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<document id="F8D5354182776661095D4EF18060D33C" ID-DOI="10.5281/zenodo.5358561" ID-ISSN="2345-7600" ID-Zenodo-Dep="5358561" ID-ZooBank="23CA96C3-FA71-4ECA-BF05-A78EA94CAD73" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630224835951" checkinUser="valdenar" docAuthor="Sidabalok, Conni M. &amp; Bruce, Niel L." docDate="2018" docId="03E98795FF9E1801FEF9F80432A7FA1D" docLanguage="en" docName="66rbz177-207.pdf" docOrigin="Raffles Bulletin of Zoology 66" docStyle="DocumentStyle:F2D34FF25DBF9998BB8DF6F24A5B097A.5:RafflesBZool.2014-2018.journal_article" docStyleId="F2D34FF25DBF9998BB8DF6F24A5B097A" docStyleName="RafflesBZool.2014-2018.journal_article" docStyleVersion="5" docTitle="Cirolana trulla Sidabalok &amp; Bruce 2018, new species" docType="treatment" docUuid="C184CEB7-9F0E-4390-A044-6DC665D31156" docUuidSource="ZooBank" docVersion="4" lastPageNumber="191" masterDocId="FFD0FFEDFF97180FFFEAFF993623FF88" masterDocTitle="Review of the Cirolana ‘ pleonastica-group’ (Crustacea: Isopoda: Cirolanidae) with description of four new species from the Indo- Malaysian region" masterLastPageNumber="207" masterPageNumber="177" pageNumber="186" updateTime="1699191830796" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
<mods:mods id="5FDA8155328538E0F3518D8620AA015E" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="F00CD071E2B88201F78A8815BD79D7B9">
<mods:title id="C6B1EEC2FF5E0CDE0E644AC500320E0A">Review of the Cirolana ‘ pleonastica-group’ (Crustacea: Isopoda: Cirolanidae) with description of four new species from the Indo- Malaysian region</mods:title>
</mods:titleInfo>
<mods:name id="0182B5BCEC6ADF8035862AF83596CD6C" type="personal">
<mods:role id="8D6AD4699CB825893266098F045E3AA1">
<mods:roleTerm id="20F8C0363640B80CA3A68B9E665D7F63">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="A4FEB833CB2DAAC71150A0A622CABD13">Sidabalok, Conni M.</mods:namePart>
</mods:name>
<mods:name id="632650FD673F4D4FD551BDA573AF7D67" type="personal">
<mods:role id="CF5BD3B18FE49CE2C6D00C84138A6805">
<mods:roleTerm id="A65FD6B5DCE539EE8C346FCBABA53E22">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="68F2DB357EB0927E5900A2987EBF5F8A">Bruce, Niel L.</mods:namePart>
</mods:name>
<mods:typeOfResource id="F04B66875E7B1929A5C9ED3336F666B6">text</mods:typeOfResource>
<mods:relatedItem id="F5E002F703DF0E2421FADED5F11BBBCB" type="host">
<mods:titleInfo id="EDB3CF9C92A1969DB82E3D6DFADA58AF">
<mods:title id="7FA2CB3302BF9C7BFF3DE47B3C5D5A89">Raffles Bulletin of Zoology</mods:title>
</mods:titleInfo>
<mods:part id="4168D22B2C0B372DDB5B9C4AF15B675C">
<mods:date id="10615C575DD62C2263E40B2A28B8608E">2018</mods:date>
<mods:detail id="8F3162FD02796C105BA0C26A7D1E5590" type="pubDate">
<mods:number id="D8D57C283DCE502EAAB311359C3E2BF7">2018-03-20</mods:number>
</mods:detail>
<mods:detail id="6FDC247C8CD4C0FE5279B7957D9FFCBC" type="volume">
<mods:number id="CDB464F52C165E4C44FACC3B9D2BB86A">66</mods:number>
</mods:detail>
<mods:extent id="BB577E457191368562C227D428DB3E7B" unit="page">
<mods:start id="C9BE07CA56359986A3477103D435B62C">177</mods:start>
