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<document id="165BA34806ECB3A253B18259979DB0B6" ID-CLB-Dataset="26202" ID-DOI="10.11646/zootaxa.4630.1.1" ID-GBIF-Dataset="db6a16a6-0fe8-4987-8ad5-42223825fcd7" ID-ISSN="1175-5326" ID-Zenodo-Dep="3336635" ID-ZooBank="8D375836-CCBA-473C-836F-6ABD44B4F881" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1563153150055" checkinUser="plazi" docAuthor="Olmi, Massimo, Copeland, Robert S. &amp; Noort, Simon Van" docDate="2019" docId="03E987E2FEAC2B4BFF3ED44FFD25F918" docLanguage="en" docName="zootaxa.4630.1.1.pdf.imf" docOrigin="Zootaxa 4630 (1)" docStyle="DocumentStyle:D239614CE4198176A422035174489AB1.4:Zootaxa.2009-2012.monograph" docStyleId="D239614CE4198176A422035174489AB1" docStyleName="Zootaxa.2009-2012.monograph" docStyleVersion="4" docTitle="Bocchinae Richards. 1939" docType="treatment" docVersion="39" lastPageNumber="260" masterDocId="FFD0FF9AFFAF2A48FFA9D64EFFE1FF87" masterDocTitle="Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea)" masterLastPageNumber="619" masterPageNumber="1" pageNumber="260" updateTime="1698741107231" updateUser="ExternalLinkService">
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<mods:title id="3EE4A864827A2A9AE972225F75DA3970">Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea)</mods:title>
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<mods:namePart id="00AF923AF4AB47034FC476C6C2BD670A">Copeland, Robert S.</mods:namePart>
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<mods:namePart id="27AC9F7E048189D50FDAF4DF0AC289DC">Noort, Simon Van</mods:namePart>
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IV. Subfamily
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Bocchinae
<bibRefCitation id="EFD14B05FEAC2B4BFE7AD44FFD6EFD9C" author="Richards, O. W." box="[467,655,513,539]" pageId="259" pageNumber="260" pagination="185 - 344" refId="ref526240" refString="Richards, O. W. (1939) The British Bethylidae (s. l.) (Hymenoptera). The Transactions of the Royal entomological Society of London, 89, 185 - 344. https: // doi. org / 10.1111 / j. 1365 - 2311.1939. tb 00740. x" type="journal article" year="1939">Richards, 1939</bibRefCitation>
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<paragraph id="8BFF36F4FEAC2B4BFF3ED40BFC04FDDA" blockId="259.[151,1436,581,698]" box="[151,997,581,606]" pageId="259" pageNumber="260">
Bocchini
<bibRefCitation id="EFD14B05FEAC2B4BFF50D40BFE23FDDA" author="Richards, O. W." box="[249,450,581,606]" pageId="259" pageNumber="260" pagination="185 - 344" refId="ref526240" refString="Richards, O. W. (1939) The British Bethylidae (s. l.) (Hymenoptera). The Transactions of the Royal entomological Society of London, 89, 185 - 344. https: // doi. org / 10.1111 / j. 1365 - 2311.1939. tb 00740. x" type="journal article" year="1939">Richards 1939: 189</bibRefCitation>
;
<bibRefCitation id="EFD14B05FEAC2B4BFE64D40BFCD1FDDA" author="Muesebeck, C. F. W. &amp; Walkley, L. M." box="[461,816,581,606]" pageId="259" pageNumber="260" pagination="1034 - 1043" refId="ref520885" refString="Muesebeck, C. F. W. &amp; Walkley, L. M. (1951) Family Dryinidae. In: Muesebeck, C. F. W., Krombein, K. V. &amp; Townes, H. K. (Eds.), Hymenoptera of America North of Mexico. Synoptic catalogue. Agriculture Monograph 2. U. S. Department of Agriculture, Washington, D. C., pp. 1034 - 1043." type="book chapter" year="1951">Muesebeck &amp; Walkley 1951: 1039</bibRefCitation>
;
<bibRefCitation id="EFD14B05FEAC2B4BFC93D408FC00FDDA" author="Nagy, C." box="[826,993,581,606]" pageId="259" pageNumber="260" pagination="331 - 337" refId="ref521025" refString="Nagy, C. (1967) Contribution l'etude de la fam. Dryinidae (Hym.) dans la faune de la Roumanie. Travaux du Museum d'Histoire Naturelle Grigore Antipa, 7, 331 - 337." type="journal article" year="1967">Nagy 1967: 334</bibRefCitation>
.
