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<document id="29B4B599CA231D52B9940588B95A6F55" ID-DOI="10.6620/ZS.2021.60-04" ID-ISSN="1810-522X" ID-PMC="PMC8292842" ID-PubMed="34322170" ID-Zenodo-Dep="12823473" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1721924749745" checkinUser="felipe" docAuthor="Barahona-Segovia, Rodrigo M., Guzmán, Vicente Valdés &amp; Pañinao-Monsálvez, Laura" docDate="2021" docId="03E987A8FFA7FF94FCB94D14F17C5802" docLanguage="en" docName="ZoolStud.60.1.1-10.pdf" docOrigin="Zoological Studies 60 (4)" docSource="http://dx.doi.org/10.5281/zenodo.8055920" docStyle="DocumentStyle:E2601EF9E8BBDC4537EE724C10FF512F.1:ZoolStud.2019-.journal_article" docStyleId="E2601EF9E8BBDC4537EE724C10FF512F" docStyleName="ZoolStud.2019-.journal_article" docStyleVersion="1" docTitle="Epicepon indicum Nierstrasz &amp; Brender" docType="treatment" docVersion="2" lastPageNumber="9" masterDocId="FFD0FFD0FFA4FF9CFFEA4A09F0005B37" masterDocTitle="Holops pullomen Barahona-Segovia &amp; Guzmán &amp; Pañinao-Monsálvez 2021, sp. nov." masterLastPageNumber="10" masterPageNumber="1" pageNumber="4" updateTime="1721926840148" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="B71DDD439260F6A079419401E3A92061">Holops pullomen Barahona-Segovia &amp; Guzmán &amp; Pañinao-Monsálvez 2021, sp. nov.</mods:title>
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<mods:namePart id="1488D1829CF9AC0665AADF36524E2370">Barahona-Segovia, Rodrigo M.</mods:namePart>
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<mods:namePart id="940FEBD6C4703323C7E612A69A2AF9DD">Guzmán, Vicente Valdés</mods:namePart>
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<mods:namePart id="391A6DB5120052F0B683539B8961CD9E">Pañinao-Monsálvez, Laura</mods:namePart>
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<mods:title id="0A1D2EB2C7A201436748043E79E0087F">Zoological Studies</mods:title>
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<mods:date id="8891051D68578359EDD33D99B396AA72">2021</mods:date>
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<treatment id="03E987A8FFA7FF94FCB94D14F17C5802" LSID="urn:lsid:plazi:treatment:03E987A8FFA7FF94FCB94D14F17C5802" httpUri="http://treatment.plazi.org/id/03E987A8FFA7FF94FCB94D14F17C5802" lastPageId="8" lastPageNumber="9" pageId="3" pageNumber="4">
<subSubSection id="C35A6535FFA7FF9FFCB94D14F4815C42" pageId="3" pageNumber="4" type="nomenclature">
<paragraph id="8BFF36BEFFA7FF9FFCB94D14F4815C42" blockId="3.[851,1376,1820,1909]" pageId="3" pageNumber="4">
<emphasis id="B934EAACFFA7FF9FFCB94D14F4AF5C61" bold="true" pageId="3" pageNumber="4">
<taxonomicName id="4C404D3DFFA7FF9FFCB94D14F5495C01" ID-CoL="6FR87" authority="Nierstrasz &amp; Brender" authorityName="Nierstrasz &amp; Brender" box="[851,1353,1821,1846]" class="Malacostraca" family="Bopyridae" genus="Epicepon" kingdom="Animalia" order="Isopoda" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="indicum">
<emphasis id="B934EAACFFA7FF9FFCB94D14F4385C01" bold="true" box="[851,1080,1821,1846]" italics="true" pageId="3" pageNumber="4">Epicepon indicum</emphasis>
Nierstrasz &amp; Brender
</taxonomicName>
à Brandis, 1931
</emphasis>
(
<figureCitation id="137B2A3BFFA7FF9FFBD34D54F4795C42" box="[1081,1145,1885,1909]" captionStart="Fig" captionStartId="4.[174,205,1842,1861]" captionTargetBox="[177,1429,917,1805]" captionTargetId="figure-401@4.[174,1454,913,1815]" captionTargetPageId="4" captionText="Fig. 2. Epicepon indicum Nierstrasz and Brender à Brandis, 1931, female syntype, ZMUC CR-6788 (A–H) and male syntype, CR-6788 (I–J). A, dorsal view. B, left antennule (A1) and antenna (A2). C, right pereopod 1. D, right pereopod 7. E, left side of barbula. F, right oostegite 1, internal view. G, right maxilliped, external view with outer projection of barbula (OB). H, pleon, ventral view showing uropods (Ur), fifth lateral plates (Lp5), exopod (Ex5) and endopod (En5). I, dorsal view. J, lateral view. Scale bars: B = 25 µm, C, D = 50 µm, E = 100 µm, F, G = 200 µm. A, H–J, from Nierstrasz and Brender à Brandis (1931); not to scale." figureDoi="http://doi.org/10.5281/zenodo.12823477" httpUri="https://zenodo.org/record/12823477/files/figure.png" pageId="3" pageNumber="4">Fig. 2</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C35A6535FFA7FF94FCC74D90F17C5802" lastPageId="8" lastPageNumber="9" pageId="3" pageNumber="4" type="description">
<paragraph id="8BFF36BEFFA7FF9FFCC74D90F5865C9C" blockId="3.[813,1414,1944,1963]" box="[813,1414,1944,1963]" pageId="3" pageNumber="4">
<taxonomicName id="4C404D3DFFA7FF9FFCC74D90F4925C9C" authority="Nierstrasz &amp; Brender" authorityName="Nierstrasz &amp; Brender" box="[813,1170,1944,1963]" class="Malacostraca" family="Bopyridae" genus="Epicepon" kingdom="Animalia" order="Isopoda" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="indicum">
<emphasis id="B934EAACFFA7FF9FFCC74D90F3CC5C9C" box="[813,972,1944,1963]" italics="true" pageId="3" pageNumber="4">Epicepon indicum</emphasis>
Nierstrasz &amp; Brender
</taxonomicName>
à Brandis, 1931: 189–191,
</paragraph>
<paragraph id="8BFF36BEFFA0FF98FF314AF4F2395A27" blockId="4.[219,569,253,272]" box="[219,569,253,272]" pageId="4" pageNumber="5">
figs. 74–77;
<bibRefCitation id="EFD14B4FFFA0FF98FEAF4AF4F2045A27" author="Dollfus R-P &amp; Neveu-Lemaire M &amp; Galliard H." box="[325,516,253,272]" pageId="4" pageNumber="5" pagination="70 - 80" refId="ref5905" refString="Dollfus R-P, Neveu-Lemaire M, Galliard H. 1933. Repertoire des especes et des genres nouveaux. Ann Parasit Hum Comp 11 (1): 70 - 80." type="journal article" year="1933">Dollfus et al. 1933: 76</bibRefCitation>
[list].
</paragraph>
<paragraph id="8BFF36BEFFA0FF98FF004B31F1205AE7" blockId="4.[174,775,312,848]" pageId="4" pageNumber="5">
<emphasis id="B934EAACFFA0FF98FF004B31F1B55A67" box="[234,437,312,336]" italics="true" pageId="4" pageNumber="5">Material examined</emphasis>
: Female
<typeStatus id="54FB881CFFA0FF98FDF04B30F26D5A66" box="[538,621,313,337]" pageId="4" pageNumber="5" type="syntype">syntype</typeStatus>
(
<quantity id="4CB89B5BFFA0FF98FD964B31F2CF5A67" box="[636,719,312,336]" metricMagnitude="-3" metricUnit="m" metricValue="2.8" pageId="4" pageNumber="5" unit="mm" value="2.8">2.8 mm</quantity>
TL), parasitizing
<taxonomicName id="4C404D3DFFA0FF98FED94B51F1B35A47" box="[307,435,344,368]" class="Malacostraca" family="Cyclodorippidae" genus="Tymolus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B934EAACFFA0FF98FED94B51F18B5A47" box="[307,395,344,368]" italics="true" pageId="4" pageNumber="5">Tymolus</emphasis>
sp.
</taxonomicName>
, Kei Islands,
<collectingCountry id="F357762EFFA0FF98FDA04B51F2B45A47" box="[586,692,344,368]" name="Indonesia" pageId="4" pageNumber="5">Indonesia</collectingCountry>
, St. 56,
<geoCoordinate id="EE745079FFA0FF98FF444B71F1065AA7" box="[174,262,376,400]" degrees="05" direction="south" minutes="30" orientation="latitude" pageId="4" pageNumber="5" precision="925" value="-5.5">05°30'S</geoCoordinate>
,
<geoCoordinate id="EE745079FFA0FF98FEF94B71F17A5AA7" box="[275,378,376,400]" degrees="132" direction="east" minutes="51" orientation="longitude" pageId="4" pageNumber="5" precision="925" value="132.85">132°51'E</geoCoordinate>
,
<quantity id="4CB89B5BFFA0FF98FE624B71F1D05AA7" box="[392,464,376,400]" metricMagnitude="2" metricUnit="m" metricValue="3.45" pageId="4" pageNumber="5" unit="m" value="345.0">345 m</quantity>
, coll. T. Mortensen, Danish Expedition to the Kei Islands,
<date id="FFFE107EFFA0FF98FDE74B91F2A55A87" box="[525,677,408,433]" pageId="4" pageNumber="5" value="1922-05-10">10 May 1922</date>
(ZMUC CR-6788).
</paragraph>
<paragraph id="8BFF36BEFFA0FF98FF004BD1F1A95947" blockId="4.[174,775,312,848]" pageId="4" pageNumber="5">
<emphasis id="B934EAACFFA0FF98FF004BD1F1965AC7" box="[234,406,472,496]" italics="true" pageId="4" pageNumber="5">Redescription</emphasis>
: Female (
<figureCitation id="137B2A3BFFA0FF98FDF64BD1F2A55AC7" box="[540,677,472,496]" captionStart="Fig" captionStartId="4.[174,205,1842,1861]" captionTargetBox="[177,1429,917,1805]" captionTargetId="figure-401@4.[174,1454,913,1815]" captionTargetPageId="4" captionText="Fig. 2. Epicepon indicum Nierstrasz and Brender à Brandis, 1931, female syntype, ZMUC CR-6788 (A–H) and male syntype, CR-6788 (I–J). A, dorsal view. B, left antennule (A1) and antenna (A2). C, right pereopod 1. D, right pereopod 7. E, left side of barbula. F, right oostegite 1, internal view. G, right maxilliped, external view with outer projection of barbula (OB). H, pleon, ventral view showing uropods (Ur), fifth lateral plates (Lp5), exopod (Ex5) and endopod (En5). I, dorsal view. J, lateral view. Scale bars: B = 25 µm, C, D = 50 µm, E = 100 µm, F, G = 200 µm. A, H–J, from Nierstrasz and Brender à Brandis (1931); not to scale." figureDoi="http://doi.org/10.5281/zenodo.12823477" httpUri="https://zenodo.org/record/12823477/files/figure.png" pageId="4" pageNumber="5">Fig. 2A–H</figureCitation>
) length
<quantity id="4CB89B5BFFA0FF98FF444BF1F1035927" box="[174,259,504,528]" metricMagnitude="-3" metricUnit="m" metricValue="2.8" pageId="4" pageNumber="5" unit="mm" value="2.8">2.8 mm</quantity>
; maximum width (across pereomere 3)
<quantity id="4CB89B5BFFA0FF98FD474BF1F3035927" box="[685,771,504,528]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" pageId="4" pageNumber="5" unit="mm" value="1.3">1.3 mm</quantity>
; head length
<quantity id="4CB89B5BFFA0FF98FEDF4811F18D5907" box="[309,397,536,560]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="4" pageNumber="5" unit="mm" value="0.5">0.5 mm</quantity>
; head width
<quantity id="4CB89B5BFFA0FF98FDF14811F2735907" box="[539,627,536,560]" metricMagnitude="-4" metricUnit="m" metricValue="7.0" pageId="4" pageNumber="5" unit="mm" value="0.7">0.7 mm</quantity>
; pleon length
<quantity id="4CB89B5BFFA0FF98FF444831F1065967" box="[174,262,568,592]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="4" pageNumber="5" unit="mm" value="1.2">1.2 mm</quantity>
. Body linear (no asymmetry); all segments of body distinct (
<figureCitation id="137B2A3BFFA0FF98FEAD4851F19D5947" box="[327,413,600,624]" captionStart="Fig" captionStartId="4.[174,205,1842,1861]" captionTargetBox="[177,1429,917,1805]" captionTargetId="figure-401@4.[174,1454,913,1815]" captionTargetPageId="4" captionText="Fig. 2. Epicepon indicum Nierstrasz and Brender à Brandis, 1931, female syntype, ZMUC CR-6788 (A–H) and male syntype, CR-6788 (I–J). A, dorsal view. B, left antennule (A1) and antenna (A2). C, right pereopod 1. D, right pereopod 7. E, left side of barbula. F, right oostegite 1, internal view. G, right maxilliped, external view with outer projection of barbula (OB). H, pleon, ventral view showing uropods (Ur), fifth lateral plates (Lp5), exopod (Ex5) and endopod (En5). I, dorsal view. J, lateral view. Scale bars: B = 25 µm, C, D = 50 µm, E = 100 µm, F, G = 200 µm. A, H–J, from Nierstrasz and Brender à Brandis (1931); not to scale." figureDoi="http://doi.org/10.5281/zenodo.12823477" httpUri="https://zenodo.org/record/12823477/files/figure.png" pageId="4" pageNumber="5">Fig. 2A</figureCitation>
).
