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<document id="2EB0F0C61EC68EFD63BF04BD1C0429FF" ID-CLB-Dataset="26202" ID-DOI="10.11646/zootaxa.4630.1.1" ID-GBIF-Dataset="db6a16a6-0fe8-4987-8ad5-42223825fcd7" ID-ISSN="1175-5326" ID-Zenodo-Dep="3336635" ID-ZooBank="8D375836-CCBA-473C-836F-6ABD44B4F881" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1563153150055" checkinUser="plazi" docAuthor="Olmi, Massimo, Copeland, Robert S. &amp; Noort, Simon Van" docDate="2019" docId="03E987E2FDEE280AFF3ED5B7FA7DFBE5" docLanguage="en" docName="zootaxa.4630.1.1.pdf.imf" docOrigin="Zootaxa 4630 (1)" docStyle="DocumentStyle:D239614CE4198176A422035174489AB1.4:Zootaxa.2009-2012.monograph" docStyleId="D239614CE4198176A422035174489AB1" docStyleName="Zootaxa.2009-2012.monograph" docStyleVersion="4" docTitle="Madecadryinus Olmi 2007" docType="treatment" docVersion="39" lastPageNumber="579" masterDocId="FFD0FF9AFFAF2A48FFA9D64EFFE1FF87" masterDocTitle="Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea)" masterLastPageNumber="619" masterPageNumber="1" pageNumber="578" updateTime="1698741107231" updateUser="ExternalLinkService">
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<mods:titleInfo id="D4D0916408CCA3ACDFE5019200B0FF88">
<mods:title id="9310D0559B1395F7E2589E08AD916F18">Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea)</mods:title>
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<mods:name id="2F171CF91164DE4C64D123564C7EBD93" type="personal">
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<mods:namePart id="FCB1BA3641BD012BA43C66D1F0312CBF">Olmi, Massimo</mods:namePart>
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<mods:namePart id="DB2B0375D5BE76EC4198673743FA9E5E">Copeland, Robert S.</mods:namePart>
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<mods:namePart id="F2BC2BB35EE8032CBBB5EF45BA5A29F9">Noort, Simon Van</mods:namePart>
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<mods:title id="AEF2781E4A1EC548A456D0F1DE972555">Zootaxa</mods:title>
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<mods:date id="D4A4D080798BE29F84A9504D545318AF">2019</mods:date>
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<mods:number id="580A86C5CF3D90A40EB0968D50589EB5">2019-07-10</mods:number>
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<mods:number id="0C3CF22848C0A6B8C0882EC081676F0E">4630</mods:number>
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<mods:classification id="94548A500C0227AD85823E597B22D299">journal article</mods:classification>
<mods:identifier id="6B787FF243BAF75816FD4B88E70243E0" type="CLB-Dataset">26202</mods:identifier>
<mods:identifier id="B585E57F2C251EF13BC58B59E2D44525" type="DOI">10.11646/zootaxa.4630.1.1</mods:identifier>
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<treatment id="03E987E2FDEE280AFF3ED5B7FA7DFBE5" ID-GBIF-Taxon="158673003" LSID="urn:lsid:plazi:treatment:03E987E2FDEE280AFF3ED5B7FA7DFBE5" httpUri="http://treatment.plazi.org/id/03E987E2FDEE280AFF3ED5B7FA7DFBE5" lastPageId="578" lastPageNumber="579" pageId="577" pageNumber="578">
<subSubSection id="C35A657FFDEE2809FF3ED5B7FD87FB94" box="[151,614,1017,1044]" pageId="577" pageNumber="578" type="nomenclature">
<paragraph id="8BFF36F4FDEE2809FF3ED5B7FD87FB94" blockId="577.[151,614,1017,1044]" box="[151,614,1017,1044]" pageId="577" pageNumber="578">
<heading id="D0B78198FDEE2809FF3ED5B7FD87FB94" bold="true" box="[151,614,1017,1044]" fontSize="11" level="1" pageId="577" pageNumber="578" reason="1">
<emphasis id="B934EAE6FDEE2809FF3ED5B7FD87FB94" bold="true" box="[151,614,1017,1044]" pageId="577" pageNumber="578">
21. Genus
<taxonomicName id="4C404D77FDEE2809FEB3D5B7FD87FB94" authority="Olmi, 2007" authorityName="Olmi" authorityYear="2007" box="[282,614,1017,1044]" class="Insecta" family="Dryinidae" genus="Madecadryinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="577" pageNumber="578" phylum="Arthropoda" rank="genus">
<emphasis id="B934EAE6FDEE2809FEB3D5B7FE34FB93" bold="true" box="[282,469,1017,1044]" italics="true" pageId="577" pageNumber="578">Madecadryinus</emphasis>
Olmi, 2007
</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C35A657FFDEE2809FF3ED20FFCCAFBDC" box="[151,811,1088,1116]" pageId="577" pageNumber="578" type="reference_group">
<paragraph id="8BFF36F4FDEE2809FF3ED20FFCCAFBDC" blockId="577.[151,811,1088,1116]" box="[151,811,1088,1116]" pageId="577" pageNumber="578">
<taxonomicName id="4C404D77FDEE2809FF3ED20FFE1EFBDC" authority="Olmi 2007 c: 17" authorityName="Olmi" authorityPageNumber="17" authorityYear="2007" box="[151,511,1089,1115]" class="Insecta" family="Dryinidae" genus="Madecadryinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="577" pageNumber="578" phylum="Arthropoda" rank="genus">
<emphasis id="B934EAE6FDEE2809FF3ED20FFEA4FBDC" box="[151,325,1089,1115]" italics="true" pageId="577" pageNumber="578">Madecadryinus</emphasis>
<bibRefCitation id="EFD14B05FDEE2809FEE5D20FFE1EFBDC" author="Olmi, M." box="[332,511,1089,1115]" pageId="577" pageNumber="578" pagination="1 - 46" refId="ref523177" refString="Olmi, M. (2007 c) Apodryininae of Madagascar and South Africa (Hymenoptera: Dryinidae). Frustula entomologica, New Series, 30, 1 - 46." type="journal article" year="2007">Olmi 2007c: 17</bibRefCitation>
</taxonomicName>
;
<bibRefCitation id="EFD14B05FDEE2809FDA0D20FFCC6FBDC" author="Azevedo, C. O. &amp; Madl, M. &amp; Olmi, M." box="[521,807,1088,1116]" pageId="577" pageNumber="578" pagination="845 - 918" refId="ref514944" refString="Azevedo, C. O., Madl, M. &amp; Olmi, M. (2010) A Catalogue of the Bethylidae, Chrysididae, Dryinidae, Embolemidae, Sclerogibbidae and Scolebythidae (Hymenoptera: Chrysidoidea) of the Malagasy Subregion. Linzer biologische Beitrage, 42, 845 - 918." type="journal article" year="2010">
Azevedo
<emphasis id="B934EAE6FDEE2809FDDDD20CFD44FBDC" box="[628,677,1089,1115]" italics="true" pageId="577" pageNumber="578">et al</emphasis>
. 2010: 885
</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C35A657FFDEE2809FF3ED2CAFBA5F959" pageId="577" pageNumber="578" type="nomenclature">
<paragraph id="8BFF36F4FDEE2809FF3ED2CAFB60FB19" blockId="577.[151,1437,1155,2011]" box="[151,1153,1155,1182]" pageId="577" pageNumber="578">
<emphasis id="B934EAE6FDEE2809FF3ED2CAFECAFB19" bold="true" box="[151,299,1155,1182]" pageId="577" pageNumber="578">
<typeStatus id="54FB8856FDEE2809FF3ED2CAFF33FB19" box="[151,210,1156,1182]" pageId="577" pageNumber="578">Type</typeStatus>
species
</emphasis>
.
<taxonomicName id="4C404D77FDEE2809FE90D2CAFD25FB19" authority="Olmi 2007" authorityName="Olmi" authorityYear="2007" box="[313,708,1156,1182]" class="Insecta" family="Dryinidae" genus="Madecadryinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="577" pageNumber="578" phylum="Arthropoda" rank="species" species="relictus">
<emphasis id="B934EAE6FDEE2809FE90D2CAFDA0FB19" box="[313,577,1156,1182]" italics="true" pageId="577" pageNumber="578">Madecadryinus relictus</emphasis>
Olmi 2007
</taxonomicName>
, by monotypy and original designation.