</paragraph>
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<taxonomicName id="4C404D77FEAC2B4BFF3ED42DFE48FDFB" authority="Olmi 1984: 599" authorityName="Olmi" authorityPageNumber="599" authorityYear="1984" box="[151,425,611,636]" class="Insecta" family="Dryinidae" kingdom="Animalia" order="Hymenoptera" pageId="259" pageNumber="260" phylum="Arthropoda" rank="subFamily" subFamily="Bocchinae">
Bocchinae
<bibRefCitation id="EFD14B05FEAC2B4BFEA1D42DFE48FDFB" author="Olmi, M." box="[264,425,611,636]" pageId="259" pageNumber="260" pagination="1 - 1913" refId="ref521356" refString="Olmi, M. (1984) A revision of the Dryinidae (Hymenoptera). Memoirs of the American Entomological Institute, 37, I-XII + 1 - 1913." type="journal article" year="1984">Olmi 1984: 599</bibRefCitation>
</taxonomicName>
;
<bibRefCitation id="EFD14B05FEAC2B4BFE1AD42AFDC7FDFB" author="Olmi, M." box="[435,550,612,636]" pageId="259" pageNumber="260" pagination="186 - 200" refId="ref521771" refString="Olmi, M. (1993 a) Dryinidae di Costa Rica: catalogo e considerazioni biogeografiche ed evolutive (Hym. Chrysidoidea). Bollettino della Societa entomologica italiana, Genova, 124, 186 - 200." type="journal article" year="1993">1993a: 193</bibRefCitation>
;
<bibRefCitation id="EFD14B05FEAC2B4BFD99D42AFD77FDFB" author="Olmi, M." box="[560,662,611,636]" pageId="259" pageNumber="260" pagination="19 - 62" refId="ref521856" refString="Olmi, M. (1993 c [&quot; 1992 &quot;]) Descriptions of new taxa of Dryinidae (Hymenoptera Chrysidoidea). Frustula entomologica, New Series, 15, 19 - 62." type="journal article" year="1993">1993c: 46</bibRefCitation>
;
<bibRefCitation id="EFD14B05FEAC2B4BFD09D42DFCF4FDFB" author="Olmi, M." box="[672,789,611,636]" pageId="259" pageNumber="260" pagination="493 - 503" refId="ref522044" refString="Olmi, M. (1995 b) Dryinidae. In: Hanson, P. E. &amp; Gauld, I. D. (Eds.), The Hymenoptera of Costa Rica. Oxford University Press, Oxford, pp. 493 - 503." type="book chapter" year="1995">1995b: 502</bibRefCitation>
;
<bibRefCitation id="EFD14B05FEAC2B4BFCB6D42AFC66FDFB" author="Olmi, M." box="[799,903,611,636]" pageId="259" pageNumber="260" refId="ref522549" refString="Olmi, M. (1999) Hymenoptera Dryinidae-Embolemidae. Fauna d'Italia. Vol. 37. Edizioni Calderini, Bologna, 425 pp." type="book" year="1999">1999: 152</bibRefCitation>
;
<bibRefCitation id="EFD14B05FEAC2B4BFC39D42DFB6DFDFB" author="Olmi, M. &amp; Bechly, G." box="[912,1164,611,636]" pageId="259" pageNumber="260" pagination="1 - 58" refId="ref523628" refString="Olmi, M. &amp; Bechly, G. (2001) New parasitic wasps from Baltic amber (Insecta: Hymenoptera: Dryinidae). Stuttgarter Beitrage zur Naturkunde, Serie B, Nr. 306, 1 - 58." type="journal article" year="2001">Olmi &amp; Bechly 2001: 41</bibRefCitation>
;
<bibRefCitation id="EFD14B05FEAC2B4BFB3FD42AFABFFDFB" author="He, J. &amp; Xu, Z." box="[1174,1374,612,636]" pageId="259" pageNumber="260" refId="ref518576" refString="He, J. &amp; Xu, Z. (2002) Hymenoptera Dryinidae. Fauna Sinica. Vol. 29. Science Press, Beijing, 464 pp." type="book" year="2002">He &amp; Xu 2002: 237</bibRefCitation>
;
<bibRefCitation id="EFD14B05FEAC2B4BFAC1D42DFE62FD1C" author="Olmi, M. &amp; Virla, E. G." pageId="259" pageNumber="260" pagination="401 - 418" refId="ref524721" refString="Olmi, M. &amp; Virla, E. G. (2006) Familia Dryinidae. In: Fernandez, F. &amp; Sharkey, M. J. (Eds.), Introduccion a los Hymenoptera de la Region Neotropical. Sociedad Colombiana de Entomologia y Universidad Nacional de Colombia, Bogota D. C., pp. 401 - 418." type="book chapter" year="2006">Olmi &amp; Virla 2006: 408</bibRefCitation>
;
<bibRefCitation id="EFD14B05FEAC2B4BFE24D4CCFD64FD1C" author="Virla, E. G. &amp; Olmi, M." box="[397,645,642,667]" pageId="259" pageNumber="260" pagination="357 - 372" refId="ref526905" refString="Virla, E. G. &amp; Olmi, M. (2008) Dryinidae. In: Claps, L. E., Debandi, G. &amp; Roig-Junent, S. (Eds.), Biodiversidad de Artropodos Argentinos. Vol. 2. Sociedad Entomologica Argentina, Mendoza, pp. 357 - 372." type="book chapter" year="2008">Virla &amp; Olmi 2008: 370</bibRefCitation>
;
<bibRefCitation id="EFD14B05FEAC2B4BFD26D4CDFCB7FD1C" author="Xu, Z. &amp; Olmi, M. &amp; He, J." box="[655,854,642,667]" pageId="259" pageNumber="260" pagination="1 - 460" refId="ref527627" refString="Xu, Z., Olmi, M. &amp; He, J. (2013) Dryinidae of the Oriental region (Hymenoptera: Chrysidoidea). Zootaxa, 3614 (1), 1 - 460. https: // doi. org / 10.11646 / zootaxa. 3614.1.1" type="journal article" year="2013">Xu et al. 2013: 223</bibRefCitation>
;
<bibRefCitation id="EFD14B05FEAC2B4BFCC8D4CCFBB6FD1C" author="Olmi, M. &amp; Virla, E. G." box="[865,1111,642,667]" pageId="259" pageNumber="260" pagination="1 - 534" refId="ref524787" refString="Olmi, M. &amp; Virla, E. G. (2014) Dryinidae of the Neotropical region (Hymenoptera: Chrysidoidea). Zootaxa, 3792, 1 - 534. https: // doi. org / 10.11646 / zootaxa. 3792.1.1" type="journal article" year="2014">Olmi &amp; Virla 2014: 206</bibRefCitation>
;
<bibRefCitation id="EFD14B05FEAC2B4BFBCBD4CCFAA5FD1C" author="Olmi, M. &amp; Xu, Z." box="[1122,1348,642,667]" pageId="259" pageNumber="260" pagination="1 - 253" refId="ref524873" refString="Olmi, M. &amp; Xu, Z. (2015) Dryinidae of the Eastern Palaearctic region (Hymenoptera: Chrysidoidea). Zootaxa, 3996 (1), 1 - 253. https: // doi. org / 10.11646 / zootaxa. 3996.1.1" type="journal article" year="2015">Olmi &amp; Xu 2015: 116</bibRefCitation>
;
<bibRefCitation id="EFD14B05FEAC2B4BFAE6D4CCFEA5FD3E" author="Olmi, M. &amp; van Noort, S. &amp; Guglielmino, A." pageId="259" pageNumber="260" pagination="45 - 95" refId="ref524656" refString="Olmi, M., van Noort, S. &amp; Guglielmino, A. (2016) Contribution to the knowledge of Afrotropical Dryinidae, Embolemidae and Sclerogibbidae (Hymenoptera), with description of new species from Central African Republic and Uganda. ZooKeys, 578, 45 - 95. https: // doi. org / 10.3897 / zookeys. 578.7820" type="journal article" year="2016">
Olmi
<emphasis id="B934EAE6FEAC2B4BFA23D4CDFF3AFD3E" italics="true" pageId="259" pageNumber="260">et al</emphasis>
. 2016: 71
</bibRefCitation>
.