</paragraph>
<paragraph id="8BFF36BEFFA0FF98FF004871F53B5AA1" blockId="4.[174,775,312,848]" lastBlockId="4.[850,1452,253,854]" pageId="4" pageNumber="5">
Head ovate, wider than long, frontal margin rounded medially. Frontal lamina relatively thin and extending beyond sides and frontal margin of head (
<figureCitation id="137B2A3BFFA0FF98FF5C48D1F10E59C7" box="[182,270,728,752]" captionStart="Fig" captionStartId="4.[174,205,1842,1861]" captionTargetBox="[177,1429,917,1805]" captionTargetId="figure-401@4.[174,1454,913,1815]" captionTargetPageId="4" captionText="Fig. 2. Epicepon indicum Nierstrasz and Brender à Brandis, 1931, female syntype, ZMUC CR-6788 (A–H) and male syntype, CR-6788 (I–J). A, dorsal view. B, left antennule (A1) and antenna (A2). C, right pereopod 1. D, right pereopod 7. E, left side of barbula. F, right oostegite 1, internal view. G, right maxilliped, external view with outer projection of barbula (OB). H, pleon, ventral view showing uropods (Ur), fifth lateral plates (Lp5), exopod (Ex5) and endopod (En5). I, dorsal view. J, lateral view. Scale bars: B = 25 µm, C, D = 50 µm, E = 100 µm, F, G = 200 µm. A, H–J, from Nierstrasz and Brender à Brandis (1931); not to scale." figureDoi="http://doi.org/10.5281/zenodo.12823477" httpUri="https://zenodo.org/record/12823477/files/figure.png" pageId="4" pageNumber="5">Fig. 2A</figureCitation>
). Eyes lacking. Antennules and antennae with three and four articles each, respectively, distalmost article of antennule and two distalmost articles of antenna terminally setose (
<figureCitation id="137B2A3BFFA0FF98FE334931F2315867" box="[473,561,824,848]" captionStart="Fig" captionStartId="4.[174,205,1842,1861]" captionTargetBox="[177,1429,917,1805]" captionTargetId="figure-401@4.[174,1454,913,1815]" captionTargetPageId="4" captionText="Fig. 2. Epicepon indicum Nierstrasz and Brender à Brandis, 1931, female syntype, ZMUC CR-6788 (A–H) and male syntype, CR-6788 (I–J). A, dorsal view. B, left antennule (A1) and antenna (A2). C, right pereopod 1. D, right pereopod 7. E, left side of barbula. F, right oostegite 1, internal view. G, right maxilliped, external view with outer projection of barbula (OB). H, pleon, ventral view showing uropods (Ur), fifth lateral plates (Lp5), exopod (Ex5) and endopod (En5). I, dorsal view. J, lateral view. Scale bars: B = 25 µm, C, D = 50 µm, E = 100 µm, F, G = 200 µm. A, H–J, from Nierstrasz and Brender à Brandis (1931); not to scale." figureDoi="http://doi.org/10.5281/zenodo.12823477" httpUri="https://zenodo.org/record/12823477/files/figure.png" pageId="4" pageNumber="5">Fig. 2B</figureCitation>
). Barbula with one slender, distally rounded lateral projection on each side, middle region smooth with low lateral lobe (
<figureCitation id="137B2A3BFFA0FF98FABA4B14F3695A61" captionStart="Fig" captionStartId="4.[174,205,1842,1861]" captionTargetBox="[177,1429,917,1805]" captionTargetId="figure-401@4.[174,1454,913,1815]" captionTargetPageId="4" captionText="Fig. 2. Epicepon indicum Nierstrasz and Brender à Brandis, 1931, female syntype, ZMUC CR-6788 (A–H) and male syntype, CR-6788 (I–J). A, dorsal view. B, left antennule (A1) and antenna (A2). C, right pereopod 1. D, right pereopod 7. E, left side of barbula. F, right oostegite 1, internal view. G, right maxilliped, external view with outer projection of barbula (OB). H, pleon, ventral view showing uropods (Ur), fifth lateral plates (Lp5), exopod (Ex5) and endopod (En5). I, dorsal view. J, lateral view. Scale bars: B = 25 µm, C, D = 50 µm, E = 100 µm, F, G = 200 µm. A, H–J, from Nierstrasz and Brender à Brandis (1931); not to scale." figureDoi="http://doi.org/10.5281/zenodo.12823477" httpUri="https://zenodo.org/record/12823477/files/figure.png" pageId="4" pageNumber="5">Fig. 2E, G</figureCitation>
). Maxilliped with prominent anterior segment, palp as low non-articulated projection with few distal setae, plectron triangular, short and broad (
<figureCitation id="137B2A3BFFA0FF98FB334B74F52F5AA1" box="[1241,1327,381,406]" captionStart="Fig" captionStartId="4.[174,205,1842,1861]" captionTargetBox="[177,1429,917,1805]" captionTargetId="figure-401@4.[174,1454,913,1815]" captionTargetPageId="4" captionText="Fig. 2. Epicepon indicum Nierstrasz and Brender à Brandis, 1931, female syntype, ZMUC CR-6788 (A–H) and male syntype, CR-6788 (I–J). A, dorsal view. B, left antennule (A1) and antenna (A2). C, right pereopod 1. D, right pereopod 7. E, left side of barbula. F, right oostegite 1, internal view. G, right maxilliped, external view with outer projection of barbula (OB). H, pleon, ventral view showing uropods (Ur), fifth lateral plates (Lp5), exopod (Ex5) and endopod (En5). I, dorsal view. J, lateral view. Scale bars: B = 25 µm, C, D = 50 µm, E = 100 µm, F, G = 200 µm. A, H–J, from Nierstrasz and Brender à Brandis (1931); not to scale." figureDoi="http://doi.org/10.5281/zenodo.12823477" httpUri="https://zenodo.org/record/12823477/files/figure.png" pageId="4" pageNumber="5">Fig. 2G</figureCitation>
).
</paragraph>
<paragraph id="8BFF36BEFFA0FF98FC654B97F4505861" blockId="4.[850,1452,253,854]" pageId="4" pageNumber="5">
Pereon broadest across pereomere 3. Irregularly shaped tergal projections prominent on pereomeres 1–5 on right side, 2–4 on left side, ovate dorsolateral bosses on pereomeres 1–5 on right side, 2–3 on left side, thin coxal plates on both sides of pereomeres 1–5 (
<figureCitation id="137B2A3BFFA0FF98FA974814F3785961" captionStart="Fig" captionStartId="4.[174,205,1842,1861]" captionTargetBox="[177,1429,917,1805]" captionTargetId="figure-401@4.[174,1454,913,1815]" captionTargetPageId="4" captionText="Fig. 2. Epicepon indicum Nierstrasz and Brender à Brandis, 1931, female syntype, ZMUC CR-6788 (A–H) and male syntype, CR-6788 (I–J). A, dorsal view. B, left antennule (A1) and antenna (A2). C, right pereopod 1. D, right pereopod 7. E, left side of barbula. F, right oostegite 1, internal view. G, right maxilliped, external view with outer projection of barbula (OB). H, pleon, ventral view showing uropods (Ur), fifth lateral plates (Lp5), exopod (Ex5) and endopod (En5). I, dorsal view. J, lateral view. Scale bars: B = 25 µm, C, D = 50 µm, E = 100 µm, F, G = 200 µm. A, H–J, from Nierstrasz and Brender à Brandis (1931); not to scale." figureDoi="http://doi.org/10.5281/zenodo.12823477" httpUri="https://zenodo.org/record/12823477/files/figure.png" pageId="4" pageNumber="5">Fig. 2A</figureCitation>
). No mid-dorsal projections on any pereomeres. Oostegites enclosing brood pouch, oostegite 1 (
<figureCitation id="137B2A3BFFA0FF98FAB94854F5A45941" box="[1363,1444,605,630]" captionStart="Fig" captionStartId="4.[174,205,1842,1861]" captionTargetBox="[177,1429,917,1805]" captionTargetId="figure-401@4.[174,1454,913,1815]" captionTargetPageId="4" captionText="Fig. 2. Epicepon indicum Nierstrasz and Brender à Brandis, 1931, female syntype, ZMUC CR-6788 (A–H) and male syntype, CR-6788 (I–J). A, dorsal view. B, left antennule (A1) and antenna (A2). C, right pereopod 1. D, right pereopod 7. E, left side of barbula. F, right oostegite 1, internal view. G, right maxilliped, external view with outer projection of barbula (OB). H, pleon, ventral view showing uropods (Ur), fifth lateral plates (Lp5), exopod (Ex5) and endopod (En5). I, dorsal view. J, lateral view. Scale bars: B = 25 µm, C, D = 50 µm, E = 100 µm, F, G = 200 µm. A, H–J, from Nierstrasz and Brender à Brandis (1931); not to scale." figureDoi="http://doi.org/10.5281/zenodo.12823477" httpUri="https://zenodo.org/record/12823477/files/figure.png" pageId="4" pageNumber="5">Fig. 2F</figureCitation>
) with ovate anterior article, longer than posterior article, internal ridge smooth, posterior article rounded on lateral margin, extending to triangular point posteriorly. Pereopods subequal in structure, first four shorter than others (
<figureCitation id="137B2A3BFFA0FF98FC4848F4F3F85821" box="[930,1016,765,790]" captionStart="Fig" captionStartId="4.[174,205,1842,1861]" captionTargetBox="[177,1429,917,1805]" captionTargetId="figure-401@4.[174,1454,913,1815]" captionTargetPageId="4" captionText="Fig. 2. Epicepon indicum Nierstrasz and Brender à Brandis, 1931, female syntype, ZMUC CR-6788 (A–H) and male syntype, CR-6788 (I–J). A, dorsal view. B, left antennule (A1) and antenna (A2). C, right pereopod 1. D, right pereopod 7. E, left side of barbula. F, right oostegite 1, internal view. G, right maxilliped, external view with outer projection of barbula (OB). H, pleon, ventral view showing uropods (Ur), fifth lateral plates (Lp5), exopod (Ex5) and endopod (En5). I, dorsal view. J, lateral view. Scale bars: B = 25 µm, C, D = 50 µm, E = 100 µm, F, G = 200 µm. A, H–J, from Nierstrasz and Brender à Brandis (1931); not to scale." figureDoi="http://doi.org/10.5281/zenodo.12823477" httpUri="https://zenodo.org/record/12823477/files/figure.png" pageId="4" pageNumber="5">Fig. 2A</figureCitation>
). All pereopods with short, blunt dactyli, cylindrical propodi, carpi and meri fused in posterior pereopods (
<figureCitation id="137B2A3BFFA0FF98FC254934F4435861" box="[975,1091,829,854]" captionStart="Fig" captionStartId="4.[174,205,1842,1861]" captionTargetBox="[177,1429,917,1805]" captionTargetId="figure-401@4.[174,1454,913,1815]" captionTargetPageId="4" captionText="Fig. 2. Epicepon indicum Nierstrasz and Brender à Brandis, 1931, female syntype, ZMUC CR-6788 (A–H) and male syntype, CR-6788 (I–J). A, dorsal view. B, left antennule (A1) and antenna (A2). C, right pereopod 1. D, right pereopod 7. E, left side of barbula. F, right oostegite 1, internal view. G, right maxilliped, external view with outer projection of barbula (OB). H, pleon, ventral view showing uropods (Ur), fifth lateral plates (Lp5), exopod (Ex5) and endopod (En5). I, dorsal view. J, lateral view. Scale bars: B = 25 µm, C, D = 50 µm, E = 100 µm, F, G = 200 µm. A, H–J, from Nierstrasz and Brender à Brandis (1931); not to scale." figureDoi="http://doi.org/10.5281/zenodo.12823477" httpUri="https://zenodo.org/record/12823477/files/figure.png" pageId="4" pageNumber="5">Fig. 2C, D</figureCitation>
).