</paragraph>
<paragraph id="8BFF36F4FDEE2809FF6ED2E9FBA5F959" blockId="577.[151,1437,1155,2011]" pageId="577" pageNumber="578">
<emphasis id="B934EAE6FDEE2809FF6ED2E9FEA3FB45" bold="true" box="[199,322,1191,1218]" pageId="577" pageNumber="578">Diagnosis.</emphasis>
<specimenCount id="9D46FD7DFDEE2809FEE3D2E6FE80FB45" box="[330,353,1192,1218]" pageId="577" pageNumber="578" type="female">♀</specimenCount>
: apterous (
<figureCitation id="137B2A71FDEE2809FE76D2E6FDBBFB45" box="[479,602,1191,1218]" captionStart="FIGURE 257" captionStartId="580.[151,250,1862,1887]" captionTargetBox="[176,1411,575,1812]" captionTargetId="figure@580.[151,1436,589,1850]" captionTargetPageId="580" captionText="FIGURE 257. Habitus of Madecadryinus species. A: M. humicolus Olmi: ♀ holotype in dorsal view. B: M. opacus Olmi: ♀ holotype in dorsal view. C: M. politus Olmi: ♀ holotype in dorsal view. D, E: M. relictus Olmi: ♀ holotype in dorsal (D) and lateral (E) view." httpUri="https://zenodo.org/record/3337192/files/figure.png" pageId="577" pageNumber="578">Figs 257A</figureCitation>
,
<figureCitation id="137B2A71FDEE2809FDCED2E6FD4AFB44" box="[615,683,1192,1219]" captionStart="FIGURE 260" captionStartId="584.[151,250,1544,1569]" captionTargetBox="[187,1403,194,1508]" captionTargetId="figure@584.[169,1419,174,1533]" captionTargetPageId="584" captionText="FIGURE 260. Madecadryinus relictus Olmi: A, B: ♀ holotype in dorsal (A) and lateral (B) view; ♂ paratype in dorsal (C) and lateral (D) view. Scale bar: A: 1.54 mm; B: 1.17 mm; C: 1.74 mm; D: 1.25 mm." httpUri="https://zenodo.org/record/3337198/files/figure.png" pageId="577" pageNumber="578">260A</figureCitation>
): palpal formula 2/2; maxillary palpi smaller than labial palpi; an- tenna geniculated; antennomeres 4–10 with ADOs (one per antennomere, except two ADOs in 10
<superScript id="7C359BBCFDEE2809FB4FD284FB12FB5F" attach="left" box="[1254,1267,1226,1240]" fontSize="6" pageId="577" pageNumber="578">th</superScript>
antennomere); occipital carina incomplete (complete on dorsal side of head and incomplete on temple); ocelli absent; mesosoma (
<figureCitation id="137B2A71FDEE2809FF36D35AFEFBFAA9" box="[159,282,1299,1326]" captionStart="FIGURE 257" captionStartId="580.[151,250,1862,1887]" captionTargetBox="[176,1411,575,1812]" captionTargetId="figure@580.[151,1436,589,1850]" captionTargetPageId="580" captionText="FIGURE 257. Habitus of Madecadryinus species. A: M. humicolus Olmi: ♀ holotype in dorsal view. B: M. opacus Olmi: ♀ holotype in dorsal view. C: M. politus Olmi: ♀ holotype in dorsal view. D, E: M. relictus Olmi: ♀ holotype in dorsal (D) and lateral (E) view." httpUri="https://zenodo.org/record/3337192/files/figure.png" pageId="577" pageNumber="578">Figs 257A</figureCitation>
,
<figureCitation id="137B2A71FDEE2809FE8ED35AFE86FAA8" box="[295,359,1300,1327]" captionStart="FIGURE 260" captionStartId="584.[151,250,1544,1569]" captionTargetBox="[187,1403,194,1508]" captionTargetId="figure@584.[169,1419,174,1533]" captionTargetPageId="584" captionText="FIGURE 260. Madecadryinus relictus Olmi: A, B: ♀ holotype in dorsal (A) and lateral (B) view; ♂ paratype in dorsal (C) and lateral (D) view. Scale bar: A: 1.54 mm; B: 1.17 mm; C: 1.74 mm; D: 1.25 mm." httpUri="https://zenodo.org/record/3337198/files/figure.png" pageId="577" pageNumber="578">260A</figureCitation>
) composed almost completely of fused segment; only propleura mobile and articulated; rest of mesosoma laterally with incomplete suture between pronotum and mesothorax and incomplete meso-metapleural suture; posterior surface of mesosoma (corresponding to propodeal declivity) completely surrounded by strong suture; anterior dorsal region of mesosoma (corresponding to pronotum + mesonotum + metanotum + metapectalpropodeal disc) laterally with incomplete suture (always absent in anterior third and occasionally absent behind propodeal spiracles); apparent pronotum not crossed by transverse impression (
<figureCitation id="137B2A71FDEE2809FB84D386FB48FA65" box="[1069,1193,1479,1506]" captionStart="FIGURE 257" captionStartId="580.[151,250,1862,1887]" captionTargetBox="[176,1411,575,1812]" captionTargetId="figure@580.[151,1436,589,1850]" captionTargetPageId="580" captionText="FIGURE 257. Habitus of Madecadryinus species. A: M. humicolus Olmi: ♀ holotype in dorsal view. B: M. opacus Olmi: ♀ holotype in dorsal view. C: M. politus Olmi: ♀ holotype in dorsal view. D, E: M. relictus Olmi: ♀ holotype in dorsal (D) and lateral (E) view." httpUri="https://zenodo.org/record/3337192/files/figure.png" pageId="577" pageNumber="578">Figs 257A</figureCitation>
,
<figureCitation id="137B2A71FDEE2809FB11D386FB1AFA64" box="[1208,1275,1480,1507]" captionStart="FIGURE 260" captionStartId="584.[151,250,1544,1569]" captionTargetBox="[187,1403,194,1508]" captionTargetId="figure@584.[169,1419,174,1533]" captionTargetPageId="584" captionText="FIGURE 260. Madecadryinus relictus Olmi: A, B: ♀ holotype in dorsal (A) and lateral (B) view; ♂ paratype in dorsal (C) and lateral (D) view. Scale bar: A: 1.54 mm; B: 1.17 mm; C: 1.74 mm; D: 1.25 mm." httpUri="https://zenodo.org/record/3337198/files/figure.png" pageId="577" pageNumber="578">260A</figureCitation>
); epicnemium concealed; enlarged claw (
<figureCitation id="137B2A71FDEE2809FE14D3A2FDCFF981" box="[445,558,1515,1542]" captionStart="FIGURE 258" captionStartId="581.[151,250,1146,1171]" captionTargetBox="[162,1416,190,1114]" captionTargetId="figure@581.[151,1436,174,1134]" captionTargetPageId="581" captionText="FIGURE 258. A, B: Madecadryinus humicolus Olmi, ♀ holotype: head in lateral view (A); chela (B). C, D: Madecadryinus opacus Olmi, ♀ holotype: head in lateral view (C); chela (D). E, G: Madecadryinus politus Olmi, ♀ holotype: head in lateral view (E); chela (G). F, H: Madecadryinus ranomafanensis Olmi, Copeland &amp; van Noort sp. nov., ♂ holotype: head in dorsal view (F); genitalia (H). Scale bar: A: 0.13 mm; B, D: 0.03 mm; C, H: 0.08 mm; E: 0.15 mm; F: 0.17 mm; G: 0.04 mm." httpUri="https://zenodo.org/record/3337194/files/figure.png" pageId="577" pageNumber="578">Fig. 258B</figureCitation>
) with small subapical and medial teeth and few bristles or slender lamellae (occasionally without teeth (
<figureCitation id="137B2A71FDEE2809FE06D05EFDC3F9AC" box="[431,546,1552,1579]" captionStart="FIGURE 261" captionStartId="585.[151,250,1459,1484]" captionTargetBox="[162,1424,560,1415]" captionTargetId="figure@585.[151,1436,553,1447]" captionTargetPageId="585" captionText="FIGURE 261. A–C: Madecadryinus relictus Olmi: A: head and mesosoma of ♀ holotype in lateral view (A); chela of holotype (B); genitalia of ♂ paratype (C). D, E, G: Madecadryinus semicaecus Olmi: head of ♀ holotype in lateral view (D); head and mesosoma of ♀ holotype in lateral view (E); chela of holotype (G). F: Madecadryinus silvanus Olmi: genitalia of ♂ from Mada- gascar, 16°27.18’S 49°47.15’E. Scale bar: A: 0.77 mm; B: 0.05 mm; C: 0.10 mm; D, F: 0.12 mm; E: 0.43 mm; G: 0.04 mm." httpUri="https://zenodo.org/record/3337200/files/figure.png" pageId="577" pageNumber="578">Fig. 261B</figureCitation>
)); protarsomere 5 (
<figureCitation id="137B2A71FDEE2809FD5CD05EFC8AF9AD" box="[757,875,1551,1578]" captionStart="FIGURE 258" captionStartId="581.[151,250,1146,1171]" captionTargetBox="[162,1416,190,1114]" captionTargetId="figure@581.[151,1436,174,1134]" captionTargetPageId="581" captionText="FIGURE 258. A, B: Madecadryinus humicolus Olmi, ♀ holotype: head in lateral view (A); chela (B). C, D: Madecadryinus opacus Olmi, ♀ holotype: head in lateral view (C); chela (D). E, G: Madecadryinus politus Olmi, ♀ holotype: head in lateral view (E); chela (G). F, H: Madecadryinus ranomafanensis Olmi, Copeland &amp; van Noort sp. nov., ♂ holotype: head in dorsal view (F); genitalia (H). Scale bar: A: 0.13 mm; B, D: 0.03 mm; C, H: 0.08 mm; E: 0.15 mm; F: 0.17 mm; G: 0.04 mm." httpUri="https://zenodo.org/record/3337194/files/figure.png" pageId="577" pageNumber="578">Figs 258B</figureCitation>
,
<figureCitation id="137B2A71FDEE2809FCDFD05EFC55F9AC" box="[886,948,1552,1579]" captionStart="FIGURE 261" captionStartId="585.[151,250,1459,1484]" captionTargetBox="[162,1424,560,1415]" captionTargetId="figure@585.[151,1436,553,1447]" captionTargetPageId="585" captionText="FIGURE 261. A–C: Madecadryinus relictus Olmi: A: head and mesosoma of ♀ holotype in lateral view (A); chela of holotype (B); genitalia of ♂ paratype (C). D, E, G: Madecadryinus semicaecus Olmi: head of ♀ holotype in lateral view (D); head and mesosoma of ♀ holotype in lateral view (E); chela of holotype (G). F: Madecadryinus silvanus Olmi: genitalia of ♂ from Mada- gascar, 16°27.18’S 49°47.15’E. Scale bar: A: 0.77 mm; B: 0.05 mm; C: 0.10 mm; D, F: 0.12 mm; E: 0.43 mm; G: 0.04 mm." httpUri="https://zenodo.org/record/3337200/files/figure.png" pageId="577" pageNumber="578">261B</figureCitation>
) with rows of lamellae; tibial spurs 1/0/1.
