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<document id="A7392215A8B7085249C7831102891EAE" ID-DOI="10.1590/2358-2936e2023006" ID-ISSN="2358-2936" ID-Zenodo-Dep="10716764" ID-ZooBank="4970E3C2-A1DB-4AAE8961-FBAA26505ED4AB" IM.bibliography_approvedBy="jonas" IM.illustrations_approvedBy="jonas" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="jonas" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="jonas" checkinTime="1709038466501" checkinUser="felipe" docAuthor="Rodríguez-Cabrera, Tomás M. &amp; de Armas, Luis F." docDate="2023" docId="03E987EBFF0FFFFBFEBEFCAEFB2FFD52" docLanguage="en" docName="Nauplius.31.e2023006.pdf" docOrigin="Nauplius (e 2023006) 31" docSource="http://dx.doi.org/10.1590/2358-2936e2023006" docStyle="DocumentStyle:64D98C772C5FBF4D3A503C51F9604E62.1:Nauplius.2017-.journal_article" docStyleId="64D98C772C5FBF4D3A503C51F9604E62" docStyleName="Nauplius.2017-.journal_article" docStyleVersion="1" docTitle="Acanthoniscus richardsonae Rodríguez-Cabrera &amp; de Armas 2023, sp. nov." docType="treatment" docVersion="3" lastPageNumber="18" masterDocId="FFD0FF93FF06FFEAFFA3FFD4FFBBFF89" masterDocTitle="Taxonomy of the enigmatic genus Acanthoniscus Gosse, 1851 (Isopoda: Oniscidea: Armadillidae), from Jamaica, with the description of a new species" masterLastPageNumber="25" masterPageNumber="1" pageNumber="10" updateTime="1712233860446" updateUser="jonas" zenodo-license-document="CC-BY-NC-4.0">
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<mods:title id="6E14E9740DA3D753921104F10C8FB27F">Taxonomy of the enigmatic genus Acanthoniscus Gosse, 1851 (Isopoda: Oniscidea: Armadillidae), from Jamaica, with the description of a new species</mods:title>
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<mods:namePart id="56BF4582CE68180AFB60DF5F216C3A26">Rodríguez-Cabrera, Tomás M.</mods:namePart>
<mods:affiliation id="B4CDC3A89D632056EF8804C781D13F3C">Instituto de Ecología y Sistemática. Boyeros, La Habana 11900, Cuba. E-mail: tomasmichel. rodriguez @ gmail. com &amp; DB-</mods:affiliation>
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<mods:namePart id="A5C880DAB63754482019813304BE91BD">de Armas, Luis F.</mods:namePart>
<mods:affiliation id="C9701263708AAFB8322FE4430FF43AAD">Sociedad Cubana de Zoología. San Antonio de los Baños, Artemisa 38100, Cuba. E-mail: luisdearmas 1945 @ gmail. com &amp; - A</mods:affiliation>
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<heading id="D0B78191FF0FFFE3FEBEFCAEFD59FC1E" bold="true" box="[285,738,890,919]" fontSize="12" level="2" pageId="9" pageNumber="10" reason="2">
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<taxonomicName id="4C404D7EFF0FFFE3FEBEFCAEFDC6FC1E" authority="Rodríguez-Cabrera &amp; de Armas, 2023" authorityName="Rodríguez-Cabrera &amp; de Armas" authorityYear="2023" box="[285,637,890,919]" class="Malacostraca" family="Delatorreiidae" genus="Acanthoniscus" kingdom="Animalia" order="Isopoda" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="richardsonae" status="sp. nov.">
<emphasis id="B934EAEFFF0FFFE3FEBEFCAEFDC6FC1E" bold="true" box="[285,637,890,919]" italics="true" pageId="9" pageNumber="10">Acanthoniscus richardsonae</emphasis>
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(
<figureCitation id="137B2A78FF0FFFE3FE05FC76FE48FC36" box="[422,499,930,959]" captionStart="Figure 1" captionStartId="3.[181,252,1611,1634]" captionTargetBox="[181,1467,151,1587]" captionTargetId="figure-19@3.[181,1467,151,1587]" captionTargetPageId="3" captionText="Figure 1. Distribution of the genus Acanthoniscus in Jamaica. Above: type locality of A. spiniger at the hills near Bluefields (yellow dot), the type locality of A.richardsonae sp.nov.in the Blue Mountains (red dot), and a second location in the Blue Mountains where a specimen presumably of A. richardsonae sp. nov. was photographed (orange dot). The main collection sites of Henry G. Hubbard in 1877 are depicted by green dots, notice the proximity of Stony Hill to the distribution of A. richardsonae sp.nov.Below: The hills known as Surinam Quarters near Bluefields, referred by Gosse (1851) as “Bluefields Mountain,” Westmoreland Parish, constitute the type locality of A. spiniger." figureDoi="http://doi.org/10.5281/zenodo.10716766" httpUri="https://zenodo.org/record/10716766/files/figure.png" pageId="9" pageNumber="10">Figs. 1</figureCitation>
, 
<figureCitation id="137B2A78FF0FFFE3FDA3FC76FD89FC36" box="[512,562,930,959]" captionStart-0="Figure 3" captionStart-1="Figure 4" captionStart-2="Figure 5" captionStart-3="Figure 6" captionStart-4="Figure 7" captionStart-5="Figure 8" captionStartId-0="10.[843,913,586,609]" captionStartId-1="11.[181,252,1628,1651]" captionStartId-2="12.[182,251,1692,1715]" captionStartId-3="13.[181,252,1788,1811]" captionStartId-4="14.[181,252,1786,1809]" captionStartId-5="15.[181,252,1523,1546]" captionTargetBox-0="[843,1467,151,563]" captionTargetBox-1="[246,1407,152,1593]" captionTargetBox-2="[312,1351,473,1658]" captionTargetBox-3="[267,1379,821,1745]" captionTargetBox-4="[387,1266,935,1753]" captionTargetBox-5="[192,1460,173,1475]" captionTargetId-0="figure-639@10.[843,1467,148,565]" captionTargetId-1="figure-20@11.[229,1420,148,1608]" captionTargetId-2="figure-134@12.[266,1382,465,1671]" captionTargetId-3="figure-164@13.[232,1415,817,1764]" captionTargetId-4="figure-324@14.[356,1292,932,1765]" captionTargetId-5="figure-248@15.[181,1466,151,1499]" captionTargetPageId-0="10" captionTargetPageId-1="11" captionTargetPageId-2="12" captionTargetPageId-3="13" captionTargetPageId-4="14" captionTargetPageId-5="15" captionText-0="Figure 3. Right lateral view of a female paratype Acanthoniscus richardsonaesp.nov.showingcolorationinalcohol.Scale bar=1 mm." captionText-1="Figure 4. Body parts of Acanthoniscus richardsonae sp. nov. A, Overall, dorsal view; B, detail of the microsculpture of the dorsal tegument (left) and tricorn-like setae (right); C, lateral view of the anterior region of the body;D, lateral view of the posterior region of the body; E, right pleon-epimera 3–5 in ventral view; F, right pereon-epimera 1–7 in ventral view; G, detail of pereon-epimera 1–2 in a more ventral position; H, cephalothorax in dorsal view; I, cephalothorax in frontal view (the tiny sensory setae inserted between the ommatidia are illustrated under a magnification of 400×: a, b, c); J, cephalothorax in ventral view; K, pleotelson and uropods in dorsal view (the uropod exopodites are slightly pointing upward); L, pleotelson and uropods in ventral view; M, right uropod in ventral view; N, right uropod in dorsal view; O, drawing of the overall dorsal view modified from Richardson (1909); P, arrangement of dorsal spines, the diametersof the circles correspond tothe relative size of the spines.Drawings of the corresponding structures modified from Richardson (1909)are included in I,J (the question mark means that the position representing thedrawing is not totally clear) and in M. Abbreviations: sympodite (sy), endopodite (en), and exopodite (ex). Scale bars = 1 mm (A–L), 0.5 mm (M–N); in some cases, a single scale bar applies to more than one figure (indicated by the letters above the scale bar)." captionText-2="Figure 5.Otherappendagesof Acanthoniscus richardsonae sp.nov.A, Second antenna;B,detail offlagellum of secondantennadepicting position of aesthetasc groups (small arrows); C, detail of the distal article of the flagellum of the second antenna slightly turned over showing the aesthetasc groups; D, detail of the distal-most portion of the flagellum of the second antenna showing the apical cone; E, first antenna; F, detail of the third article of the first antenna showing the aesthetascs; G, male left seventh pereiopod; H, detail of ischium ofmale seventh pereiopod; I, detailof dactylus ofmale seventh pereiopodin anterior view; J, detailof dactylus ofmale seventh pereiopod in posterior view; K, male left first pereiopod, showing the areas on the carpus and propodus covered with the cleaning apparatus or antennal brush (dashed outlines); L, detail of carpus ofmale first pereiopod (anterior surface).Abbreviations:basis (bs), ischium(is),merus(me),carpus (ca),propodus (pr),flagellum(fl),and dactylus(da).Drawingofthe corresponding structuremodified from Richardson (1909) is included inA. Scale bars = 1 mm (A), 0.5 mm (G,K), 0.3mm (B), 0.1 mm (C–E, H–J,L),0.05 mm (F)." captionText-3="Figure 6.Mouthparts of Acanthoniscus richardsonae sp. nov. A, Distal portion of right (R) and left (L) mandibles in anterior views, showing lacinia mobilis (lm), hairy lobe (hl), penicils (pn), and tuft of setae (ts); B, first maxilla, with detailof distalportion of outer endite (ex) and inner endite (en); C, second maxilla; D, detail of the distal-most portion of the second maxilla; E, hypopharynx; F, detail of the median region of hypopharynx; G, maxilliped; H, detail of the distal-most portion of maxilliped in posterior view; I, detail of the distal-most portion of maxilliped inanterior view.Drawings of the corresponding structures modified from Richardson (1909) are included in B, C, and G. Scale bars = 0.25 mm." captionText-4="Figure 7.Pleopods of a male Acanthoniscus richardsonae sp.nov.A, First pleopod; B, detail of first pleopod endopodite tip; C, detail of first pleopod exopodite; D, second pleopod, with detail of the two setae on exopodite outer margin (arrow); E, third pleopod exopodite;F, detail ofthe outer-basal regionof the thirdpleopod exopoditeshowing the respiratory structures (thick arrow indicates the position of the spiracle); G, detail of the outer-basal region of the third pleopod exopodite showing the spiracle partially closed from above; H, fourth pleopod exopodite; I, fifth pleopod exopodite. Abbreviations: endopodite (en) and exopodite (ex). Scale bars = 0.5 mm (A–H), 0.25 mm (I)." captionText-5="Figure 8. Selected body parts of Acanthoniscus for comparison with other taxa within Armadillidae. Cubaris murina Brandt, 1833, showing the ground pattern in Armadillidae (A–E): A, Posterior region of the body in dorsal view; B, posterior region of the body in ventral view; C, pleotelson in dorsal view; D, uropod in dorsal view; E, middle region of the sternite 7 of a male showing the bilobed caudal process on the posterior margin, which is considered an autapomorphy of Armadillidae.Acanthoniscus richardsonae sp. nov.(F–H): F, pleotelson in dorsal view; G, uropod in dorsal view; H, middle region of the sternite 7 of a male.Drawings of the pleotelson and uropods in dorsal views of some morphologically divergent taxa within Armadillidae. I, Echinodillo cavaticus Green, 1963; J, Echinodillo montanus Jackson, 1933; K, Laureola vietnamensis Kwon, Ferrara and Taiti, 1992; L, Laureola dubia Schmalfuss and Ferrara,1983; M, Schismadillo rouxi Verhoeff,1926; N, Merulanella wahrbergi Verhoeff,1926; O, Nesodillo incisus Verhoeff, 1926. Drawings modified from Verhoeff (1926) (M–O), Jackson (1933) (J), Green (1963) (I), Schmalfuss and Ferrara (1983) (L), and Kwon et al. (1992) (K). Abbreviations: endopodite (en), exopodite (ex), sympodite (sy)." figureDoi-0="http://doi.org/10.5281/zenodo.10716770" figureDoi-1="http://doi.org/10.5281/zenodo.10716774" figureDoi-2="http://doi.org/10.5281/zenodo.10716776" figureDoi-3="http://doi.org/10.5281/zenodo.10716778" figureDoi-4="http://doi.org/10.5281/zenodo.10716780" figureDoi-5="http://doi.org/10.5281/zenodo.10716782" httpUri-0="https://zenodo.org/record/10716770/files/figure.png" httpUri-1="https://zenodo.org/record/10716774/files/figure.png" httpUri-2="https://zenodo.org/record/10716776/files/figure.png" httpUri-3="https://zenodo.org/record/10716778/files/figure.png" httpUri-4="https://zenodo.org/record/10716780/files/figure.png" httpUri-5="https://zenodo.org/record/10716782/files/figure.png" pageId="9" pageNumber="10">3–8</figureCitation>
, 
<figureCitation id="137B2A78FF0FFFE3FD9EFC76FDE7FC36" box="[573,604,930,959]" captionStart="Figure 10" captionStartId="17.[180,251,963,986]" captionTargetBox="[181,805,151,940]" captionTargetId="figure-510@17.[181,805,148,943]" captionTargetPageId="17" captionText="Figure 10. Specimen of Acanthoniscus cf. richardsonae sp. nov. observed near Hardwar Gap, Blue and John Crow Mountains National Park, Portland Parish. These are the only available photographs of a live specimen of the genus Acanthoniscus. Photos by Sarah C. Crews." figureDoi="http://doi.org/10.5281/zenodo.10716786" httpUri="https://zenodo.org/record/10716786/files/figure.png" pageId="9" pageNumber="10">10</figureCitation>
)
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ZooBank: 
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urn:lsid:zoobank.org:act: 
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<paragraph id="8BFF36FDFF0FFFE3FF16FBB0FE11FA1B" blockId="9.[179,808,1124,1737]" pageId="9" pageNumber="10">
<taxonomicName id="4C404D7EFF0FFFE3FF16FBB0FE05FB08" authorityName=", White, List" authorityYear="1847" box="[181,446,1124,1153]" class="Malacostraca" family="Delatorreiidae" genus="Acanthoniscus" kingdom="Animalia" order="Isopoda" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="spiniger">
<emphasis id="B934EAEFFF0FFFE3FF16FBB0FE05FB08" box="[181,446,1124,1153]" italics="true" pageId="9" pageNumber="10">Acanthoniscus spiniger</emphasis>
</taxonomicName>
(partim): 
<bibRefCitation id="EFD14B0CFF0FFFE3FDE0FBB0FC9EFB0B" author="Richardson H" box="[579,805,1124,1154]" pageId="9" pageNumber="10" pagination="431 - 434" refId="ref17042" refString="Richardson H 1909. The isopod crustacean Acanthoniscus spiniger Kinahan redescribed. Proceedings of the United States National Museum, 36: 431 - 434. DOI: https: // www. doi. org / 10.5479 / si. 00963801.36 - 1678.431" type="journal article" year="1909">Richardson, 1909:</bibRefCitation>
431–434 (misidentification). — 
<bibRefCitation id="EFD14B0CFF0FFFE3FD23FB5FFC93FB21" author="Budde-Lund G" box="[640,808,1163,1192]" pageId="9" pageNumber="10" pagination="1905 - 1906" refId="ref14713" refString="Budde-Lund G 1910. Crustacea, 2. Isopoda. In: Wissenschaftliche Ergebnisse der schwedischen zoologischen Expedition nach dem Kilimandjaro, dem Meru und den umgebenden Massaisteppen Deutsch-Ostafrikas, 1905 - 1906. p. 3 - 20, pls. 1 - 2. Stockholm, P. Palmquists Aktebolag. https: // www. biodiversitylibrary. org / item / 16983 # page / 651 / mode / 1 up" type="book chapter" year="1910">Budde-Lund,</bibRefCitation>
1910: 11 (partim). — 
<bibRefCitation id="EFD14B0CFF0FFFE3FDA3FB66FC9EFB46" author="Richardson H" box="[512,805,1202,1231]" pageId="9" pageNumber="10" pagination="495 - 497" refId="ref17086" refString="Richardson H 1910. Description of a new terrestrial isopod from Guatemala. Proceedings of the U. S. National Museum, 37: 495 - 497. DOI: https: // www. doi. org / 10.5479 / si. 00963801.37 - 1718.495" type="journal article" year="1910">Richardson, 1910: 495</bibRefCitation>
(partim). — 
<bibRefCitation id="EFD14B0CFF0FFFE3FED3FB0DFDE6FB7F" author="Arcangeli A" box="[368,605,1241,1270]" pageId="9" pageNumber="10" pagination="126 - 141" refId="ref14187" refString="Arcangeli A 1927. Revisione dei generi degli isopodi terrestri. I. a Nota. - Sopra alcuni generi di Africa e di America. Atti della Societa Italiana di Scienze Naturali e del Museo Civico di Storia Naturale in Milano, 66: 126 - 141. https: // www. biodiversitylibrary. org / item / 264438 - page / 144 / mode / 1 up" type="journal article" year="1927">Arcangeli, 1927:135</bibRefCitation>
(partim). — 
<bibRefCitation id="EFD14B0CFF0FFFE3FD55FB0EFC9EFB7F" author="Van Name W" box="[758,805,1242,1270]" pageId="9" pageNumber="10" pagination="1 - 535" refId="ref17876" refString="Van Name W 1936. The American land and freshwater isopod Crustacea. Bulletin of the American Museum of natural History, 71: 1 - 535. https: // hdl. handle. net / 2246 / 1185" type="journal article" year="1936">Van</bibRefCitation>
Name, 1936: 402–403 (partim). — 
<bibRefCitation id="EFD14B0CFF0FFFE3FD33FAD4FC98FA94" author="Schmalfuss H" box="[656,803,1280,1309]" pageId="9" pageNumber="10" refId="ref17197" refString="Schmalfuss H 2003. World catalog of terrestrial isopods (Isopoda: Oniscidea). Stuttgarter Beitrage zur Naturkunde, Serie A, 654: 341 p. https: // www. biodiversitylibrary. org / item / 293136 - page / 1 / mode / 1 up" type="book" year="2003">Schmalfuss,</bibRefCitation>
2003: 4 (partim). — 
<bibRefCitation id="EFD14B0CFF0FFFE3FE57FAF3FC9EFACD" author="Schmidt C &amp; Leistikow, A" box="[500,805,1319,1348]" pageId="9" pageNumber="10" pagination="1 - 118" refId="ref17528" refString="Schmidt C and Leistikow, A 2004. Catalogue of genera of the terrestrial Isopoda (Crustacea: Isopoda: Oniscidea). Steenstrupia, 28: 1 - 118." type="journal article" year="2004">Schmidt and Leistikow,</bibRefCitation>
2004: 4 (partim). — 
<bibRefCitation id="EFD14B0CFF0FFFE3FE7DFA9BFC9EFAE2" author="Jass J &amp; Klausmeier B" box="[478,805,1358,1387]" pageId="9" pageNumber="10" pagination="1 - 36" refId="ref16111" refString="Jass J and Klausmeier B 2006. Terrestrial Isopods (Crustacea: Isopoda: Oniscidea) of the West Indies. Insight, A Milwaukee Public Museum Series in Natural History, 3: 1 - 36. https: // decapoda. nhm. org / pdfs / 30354 / 30354. pdf" type="journal article" year="2006">Jass and Klausmeier, 2006:</bibRefCitation>
4, 6, 25 (partim).
</paragraph>
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<paragraph id="8BFF36FDFF0FFFE3FF78FA49FE4FF940" blockId="9.[179,808,1124,1737]" pageId="9" pageNumber="10">
<emphasis id="B934EAEFFF0FFFE3FF78FA49FE3FFA30" box="[219,388,1436,1465]" italics="true" pageId="9" pageNumber="10">Type material.</emphasis>
<materialsCitation id="3B283CA0FF0FFFE3FE28FA48FC98F9F2" collectingDate="2013-11-15" collectionCode="CZACC" country="Jamaica" county="Franklyn Cala Riquelme" elevation="1250" latitude="18.0535" location="Blue Mountains" longLatPrecision="1" longitude="-76.56311" municipality="Abel Perez-Gonzalez" pageId="9" pageNumber="10" specimenCode="CZACC 5.0500" specimenCount="1" specimenCount-male="1" stateProvince="Saint Thomas" typeStatus="holotype">
<typeStatus id="54FB885FFF0FFFE3FE28FA48FDBEFA30" box="[395,517,1436,1465]" pageId="9" pageNumber="10" type="holotype">Holotype</typeStatus>
: 
<specimenCount id="9D46FD74FF0FFFE3FDB3FA48FD85FA35" box="[528,574,1436,1468]" count="1" pageId="9" pageNumber="10" type="male">1 ♂</specimenCount>
(
<specimenCode id="DBE69E86FF0FFFE3FDEDFA48FCA2FA30" box="[590,793,1436,1465]" collectionCode="CZACC" country="Cuba" httpUri="http://grbio.org/cool/b49z-cz3a" name="Coleccion Zoologia, Academia de Ciencias de Cuba" pageId="9" pageNumber="10">CZACC 5.0500</specimenCode>
), around the ranger station (
<geoCoordinate id="EE74503AFF0FFFE3FE42FA17FD3AFA69" box="[481,641,1475,1504]" degrees="18" direction="north" minutes="03" orientation="latitude" pageId="9" pageNumber="10" precision="1" seconds="12.6" value="18.0535">18°03’12.6”N</geoCoordinate>
<geoCoordinate id="EE74503AFF0FFFE3FD26FA17FC99FA69" box="[645,802,1475,1504]" degrees="76" direction="west" minutes="33" orientation="longitude" pageId="9" pageNumber="10" precision="1" seconds="47.2" value="-76.56311">76°33’47.2”W</geoCoordinate>
, approximately 
<elevation id="006DD1CEFF0FFFE3FEC5FA3EFDB9F98E" box="[358,514,1514,1543]" metricMagnitude="3" metricUnit="m" metricValue="1.25" pageId="9" pageNumber="10" unit="m" value="1250.0">
<quantity id="4CB89B18FF0FFFE3FEC5FA3EFE7CF98F" box="[358,455,1514,1543]" metricMagnitude="3" metricUnit="m" metricValue="1.25" pageId="9" pageNumber="10" unit="m" value="1250.0">1,250 m</quantity>
a.s.l.