</paragraph>
<caption id="DF3F6636FFA0FF98FF444D3BF27A5C82" ID-DOI="http://doi.org/10.5281/zenodo.12823477" ID-Zenodo-Dep="12823477" httpUri="https://zenodo.org/record/12823477/files/figure.png" pageId="4" pageNumber="5" startId="4.[174,205,1842,1861]" targetBox="[177,1429,917,1805]" targetPageId="4" targetType="figure">
<paragraph id="8BFF36BEFFA0FF98FF444D3BF27A5C82" blockId="4.[174,1452,1842,1974]" pageId="4" pageNumber="5">
<emphasis id="B934EAACFFA0FF98FF444D3BF0E85C72" bold="true" box="[174,232,1842,1861]" pageId="4" pageNumber="5">Fig. 2.</emphasis>
<taxonomicName id="4C404D3DFFA0FF98FF1C4D3AF2625C72" authority="Nierstrasz and Brender" authorityName="Nierstrasz and Brender" box="[246,610,1842,1861]" class="Malacostraca" family="Bopyridae" genus="Epicepon" kingdom="Animalia" order="Isopoda" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="indicum">
<emphasis id="B934EAACFFA0FF98FF1C4D3AF1935C72" box="[246,403,1842,1861]" italics="true" pageId="4" pageNumber="5">Epicepon indicum</emphasis>
Nierstrasz and Brender
</taxonomicName>
à Brandis, 1931, female syntype, ZMUC CR-6788 (A–H) and male syntype, CR-6788 (I–J). A, dorsal view. B, left antennule (A1) and antenna (A2). C, right pereopod 1. D, right pereopod 7. E, left side of barbula. F, right oostegite 1, internal view. G, right maxilliped, external view with outer projection of barbula (OB). H, pleon, ventral view showing uropods (Ur), fifth lateral plates (Lp5), exopod (Ex5) and endopod (En5). I, dorsal view. J, lateral view. Scale bars: B = 25 µm, C, D = 50 µm, E = 100 µm, F, G = 200 µm. A, H–J, from Nierstrasz and Brender à Brandis (1931); not to scale.
</paragraph>
</caption>
<paragraph id="8BFF36BEFFA1FF99FF2F4AF4F2765901" blockId="5.[136,738,253,1621]" pageId="5" pageNumber="6">
Pleon with six segments, first five pleomeres each bearing five pairs of biramous pleopods and longer uniramous lateral plates, all margins with slight undulation but not digitate, surfaces smooth (
<figureCitation id="137B2A3BFFA1FF99FF7A4B74F1115AA1" box="[144,273,381,406]" captionStart="Fig" captionStartId="4.[174,205,1842,1861]" captionTargetBox="[177,1429,917,1805]" captionTargetId="figure-401@4.[174,1454,913,1815]" captionTargetPageId="4" captionText="Fig. 2. Epicepon indicum Nierstrasz and Brender à Brandis, 1931, female syntype, ZMUC CR-6788 (A–H) and male syntype, CR-6788 (I–J). A, dorsal view. B, left antennule (A1) and antenna (A2). C, right pereopod 1. D, right pereopod 7. E, left side of barbula. F, right oostegite 1, internal view. G, right maxilliped, external view with outer projection of barbula (OB). H, pleon, ventral view showing uropods (Ur), fifth lateral plates (Lp5), exopod (Ex5) and endopod (En5). I, dorsal view. J, lateral view. Scale bars: B = 25 µm, C, D = 50 µm, E = 100 µm, F, G = 200 µm. A, H–J, from Nierstrasz and Brender à Brandis (1931); not to scale." figureDoi="http://doi.org/10.5281/zenodo.12823477" httpUri="https://zenodo.org/record/12823477/files/figure.png" pageId="5" pageNumber="6">Fig. 2A, H</figureCitation>
). Short digitiform endopods of pleopods approximately half as long as slender, distally tapering exopods (
<figureCitation id="137B2A3BFFA1FF99FEEA4BB4F1645AE1" box="[256,356,445,470]" captionStart="Fig" captionStartId="4.[174,205,1842,1861]" captionTargetBox="[177,1429,917,1805]" captionTargetId="figure-401@4.[174,1454,913,1815]" captionTargetPageId="4" captionText="Fig. 2. Epicepon indicum Nierstrasz and Brender à Brandis, 1931, female syntype, ZMUC CR-6788 (A–H) and male syntype, CR-6788 (I–J). A, dorsal view. B, left antennule (A1) and antenna (A2). C, right pereopod 1. D, right pereopod 7. E, left side of barbula. F, right oostegite 1, internal view. G, right maxilliped, external view with outer projection of barbula (OB). H, pleon, ventral view showing uropods (Ur), fifth lateral plates (Lp5), exopod (Ex5) and endopod (En5). I, dorsal view. J, lateral view. Scale bars: B = 25 µm, C, D = 50 µm, E = 100 µm, F, G = 200 µm. A, H–J, from Nierstrasz and Brender à Brandis (1931); not to scale." figureDoi="http://doi.org/10.5281/zenodo.12823477" httpUri="https://zenodo.org/record/12823477/files/figure.png" pageId="5" pageNumber="6">Fig. 2H</figureCitation>
). Terminal pleomere ending in uniramous uropods, similar to but slightly longer than fifth pleopodal exopods, surfaces smooth, margins with slight undulation but not digitate (
<figureCitation id="137B2A3BFFA1FF99FE194814F26A5901" box="[499,618,541,566]" captionStart="Fig" captionStartId="4.[174,205,1842,1861]" captionTargetBox="[177,1429,917,1805]" captionTargetId="figure-401@4.[174,1454,913,1815]" captionTargetPageId="4" captionText="Fig. 2. Epicepon indicum Nierstrasz and Brender à Brandis, 1931, female syntype, ZMUC CR-6788 (A–H) and male syntype, CR-6788 (I–J). A, dorsal view. B, left antennule (A1) and antenna (A2). C, right pereopod 1. D, right pereopod 7. E, left side of barbula. F, right oostegite 1, internal view. G, right maxilliped, external view with outer projection of barbula (OB). H, pleon, ventral view showing uropods (Ur), fifth lateral plates (Lp5), exopod (Ex5) and endopod (En5). I, dorsal view. J, lateral view. Scale bars: B = 25 µm, C, D = 50 µm, E = 100 µm, F, G = 200 µm. A, H–J, from Nierstrasz and Brender à Brandis (1931); not to scale." figureDoi="http://doi.org/10.5281/zenodo.12823477" httpUri="https://zenodo.org/record/12823477/files/figure.png" pageId="5" pageNumber="6">Fig. 2A, H</figureCitation>
).
</paragraph>
<paragraph id="8BFF36BEFFA1FF99FF2F4834F1485942" blockId="5.[136,738,253,1621]" pageId="5" pageNumber="6">
Male (
<figureCitation id="137B2A3BFFA1FF99FEFC4834F1895961" box="[278,393,573,598]" captionStart="Fig" captionStartId="4.[174,205,1842,1861]" captionTargetBox="[177,1429,917,1805]" captionTargetId="figure-401@4.[174,1454,913,1815]" captionTargetPageId="4" captionText="Fig. 2. Epicepon indicum Nierstrasz and Brender à Brandis, 1931, female syntype, ZMUC CR-6788 (A–H) and male syntype, CR-6788 (I–J). A, dorsal view. B, left antennule (A1) and antenna (A2). C, right pereopod 1. D, right pereopod 7. E, left side of barbula. F, right oostegite 1, internal view. G, right maxilliped, external view with outer projection of barbula (OB). H, pleon, ventral view showing uropods (Ur), fifth lateral plates (Lp5), exopod (Ex5) and endopod (En5). I, dorsal view. J, lateral view. Scale bars: B = 25 µm, C, D = 50 µm, E = 100 µm, F, G = 200 µm. A, H–J, from Nierstrasz and Brender à Brandis (1931); not to scale." figureDoi="http://doi.org/10.5281/zenodo.12823477" httpUri="https://zenodo.org/record/12823477/files/figure.png" pageId="5" pageNumber="6">Fig. 2I–J</figureCitation>
) with all body regions and segments distinct.
</paragraph>
<paragraph id="8BFF36BEFFA1FF99FF2F4877F1095802" blockId="5.[136,738,253,1621]" pageId="5" pageNumber="6">
Head broadly rounded distally, extending posteriorly to rounded point overlapping approximately 1/3 length of pereomere 1 (
<figureCitation id="137B2A3BFFA1FF99FE5048B4F20859E1" box="[442,520,701,726]" captionStart="Fig" captionStartId="4.[174,205,1842,1861]" captionTargetBox="[177,1429,917,1805]" captionTargetId="figure-401@4.[174,1454,913,1815]" captionTargetPageId="4" captionText="Fig. 2. Epicepon indicum Nierstrasz and Brender à Brandis, 1931, female syntype, ZMUC CR-6788 (A–H) and male syntype, CR-6788 (I–J). A, dorsal view. B, left antennule (A1) and antenna (A2). C, right pereopod 1. D, right pereopod 7. E, left side of barbula. F, right oostegite 1, internal view. G, right maxilliped, external view with outer projection of barbula (OB). H, pleon, ventral view showing uropods (Ur), fifth lateral plates (Lp5), exopod (Ex5) and endopod (En5). I, dorsal view. J, lateral view. Scale bars: B = 25 µm, C, D = 50 µm, E = 100 µm, F, G = 200 µm. A, H–J, from Nierstrasz and Brender à Brandis (1931); not to scale." figureDoi="http://doi.org/10.5281/zenodo.12823477" httpUri="https://zenodo.org/record/12823477/files/figure.png" pageId="5" pageNumber="6">Fig. 2I</figureCitation>
). Small, ovate dark eyes near posterolateral margins of head. Antennules and antennae not visible beyond margins of head in dorsal view.
</paragraph>
<paragraph id="8BFF36BEFFA1FF99FF2F4937F0E958E1" blockId="5.[136,738,253,1621]" pageId="5" pageNumber="6">
Pereomeres 2 – 4 subequal in width, other pereomeres only slightly narrower (
<figureCitation id="137B2A3BFFA1FF99FDA34954F2A25841" box="[585,674,861,886]" captionStart="Fig" captionStartId="4.[174,205,1842,1861]" captionTargetBox="[177,1429,917,1805]" captionTargetId="figure-401@4.[174,1454,913,1815]" captionTargetPageId="4" captionText="Fig. 2. Epicepon indicum Nierstrasz and Brender à Brandis, 1931, female syntype, ZMUC CR-6788 (A–H) and male syntype, CR-6788 (I–J). A, dorsal view. B, left antennule (A1) and antenna (A2). C, right pereopod 1. D, right pereopod 7. E, left side of barbula. F, right oostegite 1, internal view. G, right maxilliped, external view with outer projection of barbula (OB). H, pleon, ventral view showing uropods (Ur), fifth lateral plates (Lp5), exopod (Ex5) and endopod (En5). I, dorsal view. J, lateral view. Scale bars: B = 25 µm, C, D = 50 µm, E = 100 µm, F, G = 200 µm. A, H–J, from Nierstrasz and Brender à Brandis (1931); not to scale." figureDoi="http://doi.org/10.5281/zenodo.12823477" httpUri="https://zenodo.org/record/12823477/files/figure.png" pageId="5" pageNumber="6">Fig. 2I</figureCitation>
). All pereomeres tapered laterally, lacking mid-ventral tubercles. All pereopods subequal in structure and size (
<figureCitation id="137B2A3BFFA1FF99FF7A49B4F0DD58E1" box="[144,221,957,982]" captionStart="Fig" captionStartId="4.[174,205,1842,1861]" captionTargetBox="[177,1429,917,1805]" captionTargetId="figure-401@4.[174,1454,913,1815]" captionTargetPageId="4" captionText="Fig. 2. Epicepon indicum Nierstrasz and Brender à Brandis, 1931, female syntype, ZMUC CR-6788 (A–H) and male syntype, CR-6788 (I–J). A, dorsal view. B, left antennule (A1) and antenna (A2). C, right pereopod 1. D, right pereopod 7. E, left side of barbula. F, right oostegite 1, internal view. G, right maxilliped, external view with outer projection of barbula (OB). H, pleon, ventral view showing uropods (Ur), fifth lateral plates (Lp5), exopod (Ex5) and endopod (En5). I, dorsal view. J, lateral view. Scale bars: B = 25 µm, C, D = 50 µm, E = 100 µm, F, G = 200 µm. A, H–J, from Nierstrasz and Brender à Brandis (1931); not to scale." figureDoi="http://doi.org/10.5281/zenodo.12823477" httpUri="https://zenodo.org/record/12823477/files/figure.png" pageId="5" pageNumber="6">Fig. 2J</figureCitation>
).