<specimenCount id="9D46FD7DFDEE2809FA2ED05EFA7DF9AD" box="[1415,1436,1552,1578]" pageId="577" pageNumber="578" type="male">♂</specimenCount>
(
<figureCitation id="137B2A71FDEE2809FF36D07AFEE4F9C9" box="[159,261,1587,1614]" captionStart="FIGURE 259" captionStartId="583.[151,250,1404,1429]" captionTargetBox="[175,1429,429,1369]" captionTargetId="figure@583.[151,1436,409,1392]" captionTargetPageId="583" captionText="FIGURE 259. Madecadryinus ranomafanensis Olmi, Copeland &amp; van Noort sp. nov., ♂ holotype: habitus in dorsal (A) and lateral (B) view; head in frontal view (C); forewing (D)." httpUri="https://zenodo.org/record/3337196/files/figure.png" pageId="577" pageNumber="578">Figs 259</figureCitation>
,
<figureCitation id="137B2A71FDEE2809FEB9D07AFEB0F9C8" box="[272,337,1588,1615]" captionStart="FIGURE 260" captionStartId="584.[151,250,1544,1569]" captionTargetBox="[187,1403,194,1508]" captionTargetId="figure@584.[169,1419,174,1533]" captionTargetPageId="584" captionText="FIGURE 260. Madecadryinus relictus Olmi: A, B: ♀ holotype in dorsal (A) and lateral (B) view; ♂ paratype in dorsal (C) and lateral (D) view. Scale bar: A: 1.54 mm; B: 1.17 mm; C: 1.74 mm; D: 1.25 mm." httpUri="https://zenodo.org/record/3337198/files/figure.png" pageId="577" pageNumber="578">260C</figureCitation>
): fully winged; forewing with C, R and 1Cu cells completely enclosed by pigmented veins; palpal formula 6/3; antenna very long, much longer than body; mandible with four irregular teeth; occipital carina complete; temple very long, much shorter than eye (
<figureCitation id="137B2A71FDEE2809FD00D032FCFFF910" box="[681,798,1660,1687]" captionStart="FIGURE 260" captionStartId="584.[151,250,1544,1569]" captionTargetBox="[187,1403,194,1508]" captionTargetId="figure@584.[169,1419,174,1533]" captionTargetPageId="584" captionText="FIGURE 260. Madecadryinus relictus Olmi: A, B: ♀ holotype in dorsal (A) and lateral (B) view; ♂ paratype in dorsal (C) and lateral (D) view. Scale bar: A: 1.54 mm; B: 1.17 mm; C: 1.74 mm; D: 1.25 mm." httpUri="https://zenodo.org/record/3337198/files/figure.png" pageId="577" pageNumber="578">Fig. 260C</figureCitation>
); occiput deeply excavated (
<figureCitation id="137B2A71FDEE2809FBF1D032FB5BF911" box="[1112,1210,1659,1686]" captionStart="FIGURE 259" captionStartId="583.[151,250,1404,1429]" captionTargetBox="[175,1429,429,1369]" captionTargetId="figure@583.[151,1436,409,1392]" captionTargetPageId="583" captionText="FIGURE 259. Madecadryinus ranomafanensis Olmi, Copeland &amp; van Noort sp. nov., ♂ holotype: habitus in dorsal (A) and lateral (B) view; head in frontal view (C); forewing (D)." httpUri="https://zenodo.org/record/3337196/files/figure.png" pageId="577" pageNumber="578">Fig. 259</figureCitation>
); minimum distance between eyes and occipital carina much shorter than OOL (
<figureCitation id="137B2A71FDEE2809FCEED0EEFC5EF93C" box="[839,959,1696,1723]" captionStart="FIGURE 260" captionStartId="584.[151,250,1544,1569]" captionTargetBox="[187,1403,194,1508]" captionTargetId="figure@584.[169,1419,174,1533]" captionTargetPageId="584" captionText="FIGURE 260. Madecadryinus relictus Olmi: A, B: ♀ holotype in dorsal (A) and lateral (B) view; ♂ paratype in dorsal (C) and lateral (D) view. Scale bar: A: 1.54 mm; B: 1.17 mm; C: 1.74 mm; D: 1.25 mm." httpUri="https://zenodo.org/record/3337198/files/figure.png" pageId="577" pageNumber="578">Fig. 260C</figureCitation>
); epicnemium exposed; notauli complete; paramere without inner slender membranous process (
<figureCitation id="137B2A71FDEE2809FD5FD08AFC8FF959" box="[758,878,1731,1758]" captionStart="FIGURE 258" captionStartId="581.[151,250,1146,1171]" captionTargetBox="[162,1416,190,1114]" captionTargetId="figure@581.[151,1436,174,1134]" captionTargetPageId="581" captionText="FIGURE 258. A, B: Madecadryinus humicolus Olmi, ♀ holotype: head in lateral view (A); chela (B). C, D: Madecadryinus opacus Olmi, ♀ holotype: head in lateral view (C); chela (D). E, G: Madecadryinus politus Olmi, ♀ holotype: head in lateral view (E); chela (G). F, H: Madecadryinus ranomafanensis Olmi, Copeland &amp; van Noort sp. nov., ♂ holotype: head in dorsal view (F); genitalia (H). Scale bar: A: 0.13 mm; B, D: 0.03 mm; C, H: 0.08 mm; E: 0.15 mm; F: 0.17 mm; G: 0.04 mm." httpUri="https://zenodo.org/record/3337194/files/figure.png" pageId="577" pageNumber="578">Fig. 258H</figureCitation>
); tibial spurs 1/1/2.
</paragraph>
</subSubSection>
<subSubSection id="C35A657FFDEE2809FF6ED0A9FDA7F885" box="[199,582,1767,1794]" pageId="577" pageNumber="578" type="distribution">
<paragraph id="8BFF36F4FDEE2809FF6ED0A9FDA7F885" blockId="577.[151,1437,1155,2011]" box="[199,582,1767,1794]" pageId="577" pageNumber="578">
<emphasis id="B934EAE6FDEE2809FF6ED0A9FEB8F885" bold="true" box="[199,345,1767,1794]" pageId="577" pageNumber="578">Distribution</emphasis>
. Afrotropical region.