<mods:end id="D5E6FC3E22381982152B7450EF7CA177">207</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="815582A78ED4DAAFF4F3CB9D73C8D51A">journal article</mods:classification>
<mods:identifier id="ED63DE2CB72DE8BF9215F8F6F0683AB8" type="DOI">10.5281/zenodo.5358561</mods:identifier>
<mods:identifier id="D4B2D3BFE17967701B7700C8A937048F" type="ISSN">2345-7600</mods:identifier>
<mods:identifier id="539F00D880A219ACB5AF3FDBC0D2C453" type="Zenodo-Dep">5358561</mods:identifier>
<mods:identifier id="4DA212B1CDF9E8FA6EA7B197FCDEA91D" type="ZooBank">23CA96C3-FA71-4ECA-BF05-A78EA94CAD73</mods:identifier>
</mods:mods>
<treatment id="03E98795FF9E1801FEF9F80432A7FA1D" LSID="urn:lsid:zoobank.org:act:C184CEB7-9F0E-4390-A044-6DC665D31156" httpUri="http://treatment.plazi.org/id/03E98795FF9E1801FEF9F80432A7FA1D" lastPageId="14" lastPageNumber="191" pageId="9" pageNumber="186">
<subSubSection id="C35A6508FF9E1806FEF9F804346CF83D" box="[275,591,1949,1973]" pageId="9" pageNumber="186" type="nomenclature">
<paragraph id="8BFF3683FF9E1806FEF9F804346CF83D" blockId="9.[275,591,1949,2005]" box="[275,591,1949,1973]" pageId="9" pageNumber="186">
<heading id="D0B781EFFF9E1806FEF9F804346CF83D" bold="true" box="[275,591,1949,1973]" centered="true" fontSize="10" level="2" pageId="9" pageNumber="186" reason="2">
<emphasis id="B934EA91FF9E1806FEF9F804346CF83D" bold="true" box="[275,591,1949,1973]" pageId="9" pageNumber="186">
<taxonomicName id="4C404D00FF9E1806FEF9F8043799F83D" ID-CoL="VGGL" authority="Sidabalok &amp; Bruce, 2018" authorityName="Sidabalok &amp; Bruce" authorityYear="2018" box="[275,442,1949,1973]" class="Malacostraca" family="Cirolanidae" genus="Cirolana" kingdom="Animalia" order="Isopoda" pageId="9" pageNumber="186" phylum="Arthropoda" rank="species" species="trulla" status="sp. nov.">
<emphasis id="B934EA91FF9E1806FEF9F8043799F83D" bold="true" box="[275,442,1949,1973]" italics="true" pageId="9" pageNumber="186">Cirolana trulla</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A20757EAFF9E1806FE23F804346CF83D" box="[457,591,1949,1973]" pageId="9" pageNumber="186" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C35A6508FF9E1806FE9FF82437CFF85D" box="[373,492,1981,2005]" pageId="9" pageNumber="186" type="description">
<paragraph id="8BFF3683FF9E1806FE9FF82437CFF85D" blockId="9.[275,591,1949,2005]" box="[373,492,1981,2005]" pageId="9" pageNumber="186">
(
<figureCitation id="137B2A06FF9E1806FE97F82437C7F85D" box="[381,484,1981,2005]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStart-3="Fig" captionStartId-0="10.[135,170,2035,2057]" captionStartId-1="11.[113,148,1417,1439]" captionStartId-2="12.[135,170,2064,2086]" captionStartId-3="13.[113,148,1933,1955]" captionTargetBox-0="[169,1442,191,2020]" captionTargetBox-1="[114,1451,192,1399]" captionTargetBox-2="[168,1442,189,2047]" captionTargetBox-3="[114,1451,320,1917]" captionTargetId-0="figure-8@10.[121,1442,187,2021]" captionTargetId-1="figure-10@11.[114,1451,191,1400]" captionTargetId-2="figure-8@12.[168,1442,189,2047]" captionTargetId-3="figure-10@13.