</paragraph>
<paragraph id="8BFF36BEFFA1FF99FF2F49D4F23F5F81" blockId="5.[136,738,253,1621]" pageId="5" pageNumber="6">
Pleon with six pleomeres, rounded laterally, each slightly narrower than preceding one (
<figureCitation id="137B2A3BFFA1FF99FDAB49F4F2935F21" box="[577,659,1021,1046]" captionStart="Fig" captionStartId="4.[174,205,1842,1861]" captionTargetBox="[177,1429,917,1805]" captionTargetId="figure-401@4.[174,1454,913,1815]" captionTargetPageId="4" captionText="Fig. 2. Epicepon indicum Nierstrasz and Brender à Brandis, 1931, female syntype, ZMUC CR-6788 (A–H) and male syntype, CR-6788 (I–J). A, dorsal view. B, left antennule (A1) and antenna (A2). C, right pereopod 1. D, right pereopod 7. E, left side of barbula. F, right oostegite 1, internal view. G, right maxilliped, external view with outer projection of barbula (OB). H, pleon, ventral view showing uropods (Ur), fifth lateral plates (Lp5), exopod (Ex5) and endopod (En5). I, dorsal view. J, lateral view. Scale bars: B = 25 µm, C, D = 50 µm, E = 100 µm, F, G = 200 µm. A, H–J, from Nierstrasz and Brender à Brandis (1931); not to scale." figureDoi="http://doi.org/10.5281/zenodo.12823477" httpUri="https://zenodo.org/record/12823477/files/figure.png" pageId="5" pageNumber="6">Fig. 2I</figureCitation>
). Mid-ventral tubercles lacking, pleopods as ventrolateral swelling on pleomeres 1–5 (
<figureCitation id="137B2A3BFFA1FF99FE0F4E34F23F5F61" box="[485,575,1085,1110]" captionStart="Fig" captionStartId="4.[174,205,1842,1861]" captionTargetBox="[177,1429,917,1805]" captionTargetId="figure-401@4.[174,1454,913,1815]" captionTargetPageId="4" captionText="Fig. 2. Epicepon indicum Nierstrasz and Brender à Brandis, 1931, female syntype, ZMUC CR-6788 (A–H) and male syntype, CR-6788 (I–J). A, dorsal view. B, left antennule (A1) and antenna (A2). C, right pereopod 1. D, right pereopod 7. E, left side of barbula. F, right oostegite 1, internal view. G, right maxilliped, external view with outer projection of barbula (OB). H, pleon, ventral view showing uropods (Ur), fifth lateral plates (Lp5), exopod (Ex5) and endopod (En5). I, dorsal view. J, lateral view. Scale bars: B = 25 µm, C, D = 50 µm, E = 100 µm, F, G = 200 µm. A, H–J, from Nierstrasz and Brender à Brandis (1931); not to scale." figureDoi="http://doi.org/10.5281/zenodo.12823477" httpUri="https://zenodo.org/record/12823477/files/figure.png" pageId="5" pageNumber="6">Fig. 2J</figureCitation>
). Pleomere 6 without uropods, posterolaterally extended into two slender rami, each as long as rest of pleotelson; minute anal cone present distomedially (
<figureCitation id="137B2A3BFFA1FF99FE024E94F2335F81" box="[488,563,1181,1206]" captionStart="Fig" captionStartId="4.[174,205,1842,1861]" captionTargetBox="[177,1429,917,1805]" captionTargetId="figure-401@4.[174,1454,913,1815]" captionTargetPageId="4" captionText="Fig. 2. Epicepon indicum Nierstrasz and Brender à Brandis, 1931, female syntype, ZMUC CR-6788 (A–H) and male syntype, CR-6788 (I–J). A, dorsal view. B, left antennule (A1) and antenna (A2). C, right pereopod 1. D, right pereopod 7. E, left side of barbula. F, right oostegite 1, internal view. G, right maxilliped, external view with outer projection of barbula (OB). H, pleon, ventral view showing uropods (Ur), fifth lateral plates (Lp5), exopod (Ex5) and endopod (En5). I, dorsal view. J, lateral view. Scale bars: B = 25 µm, C, D = 50 µm, E = 100 µm, F, G = 200 µm. A, H–J, from Nierstrasz and Brender à Brandis (1931); not to scale." figureDoi="http://doi.org/10.5281/zenodo.12823477" httpUri="https://zenodo.org/record/12823477/files/figure.png" pageId="5" pageNumber="6">Fig. 2I</figureCitation>
).
</paragraph>
<paragraph id="8BFF36BEFFA1FF99FF2F4EB7F2B45FC2" blockId="5.[136,738,253,1621]" pageId="5" pageNumber="6">
<emphasis id="B934EAACFFA1FF99FF2F4EB7F10B5FE2" box="[197,267,1214,1237]" italics="true" pageId="5" pageNumber="6">Range</emphasis>
: Known only from the
<typeStatus id="54FB881CFFA1FF99FDF74EB7F24D5FE1" box="[541,589,1214,1238]" pageId="5" pageNumber="6">type</typeStatus>
locality, Kei Islands,
<collectingCountry id="F357762EFFA1FF99FF0B4ED4F14B5FC2" box="[225,331,1245,1269]" name="Indonesia" pageId="5" pageNumber="6">Indonesia</collectingCountry>
,
<geoCoordinate id="EE745079FFA1FF99FEBC4ED4F1AB5FC2" box="[342,427,1245,1269]" degrees="05" direction="south" minutes="30" orientation="latitude" pageId="5" pageNumber="6" precision="925" value="-5.5">05°30'S</geoCoordinate>
,
<geoCoordinate id="EE745079FFA1FF99FE5D4ED4F21B5FC2" box="[439,539,1245,1269]" degrees="132" direction="east" minutes="51" orientation="longitude" pageId="5" pageNumber="6" precision="925" value="132.85">132°51'E</geoCoordinate>
,
<quantity id="4CB89B5BFFA1FF99FDCD4ED4F26A5FC2" box="[551,618,1245,1269]" metricMagnitude="2" metricUnit="m" metricValue="3.45" pageId="5" pageNumber="6" unit="m" value="345.0">345 m</quantity>
depth.
</paragraph>
<paragraph id="8BFF36BEFFA1FF99FF2F4EF4F17C5D62" blockId="5.[136,738,253,1621]" pageId="5" pageNumber="6">
<emphasis id="B934EAACFFA1FF99FF2F4EF4F1285E22" box="[197,296,1277,1301]" italics="true" pageId="5" pageNumber="6">Remarks</emphasis>
: The male
<typeStatus id="54FB881CFFA1FF99FE584EF7F20B5E21" box="[434,523,1278,1302]" pageId="5" pageNumber="6" type="syntype">syntype</typeStatus>
was incompletely described and figured by Nierstrasz &amp; Brender à Brandis (1931) but is no longer in the vial with the female
<typeStatus id="54FB881CFFA1FF99FF624F57F0E15E41" box="[136,225,1374,1398]" pageId="5" pageNumber="6" type="syntype">syntype</typeStatus>
and the above redescription of the male is therefore incomplete. The female
<typeStatus id="54FB881CFFA1FF99FE154F77F2535EA1" box="[511,595,1406,1430]" pageId="5" pageNumber="6" type="syntype">syntype</typeStatus>
superficially appears to be immature based on the linear shape of the body but the oostegites are well-developed. The specific identity of the host is unknown as it was not retained with the parasites and the branchial chamber from which the parasites were extracted is not clear as the female is nearly linear.
</paragraph>
<paragraph id="8BFF36BEFFA1FF99FECA4C74F2065DDE" blockId="5.[165,707,1661,1769]" pageId="5" pageNumber="6">
<emphasis id="B934EAACFFA1FF99FECA4C74F2495DA1" bold="true" box="[288,585,1661,1686]" pageId="5" pageNumber="6">
<taxonomicName id="4C404D3DFFA1FF99FECA4C74F1FC5DA1" authorityName="Barahona-Segovia &amp; Guzmán &amp; Pañinao-Monsálvez" authorityYear="2021" box="[288,508,1661,1686]" class="Malacostraca" family="Bopyridae" genus="Epicepon" kingdom="Animalia" order="Isopoda" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="belema" status="sp. nov.">
<emphasis id="B934EAACFFA1FF99FECA4C74F1FC5DA1" bold="true" box="[288,508,1661,1686]" italics="true" pageId="5" pageNumber="6">Epicepon belema</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20757D7FFA1FF99FDE94C74F2495DA1" box="[515,585,1661,1686]" pageId="5" pageNumber="6" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="137B2A3BFFA1FF99FE684C94F1CD5D82" box="[386,461,1693,1717]" captionStart="Fig" captionStartId="6.[174,205,1840,1859]" captionTargetBox="[174,1451,484,1812]" captionTargetId="figure-217@6.[174,1451,484,1812]" captionTargetPageId="6" captionText="Fig. 3. Epicepon belema, new species, female holotype, ZRC 2016.0424. A, dorsal view. B, right antennule (A1) and antenna (A2). C, left pereopod 1. D, left pereopod 7. E, left oostegite 1, external view. F, left oostegite 1, internal view. G, left maxilliped, external view (with outer lobe of barbula indicated with arrow). H, barbula (median lobes only). I, right side of pleon showing uropods (Ur), fourth and fifth lateral plates (Lp4, Lp5), fourth exopod (Ex4) and fourth endopod (En4). J, right pleomere 5 showing lateral plate (Lp), exopod (Ex) and endopod (En). Scale bars:A, E, F = 250 µm, B–D, J = 50 µm, G, I = 200 µm, H = 100 µm." figureDoi="http://doi.org/10.5281/zenodo.12823479" httpUri="https://zenodo.org/record/12823479/files/figure.png" pageId="5" pageNumber="6">Figs. 3</figureCitation>
,
<figureCitation id="137B2A3BFFA1FF99FE334C94F1E75D82" box="[473,487,1693,1717]" captionStart="Fig" captionStartId="7.[136,168,1896,1915]" captionTargetBox="[136,1413,435,1868]" captionTargetId="figure-5@7.[136,1413,383,1869]" captionTargetPageId="7" captionText="Fig. 4. Epicepon belema, new species, male allotype, ZRC 2016.0424. A, dorsal view. B, ventral view. C, left antennule (A1), antenna (A2), pereopod 1 and mouthparts. D, right pereopod 7 and pleomere 1 showing pleopod 1 (P1). E, pleomere 5 with pleopods (P5) and uropods (Ur). Scale bars: A, B = 250 µm, C–E = 50 µm." figureDoi="http://doi.org/10.5281/zenodo.12823481" httpUri="https://zenodo.org/record/12823481/files/figure.png" pageId="5" pageNumber="6">4</figureCitation>
)
<uri id="FFD13ABCFFA1FF99FF4F4CB1F2065DDE" pageId="5" pageNumber="6">
urn:lsid:zoobank.org:act:
<uuid id="FFE60C6BFFA1FF99FE784CB1F2065DDE" pageId="5" pageNumber="6">975C18A3-DE0C-4892-AEF0- D47ACEE92297</uuid>
</uri>
</paragraph>
<paragraph id="8BFF36BEFFA1FF99FF2F4D19F54F5A02" blockId="5.[136,737,1808,1961]" lastBlockId="5.[813,1414,253,1974]" pageId="5" pageNumber="6">
<emphasis id="B934EAACFFA1FF99FF2F4D19F1A05C1F" box="[197,416,1808,1832]" italics="true" pageId="5" pageNumber="6">Material examined</emphasis>
: Female
<typeStatus id="54FB881CFFA1FF99FDFA4D19F2745C1F" box="[528,628,1808,1832]" pageId="5" pageNumber="6" type="holotype">holotype</typeStatus>
(2.0 mm TL including uropods), male
<typeStatus id="54FB881CFFA1FF99FE094D39F23F5C7F" box="[483,575,1840,1864]" pageId="5" pageNumber="6" type="allotype">allotype</typeStatus>
(
<quantity id="4CB89B5BFFA1FF99FDB94D38F2AB5C7F" box="[595,683,1841,1865]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" pageId="5" pageNumber="6" unit="mm" value="1.1">1.1 mm</quantity>
TL) parasitizing right branchial chamber of
<taxonomicName id="4C404D3DFFA1FF99FD944D59F1DF5CBE" authority="Tan &amp; Huang, 2000" authorityName="Tan &amp; Huang" authorityYear="2000" class="Malacostraca" family="Cyclodorippidae" genus="Tymolus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="hirtipes">
<emphasis id="B934EAACFFA1FF99FD944D59F0E25CBF" italics="true" pageId="5" pageNumber="6">Tymolus hirtipes</emphasis>
<bibRefCitation id="EFD14B4FFFA1FF99FF074D78F1DF5CBE" author="Tan SH &amp; Huang J-F." box="[237,479,1905,1929]" pageId="5" pageNumber="6" pagination="135 - 140" refId="ref7046" refString="Tan SH, Huang J-F. 2000. Description of a new species of Tymolus (Crustacea: Decapoda: Brachyura: Cyclodorippidae) from Taiwan. Natl Mus Taiwan Spec Pub Ser 10: 135 - 140." type="journal article" year="2000">Tan &amp; Huang, 2000</bibRefCitation>
</taxonomicName>
, north of Panan, Pott (Phwoc)
<collectingCountry id="F357762EFFA1FF99FF1B4D99F1395C9F" box="[241,313,1936,1960]" name="Iceland" pageId="5" pageNumber="6">Island</collectingCountry>
, Belep Islands,
<collectingCountry id="F357762EFFA1FF99FE1E4D98F2A85C9F" box="[500,680,1936,1960]" name="New Caledonia" pageId="5" pageNumber="6">New Caledonia</collectingCountry>
, Sta. CP172,
<geoCoordinate id="EE745079FFA1FF99FC624AF7F4105A21" box="[904,1040,254,278]" degrees="19" direction="south" minutes="01" orientation="latitude" pageId="5" pageNumber="6" precision="1" seconds="1.2" value="-19.017">19°01'1.2&quot;S</geoCoordinate>
,
<geoCoordinate id="EE745079FFA1FF99FBF54AF4F4B65A22" box="[1055,1206,253,277]" degrees="163" direction="east" minutes="16" orientation="longitude" pageId="5" pageNumber="6" precision="1" seconds="1.2" value="163.267">163°16'1.2&quot;E</geoCoordinate>
,
<quantity id="4CB89B5BFFA1FF99FB2E4AF4F5475A22" box="[1220,1351,253,277]" metricMagnitude="2" metricUnit="m" metricValue="3.025" metricValueMax="3.3" metricValueMin="2.75" pageId="5" pageNumber="6" unit="m" value="302.5" valueMax="330.0" valueMin="275.0">275–330 m</quantity>
, coll. MUSORSTOM 4,
<date id="FFFE107EFFA1FF99FC1C4B17F4835A02" box="[1014,1155,285,310]" pageId="5" pageNumber="6" value="1985-09-17">17 Sept 1985</date>
(ZRC 2016.0424).