</paragraph>
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<emphasis id="B934EAE6FEAC2B4BFF3ED491FEF0FD70" bold="true" box="[151,273,735,759]" pageId="259" pageNumber="260">
<typeStatus id="54FB8856FEAC2B4BFF3ED491FF2DFD70" box="[151,204,735,759]" pageId="259" pageNumber="260">Type</typeStatus>
genus
</emphasis>
.
<taxonomicName id="4C404D77FEAC2B4BFEB4D491FDF2FD70" authority="Ashmead 1893" authorityName="Ashmead" authorityYear="1893" box="[285,531,734,759]" class="Insecta" family="Dryinidae" genus="Bocchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="259" pageNumber="260" phylum="Arthropoda" rank="genus">
<emphasis id="B934EAE6FEAC2B4BFEB4D491FE92FD70" box="[285,371,735,759]" italics="true" pageId="259" pageNumber="260">Bocchus</emphasis>
<bibRefCitation id="EFD14B05FEAC2B4BFED1D490FDF2FD70" author="Ashmead, W. H." box="[376,531,734,759]" pageId="259" pageNumber="260" pagination="1 - 472" refId="ref514814" refString="Ashmead, W. H. (1893) Monograph of the North American Proctotrypidae. Bulletin of the United States National Museum, 45, 1 - 472. https: // doi. org / 10.5479 / si. 03629236.45.1" type="journal article" year="1893">Ashmead 1893</bibRefCitation>
</taxonomicName>
, designated by
<bibRefCitation id="EFD14B05FEAC2B4BFD19D490FCD0FD70" author="Olmi, M." box="[688,817,734,759]" pageId="259" pageNumber="260" pagination="1 - 1913" refId="ref521356" refString="Olmi, M. (1984) A revision of the Dryinidae (Hymenoptera). Memoirs of the American Entomological Institute, 37, I-XII + 1 - 1913." type="journal article" year="1984">Olmi (1984)</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C35A657FFEAC2B4BFF3ED56AFE66FAF8" pageId="259" pageNumber="260" type="diagnosis">
<paragraph id="8BFF36F4FEAC2B4BFF3ED56AFE66FAF8" blockId="259.[151,1437,804,1695]" pageId="259" pageNumber="260">
<emphasis id="B934EAE6FEAC2B4BFF3ED56AFEEBFCB8" bold="true" box="[151,266,804,831]" pageId="259" pageNumber="260">Diagnosis</emphasis>
.
<specimenCount id="9D46FD7DFEAC2B4BFEBED56BFECDFCB8" box="[279,300,805,831]" pageId="259" pageNumber="260" type="female">♀</specimenCount>
(
<figureCitation id="137B2A71FEAC2B4BFE93D56BFE33FCB8" box="[314,466,805,831]" captionStart="FIGURE 100" captionStartId="260.[151,250,1618,1643]" captionTargetBox="[151,1436,529,1605]" captionTargetId="figure@260.[151,1436,529,1606]" captionTargetPageId="260" captionText="FIGURE 100. Habitus of Bocchus species. ♀ in dorsal (A) and lateral (B) view. ♂ in lateral (C) and dorsal (D) view. Scale bar: A: 3.24 mm; B: 1.88 mm; C: 1.55 mm; D: 2.64 mm." httpUri="https://zenodo.org/record/3336870/files/figure.png" pageId="259" pageNumber="260">Figs 100A, B</figureCitation>
): fully winged (
<figureCitation id="137B2A71FEAC2B4BFD29D56BFCF9FCB8" box="[640,792,805,831]" captionStart="FIGURE 100" captionStartId="260.[151,250,1618,1643]" captionTargetBox="[151,1436,529,1605]" captionTargetId="figure@260.[151,1436,529,1606]" captionTargetPageId="260" captionText="FIGURE 100. Habitus of Bocchus species. ♀ in dorsal (A) and lateral (B) view. ♂ in lateral (C) and dorsal (D) view. Scale bar: A: 3.24 mm; B: 1.88 mm; C: 1.55 mm; D: 2.64 mm." httpUri="https://zenodo.org/record/3336870/files/figure.png" pageId="259" pageNumber="260">Figs 100A, B</figureCitation>
); occasionally brachypterous or micropterous or apterous; occipital carina complete; ocelli present; mandible with one, two, three or four teeth (
<figureCitation id="137B2A71FEAC2B4BFBC2D507FAD5FCE4" box="[1131,1332,841,867]" captionStart="FIGURE 8" captionStartId="20.[151,250,1364,1389]" captionTargetBox="[164,1424,185,1344]" captionTargetId="figure@20.[151,1436,174,1352]" captionTargetPageId="20" captionText="FIGURE 8. A: tridentate mandible progressing larger from anterior one to posterior. B: quadridentate mandible, with intermediate rudimentary tooth (i). C: unidentate mandible. D: tridentate mandible, with intermediate rudimentary tooth. E: ♂ genitalia of Gonatopus sp.. F: ♂ genitalia of Dryinus sp. G: hypopygium umbrella-shaped. H: hypopygium not umbrella-shaped. b: basivolsella; d: distivolsella; k: dorsal process of paramere; o: paramere; p: penis." httpUri="https://zenodo.org/record/3336666/files/figure.png" pageId="259" pageNumber="260">Figs 8A, B, C, D</figureCitation>
); quadridentate mandible usually with three large teeth and one rudimentary tooth between two posterior teeth (