</paragraph>
</subSubSection>
<subSubSection id="C35A657FFDEE2809FF6ED142FE6FF8A1" box="[199,398,1804,1830]" pageId="577" pageNumber="578" type="biology_ecology">
<paragraph id="8BFF36F4FDEE2809FF6ED142FE6FF8A1" blockId="577.[151,1437,1155,2011]" box="[199,398,1804,1830]" pageId="577" pageNumber="578">
<emphasis id="B934EAE6FDEE2809FF6ED142FEF0F8A1" bold="true" box="[199,273,1804,1830]" pageId="577" pageNumber="578">Hosts.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="C35A657FFDEE2809FF6ED17EFC0BF8CD" box="[199,1002,1839,1866]" pageId="577" pageNumber="578" type="discussion">
<paragraph id="8BFF36F4FDEE2809FF6ED17EFC0BF8CD" blockId="577.[151,1437,1155,2011]" box="[199,1002,1839,1866]" pageId="577" pageNumber="578">
<emphasis id="B934EAE6FDEE2809FF6ED17EFE8DF8CD" bold="true" box="[199,364,1839,1866]" pageId="577" pageNumber="578">World species</emphasis>
. Seven species are known, all in the Afrotropical region.
</paragraph>
</subSubSection>
<subSubSection id="C35A657FFDEE280AFF6ED11AFAFAFE37" lastPageId="578" lastPageNumber="579" pageId="577" pageNumber="578" type="discussion">
<paragraph id="8BFF36F4FDEE280AFF6ED11AFAFAFE37" blockId="577.[151,1437,1155,2011]" lastBlockId="578.[151,1437,153,432]" lastPageId="578" lastPageNumber="579" pageId="577" pageNumber="578">
<emphasis id="B934EAE6FDEE2809FF6ED11AFED5F8E9" bold="true" box="[199,308,1876,1902]" pageId="577" pageNumber="578">Remarks</emphasis>
. Based on the presence of an incomplete suture that is only laterally visible between the pronotum and mesothorax in
<specimenCount id="9D46FD7DFDEE2809FED9D136FE7AF815" box="[368,411,1912,1938]" pageId="577" pageNumber="578" type="female">♀♀</specimenCount>
, the genus
<taxonomicName id="4C404D77FDEE2809FDB5D136FD2BF815" authorityName="Olmi" authorityYear="2007" box="[540,714,1912,1938]" class="Insecta" family="Dryinidae" genus="Madecadryinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="577" pageNumber="578" phylum="Arthropoda" rank="genus">
<emphasis id="B934EAE6FDEE2809FDB5D136FD2BF815" box="[540,714,1912,1938]" italics="true" pageId="577" pageNumber="578">Madecadryinus</emphasis>
</taxonomicName>
is very similar to the Neotropical genus
<taxonomicName id="4C404D77FDEE2809FB33D136FA7DF815" authority="Olmi 1984" authorityName="Olmi" authorityYear="1984" box="[1178,1436,1912,1938]" class="Insecta" family="Dryinidae" genus="Apodryinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="577" pageNumber="578" phylum="Arthropoda" rank="genus">
<emphasis id="B934EAE6FDEE2809FB33D136FAFAF815" box="[1178,1307,1912,1938]" italics="true" pageId="577" pageNumber="578">Apodryinus</emphasis>
<bibRefCitation id="EFD14B05FDEE2809FA8AD136FA7DF815" author="Olmi, M." box="[1315,1436,1912,1938]" pageId="577" pageNumber="578" pagination="1 - 1913" refId="ref521356" refString="Olmi, M. (1984) A revision of the Dryinidae (Hymenoptera). Memoirs of the American Entomological Institute, 37, I-XII + 1 - 1913." type="journal article" year="1984">Olmi 1984</bibRefCitation>
</taxonomicName>
(in contrast the Australian genus
<taxonomicName id="4C404D77FDEE2809FDACD1D2FCE3F831" authority="Olmi 1991" authorityName="Olmi" authorityYear="1991" box="[517,770,1948,1974]" class="Insecta" family="Dryinidae" genus="Bocchopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="577" pageNumber="578" phylum="Arthropoda" rank="genus">
<emphasis id="B934EAE6FDEE2809FDACD1D2FD64F831" box="[517,645,1948,1974]" italics="true" pageId="577" pageNumber="578">Bocchopsis</emphasis>
<bibRefCitation id="EFD14B05FDEE2809FD22D1D2FCE3F831" author="Olmi, M." box="[651,770,1948,1974]" pageId="577" pageNumber="578" pagination="109 - 395" refId="ref521701" refString="Olmi, M. (1991 [&quot; 1989 &quot;]) Supplement to the revision of the world Dryinidae (Hymenoptera Chrysidoidea). Frustula entomologica, New Series, 12, 109 - 395." type="journal article" year="1991">Olmi 1991</bibRefCitation>
</taxonomicName>
has the suture between pronotum and mesothorax complete and visible not only laterally, but also dorsally). The main differences between
<taxonomicName id="4C404D77FDEE2809FBA4D18EFB6FF85D" authorityName="Olmi" authorityYear="1984" box="[1037,1166,1984,2010]" class="Insecta" family="Dryinidae" genus="Apodryinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="577" pageNumber="578" phylum="Arthropoda" rank="genus">
<emphasis id="B934EAE6FDEE2809FBA4D18EFB6FF85D" box="[1037,1166,1984,2010]" italics="true" pageId="577" pageNumber="578">Apodryinus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C404D77FDEE2809FB6CD18EFA92F85D" authorityName="Olmi" authorityYear="2007" box="[1221,1395,1984,2010]" class="Insecta" family="Dryinidae" genus="Madecadryinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="577" pageNumber="578" phylum="Arthropoda" rank="genus">
<emphasis id="B934EAE6FDEE2809FB6CD18EFA92F85D" box="[1221,1395,1984,2010]" italics="true" pageId="577" pageNumber="578">Madecadryinus</emphasis>
</taxonomicName>
are in respect of the meso-metapleural suture (completely absent in
<taxonomicName id="4C404D77FDED280AFCCCD6D7FC07FF34" authorityName="Olmi" authorityYear="1984" box="[869,998,153,179]" class="Insecta" family="Dryinidae" genus="Apodryinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="578" pageNumber="579" phylum="Arthropoda" rank="genus">
<emphasis id="B934EAE6FDED280AFCCCD6D7FC07FF34" box="[869,998,153,179]" italics="true" pageId="578" pageNumber="579">Apodryinus</emphasis>
</taxonomicName>
, laterally partly present in
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<emphasis id="B934EAE6FDED280AFABAD6D7FF27FF50" italics="true" pageId="578" pageNumber="579">Madecadryinus</emphasis>
</taxonomicName>
), the occipital carina (absent in
<taxonomicName id="4C404D77FDED280AFD86D6F3FD51FF50" authorityName="Olmi" authorityYear="1984" box="[559,688,189,215]" class="Insecta" family="Dryinidae" genus="Apodryinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="578" pageNumber="579" phylum="Arthropoda" rank="genus">
<emphasis id="B934EAE6FDED280AFD86D6F3FD51FF50" box="[559,688,189,215]" italics="true" pageId="578" pageNumber="579">Apodryinus</emphasis>
</taxonomicName>
, almost complete in
<taxonomicName id="4C404D77FDED280AFC35D6F3FBABFF50" authorityName="Olmi" authorityYear="2007" box="[924,1098,189,215]" class="Insecta" family="Dryinidae" genus="Madecadryinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="578" pageNumber="579" phylum="Arthropoda" rank="genus">
<emphasis id="B934EAE6FDED280AFC35D6F3FBABFF50" box="[924,1098,189,215]" italics="true" pageId="578" pageNumber="579">Madecadryinus</emphasis>
</taxonomicName>
), the shape of the head (spheroidal in
<taxonomicName id="4C404D77FDED280AFF56D6AFFE61FF7C" authorityName="Olmi" authorityYear="1984" box="[255,384,225,251]" class="Insecta" family="Dryinidae" genus="Apodryinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="578" pageNumber="579" phylum="Arthropoda" rank="genus">
<emphasis id="B934EAE6FDED280AFF56D6AFFE61FF7C" box="[255,384,225,251]" italics="true" pageId="578" pageNumber="579">Apodryinus</emphasis>
</taxonomicName>
, cuboidal in
<taxonomicName id="4C404D77FDED280AFDBED6AFFD24FF7C" authorityName="Olmi" authorityYear="2007" box="[535,709,225,251]" class="Insecta" family="Dryinidae" genus="Madecadryinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="578" pageNumber="579" phylum="Arthropoda" rank="genus">
<emphasis id="B934EAE6FDED280AFDBED6AFFD24FF7C" box="[535,709,225,251]" italics="true" pageId="578" pageNumber="579">Madecadryinus</emphasis>
</taxonomicName>
), the eye (larger in
<taxonomicName id="4C404D77FDED280AFC00D6AFFBCBFF7C" authorityName="Olmi" authorityYear="1984" box="[937,1066,225,251]" class="Insecta" family="Dryinidae" genus="Apodryinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="578" pageNumber="579" phylum="Arthropoda" rank="genus">