</elevation>
), 
<location id="8E9F6026FF0FFFE3FDB4FA3EFD61F98E" LSID="urn:lsid:plazi:treatment:03E987EBFF0FFFFBFEBEFCAEFB2FFD52:8E9F6026FF0FFFE3FDB4FA3EFD61F98E" box="[535,730,1514,1543]" country="Jamaica" county="Franklyn Cala Riquelme" latitude="18.0535" longLatPrecision="1" longitude="-76.56311" municipality="Abel Perez-Gonzalez" name="Blue Mountains" pageId="9" pageNumber="10" stateProvince="Saint Thomas">Blue Mountains</location>
, 
<collectingRegion id="4984F81FFF0FFFE3FD47FA3EFED6F9A7" country="Jamaica" name="Saint Thomas" pageId="9" pageNumber="10">Saint Thomas Parish</collectingRegion>
, 
<collectingCountry id="F357766DFF0FFFE3FED5F9C5FE6EF9A7" box="[374,469,1553,1582]" name="Jamaica" pageId="9" pageNumber="10">Jamaica</collectingCountry>
(
<figureCitation id="137B2A78FF0FFFE3FE45F9C5FD9EF9A7" box="[486,549,1553,1582]" captionStart="Figure 1" captionStartId="3.[181,252,1611,1634]" captionTargetBox="[181,1467,151,1587]" captionTargetId="figure-19@3.[181,1467,151,1587]" captionTargetPageId="3" captionText="Figure 1. Distribution of the genus Acanthoniscus in Jamaica. Above: type locality of A. spiniger at the hills near Bluefields (yellow dot), the type locality of A.richardsonae sp.nov.in the Blue Mountains (red dot), and a second location in the Blue Mountains where a specimen presumably of A. richardsonae sp. nov. was photographed (orange dot). The main collection sites of Henry G. Hubbard in 1877 are depicted by green dots, notice the proximity of Stony Hill to the distribution of A. richardsonae sp.nov.Below: The hills known as Surinam Quarters near Bluefields, referred by Gosse (1851) as “Bluefields Mountain,” Westmoreland Parish, constitute the type locality of A. spiniger." figureDoi="http://doi.org/10.5281/zenodo.10716766" httpUri="https://zenodo.org/record/10716766/files/figure.png" pageId="9" pageNumber="10">Fig. 1</figureCitation>
), 
<date id="FFFE103DFF0FFFE3FD98F9C4FC9AF9A7" box="[571,801,1552,1582]" pageId="9" pageNumber="10" value="2013-11-15">
<collectingDate id="EFBAE9D5FF0FFFE3FD98F9C4FC9AF9A7" box="[571,801,1552,1582]" pageId="9" pageNumber="10" value="2013-11-15">15 November 2013</collectingDate>
</date>
, 
<collectingMunicipality id="6B9BAC87FF0FFFE3FF17F9ECFE0EF9DC" box="[180,437,1592,1621]" pageId="9" pageNumber="10">Abel Pérez-González</collectingMunicipality>
and 
<collectingCounty id="629E4E71FF0FFFE3FE50F9ECFC98F9DC" box="[499,803,1592,1621]" pageId="9" pageNumber="10">Franklyn Cala Riquelme</collectingCounty>
(collectors’ code: “JA 10-2”, “1671Caribbean”)
</materialsCitation>
. 
<materialsCitation id="3B283CA0FF0FFFE3FF16F952FE4BF940" collectingDate="2013-11-15" collectionCode="CZACC" country="Jamaica" county="Franklyn Cala Riquelme" elevation="1250" latitude="18.0535" location="Blue Mountains" longLatPrecision="1" longitude="-76.56311" municipality="Abel Perez-Gonzalez" pageId="9" pageNumber="10" specimenCode="CZACC 5.0501" specimenCount="8" specimenCount-female="6" specimenCount-male="2" stateProvince="Saint Thomas" typeStatus="paratype">
<typeStatus id="54FB885FFF0FFFE3FF16F952FE8FF92B" box="[181,308,1670,1698]" pageId="9" pageNumber="10" type="paratype">Paratypes</typeStatus>
: 
<specimenCount id="9D46FD74FF0FFFE3FEE2F952FE33F92C" box="[321,392,1670,1701]" count="2" pageId="9" pageNumber="10" type="male">2 ♂♂</specimenCount>
, 
<specimenCount id="9D46FD74FF0FFFE3FE35F951FE67F92C" box="[406,476,1669,1701]" count="6" pageId="9" pageNumber="10" type="female">6 ♀♀</specimenCount>
, 2 juvs. (
<specimenCode id="DBE69E86FF0FFFE3FDEEF952FCACF92B" box="[589,791,1669,1699]" collectionCode="CZACC" country="Cuba" httpUri="http://grbio.org/cool/b49z-cz3a" name="Coleccion Zoologia, Academia de Ciencias de Cuba" pageId="9" pageNumber="10">CZACC 5.0501</specimenCode>
), same data as the holotype
</materialsCitation>
.
</paragraph>
<paragraph id="8BFF36FDFF0FFFE3FF78F92EFA06FED9" blockId="9.[181,805,1786,1971]" lastBlockId="9.[843,1469,151,336]" pageId="9" pageNumber="10">
<emphasis id="B934EAEFFF0FFFE3FF78F92EFE63F89E" box="[219,472,1786,1815]" italics="true" pageId="9" pageNumber="10">Additional material</emphasis>
(not 
<typeStatus id="54FB885FFF0FFFE3FD8FF92EFD1FF89E" box="[556,676,1786,1815]" pageId="9" pageNumber="10" type="paratype">paratype</typeStatus>
). 
<collectingCountry id="F357766DFF0FFFE3FD1CF92EFC9FF89E" box="[703,804,1786,1815]" name="Jamaica" pageId="9" pageNumber="10">Jamaica</collectingCountry>
(unknown locality), a single specimen collected by Hubbard in 1877, the original label reads: “ 
<taxonomicName id="4C404D7EFF0FFFE3FF63F8A4FE35F805" box="[192,398,1904,1932]" class="Malacostraca" family="Oniscidae" genus="Oniscus" kingdom="Animalia" order="Isopoda" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="spiniger">
<emphasis id="B934EAEFFF0FFFE3FF63F8A4FE35F805" box="[192,398,1904,1932]" italics="true" pageId="9" pageNumber="10">Oniscus spiniger</emphasis>
</taxonomicName>
. 
<collectingCountry id="F357766DFF0FFFE3FE00F8BBFDB1F805" box="[419,522,1903,1932]" name="Jamaica" pageId="9" pageNumber="10">Jamaica</collectingCountry>
” (
<bibRefCitation id="EFD14B0CFF0FFFE3FD8FF8BBFF5FF83A" author="Richardson H" pageId="9" pageNumber="10" pagination="431 - 434" refId="ref17042" refString="Richardson H 1909. The isopod crustacean Acanthoniscus spiniger Kinahan redescribed. Proceedings of the United States National Museum, 36: 431 - 434. DOI: https: // www. doi. org / 10.5479 / si. 00963801.36 - 1678.431" type="journal article" year="1909">Richardson, 1909: 431</bibRefCitation>
). The current catalog number is: 
<collectionCode id="ED51AE38FF0FFFE3FDD9F842FD6FF83A" box="[634,724,1942,1971]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="9" pageNumber="10" type="Museum">USNM</collectionCode>
41501 (https://n 
<quantity id="4CB89B18FF0FFFE3FC18FF43FC68FF3D" box="[955,979,151,180]" metricMagnitude="3" metricUnit="kg" metricValue="2.0" pageId="9" pageNumber="10" unit="t" value="2.0">2t</quantity>
.net/ark:/65665/3ac606ed9-99f9-4214- 891d-1e92ed7ccbe4). This specimen served as the basis for 
<bibRefCitation id="EFD14B0CFF0FFFE3FC1AFF31FB0BFE8B" author="Richardson H" box="[953,1200,229,258]" pageId="9" pageNumber="10" pagination="431 - 434" refId="ref17042" refString="Richardson H 1909. The isopod crustacean Acanthoniscus spiniger Kinahan redescribed. Proceedings of the United States National Museum, 36: 431 - 434. DOI: https: // www. doi. org / 10.5479 / si. 00963801.36 - 1678.431" type="journal article" year="1909">Richardson’s (1909)</bibRefCitation>
description, assumed as a redescription of 
<taxonomicName id="4C404D7EFF0FFFE3FBEFFED8FB7DFEA0" box="[1100,1222,268,297]" class="Malacostraca" family="Oniscidae" genus="Oniscus" kingdom="Animalia" order="Isopoda" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="spiniger">
<emphasis id="B934EAEFFF0FFFE3FBEFFED8FB7DFEA0" box="[1100,1222,268,297]" italics="true" pageId="9" pageNumber="10">A. spiniger</emphasis>
</taxonomicName>
. It was not possible for us to review this specimen directly or by pictures.
</paragraph>
</subSubSection>
<subSubSection id="C35A6576FF0FFFE3FCD2FE56FB7FF83A" pageId="9" pageNumber="10" type="diagnosis">
<paragraph id="8BFF36FDFF0FFFE3FCD2FE56FB7FF83A" blockId="9.[841,1468,385,1971]" pageId="9" pageNumber="10">
<emphasis id="B934EAEFFF0FFFE3FCD2FE56FC53FE17" box="[881,1000,386,414]" italics="true" pageId="9" pageNumber="10">Diagnosis</emphasis>
(Diagnosis and description are the same for the 
<typeStatus id="54FB885FFF0FFFE3FC55FE7CFBDCFE4C" box="[1014,1127,424,453]" pageId="9" pageNumber="10" type="holotype">holotype</typeStatus>
and the 
<typeStatus id="54FB885FFF0FFFE3FB7EFE7CFAE7FE4D" box="[1245,1372,424,452]" pageId="9" pageNumber="10" type="paratype">paratypes</typeStatus>
, unless specified). Maximum total length: 
<quantity id="4CB89B18FF0FFFE3FB50FE1BFAF4FE62" box="[1267,1359,463,492]" metricMagnitude="-3" metricUnit="m" metricValue="7.2" pageId="9" pageNumber="10" unit="mm" value="7.2">7.2 mm</quantity>
in males (
<typeStatus id="54FB885FFF0FFFE3FCF0FE21FC7EFD9B" box="[851,965,501,530]" pageId="9" pageNumber="10" type="holotype">holotype</typeStatus>
) and 
<quantity id="4CB89B18FF0FFFE3FBADFE21FBD4FD9B" box="[1038,1135,501,530]" metricMagnitude="-3" metricUnit="m" metricValue="8.5" pageId="9" pageNumber="10" unit="mm" value="8.5">8.5 mm</quantity>
in females (see Variation). Cephalothorax with rounded lateral lobes, and a well-developed, deeply notched median lobe, forming two triangular projections, dorsal surface with a pair of small paramedian spiniform tubercles and three medium-sized, club-like, posteromedian spines (
<figureCitation id="137B2A78FF0FFFE3FC0AFD0BFBFCFD75" box="[937,1095,735,764]" captionStart="Figure 4" captionStartId="11.[181,252,1628,1651]" captionTargetBox="[246,1407,152,1593]" captionTargetId="figure-20@11.[229,1420,148,1608]" captionTargetPageId="11" captionText="Figure 4. Body parts of Acanthoniscus richardsonae sp. nov. A, Overall, dorsal view; B, detail of the microsculpture of the dorsal tegument (left) and tricorn-like setae (right); C, lateral view of the anterior region of the body;D, lateral view of the posterior region of the body; E, right pleon-epimera 3–5 in ventral view; F, right pereon-epimera 1–7 in ventral view; G, detail of pereon-epimera 1–2 in a more ventral position; H, cephalothorax in dorsal view; I, cephalothorax in frontal view (the tiny sensory setae inserted between the ommatidia are illustrated under a magnification of 400×: a, b, c); J, cephalothorax in ventral view; K, pleotelson and uropods in dorsal view (the uropod exopodites are slightly pointing upward); L, pleotelson and uropods in ventral view; M, right uropod in ventral view; N, right uropod in dorsal view; O, drawing of the overall dorsal view modified from Richardson (1909); P, arrangement of dorsal spines, the diametersof the circles correspond tothe relative size of the spines.Drawings of the corresponding structures modified from Richardson (1909)are included in I,J (the question mark means that the position representing thedrawing is not totally clear) and in M. Abbreviations: sympodite (sy), endopodite (en), and exopodite (ex). Scale bars = 1 mm (A–L), 0.5 mm (M–N); in some cases, a single scale bar applies to more than one figure (indicated by the letters above the scale bar)." figureDoi="http://doi.org/10.5281/zenodo.10716774" httpUri="https://zenodo.org/record/10716774/files/figure.png" pageId="9" pageNumber="10">Fig. 4C, H–J</figureCitation>
). Pereonite 1 with two pairs of paramedian spiniform tubercles and three medium-sized posteromedian spines; pereonites 2–7 with a pair of paramedian spiniform tubercles and three medium-sized posteromedian spines (
<figureCitation id="137B2A78FF0FFFE3FAB5FCAFFC32FC36" captionStart="Figure 4" captionStartId="11.[181,252,1628,1651]" captionTargetBox="[246,1407,152,1593]" captionTargetId="figure-20@11.[229,1420,148,1608]" captionTargetPageId="11" captionText="Figure 4. Body parts of Acanthoniscus richardsonae sp. nov. A, Overall, dorsal view; B, detail of the microsculpture of the dorsal tegument (left) and tricorn-like setae (right); C, lateral view of the anterior region of the body;D, lateral view of the posterior region of the body; E, right pleon-epimera 3–5 in ventral view; F, right pereon-epimera 1–7 in ventral view; G, detail of pereon-epimera 1–2 in a more ventral position; H, cephalothorax in dorsal view; I, cephalothorax in frontal view (the tiny sensory setae inserted between the ommatidia are illustrated under a magnification of 400×: a, b, c); J, cephalothorax in ventral view; K, pleotelson and uropods in dorsal view (the uropod exopodites are slightly pointing upward); L, pleotelson and uropods in ventral view; M, right uropod in ventral view; N, right uropod in dorsal view; O, drawing of the overall dorsal view modified from Richardson (1909); P, arrangement of dorsal spines, the diametersof the circles correspond tothe relative size of the spines.Drawings of the corresponding structures modified from Richardson (1909)are included in I,J (the question mark means that the position representing thedrawing is not totally clear) and in M. Abbreviations: sympodite (sy), endopodite (en), and exopodite (ex). Scale bars = 1 mm (A–L), 0.5 mm (M–N); in some cases, a single scale bar applies to more than one figure (indicated by the letters above the scale bar)." figureDoi="http://doi.org/10.5281/zenodo.10716774" httpUri="https://zenodo.org/record/10716774/files/figure.png" pageId="9" pageNumber="10">Fig. 4A, C–D, O–P</figureCitation>
); epimera more or less quadrangular (except epimera 4: subtriangular, very narrow and sharp-pointed), with anterior (epimera 2–3) or posterior (epimera 5–7) margins widely convex and produced (forming an almost square angle) (
<figureCitation id="137B2A78FF0FFFE3FB4FFBE9FADDFBD2" box="[1260,1382,1085,1115]" captionStart="Figure 4" captionStartId="11.[181,252,1628,1651]" captionTargetBox="[246,1407,152,1593]" captionTargetId="figure-20@11.[229,1420,148,1608]" captionTargetPageId="11" captionText="Figure 4. Body parts of Acanthoniscus richardsonae sp. nov. A, Overall, dorsal view; B, detail of the microsculpture of the dorsal tegument (left) and tricorn-like setae (right); C, lateral view of the anterior region of the body;D, lateral view of the posterior region of the body; E, right pleon-epimera 3–5 in ventral view; F, right pereon-epimera 1–7 in ventral view; G, detail of pereon-epimera 1–2 in a more ventral position; H, cephalothorax in dorsal view; I, cephalothorax in frontal view (the tiny sensory setae inserted between the ommatidia are illustrated under a magnification of 400×: a, b, c); J, cephalothorax in ventral view; K, pleotelson and uropods in dorsal view (the uropod exopodites are slightly pointing upward); L, pleotelson and uropods in ventral view; M, right uropod in ventral view; N, right uropod in dorsal view; O, drawing of the overall dorsal view modified from Richardson (1909); P, arrangement of dorsal spines, the diametersof the circles correspond tothe relative size of the spines.Drawings of the corresponding structures modified from Richardson (1909)are included in I,J (the question mark means that the position representing thedrawing is not totally clear) and in M. Abbreviations: sympodite (sy), endopodite (en), and exopodite (ex). Scale bars = 1 mm (A–L), 0.5 mm (M–N); in some cases, a single scale bar applies to more than one figure (indicated by the letters above the scale bar)." figureDoi="http://doi.org/10.5281/zenodo.10716774" httpUri="https://zenodo.org/record/10716774/files/figure.png" pageId="9" pageNumber="10">Fig. 4C, F</figureCitation>
); coxal plates 5–7 with a medium-sized tooth-like process, in medial position, forming a schisma and extending anteriorly into an oblique, very shallow, longitudinal groove, and with a very small submarginal tooth-like process, in basal position, just posterior to pereiopod’s junction, forming a second very small schisma; latter two schisma forming a continuum engaging with the contiguous posterior pereon epimera during conglobation (
<figureCitation id="137B2A78FF0FFFE3FBA7FA48FBDAFA30" box="[1028,1121,1436,1465]" captionStart="Figure 4" captionStartId="11.[181,252,1628,1651]" captionTargetBox="[246,1407,152,1593]" captionTargetId="figure-20@11.[229,1420,148,1608]" captionTargetPageId="11" captionText="Figure 4. Body parts of Acanthoniscus richardsonae sp. nov. A, Overall, dorsal view; B, detail of the microsculpture of the dorsal tegument (left) and tricorn-like setae (right); C, lateral view of the anterior region of the body;D, lateral view of the posterior region of the body; E, right pleon-epimera 3–5 in ventral view; F, right pereon-epimera 1–7 in ventral view; G, detail of pereon-epimera 1–2 in a more ventral position; H, cephalothorax in dorsal view; I, cephalothorax in frontal view (the tiny sensory setae inserted between the ommatidia are illustrated under a magnification of 400×: a, b, c); J, cephalothorax in ventral view; K, pleotelson and uropods in dorsal view (the uropod exopodites are slightly pointing upward); L, pleotelson and uropods in ventral view; M, right uropod in ventral view; N, right uropod in dorsal view; O, drawing of the overall dorsal view modified from Richardson (1909); P, arrangement of dorsal spines, the diametersof the circles correspond tothe relative size of the spines.Drawings of the corresponding structures modified from Richardson (1909)are included in I,J (the question mark means that the position representing thedrawing is not totally clear) and in M. Abbreviations: sympodite (sy), endopodite (en), and exopodite (ex). Scale bars = 1 mm (A–L), 0.5 mm (M–N); in some cases, a single scale bar applies to more than one figure (indicated by the letters above the scale bar)." figureDoi="http://doi.org/10.5281/zenodo.10716774" httpUri="https://zenodo.org/record/10716774/files/figure.png" pageId="9" pageNumber="10">Fig. 4F</figureCitation>
). Pleonites 3–4 with a pair of small posteromedian spines; pleon epimera 3–5 with anterior margin widely convex and produced (forming an almost square angle) (
<figureCitation id="137B2A78FF0FFFE3FB78F9C5FA94F9A7" box="[1243,1327,1553,1582]" captionStart="Figure 4" captionStartId="11.[181,252,1628,1651]" captionTargetBox="[246,1407,152,1593]" captionTargetId="figure-20@11.[229,1420,148,1608]" captionTargetPageId="11" captionText="Figure 4. Body parts of Acanthoniscus richardsonae sp. nov. A, Overall, dorsal view; B, detail of the microsculpture of the dorsal tegument (left) and tricorn-like setae (right); C, lateral view of the anterior region of the body;D, lateral view of the posterior region of the body; E, right pleon-epimera 3–5 in ventral view; F, right pereon-epimera 1–7 in ventral view; G, detail of pereon-epimera 1–2 in a more ventral position; H, cephalothorax in dorsal view; I, cephalothorax in frontal view (the tiny sensory setae inserted between the ommatidia are illustrated under a magnification of 400×: a, b, c); J, cephalothorax in ventral view; K, pleotelson and uropods in dorsal view (the uropod exopodites are slightly pointing upward); L, pleotelson and uropods in ventral view; M, right uropod in ventral view; N, right uropod in dorsal view; O, drawing of the overall dorsal view modified from Richardson (1909); P, arrangement of dorsal spines, the diametersof the circles correspond tothe relative size of the spines.Drawings of the corresponding structures modified from Richardson (1909)are included in I,J (the question mark means that the position representing thedrawing is not totally clear) and in M. Abbreviations: sympodite (sy), endopodite (en), and exopodite (ex). Scale bars = 1 mm (A–L), 0.5 mm (M–N); in some cases, a single scale bar applies to more than one figure (indicated by the letters above the scale bar)." figureDoi="http://doi.org/10.5281/zenodo.10716774" httpUri="https://zenodo.org/record/10716774/files/figure.png" pageId="9" pageNumber="10">Fig. 4E</figureCitation>
). Pleotelson with posterior margin ending in two small triangular projections separated by a shallow notch (