<figureCitation id="137B2A71FEAC2B4BFA9CD523FA6FFC00" box="[1333,1422,877,903]" captionStart="FIGURE 8" captionStartId="20.[151,250,1364,1389]" captionTargetBox="[164,1424,185,1344]" captionTargetId="figure@20.[151,1436,174,1352]" captionTargetPageId="20" captionText="FIGURE 8. A: tridentate mandible progressing larger from anterior one to posterior. B: quadridentate mandible, with intermediate rudimentary tooth (i). C: unidentate mandible. D: tridentate mandible, with intermediate rudimentary tooth. E: ♂ genitalia of Gonatopus sp.. F: ♂ genitalia of Dryinus sp. G: hypopygium umbrella-shaped. H: hypopygium not umbrella-shaped. b: basivolsella; d: distivolsella; k: dorsal process of paramere; o: paramere; p: penis." httpUri="https://zenodo.org/record/3336666/files/figure.png" pageId="259" pageNumber="260">Fig. 8B</figureCitation>
); rarely quadridentate mandible with teeth becoming regularly progressively larger from dorsal to ventral tooth (as in
<figureCitation id="137B2A71FEAC2B4BFF3ED5FBFF0CFC57" box="[151,237,949,976]" captionStart="FIGURE 6" captionStartId="18.[151,250,1921,1946]" captionTargetBox="[161,1429,526,1897]" captionTargetId="figure@18.[151,1436,517,1909]" captionTargetPageId="18" captionText="FIGURE 6. A: antennal ADOs of Gonatopus sp. ♀. B: quadridentate mandible of Anteon sp. C: occiput of Gonatopus sp. ♂. D: micropterous ♀ of Deinodryinus sp. E: apterous ♀ of Gonatopus sp. a: propodeal declivity; c: mesoscutellum; m: metanotum; p: metapectal-propodeal disc; r: pronotum; s: mesoscutum." httpUri="https://zenodo.org/record/3336662/files/figure.png" pageId="259" pageNumber="260">Fig. 6B</figureCitation>
); palpal formula 6/3; antenna without tufts of long setae and without ADOs; pronotal tubercle present; epicnemium not visible, because lateral regions of prothorax continuous with mesopleura (as in
<figureCitation id="137B2A71FEAC2B4BFB03D597FAE3FC74" box="[1194,1282,985,1011]" captionStart="FIGURE 7" captionStartId="19.[151,250,1473,1498]" captionTargetBox="[163,1425,188,1453]" captionTargetId="figure@19.[151,1436,174,1461]" captionTargetPageId="19" captionText="FIGURE 7. A, B: chelae. C, D: mesosoma of Bocchinae (C) and Dryininae (D) ♂♂ in lateral view. a: arolium; e: enlarged claw; m: mesopleuron; p: pronotum; r: rudimentary claw; s: protarsomere 5; t: tegula; z: epicnemium. E: forewing of fictitious specimen (C: costal cell; cv: costal vein; Pt: pterostigma; R: radial cell; 1Cu: first cubital cell; 2Cu: second cubital cell; 2R1: second radial 1 cell; 2r-rs&amp;Rs: second radial cross &amp; radial sector vein; PostabR1: poststigmal abscissa of Radial1 vein)." httpUri="https://zenodo.org/record/3336664/files/figure.png" pageId="259" pageNumber="260">Fig. 7C</figureCitation>
); forewing of fully winged specimens with three cells enclosed by pigmented veins (C, R and 1Cu) (
<figureCitation id="137B2A71FEAC2B4BFBFBD5B3FB0BFB90" box="[1106,1258,1021,1047]" captionStart="FIGURE 100" captionStartId="260.[151,250,1618,1643]" captionTargetBox="[151,1436,529,1605]" captionTargetId="figure@260.[151,1436,529,1606]" captionTargetPageId="260" captionText="FIGURE 100. Habitus of Bocchus species. ♀ in dorsal (A) and lateral (B) view. ♂ in lateral (C) and dorsal (D) view. Scale bar: A: 3.24 mm; B: 1.88 mm; C: 1.55 mm; D: 2.64 mm." httpUri="https://zenodo.org/record/3336870/files/figure.png" pageId="259" pageNumber="260">Figs 100A, B</figureCitation>
); forewing with 2r-rs&amp;Rs vein (
<figureCitation id="137B2A71FEAC2B4BFEE1D26FFE07FBBC" box="[328,486,1057,1083]" captionStart="FIGURE 100" captionStartId="260.[151,250,1618,1643]" captionTargetBox="[151,1436,529,1605]" captionTargetId="figure@260.[151,1436,529,1606]" captionTargetPageId="260" captionText="FIGURE 100. Habitus of Bocchus species. ♀ in dorsal (A) and lateral (B) view. ♂ in lateral (C) and dorsal (D) view. Scale bar: A: 3.24 mm; B: 1.88 mm; C: 1.55 mm; D: 2.64 mm." httpUri="https://zenodo.org/record/3336870/files/figure.png" pageId="259" pageNumber="260">Figs 100A, B</figureCitation>
); rarely 2r-rs&amp;Rs vein absent; pterostigma of fully winged specimens present (
<figureCitation id="137B2A71FEAC2B4BFAC6D26FFF36FBD8" captionStart="FIGURE 100" captionStartId="260.[151,250,1618,1643]" captionTargetBox="[151,1436,529,1605]" captionTargetId="figure@260.[151,1436,529,1606]" captionTargetPageId="260" captionText="FIGURE 100. Habitus of Bocchus species. ♀ in dorsal (A) and lateral (B) view. ♂ in lateral (C) and dorsal (D) view. Scale bar: A: 3.24 mm; B: 1.88 mm; C: 1.55 mm; D: 2.64 mm." httpUri="https://zenodo.org/record/3336870/files/figure.png" pageId="259" pageNumber="260">Fig. 100B</figureCitation>
), often very reduced (