<emphasis id="B934EAE6FDED280AFC00D6AFFBCBFF7C" box="[937,1066,225,251]" italics="true" pageId="578" pageNumber="579">Apodryinus</emphasis>
</taxonomicName>
, smaller in
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<emphasis id="B934EAE6FDED280AFB1AD6AFFA80FF7C" box="[1203,1377,225,251]" italics="true" pageId="578" pageNumber="579">Madecadryinus</emphasis>
</taxonomicName>
), the palpal formula (4/
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<taxonomicName id="4C404D77FDED280AFE26D74BFDF1FE98" authorityName="Olmi" authorityYear="1984" box="[399,528,261,287]" class="Insecta" family="Dryinidae" genus="Apodryinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="578" pageNumber="579" phylum="Arthropoda" rank="genus">
<emphasis id="B934EAE6FDED280AFE26D74BFDF1FE98" box="[399,528,261,287]" italics="true" pageId="578" pageNumber="579">Apodryinus</emphasis>
</taxonomicName>
, 2/
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<taxonomicName id="4C404D77FDED280AFDCBD74BFCF1FE98" authorityName="Olmi" authorityYear="2007" box="[610,784,261,287]" class="Insecta" family="Dryinidae" genus="Madecadryinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="578" pageNumber="579" phylum="Arthropoda" rank="genus">
<emphasis id="B934EAE6FDED280AFDCBD74BFCF1FE98" box="[610,784,261,287]" italics="true" pageId="578" pageNumber="579">Madecadryinus</emphasis>
</taxonomicName>
), the shape of the antennae (slenderer in
<taxonomicName id="4C404D77FDED280AFB7CD74BFAB7FE98" authorityName="Olmi" authorityYear="1984" box="[1237,1366,261,287]" class="Insecta" family="Dryinidae" genus="Apodryinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="578" pageNumber="579" phylum="Arthropoda" rank="genus">
<emphasis id="B934EAE6FDED280AFB7CD74BFAB7FE98" box="[1237,1366,261,287]" italics="true" pageId="578" pageNumber="579">Apodryinus</emphasis>
</taxonomicName>
, more thickened in
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<emphasis id="B934EAE6FDED280AFE8FD767FE35FEC4" box="[294,468,297,323]" italics="true" pageId="578" pageNumber="579">Madecadryinus</emphasis>
</taxonomicName>
), the ADOs (present in antennomeres
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<taxonomicName id="4C404D77FDED280AFC70D767FBBBFEC4" authorityName="Olmi" authorityYear="1984" box="[985,1114,297,323]" class="Insecta" family="Dryinidae" genus="Apodryinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="578" pageNumber="579" phylum="Arthropoda" rank="genus">
<emphasis id="B934EAE6FDED280AFC70D767FBBBFEC4" box="[985,1114,297,323]" italics="true" pageId="578" pageNumber="579">Apodryinus</emphasis>
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, in
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<taxonomicName id="4C404D77FDED280AFB77D767FA6DFEC4" authorityName="Olmi" authorityYear="2007" box="[1246,1420,297,323]" class="Insecta" family="Dryinidae" genus="Madecadryinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="578" pageNumber="579" phylum="Arthropoda" rank="genus">
<emphasis id="B934EAE6FDED280AFB77D767FA6DFEC4" box="[1246,1420,297,323]" italics="true" pageId="578" pageNumber="579">Madecadryinus</emphasis>
</taxonomicName>
), the enlarged claw (with lamellae and one subapical tooth in
<taxonomicName id="4C404D77FDED280AFC95D703FC5CFEE0" authorityName="Olmi" authorityYear="1984" box="[828,957,333,359]" class="Insecta" family="Dryinidae" genus="Apodryinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="578" pageNumber="579" phylum="Arthropoda" rank="genus">
<emphasis id="B934EAE6FDED280AFC95D703FC5CFEE0" box="[828,957,333,359]" italics="true" pageId="578" pageNumber="579">Apodryinus</emphasis>
</taxonomicName>
, with bristles or very slender lamellae and with small subapical and medial teeth in
<taxonomicName id="4C404D77FDED280AFDCCD73FFCF2FE0C" authorityName="Olmi" authorityYear="2007" box="[613,787,369,395]" class="Insecta" family="Dryinidae" genus="Madecadryinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="578" pageNumber="579" phylum="Arthropoda" rank="genus">
<emphasis id="B934EAE6FDED280AFDCCD73FFCF2FE0C" box="[613,787,369,395]" italics="true" pageId="578" pageNumber="579">Madecadryinus</emphasis>
</taxonomicName>
). Based on the absence of an inner slender process of the parameres, the
<specimenCount id="9D46FD7DFDED280AFEE8D7D8FEB7FE37" box="[321,342,406,432]" pageId="578" pageNumber="579" type="male">♂</specimenCount>
of
<taxonomicName id="4C404D77FDED280AFED5D7DBFDCBFE28" authorityName="Olmi" authorityYear="2007" box="[380,554,405,431]" class="Insecta" family="Dryinidae" genus="Madecadryinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="578" pageNumber="579" phylum="Arthropoda" rank="genus">
<emphasis id="B934EAE6FDED280AFED5D7DBFDCBFE28" box="[380,554,405,431]" italics="true" pageId="578" pageNumber="579">Madecadryinus</emphasis>
</taxonomicName>
is more similar to the
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of
<taxonomicName id="4C404D77FDED280AFCDCD7DBFC17FE28" authorityName="Olmi" authorityYear="1984" box="[885,1014,405,431]" class="Insecta" family="Dryinidae" genus="Apodryinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="578" pageNumber="579" phylum="Arthropoda" rank="genus">
<emphasis id="B934EAE6FDED280AFCDCD7DBFC17FE28" box="[885,1014,405,431]" italics="true" pageId="578" pageNumber="579">Apodryinus</emphasis>
</taxonomicName>
than those of
<taxonomicName id="4C404D77FDED280AFB3CD7DBFAF4FE28" authorityName="Olmi" authorityYear="1991" box="[1173,1301,405,431]" class="Insecta" family="Dryinidae" genus="Bocchopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="578" pageNumber="579" phylum="Arthropoda" rank="genus">
<emphasis id="B934EAE6FDED280AFB3CD7DBFAF4FE28" box="[1173,1301,405,431]" italics="true" pageId="578" pageNumber="579">Bocchopsis</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C35A657FFDED280AFF3ED44FFA7DFBE5" pageId="578" pageNumber="579" type="key">
<paragraph id="8BFF36F4FDED280AFF3ED44FFDC8FD9B" blockId="578.[151,553,513,540]" box="[151,553,513,540]" pageId="578" pageNumber="579">
<heading id="D0B78198FDED280AFF3ED44FFDC8FD9B" bold="true" box="[151,553,513,540]" fontSize="11" level="1" pageId="578" pageNumber="579" reason="1">
<emphasis id="B934EAE6FDED280AFF3ED44FFDC8FD9B" bold="true" box="[151,553,513,540]" pageId="578" pageNumber="579">
Key to species of
<taxonomicName id="4C404D77FDED280AFEC7D44FFDC8FD9B" authorityName="Olmi" authorityYear="2007" box="[366,553,513,540]" class="Insecta" family="Dryinidae" genus="Madecadryinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="578" pageNumber="579" phylum="Arthropoda" rank="genus">
<emphasis id="B934EAE6FDED280AFEC7D44FFDC8FD9B" bold="true" box="[366,553,513,540]" italics="true" pageId="578" pageNumber="579">Madecadryinus</emphasis>
</taxonomicName>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BFF36F4FDED280AFF3ED404FD17FDE3" blockId="578.[151,1437,585,904]" box="[151,758,585,612]" pageId="578" pageNumber="579">
<specimenCount id="9D46FD7DFDED280AFF3ED404FF20FDE3" box="[151,193,586,612]" pageId="578" pageNumber="579" type="female">♀♀</specimenCount>
(unknown in
<taxonomicName id="4C404D77FDED280AFEF7D404FDD7FDE4" authorityName="Olmi &amp; Copeland &amp; Noort" authorityYear="2019" box="[350,566,585,611]" class="Liliopsida" family="Araceae" genus="Gonatopus" kingdom="Plantae" order="Alismatales" pageId="578" pageNumber="579" phylum="Tracheophyta" rank="species" species="ranomafanensis">