<figureCitation id="137B2A78FF0FFFE3FAEDF98AFC3EF92B" captionStart="Figure 4" captionStartId="11.[181,252,1628,1651]" captionTargetBox="[246,1407,152,1593]" captionTargetId="figure-20@11.[229,1420,148,1608]" captionTargetPageId="11" captionText="Figure 4. Body parts of Acanthoniscus richardsonae sp. nov. A, Overall, dorsal view; B, detail of the microsculpture of the dorsal tegument (left) and tricorn-like setae (right); C, lateral view of the anterior region of the body;D, lateral view of the posterior region of the body; E, right pleon-epimera 3–5 in ventral view; F, right pereon-epimera 1–7 in ventral view; G, detail of pereon-epimera 1–2 in a more ventral position; H, cephalothorax in dorsal view; I, cephalothorax in frontal view (the tiny sensory setae inserted between the ommatidia are illustrated under a magnification of 400×: a, b, c); J, cephalothorax in ventral view; K, pleotelson and uropods in dorsal view (the uropod exopodites are slightly pointing upward); L, pleotelson and uropods in ventral view; M, right uropod in ventral view; N, right uropod in dorsal view; O, drawing of the overall dorsal view modified from Richardson (1909); P, arrangement of dorsal spines, the diametersof the circles correspond tothe relative size of the spines.Drawings of the corresponding structures modified from Richardson (1909)are included in I,J (the question mark means that the position representing thedrawing is not totally clear) and in M. Abbreviations: sympodite (sy), endopodite (en), and exopodite (ex). Scale bars = 1 mm (A–L), 0.5 mm (M–N); in some cases, a single scale bar applies to more than one figure (indicated by the letters above the scale bar)." figureDoi="http://doi.org/10.5281/zenodo.10716774" httpUri="https://zenodo.org/record/10716774/files/figure.png" pageId="9" pageNumber="10">Figs. 4D, K–L</figureCitation>
, 
<figureCitation id="137B2A78FF0FFFE3FC2EF952FC0BF92A" box="[909,944,1670,1699]" captionStart="Figure 8" captionStartId="15.[181,252,1523,1546]" captionTargetBox="[192,1460,173,1475]" captionTargetId="figure-248@15.[181,1466,151,1499]" captionTargetPageId="15" captionText="Figure 8. Selected body parts of Acanthoniscus for comparison with other taxa within Armadillidae. Cubaris murina Brandt, 1833, showing the ground pattern in Armadillidae (A–E): A, Posterior region of the body in dorsal view; B, posterior region of the body in ventral view; C, pleotelson in dorsal view; D, uropod in dorsal view; E, middle region of the sternite 7 of a male showing the bilobed caudal process on the posterior margin, which is considered an autapomorphy of Armadillidae.Acanthoniscus richardsonae sp. nov.(F–H): F, pleotelson in dorsal view; G, uropod in dorsal view; H, middle region of the sternite 7 of a male.Drawings of the pleotelson and uropods in dorsal views of some morphologically divergent taxa within Armadillidae. I, Echinodillo cavaticus Green, 1963; J, Echinodillo montanus Jackson, 1933; K, Laureola vietnamensis Kwon, Ferrara and Taiti, 1992; L, Laureola dubia Schmalfuss and Ferrara,1983; M, Schismadillo rouxi Verhoeff,1926; N, Merulanella wahrbergi Verhoeff,1926; O, Nesodillo incisus Verhoeff, 1926. Drawings modified from Verhoeff (1926) (M–O), Jackson (1933) (J), Green (1963) (I), Schmalfuss and Ferrara (1983) (L), and Kwon et al. (1992) (K). Abbreviations: endopodite (en), exopodite (ex), sympodite (sy)." figureDoi="http://doi.org/10.5281/zenodo.10716782" httpUri="https://zenodo.org/record/10716782/files/figure.png" pageId="9" pageNumber="10">8F</figureCitation>
). Uropod sympodite “subtrapezoidal,” outer margin widely convex and produced (forming an almost square angle), medial-posterior angle narrow, and approximately at same level or slightly surpassing the distal margin of pleotelson; endopodite surpassing the insertion of exopodite, but not surpassing either the posterior margin of sympodite or the distal margin of pleotelson (
<figureCitation id="137B2A78FF0FFFE3FC5BF842FB3EF83A" box="[1016,1157,1942,1971]" captionStart="Figure 4" captionStartId="11.[181,252,1628,1651]" captionTargetBox="[246,1407,152,1593]" captionTargetId="figure-20@11.[229,1420,148,1608]" captionTargetPageId="11" captionText="Figure 4. Body parts of Acanthoniscus richardsonae sp. nov. A, Overall, dorsal view; B, detail of the microsculpture of the dorsal tegument (left) and tricorn-like setae (right); C, lateral view of the anterior region of the body;D, lateral view of the posterior region of the body; E, right pleon-epimera 3–5 in ventral view; F, right pereon-epimera 1–7 in ventral view; G, detail of pereon-epimera 1–2 in a more ventral position; H, cephalothorax in dorsal view; I, cephalothorax in frontal view (the tiny sensory setae inserted between the ommatidia are illustrated under a magnification of 400×: a, b, c); J, cephalothorax in ventral view; K, pleotelson and uropods in dorsal view (the uropod exopodites are slightly pointing upward); L, pleotelson and uropods in ventral view; M, right uropod in ventral view; N, right uropod in dorsal view; O, drawing of the overall dorsal view modified from Richardson (1909); P, arrangement of dorsal spines, the diametersof the circles correspond tothe relative size of the spines.Drawings of the corresponding structures modified from Richardson (1909)are included in I,J (the question mark means that the position representing thedrawing is not totally clear) and in M. Abbreviations: sympodite (sy), endopodite (en), and exopodite (ex). Scale bars = 1 mm (A–L), 0.5 mm (M–N); in some cases, a single scale bar applies to more than one figure (indicated by the letters above the scale bar)." figureDoi="http://doi.org/10.5281/zenodo.10716774" httpUri="https://zenodo.org/record/10716774/files/figure.png" pageId="9" pageNumber="10">Figs. 4K–N</figureCitation>
, 
<figureCitation id="137B2A78FF0FFFE3FB2DF842FB0CF83A" box="[1166,1207,1942,1971]" captionStart="Figure 8" captionStartId="15.[181,252,1523,1546]" captionTargetBox="[192,1460,173,1475]" captionTargetId="figure-248@15.[181,1466,151,1499]" captionTargetPageId="15" captionText="Figure 8. Selected body parts of Acanthoniscus for comparison with other taxa within Armadillidae. Cubaris murina Brandt, 1833, showing the ground pattern in Armadillidae (A–E): A, Posterior region of the body in dorsal view; B, posterior region of the body in ventral view; C, pleotelson in dorsal view; D, uropod in dorsal view; E, middle region of the sternite 7 of a male showing the bilobed caudal process on the posterior margin, which is considered an autapomorphy of Armadillidae.Acanthoniscus richardsonae sp. nov.(F–H): F, pleotelson in dorsal view; G, uropod in dorsal view; H, middle region of the sternite 7 of a male.Drawings of the pleotelson and uropods in dorsal views of some morphologically divergent taxa within Armadillidae. I, Echinodillo cavaticus Green, 1963; J, Echinodillo montanus Jackson, 1933; K, Laureola vietnamensis Kwon, Ferrara and Taiti, 1992; L, Laureola dubia Schmalfuss and Ferrara,1983; M, Schismadillo rouxi Verhoeff,1926; N, Merulanella wahrbergi Verhoeff,1926; O, Nesodillo incisus Verhoeff, 1926. Drawings modified from Verhoeff (1926) (M–O), Jackson (1933) (J), Green (1963) (I), Schmalfuss and Ferrara (1983) (L), and Kwon et al. (1992) (K). Abbreviations: endopodite (en), exopodite (ex), sympodite (sy)." figureDoi="http://doi.org/10.5281/zenodo.10716782" httpUri="https://zenodo.org/record/10716782/files/figure.png" pageId="9" pageNumber="10">8G</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C35A6576FF0CFFFAFF78FF43FCB1FB7F" lastPageId="16" lastPageNumber="17" pageId="10" pageNumber="11" type="description">
<paragraph id="8BFF36FDFF0CFFE0FF78FF43FC9EFDB3" blockId="10.[179,807,151,1971]" pageId="10" pageNumber="11">
Cephalothorax (
<figureCitation id="137B2A78FF0CFFE0FE17FF43FDDFFF3D" box="[436,612,151,180]" captionStart="Figure 4" captionStartId="11.[181,252,1628,1651]" captionTargetBox="[246,1407,152,1593]" captionTargetId="figure-20@11.[229,1420,148,1608]" captionTargetPageId="11" captionText="Figure 4. Body parts of Acanthoniscus richardsonae sp. nov. A, Overall, dorsal view; B, detail of the microsculpture of the dorsal tegument (left) and tricorn-like setae (right); C, lateral view of the anterior region of the body;D, lateral view of the posterior region of the body; E, right pleon-epimera 3–5 in ventral view; F, right pereon-epimera 1–7 in ventral view; G, detail of pereon-epimera 1–2 in a more ventral position; H, cephalothorax in dorsal view; I, cephalothorax in frontal view (the tiny sensory setae inserted between the ommatidia are illustrated under a magnification of 400×: a, b, c); J, cephalothorax in ventral view; K, pleotelson and uropods in dorsal view (the uropod exopodites are slightly pointing upward); L, pleotelson and uropods in ventral view; M, right uropod in ventral view; N, right uropod in dorsal view; O, drawing of the overall dorsal view modified from Richardson (1909); P, arrangement of dorsal spines, the diametersof the circles correspond tothe relative size of the spines.Drawings of the corresponding structures modified from Richardson (1909)are included in I,J (the question mark means that the position representing thedrawing is not totally clear) and in M. Abbreviations: sympodite (sy), endopodite (en), and exopodite (ex). Scale bars = 1 mm (A–L), 0.5 mm (M–N); in some cases, a single scale bar applies to more than one figure (indicated by the letters above the scale bar)." figureDoi="http://doi.org/10.5281/zenodo.10716774" httpUri="https://zenodo.org/record/10716774/files/figure.png" pageId="10" pageNumber="11">Fig. 4C, H–J</figureCitation>
). Nearly three times (2.7) as wide as long, with rounded lateral lobes, and well-developed, deeply notched median lobe, forming 2 triangular projections slightly bent upward. Dorsal surface with pair of small paramedian spiniform tubercles and 3 medium-sized, club-like, posteromedian spines, one on the median line and one on either side close to the eyes; lateral spines about half length of cephalothorax, median spine smaller than lateral ones. Eyes with tiny scale-setae inserted on inter-ommatidia space (observable only at&gt; 80×).
</paragraph>
<paragraph id="8BFF36FDFF0CFFE0FF78FD90FBFCFB46" blockId="10.[179,807,151,1971]" lastBlockId="10.[840,1468,696,1971]" pageId="10" pageNumber="11">
Pereon (
<figureCitation id="137B2A78FF0CFFE0FE9CFD97FE30FDE8" box="[319,395,579,609]" captionStart="Figure 3" captionStartId="10.[843,913,586,609]" captionTargetBox="[843,1467,151,563]" captionTargetId="figure-639@10.[843,1467,148,565]" captionTargetPageId="10" captionText="Figure 3. Right lateral view of a female paratype Acanthoniscus richardsonaesp.nov.showingcolorationinalcohol.Scale bar=1 mm." figureDoi="http://doi.org/10.5281/zenodo.10716770" httpUri="https://zenodo.org/record/10716770/files/figure.png" pageId="10" pageNumber="11">Figs.3</figureCitation>
, 
<figureCitation id="137B2A78FF0CFFE0FE36FD90FD33FDE9" box="[405,648,579,609]" captionStart="Figure 4" captionStartId="11.[181,252,1628,1651]" captionTargetBox="[246,1407,152,1593]" captionTargetId="figure-20@11.[229,1420,148,1608]" captionTargetPageId="11" captionText="Figure 4. Body parts of Acanthoniscus richardsonae sp. nov. A, Overall, dorsal view; B, detail of the microsculpture of the dorsal tegument (left) and tricorn-like setae (right); C, lateral view of the anterior region of the body;D, lateral view of the posterior region of the body; E, right pleon-epimera 3–5 in ventral view; F, right pereon-epimera 1–7 in ventral view; G, detail of pereon-epimera 1–2 in a more ventral position; H, cephalothorax in dorsal view; I, cephalothorax in frontal view (the tiny sensory setae inserted between the ommatidia are illustrated under a magnification of 400×: a, b, c); J, cephalothorax in ventral view; K, pleotelson and uropods in dorsal view (the uropod exopodites are slightly pointing upward); L, pleotelson and uropods in ventral view; M, right uropod in ventral view; N, right uropod in dorsal view; O, drawing of the overall dorsal view modified from Richardson (1909); P, arrangement of dorsal spines, the diametersof the circles correspond tothe relative size of the spines.Drawings of the corresponding structures modified from Richardson (1909)are included in I,J (the question mark means that the position representing thedrawing is not totally clear) and in M. Abbreviations: sympodite (sy), endopodite (en), and exopodite (ex). Scale bars = 1 mm (A–L), 0.5 mm (M–N); in some cases, a single scale bar applies to more than one figure (indicated by the letters above the scale bar)." figureDoi="http://doi.org/10.5281/zenodo.10716774" httpUri="https://zenodo.org/record/10716774/files/figure.png" pageId="10" pageNumber="11">4A,C–D, F–G, O–P</figureCitation>
). Pereonite 1, 3 times as wide and 1.5times as long as cephalothorax, with 2 very long, slightly sinuous dorsolateral spines (twice the length of pereonite1), 2 pairs of paramedian spiniform tubercles (2 on either side of median line), 3 medium-sized posteromedian spines (0.3 times as long as pereonite 1, 1 on the median line), and smaller lateral spine on either side near posterior margin, directed backward and slightly bent upward; pereonites 2–7 similar, four times as wide as long (pereonite 7 slightly narrower), with 2 very long slightly sinuous dorsolateral spines (twice the length of pereonite 1), pair of paramedian spiniform tubercles (one on either side of median line), 3 medium-sized posteromedian spines, smaller lateral spine on each side near posterior margin, straight and directed backward and slightly outward (gradually increasing in length posteriorly, being lateral spines of seventh tergite about half as long as dorsolateral spines), and spiniform tubercle near anterior margin and directed forward (engaging with previous tergite during conglobation, decreasing in size posteriorly, vestigial on pereonite 7). Epimera 1 with posterior angle and outer margin broadly rounded, anterior angle more acutely produced, dorsal surface concave; epimera 2–3 more or less quadrangular and narrow, with anterior margin concave and posterior margin widely convex, produced distally and then straight toward median region, outer angle acutely produced and directed forward; epimera 4 subtriangular, very narrow and sharp-pointed, and strongly differentiated from epimera 2–3; epimera 5–7 more or less quadrangular, with anterior margin widely convex and produced distally (forming almost square angle), posterior margin concave, and outer angle acutely produced (sharp-pointed) and directed backward. Coxal plates 1–3 with small inner tooth-like process near posterior margin and directed backward, in medial position, forming small schisma and, in coxal plate 1, extending into shallow transverse groove toward base of coxa; coxal plate 4 without tooth-like processes; coxal plates 5–7 with medium-sized tooth-like process, in medial position and near posterior margin, directed backward, forming schisma and extending anteriorly into oblique, very shallow, longitudinal groove, and with very small submarginal tooth-like process, in basal position, just posterior to pereiopod junction, forming second very small schisma; the latter 2 schisma forming continuum engaging with contiguous posterior pereon epimera during conglobation.
</paragraph>
<caption id="DF3F6675FF0CFFE0FCE8FD9EFA07FD08" ID-DOI="http://doi.org/10.5281/zenodo.10716770" ID-Zenodo-Dep="10716770" httpUri="https://zenodo.org/record/10716770/files/figure.png" pageId="10" pageNumber="11" startId="10.[843,913,586,609]" targetBox="[843,1467,151,563]" targetPageId="10" targetType="figure">
<paragraph id="8BFF36FDFF0CFFE0FCE8FD9EFA07FD08" blockId="10.[843,1468,585,641]" pageId="10" pageNumber="11">
<emphasis id="B934EAEFFF0CFFE0FCE8FD9EFC14FDE8" bold="true" box="[843,943,586,609]" pageId="10" pageNumber="11">Figure 3.</emphasis>
Right lateral view of a female paratype 
<taxonomicName id="4C404D7EFF0CFFE0FA96FD9DFC00FD08" authority="Rodríguez-Cabrera &amp; de Armas, 2023" authorityName="Rodríguez-Cabrera &amp; de Armas" authorityYear="2023" class="Malacostraca" family="Delatorreiidae" genus="Acanthoniscus" kingdom="Animalia" order="Isopoda" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="richardsonae" status="sp. nov.">
<emphasis id="B934EAEFFF0CFFE0FA96FD9DFC00FD08" italics="true" pageId="10" pageNumber="11">Acanthoniscus richardsonae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2075794FF0CFFE0FC1DFDBEFBB9FD08" box="[958,1026,618,641]" pageId="10" pageNumber="11" rank="species">sp.nov.</taxonomicNameLabel>
showing coloration in alcohol.Scale bar = 1 mm.
</paragraph>
</caption>
<paragraph id="8BFF36FDFF0CFFE0FCD2FB0DFACCF98F" blockId="10.[840,1468,696,1971]" pageId="10" pageNumber="11">
Pleon (
<figureCitation id="137B2A78FF0CFFE0FC64FB0DFBDDFB7F" box="[967,1126,1241,1270]" captionStart="Figure 4" captionStartId="11.[181,252,1628,1651]" captionTargetBox="[246,1407,152,1593]" captionTargetId="figure-20@11.[229,1420,148,1608]" captionTargetPageId="11" captionText="Figure 4. Body parts of Acanthoniscus richardsonae sp. nov. A, Overall, dorsal view; B, detail of the microsculpture of the dorsal tegument (left) and tricorn-like setae (right); C, lateral view of the anterior region of the body;D, lateral view of the posterior region of the body; E, right pleon-epimera 3–5 in ventral view; F, right pereon-epimera 1–7 in ventral view; G, detail of pereon-epimera 1–2 in a more ventral position; H, cephalothorax in dorsal view; I, cephalothorax in frontal view (the tiny sensory setae inserted between the ommatidia are illustrated under a magnification of 400×: a, b, c); J, cephalothorax in ventral view; K, pleotelson and uropods in dorsal view (the uropod exopodites are slightly pointing upward); L, pleotelson and uropods in ventral view; M, right uropod in ventral view; N, right uropod in dorsal view; O, drawing of the overall dorsal view modified from Richardson (1909); P, arrangement of dorsal spines, the diametersof the circles correspond tothe relative size of the spines.Drawings of the corresponding structures modified from Richardson (1909)are included in I,J (the question mark means that the position representing thedrawing is not totally clear) and in M. Abbreviations: sympodite (sy), endopodite (en), and exopodite (ex). Scale bars = 1 mm (A–L), 0.5 mm (M–N); in some cases, a single scale bar applies to more than one figure (indicated by the letters above the scale bar)." figureDoi="http://doi.org/10.5281/zenodo.10716774" httpUri="https://zenodo.org/record/10716774/files/figure.png" pageId="10" pageNumber="11">Fig. 4A, D–E</figureCitation>
). Tergites 1, 2, and 5 without spines; tergites 3–4 with pair of small posteromedian spines (one on either side of the median line) as long as pleonite length; epimera of pleonites 3–5 acutely produced, similar in shape and size, with anterior margin widely convex and produced medially(forming almost square angle), posterior margin straight, and outer angle acutely produced (sharp-pointed).