<figureCitation id="137B2A71FEAC2B4BFE63D20BFDA1FBD8" box="[458,576,1093,1119]" captionStart="FIGURE 100" captionStartId="260.[151,250,1618,1643]" captionTargetBox="[151,1436,529,1605]" captionTargetId="figure@260.[151,1436,529,1606]" captionTargetPageId="260" captionText="FIGURE 100. Habitus of Bocchus species. ♀ in dorsal (A) and lateral (B) view. ♂ in lateral (C) and dorsal (D) view. Scale bar: A: 3.24 mm; B: 1.88 mm; C: 1.55 mm; D: 2.64 mm." httpUri="https://zenodo.org/record/3336870/files/figure.png" pageId="259" pageNumber="260">Fig. 100A</figureCitation>
); protarsus chelate (
<figureCitation id="137B2A71FEAC2B4BFCB7D20BFC56FBD8" box="[798,951,1093,1119]" captionStart="FIGURE 100" captionStartId="260.[151,250,1618,1643]" captionTargetBox="[151,1436,529,1605]" captionTargetId="figure@260.[151,1436,529,1606]" captionTargetPageId="260" captionText="FIGURE 100. Habitus of Bocchus species. ♀ in dorsal (A) and lateral (B) view. ♂ in lateral (C) and dorsal (D) view. Scale bar: A: 3.24 mm; B: 1.88 mm; C: 1.55 mm; D: 2.64 mm." httpUri="https://zenodo.org/record/3336870/files/figure.png" pageId="259" pageNumber="260">Figs 100A, B</figureCitation>
); chela with rudimentary claw (
<figureCitation id="137B2A71FEAC2B4BFAB0D20BFA6EFBD8" box="[1305,1423,1093,1119]" captionStart="FIGURE 101" captionStartId="265.[151,250,1471,1496]" captionTargetBox="[155,1426,289,1451]" captionTargetId="figure@265.[151,1436,301,1459]" captionTargetPageId="265" captionText="FIGURE 101. Chelae of Bocchus species.A: B. bini Olmi, ♀ from Nigeria, Ibadan; B: B. botswanensis Olmi, ♀ from Botswa- na, Serowe; C: B. brooksi Olmi, holotype; D: B. hyalinus Olmi, holotype; E: B. kirkspriggsi Olmi, holotype; F: B. kogelbergensis Olmi, holotype. Scale bar: A, F: 0.08 mm; B, C: 0.09 mm; D: 0.07 mm; E: 0.10 mm." httpUri="https://zenodo.org/record/3336872/files/figure.png" pageId="259" pageNumber="260">Fig. 101A</figureCitation>
); protrochanter short and slightly longer than broad; tibial spurs 1/1/1, or 1/1/2.
<specimenCount id="9D46FD7DFEAC2B4BFB83D227FBDEFB04" box="[1066,1087,1129,1155]" pageId="259" pageNumber="260" type="male">♂</specimenCount>
(
<figureCitation id="137B2A71FEAC2B4BFBF8D227FB12FB04" box="[1105,1267,1129,1155]" captionStart="FIGURE 100" captionStartId="260.[151,250,1618,1643]" captionTargetBox="[151,1436,529,1605]" captionTargetId="figure@260.[151,1436,529,1606]" captionTargetPageId="260" captionText="FIGURE 100. Habitus of Bocchus species. ♀ in dorsal (A) and lateral (B) view. ♂ in lateral (C) and dorsal (D) view. Scale bar: A: 3.24 mm; B: 1.88 mm; C: 1.55 mm; D: 2.64 mm." httpUri="https://zenodo.org/record/3336870/files/figure.png" pageId="259" pageNumber="260">Figs 100C, D</figureCitation>
): fully winged (
<figureCitation id="137B2A71FEAC2B4BFF36D2C3FEDCFB20" box="[159,317,1165,1191]" captionStart="FIGURE 100" captionStartId="260.[151,250,1618,1643]" captionTargetBox="[151,1436,529,1605]" captionTargetId="figure@260.[151,1436,529,1606]" captionTargetPageId="260" captionText="FIGURE 100. Habitus of Bocchus species. ♀ in dorsal (A) and lateral (B) view. ♂ in lateral (C) and dorsal (D) view. Scale bar: A: 3.24 mm; B: 1.88 mm; C: 1.55 mm; D: 2.64 mm." httpUri="https://zenodo.org/record/3336870/files/figure.png" pageId="259" pageNumber="260">Figs 100C, D</figureCitation>
), rarely micropterous (
<figureCitation id="137B2A71FEAC2B4BFDE8D2C3FD52FB20" box="[577,691,1165,1191]" captionStart="FIGURE 107" captionStartId="277.[151,250,1650,1675]" captionTargetBox="[157,1430,178,1632]" captionTargetId="figure@277.[151,1436,174,1639]" captionTargetPageId="277" captionText="FIGURE 107. Habitus of Bocchus species. A–C: B. kibalensis Olmi, van Noort &amp; Guglielmino, ♂ holotype. D–F: B. kogelbergensis Olmi, ♀ from South Africa, Walker Bay. Scale bar: D: 1.48 mm." httpUri="https://zenodo.org/record/3336886/files/figure.png" pageId="259" pageNumber="260">Fig. 107E</figureCitation>
) or apterous (
<taxonomicName id="4C404D77FEAC2B4BFCFDD2C3FB14FB20" authority="Ponomarenko" authorityName="Ponomarenko" authorityYear="2000" box="[852,1269,1165,1191]" class="Insecta" family="Dryinidae" genus="Mystrophorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="259" pageNumber="260" phylum="Arthropoda" rank="species" species="apterus">
<emphasis id="B934EAE6FEAC2B4BFCFDD2C3FBACFB20" box="[852,1101,1165,1191]" italics="true" pageId="259" pageNumber="260">Mystrophorus apterus</emphasis>
Ponomarenko
</taxonomicName>
, not present in the Afrotropical region); occipital carina complete; mandible with one, two, three or four teeth (
