD24B466065FFA757BBB5D275844FB6" author="Hirschmann" box="[1259,1436,151,173]" pageId="6" pageNumber="278" refString="Hirschmann, W. (1979) Stadiensystematik der Parasitiformes Teil 1. Stadienfamilien und Stadiengattungen der Atrichopygidiina, erstellt im Vergleich zum Gangsystem Hirschmann 1979. Acarologie. Schriftenreihe fur Vergleichende Milbenkunde, 26, 57 - 70." type="journal article" year="1979">Hirschmann, 1979</bibRefCitation>
</keyLead>
</paragraph>
<paragraph id="8BFC36B76065FFA753C7B5F075844FD1" blockId="6.[151,1436,151,202]" box="[151,1436,180,202]" pageId="6" pageNumber="278">
<keyLead id="30B295826065FFA753C7B5F075844FD1" box="[151,1436,180,202]" pageId="6" pageNumber="278">
- C-shaped dorsal strongly sclerotised structure small...............................
<emphasis id="B937EAA56065FFA75706B5F074FC4FD1" box="[1110,1252,181,202]" italics="true" pageId="6" pageNumber="278">Graecatrachys</emphasis>
<bibRefCitation id="EFD24B466065FFA757BBB5F175844FD1" author="Hirschmann" box="[1259,1436,180,202]" pageId="6" pageNumber="278" refString="Hirschmann, W. (1979) Stadiensystematik der Parasitiformes Teil 1. Stadienfamilien und Stadiengattungen der Atrichopygidiina, erstellt im Vergleich zum Gangsystem Hirschmann 1979. Acarologie. Schriftenreihe fur Vergleichende Milbenkunde, 26, 57 - 70." type="journal article" year="1979">Hirschmann, 1979</bibRefCitation>
</keyLead>
</paragraph>
</keyStep>
</subSubSection>
</treatment>
</document>