</paragraph>
<paragraph id="8BFF36BEFFA1FF99FC834B37F3EC5AE1" blockId="5.[813,1414,253,1974]" pageId="5" pageNumber="6">
<emphasis id="B934EAACFFA1FF99FC834B37F3F45A62" box="[873,1012,318,341]" italics="true" pageId="5" pageNumber="6">Description</emphasis>
: Female (
<figureCitation id="137B2A3BFFA1FF99FB9B4B34F4BA5A61" box="[1137,1210,317,342]" captionStart="Fig" captionStartId="6.[174,205,1840,1859]" captionTargetBox="[174,1451,484,1812]" captionTargetId="figure-217@6.[174,1451,484,1812]" captionTargetPageId="6" captionText="Fig. 3. Epicepon belema, new species, female holotype, ZRC 2016.0424. A, dorsal view. B, right antennule (A1) and antenna (A2). C, left pereopod 1. D, left pereopod 7. E, left oostegite 1, external view. F, left oostegite 1, internal view. G, left maxilliped, external view (with outer lobe of barbula indicated with arrow). H, barbula (median lobes only). I, right side of pleon showing uropods (Ur), fourth and fifth lateral plates (Lp4, Lp5), fourth exopod (Ex4) and fourth endopod (En4). J, right pleomere 5 showing lateral plate (Lp), exopod (Ex) and endopod (En). Scale bars:A, E, F = 250 µm, B–D, J = 50 µm, G, I = 200 µm, H = 100 µm." figureDoi="http://doi.org/10.5281/zenodo.12823479" httpUri="https://zenodo.org/record/12823479/files/figure.png" pageId="5" pageNumber="6">Fig. 3</figureCitation>
) length 2.0 mm; maximum width (across pereomere 3)
<quantity id="4CB89B5BFFA1FF99FB034B57F5435A42" box="[1257,1347,350,374]" metricMagnitude="-3" metricUnit="m" metricValue="1.4" pageId="5" pageNumber="6" unit="mm" value="1.4">1.4 mm</quantity>
; head length
<quantity id="4CB89B5BFFA1FF99FC684B77F3E05AA2" box="[898,992,382,406]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" pageId="5" pageNumber="6" unit="mm" value="0.4">0.4 mm</quantity>
; head width
<quantity id="4CB89B5BFFA1FF99FB6B4B77F4DF5AA2" box="[1153,1247,382,406]" metricMagnitude="-4" metricUnit="m" metricValue="7.0" pageId="5" pageNumber="6" unit="mm" value="0.7">0.7 mm</quantity>
; pleon length
<quantity id="4CB89B5BFFA1FF99FCC74B97F3875A82" box="[813,903,414,438]" metricMagnitude="-4" metricUnit="m" metricValue="7.0" pageId="5" pageNumber="6" unit="mm" value="0.7">0.7 mm</quantity>
. Body slightly dextral; all segments of body distinct (
<figureCitation id="137B2A3BFFA1FF99FC604BB4F3E05AE1" box="[906,992,445,470]" captionStart="Fig" captionStartId="6.[174,205,1840,1859]" captionTargetBox="[174,1451,484,1812]" captionTargetId="figure-217@6.[174,1451,484,1812]" captionTargetPageId="6" captionText="Fig. 3. Epicepon belema, new species, female holotype, ZRC 2016.0424. A, dorsal view. B, right antennule (A1) and antenna (A2). C, left pereopod 1. D, left pereopod 7. E, left oostegite 1, external view. F, left oostegite 1, internal view. G, left maxilliped, external view (with outer lobe of barbula indicated with arrow). H, barbula (median lobes only). I, right side of pleon showing uropods (Ur), fourth and fifth lateral plates (Lp4, Lp5), fourth exopod (Ex4) and fourth endopod (En4). J, right pleomere 5 showing lateral plate (Lp), exopod (Ex) and endopod (En). Scale bars:A, E, F = 250 µm, B–D, J = 50 µm, G, I = 200 µm, H = 100 µm." figureDoi="http://doi.org/10.5281/zenodo.12823479" httpUri="https://zenodo.org/record/12823479/files/figure.png" pageId="5" pageNumber="6">Fig. 3A</figureCitation>
).
</paragraph>
<paragraph id="8BFF36BEFFA1FF99FC834BD4F5025841" blockId="5.[813,1414,253,1974]" pageId="5" pageNumber="6">
Head oblong, wider than long, extending into rounded lateral projections, frontal margin entire medially. Frontal lamina thin and extending slightly beyond frontal margin of head (
<figureCitation id="137B2A3BFFA1FF99FB514834F51F5961" box="[1211,1311,573,598]" captionStart="Fig" captionStartId="6.[174,205,1840,1859]" captionTargetBox="[174,1451,484,1812]" captionTargetId="figure-217@6.[174,1451,484,1812]" captionTargetPageId="6" captionText="Fig. 3. Epicepon belema, new species, female holotype, ZRC 2016.0424. A, dorsal view. B, right antennule (A1) and antenna (A2). C, left pereopod 1. D, left pereopod 7. E, left oostegite 1, external view. F, left oostegite 1, internal view. G, left maxilliped, external view (with outer lobe of barbula indicated with arrow). H, barbula (median lobes only). I, right side of pleon showing uropods (Ur), fourth and fifth lateral plates (Lp4, Lp5), fourth exopod (Ex4) and fourth endopod (En4). J, right pleomere 5 showing lateral plate (Lp), exopod (Ex) and endopod (En). Scale bars:A, E, F = 250 µm, B–D, J = 50 µm, G, I = 200 µm, H = 100 µm." figureDoi="http://doi.org/10.5281/zenodo.12823479" httpUri="https://zenodo.org/record/12823479/files/figure.png" pageId="5" pageNumber="6">Fig. 3A</figureCitation>
). Small, irregular black eyes present mediolaterally (
<figureCitation id="137B2A3BFFA1FF99FAF44854F5785941" box="[1310,1400,605,630]" captionStart="Fig" captionStartId="6.[174,205,1840,1859]" captionTargetBox="[174,1451,484,1812]" captionTargetId="figure-217@6.[174,1451,484,1812]" captionTargetPageId="6" captionText="Fig. 3. Epicepon belema, new species, female holotype, ZRC 2016.0424. A, dorsal view. B, right antennule (A1) and antenna (A2). C, left pereopod 1. D, left pereopod 7. E, left oostegite 1, external view. F, left oostegite 1, internal view. G, left maxilliped, external view (with outer lobe of barbula indicated with arrow). H, barbula (median lobes only). I, right side of pleon showing uropods (Ur), fourth and fifth lateral plates (Lp4, Lp5), fourth exopod (Ex4) and fourth endopod (En4). J, right pleomere 5 showing lateral plate (Lp), exopod (Ex) and endopod (En). Scale bars:A, E, F = 250 µm, B–D, J = 50 µm, G, I = 200 µm, H = 100 µm." figureDoi="http://doi.org/10.5281/zenodo.12823479" httpUri="https://zenodo.org/record/12823479/files/figure.png" pageId="5" pageNumber="6">Fig. 3A</figureCitation>
). Antennules and antennae with two and three articles each, respectively, distalmost article of each terminally setose (
<figureCitation id="137B2A3BFFA1FF99FC6948B4F3DD59E1" box="[899,989,701,726]" captionStart="Fig" captionStartId="6.[174,205,1840,1859]" captionTargetBox="[174,1451,484,1812]" captionTargetId="figure-217@6.[174,1451,484,1812]" captionTargetPageId="6" captionText="Fig. 3. Epicepon belema, new species, female holotype, ZRC 2016.0424. A, dorsal view. B, right antennule (A1) and antenna (A2). C, left pereopod 1. D, left pereopod 7. E, left oostegite 1, external view. F, left oostegite 1, internal view. G, left maxilliped, external view (with outer lobe of barbula indicated with arrow). H, barbula (median lobes only). I, right side of pleon showing uropods (Ur), fourth and fifth lateral plates (Lp4, Lp5), fourth exopod (Ex4) and fourth endopod (En4). J, right pleomere 5 showing lateral plate (Lp), exopod (Ex) and endopod (En). Scale bars:A, E, F = 250 µm, B–D, J = 50 µm, G, I = 200 µm, H = 100 µm." figureDoi="http://doi.org/10.5281/zenodo.12823479" httpUri="https://zenodo.org/record/12823479/files/figure.png" pageId="5" pageNumber="6">Fig. 3B</figureCitation>
). Barbula with hook-shaped rounded lateral projection (
<figureCitation id="137B2A3BFFA1FF99FBE148D4F46859C1" box="[1035,1128,733,758]" captionStart="Fig" captionStartId="6.[174,205,1840,1859]" captionTargetBox="[174,1451,484,1812]" captionTargetId="figure-217@6.[174,1451,484,1812]" captionTargetPageId="6" captionText="Fig. 3. Epicepon belema, new species, female holotype, ZRC 2016.0424. A, dorsal view. B, right antennule (A1) and antenna (A2). C, left pereopod 1. D, left pereopod 7. E, left oostegite 1, external view. F, left oostegite 1, internal view. G, left maxilliped, external view (with outer lobe of barbula indicated with arrow). H, barbula (median lobes only). I, right side of pleon showing uropods (Ur), fourth and fifth lateral plates (Lp4, Lp5), fourth exopod (Ex4) and fourth endopod (En4). J, right pleomere 5 showing lateral plate (Lp), exopod (Ex) and endopod (En). Scale bars:A, E, F = 250 µm, B–D, J = 50 µm, G, I = 200 µm, H = 100 µm." figureDoi="http://doi.org/10.5281/zenodo.12823479" httpUri="https://zenodo.org/record/12823479/files/figure.png" pageId="5" pageNumber="6">Fig. 3G</figureCitation>
) and one rounded lobe, middle region smooth with low lateral lobe (
<figureCitation id="137B2A3BFFA1FF99FACA48F4F5795821" box="[1312,1401,765,790]" captionStart="Fig" captionStartId="6.[174,205,1840,1859]" captionTargetBox="[174,1451,484,1812]" captionTargetId="figure-217@6.[174,1451,484,1812]" captionTargetPageId="6" captionText="Fig. 3. Epicepon belema, new species, female holotype, ZRC 2016.0424. A, dorsal view. B, right antennule (A1) and antenna (A2). C, left pereopod 1. D, left pereopod 7. E, left oostegite 1, external view. F, left oostegite 1, internal view. G, left maxilliped, external view (with outer lobe of barbula indicated with arrow). H, barbula (median lobes only). I, right side of pleon showing uropods (Ur), fourth and fifth lateral plates (Lp4, Lp5), fourth exopod (Ex4) and fourth endopod (En4). J, right pleomere 5 showing lateral plate (Lp), exopod (Ex) and endopod (En). Scale bars:A, E, F = 250 µm, B–D, J = 50 µm, G, I = 200 µm, H = 100 µm." figureDoi="http://doi.org/10.5281/zenodo.12823479" httpUri="https://zenodo.org/record/12823479/files/figure.png" pageId="5" pageNumber="6">Fig. 3H</figureCitation>
). Maxilliped with prominent anterior segment, palp as low non-articulated projection with few distal setae, plectron triangular, short and thin (
<figureCitation id="137B2A3BFFA1FF99FB4A4954F4F65841" box="[1184,1270,861,886]" captionStart="Fig" captionStartId="6.[174,205,1840,1859]" captionTargetBox="[174,1451,484,1812]" captionTargetId="figure-217@6.[174,1451,484,1812]" captionTargetPageId="6" captionText="Fig. 3. Epicepon belema, new species, female holotype, ZRC 2016.0424. A, dorsal view. B, right antennule (A1) and antenna (A2). C, left pereopod 1. D, left pereopod 7. E, left oostegite 1, external view. F, left oostegite 1, internal view. G, left maxilliped, external view (with outer lobe of barbula indicated with arrow). H, barbula (median lobes only). I, right side of pleon showing uropods (Ur), fourth and fifth lateral plates (Lp4, Lp5), fourth exopod (Ex4) and fourth endopod (En4). J, right pleomere 5 showing lateral plate (Lp), exopod (Ex) and endopod (En). Scale bars:A, E, F = 250 µm, B–D, J = 50 µm, G, I = 200 µm, H = 100 µm." figureDoi="http://doi.org/10.5281/zenodo.12823479" httpUri="https://zenodo.org/record/12823479/files/figure.png" pageId="5" pageNumber="6">Fig. 3G</figureCitation>
).