<figureCitation id="137B2A71FEAC2B4BFB62D2FFFA6FFB4C" box="[1227,1422,1201,1227]" captionStart="FIGURE 8" captionStartId="20.[151,250,1364,1389]" captionTargetBox="[164,1424,185,1344]" captionTargetId="figure@20.[151,1436,174,1352]" captionTargetPageId="20" captionText="FIGURE 8. A: tridentate mandible progressing larger from anterior one to posterior. B: quadridentate mandible, with intermediate rudimentary tooth (i). C: unidentate mandible. D: tridentate mandible, with intermediate rudimentary tooth. E: ♂ genitalia of Gonatopus sp.. F: ♂ genitalia of Dryinus sp. G: hypopygium umbrella-shaped. H: hypopygium not umbrella-shaped. b: basivolsella; d: distivolsella; k: dorsal process of paramere; o: paramere; p: penis." httpUri="https://zenodo.org/record/3336666/files/figure.png" pageId="259" pageNumber="260">Figs 8A, B, C, D</figureCitation>
); quadridentate mandible usually with three large teeth and one rudimentary tooth between two posterior teeth (
<figureCitation id="137B2A71FEAC2B4BFAC6D29BFF5BFA94" captionStart="FIGURE 8" captionStartId="20.[151,250,1364,1389]" captionTargetBox="[164,1424,185,1344]" captionTargetId="figure@20.[151,1436,174,1352]" captionTargetPageId="20" captionText="FIGURE 8. A: tridentate mandible progressing larger from anterior one to posterior. B: quadridentate mandible, with intermediate rudimentary tooth (i). C: unidentate mandible. D: tridentate mandible, with intermediate rudimentary tooth. E: ♂ genitalia of Gonatopus sp.. F: ♂ genitalia of Dryinus sp. G: hypopygium umbrella-shaped. H: hypopygium not umbrella-shaped. b: basivolsella; d: distivolsella; k: dorsal process of paramere; o: paramere; p: penis." httpUri="https://zenodo.org/record/3336666/files/figure.png" pageId="259" pageNumber="260">Fig. 8B</figureCitation>
); palpal formula 6/3; epicnemium invisible, because lateral regions of prothorax continuous with mesopleura (
<figureCitation id="137B2A71FEAC2B4BFF09D353FF18FAB0" box="[160,249,1309,1335]" captionStart="FIGURE 7" captionStartId="19.[151,250,1473,1498]" captionTargetBox="[163,1425,188,1453]" captionTargetId="figure@19.[151,1436,174,1461]" captionTargetPageId="19" captionText="FIGURE 7. A, B: chelae. C, D: mesosoma of Bocchinae (C) and Dryininae (D) ♂♂ in lateral view. a: arolium; e: enlarged claw; m: mesopleuron; p: pronotum; r: rudimentary claw; s: protarsomere 5; t: tegula; z: epicnemium. E: forewing of fictitious specimen (C: costal cell; cv: costal vein; Pt: pterostigma; R: radial cell; 1Cu: first cubital cell; 2Cu: second cubital cell; 2R1: second radial 1 cell; 2r-rs&amp;Rs: second radial cross &amp; radial sector vein; PostabR1: poststigmal abscissa of Radial1 vein)." httpUri="https://zenodo.org/record/3336664/files/figure.png" pageId="259" pageNumber="260">Fig. 7C</figureCitation>
); in fully winged specimens forewing with three cells enclosed by pigmented veins (C, R and 1Cu) (
<figureCitation id="137B2A71FEAC2B4BFAC3D353FF19FADC" captionStart="FIGURE 100" captionStartId="260.[151,250,1618,1643]" captionTargetBox="[151,1436,529,1605]" captionTargetId="figure@260.[151,1436,529,1606]" captionTargetPageId="260" captionText="FIGURE 100. Habitus of Bocchus species. ♀ in dorsal (A) and lateral (B) view. ♂ in lateral (C) and dorsal (D) view. Scale bar: A: 3.24 mm; B: 1.88 mm; C: 1.55 mm; D: 2.64 mm." httpUri="https://zenodo.org/record/3336870/files/figure.png" pageId="259" pageNumber="260">Figs 100C, D</figureCitation>
); in fully winged specimens forewing with 2r-rs&amp;Rs vein (
<figureCitation id="137B2A71FEAC2B4BFC20D30FFBC3FADC" box="[905,1058,1345,1371]" captionStart="FIGURE 100" captionStartId="260.[151,250,1618,1643]" captionTargetBox="[151,1436,529,1605]" captionTargetId="figure@260.[151,1436,529,1606]" captionTargetPageId="260" captionText="FIGURE 100. Habitus of Bocchus species. ♀ in dorsal (A) and lateral (B) view. ♂ in lateral (C) and dorsal (D) view. Scale bar: A: 3.24 mm; B: 1.88 mm; C: 1.55 mm; D: 2.64 mm." httpUri="https://zenodo.org/record/3336870/files/figure.png" pageId="259" pageNumber="260">Figs 100C, D</figureCitation>
); pterostigma present (
<figureCitation id="137B2A71FEAC2B4BFA89D30FFF4FFAF8" captionStart="FIGURE 100" captionStartId="260.[151,250,1618,1643]" captionTargetBox="[151,1436,529,1605]" captionTargetId="figure@260.[151,1436,529,1606]" captionTargetPageId="260" captionText="FIGURE 100. Habitus of Bocchus species. ♀ in dorsal (A) and lateral (B) view. ♂ in lateral (C) and dorsal (D) view. Scale bar: A: 3.24 mm; B: 1.88 mm; C: 1.55 mm; D: 2.64 mm." httpUri="https://zenodo.org/record/3336870/files/figure.png" pageId="259" pageNumber="260">Figs 100C, D</figureCitation>
); tibial spurs 1/1/2.