</paragraph>
<paragraph id="8BFF36FDFF0CFFE0FCD2F9C4FAA6F979" blockId="10.[840,1468,696,1971]" pageId="10" pageNumber="11">
Pleotelson(
<figureCitation id="137B2A78FF0CFFE0FC5FF9C4FB26F9A4" box="[1020,1181,1552,1582]" captionStart="Figure 4" captionStartId="11.[181,252,1628,1651]" captionTargetBox="[246,1407,152,1593]" captionTargetId="figure-20@11.[229,1420,148,1608]" captionTargetPageId="11" captionText="Figure 4. Body parts of Acanthoniscus richardsonae sp. nov. A, Overall, dorsal view; B, detail of the microsculpture of the dorsal tegument (left) and tricorn-like setae (right); C, lateral view of the anterior region of the body;D, lateral view of the posterior region of the body; E, right pleon-epimera 3–5 in ventral view; F, right pereon-epimera 1–7 in ventral view; G, detail of pereon-epimera 1–2 in a more ventral position; H, cephalothorax in dorsal view; I, cephalothorax in frontal view (the tiny sensory setae inserted between the ommatidia are illustrated under a magnification of 400×: a, b, c); J, cephalothorax in ventral view; K, pleotelson and uropods in dorsal view (the uropod exopodites are slightly pointing upward); L, pleotelson and uropods in ventral view; M, right uropod in ventral view; N, right uropod in dorsal view; O, drawing of the overall dorsal view modified from Richardson (1909); P, arrangement of dorsal spines, the diametersof the circles correspond tothe relative size of the spines.Drawings of the corresponding structures modified from Richardson (1909)are included in I,J (the question mark means that the position representing thedrawing is not totally clear) and in M. Abbreviations: sympodite (sy), endopodite (en), and exopodite (ex). Scale bars = 1 mm (A–L), 0.5 mm (M–N); in some cases, a single scale bar applies to more than one figure (indicated by the letters above the scale bar)." figureDoi="http://doi.org/10.5281/zenodo.10716774" httpUri="https://zenodo.org/record/10716774/files/figure.png" pageId="10" pageNumber="11">Figs.4D, K–L</figureCitation>
, 
<figureCitation id="137B2A78FF0CFFE0FB05F9C5FB73F9A7" box="[1190,1224,1553,1582]" captionStart="Figure 8" captionStartId="15.[181,252,1523,1546]" captionTargetBox="[192,1460,173,1475]" captionTargetId="figure-248@15.[181,1466,151,1499]" captionTargetPageId="15" captionText="Figure 8. Selected body parts of Acanthoniscus for comparison with other taxa within Armadillidae. Cubaris murina Brandt, 1833, showing the ground pattern in Armadillidae (A–E): A, Posterior region of the body in dorsal view; B, posterior region of the body in ventral view; C, pleotelson in dorsal view; D, uropod in dorsal view; E, middle region of the sternite 7 of a male showing the bilobed caudal process on the posterior margin, which is considered an autapomorphy of Armadillidae.Acanthoniscus richardsonae sp. nov.(F–H): F, pleotelson in dorsal view; G, uropod in dorsal view; H, middle region of the sternite 7 of a male.Drawings of the pleotelson and uropods in dorsal views of some morphologically divergent taxa within Armadillidae. I, Echinodillo cavaticus Green, 1963; J, Echinodillo montanus Jackson, 1933; K, Laureola vietnamensis Kwon, Ferrara and Taiti, 1992; L, Laureola dubia Schmalfuss and Ferrara,1983; M, Schismadillo rouxi Verhoeff,1926; N, Merulanella wahrbergi Verhoeff,1926; O, Nesodillo incisus Verhoeff, 1926. Drawings modified from Verhoeff (1926) (M–O), Jackson (1933) (J), Green (1963) (I), Schmalfuss and Ferrara (1983) (L), and Kwon et al. (1992) (K). Abbreviations: endopodite (en), exopodite (ex), sympodite (sy)." figureDoi="http://doi.org/10.5281/zenodo.10716782" httpUri="https://zenodo.org/record/10716782/files/figure.png" pageId="10" pageNumber="11">8F</figureCitation>
). Basal part 1.7 times wider than distal part, 1.4 times as wide as long, with both dorsal spines 1.3–1.5 times as long as pleotelson; distal margin ending in 2 small triangular projections separated by shallow notch (0.3 times as deep as the median diameter of pleotelson spines).
</paragraph>
<paragraph id="8BFF36FDFF0CFFE6FCD2F92EFD28FE17" blockId="10.[840,1468,696,1971]" lastBlockId="12.[180,807,151,414]" lastPageId="12" lastPageNumber="13" pageId="10" pageNumber="11">
Uropods(
<figureCitation id="137B2A78FF0CFFE0FC42F92EFBE7F89E" box="[993,1116,1786,1815]" captionStart="Figure 4" captionStartId="11.[181,252,1628,1651]" captionTargetBox="[246,1407,152,1593]" captionTargetId="figure-20@11.[229,1420,148,1608]" captionTargetPageId="11" captionText="Figure 4. Body parts of Acanthoniscus richardsonae sp. nov. A, Overall, dorsal view; B, detail of the microsculpture of the dorsal tegument (left) and tricorn-like setae (right); C, lateral view of the anterior region of the body;D, lateral view of the posterior region of the body; E, right pleon-epimera 3–5 in ventral view; F, right pereon-epimera 1–7 in ventral view; G, detail of pereon-epimera 1–2 in a more ventral position; H, cephalothorax in dorsal view; I, cephalothorax in frontal view (the tiny sensory setae inserted between the ommatidia are illustrated under a magnification of 400×: a, b, c); J, cephalothorax in ventral view; K, pleotelson and uropods in dorsal view (the uropod exopodites are slightly pointing upward); L, pleotelson and uropods in ventral view; M, right uropod in ventral view; N, right uropod in dorsal view; O, drawing of the overall dorsal view modified from Richardson (1909); P, arrangement of dorsal spines, the diametersof the circles correspond tothe relative size of the spines.Drawings of the corresponding structures modified from Richardson (1909)are included in I,J (the question mark means that the position representing thedrawing is not totally clear) and in M. Abbreviations: sympodite (sy), endopodite (en), and exopodite (ex). Scale bars = 1 mm (A–L), 0.5 mm (M–N); in some cases, a single scale bar applies to more than one figure (indicated by the letters above the scale bar)." figureDoi="http://doi.org/10.5281/zenodo.10716774" httpUri="https://zenodo.org/record/10716774/files/figure.png" pageId="10" pageNumber="11">Figs.4K–N</figureCitation>
, 
<figureCitation id="137B2A78FF0CFFE0FBC1F92EFB32F89E" box="[1122,1161,1786,1815]" captionStart="Figure 8" captionStartId="15.[181,252,1523,1546]" captionTargetBox="[192,1460,173,1475]" captionTargetId="figure-248@15.[181,1466,151,1499]" captionTargetPageId="15" captionText="Figure 8. Selected body parts of Acanthoniscus for comparison with other taxa within Armadillidae. Cubaris murina Brandt, 1833, showing the ground pattern in Armadillidae (A–E): A, Posterior region of the body in dorsal view; B, posterior region of the body in ventral view; C, pleotelson in dorsal view; D, uropod in dorsal view; E, middle region of the sternite 7 of a male showing the bilobed caudal process on the posterior margin, which is considered an autapomorphy of Armadillidae.Acanthoniscus richardsonae sp. nov.(F–H): F, pleotelson in dorsal view; G, uropod in dorsal view; H, middle region of the sternite 7 of a male.Drawings of the pleotelson and uropods in dorsal views of some morphologically divergent taxa within Armadillidae. I, Echinodillo cavaticus Green, 1963; J, Echinodillo montanus Jackson, 1933; K, Laureola vietnamensis Kwon, Ferrara and Taiti, 1992; L, Laureola dubia Schmalfuss and Ferrara,1983; M, Schismadillo rouxi Verhoeff,1926; N, Merulanella wahrbergi Verhoeff,1926; O, Nesodillo incisus Verhoeff, 1926. Drawings modified from Verhoeff (1926) (M–O), Jackson (1933) (J), Green (1963) (I), Schmalfuss and Ferrara (1983) (L), and Kwon et al. (1992) (K). Abbreviations: endopodite (en), exopodite (ex), sympodite (sy)." figureDoi="http://doi.org/10.5281/zenodo.10716782" httpUri="https://zenodo.org/record/10716782/files/figure.png" pageId="10" pageNumber="11">8G</figureCitation>
).Sympoditesubtrapezoidal, 1.3times as long as wide, ventral surface flat, outer margin widely convex and produced (forming almost square angle), medial-posterior angle narrow and approximately at same level, or slightly surpassing, distal margin of pleotelson, medial margin with very small inner lobe above and basal with respect to insertion of exopodite; endopodite surpassing insertion of exopodite, but not surpassing either posterior margin of sympodite or distal margin of pleotelson (not visible dorsally); exopodite 1.1 times as long as sympodite and surpassing posterior margin of sympodite by 60% of its length.
</paragraph>
<caption id="DF3F6675FF0DFFE1FF16F988FAD0F83A" ID-DOI="http://doi.org/10.5281/zenodo.10716774" ID-Zenodo-Dep="10716774" httpUri="https://zenodo.org/record/10716774/files/figure.png" pageId="11" pageNumber="12" startId="11.[181,252,1628,1651]" targetBox="[246,1407,152,1593]" targetPageId="11" targetType="figure">
<paragraph id="8BFF36FDFF0DFFE1FF16F988FAD0F83A" blockId="11.[180,1467,1627,1971]" pageId="11" pageNumber="12">
<emphasis id="B934EAEFFF0DFFE1FF16F988FEA0F9FA" bold="true" box="[181,283,1628,1651]" pageId="11" pageNumber="12">Figure 4.</emphasis>
Body parts of 
<taxonomicName id="4C404D7EFF0DFFE1FE13F98FFD03F9FA" authority="Rodríguez-Cabrera &amp; de Armas, 2023" authorityName="Rodríguez-Cabrera &amp; de Armas" authorityYear="2023" box="[432,696,1627,1651]" class="Malacostraca" family="Delatorreiidae" genus="Acanthoniscus" kingdom="Animalia" order="Isopoda" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="richardsonae" status="sp. nov.">
<emphasis id="B934EAEFFF0DFFE1FE13F98FFD03F9FA" box="[432,696,1627,1651]" italics="true" pageId="11" pageNumber="12">Acanthoniscus richardsonae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2075794FF0DFFE1FD1DF988FCB1F9FA" box="[702,778,1628,1651]" pageId="11" pageNumber="12" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="B934EAEFFF0DFFE1FCACF98FFCA5F9FA" bold="true" box="[783,798,1627,1651]" pageId="11" pageNumber="12">A</emphasis>
, Overall, dorsal view; 
<emphasis id="B934EAEFFF0DFFE1FBA2F98FFBA9F9FA" bold="true" box="[1025,1042,1627,1651]" pageId="11" pageNumber="12">B</emphasis>
, detail of the microsculpture of the dorsal tegument (left) and tricorn-like setae (right); 
<emphasis id="B934EAEFFF0DFFE1FDD6F9AFFD3CF91A" bold="true" box="[629,647,1659,1683]" pageId="11" pageNumber="12">C</emphasis>
, lateral view of the anterior region of the body; 
<emphasis id="B934EAEFFF0DFFE1FBF3F9AFFBD8F91A" bold="true" box="[1104,1123,1659,1683]" pageId="11" pageNumber="12">D</emphasis>
, lateral view of the posterior region of the body; 
<emphasis id="B934EAEFFF0DFFE1FE97F94FFEFFF93A" bold="true" box="[308,324,1691,1715]" pageId="11" pageNumber="12">E</emphasis>
, right pleon-epimera 3–5 in ventral view; 
<emphasis id="B934EAEFFF0DFFE1FD45F94FFD4EF93A" bold="true" box="[742,757,1691,1715]" pageId="11" pageNumber="12">F</emphasis>
, right pereon-epimera 1–7 in ventral view; 
<emphasis id="B934EAEFFF0DFFE1FB04F94FFB01F93A" bold="true" box="[1191,1210,1691,1715]" pageId="11" pageNumber="12">G</emphasis>
, detail of pereon-epimera 1–2 in a more ventral position; 
<emphasis id="B934EAEFFF0DFFE1FE50F96FFDBCF95A" bold="true" box="[499,519,1723,1747]" pageId="11" pageNumber="12">H</emphasis>
, cephalothorax in dorsal view; 
<emphasis id="B934EAEFFF0DFFE1FCE3F96FFCF1F95A" bold="true" box="[832,842,1723,1747]" pageId="11" pageNumber="12">I</emphasis>
, cephalothorax in frontal view (the tiny sensory setae inserted between the ommatidia are illustrated under a magnification of 400×: a, b, c); 
<emphasis id="B934EAEFFF0DFFE1FC61F90FFC77F97A" bold="true" box="[962,972,1755,1779]" pageId="11" pageNumber="12">J</emphasis>
, cephalothorax in ventral view; 
<emphasis id="B934EAEFFF0DFFE1FAA9F90FFAA7F97A" bold="true" box="[1290,1308,1755,1779]" pageId="11" pageNumber="12">K</emphasis>
, pleotelson and uropods in dorsal view (the uropod exopodites are slightly pointing upward); 
<emphasis id="B934EAEFFF0DFFE1FC1DF92FFC76F89A" bold="true" box="[958,973,1787,1811]" pageId="11" pageNumber="12">L</emphasis>
, pleotelson and uropods in ventral view; 
<emphasis id="B934EAEFFF0DFFE1FACAF92FFAC4F89A" bold="true" box="[1385,1407,1787,1811]" pageId="11" pageNumber="12">M</emphasis>
, right uropod in ventral view; 
<emphasis id="B934EAEFFF0DFFE1FE02F8CFFE0FF8BA" bold="true" box="[417,436,1819,1843]" pageId="11" pageNumber="12">N</emphasis>
, right uropod in dorsal view; 
<emphasis id="B934EAEFFF0DFFE1FD75F8CFFD51F8BA" bold="true" box="[726,746,1819,1843]" pageId="11" pageNumber="12">O</emphasis>
, drawing of the overall dorsal view modified from 
<bibRefCitation id="EFD14B0CFF0DFFE1FB7AF8CFFA27F8BA" author="Richardson H" box="[1241,1436,1819,1843]" pageId="11" pageNumber="12" pagination="431 - 434" refId="ref17042" refString="Richardson H 1909. The isopod crustacean Acanthoniscus spiniger Kinahan redescribed. Proceedings of the United States National Museum, 36: 431 - 434. DOI: https: // www. doi. org / 10.5479 / si. 00963801.36 - 1678.431" type="journal article" year="1909">Richardson (1909)</bibRefCitation>
; 
<emphasis id="B934EAEFFF0DFFE1FA06F8CFFA0EF8BA" bold="true" box="[1445,1461,1819,1843]" pageId="11" pageNumber="12">P</emphasis>
, arrangement of dorsal spines, the diameters of the circles correspond to the relative size of the spines. Drawings of the corresponding structures modified from 
<bibRefCitation id="EFD14B0CFF0DFFE1FE0FF88FFDD1F8FA" author="Richardson H" box="[428,618,1883,1907]" pageId="11" pageNumber="12" pagination="431 - 434" refId="ref17042" refString="Richardson H 1909. The isopod crustacean Acanthoniscus spiniger Kinahan redescribed. Proceedings of the United States National Museum, 36: 431 - 434. DOI: https: // www. doi. org / 10.5479 / si. 00963801.36 - 1678.431" type="journal article" year="1909">Richardson (1909)</bibRefCitation>
are included in 
<emphasis id="B934EAEFFF0DFFE1FCA3F88FFCA5F8FA" bold="true" box="[768,798,1883,1907]" pageId="11" pageNumber="12">I, J</emphasis>
(the question mark means that the position representing the drawing is not totally clear) and in 
<emphasis id="B934EAEFFF0DFFE1FE19F8AFFE6BF81A" bold="true" box="[442,464,1915,1939]" pageId="11" pageNumber="12">M</emphasis>
. Abbreviations: sympodite (
<emphasis id="B934EAEFFF0DFFE1FD4CF8A8FCBAF81A" box="[751,769,1916,1939]" italics="true" pageId="11" pageNumber="12">sy</emphasis>
), endopodite (
<emphasis id="B934EAEFFF0DFFE1FC38F8A8FC0AF81A" box="[923,945,1916,1939]" italics="true" pageId="11" pageNumber="12">en</emphasis>
), and exopodite (
<emphasis id="B934EAEFFF0DFFE1FBC6F8A8FBC1F81A" box="[1125,1146,1916,1939]" italics="true" pageId="11" pageNumber="12">ex</emphasis>
). Scale bars = 1 mm (
<emphasis id="B934EAEFFF0DFFE1FAFAF8AFFA3DF81A" bold="true" box="[1369,1414,1915,1939]" pageId="11" pageNumber="12">A–L</emphasis>
), 0.5 mm (
<emphasis id="B934EAEFFF0DFFE1FF4EF84FFE9FF83A" bold="true" box="[237,292,1947,1971]" pageId="11" pageNumber="12">M–N</emphasis>
); in some cases, a single scale bar applies to more than one figure (indicated by the letters above the scale bar).
</paragraph>
</caption>
<paragraph id="8BFF36FDFF0AFFE7FCD2FF43FD31FDB0" blockId="12.[841,1468,151,414]" lastBlockId="13.[180,807,151,764]" lastPageId="13" lastPageNumber="14" pageId="12" pageNumber="13">
Antennae (
<figureCitation id="137B2A78FF0AFFE6FC50FF43FBFCFF3D" box="[1011,1095,151,180]" captionStart="Figure 4" captionStartId="11.[181,252,1628,1651]" captionTargetBox="[246,1407,152,1593]" captionTargetId="figure-20@11.[229,1420,148,1608]" captionTargetPageId="11" captionText="Figure 4. Body parts of Acanthoniscus richardsonae sp. nov. A, Overall, dorsal view; B, detail of the microsculpture of the dorsal tegument (left) and tricorn-like setae (right); C, lateral view of the anterior region of the body;D, lateral view of the posterior region of the body; E, right pleon-epimera 3–5 in ventral view; F, right pereon-epimera 1–7 in ventral view; G, detail of pereon-epimera 1–2 in a more ventral position; H, cephalothorax in dorsal view; I, cephalothorax in frontal view (the tiny sensory setae inserted between the ommatidia are illustrated under a magnification of 400×: a, b, c); J, cephalothorax in ventral view; K, pleotelson and uropods in dorsal view (the uropod exopodites are slightly pointing upward); L, pleotelson and uropods in ventral view; M, right uropod in ventral view; N, right uropod in dorsal view; O, drawing of the overall dorsal view modified from Richardson (1909); P, arrangement of dorsal spines, the diametersof the circles correspond tothe relative size of the spines.Drawings of the corresponding structures modified from Richardson (1909)are included in I,J (the question mark means that the position representing thedrawing is not totally clear) and in M. Abbreviations: sympodite (sy), endopodite (en), and exopodite (ex). Scale bars = 1 mm (A–L), 0.5 mm (M–N); in some cases, a single scale bar applies to more than one figure (indicated by the letters above the scale bar)." figureDoi="http://doi.org/10.5281/zenodo.10716774" httpUri="https://zenodo.org/record/10716774/files/figure.png" pageId="12" pageNumber="13">Figs.4J</figureCitation>
, 
<figureCitation id="137B2A78FF0AFFE6FBF3FF43FB20FF3D" box="[1104,1179,151,180]" captionStart="Figure 5" captionStartId="12.[182,251,1692,1715]" captionTargetBox="[312,1351,473,1658]" captionTargetId="figure-134@12.[266,1382,465,1671]" captionTargetPageId="12" captionText="Figure 5.Otherappendagesof Acanthoniscus richardsonae sp.nov.A, Second antenna;B,detail offlagellum of secondantennadepicting position of aesthetasc groups (small arrows); C, detail of the distal article of the flagellum of the second antenna slightly turned over showing the aesthetasc groups; D, detail of the distal-most portion of the flagellum of the second antenna showing the apical cone; E, first antenna; F, detail of the third article of the first antenna showing the aesthetascs; G, male left seventh pereiopod; H, detail of ischium ofmale seventh pereiopod; I, detailof dactylus ofmale seventh pereiopodin anterior view; J, detailof dactylus ofmale seventh pereiopod in posterior view; K, male left first pereiopod, showing the areas on the carpus and propodus covered with the cleaning apparatus or antennal brush (dashed outlines); L, detail of carpus ofmale first pereiopod (anterior surface).Abbreviations:basis (bs), ischium(is),merus(me),carpus (ca),propodus (pr),flagellum(fl),and dactylus(da).Drawingofthe corresponding structuremodified from Richardson (1909) is included inA. Scale bars = 1 mm (A), 0.5 mm (G,K), 0.3mm (B), 0.1 mm (C–E, H–J,L),0.05 mm (F)." figureDoi="http://doi.org/10.5281/zenodo.10716776" httpUri="https://zenodo.org/record/10716776/files/figure.png" pageId="12" pageNumber="13">5A–D</figureCitation>
). First antenna 3-jointed; first article thick, sub-cylindrical, twice the length of second article and 1.6 times as long as third article; second article shortest and sub-conical in shape; third article narrow and asymmetric, slightly curved, with row of eight aesthetascs in subapical position.Second antenna flagellum with distal article 3 times longer than basal article, with aesthetasc group on basal-most quarter and other near middle region (formula: 1–3), and tuft of 3 small setae near distal end, apical cone as long as basal article of flagellum and hence 3 times shorter than distal article, with 2 long lateral free sensilla inserted on basal-most quarter and nearly reaching tip of cone; ischium nearly as long as merus, carpus 1.3 times longer than merus, propodus 1.5 times longer than carpus and 1.3 times longer than flagellum; ischium, merus, carpus,and propodus with deep groove along entire inner surface.