</paragraph>
<paragraph id="8BFF36BEFFA1FF99FC834977F4675E01" blockId="5.[813,1414,253,1974]" pageId="5" pageNumber="6">
Pereon broadest across pereomere 3. Irregularly shaped tergal projections prominent on pereomeres 1–5 on right side and 2–4 on left side, kidney-shaped dorsolateral bosses on both sides of pereomeres 1–4, thin coxal plates on both sides of pereomeres 1–4 (
<figureCitation id="137B2A3BFFA1FF99FAB049F4F3525F01" captionStart="Fig" captionStartId="6.[174,205,1840,1859]" captionTargetBox="[174,1451,484,1812]" captionTargetId="figure-217@6.[174,1451,484,1812]" captionTargetPageId="6" captionText="Fig. 3. Epicepon belema, new species, female holotype, ZRC 2016.0424. A, dorsal view. B, right antennule (A1) and antenna (A2). C, left pereopod 1. D, left pereopod 7. E, left oostegite 1, external view. F, left oostegite 1, internal view. G, left maxilliped, external view (with outer lobe of barbula indicated with arrow). H, barbula (median lobes only). I, right side of pleon showing uropods (Ur), fourth and fifth lateral plates (Lp4, Lp5), fourth exopod (Ex4) and fourth endopod (En4). J, right pleomere 5 showing lateral plate (Lp), exopod (Ex) and endopod (En). Scale bars:A, E, F = 250 µm, B–D, J = 50 µm, G, I = 200 µm, H = 100 µm." figureDoi="http://doi.org/10.5281/zenodo.12823479" httpUri="https://zenodo.org/record/12823479/files/figure.png" pageId="5" pageNumber="6">Fig. 3A</figureCitation>
). No mid-dorsal projections on any pereomeres. Oostegites enclosing brood pouch; oostegite 1 (
<figureCitation id="137B2A3BFFA1FF99FAB24E34F3695F41" captionStart="Fig" captionStartId="6.[174,205,1840,1859]" captionTargetBox="[174,1451,484,1812]" captionTargetId="figure-217@6.[174,1451,484,1812]" captionTargetPageId="6" captionText="Fig. 3. Epicepon belema, new species, female holotype, ZRC 2016.0424. A, dorsal view. B, right antennule (A1) and antenna (A2). C, left pereopod 1. D, left pereopod 7. E, left oostegite 1, external view. F, left oostegite 1, internal view. G, left maxilliped, external view (with outer lobe of barbula indicated with arrow). H, barbula (median lobes only). I, right side of pleon showing uropods (Ur), fourth and fifth lateral plates (Lp4, Lp5), fourth exopod (Ex4) and fourth endopod (En4). J, right pleomere 5 showing lateral plate (Lp), exopod (Ex) and endopod (En). Scale bars:A, E, F = 250 µm, B–D, J = 50 µm, G, I = 200 µm, H = 100 µm." figureDoi="http://doi.org/10.5281/zenodo.12823479" httpUri="https://zenodo.org/record/12823479/files/figure.png" pageId="5" pageNumber="6">Fig. 3E, F</figureCitation>
) with ovate anterior article, subequal in length to posterior article, internal ridge smooth, posterior article rounded distally (smaller posterior lobe is curled and has slight damage in
<typeStatus id="54FB881CFFA1FF99FBF24EB4F47B5FE2" box="[1048,1147,1213,1237]" pageId="5" pageNumber="6" type="holotype">holotype</typeStatus>
). Pereopods subequal in structure and size (
<figureCitation id="137B2A3BFFA1FF99FC154ED4F4585FC1" box="[1023,1112,1245,1270]" captionStart="Fig" captionStartId="6.[174,205,1840,1859]" captionTargetBox="[174,1451,484,1812]" captionTargetId="figure-217@6.[174,1451,484,1812]" captionTargetPageId="6" captionText="Fig. 3. Epicepon belema, new species, female holotype, ZRC 2016.0424. A, dorsal view. B, right antennule (A1) and antenna (A2). C, left pereopod 1. D, left pereopod 7. E, left oostegite 1, external view. F, left oostegite 1, internal view. G, left maxilliped, external view (with outer lobe of barbula indicated with arrow). H, barbula (median lobes only). I, right side of pleon showing uropods (Ur), fourth and fifth lateral plates (Lp4, Lp5), fourth exopod (Ex4) and fourth endopod (En4). J, right pleomere 5 showing lateral plate (Lp), exopod (Ex) and endopod (En). Scale bars:A, E, F = 250 µm, B–D, J = 50 µm, G, I = 200 µm, H = 100 µm." figureDoi="http://doi.org/10.5281/zenodo.12823479" httpUri="https://zenodo.org/record/12823479/files/figure.png" pageId="5" pageNumber="6">Fig. 3A</figureCitation>
). All pereopods with short, blunt dactyli, cylindrical propodi, carpi and meri distinct in all pereopods (
<figureCitation id="137B2A3BFFA1FF99FC0C4F14F45A5E01" box="[998,1114,1309,1334]" captionStart="Fig" captionStartId="6.[174,205,1840,1859]" captionTargetBox="[174,1451,484,1812]" captionTargetId="figure-217@6.[174,1451,484,1812]" captionTargetPageId="6" captionText="Fig. 3. Epicepon belema, new species, female holotype, ZRC 2016.0424. A, dorsal view. B, right antennule (A1) and antenna (A2). C, left pereopod 1. D, left pereopod 7. E, left oostegite 1, external view. F, left oostegite 1, internal view. G, left maxilliped, external view (with outer lobe of barbula indicated with arrow). H, barbula (median lobes only). I, right side of pleon showing uropods (Ur), fourth and fifth lateral plates (Lp4, Lp5), fourth exopod (Ex4) and fourth endopod (En4). J, right pleomere 5 showing lateral plate (Lp), exopod (Ex) and endopod (En). Scale bars:A, E, F = 250 µm, B–D, J = 50 µm, G, I = 200 µm, H = 100 µm." figureDoi="http://doi.org/10.5281/zenodo.12823479" httpUri="https://zenodo.org/record/12823479/files/figure.png" pageId="5" pageNumber="6">Fig. 3C, D</figureCitation>
).
</paragraph>
<paragraph id="8BFF36BEFFA1FF99FC834F34F5465D41" blockId="5.[813,1414,253,1974]" pageId="5" pageNumber="6">
Pleon with six segments, first five pleomeres each bearing pair of biramous pleopods and longer uniramous lateral plates, all margins with slight undulation but not digitate, surfaces smooth (
<figureCitation id="137B2A3BFFA1FF99FB644F94F5055E81" box="[1166,1285,1437,1462]" captionStart="Fig" captionStartId="6.[174,205,1840,1859]" captionTargetBox="[174,1451,484,1812]" captionTargetId="figure-217@6.[174,1451,484,1812]" captionTargetPageId="6" captionText="Fig. 3. Epicepon belema, new species, female holotype, ZRC 2016.0424. A, dorsal view. B, right antennule (A1) and antenna (A2). C, left pereopod 1. D, left pereopod 7. E, left oostegite 1, external view. F, left oostegite 1, internal view. G, left maxilliped, external view (with outer lobe of barbula indicated with arrow). H, barbula (median lobes only). I, right side of pleon showing uropods (Ur), fourth and fifth lateral plates (Lp4, Lp5), fourth exopod (Ex4) and fourth endopod (En4). J, right pleomere 5 showing lateral plate (Lp), exopod (Ex) and endopod (En). Scale bars:A, E, F = 250 µm, B–D, J = 50 µm, G, I = 200 µm, H = 100 µm." figureDoi="http://doi.org/10.5281/zenodo.12823479" httpUri="https://zenodo.org/record/12823479/files/figure.png" pageId="5" pageNumber="6">Fig. 3A, I</figureCitation>
). Stub-like endopods of pleopods no more than half length of short, distally rounded exopods (
<figureCitation id="137B2A3BFFA1FF99FBA64FD4F4B15EC1" box="[1100,1201,1501,1526]" captionStart="Fig" captionStartId="6.[174,205,1840,1859]" captionTargetBox="[174,1451,484,1812]" captionTargetId="figure-217@6.[174,1451,484,1812]" captionTargetPageId="6" captionText="Fig. 3. Epicepon belema, new species, female holotype, ZRC 2016.0424. A, dorsal view. B, right antennule (A1) and antenna (A2). C, left pereopod 1. D, left pereopod 7. E, left oostegite 1, external view. F, left oostegite 1, internal view. G, left maxilliped, external view (with outer lobe of barbula indicated with arrow). H, barbula (median lobes only). I, right side of pleon showing uropods (Ur), fourth and fifth lateral plates (Lp4, Lp5), fourth exopod (Ex4) and fourth endopod (En4). J, right pleomere 5 showing lateral plate (Lp), exopod (Ex) and endopod (En). Scale bars:A, E, F = 250 µm, B–D, J = 50 µm, G, I = 200 µm, H = 100 µm." figureDoi="http://doi.org/10.5281/zenodo.12823479" httpUri="https://zenodo.org/record/12823479/files/figure.png" pageId="5" pageNumber="6">Fig. 3I, J</figureCitation>
); exopods less than ½ length of lateral plates (
<figureCitation id="137B2A3BFFA1FF99FBAF4FF4F4A85D21" box="[1093,1192,1533,1558]" captionStart="Fig" captionStartId="6.[174,205,1840,1859]" captionTargetBox="[174,1451,484,1812]" captionTargetId="figure-217@6.[174,1451,484,1812]" captionTargetPageId="6" captionText="Fig. 3. Epicepon belema, new species, female holotype, ZRC 2016.0424. A, dorsal view. B, right antennule (A1) and antenna (A2). C, left pereopod 1. D, left pereopod 7. E, left oostegite 1, external view. F, left oostegite 1, internal view. G, left maxilliped, external view (with outer lobe of barbula indicated with arrow). H, barbula (median lobes only). I, right side of pleon showing uropods (Ur), fourth and fifth lateral plates (Lp4, Lp5), fourth exopod (Ex4) and fourth endopod (En4). J, right pleomere 5 showing lateral plate (Lp), exopod (Ex) and endopod (En). Scale bars:A, E, F = 250 µm, B–D, J = 50 µm, G, I = 200 µm, H = 100 µm." figureDoi="http://doi.org/10.5281/zenodo.12823479" httpUri="https://zenodo.org/record/12823479/files/figure.png" pageId="5" pageNumber="6">Fig. 3I, J</figureCitation>
). Terminal pleomere ending in large uniramous uropods, similar to but longer than fifth pleopodal exopods, surfaces smooth, margins with slight undulation but not digitate (
<figureCitation id="137B2A3BFFA1FF99FB274C54F5385D41" box="[1229,1336,1629,1654]" captionStart="Fig" captionStartId="6.[174,205,1840,1859]" captionTargetBox="[174,1451,484,1812]" captionTargetId="figure-217@6.[174,1451,484,1812]" captionTargetPageId="6" captionText="Fig. 3. Epicepon belema, new species, female holotype, ZRC 2016.0424. A, dorsal view. B, right antennule (A1) and antenna (A2). C, left pereopod 1. D, left pereopod 7. E, left oostegite 1, external view. F, left oostegite 1, internal view. G, left maxilliped, external view (with outer lobe of barbula indicated with arrow). H, barbula (median lobes only). I, right side of pleon showing uropods (Ur), fourth and fifth lateral plates (Lp4, Lp5), fourth exopod (Ex4) and fourth endopod (En4). J, right pleomere 5 showing lateral plate (Lp), exopod (Ex) and endopod (En). Scale bars:A, E, F = 250 µm, B–D, J = 50 µm, G, I = 200 µm, H = 100 µm." figureDoi="http://doi.org/10.5281/zenodo.12823479" httpUri="https://zenodo.org/record/12823479/files/figure.png" pageId="5" pageNumber="6">Fig. 3A, I</figureCitation>
).