<emphasis id="B934EAE6FDED280AFEF7D404FDD7FDE4" box="[350,566,585,611]" italics="true" pageId="578" pageNumber="579">M. ranomafanensis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C404D77FDED280AFDC5D404FD0CFDE4" authorityName="Olmi" authorityYear="2007" box="[620,749,585,611]" class="Insecta" family="Dryinidae" genus="Madecadryinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="578" pageNumber="579" phylum="Arthropoda" rank="species" species="silvanus">
<emphasis id="B934EAE6FDED280AFDC5D404FD0CFDE4" box="[620,749,585,611]" italics="true" pageId="578" pageNumber="579">M. silvanus</emphasis>
</taxonomicName>
)
</paragraph>
<keyStep id="30B42E51FDED280AFF3ED424FA7DFD50" pageId="578" pageNumber="579">
<paragraph id="8BFF36F4FDED280AFF3ED424FA7DFD1A" blockId="578.[151,1437,585,904]" pageId="578" pageNumber="579">
<keyLead id="30B195C1FDED280AFF3ED424FA7DFD1A" pageId="578" pageNumber="579">
1. Lateral suture of dorsal surface of mesosoma not present from propodeal spiracles to lateral corners of keel surrounding posterior inclined region (
<figureCitation id="137B2A71FDED280AFE01D4C9FDEAFD1A" box="[424,523,647,669]" captionStart="FIGURE 261" captionStartId="585.[151,250,1459,1484]" captionTargetBox="[162,1424,560,1415]" captionTargetId="figure@585.[151,1436,553,1447]" captionTargetPageId="585" captionText="FIGURE 261. A–C: Madecadryinus relictus Olmi: A: head and mesosoma of ♀ holotype in lateral view (A); chela of holotype (B); genitalia of ♂ paratype (C). D, E, G: Madecadryinus semicaecus Olmi: head of ♀ holotype in lateral view (D); head and mesosoma of ♀ holotype in lateral view (E); chela of holotype (G). F: Madecadryinus silvanus Olmi: genitalia of ♂ from Mada- gascar, 16°27.18’S 49°47.15’E. Scale bar: A: 0.77 mm; B: 0.05 mm; C: 0.10 mm; D, F: 0.12 mm; E: 0.43 mm; G: 0.04 mm." httpUri="https://zenodo.org/record/3337200/files/figure.png" pageId="578" pageNumber="579">Fig. 261A</figureCitation>
).............................................................
<taxonomicName id="4C404D77FDED280AFB57D4C6FA7DFD1A" authority="Olmi" authorityName="Olmi" authorityYear="2007" box="[1278,1436,647,669]" class="Insecta" family="Dryinidae" genus="Madecadryinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="578" pageNumber="579" phylum="Arthropoda" rank="species" species="relictus">
<emphasis id="B934EAE6FDED280AFB57D4C6FA84FD1B" box="[1278,1381,647,669]" italics="true" pageId="578" pageNumber="579">M. relictus</emphasis>
Olmi
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BFF36F4FDED280AFF3ED4EBFA7DFD50" blockId="578.[151,1437,585,904]" pageId="578" pageNumber="579">
<keyLead id="30B195C1FDED280AFF3ED4EBFA7DFD50" pageId="578" pageNumber="579">
- Lateral suture of dorsal surface of mesosoma present from propodeal spiracles to lateral corners of keel surrounding posterior inclined region (
<figureCitation id="137B2A71FDED280AFE29D48CFE00FD5F" box="[384,481,706,728]" captionStart="FIGURE 261" captionStartId="585.[151,250,1459,1484]" captionTargetBox="[162,1424,560,1415]" captionTargetId="figure@585.[151,1436,553,1447]" captionTargetPageId="585" captionText="FIGURE 261. A–C: Madecadryinus relictus Olmi: A: head and mesosoma of ♀ holotype in lateral view (A); chela of holotype (B); genitalia of ♂ paratype (C). D, E, G: Madecadryinus semicaecus Olmi: head of ♀ holotype in lateral view (D); head and mesosoma of ♀ holotype in lateral view (E); chela of holotype (G). F: Madecadryinus silvanus Olmi: genitalia of ♂ from Mada- gascar, 16°27.18’S 49°47.15’E. Scale bar: A: 0.77 mm; B: 0.05 mm; C: 0.10 mm; D, F: 0.12 mm; E: 0.43 mm; G: 0.04 mm." httpUri="https://zenodo.org/record/3337200/files/figure.png" pageId="578" pageNumber="579">Fig. 261E</figureCitation>
)............................................................................. 2
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30B42E51FDED280AFF3ED491FA7DFC96" pageId="578" pageNumber="579">
<paragraph id="8BFF36F4FDED280AFF3ED491FA7DFD72" blockId="578.[151,1437,585,904]" box="[151,1436,735,757]" pageId="578" pageNumber="579">
<keyLead id="30B195C1FDED280AFF3ED491FA7DFD72" box="[151,1436,735,757]" pageId="578" pageNumber="579">
2. Eye very small (
<figureCitation id="137B2A71FDED280AFED6D491FE04FD72" box="[383,485,735,757]" captionStart="FIGURE 261" captionStartId="585.[151,250,1459,1484]" captionTargetBox="[162,1424,560,1415]" captionTargetId="figure@585.[151,1436,553,1447]" captionTargetPageId="585" captionText="FIGURE 261. A–C: Madecadryinus relictus Olmi: A: head and mesosoma of ♀ holotype in lateral view (A); chela of holotype (B); genitalia of ♂ paratype (C). D, E, G: Madecadryinus semicaecus Olmi: head of ♀ holotype in lateral view (D); head and mesosoma of ♀ holotype in lateral view (E); chela of holotype (G). F: Madecadryinus silvanus Olmi: genitalia of ♂ from Mada- gascar, 16°27.18’S 49°47.15’E. Scale bar: A: 0.77 mm; B: 0.05 mm; C: 0.10 mm; D, F: 0.12 mm; E: 0.43 mm; G: 0.04 mm." httpUri="https://zenodo.org/record/3337200/files/figure.png" pageId="578" pageNumber="579">Fig. 261D</figureCitation>
); head more than eight × as long as eye...............................
<taxonomicName id="4C404D77FDED280AFB71D4AEFA7DFD72" authority="Olmi" authorityName="Olmi" authorityYear="2007" box="[1240,1436,735,757]" class="Insecta" family="Dryinidae" genus="Madecadryinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="578" pageNumber="579" phylum="Arthropoda" rank="species" species="semicaecus">
<emphasis id="B934EAE6FDED280AFB71D4AEFA84FD72" box="[1240,1381,736,757]" italics="true" pageId="578" pageNumber="579">M. semicaecus</emphasis>
Olmi
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BFF36F4FDED280AFF3ED4B3FA7DFC96" blockId="578.[151,1437,585,904]" box="[151,1436,764,786]" pageId="578" pageNumber="579">
<keyLead id="30B195C1FDED280AFF3ED4B3FA7DFC96" box="[151,1436,764,786]" pageId="578" pageNumber="579">
- Eye larger (
<figureCitation id="137B2A71FDED280AFEF8D4B2FE37FC95" box="[337,470,764,786]" captionStart="FIGURE 258" captionStartId="581.[151,250,1146,1171]" captionTargetBox="[162,1416,190,1114]" captionTargetId="figure@581.[151,1436,174,1134]" captionTargetPageId="581" captionText="FIGURE 258. A, B: Madecadryinus humicolus Olmi, ♀ holotype: head in lateral view (A); chela (B). C, D: Madecadryinus opacus Olmi, ♀ holotype: head in lateral view (C); chela (D). E, G: Madecadryinus politus Olmi, ♀ holotype: head in lateral view (E); chela (G). F, H: Madecadryinus ranomafanensis Olmi, Copeland &amp; van Noort sp. nov., ♂ holotype: head in dorsal view (F); genitalia (H). Scale bar: A: 0.13 mm; B, D: 0.03 mm; C, H: 0.08 mm; E: 0.15 mm; F: 0.17 mm; G: 0.04 mm." httpUri="https://zenodo.org/record/3337194/files/figure.png" pageId="578" pageNumber="579">Figs 258A, C</figureCitation>
); head less than eight × as long as eye................................................ 3
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30B42E51FDED280AFF3ED554FA7DFCCB" pageId="578" pageNumber="579">
<paragraph id="8BFF36F4FDED280AFF3ED554FA7DFCB7" blockId="578.[151,1437,585,904]" box="[151,1436,793,816]" pageId="578" pageNumber="579">
<keyLead id="30B195C1FDED280AFF3ED554FA7DFCB7" box="[151,1436,793,816]" pageId="578" pageNumber="579">
3. Antenna slender (
<figureCitation id="137B2A71FDED280AFE25D554FE13FCA8" box="[396,498,793,816]" captionStart="FIGURE 257" captionStartId="580.[151,250,1862,1887]" captionTargetBox="[176,1411,575,1812]" captionTargetId="figure@580.[151,1436,589,1850]" captionTargetPageId="580" captionText="FIGURE 257. Habitus of Madecadryinus species. A: M. humicolus Olmi: ♀ holotype in dorsal view. B: M. opacus Olmi: ♀ holotype in dorsal view. C: M. politus Olmi: ♀ holotype in dorsal view. D, E: M. relictus Olmi: ♀ holotype in dorsal (D) and lateral (E) view." httpUri="https://zenodo.org/record/3337192/files/figure.png" pageId="578" pageNumber="579">Fig. 257A</figureCitation>
), with antennomere 6 longer than broad..............................