</paragraph>
<caption id="DF3F6675FF0AFFE6FF15F948FA00F83A" ID-DOI="http://doi.org/10.5281/zenodo.10716776" ID-Zenodo-Dep="10716776" httpUri="https://zenodo.org/record/10716776/files/figure.png" pageId="12" pageNumber="13" startId="12.[182,251,1692,1715]" targetBox="[312,1351,473,1658]" targetPageId="12" targetType="figure">
<paragraph id="8BFF36FDFF0AFFE6FF15F948FA00F83A" blockId="12.[180,1467,1691,1971]" pageId="12" pageNumber="13">
<emphasis id="B934EAEFFF0AFFE6FF15F948FEAFF93A" bold="true" box="[182,276,1692,1715]" pageId="12" pageNumber="13">Figure 5.</emphasis>
Other appendages of 
<taxonomicName id="4C404D7EFF0AFFE6FE7CF94FFD60F93A" authority="Rodríguez-Cabrera &amp; de Armas, 2023" authorityName="Rodríguez-Cabrera &amp; de Armas" authorityYear="2023" box="[479,731,1691,1715]" class="Malacostraca" family="Delatorreiidae" genus="Acanthoniscus" kingdom="Animalia" order="Isopoda" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="richardsonae" status="sp. nov.">
<emphasis id="B934EAEFFF0AFFE6FE7CF94FFD60F93A" box="[479,731,1691,1715]" italics="true" pageId="12" pageNumber="13">Acanthoniscus richardsonae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2075794FF0AFFE6FD7CF948FC9EF93A" box="[735,805,1692,1715]" pageId="12" pageNumber="13" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="B934EAEFFF0AFFE6FC8BF94FFC82F93A" bold="true" box="[808,825,1691,1715]" pageId="12" pageNumber="13">A</emphasis>
, Second antenna; 
<emphasis id="B934EAEFFF0AFFE6FC44F94FFC4CF93A" bold="true" box="[999,1015,1691,1715]" pageId="12" pageNumber="13">B</emphasis>
, detail of flagellum of second antenna depicting position of aesthetasc groups (small arrows); 
<emphasis id="B934EAEFFF0AFFE6FDD3F96FFD39F95A" bold="true" box="[624,642,1723,1747]" pageId="12" pageNumber="13">C</emphasis>
, detail of the distal article of the flagellum of the second antenna slightly turned over showing the aesthetasc groups; 
<emphasis id="B934EAEFFF0AFFE6FE4EF90FFDBBF97A" bold="true" box="[493,512,1755,1779]" pageId="12" pageNumber="13">D</emphasis>
, detail of the distal-most portion of the flagellum of the second antenna showing the apical cone; 
<emphasis id="B934EAEFFF0AFFE6FF16F92FFF7FF89A" bold="true" box="[181,196,1787,1811]" pageId="12" pageNumber="13">E</emphasis>
, first antenna; 
<emphasis id="B934EAEFFF0AFFE6FEF5F92FFEDEF89A" bold="true" box="[342,357,1787,1811]" pageId="12" pageNumber="13">F</emphasis>
, detail of the third article of the first antenna showing the aesthetascs; 
<emphasis id="B934EAEFFF0AFFE6FBB0F92FFB9DF89A" bold="true" box="[1043,1062,1787,1811]" pageId="12" pageNumber="13">G</emphasis>
, male left seventh pereiopod; 
<emphasis id="B934EAEFFF0AFFE6FAEFF92FFADBF89A" bold="true" box="[1356,1376,1787,1811]" pageId="12" pageNumber="13">H</emphasis>
, detail of ischium of male seventh pereiopod; 
<emphasis id="B934EAEFFF0AFFE6FDB3F8CFFDA1F8BA" bold="true" box="[528,538,1819,1843]" pageId="12" pageNumber="13">I</emphasis>
, detail of dactylus of male seventh pereiopod in anterior view; 
<emphasis id="B934EAEFFF0AFFE6FBCFF8CFFBCEF8BA" bold="true" box="[1132,1141,1819,1843]" pageId="12" pageNumber="13">J</emphasis>
, detail of dactylus of male seventh pereiopod in posterior view; 
<emphasis id="B934EAEFFF0AFFE6FE77F8EFFE5DF8DA" bold="true" box="[468,486,1851,1875]" pageId="12" pageNumber="13">K</emphasis>
, male left first pereiopod, showing the areas on the carpus and propodus covered with the cleaning apparatus or antennal brush (dashed outlines); 
<emphasis id="B934EAEFFF0AFFE6FDDCF88FFD35F8FA" bold="true" box="[639,654,1883,1907]" pageId="12" pageNumber="13">L</emphasis>
, detail of carpus of male first pereiopod (anterior surface). Abbreviations: basis (
<emphasis id="B934EAEFFF0AFFE6FA34F88FFA10F8FA" box="[1431,1451,1883,1907]" italics="true" pageId="12" pageNumber="13">bs</emphasis>
), ischium (
<emphasis id="B934EAEFFF0AFFE6FEB3F8A8FEA5F81A" box="[272,286,1916,1939]" italics="true" pageId="12" pageNumber="13">is</emphasis>
), merus (
<emphasis id="B934EAEFFF0AFFE6FED8F8A8FE2DF81A" box="[379,406,1916,1939]" italics="true" pageId="12" pageNumber="13">me</emphasis>
), carpus (
<emphasis id="B934EAEFFF0AFFE6FE54F8A8FDB0F81A" box="[503,523,1916,1939]" italics="true" pageId="12" pageNumber="13">ca</emphasis>
), propodus (
<emphasis id="B934EAEFFF0AFFE6FD2AF8A8FD26F81A" box="[649,669,1916,1939]" italics="true" pageId="12" pageNumber="13">pr</emphasis>
), flagellum (
<emphasis id="B934EAEFFF0AFFE6FCBBF8AFFC9FF81A" box="[792,804,1915,1939]" italics="true" pageId="12" pageNumber="13">fl</emphasis>
), and dactylus (
<emphasis id="B934EAEFFF0AFFE6FC1EF8AFFC6FF81A" box="[957,980,1915,1939]" italics="true" pageId="12" pageNumber="13">da</emphasis>
). Drawing of the corresponding structure modified from 
<bibRefCitation id="EFD14B0CFF0AFFE6FF4AF84FFE1DF83A" author="Richardson H" box="[233,422,1947,1971]" pageId="12" pageNumber="13" pagination="431 - 434" refId="ref17042" refString="Richardson H 1909. The isopod crustacean Acanthoniscus spiniger Kinahan redescribed. Proceedings of the United States National Museum, 36: 431 - 434. DOI: https: // www. doi. org / 10.5479 / si. 00963801.36 - 1678.431" type="journal article" year="1909">Richardson (1909)</bibRefCitation>
is included in 
<emphasis id="B934EAEFFF0AFFE6FD93F84FFDFAF83A" bold="true" box="[560,577,1947,1971]" pageId="12" pageNumber="13">A</emphasis>
. Scale bars = 1 mm (
<emphasis id="B934EAEFFF0AFFE6FCA8F84FFCA7F83A" bold="true" box="[779,796,1947,1971]" pageId="12" pageNumber="13">A</emphasis>
), 0.5 mm (
<emphasis id="B934EAEFFF0AFFE6FC2AF84FFC03F83A" bold="true" box="[905,952,1947,1971]" pageId="12" pageNumber="13">G, K</emphasis>
), 0.3 mm (
<emphasis id="B934EAEFFF0AFFE6FB85F84FFB8CF83A" bold="true" box="[1062,1079,1947,1971]" pageId="12" pageNumber="13">B</emphasis>
), 0.1 mm (
<emphasis id="B934EAEFFF0AFFE6FB06F84FFA9AF83A" bold="true" box="[1189,1313,1947,1971]" pageId="12" pageNumber="13">C–E, H–J, L</emphasis>
), 0.05 mm (
<emphasis id="B934EAEFFF0AFFE6FA3FF84FFA10F83A" bold="true" box="[1436,1451,1947,1971]" pageId="12" pageNumber="13">F</emphasis>
).
</paragraph>
</caption>
<paragraph id="8BFF36FDFF0BFFE4FF78FD97FD96FCF8" blockId="13.[180,807,151,764]" lastBlockId="14.[180,808,151,881]" lastPageId="14" lastPageNumber="15" pageId="13" pageNumber="14">
Mouthparts (
<figureCitation id="137B2A78FF0BFFE7FED4FD97FE03FDE9" box="[375,440,579,608]" captionStart="Figure 6" captionStartId="13.[181,252,1788,1811]" captionTargetBox="[267,1379,821,1745]" captionTargetId="figure-164@13.[232,1415,817,1764]" captionTargetPageId="13" captionText="Figure 6.Mouthparts of Acanthoniscus richardsonae sp. nov. A, Distal portion of right (R) and left (L) mandibles in anterior views, showing lacinia mobilis (lm), hairy lobe (hl), penicils (pn), and tuft of setae (ts); B, first maxilla, with detailof distalportion of outer endite (ex) and inner endite (en); C, second maxilla; D, detail of the distal-most portion of the second maxilla; E, hypopharynx; F, detail of the median region of hypopharynx; G, maxilliped; H, detail of the distal-most portion of maxilliped in posterior view; I, detail of the distal-most portion of maxilliped inanterior view.Drawings of the corresponding structures modified from Richardson (1909) are included in B, C, and G. Scale bars = 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.10716778" httpUri="https://zenodo.org/record/10716778/files/figure.png" pageId="13" pageNumber="14">Fig.6</figureCitation>
). Left mandible in anterior view (in distal-proximal order) with a narrow quadricuspid incisor process, lacinia mobilis bicuspid with smaller inner lobe, hairy lobe bearing 2 penicils, 1 free penicil, and tuft of hairy setae as molar process.Right mandible in anterior view (in distal-proximal order) with wider quadricuspid incisor process, oval hyaline process, hairy lobe bearing 2 penicils, and tuft of hairy setae as molar process. Hypopharynx with middle lobe notched at tip, with groups of relatively large setae along outer margin of lateral lobes, inner margin of lateral lobes densely hirsute. First maxilla outer endite with outer margin deeply curved distally and with marginal fringe of small setae and 10 simple tooth-like setae of approximately equal in length: outer group composed of 4 strong setae (lateral-most considerably stronger) and small triangular lobe, inner group composed of 6 slender setae, with very small subapical seta between both groups of tooth setae; inner endite with 2 subapical thickset penicils and slightly produced, rounded laterodistal corner. Second maxilla outer lobe wider and sensilla scattered, with outer notch; inner lobe narrower, with sensilla concentrated in dense field on mediodistal corner; with 2–3 differentiated setae in incision between both lobes, on medial margin of inner lobe. Maxilliped basipodite with nearly parallel outer and inner margins and wide lamella on distal portion of outer margin, with scaly surface and scattered small setae; endite apically truncate, with 3 enlarged subapical setae on posterior surface and outer corner with 2 small rounded lobes, one of them ending in tiny buttonlike seta; palpus with middle and distal article fused; basal article bearing 2 strong setae, outer one nearly reaching distal margin of endite; middle article with 2 “tufts” on medial margin bearing small single seta each (basal-most one on produced lobe), and single seta on outer-distal margin; apical article narrow and bent inward, ending in tuft of setae and with tiny subapical seta on outer margin.
</paragraph>
<caption id="DF3F6675FF0BFFE7FF16F928FD4EF83A" ID-DOI="http://doi.org/10.5281/zenodo.10716778" ID-Zenodo-Dep="10716778" httpUri="https://zenodo.org/record/10716778/files/figure.png" pageId="13" pageNumber="14" startId="13.[181,252,1788,1811]" targetBox="[267,1379,821,1745]" targetPageId="13" targetType="figure">
<paragraph id="8BFF36FDFF0BFFE7FF16F928FD4EF83A" blockId="13.[180,1467,1787,1971]" pageId="13" pageNumber="14">
<emphasis id="B934EAEFFF0BFFE7FF16F928FEA3F89A" bold="true" box="[181,280,1788,1811]" pageId="13" pageNumber="14">Figure 6.</emphasis>
Mouthparts of 
<taxonomicName id="4C404D7EFF0BFFE7FE13F92FFD0EF89A" authority="Rodríguez-Cabrera &amp; de Armas, 2023" authorityName="Rodríguez-Cabrera &amp; de Armas" authorityYear="2023" box="[432,693,1787,1811]" class="Malacostraca" family="Delatorreiidae" genus="Acanthoniscus" kingdom="Animalia" order="Isopoda" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="richardsonae" status="sp. nov.">
<emphasis id="B934EAEFFF0BFFE7FE13F92FFD0EF89A" box="[432,693,1787,1811]" italics="true" pageId="13" pageNumber="14">Acanthoniscus richardsonae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2075794FF0BFFE7FD19F928FCBFF89A" box="[698,772,1788,1811]" pageId="13" pageNumber="14" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="B934EAEFFF0BFFE7FCABF92FFCA2F89A" bold="true" box="[776,793,1787,1811]" pageId="13" pageNumber="14">A</emphasis>
, Distal portion of right (
<emphasis id="B934EAEFFF0BFFE7FBAFF928FBA0F89A" box="[1036,1051,1788,1811]" italics="true" pageId="13" pageNumber="14">R</emphasis>
) and left (
<emphasis id="B934EAEFFF0BFFE7FB25F928FB2FF89A" box="[1158,1172,1788,1811]" italics="true" pageId="13" pageNumber="14">L</emphasis>
) mandibles in anterior views, showing lacinia mobilis (
<emphasis id="B934EAEFFF0BFFE7FE08F8CFFE7FF8BA" box="[427,452,1819,1843]" italics="true" pageId="13" pageNumber="14">lm</emphasis>
), hairy lobe (
<emphasis id="B934EAEFFF0BFFE7FDEAF8CFFDE7F8BA" box="[585,604,1819,1843]" italics="true" pageId="13" pageNumber="14">hl</emphasis>
), penicils (
<emphasis id="B934EAEFFF0BFFE7FD68F8C8FD5FF8BA" box="[715,740,1820,1843]" italics="true" pageId="13" pageNumber="14">pn</emphasis>
), and tuft of setae (ts); 
<emphasis id="B934EAEFFF0BFFE7FC64F8CFFC63F8BA" bold="true" box="[967,984,1819,1843]" pageId="13" pageNumber="14">B</emphasis>
, first maxilla, with detail of distal portion of outer endite (
<emphasis id="B934EAEFFF0BFFE7FEA0F8E8FEA3F8DA" box="[259,280,1852,1875]" italics="true" pageId="13" pageNumber="14">ex</emphasis>
) and inner endite (
<emphasis id="B934EAEFFF0BFFE7FE79F8E8FE4BF8DA" box="[474,496,1852,1875]" italics="true" pageId="13" pageNumber="14">en</emphasis>
); 
<emphasis id="B934EAEFFF0BFFE7FDA5F8EFFDA3F8DA" bold="true" box="[518,536,1851,1875]" pageId="13" pageNumber="14">C</emphasis>
, second maxilla; 
<emphasis id="B934EAEFFF0BFFE7FD62F8EFFD6FF8DA" bold="true" box="[705,724,1851,1875]" pageId="13" pageNumber="14">D</emphasis>
, detail of the distal-most portion of the second maxilla; 
<emphasis id="B934EAEFFF0BFFE7FB59F8EFFAB1F8DA" bold="true" box="[1274,1290,1851,1875]" pageId="13" pageNumber="14">E</emphasis>
, hypopharynx; 
<emphasis id="B934EAEFFF0BFFE7FA05F8EFFA0EF8DA" bold="true" box="[1446,1461,1851,1875]" pageId="13" pageNumber="14">F</emphasis>
, detail of the median region of hypopharynx; 
<emphasis id="B934EAEFFF0BFFE7FDD6F88FFD33F8FA" bold="true" box="[629,648,1883,1907]" pageId="13" pageNumber="14">G</emphasis>
, maxilliped; 
<emphasis id="B934EAEFFF0BFFE7FCA9F88FFCA5F8FA" bold="true" box="[778,798,1883,1907]" pageId="13" pageNumber="14">H</emphasis>
, detail of the distal-most portion of maxilliped in posterior view; 
<emphasis id="B934EAEFFF0BFFE7FA0FF88FFA0DF8FA" bold="true" box="[1452,1462,1883,1907]" pageId="13" pageNumber="14">I</emphasis>
, detail of the distal-most portion of maxilliped in anterior view. Drawings of the corresponding structures modified from 
<bibRefCitation id="EFD14B0CFF0BFFE7FAEBF8AFFF46F83A" author="Richardson H" pageId="13" pageNumber="14" pagination="431 - 434" refId="ref17042" refString="Richardson H 1909. The isopod crustacean Acanthoniscus spiniger Kinahan redescribed. Proceedings of the United States National Museum, 36: 431 - 434. DOI: https: // www. doi. org / 10.5479 / si. 00963801.36 - 1678.431" type="journal article" year="1909">Richardson (1909)</bibRefCitation>
are included in 
<emphasis id="B934EAEFFF0BFFE7FE39F84FFE73F83A" bold="true" box="[410,456,1947,1971]" pageId="13" pageNumber="14">B, C</emphasis>
, and 
<emphasis id="B934EAEFFF0BFFE7FE5EF84FFDABF83A" bold="true" box="[509,528,1947,1971]" pageId="13" pageNumber="14">G</emphasis>
. Scale bars = 0.25 mm.
</paragraph>
</caption>
<paragraph id="8BFF36FDFF08FFE4FCD2FF43FAC0FDE9" blockId="14.[841,1468,151,881]" pageId="14" pageNumber="15">
Pereiopods (
<figureCitation id="137B2A78FF08FFE4FBA4FF43FB38FF3D" box="[1031,1155,151,180]" captionStart="Figure 5" captionStartId="12.[182,251,1692,1715]" captionTargetBox="[312,1351,473,1658]" captionTargetId="figure-134@12.[266,1382,465,1671]" captionTargetPageId="12" captionText="Figure 5.Otherappendagesof Acanthoniscus richardsonae sp.nov.A, Second antenna;B,detail offlagellum of secondantennadepicting position of aesthetasc groups (small arrows); C, detail of the distal article of the flagellum of the second antenna slightly turned over showing the aesthetasc groups; D, detail of the distal-most portion of the flagellum of the second antenna showing the apical cone; E, first antenna; F, detail of the third article of the first antenna showing the aesthetascs; G, male left seventh pereiopod; H, detail of ischium ofmale seventh pereiopod; I, detailof dactylus ofmale seventh pereiopodin anterior view; J, detailof dactylus ofmale seventh pereiopod in posterior view; K, male left first pereiopod, showing the areas on the carpus and propodus covered with the cleaning apparatus or antennal brush (dashed outlines); L, detail of carpus ofmale first pereiopod (anterior surface).Abbreviations:basis (bs), ischium(is),merus(me),carpus (ca),propodus (pr),flagellum(fl),and dactylus(da).Drawingofthe corresponding structuremodified from Richardson (1909) is included inA. Scale bars = 1 mm (A), 0.5 mm (G,K), 0.3mm (B), 0.1 mm (C–E, H–J,L),0.05 mm (F)." figureDoi="http://doi.org/10.5281/zenodo.10716776" httpUri="https://zenodo.org/record/10716776/files/figure.png" pageId="14" pageNumber="15">Fig. 5G–L</figureCitation>
). Basis of pereiopod 1 with dorsal ridge and dorsal surface with longitudinal groove; carpus ventral surface with 9 multi-apex setae, with one in subapical position nearly duplicating size of others; cleaning apparatus, antennal brush, or antenna-grooming brush broad and covering slightly over half of carpus anterior surface and also present on ventral surface of basal-most third of propodus. Male pereiopod 7 without special modifications, with 6 multi-apex setae on ventral surface of carpus, one in subapical position duplicating size of the others; ischium sub-conical, 2.3 times as long as wide.