</paragraph>
<paragraph id="8BFF36BEFFA1FF99FC834C74F4565DC1" blockId="5.[813,1414,253,1974]" pageId="5" pageNumber="6">
Male (
<figureCitation id="137B2A3BFFA1FF99FC5F4C74F3FC5DA1" box="[949,1020,1661,1686]" captionStart="Fig" captionStartId="7.[136,168,1896,1915]" captionTargetBox="[136,1413,435,1868]" captionTargetId="figure-5@7.[136,1413,383,1869]" captionTargetPageId="7" captionText="Fig. 4. Epicepon belema, new species, male allotype, ZRC 2016.0424. A, dorsal view. B, ventral view. C, left antennule (A1), antenna (A2), pereopod 1 and mouthparts. D, right pereopod 7 and pleomere 1 showing pleopod 1 (P1). E, pleomere 5 with pleopods (P5) and uropods (Ur). Scale bars: A, B = 250 µm, C–E = 50 µm." figureDoi="http://doi.org/10.5281/zenodo.12823481" httpUri="https://zenodo.org/record/12823481/files/figure.png" pageId="5" pageNumber="6">Fig. 4</figureCitation>
) length
<quantity id="4CB89B5BFFA1FF99FB8A4C77F4BB5DA2" box="[1120,1211,1662,1686]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" pageId="5" pageNumber="6" unit="mm" value="1.1">1.1 mm</quantity>
; maximum width (across pereomeres 3 and 4)
<quantity id="4CB89B5BFFA1FF99FB6E4C97F4F05D82" box="[1156,1264,1694,1718]" metricMagnitude="-4" metricUnit="m" metricValue="3.2" pageId="5" pageNumber="6" unit="mm" value="0.32">0.32 mm</quantity>
; head length
<quantity id="4CB89B5BFFA1FF99FCC74CB7F3875DE2" box="[813,903,1726,1750]" metricMagnitude="-4" metricUnit="m" metricValue="2.0" pageId="5" pageNumber="6" unit="mm" value="0.2">0.2 mm</quantity>
; head width
<quantity id="4CB89B5BFFA1FF99FBF44CB4F4865DE2" box="[1054,1158,1725,1749]" metricMagnitude="-4" metricUnit="m" metricValue="2.5" pageId="5" pageNumber="6" unit="mm" value="0.25">0.25 mm</quantity>
. All body regions and segments distinct (
<figureCitation id="137B2A3BFFA1FF99FC1E4CD4F44A5DC1" box="[1012,1098,1757,1782]" captionStart="Fig" captionStartId="7.[136,168,1896,1915]" captionTargetBox="[136,1413,435,1868]" captionTargetId="figure-5@7.[136,1413,383,1869]" captionTargetPageId="7" captionText="Fig. 4. Epicepon belema, new species, male allotype, ZRC 2016.0424. A, dorsal view. B, ventral view. C, left antennule (A1), antenna (A2), pereopod 1 and mouthparts. D, right pereopod 7 and pleomere 1 showing pleopod 1 (P1). E, pleomere 5 with pleopods (P5) and uropods (Ur). Scale bars: A, B = 250 µm, C–E = 50 µm." figureDoi="http://doi.org/10.5281/zenodo.12823481" httpUri="https://zenodo.org/record/12823481/files/figure.png" pageId="5" pageNumber="6">Fig. 4A</figureCitation>
).
</paragraph>
<paragraph id="8BFF36BEFFA1FF99FC834CF4F4F45C81" blockId="5.[813,1414,253,1974]" pageId="5" pageNumber="6">
Head expanded distally, rounded posteriorly, distinctly separated from first pereomere (
<figureCitation id="137B2A3BFFA1FF99FAF04D14F5795C01" box="[1306,1401,1821,1846]" captionStart="Fig" captionStartId="7.[136,168,1896,1915]" captionTargetBox="[136,1413,435,1868]" captionTargetId="figure-5@7.[136,1413,383,1869]" captionTargetPageId="7" captionText="Fig. 4. Epicepon belema, new species, male allotype, ZRC 2016.0424. A, dorsal view. B, ventral view. C, left antennule (A1), antenna (A2), pereopod 1 and mouthparts. D, right pereopod 7 and pleomere 1 showing pleopod 1 (P1). E, pleomere 5 with pleopods (P5) and uropods (Ur). Scale bars: A, B = 250 µm, C–E = 50 µm." figureDoi="http://doi.org/10.5281/zenodo.12823481" httpUri="https://zenodo.org/record/12823481/files/figure.png" pageId="5" pageNumber="6">Fig. 4A</figureCitation>
). Large, irregular dark eyes near posterolateral margins of head. Antennules and antennae not visible beyond margins of head in dorsal view, of three articles each; both bearing setae on all articles (
<figureCitation id="137B2A3BFFA1FF99FB7E4D94F4E85C81" box="[1172,1256,1949,1974]" captionStart="Fig" captionStartId="7.[136,168,1896,1915]" captionTargetBox="[136,1413,435,1868]" captionTargetId="figure-5@7.[136,1413,383,1869]" captionTargetPageId="7" captionText="Fig. 4. Epicepon belema, new species, male allotype, ZRC 2016.0424. A, dorsal view. B, ventral view. C, left antennule (A1), antenna (A2), pereopod 1 and mouthparts. D, right pereopod 7 and pleomere 1 showing pleopod 1 (P1). E, pleomere 5 with pleopods (P5) and uropods (Ur). Scale bars: A, B = 250 µm, C–E = 50 µm." figureDoi="http://doi.org/10.5281/zenodo.12823481" httpUri="https://zenodo.org/record/12823481/files/figure.png" pageId="5" pageNumber="6">Fig. 4C</figureCitation>
).
</paragraph>
<paragraph id="8BFF36BEFFA2FF9AFF004AF7F5425A41" blockId="6.[174,775,253,438]" lastBlockId="6.[850,1451,253,438]" pageId="6" pageNumber="7">
Pereomeres 3 and 4 subequal in width, other pereomeres only slightly narrower except pereomere 7 which is much narrower (
<figureCitation id="137B2A3BFFA2FF9AFE004B34F2475A61" box="[490,583,317,342]" captionStart="Fig" captionStartId="7.[136,168,1896,1915]" captionTargetBox="[136,1413,435,1868]" captionTargetId="figure-5@7.[136,1413,383,1869]" captionTargetPageId="7" captionText="Fig. 4. Epicepon belema, new species, male allotype, ZRC 2016.0424. A, dorsal view. B, ventral view. C, left antennule (A1), antenna (A2), pereopod 1 and mouthparts. D, right pereopod 7 and pleomere 1 showing pleopod 1 (P1). E, pleomere 5 with pleopods (P5) and uropods (Ur). Scale bars: A, B = 250 µm, C–E = 50 µm." figureDoi="http://doi.org/10.5281/zenodo.12823481" httpUri="https://zenodo.org/record/12823481/files/figure.png" pageId="6" pageNumber="7">Fig. 4A</figureCitation>
). All pereomeres rounded laterally, lacking mid-ventral tubercles. First pereopod larger than second, both larger than other five (
<figureCitation id="137B2A3BFFA2FF9AFF0C4B94F17D5A81" box="[230,381,413,438]" captionStart="Fig" captionStartId="7.[136,168,1896,1915]" captionTargetBox="[136,1413,435,1868]" captionTargetId="figure-5@7.[136,1413,383,1869]" captionTargetPageId="7" captionText="Fig. 4. Epicepon belema, new species, male allotype, ZRC 2016.0424. A, dorsal view. B, ventral view. C, left antennule (A1), antenna (A2), pereopod 1 and mouthparts. D, right pereopod 7 and pleomere 1 showing pleopod 1 (P1). E, pleomere 5 with pleopods (P5) and uropods (Ur). Scale bars: A, B = 250 µm, C–E = 50 µm." figureDoi="http://doi.org/10.5281/zenodo.12823481" httpUri="https://zenodo.org/record/12823481/files/figure.png" pageId="6" pageNumber="7">Fig. 4B, C, D</figureCitation>
); all carpi with setae on distoventral ventral surface. Dactyli of pereopods 1 and 2 elongate, reaching distal margins of carpi (
<figureCitation id="137B2A3BFFA2FF9AFB504B14F52F5A01" box="[1210,1327,285,310]" captionStart="Fig" captionStartId="7.[136,168,1896,1915]" captionTargetBox="[136,1413,435,1868]" captionTargetId="figure-5@7.[136,1413,383,1869]" captionTargetPageId="7" captionText="Fig. 4. Epicepon belema, new species, male allotype, ZRC 2016.0424. A, dorsal view. B, ventral view. C, left antennule (A1), antenna (A2), pereopod 1 and mouthparts. D, right pereopod 7 and pleomere 1 showing pleopod 1 (P1). E, pleomere 5 with pleopods (P5) and uropods (Ur). Scale bars: A, B = 250 µm, C–E = 50 µm." figureDoi="http://doi.org/10.5281/zenodo.12823481" httpUri="https://zenodo.org/record/12823481/files/figure.png" pageId="6" pageNumber="7">Fig. 4B, C</figureCitation>
), dactyli of pereopods 3–7 short, inserted into cup-like indentations at distoventral margin of propodi (
<figureCitation id="137B2A3BFFA2FF9AFB2B4B54F5355A41" box="[1217,1333,349,374]" captionStart="Fig" captionStartId="7.[136,168,1896,1915]" captionTargetBox="[136,1413,435,1868]" captionTargetId="figure-5@7.[136,1413,383,1869]" captionTargetPageId="7" captionText="Fig. 4. Epicepon belema, new species, male allotype, ZRC 2016.0424. A, dorsal view. B, ventral view. C, left antennule (A1), antenna (A2), pereopod 1 and mouthparts. D, right pereopod 7 and pleomere 1 showing pleopod 1 (P1). E, pleomere 5 with pleopods (P5) and uropods (Ur). Scale bars: A, B = 250 µm, C–E = 50 µm." figureDoi="http://doi.org/10.5281/zenodo.12823481" httpUri="https://zenodo.org/record/12823481/files/figure.png" pageId="6" pageNumber="7">Fig. 4B, D</figureCitation>
).