</paragraph>
</subSubSection>
<subSubSection id="C35A657FFEAC2B4BFF6ED3C6FE0BFA24" box="[199,490,1416,1443]" pageId="259" pageNumber="260" type="distribution">
<paragraph id="8BFF36F4FEAC2B4BFF6ED3C6FE0BFA24" blockId="259.[151,1437,804,1695]" box="[199,490,1416,1443]" pageId="259" pageNumber="260">
<emphasis id="B934EAE6FEAC2B4BFF6ED3C6FEB8FA24" bold="true" box="[199,345,1416,1443]" pageId="259" pageNumber="260">Distribution</emphasis>
. Worldwide.
</paragraph>
</subSubSection>
<subSubSection id="C35A657FFEAC2B4BFF6ED3E3FE62FA6C" pageId="259" pageNumber="260" type="biology_ecology">
<paragraph id="8BFF36F4FEAC2B4BFF6ED3E3FE62FA6C" blockId="259.[151,1437,804,1695]" pageId="259" pageNumber="260">
<emphasis id="B934EAE6FEAC2B4BFF6ED3E3FEEBFA40" bold="true" box="[199,266,1453,1479]" pageId="259" pageNumber="260">Hosts</emphasis>
.
<taxonomicName id="4C404D77FEAC2B4BFEB0D3E3FE44FA40" authorityName="Amyot &amp; Serville" authorityYear="1834" box="[281,421,1453,1479]" class="Insecta" family="Caliscelidae" kingdom="Animalia" order="Hemiptera" pageId="259" pageNumber="260" phylum="Arthropoda" rank="family">Caliscelidae</taxonomicName>
,
<taxonomicName id="4C404D77FEAC2B4BFE19D3E3FDB4FA40" baseAuthorityName="Guglielmino et al." baseAuthorityYear="2013" box="[432,597,1453,1479]" class="Insecta" family="Tropiduchidae" kingdom="Animalia" order="Hemiptera" pageId="259" pageNumber="260" phylum="Arthropoda" rank="family">Tropiduchidae</taxonomicName>
,
<taxonomicName id="4C404D77FEAC2B4BFDC8D3E3FD0CFA40" baseAuthorityName="Guglielmino et al." baseAuthorityYear="2013" box="[609,749,1453,1479]" class="Insecta" family="Cicadellidae" kingdom="Animalia" order="Hemiptera" pageId="259" pageNumber="260" phylum="Arthropoda" rank="family">Cicadellidae</taxonomicName>
(except
<taxonomicName id="4C404D77FEAC2B4BFCE7D3E3FC2EFA40" authorityName="C.F.Baker" authorityYear="1915" box="[846,975,1453,1479]" class="Insecta" family="Cicadellidae" kingdom="Animalia" order="Hemiptera" pageId="259" pageNumber="260" phylum="Arthropoda" rank="subFamily" subFamily="Idiocerinae">Idiocerinae</taxonomicName>
,
<taxonomicName id="4C404D77FEAC2B4BFC72D3E3FB8DFA40" authorityName="Evans" authorityYear="1935" box="[987,1132,1453,1479]" class="Insecta" family="Cicadellidae" kingdom="Animalia" order="Hemiptera" pageId="259" pageNumber="260" phylum="Arthropoda" rank="subFamily" subFamily="Macropsinae">Macropsinae</taxonomicName>
and
<taxonomicName id="4C404D77FEAC2B4BFB0BD3E3FAABFA4F" authorityName="Kirschbaum" authorityYear="1868" box="[1186,1354,1453,1480]" class="Insecta" family="Cicadellidae" kingdom="Animalia" order="Hemiptera" pageId="259" pageNumber="260" phylum="Arthropoda" rank="subFamily" subFamily="Typhlocybinae">Typhlocybinae</taxonomicName>
) (
<bibRefCitation id="EFD14B05FEAC2B4BFAC8D3E3FE96FA6C" author="Guglielmino, A. &amp; Olmi, M. &amp; Buckle, C." pageId="259" pageNumber="260" pagination="1 - 113" refId="ref517841" refString="Guglielmino, A., Olmi, M. &amp; Buckle, C. (2013) An updated host-parasite catalogue of world Dryinidae (Hymenoptera: Chrysidoidea). Zootaxa, 3740 (1), 1 - 113. https: // doi. org / 10.11646 / zootaxa. 3740.1.1" type="journal article" year="2013">
Guglielmino
<emphasis id="B934EAE6FEAC2B4BFF55D39CFECCFA6C" box="[252,301,1489,1515]" italics="true" pageId="259" pageNumber="260">et al</emphasis>
. 2013
</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C35A657FFEAC2B4BFF6ED3BBFC0AF9B4" pageId="259" pageNumber="260" type="distribution">
<paragraph id="8BFF36F4FEAC2B4BFF6ED3BBFC2FF988" blockId="259.[151,1437,804,1695]" box="[199,974,1524,1551]" pageId="259" pageNumber="260">
<emphasis id="B934EAE6FEAC2B4BFF6ED3BBFE8DF988" bold="true" box="[199,364,1524,1551]" pageId="259" pageNumber="260">World species</emphasis>
. 112 species are known,
<quantity id="4CB89B11FEAC2B4BFD2FD3BBFD5EF988" box="[646,703,1525,1551]" metricMagnitude="-1" metricUnit="m" metricValue="8.382" pageId="259" pageNumber="260" unit="in" value="33.0">33 in</quantity>
the Afrotropical region.