</paragraph>
<paragraph id="8BFF36FDFF08FFFAFCD2FDBEFE59FD0E" blockId="14.[841,1468,151,881]" lastBlockId="16.[179,808,151,842]" lastPageId="16" lastPageNumber="17" pageId="14" pageNumber="15">
Pleopods (
<figureCitation id="137B2A78FF08FFE4FC56FDBEFB87FD0E" box="[1013,1084,618,647]" captionStart="Figure 7" captionStartId="14.[181,252,1786,1809]" captionTargetBox="[387,1266,935,1753]" captionTargetId="figure-324@14.[356,1292,932,1765]" captionTargetPageId="14" captionText="Figure 7.Pleopods of a male Acanthoniscus richardsonae sp.nov.A, First pleopod; B, detail of first pleopod endopodite tip; C, detail of first pleopod exopodite; D, second pleopod, with detail of the two setae on exopodite outer margin (arrow); E, third pleopod exopodite;F, detail ofthe outer-basal regionof the thirdpleopod exopoditeshowing the respiratory structures (thick arrow indicates the position of the spiracle); G, detail of the outer-basal region of the third pleopod exopodite showing the spiracle partially closed from above; H, fourth pleopod exopodite; I, fifth pleopod exopodite. Abbreviations: endopodite (en) and exopodite (ex). Scale bars = 0.5 mm (A–H), 0.25 mm (I)." figureDoi="http://doi.org/10.5281/zenodo.10716780" httpUri="https://zenodo.org/record/10716780/files/figure.png" pageId="14" pageNumber="15">Fig. 7</figureCitation>
). Exopodites 1–5 with tracheal structures and single elongate spiracle; first female endopodite absent, first exopodite vestigial. Endopodite 
<quantity id="4CB89B18FF08FFE4FC4EFD0BFB9AFD75" box="[1005,1057,735,764]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="14" pageNumber="15" unit="in" value="1.0">1 in</quantity>
males slightly curved outward distally, 3.7 times as long as wide, distal part without spines; exopodite 
<quantity id="4CB89B18FF08FFE4FBBEFCF9FBF3FCC3" box="[1053,1096,813,842]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="14" pageNumber="15" unit="in" value="1.0">1 in</quantity>
males asymmetric, 1.5 times as long as wide, with narrower inner margin, tiny setae distributed along entire margin, 1 larger apical seta on inner narrow margin, 3 subapical setae, 1 seta on outer margin and 1 submarginal seta one-third before apex. Exopodite 
<quantity id="4CB89B18FF16FFFAFE9AFED8FED3FEA0" box="[313,360,268,297]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="16" pageNumber="17" unit="in" value="2.0">2 in</quantity>
males very wide basally and narrow distally, 1.8 times as long as wide, with 5 submarginal setae; inner margin with tiny setae and notch in middle region; endopodite 
<quantity id="4CB89B18FF16FFFAFE0EFE55FE64FE17" box="[429,479,385,414]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="16" pageNumber="17" unit="in" value="2.0">2 in</quantity>
males slightly surpassing tip of exopodite 2 and with 2 setae on outer margin near median region (one of them long and slender and the other short and spatulate). Exopodites 3–4 with 3 submarginal setae near outer-distal margin. Exopodite 5 widely convex, with 4 submarginal setae near outer-distal margin.
</paragraph>
<caption id="DF3F6675FF08FFE4FF16F92EFDA5F838" ID-DOI="http://doi.org/10.5281/zenodo.10716780" ID-Zenodo-Dep="10716780" httpUri="https://zenodo.org/record/10716780/files/figure.png" pageId="14" pageNumber="15" startId="14.[181,252,1786,1809]" targetBox="[387,1266,935,1753]" targetPageId="14" targetType="figure">
<paragraph id="8BFF36FDFF08FFE4FF16F92EFDA5F838" blockId="14.[180,1467,1785,1969]" pageId="14" pageNumber="15">
<emphasis id="B934EAEFFF08FFE4FF16F92EFEACF898" bold="true" box="[181,279,1786,1809]" pageId="14" pageNumber="15">Figure 7.</emphasis>
Pleopods of a male 
<taxonomicName id="4C404D7EFF08FFE4FE75F92DFD61F898" authority="Rodríguez-Cabrera &amp; de Armas, 2023" authorityName="Rodríguez-Cabrera &amp; de Armas" authorityYear="2023" box="[470,730,1785,1809]" class="Malacostraca" family="Delatorreiidae" genus="Acanthoniscus" kingdom="Animalia" order="Isopoda" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="richardsonae" status="sp. nov.">
<emphasis id="B934EAEFFF08FFE4FE75F92DFD61F898" box="[470,730,1785,1809]" italics="true" pageId="14" pageNumber="15">Acanthoniscus richardsonae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2075794FF08FFE4FD7CF92EFC93F898" box="[735,808,1786,1809]" pageId="14" pageNumber="15" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="B934EAEFFF08FFE4FC88F92DFC87F898" bold="true" box="[811,828,1785,1809]" pageId="14" pageNumber="15">A</emphasis>
, First pleopod; 
<emphasis id="B934EAEFFF08FFE4FC75F92DFC5CF898" bold="true" box="[982,999,1785,1809]" pageId="14" pageNumber="15">B</emphasis>
, detail of first pleopod endopodite tip; 
<emphasis id="B934EAEFFF08FFE4FAC5F92DFAC3F898" bold="true" box="[1382,1400,1785,1809]" pageId="14" pageNumber="15">C</emphasis>
, detail of first pleopod exopodite; 
<emphasis id="B934EAEFFF08FFE4FE64F8CDFE61F8B8" bold="true" box="[455,474,1817,1841]" pageId="14" pageNumber="15">D</emphasis>
, second pleopod, with detail of the two setae on exopodite outer margin (arrow); 
<emphasis id="B934EAEFFF08FFE4FAB5F8CDFA9EF8B8" bold="true" box="[1302,1317,1817,1841]" pageId="14" pageNumber="15">E</emphasis>
, third pleopod exopodite; 
<emphasis id="B934EAEFFF08FFE4FE80F8EDFE89F8D8" bold="true" box="[291,306,1849,1873]" pageId="14" pageNumber="15">F</emphasis>
, detail of the outer-basal region of the third pleopod exopodite showing the respiratory structures (thick arrow indicates the position of the spiracle); 
<emphasis id="B934EAEFFF08FFE4FE72F88DFE5FF8F8" bold="true" box="[465,484,1881,1905]" pageId="14" pageNumber="15">G</emphasis>
, detail of the outer-basal region of the third pleopod exopodite showing the spiracle partially closed from above; 
<emphasis id="B934EAEFFF08FFE4FE96F8ADFEF2F818" bold="true" box="[309,329,1913,1937]" pageId="14" pageNumber="15">H</emphasis>
, fourth pleopod exopodite; I, fifth pleopod exopodite. Abbreviations: endopodite (
<emphasis id="B934EAEFFF08FFE4FB30F8AEFB12F818" box="[1171,1193,1914,1937]" italics="true" pageId="14" pageNumber="15">en</emphasis>
) and exopodite (
<emphasis id="B934EAEFFF08FFE4FAF8F8AEFACBF818" box="[1371,1392,1914,1937]" italics="true" pageId="14" pageNumber="15">ex</emphasis>
). Scale bars = 0.5 mm (
<emphasis id="B934EAEFFF08FFE4FEF0F84DFE3EF838" bold="true" box="[339,389,1945,1969]" pageId="14" pageNumber="15">A–H</emphasis>
), 0.25 mm (
<emphasis id="B934EAEFFF08FFE4FDA0F84DFDB6F838" bold="true" box="[515,525,1945,1969]" pageId="14" pageNumber="15">I</emphasis>
).
</paragraph>
</caption>
<caption id="DF3F6675FF09FFE5FF16FA27FB91F8A3" ID-DOI="http://doi.org/10.5281/zenodo.10716782" ID-Zenodo-Dep="10716782" httpUri="https://zenodo.org/record/10716782/files/figure.png" pageId="15" pageNumber="16" startId="15.[181,252,1523,1546]" targetBox="[192,1460,173,1475]" targetPageId="15" targetType="figure">
<paragraph id="8BFF36FDFF09FFE5FF16FA27FB91F8A3" blockId="15.[180,1467,1522,1834]" pageId="15" pageNumber="16">
<emphasis id="B934EAEFFF09FFE5FF16FA27FEA2F983" bold="true" box="[181,281,1523,1546]" pageId="15" pageNumber="16">Figure 8.</emphasis>
Selected body parts of 
<taxonomicName id="4C404D7EFF09FFE5FE5CFA26FD3DF983" box="[511,646,1522,1546]" class="Malacostraca" family="Delatorreiidae" genus="Acanthoniscus" kingdom="Animalia" order="Isopoda" pageId="15" pageNumber="16" phylum="Arthropoda" rank="genus">
<emphasis id="B934EAEFFF09FFE5FE5CFA26FD3DF983" box="[511,646,1522,1546]" italics="true" pageId="15" pageNumber="16">Acanthoniscus</emphasis>
</taxonomicName>
for comparison with other taxa within 
<taxonomicName id="4C404D7EFF09FFE5FBA8FA26FB34F983" authorityName="Brandt" authorityYear="1831" box="[1035,1167,1522,1546]" class="Malacostraca" family="Armadillidae" kingdom="Animalia" order="Isopoda" pageId="15" pageNumber="16" phylum="Arthropoda" rank="family">Armadillidae</taxonomicName>
. 
<taxonomicName id="4C404D7EFF09FFE5FB34FA26FA0DF983" authority="Brandt, 1833" authorityName="Brandt" authorityYear="1833" box="[1175,1462,1522,1546]" class="Malacostraca" family="Armadillidae" genus="Cubaris" kingdom="Animalia" order="Isopoda" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="murina">
<emphasis id="B934EAEFFF09FFE5FB34FA26FA95F983" box="[1175,1326,1522,1546]" italics="true" pageId="15" pageNumber="16">Cubaris murina</emphasis>
Brandt, 1833
</taxonomicName>
, showing the ground pattern in 
<taxonomicName id="4C404D7EFF09FFE5FE45F9C6FDD3F9A3" authorityName="Brandt" authorityYear="1831" box="[486,616,1554,1578]" class="Malacostraca" family="Armadillidae" kingdom="Animalia" order="Isopoda" pageId="15" pageNumber="16" phylum="Arthropoda" rank="family">Armadillidae</taxonomicName>
(
<emphasis id="B934EAEFFF09FFE5FDDBF9C6FD1DF9A3" bold="true" box="[632,678,1554,1578]" pageId="15" pageNumber="16">A–E</emphasis>
): 
<emphasis id="B934EAEFFF09FFE5FD1EF9C6FD75F9A3" bold="true" box="[701,718,1554,1578]" pageId="15" pageNumber="16">A</emphasis>
, Posterior region of the body in dorsal view; 
<emphasis id="B934EAEFFF09FFE5FB2AF9C6FB21F9A3" bold="true" box="[1161,1178,1554,1578]" pageId="15" pageNumber="16">B</emphasis>
, posterior region of the body in ventral view; 
<emphasis id="B934EAEFFF09FFE5FEF4F9E6FED2F9C3" bold="true" box="[343,361,1586,1610]" pageId="15" pageNumber="16">C</emphasis>
, pleotelson in dorsal view; 
<emphasis id="B934EAEFFF09FFE5FDDDF9E6FD2AF9C3" bold="true" box="[638,657,1586,1610]" pageId="15" pageNumber="16">D</emphasis>
, uropod in dorsal view; 
<emphasis id="B934EAEFFF09FFE5FC2BF9E6FC23F9C3" bold="true" box="[904,920,1586,1610]" pageId="15" pageNumber="16">E</emphasis>
, middle region of the sternite 7 of a male showing the bilobed caudal process on the posterior margin, which is considered an autapomorphy of 
<taxonomicName id="4C404D7EFF09FFE5FB89F986FB15F9E3" authorityName="Brandt" authorityYear="1831" box="[1066,1198,1618,1642]" class="Malacostraca" family="Armadillidae" kingdom="Animalia" order="Isopoda" pageId="15" pageNumber="16" phylum="Arthropoda" rank="family">Armadillidae</taxonomicName>
. 
<taxonomicName id="4C404D7EFF09FFE5FB16F986FA00F9E3" authority="Rodríguez-Cabrera &amp; de Armas, 2023" authorityName="Rodríguez-Cabrera &amp; de Armas" authorityYear="2023" box="[1205,1467,1618,1642]" class="Malacostraca" family="Delatorreiidae" genus="Acanthoniscus" kingdom="Animalia" order="Isopoda" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="richardsonae" status="sp. nov.">
<emphasis id="B934EAEFFF09FFE5FB16F986FA00F9E3" box="[1205,1467,1618,1642]" italics="true" pageId="15" pageNumber="16">Acanthoniscus richardsonae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2075794FF09FFE5FF16F9A7FF44F903" box="[181,255,1651,1674]" pageId="15" pageNumber="16" rank="species">sp. nov.</taxonomicNameLabel>
(
<emphasis id="B934EAEFFF09FFE5FEAEF9A6FE86F903" bold="true" box="[269,317,1650,1674]" pageId="15" pageNumber="16">F–H</emphasis>
): 
<emphasis id="B934EAEFFF09FFE5FEF7F9A6FED8F903" bold="true" box="[340,355,1650,1674]" pageId="15" pageNumber="16">F</emphasis>
, pleotelson in dorsal view; 
<emphasis id="B934EAEFFF09FFE5FDCDF9A6FD3AF903" bold="true" box="[622,641,1650,1674]" pageId="15" pageNumber="16">G</emphasis>
, uropod in dorsal view; 
<emphasis id="B934EAEFFF09FFE5FCCCF9A6FC38F903" bold="true" box="[879,899,1650,1674]" pageId="15" pageNumber="16">H</emphasis>
, middle region of the sternite 7 of a male. Drawings of the pleotelson and uropods in dorsal views of some morphologically divergent taxa within 
<taxonomicName id="4C404D7EFF09FFE5FBAEF946FB2AF923" authorityName="Brandt" authorityYear="1831" box="[1037,1169,1682,1706]" class="Malacostraca" family="Armadillidae" kingdom="Animalia" order="Isopoda" pageId="15" pageNumber="16" phylum="Arthropoda" rank="family">Armadillidae</taxonomicName>
. 
<emphasis id="B934EAEFFF09FFE5FB3AF946FB18F923" bold="true" box="[1177,1187,1682,1706]" pageId="15" pageNumber="16">I</emphasis>
, 
<taxonomicName id="4C404D7EFF09FFE5FB0EF946FF50F943" authority="Green, 1963" authorityName="Green" authorityYear="1963" class="Malacostraca" family="Armadillidae" genus="Echinodillo" kingdom="Animalia" order="Isopoda" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="cavaticus">
<emphasis id="B934EAEFFF09FFE5FB0EF946FAC9F923" box="[1197,1394,1682,1706]" italics="true" pageId="15" pageNumber="16">Echinodillo cavaticus</emphasis>
<bibRefCitation id="EFD14B0CFF09FFE5FAD5F946FF50F943" author="Green A" pageId="15" pageNumber="16" pagination="77 - 80" refId="ref15608" refString="Green A 1963. A new species of Echinodillo (Isopoda, Oniscoidea, Armadillidae) from Flinders Island, Tasmania. Papers and Proceedings of the Royal Society of Tasmania, 97: 77 - 80. https: // eprints. utas. edu. au / 14329 / 1 / 1963 _ Green _ New _ species _ Echinodillo. pdf" type="journal article" year="1963">Green, 1963</bibRefCitation>
</taxonomicName>
; 
<emphasis id="B934EAEFFF09FFE5FF56F966FF44F943" bold="true" box="[245,255,1714,1738]" pageId="15" pageNumber="16">J</emphasis>
, 
<taxonomicName id="4C404D7EFF09FFE5FEAAF966FDD2F943" authority="Jackson, 1933" authorityName="Jackson" authorityYear="1933" box="[265,617,1714,1738]" class="Malacostraca" family="Armadillidae" genus="Echinodillo" kingdom="Animalia" order="Isopoda" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="montanus">
<emphasis id="B934EAEFFF09FFE5FEAAF966FE6DF943" box="[265,470,1714,1738]" italics="true" pageId="15" pageNumber="16">Echinodillo montanus</emphasis>
<bibRefCitation id="EFD14B0CFF09FFE5FE7FF966FDD2F943" author="Jackson H" box="[476,617,1714,1738]" pageId="15" pageNumber="16" pagination="145 - 162" refId="ref16063" refString="Jackson H 1933. Marquesan terrestrial Isopoda. Bulletin of the Bernice P. Bishop Museum, 114: 145 - 162." type="journal article" year="1933">Jackson, 1933</bibRefCitation>
</taxonomicName>
; 
<emphasis id="B934EAEFFF09FFE5FDD1F966FD3FF943" bold="true" box="[626,644,1714,1738]" pageId="15" pageNumber="16">K</emphasis>
, 
<taxonomicName id="4C404D7EFF09FFE5FD2CF966FB2AF943" authority="Kwon, Ferrara and Taiti, 1992" authorityName="Kwon, Ferrara and Taiti" authorityYear="1992" box="[655,1169,1714,1738]" class="Malacostraca" family="Armadillidae" genus="Laureola" kingdom="Animalia" order="Isopoda" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="vietnamensis">
<emphasis id="B934EAEFFF09FFE5FD2CF966FCDAF943" box="[655,865,1714,1738]" italics="true" pageId="15" pageNumber="16">Laureola vietnamensis</emphasis>
<bibRefCitation id="EFD14B0CFF09FFE5FCC5F967FB2AF943" author="Kwon D &amp; Ferrara F &amp; Taiti S" box="[870,1169,1714,1738]" pageId="15" pageNumber="16" pagination="645 - 653" refId="ref16327" refString="Kwon D; Ferrara F and Taiti S 1992. Two new species of Laureola Barnard, 1960 from India and Vietnam (Crustacea, Oniscidea, Armadillidae). Revue suisse de Zoologie, 99: 645 - 653. DOI: https: // www. doi. org / 10.5962 / bhl. part. 79844" type="journal article" year="1992">Kwon, Ferrara and Taiti, 1992</bibRefCitation>
</taxonomicName>
; 
<emphasis id="B934EAEFFF09FFE5FB38F966FB11F943" bold="true" box="[1179,1194,1714,1738]" pageId="15" pageNumber="16">L</emphasis>
, 
<taxonomicName id="4C404D7EFF09FFE5FB17F966FEDFF963" authority="Schmalfuss and Ferrara, 1983" authorityName="Schmalfuss and Ferrara" authorityYear="1983" class="Malacostraca" family="Armadillidae" genus="Laureola" kingdom="Animalia" order="Isopoda" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="dubia">
<emphasis id="B934EAEFFF09FFE5FB17F966FAFEF943" box="[1204,1349,1714,1738]" italics="true" pageId="15" pageNumber="16">Laureola dubia</emphasis>
<bibRefCitation id="EFD14B0CFF09FFE5FAE9F966FEDFF963" author="Schmalfuss H &amp; Ferrara F" pageId="15" pageNumber="16" pagination="111 - 157" refId="ref17246" refString="Schmalfuss H and Ferrara F 1983. Terrestrial isopods from West Africa, Part 3: Family Armadillidae Verhoeff, 1917. Monitore zoologico italiano, Nuova Serie, Supplemento, 18: 111 - 157. DOI: https: // www. doi. org / 10.1080 / 00269786.1983.11 758568" type="journal article" year="1983">Schmalfuss and Ferrara, 1983</bibRefCitation>
</taxonomicName>
; 
<emphasis id="B934EAEFFF09FFE5FECFF906FE39F963" bold="true" box="[364,386,1746,1770]" pageId="15" pageNumber="16">M</emphasis>
, 
<taxonomicName id="4C404D7EFF09FFE5FE2FF906FD6AF963" authority="Verhoeff, 1926" authorityName="Verhoeff" authorityYear="1926" box="[396,721,1746,1770]" class="Malacostraca" family="Armadillidae" genus="Schismadillo" kingdom="Animalia" order="Isopoda" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="rouxi">
<emphasis id="B934EAEFFF09FFE5FE2FF906FD82F963" box="[396,569,1746,1770]" italics="true" pageId="15" pageNumber="16">Schismadillo rouxi</emphasis>
Verhoeff, 1926
</taxonomicName>
; 
<emphasis id="B934EAEFFF09FFE5FD7AF906FD57F963" bold="true" box="[729,748,1746,1770]" pageId="15" pageNumber="16">N</emphasis>
, 
<taxonomicName id="4C404D7EFF09FFE5FD56F906FBD9F963" authority="Verhoeff, 1926" authorityName="Verhoeff" authorityYear="1926" box="[757,1122,1746,1770]" class="Malacostraca" family="Armadillidae" genus="Merulanella" kingdom="Animalia" order="Isopoda" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="wahrbergi">
<emphasis id="B934EAEFFF09FFE5FD56F906FC72F963" box="[757,969,1746,1770]" italics="true" pageId="15" pageNumber="16">Merulanella wahrbergi</emphasis>
Verhoeff, 1926
</taxonomicName>
; 
<emphasis id="B934EAEFFF09FFE5FBCAF906FBC6F963" bold="true" box="[1129,1149,1746,1770]" pageId="15" pageNumber="16">O</emphasis>
, 
<taxonomicName id="4C404D7EFF09FFE5FB24F906FA03F963" authority="Verhoeff, 1926" authorityName="Verhoeff" authorityYear="1926" box="[1159,1464,1746,1770]" class="Malacostraca" family="Armadillidae" genus="Nesodillo" kingdom="Animalia" order="Isopoda" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="incisus">
<emphasis id="B934EAEFFF09FFE5FB24F906FA9BF963" box="[1159,1312,1746,1770]" italics="true" pageId="15" pageNumber="16">Nesodillo incisus</emphasis>
Verhoeff, 1926
</taxonomicName>
. Drawings modified from Verhoeff (1926) (
<emphasis id="B934EAEFFF09FFE5FDFCF926FD23F883" bold="true" box="[607,664,1778,1802]" pageId="15" pageNumber="16">M–O</emphasis>
), 
<bibRefCitation id="EFD14B0CFF09FFE5FD0EF926FCFCF883" author="Jackson H" box="[685,839,1778,1802]" pageId="15" pageNumber="16" pagination="145 - 162" refId="ref16063" refString="Jackson H 1933. Marquesan terrestrial Isopoda. Bulletin of the Bernice P. Bishop Museum, 114: 145 - 162." type="journal article" year="1933">Jackson (1933)</bibRefCitation>
(
<emphasis id="B934EAEFFF09FFE5FCF5F926FCDBF883" bold="true" box="[854,864,1778,1802]" pageId="15" pageNumber="16">J</emphasis>
), 
<bibRefCitation id="EFD14B0CFF09FFE5FCD7F926FBBBF883" author="Green A" box="[884,1024,1778,1802]" pageId="15" pageNumber="16" pagination="77 - 80" refId="ref15608" refString="Green A 1963. A new species of Echinodillo (Isopoda, Oniscoidea, Armadillidae) from Flinders Island, Tasmania. Papers and Proceedings of the Royal Society of Tasmania, 97: 77 - 80. https: // eprints. utas. edu. au / 14329 / 1 / 1963 _ Green _ New _ species _ Echinodillo. pdf" type="journal article" year="1963">Green (1963)</bibRefCitation>
(
<emphasis id="B934EAEFFF09FFE5FBB3F926FBA1F883" bold="true" box="[1040,1050,1778,1802]" pageId="15" pageNumber="16">I</emphasis>
), 
<bibRefCitation id="EFD14B0CFF09FFE5FB8CF926FAD9F883" author="Schmalfuss H &amp; Ferrara F" box="[1071,1378,1778,1802]" pageId="15" pageNumber="16" pagination="111 - 157" refId="ref17246" refString="Schmalfuss H and Ferrara F 1983. Terrestrial isopods from West Africa, Part 3: Family Armadillidae Verhoeff, 1917. Monitore zoologico italiano, Nuova Serie, Supplemento, 18: 111 - 157. DOI: https: // www. doi. org / 10.1080 / 00269786.1983.11 758568" type="journal article" year="1983">Schmalfuss and Ferrara (1983)</bibRefCitation>
(
<emphasis id="B934EAEFFF09FFE5FAD2F926FA3BF883" bold="true" box="[1393,1408,1778,1802]" pageId="15" pageNumber="16">L</emphasis>
), and 
<bibRefCitation id="EFD14B0CFF09FFE5FF16F8C7FEC9F8A3" author="Kwon D &amp; Ferrara F &amp; Taiti S" box="[181,370,1810,1834]" pageId="15" pageNumber="16" pagination="645 - 653" refId="ref16327" refString="Kwon D; Ferrara F and Taiti S 1992. Two new species of Laureola Barnard, 1960 from India and Vietnam (Crustacea, Oniscidea, Armadillidae). Revue suisse de Zoologie, 99: 645 - 653. DOI: https: // www. doi. org / 10.5962 / bhl. part. 79844" type="journal article" year="1992">Kwon et al. (1992)</bibRefCitation>
(
<emphasis id="B934EAEFFF09FFE5FE22F8C6FE28F8A3" bold="true" box="[385,403,1810,1834]" pageId="15" pageNumber="16">K</emphasis>
). Abbreviations: endopodite (
<emphasis id="B934EAEFFF09FFE5FD67F8C7FD61F8A3" box="[708,730,1811,1834]" italics="true" pageId="15" pageNumber="16">en</emphasis>
), exopodite (
<emphasis id="B934EAEFFF09FFE5FCC0F8C7FCC3F8A3" box="[867,888,1811,1834]" italics="true" pageId="15" pageNumber="16">ex</emphasis>
), sympodite (
<emphasis id="B934EAEFFF09FFE5FBA4F8C7FBA2F8A3" box="[1031,1049,1811,1834]" italics="true" pageId="15" pageNumber="16">sy</emphasis>
).