</paragraph>
<paragraph id="8BFF36BEFFA2FF9BFC654B74F5515A21" blockId="6.[850,1451,253,438]" lastBlockId="7.[813,1414,253,373]" lastPageId="7" lastPageNumber="8" pageId="6" pageNumber="7">
Pleon with six pleomeres, rounded laterally, each slightly narrower than preceding one (
<figureCitation id="137B2A3BFFA2FF9AFAF14B94F59D5A81" box="[1307,1437,413,438]" captionStart="Fig" captionStartId="7.[136,168,1896,1915]" captionTargetBox="[136,1413,435,1868]" captionTargetId="figure-5@7.[136,1413,383,1869]" captionTargetPageId="7" captionText="Fig. 4. Epicepon belema, new species, male allotype, ZRC 2016.0424. A, dorsal view. B, ventral view. C, left antennule (A1), antenna (A2), pereopod 1 and mouthparts. D, right pereopod 7 and pleomere 1 showing pleopod 1 (P1). E, pleomere 5 with pleopods (P5) and uropods (Ur). Scale bars: A, B = 250 µm, C–E = 50 µm." figureDoi="http://doi.org/10.5281/zenodo.12823481" httpUri="https://zenodo.org/record/12823481/files/figure.png" pageId="6" pageNumber="7">Fig. 4A, B</figureCitation>
). Mid-ventral tubercles absent, distinct pleopods present laterally on pleomeres 1–5. Pleomere 6 without uropods, posterolaterally extended into two slender rami, each longer than rest of pleotelson, both tipped with short setae; anal cone present distomedially (
<figureCitation id="137B2A3BFFA3FF9BFB3A4AF4F5435A21" box="[1232,1347,253,278]" captionStart="Fig" captionStartId="7.[136,168,1896,1915]" captionTargetBox="[136,1413,435,1868]" captionTargetId="figure-5@7.[136,1413,383,1869]" captionTargetPageId="7" captionText="Fig. 4. Epicepon belema, new species, male allotype, ZRC 2016.0424. A, dorsal view. B, ventral view. C, left antennule (A1), antenna (A2), pereopod 1 and mouthparts. D, right pereopod 7 and pleomere 1 showing pleopod 1 (P1). E, pleomere 5 with pleopods (P5) and uropods (Ur). Scale bars: A, B = 250 µm, C–E = 50 µm." figureDoi="http://doi.org/10.5281/zenodo.12823481" httpUri="https://zenodo.org/record/12823481/files/figure.png" pageId="7" pageNumber="8">Fig. 4A, B</figureCitation>
).
</paragraph>
<caption id="DF3F6636FFA2FF9AFF444D39F2365C83" ID-DOI="http://doi.org/10.5281/zenodo.12823479" ID-Zenodo-Dep="12823479" httpUri="https://zenodo.org/record/12823479/files/figure.png" pageId="6" pageNumber="7" startId="6.[174,205,1840,1859]" targetBox="[174,1451,484,1812]" targetPageId="6" targetType="figure">
<paragraph id="8BFF36BEFFA2FF9AFF444D39F2365C83" blockId="6.[174,1452,1840,1972]" pageId="6" pageNumber="7">
<emphasis id="B934EAACFFA2FF9AFF444D39F0E65C74" bold="true" box="[174,230,1840,1859]" pageId="6" pageNumber="7">Fig. 3.</emphasis>
<taxonomicName id="4C404D3DFFA2FF9AFF184D38F1865C74" authorityName="Barahona-Segovia &amp; Guzmán &amp; Pañinao-Monsálvez" authorityYear="2021" box="[242,390,1840,1859]" class="Malacostraca" family="Bopyridae" genus="Epicepon" kingdom="Animalia" order="Isopoda" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="belema" status="sp. nov.">
<emphasis id="B934EAACFFA2FF9AFF184D38F1865C74" box="[242,390,1840,1859]" italics="true" pageId="6" pageNumber="7">Epicepon belema</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A20757D7FFA2FF9AFE7B4D38F1FA5C74" box="[401,506,1840,1860]" pageId="6" pageNumber="7" rank="species">new species</taxonomicNameLabel>
, female holotype, ZRC 2016.0424. A, dorsal view. B, right antennule (A1) and antenna (A2). C, left pereopod 1. D, left pereopod 7. E, left oostegite 1, external view. F, left oostegite 1, internal view. G, left maxilliped, external view (with outer lobe of barbula indicated with arrow). H, barbula (median lobes only). I, right side of pleon showing uropods (Ur), fourth and fifth lateral plates (Lp4, Lp5), fourth exopod (Ex4) and fourth endopod (En4). J, right pleomere 5 showing lateral plate (Lp), exopod (Ex) and endopod (En). Scale bars:A, E, F = 250 µm, B–D, J = 50 µm, G, I = 200 µm, H = 100 µm.
</paragraph>
</caption>
<paragraph id="8BFF36BEFFA3FF9BFC834B17F4265A42" blockId="7.[813,1414,253,373]" pageId="7" pageNumber="8">
<emphasis id="B934EAACFFA3FF9BFC834B17F3AE5A02" box="[873,942,286,309]" italics="true" pageId="7" pageNumber="8">Range</emphasis>
: Known only from the
<typeStatus id="54FB881CFFA3FF9BFB5C4B17F4E45A01" box="[1206,1252,286,310]" pageId="7" pageNumber="8">type</typeStatus>
locality, north of Panan, Pott (Phwoc)
<collectingCountry id="F357762EFFA3FF9BFBA34B34F4925A62" box="[1097,1170,317,341]" name="Iceland" pageId="7" pageNumber="8">Island</collectingCountry>
, Belep Islands,
<collectingCountry id="F357762EFFA3FF9BFABB4B37F39C5A42" name="New Caledonia" pageId="7" pageNumber="8">New Caledonia</collectingCountry>
,
<quantity id="4CB89B5BFFA3FF9BFC4C4B54F4225A42" box="[934,1058,349,373]" metricMagnitude="2" metricUnit="m" metricValue="3.025" metricValueMax="3.3" metricValueMin="2.75" pageId="7" pageNumber="8" unit="m" value="302.5" valueMax="330.0" valueMin="275.0">
275–
<specimenCount id="9D46FD37FFA3FF9BFC314B54F4225A42" box="[987,1058,349,373]" count="330" pageId="7" pageNumber="8" type="generic">330 m</specimenCount>
</quantity>
.
</paragraph>
<caption id="DF3F6636FFA3FF9BFF624D61F1BC5C84" ID-DOI="http://doi.org/10.5281/zenodo.12823481" ID-Zenodo-Dep="12823481" httpUri="https://zenodo.org/record/12823481/files/figure.png" pageId="7" pageNumber="8" startId="7.[136,168,1896,1915]" targetBox="[136,1413,435,1868]" targetPageId="7" targetType="figure">
<paragraph id="8BFF36BEFFA3FF9BFF624D61F1BC5C84" blockId="7.[136,1414,1896,1971]" pageId="7" pageNumber="8">
<emphasis id="B934EAACFFA3FF9BFF624D61F0C35C4C" bold="true" box="[136,195,1896,1915]" pageId="7" pageNumber="8">Fig. 4.</emphasis>
<taxonomicName id="4C404D3DFFA3FF9BFF394D60F16C5C4C" authorityName="Barahona-Segovia &amp; Guzmán &amp; Pañinao-Monsálvez" authorityYear="2021" box="[211,364,1896,1915]" class="Malacostraca" family="Bopyridae" genus="Epicepon" kingdom="Animalia" order="Isopoda" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="belema" status="sp. nov.">
<emphasis id="B934EAACFFA3FF9BFF394D60F16C5C4C" box="[211,364,1896,1915]" italics="true" pageId="7" pageNumber="8">Epicepon belema</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A20757D7FFA3FF9BFE934D61F1E75C4C" box="[377,487,1896,1915]" pageId="7" pageNumber="8" rank="species">new species</taxonomicNameLabel>
, male allotype, ZRC 2016.0424. A, dorsal view. B, ventral view. C, left antennule (A1), antenna (A2), pereopod 1 and mouthparts. D, right pereopod 7 and pleomere 1 showing pleopod 1 (P1). E, pleomere 5 with pleopods (P5) and uropods (Ur). Scale bars: A, B = 250 µm, C–E = 50 µm.
</paragraph>
</caption>
<paragraph id="8BFF36BEFFACFF94FF004AF4F0F659A2" blockId="8.[174,775,253,821]" pageId="8" pageNumber="9">
<emphasis id="B934EAACFFACFF94FF004AF4F1495A22" box="[234,329,253,277]" italics="true" pageId="8" pageNumber="9">Remarks</emphasis>
: The female of
<taxonomicName id="4C404D3DFFACFF94FDE84AF7F2C15A22" authorityName="Barahona-Segovia &amp; Guzmán &amp; Pañinao-Monsálvez" authorityYear="2021" box="[514,705,253,277]" class="Malacostraca" family="Bopyridae" genus="Epicepon" kingdom="Animalia" order="Isopoda" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="belema" status="sp. nov.">
<emphasis id="B934EAACFFACFF94FDE84AF7F2C15A22" box="[514,705,253,277]" italics="true" pageId="8" pageNumber="9">Epicepon belema</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20757D7FFACFF94FD204AF7F3075A22" box="[714,775,254,277]" pageId="8" pageNumber="9" rank="species">n. sp.</taxonomicNameLabel>
can be distinguished from those of the other two in the genus in having a total length less than 2 times the maximum width (
<taxonomicName id="4C404D3DFFACFF94FE9C4B57F1EF5A42" box="[374,495,349,373]" class="Malacostraca" family="Bopyridae" genus="Epicepon" kingdom="Animalia" order="Isopoda" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="indicum">
<emphasis id="B934EAACFFACFF94FE9C4B57F1865A42" box="[374,390,350,373]" italics="true" pageId="8" pageNumber="9">E</emphasis>
.
<emphasis id="B934EAACFFACFF94FE7D4B54F1EF5A42" box="[407,495,349,373]" italics="true" pageId="8" pageNumber="9">indicum</emphasis>
</taxonomicName>
with total length greater than 2 times the maximum width) and pleopodal endopods greater than 1/4 length of the exopods, exopods less than 1/2 length of lateral plates, and barbula without slender elongated lobes (
<taxonomicName id="4C404D3DFFACFF94FD6F4BD7F3075AC2" box="[645,775,478,501]" class="Malacostraca" family="Bopyridae" genus="Epicepon" kingdom="Animalia" order="Isopoda" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="japonica">
<emphasis id="B934EAACFFACFF94FD6F4BD7F2955AC2" box="[645,661,478,501]" italics="true" pageId="8" pageNumber="9">E</emphasis>
.
<emphasis id="B934EAACFFACFF94FD4C4BD7F3075AC2" box="[678,775,478,501]" italics="true" pageId="8" pageNumber="9">japonica</emphasis>
</taxonomicName>
with pleopodal endopods less than 1/4 length of the exopods, exopods greater than 1/2 length of lateral plates; barbula of
<taxonomicName id="4C404D3DFFACFF94FE994837F1E85962" box="[371,488,573,597]" class="Malacostraca" family="Bopyridae" genus="Epicepon" kingdom="Animalia" order="Isopoda" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="indicum">
<emphasis id="B934EAACFFACFF94FE994837F1835962" box="[371,387,574,597]" italics="true" pageId="8" pageNumber="9">E</emphasis>
.
<emphasis id="B934EAACFFACFF94FE784834F1E85962" box="[402,488,573,597]" italics="true" pageId="8" pageNumber="9">indicum</emphasis>
</taxonomicName>
with one slender elongate lobe, barbula of
<taxonomicName id="4C404D3DFFACFF94FE8A4857F1DC5942" box="[352,476,606,629]" class="Malacostraca" family="Bopyridae" genus="Epicepon" kingdom="Animalia" order="Isopoda" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="japonica">
<emphasis id="B934EAACFFACFF94FE8A4857F1705942" box="[352,368,606,629]" italics="true" pageId="8" pageNumber="9">E</emphasis>
.
<emphasis id="B934EAACFFACFF94FE954857F1DC5942" box="[383,476,606,629]" italics="true" pageId="8" pageNumber="9">japonica</emphasis>
</taxonomicName>
with two slender elongated lobes).
</paragraph>
<paragraph id="8BFF36BEFFACFF94FF004894F17C5802" blockId="8.[174,775,253,821]" pageId="8" pageNumber="9">
<emphasis id="B934EAACFFACFF94FF004894F16B5982" box="[234,363,669,693]" italics="true" pageId="8" pageNumber="9">Etymology</emphasis>
: The specific name
<taxonomicName id="4C404D3DFFACFF94FD9E4897F2D95982" authorityName="Barahona-Segovia &amp; Guzmán &amp; Pañinao-Monsálvez" authorityYear="2021" box="[628,729,669,694]" class="Malacostraca" family="Bopyridae" genus="Epicepon" kingdom="Animalia" order="Isopoda" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="belema">
“
<emphasis id="B934EAACFFACFF94FD684894F2D95982" box="[642,729,669,693]" italics="true" pageId="8" pageNumber="9">belema</emphasis>
</taxonomicName>
” is both the name inhabitants of the Belep islands call themselves as well as the adjective used for things related to the Belep Islands; the name is used here as a noun in apposition.
</paragraph>
</subSubSection>
</treatment>
</document>
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