<taxonomicName id="4C404D77FDED280AFB4BD554FA7DFCB7" authority="Olmi" authorityName="Olmi" authorityYear="2007" box="[1250,1436,794,816]" class="Insecta" family="Dryinidae" genus="Madecadryinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="578" pageNumber="579" phylum="Arthropoda" rank="species" species="humicolus">
<emphasis id="B934EAE6FDED280AFB4BD554FA84FCA8" box="[1250,1381,794,815]" italics="true" pageId="578" pageNumber="579">M. humicolus</emphasis>
Olmi
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BFF36F4FDED280AFF3ED579FA7DFCCB" blockId="578.[151,1437,585,904]" box="[151,1436,823,845]" pageId="578" pageNumber="579">
<keyLead id="30B195C1FDED280AFF3ED579FA7DFCCB" box="[151,1436,823,845]" pageId="578" pageNumber="579">
- Antenna thicker (
<figureCitation id="137B2A71FDED280AFE20D579FDEAFCCA" box="[393,523,823,845]" captionStart="FIGURE 257" captionStartId="580.[151,250,1862,1887]" captionTargetBox="[176,1411,575,1812]" captionTargetId="figure@580.[151,1436,589,1850]" captionTargetPageId="580" captionText="FIGURE 257. Habitus of Madecadryinus species. A: M. humicolus Olmi: ♀ holotype in dorsal view. B: M. opacus Olmi: ♀ holotype in dorsal view. C: M. politus Olmi: ♀ holotype in dorsal view. D, E: M. relictus Olmi: ♀ holotype in dorsal (D) and lateral (E) view." httpUri="https://zenodo.org/record/3337192/files/figure.png" pageId="578" pageNumber="579">Figs 257B, C</figureCitation>
), with antennomere 6 about as long as broad...................................... 4
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30B42E51FDED280AFF3ED51AFA7DFC0F" pageId="578" pageNumber="579">
<paragraph id="8BFF36F4FDED280AFF3ED51AFA7DFCED" blockId="578.[151,1437,585,904]" box="[151,1436,852,874]" pageId="578" pageNumber="579">
<keyLead id="30B195C1FDED280AFF3ED51AFA7DFCED" box="[151,1436,852,874]" pageId="578" pageNumber="579">
4. Head and anterior surface of mesosoma dull, granulate...........................................
<taxonomicName id="4C404D77FDED280AFAA8D51BFA7DFCED" authority="Olmi" authorityName="Olmi" box="[1281,1436,852,874]" class="Insecta" family="Dryinidae" genus="Pseudogonatopus" kingdom="Animalia" order="Hymenoptera" pageId="578" pageNumber="579" phylum="Arthropoda" rank="species" species="opacus">
<emphasis id="B934EAE6FDED280AFAA8D51BFA84FCED" box="[1281,1381,853,874]" italics="true" pageId="578" pageNumber="579">M. opacus</emphasis>
Olmi
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BFF36F4FDED280AFF3ED53CFA7DFC0F" blockId="578.[151,1437,585,904]" box="[151,1436,882,904]" pageId="578" pageNumber="579">
<keyLead id="30B195C1FDED280AFF3ED53CFA7DFC0F" box="[151,1436,882,904]" pageId="578" pageNumber="579">
- Head and anterior surface of mesosoma shiny, finely punctate and unsculptured among punctures.........
<taxonomicName id="4C404D77FDED280AFAADD53CFA7DFC0F" authority="Olmi" authorityName="Olmi" authorityYear="2007" box="[1284,1436,882,904]" class="Insecta" family="Dryinidae" genus="Madecadryinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="578" pageNumber="579" phylum="Arthropoda" rank="species" species="politus">
<emphasis id="B934EAE6FDED280AFAADD53CFA84FC00" box="[1284,1381,882,903]" italics="true" pageId="578" pageNumber="579">M. politus</emphasis>
Olmi
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<paragraph id="8BFF36F4FDED280AFF3ED5F9FC8DFC56" blockId="578.[151,1436,950,1122]" box="[151,876,950,977]" pageId="578" pageNumber="579">
<specimenCount id="9D46FD7DFDED280AFF3ED5F9FF20FC56" box="[151,193,951,977]" pageId="578" pageNumber="579" type="male">♂♂</specimenCount>
(unknown in
<taxonomicName id="4C404D77FDED280AFEF7D5F9FE17FC57" authorityName="Olmi" authorityYear="2007" box="[350,502,950,976]" class="Insecta" family="Dryinidae" genus="Madecadryinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="578" pageNumber="579" phylum="Arthropoda" rank="species" species="humicolus">
<emphasis id="B934EAE6FDED280AFEF7D5F9FE17FC57" box="[350,502,950,976]" italics="true" pageId="578" pageNumber="579">M. humicolus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C404D77FDED280AFDADD5F9FDB2FC57" authorityName="Olmi" box="[516,595,951,976]" class="Insecta" family="Dryinidae" genus="Pseudogonatopus" kingdom="Animalia" order="Hymenoptera" pageId="578" pageNumber="579" phylum="Arthropoda" rank="species" species="opacus">
<emphasis id="B934EAE6FDED280AFDADD5F9FDB2FC57" box="[516,595,951,976]" italics="true" pageId="578" pageNumber="579">opacus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C404D77FDED280AFDC8D5F8FD4CFC57" authorityName="Olmi" authorityYear="2007" box="[609,685,950,976]" class="Insecta" family="Dryinidae" genus="Madecadryinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="578" pageNumber="579" phylum="Arthropoda" rank="species" species="politus">
<emphasis id="B934EAE6FDED280AFDC8D5F8FD4CFC57" box="[609,685,950,976]" italics="true" pageId="578" pageNumber="579">politus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C404D77FDED280AFD4DD5F9FC85FC57" authorityName="Olmi" authorityYear="2007" box="[740,868,951,976]" class="Insecta" family="Dryinidae" genus="Madecadryinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="578" pageNumber="579" phylum="Arthropoda" rank="species" species="semicaecus">
<emphasis id="B934EAE6FDED280AFD4DD5F9FC85FC57" box="[740,868,951,976]" italics="true" pageId="578" pageNumber="579">semicaecus</emphasis>
</taxonomicName>
)
</paragraph>
<keyStep id="30B42E51FDED280AFF3ED599FA7DFBA0" pageId="578" pageNumber="579">
<paragraph id="8BFF36F4FDED280AFF3ED599FA7DFB8D" blockId="578.[151,1436,950,1122]" pageId="578" pageNumber="579">
<keyLead id="30B195C1FDED280AFF3ED599FA7DFB8D" pageId="578" pageNumber="579">
1. Mesoscutum unsculptured or finely punctate and unsculptured among punctures (
<figureCitation id="137B2A71FDED280AFC59D599FBB7FC6A" box="[1008,1110,983,1005]" captionStart="FIGURE 262" captionStartId="586.[151,250,1644,1669]" captionTargetBox="[167,1409,661,1632]" captionTargetId="figure@586.[151,1436,661,1632]" captionTargetPageId="586" captionText="FIGURE 262. Habitus of Madecadryinus species. A: M. relictus Olmi: ♂ from Madagascar, 23°01.31’S 47°43.20’E, in dorsal view. B, C: M. semicaecus Olmi: ♀ holotype in dorsal (B) and lateral (C) view. D: M. silvanus Olmi: ♂ holotype in dorsal view. Scale bar: A: 1.88 mm; D: 2.66 mm." httpUri="https://zenodo.org/record/3337202/files/figure.png" pageId="578" pageNumber="579">Fig. 262D</figureCitation>
); ocelli large (
<figureCitation id="137B2A71FDED280AFB76D599FAA3FC6A" box="[1247,1346,983,1005]" captionStart="FIGURE 262" captionStartId="586.[151,250,1644,1669]" captionTargetBox="[167,1409,661,1632]" captionTargetId="figure@586.[151,1436,661,1632]" captionTargetPageId="586" captionText="FIGURE 262. Habitus of Madecadryinus species. A: M. relictus Olmi: ♂ from Madagascar, 23°01.31’S 47°43.20’E, in dorsal view. B, C: M. semicaecus Olmi: ♀ holotype in dorsal (B) and lateral (C) view. D: M. silvanus Olmi: ♂ holotype in dorsal view. Scale bar: A: 1.88 mm; D: 2.66 mm." httpUri="https://zenodo.org/record/3337202/files/figure.png" pageId="578" pageNumber="579">Fig. 262D</figureCitation>
)............................................................................................