</paragraph>
</caption>
<paragraph id="8BFF36FDFF16FFFAFF78FD46FEDCFCC3" blockId="16.[179,808,151,842]" pageId="16" pageNumber="17">
Tegument (
<figureCitation id="137B2A78FF16FFFAFED8FD45FE45FD27" box="[379,510,657,686]" captionStart-0="Figure 4" captionStart-1="Figure 5" captionStart-2="Figure 6" captionStartId-0="11.[181,252,1628,1651]" captionStartId-1="12.[182,251,1692,1715]" captionStartId-2="13.[181,252,1788,1811]" captionTargetBox-0="[246,1407,152,1593]" captionTargetBox-1="[312,1351,473,1658]" captionTargetBox-2="[267,1379,821,1745]" captionTargetId-0="figure-20@11.[229,1420,148,1608]" captionTargetId-1="figure-134@12.[266,1382,465,1671]" captionTargetId-2="figure-164@13.[232,1415,817,1764]" captionTargetPageId-0="11" captionTargetPageId-1="12" captionTargetPageId-2="13" captionText-0="Figure 4. Body parts of Acanthoniscus richardsonae sp. nov. A, Overall, dorsal view; B, detail of the microsculpture of the dorsal tegument (left) and tricorn-like setae (right); C, lateral view of the anterior region of the body;D, lateral view of the posterior region of the body; E, right pleon-epimera 3–5 in ventral view; F, right pereon-epimera 1–7 in ventral view; G, detail of pereon-epimera 1–2 in a more ventral position; H, cephalothorax in dorsal view; I, cephalothorax in frontal view (the tiny sensory setae inserted between the ommatidia are illustrated under a magnification of 400×: a, b, c); J, cephalothorax in ventral view; K, pleotelson and uropods in dorsal view (the uropod exopodites are slightly pointing upward); L, pleotelson and uropods in ventral view; M, right uropod in ventral view; N, right uropod in dorsal view; O, drawing of the overall dorsal view modified from Richardson (1909); P, arrangement of dorsal spines, the diametersof the circles correspond tothe relative size of the spines.Drawings of the corresponding structures modified from Richardson (1909)are included in I,J (the question mark means that the position representing thedrawing is not totally clear) and in M. Abbreviations: sympodite (sy), endopodite (en), and exopodite (ex). Scale bars = 1 mm (A–L), 0.5 mm (M–N); in some cases, a single scale bar applies to more than one figure (indicated by the letters above the scale bar)." captionText-1="Figure 5.Otherappendagesof Acanthoniscus richardsonae sp.nov.A, Second antenna;B,detail offlagellum of secondantennadepicting position of aesthetasc groups (small arrows); C, detail of the distal article of the flagellum of the second antenna slightly turned over showing the aesthetasc groups; D, detail of the distal-most portion of the flagellum of the second antenna showing the apical cone; E, first antenna; F, detail of the third article of the first antenna showing the aesthetascs; G, male left seventh pereiopod; H, detail of ischium ofmale seventh pereiopod; I, detailof dactylus ofmale seventh pereiopodin anterior view; J, detailof dactylus ofmale seventh pereiopod in posterior view; K, male left first pereiopod, showing the areas on the carpus and propodus covered with the cleaning apparatus or antennal brush (dashed outlines); L, detail of carpus ofmale first pereiopod (anterior surface).Abbreviations:basis (bs), ischium(is),merus(me),carpus (ca),propodus (pr),flagellum(fl),and dactylus(da).Drawingofthe corresponding structuremodified from Richardson (1909) is included inA. Scale bars = 1 mm (A), 0.5 mm (G,K), 0.3mm (B), 0.1 mm (C–E, H–J,L),0.05 mm (F)." captionText-2="Figure 6.Mouthparts of Acanthoniscus richardsonae sp. nov. A, Distal portion of right (R) and left (L) mandibles in anterior views, showing lacinia mobilis (lm), hairy lobe (hl), penicils (pn), and tuft of setae (ts); B, first maxilla, with detailof distalportion of outer endite (ex) and inner endite (en); C, second maxilla; D, detail of the distal-most portion of the second maxilla; E, hypopharynx; F, detail of the median region of hypopharynx; G, maxilliped; H, detail of the distal-most portion of maxilliped in posterior view; I, detail of the distal-most portion of maxilliped inanterior view.Drawings of the corresponding structures modified from Richardson (1909) are included in B, C, and G. Scale bars = 0.25 mm." figureDoi-0="http://doi.org/10.5281/zenodo.10716774" figureDoi-1="http://doi.org/10.5281/zenodo.10716776" figureDoi-2="http://doi.org/10.5281/zenodo.10716778" httpUri-0="https://zenodo.org/record/10716774/files/figure.png" httpUri-1="https://zenodo.org/record/10716776/files/figure.png" httpUri-2="https://zenodo.org/record/10716778/files/figure.png" pageId="16" pageNumber="17">Figs. 4–6</figureCitation>
). Dorsal tegument of cephalothorax, pereon and pleon, as well as that of second antenna, pereiopods and maxilliped basipodite, bearing scaly microsculpture, with tricorn-like setae on some areas.
</paragraph>
<paragraph id="8BFF36FDFF16FFFAFF78FCA8FCB1FB7F" blockId="16.[180,807,891,1270]" pageId="16" pageNumber="17">
<emphasis id="B934EAEFFF16FFFAFF78FCA8FEF1FC11" box="[219,330,892,920]" italics="true" pageId="16" pageNumber="17">Variation</emphasis>
. Adult size varied from 
<quantity id="4CB89B18FF16FFFAFDCCFCAFFCBEFC1E" box="[623,773,891,920]" metricMagnitude="-3" metricUnit="m" metricValue="7.05" metricValueMax="8.5" metricValueMin="5.6" pageId="16" pageNumber="17" unit="mm" value="7.05" valueMax="8.5" valueMin="5.6">5.6–8.5 mm</quantity>
in total length, but males (5.6–7.2, n = 3) tend to be smaller than females (5.7–8.5, n = 6) in the sample; juveniles measured 4.8 and 5.0 mm. 
<specimenCount id="9D46FD74FF16FFFAFD02FC24FC9DFB84" box="[673,806,1008,1037]" count="2" pageId="16" pageNumber="17" type="adult">Two adult</specimenCount>
females had a small paramedian spine on one side of pleonite 5. One of the juveniles had a very small supernumerary spiniform tubercle between one of the pairs of paramedian tubercles of pereonite 1. The middle article of maxilliped palpus sometimes with two setae in distal position on the medial margin.
</paragraph>
</subSubSection>
<caption id="DF3F6675FF16FFFAFCE8FB95FA37FBF0" ID-DOI="http://doi.org/10.5281/zenodo.10716784" ID-Zenodo-Dep="10716784" httpUri="https://zenodo.org/record/10716784/files/figure.png" pageId="16" pageNumber="17" startId="16.[843,914,1089,1112]" targetBox="[843,1467,151,1065]" targetPageId="16" targetType="figure">
<paragraph id="8BFF36FDFF16FFFAFCE8FB95FA37FBF0" blockId="16.[843,1466,1089,1145]" pageId="16" pageNumber="17">
<emphasis id="B934EAEFFF16FFFAFCE8FB95FC0BFBD1" bold="true" box="[843,944,1089,1112]" pageId="16" pageNumber="17">Figure 9.</emphasis>
Habitat of 
<taxonomicName id="4C404D7EFF16FFFAFB80FB95FA97FBD0" authority="Rodríguez-Cabrera &amp; de Armas, 2023" authorityName="Rodríguez-Cabrera &amp; de Armas" authorityYear="2023" box="[1059,1324,1089,1113]" class="Malacostraca" family="Delatorreiidae" genus="Acanthoniscus" kingdom="Animalia" order="Isopoda" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="richardsonae" status="sp. nov.">
<emphasis id="B934EAEFFF16FFFAFB80FB95FA97FBD0" box="[1059,1324,1089,1113]" italics="true" pageId="16" pageNumber="17">Acanthoniscus richardsonae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2075794FF16FFFAFA91FB95FAC6FBD1" box="[1330,1405,1089,1112]" pageId="16" pageNumber="17" rank="species">sp. nov.</taxonomicNameLabel>
in the Blue Mountains, eastern Jamaica. Photo by Carlos Víquez.
</paragraph>
</caption>
<subSubSection id="C35A6576FF16FFFAFF78FAF3FC9EFA30" pageId="16" pageNumber="17" type="biology_ecology">
<paragraph id="8BFF36FDFF16FFFAFF78FAF3FC9EFA30" blockId="16.[181,805,1319,1465]" pageId="16" pageNumber="17">
<emphasis id="B934EAEFFF16FFFAFF78FAF3FE35FACD" box="[219,398,1319,1348]" italics="true" pageId="16" pageNumber="17">Natural history</emphasis>
. Specimens of 
<taxonomicName id="4C404D7EFF16FFFAFDE5FAFCFD47FACD" authority="Rodríguez-Cabrera &amp; de Armas, 2023" authorityName="Rodríguez-Cabrera &amp; de Armas" authorityYear="2023" box="[582,764,1319,1348]" class="Malacostraca" family="Delatorreiidae" genus="Acanthoniscus" kingdom="Animalia" order="Isopoda" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="richardsonae" status="sp. nov.">
<emphasis id="B934EAEFFF16FFFAFDE5FAFCFD47FACD" box="[582,764,1319,1348]" italics="true" pageId="16" pageNumber="17">A. richardsonae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2075794FF16FFFAFCA7FAFCFF53FAE2" pageId="16" pageNumber="17" rank="species">sp. nov.</taxonomicNameLabel>
were collected under rocks and in humid leaf litter in closed broadleaf forest, specifically in upper montane rainforest (F.Cala, in litt., 
<date id="FFFE103DFF16FFFAFDEAFA48FD7BFA30" box="[585,704,1436,1465]" pageId="16" pageNumber="17" value="2018-09-04">4.IX.2018</date>
) (
<figureCitation id="137B2A78FF16FFFAFD74FA48FCACFA30" box="[727,791,1436,1465]" captionStart="Figure 9" captionStartId="16.[843,914,1089,1112]" captionTargetBox="[843,1467,151,1065]" captionTargetId="figure-503@16.[843,1467,148,1065]" captionTargetPageId="16" captionText="Figure 9. Habitat of Acanthoniscus richardsonae sp. nov.in the Blue Mountains, eastern Jamaica. Photo by Carlos Víquez." figureDoi="http://doi.org/10.5281/zenodo.10716784" httpUri="https://zenodo.org/record/10716784/files/figure.png" pageId="16" pageNumber="17">Fig. 9</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C35A6576FF16FFFAFF78FA3EFDE8F83A" pageId="16" pageNumber="17" type="distribution">
<paragraph id="8BFF36FDFF16FFFAFF78FA3EFDE8F83A" blockId="16.[180,806,1514,1971]" pageId="16" pageNumber="17">
<emphasis id="B934EAEFFF16FFFAFF78FA3EFED0F98E" box="[219,363,1514,1543]" italics="true" pageId="16" pageNumber="17">Distribution</emphasis>
. Know only from the 
<typeStatus id="54FB885FFF16FFFAFD27FA3EFD00F98F" box="[644,699,1514,1542]" pageId="16" pageNumber="17">type</typeStatus>
locality, around the ranger station, Blue Mountains, 
<collectingRegion id="4984F81FFF16FFFAFD46F9C5FECCF9DC" country="Jamaica" name="Saint Thomas" pageId="16" pageNumber="17">Saint Thomas Parish</collectingRegion>
, 
<collectingCountry id="F357766DFF16FFFAFE26F9ECFE5DF9DC" box="[389,486,1592,1621]" name="Jamaica" pageId="16" pageNumber="17">Jamaica</collectingCountry>
(
<figureCitation id="137B2A78FF16FFFAFE58F9ECFDFAF9DC" box="[507,577,1592,1621]" captionStart="Figure 1" captionStartId="3.[181,252,1611,1634]" captionTargetBox="[181,1467,151,1587]" captionTargetId="figure-19@3.[181,1467,151,1587]" captionTargetPageId="3" captionText="Figure 1. Distribution of the genus Acanthoniscus in Jamaica. Above: type locality of A. spiniger at the hills near Bluefields (yellow dot), the type locality of A.richardsonae sp.nov.in the Blue Mountains (red dot), and a second location in the Blue Mountains where a specimen presumably of A. richardsonae sp. nov. was photographed (orange dot). The main collection sites of Henry G. Hubbard in 1877 are depicted by green dots, notice the proximity of Stony Hill to the distribution of A. richardsonae sp.nov.Below: The hills known as Surinam Quarters near Bluefields, referred by Gosse (1851) as “Bluefields Mountain,” Westmoreland Parish, constitute the type locality of A. spiniger." figureDoi="http://doi.org/10.5281/zenodo.10716766" httpUri="https://zenodo.org/record/10716766/files/figure.png" pageId="16" pageNumber="17">Fig. 1</figureCitation>
). Nonetheless, on 
<date id="FFFE103DFF16FFFAFF16F98AFE71F9F5" box="[181,458,1629,1660]" pageId="16" pageNumber="17" value="2013-11-19">
November 19 
<superScript id="7C359BB5FF16FFFAFEC4F989FECCF9E7" attach="right" box="[359,375,1629,1646]" fontSize="7" pageId="16" pageNumber="17">th</superScript>
, 2013
</date>
, a specimen which at first sight matched the description of 
<taxonomicName id="4C404D7EFF16FFFAFDE5F952FD47F92B" authority="Rodríguez-Cabrera &amp; de Armas, 2023" authorityName="Rodríguez-Cabrera &amp; de Armas" authorityYear="2023" box="[582,764,1669,1698]" class="Malacostraca" family="Delatorreiidae" genus="Acanthoniscus" kingdom="Animalia" order="Isopoda" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="richardsonae" status="sp. nov.">
<emphasis id="B934EAEFFF16FFFAFDE5F952FD47F92B" box="[582,764,1669,1698]" italics="true" pageId="16" pageNumber="17">A. richardsonae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2075794FF16FFFAFCA7F952FF51F940" pageId="16" pageNumber="17" rank="species">sp. nov.</taxonomicNameLabel>
, was found in a leaf litter sample, photographed alive, and posted on iNaturalist by Sarah C. Crews (https://www.inaturalist.org/observations/484229; S.C. Crews, in litt. 