</paragraph>
<paragraph id="8BFF36F4FEAC2B4BFF6ED057FC0AF9B4" blockId="259.[151,1437,804,1695]" box="[199,1003,1561,1587]" pageId="259" pageNumber="260">
<emphasis id="B934EAE6FEAC2B4BFF6ED057FE8AF9B4" bold="true" box="[199,363,1561,1587]" pageId="259" pageNumber="260">World genera</emphasis>
. Three genera are known, one in the Afrotropical region.
</paragraph>
</subSubSection>
<subSubSection id="C35A657FFEAC2B4BFF6ED073FD25F918" pageId="259" pageNumber="260" type="discussion">
<paragraph id="8BFF36F4FEAC2B4BFF6ED073FD25F918" blockId="259.[151,1437,804,1695]" pageId="259" pageNumber="260">
<emphasis id="B934EAE6FEAC2B4BFF6ED073FED5F9D0" bold="true" box="[199,308,1597,1623]" pageId="259" pageNumber="260">Remarks</emphasis>
. The following genera have not been recorded from the Afrotropical region:
<taxonomicName id="4C404D77FEAC2B4BFB01D073FF2EF9FC" authority="Forster 1856" authorityName="Forster" authorityYear="1856" class="Insecta" family="Dryinidae" genus="Mystrophorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="259" pageNumber="260" phylum="Arthropoda" rank="genus">
<emphasis id="B934EAE6FEAC2B4BFB01D073FAA5F9D0" box="[1192,1348,1597,1623]" italics="true" pageId="259" pageNumber="260">Mystrophorus</emphasis>
<bibRefCitation id="EFD14B05FEAC2B4BFAE4D073FF2EF9FC" author="Forster, A." pageId="259" pageNumber="260" refId="ref516827" refString="Forster, A. (1856) Hymenopterologischer Studien, 2. Chalcidiae und Proctotrupii. Ernst ter Meer, Aachen, 152 pp." type="book" year="1856">Förster 1856</bibRefCitation>
</taxonomicName>
(Western Palaearctic) and
<taxonomicName id="4C404D77FEAC2B4BFDA1D02FFC9BF9FC" authority="Ponomarenko 1972" authorityName="Ponomarenko" authorityYear="1972" box="[520,890,1633,1659]" class="Insecta" family="Dryinidae" genus="Mirodryinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="259" pageNumber="260" phylum="Arthropoda" rank="genus">
<emphasis id="B934EAE6FEAC2B4BFDA1D02FFD72F9FC" box="[520,659,1633,1659]" italics="true" pageId="259" pageNumber="260">Mirodryinus</emphasis>
<bibRefCitation id="EFD14B05FEAC2B4BFD34D02FFC9BF9FC" author="Ponomarenko, N. G." box="[669,890,1633,1659]" pageId="259" pageNumber="260" pagination="673 - 680" refId="ref525412" refString="Ponomarenko, N. G. (1972) [On the fauna of dryinids (Hymenoptera, Dryinidae) of the Mongolian People's Republic]. [Insects of Mongolia], 1, 673 - 680. [in Russian]" type="journal article" year="1972">Ponomarenko 1972</bibRefCitation>
</taxonomicName>
(Western Palaearctic). Larvae of
<taxonomicName id="4C404D77FEAC2B4BFAA9D02FFA7DF9FC" authorityName="Forster" authorityYear="1856" box="[1280,1436,1633,1659]" class="Insecta" family="Dryinidae" genus="Mystrophorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="259" pageNumber="260" phylum="Arthropoda" rank="genus">
<emphasis id="B934EAE6FEAC2B4BFAA9D02FFA7DF9FC" box="[1280,1436,1633,1659]" italics="true" pageId="259" pageNumber="260">Mystrophorus</emphasis>
</taxonomicName>
were described by
<bibRefCitation id="EFD14B05FEAC2B4BFEC3D0CBFD59F918" author="Guglielmino, A. &amp; Buckle, C." box="[362,696,1669,1695]" pageId="259" pageNumber="260" pagination="57 - 66" refId="ref517388" refString="Guglielmino, A. &amp; Buckle, C. (2010) Description of larval instars of Mystrophorus formicaeformis Ruthe (Hymenoptera: Dryinidae). Zootaxa, 2602 (1), 57 - 66. https: // doi. org / 10.11646 / zootaxa. 2602.1.3" type="journal article" year="2010">Guglielmino &amp; Bückle (2010</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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