<taxonomicName id="4C404D77FDED280AFB5FD5BBFA7DFB8D" authority="Olmi" authorityName="Olmi" authorityYear="2007" box="[1270,1436,1012,1034]" class="Insecta" family="Dryinidae" genus="Madecadryinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="578" pageNumber="579" phylum="Arthropoda" rank="species" species="silvanus">
<emphasis id="B934EAE6FDED280AFB5FD5BBFA84FB8E" box="[1270,1381,1012,1034]" italics="true" pageId="578" pageNumber="579">M. silvanus</emphasis>
Olmi
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BFF36F4FDED280AFF3ED25CFA7DFBA0" blockId="578.[151,1436,950,1122]" box="[151,1436,1041,1064]" pageId="578" pageNumber="579">
<keyLead id="30B195C1FDED280AFF3ED25CFA7DFBA0" box="[151,1436,1041,1064]" pageId="578" pageNumber="579">
- Mesoscutum granulate; ocelli small or large (
<figureCitation id="137B2A71FDED280AFD3DD25CFD0AFBA0" box="[660,747,1041,1064]" captionStart="FIGURE 259" captionStartId="583.[151,250,1404,1429]" captionTargetBox="[175,1429,429,1369]" captionTargetId="figure@583.[151,1436,409,1392]" captionTargetPageId="583" captionText="FIGURE 259. Madecadryinus ranomafanensis Olmi, Copeland &amp; van Noort sp. nov., ♂ holotype: habitus in dorsal (A) and lateral (B) view; head in frontal view (C); forewing (D)." httpUri="https://zenodo.org/record/3337196/files/figure.png" pageId="578" pageNumber="579">Figs 259</figureCitation>
,
<figureCitation id="137B2A71FDED280AFD5CD25CFCCDFBAF" box="[757,812,1042,1064]" captionStart="FIGURE 262" captionStartId="586.[151,250,1644,1669]" captionTargetBox="[167,1409,661,1632]" captionTargetId="figure@586.[151,1436,661,1632]" captionTargetPageId="586" captionText="FIGURE 262. Habitus of Madecadryinus species. A: M. relictus Olmi: ♂ from Madagascar, 23°01.31’S 47°43.20’E, in dorsal view. B, C: M. semicaecus Olmi: ♀ holotype in dorsal (B) and lateral (C) view. D: M. silvanus Olmi: ♂ holotype in dorsal view. Scale bar: A: 1.88 mm; D: 2.66 mm." httpUri="https://zenodo.org/record/3337202/files/figure.png" pageId="578" pageNumber="579">262A</figureCitation>
)................................................. 2
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30B42E51FDED280AFF3ED261FA7DFBE5" pageId="578" pageNumber="579">
<paragraph id="8BFF36F4FDED280AFF3ED261FA7DFBC2" blockId="578.[151,1436,950,1122]" box="[151,1436,1071,1093]" pageId="578" pageNumber="579">
<keyLead id="30B195C1FDED280AFF3ED261FA7DFBC2" box="[151,1436,1071,1093]" pageId="578" pageNumber="579">
2. POL less than twice as long as OOL (
<figureCitation id="137B2A71FDED280AFDE3D261FD4DFBC2" box="[586,684,1071,1093]" captionStart="FIGURE 260" captionStartId="584.[151,250,1544,1569]" captionTargetBox="[187,1403,194,1508]" captionTargetId="figure@584.[169,1419,174,1533]" captionTargetPageId="584" captionText="FIGURE 260. Madecadryinus relictus Olmi: A, B: ♀ holotype in dorsal (A) and lateral (B) view; ♂ paratype in dorsal (C) and lateral (D) view. Scale bar: A: 1.54 mm; B: 1.17 mm; C: 1.74 mm; D: 1.25 mm." httpUri="https://zenodo.org/record/3337198/files/figure.png" pageId="578" pageNumber="579">Fig. 260C</figureCitation>
)................................................
<taxonomicName id="4C404D77FDED280AFB57D27EFA7DFBC2" authority="Olmi" authorityName="Olmi" authorityYear="2007" box="[1278,1436,1071,1093]" class="Insecta" family="Dryinidae" genus="Madecadryinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="578" pageNumber="579" phylum="Arthropoda" rank="species" species="relictus">
<emphasis id="B934EAE6FDED280AFB57D27EFA84FBC3" box="[1278,1381,1071,1093]" italics="true" pageId="578" pageNumber="579">M. relictus</emphasis>
Olmi
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BFF36F4FDED280AFF3ED203FA7DFBE5" blockId="578.[151,1436,950,1122]" box="[151,1436,1100,1122]" pageId="578" pageNumber="579">
<keyLead id="30B195C1FDED280AFF3ED203FA7DFBE5" box="[151,1436,1100,1122]" pageId="578" pageNumber="579">
- POL about 3 × as long as OOL (
<figureCitation id="137B2A71FDED280AFDBED202FD96FBE5" box="[535,631,1100,1122]" captionStart="FIGURE 258" captionStartId="581.[151,250,1146,1171]" captionTargetBox="[162,1416,190,1114]" captionTargetId="figure@581.[151,1436,174,1134]" captionTargetPageId="581" captionText="FIGURE 258. A, B: Madecadryinus humicolus Olmi, ♀ holotype: head in lateral view (A); chela (B). C, D: Madecadryinus opacus Olmi, ♀ holotype: head in lateral view (C); chela (D). E, G: Madecadryinus politus Olmi, ♀ holotype: head in lateral view (E); chela (G). F, H: Madecadryinus ranomafanensis Olmi, Copeland &amp; van Noort sp. nov., ♂ holotype: head in dorsal view (F); genitalia (H). Scale bar: A: 0.13 mm; B, D: 0.03 mm; C, H: 0.08 mm; E: 0.15 mm; F: 0.17 mm; G: 0.04 mm." httpUri="https://zenodo.org/record/3337194/files/figure.png" pageId="578" pageNumber="579">Fig. 258F</figureCitation>
)..................
<taxonomicName id="4C404D77FDED280AFCC5D203FAA7FBE5" authority="Olmi, Copeland &amp; van Noort" authorityName="Olmi &amp; Copeland &amp; Noort" authorityYear="2019" box="[876,1350,1100,1122]" class="Liliopsida" family="Araceae" genus="Gonatopus" kingdom="Plantae" order="Alismatales" pageId="578" pageNumber="579" phylum="Tracheophyta" rank="species" species="ranomafanensis" status="sp. nov.">
<emphasis id="B934EAE6FDED280AFCC5D203FBC4FBE6" box="[876,1061,1100,1122]" italics="true" pageId="578" pageNumber="579">M. ranomafanensis</emphasis>
Olmi, Copeland &amp; van Noort
</taxonomicName>
,
<taxonomicNameLabel id="A207579DFDED280AFAF9D203FA7DFBE5" box="[1360,1436,1101,1122]" pageId="578" pageNumber="579" rank="species">
<emphasis id="B934EAE6FDED280AFAF9D203FA7DFBE5" bold="true" box="[1360,1436,1101,1122]" pageId="578" pageNumber="579">sp. nov.</emphasis>
</taxonomicNameLabel>
</keyLead>
</paragraph>
</keyStep>
</subSubSection>
</treatment>
</document>
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