<date id="FFFE103DFF16FFFAFE3BF8F5FD9DF8B7" box="[408,550,1825,1854]" pageId="16" pageNumber="17" value="2021-09-24">24.IX.2021</date>
), from near Hardwar Gap(
<geoCoordinate id="EE74503AFF16FFFAFF57F89CFE23F8EC" box="[244,408,1864,1893]" degrees="18" direction="north" minutes="08" orientation="latitude" pageId="16" pageNumber="17" precision="1" seconds="05.6" value="18.13489">18°08’05.6”N</geoCoordinate>
, 
<geoCoordinate id="EE74503AFF16FFFAFE02F89CFDFFF8EC" box="[417,580,1864,1893]" degrees="76" direction="west" minutes="43" orientation="longitude" pageId="16" pageNumber="17" precision="1" seconds="48.0" value="-76.73">76°43’48.0”W</geoCoordinate>
; approximately 
<quantity id="4CB89B18FF16FFFAFD5BF89CFF77F805" metricMagnitude="2" metricUnit="m" metricValue="9.8" pageId="16" pageNumber="17" unit="m" value="980.0">980 m</quantity>
a.s.l.), Blue and John Crow Mountains National Park, 
<collectingRegion id="4984F81FFF16FFFAFF58F842FE02F83A" box="[251,441,1942,1971]" country="Jamaica" name="Portland" pageId="16" pageNumber="17">Portland Parish</collectingRegion>
(
<figureCitation id="137B2A78FF16FFFAFE68F842FDA2F83A" box="[459,537,1942,1971]" captionStart="Figure 1" captionStartId="3.[181,252,1611,1634]" captionTargetBox="[181,1467,151,1587]" captionTargetId="figure-19@3.[181,1467,151,1587]" captionTargetPageId="3" captionText="Figure 1. Distribution of the genus Acanthoniscus in Jamaica. Above: type locality of A. spiniger at the hills near Bluefields (yellow dot), the type locality of A.richardsonae sp.nov.in the Blue Mountains (red dot), and a second location in the Blue Mountains where a specimen presumably of A. richardsonae sp. nov. was photographed (orange dot). The main collection sites of Henry G. Hubbard in 1877 are depicted by green dots, notice the proximity of Stony Hill to the distribution of A. richardsonae sp.nov.Below: The hills known as Surinam Quarters near Bluefields, referred by Gosse (1851) as “Bluefields Mountain,” Westmoreland Parish, constitute the type locality of A. spiniger." figureDoi="http://doi.org/10.5281/zenodo.10716766" httpUri="https://zenodo.org/record/10716766/files/figure.png" pageId="16" pageNumber="17">Figs. 1</figureCitation>
, 
<figureCitation id="137B2A78FF16FFFAFD86F842FDFEF83A" box="[549,581,1942,1971]" captionStart="Figure 10" captionStartId="17.[180,251,963,986]" captionTargetBox="[181,805,151,940]" captionTargetId="figure-510@17.[181,805,148,943]" captionTargetPageId="17" captionText="Figure 10. Specimen of Acanthoniscus cf. richardsonae sp. nov. observed near Hardwar Gap, Blue and John Crow Mountains National Park, Portland Parish. These are the only available photographs of a live specimen of the genus Acanthoniscus. Photos by Sarah C. Crews." figureDoi="http://doi.org/10.5281/zenodo.10716786" httpUri="https://zenodo.org/record/10716786/files/figure.png" pageId="16" pageNumber="17">10</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C35A6576FF16FFFAFCD2FB66FB2BFA56" pageId="16" pageNumber="17" type="etymology">
<paragraph id="8BFF36FDFF16FFFAFCD2FB66FB2BFA56" blockId="16.[843,1468,1202,1503]" pageId="16" pageNumber="17">
<emphasis id="B934EAEFFF16FFFAFCD2FB66FC54FB46" box="[881,1007,1202,1231]" italics="true" pageId="16" pageNumber="17">Etymology</emphasis>
. The selected epithet is a Latinized patronym honoring Harriet Richardson Searle (1874–1958), 
<collectingRegion id="4984F81FFF16FFFAFC54FAD4FB6AFA94" box="[1015,1233,1280,1309]" country="United States of America" name="District of Columbia" pageId="16" pageNumber="17">Washington D.C.</collectingRegion>
, 
<collectingCountry id="F357766DFF16FFFAFB42FAD4FA8FFA94" box="[1249,1332,1280,1309]" name="United States of America" pageId="16" pageNumber="17">U.S.A.</collectingCountry>
, known as the first lady of isopods and one of the first women carcinologists, who made important contributions to isopod taxonomy and made the first description and illustrations of 
<taxonomicName id="4C404D7EFF16FFFAFB92FA48FB5EFA31" authority="Rodríguez-Cabrera &amp; de Armas, 2023" authorityName="Rodríguez-Cabrera &amp; de Armas" authorityYear="2023" box="[1073,1253,1435,1464]" class="Malacostraca" family="Delatorreiidae" genus="Acanthoniscus" kingdom="Animalia" order="Isopoda" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="richardsonae" status="sp. nov.">
<emphasis id="B934EAEFFF16FFFAFB92FA48FB5EFA31" box="[1073,1253,1435,1464]" italics="true" pageId="16" pageNumber="17">A. richardsonae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2075794FF16FFFAFB49FA48FAFFFA31" box="[1258,1348,1436,1464]" pageId="16" pageNumber="17" rank="species">sp. nov.</taxonomicNameLabel>
, although misidentified as 
<emphasis id="B934EAEFFF16FFFAFBB2FA17FB2BFA56" box="[1041,1168,1475,1503]" italics="true" pageId="16" pageNumber="17">
<taxonomicName id="4C404D7EFF16FFFAFBB2FA17FB37FA56" box="[1041,1164,1475,1503]" class="Malacostraca" family="Oniscidae" genus="Oniscus" kingdom="Animalia" order="Isopoda" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="spiniger">A. spiniger</taxonomicName>
.
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C35A6576FF16FFFAFCD2F9C4FBADF83A" pageId="16" pageNumber="17" type="description">
<paragraph id="8BFF36FDFF16FFFAFCD2F9C4FBADF83A" blockId="16.[843,1468,1552,1971]" pageId="16" pageNumber="17">
<emphasis id="B934EAEFFF16FFFAFCD2F9C4FC09F9A4" box="[881,946,1552,1581]" italics="true" pageId="16" pageNumber="17">Color</emphasis>
(in alcohol) (
<figureCitation id="137B2A78FF16FFFAFBFDF9C4FB1FF9A4" box="[1118,1188,1552,1581]" captionStart="Figure 3" captionStartId="10.[843,913,586,609]" captionTargetBox="[843,1467,151,563]" captionTargetId="figure-639@10.[843,1467,148,565]" captionTargetPageId="10" captionText="Figure 3. Right lateral view of a female paratype Acanthoniscus richardsonaesp.nov.showingcolorationinalcohol.Scale bar=1 mm." figureDoi="http://doi.org/10.5281/zenodo.10716770" httpUri="https://zenodo.org/record/10716770/files/figure.png" pageId="16" pageNumber="17">Fig. 3</figureCitation>
). Grayish-brown, with muscle spots whitish-beige on dorsal surface of cephalothorax and on dorsolateral region of pereon tergites, around the base of largest dorsolateral spines; dorsal surface of pereon epimera and pleon tergites uniformly grayish-brown; dorsal surface of pereon epimera 1 sometimes spotted on dark gray. Second antenna uniformly light brown. Pereiopods grayish-brown irregularly spotted on whitish-beige. Apparently, the coloration in life is similar to that in alcohol (
<figureCitation id="137B2A78FF16FFFAFC16F841FBB3F83A" box="[949,1032,1941,1971]" captionStart="Figure 10" captionStartId="17.[180,251,963,986]" captionTargetBox="[181,805,151,940]" captionTargetId="figure-510@17.[181,805,148,943]" captionTargetPageId="17" captionText="Figure 10. Specimen of Acanthoniscus cf. richardsonae sp. nov. observed near Hardwar Gap, Blue and John Crow Mountains National Park, Portland Parish. These are the only available photographs of a live specimen of the genus Acanthoniscus. Photos by Sarah C. Crews." figureDoi="http://doi.org/10.5281/zenodo.10716786" httpUri="https://zenodo.org/record/10716786/files/figure.png" pageId="16" pageNumber="17">Fig. 10</figureCitation>
).
</paragraph>
</subSubSection>
<caption id="DF3F6675FF17FFFBFF17FC17FE06FBD2" ID-DOI="http://doi.org/10.5281/zenodo.10716786" ID-Zenodo-Dep="10716786" httpUri="https://zenodo.org/record/10716786/files/figure.png" pageId="17" pageNumber="18" startId="17.[180,251,963,986]" targetBox="[181,805,151,940]" targetPageId="17" targetType="figure">
<paragraph id="8BFF36FDFF17FFFBFF17FC17FE06FBD2" blockId="17.[180,806,963,1115]" pageId="17" pageNumber="18">
<emphasis id="B934EAEFFF17FFFBFF17FC17FE98FC53" bold="true" box="[180,291,963,986]" pageId="17" pageNumber="18">Figure 10.</emphasis>
Specimen of 
<taxonomicName id="4C404D7EFF17FFFBFE11FC17FD6FFC52" authority="Rodríguez-Cabrera &amp; de Armas, 2023" authorityName="Rodríguez-Cabrera &amp; de Armas" authorityYear="2023" box="[434,724,963,987]" class="Malacostraca" family="Delatorreiidae" genus="Acanthoniscus" isUncertain="true" kingdom="Animalia" order="Isopoda" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="richardsonae" status="sp. nov.">
<emphasis id="B934EAEFFF17FFFBFE11FC17FD81FC52" box="[434,570,963,987]" italics="true" pageId="17" pageNumber="18">Acanthoniscus</emphasis>
cf. 
<emphasis id="B934EAEFFF17FFFBFDF8FC17FD6FFC52" box="[603,724,963,987]" italics="true" pageId="17" pageNumber="18">richardsonae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2075794FF17FFFBFD78FC17FC9DFC53" box="[731,806,963,986]" pageId="17" pageNumber="18" rank="species">sp. nov.</taxonomicNameLabel>
observed near Hardwar Gap, Blue and John Crow Mountains National Park, Portland Parish. These are the only available photographs of a live specimen of the genus 
<taxonomicName id="4C404D7EFF17FFFBFD31FBF7FCA4FBB2" box="[658,799,1059,1083]" class="Malacostraca" family="Delatorreiidae" genus="Acanthoniscus" kingdom="Animalia" order="Isopoda" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
<emphasis id="B934EAEFFF17FFFBFD31FBF7FCA4FBB2" box="[658,799,1059,1083]" italics="true" pageId="17" pageNumber="18">Acanthoniscus</emphasis>
</taxonomicName>
. Photos by Sarah C. Crews.
</paragraph>
</caption>
<subSubSection id="C35A6576FF17FFFBFF78FB45FE3AF97E" pageId="17" pageNumber="18" type="discussion">
<paragraph id="8BFF36FDFF17FFFBFF78FB45FE3AF97E" blockId="17.[179,806,1169,1979]" pageId="17" pageNumber="18">
<emphasis id="B934EAEFFF17FFFBFF78FB45FEF8FB27" box="[219,323,1169,1198]" italics="true" pageId="17" pageNumber="18">Remarks</emphasis>
. The isopods of the genus 
<taxonomicName id="4C404D7EFF17FFFBFDDDFB45FC9EFB27" box="[638,805,1169,1198]" class="Malacostraca" family="Delatorreiidae" genus="Acanthoniscus" kingdom="Animalia" order="Isopoda" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
<emphasis id="B934EAEFFF17FFFBFDDDFB45FC9EFB27" box="[638,805,1169,1198]" italics="true" pageId="17" pageNumber="18">Acanthoniscus</emphasis>
</taxonomicName>
are very fragile, therefore, most of the available specimens are fragmented, some of them severely damaged. Our description of 
<taxonomicName id="4C404D7EFF17FFFBFDB7FAD3FD73FAAA" authority="Rodríguez-Cabrera &amp; de Armas, 2023" authorityName="Rodríguez-Cabrera &amp; de Armas" authorityYear="2023" box="[532,712,1286,1315]" class="Malacostraca" family="Delatorreiidae" genus="Acanthoniscus" kingdom="Animalia" order="Isopoda" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="richardsonae" status="sp. nov.">
<emphasis id="B934EAEFFF17FFFBFDB7FAD3FD73FAAA" box="[532,712,1286,1315]" italics="true" pageId="17" pageNumber="18">A. richardsonae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2075794FF17FFFBFD6EFAD3FC9EFAAA" box="[717,805,1287,1315]" pageId="17" pageNumber="18" rank="species">sp. nov.</taxonomicNameLabel>
largely coincides with that of 
<bibRefCitation id="EFD14B0CFF17FFFBFDAEFAF9FD49FAC3" author="Richardson H" box="[525,754,1325,1354]" pageId="17" pageNumber="18" pagination="431 - 434" refId="ref17042" refString="Richardson H 1909. The isopod crustacean Acanthoniscus spiniger Kinahan redescribed. Proceedings of the United States National Museum, 36: 431 - 434. DOI: https: // www. doi. org / 10.5479 / si. 00963801.36 - 1678.431" type="journal article" year="1909">Richardson (1909)</bibRefCitation>
, but since the precise locality of her specimen is unknown, we ignore whether its origin is the same as that of the 
<typeStatus id="54FB885FFF17FFFBFF45FA77FEA5FA36" box="[230,286,1443,1471]" pageId="17" pageNumber="18">type</typeStatus>
series of 
<taxonomicName id="4C404D7EFF17FFFBFE35FA77FDEAFA36" authority="Rodríguez-Cabrera &amp; de Armas, 2023" authorityName="Rodríguez-Cabrera &amp; de Armas" authorityYear="2023" box="[406,593,1442,1471]" class="Malacostraca" family="Delatorreiidae" genus="Acanthoniscus" kingdom="Animalia" order="Isopoda" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="richardsonae" status="sp. nov.">
<emphasis id="B934EAEFFF17FFFBFE35FA77FDEAFA36" box="[406,593,1442,1471]" italics="true" pageId="17" pageNumber="18">A. richardsonae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2075794FF17FFFBFDF8FA77FD03FA36" box="[603,696,1443,1471]" pageId="17" pageNumber="18" rank="species">sp. nov.</taxonomicNameLabel>
(eastern 
<collectingCountry id="F357766DFF17FFFBFF16FA1DFEA0FA6F" box="[181,283,1481,1510]" name="Jamaica" pageId="17" pageNumber="18">Jamaica</collectingCountry>
). 
<bibRefCitation id="EFD14B0CFF17FFFBFE92FA1DFDE7FA6F" author="Richardson H" box="[305,604,1481,1510]" pageId="17" pageNumber="18" pagination="431 - 434" refId="ref17042" refString="Richardson H 1909. The isopod crustacean Acanthoniscus spiniger Kinahan redescribed. Proceedings of the United States National Museum, 36: 431 - 434. DOI: https: // www. doi. org / 10.5479 / si. 00963801.36 - 1678.431" type="journal article" year="1909">Richardson (1909: 432)</bibRefCitation>
mentioned that her specimen had the cephalothorax “with the front emarginate and the lateral angles acutely produced” (
<figureCitation id="137B2A78FF17FFFBFF62F9EAFEA6F9D2" box="[193,285,1598,1627]" captionStart="Figure 4" captionStartId="11.[181,252,1628,1651]" captionTargetBox="[246,1407,152,1593]" captionTargetId="figure-20@11.[229,1420,148,1608]" captionTargetPageId="11" captionText="Figure 4. Body parts of Acanthoniscus richardsonae sp. nov. A, Overall, dorsal view; B, detail of the microsculpture of the dorsal tegument (left) and tricorn-like setae (right); C, lateral view of the anterior region of the body;D, lateral view of the posterior region of the body; E, right pleon-epimera 3–5 in ventral view; F, right pereon-epimera 1–7 in ventral view; G, detail of pereon-epimera 1–2 in a more ventral position; H, cephalothorax in dorsal view; I, cephalothorax in frontal view (the tiny sensory setae inserted between the ommatidia are illustrated under a magnification of 400×: a, b, c); J, cephalothorax in ventral view; K, pleotelson and uropods in dorsal view (the uropod exopodites are slightly pointing upward); L, pleotelson and uropods in ventral view; M, right uropod in ventral view; N, right uropod in dorsal view; O, drawing of the overall dorsal view modified from Richardson (1909); P, arrangement of dorsal spines, the diametersof the circles correspond tothe relative size of the spines.Drawings of the corresponding structures modified from Richardson (1909)are included in I,J (the question mark means that the position representing thedrawing is not totally clear) and in M. Abbreviations: sympodite (sy), endopodite (en), and exopodite (ex). Scale bars = 1 mm (A–L), 0.5 mm (M–N); in some cases, a single scale bar applies to more than one figure (indicated by the letters above the scale bar)." figureDoi="http://doi.org/10.5281/zenodo.10716774" httpUri="https://zenodo.org/record/10716774/files/figure.png" pageId="17" pageNumber="18">Fig. 4O</figureCitation>
). However, since all other overall characters she described coincide with the description of 
<taxonomicName id="4C404D7EFF17FFFBFCAAF9B2FEF3F920" authority="Rodríguez-Cabrera &amp; de Armas, 2023" authorityName="Rodríguez-Cabrera &amp; de Armas" authorityYear="2023" class="Malacostraca" family="Delatorreiidae" genus="Acanthoniscus" kingdom="Animalia" order="Isopoda" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="richardsonae" status="sp. nov.">
<emphasis id="B934EAEFFF17FFFBFCAAF9B2FEF3F920" italics="true" pageId="17" pageNumber="18">A. richardsonae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2075794FF17FFFBFEEEF959FE1DF920" box="[333,422,1677,1705]" pageId="17" pageNumber="18" rank="species">sp. nov.</taxonomicNameLabel>
, we suspect that a different angle while making the drawings could be the cause for such differences.
</paragraph>
</subSubSection>
<subSubSection id="C35A6576FF17FFFBFF78F8D6FB2FFD52" pageId="17" pageNumber="18" type="conservation">
<paragraph id="8BFF36FDFF17FFFBFF78F8D6FB2FFD52" blockId="17.[179,806,1169,1979]" lastBlockId="17.[841,1468,149,731]" pageId="17" pageNumber="18">
<emphasis id="B934EAEFFF17FFFBFF78F8D6FE08F897" box="[219,435,1794,1822]" italics="true" pageId="17" pageNumber="18">Conservation status</emphasis>
. Apparently, both 
<taxonomicName id="4C404D7EFF17FFFBFD20F8D6FD4EF897" box="[643,757,1794,1822]" class="Malacostraca" family="Oniscidae" genus="Oniscus" kingdom="Animalia" order="Isopoda" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="spiniger">
<emphasis id="B934EAEFFF17FFFBFD20F8D6FD4EF897" box="[643,757,1794,1822]" italics="true" pageId="17" pageNumber="18">A.spiniger</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C404D7EFF17FFFBFF10F8FDFED2F8CC" authority="Rodríguez-Cabrera &amp; de Armas, 2023" authorityName="Rodríguez-Cabrera &amp; de Armas" authorityYear="2023" box="[179,361,1832,1861]" class="Malacostraca" family="Delatorreiidae" genus="Acanthoniscus" kingdom="Animalia" order="Isopoda" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="richardsonae" status="sp. nov.">
<emphasis id="B934EAEFFF17FFFBFF10F8FDFED2F8CC" box="[179,361,1832,1861]" italics="true" pageId="17" pageNumber="18">A. richardsonae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2075794FF17FFFBFED2F8FDFE70F8CC" box="[369,459,1833,1861]" pageId="17" pageNumber="18" rank="species">sp. nov.</taxonomicNameLabel>
have restricted distributions (known only from a single or two localities), with severely fragmented and reduced habitats. However, since only three collection events (two in the mid-19 
<superScript id="7C359BB5FF17FFFBFCCAFF41FCC2FF2F" attach="left" box="[873,889,149,166]" fontSize="7" pageId="17" pageNumber="18">th</superScript>
century) and an additional observation exist, it would be premature to make an evaluation of their conservation statuses based on the available information. Therefore, we tentatively consider them as Data Deficient (
<bibRefCitation id="EFD14B0CFF17FFFBFB90FEE1FBD8FEF0" author="IUCN Standards &amp; Petitions Subcommittee" pageId="17" pageNumber="18" refId="ref16008" refString="IUCN Standards and Petitions Subcommittee 2019. Guidelines for Using the IUCN Red List Categories and Criteria. Version 14. Prepared by the Standards and Petitions Subcommittee, 113 pp. Available at https: // www. iucnredlist. org / documents / RedListGuidelines. pdf. Accessed on 1 December 2021." type="book" year="2019">IUCN Standards and Petitions Subcommittee, 2019</bibRefCitation>
). The 
<typeStatus id="54FB885FFF17FFFBFB63FE89FB41FEF0" box="[1216,1274,349,377]" pageId="17" pageNumber="18">type</typeStatus>
locality of 
<taxonomicName id="4C404D7EFF17FFFBFA3CFE88FC5BFE29" authority="Rodríguez-Cabrera &amp; de Armas, 2023" authorityName="Rodríguez-Cabrera &amp; de Armas" authorityYear="2023" class="Malacostraca" family="Delatorreiidae" genus="Acanthoniscus" kingdom="Animalia" order="Isopoda" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="richardsonae" status="sp. nov.">
<emphasis id="B934EAEFFF17FFFBFA3CFE88FC5BFE29" italics="true" pageId="17" pageNumber="18">A. richardsonae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2075794FF17FFFBFC4BFE50FBFFFE29" box="[1000,1092,388,416]" pageId="17" pageNumber="18" rank="species">sp. nov.</taxonomicNameLabel>
falls within the Blue and John Crow Mountains National Park (see Protected Areas System Master Plan: 
<collectingCountry id="F357766DFF17FFFBFBEEFE06FB16FE66" box="[1101,1197,466,495]" name="Jamaica" pageId="17" pageNumber="18">Jamaica</collectingCountry>
2013–2017; Jamaica’s National Ecological Gap Assessment Report, 2009), but that of 
<taxonomicName id="4C404D7EFF17FFFBFC6BFDF5FBFBFDB7" box="[968,1088,545,574]" class="Malacostraca" family="Oniscidae" genus="Oniscus" kingdom="Animalia" order="Isopoda" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="spiniger">
<emphasis id="B934EAEFFF17FFFBFC6BFDF5FBFBFDB7" box="[968,1088,545,574]" italics="true" pageId="17" pageNumber="18">A. spiniger</emphasis>
</taxonomicName>
is not included in any protected area.Additional surveys are required to collect data on their distributions, ecology, demographic parameters and threats, in order to make accurate assessments of their conservation statuses.
</paragraph>
</subSubSection>
</treatment>
</document>