[114,1451,320,1918]" captionTargetPageId-0="10" captionTargetPageId-1="11" captionTargetPageId-2="12" captionTargetPageId-3="13" captionText-0="Fig. 5. Cirolana trulla n. sp. A–C, E–F, holotype male (7.4 mm) (ZRC 2018.0206), Singapore. D, G, H, paratype male (6.0 mm) (ZRC 2018.0207), Singapore. I, paratype female (6.3 mm) (ZRC 2018.0207), Singapore. A, dorsal view; B, lateral view; C, head; D, antennula; E, frons; F, pleon; G, antenna peduncle; H, uropod male; I, uropod female." captionText-1="Fig. 6. Cirolana trulla n. sp. E, holotype male (7.4 mm) (ZRC 2018.0206), Singapore. A–D, F, paratype male (6.0 mm) (ZRC 2018.0207), Singapore. A, maxilliped; B, maxilla; C, maxillule; D, right mandible; E, penial process; F, pleotelson." captionText-2="Fig. 7. Cirolana trulla n. sp. A–D, paratype male (6.0 mm) (ZRC 2018.0207), Singapore. A, pereopod 1; B, pereopod 2; C, pereopod 6; D, pereopod 7." captionText-3="Fig. 8. Cirolana trulla n. sp. A–E, paratype male (6.0 mm) (ZRC 2018.0207), Singapore. A, pleopod 1; B, pleopod 2; C, pleopod 3; D, pleopod 4; E, pleopod 5." figureDoi-0="http://doi.org/10.5281/zenodo.5358571" figureDoi-1="http://doi.org/10.5281/zenodo.5358573" figureDoi-2="http://doi.org/10.5281/zenodo.5358575" figureDoi-3="http://doi.org/10.5281/zenodo.5358577" httpUri-0="https://zenodo.org/record/5358571/files/figure.png" httpUri-1="https://zenodo.org/record/5358573/files/figure.png" httpUri-2="https://zenodo.org/record/5358575/files/figure.png" httpUri-3="https://zenodo.org/record/5358577/files/figure.png" pageId="9" pageNumber="186">Figs. 5–8</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C35A6508FF9E1806FF9BF8643214FDBD" pageId="9" pageNumber="186" type="materials_examined">
<paragraph id="8BFF3683FF9E1806FF9BF86434CDF7BD" blockId="9.[112,752,2045,2101]" pageId="9" pageNumber="186">
<emphasis id="B934EA91FF9E1806FF9BF8643768F79D" bold="true" box="[113,331,2045,2069]" pageId="9" pageNumber="186">Material examined.</emphasis>
<materialsCitation id="3B283CDEFF9E1806FEA7F86434CDF7BD" collectionCode="ZRC" country="Singapore" location="St John's Island" pageId="9" pageNumber="186" specimenCode="ZRC 2018.0206" specimenCount="1" typeStatus="holotype">
All
<collectingCountry id="F3577613FF9E1806FE9EF86437C1F79D" box="[372,482,2045,2069]" name="Singapore" pageId="9" pageNumber="186">Singapore</collectingCountry>
.
<typeStatus id="54FB8821FF9E1806FE03F864346EF79D" box="[489,589,2045,2069]" pageId="9" pageNumber="186" type="holotype">Holotype</typeStatus>
, male (
<quantity id="4CB89B66FF9E1806FD7DF86434CAF79D" box="[663,745,2045,2069]" metricMagnitude="-3" metricUnit="m" metricValue="7.4" pageId="9" pageNumber="186" unit="mm" value="7.4">7.4 mm</quantity>
) (
<specimenCode id="DBE69EF8FF9E1806FF9DF784370CF7BD" box="[119,303,2077,2101]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="9" pageNumber="186">ZRC 2018.0206</specimenCode>
), between
<location id="8E9F6058FF9E1806FE43F784347EF7BD" LSID="urn:lsid:plazi:treatment:03E98795FF9E1801FEF9F80432A7FA1D:8E9F6058FF9E1806FE43F784347EF7BD" box="[425,605,2077,2101]" country="Singapore" name="St John's Island" pageId="9" pageNumber="186">St John’s Island</location>
and Lazarus
</materialsCitation>
</paragraph>
<paragraph id="8BFF3683FF9E1806FCC7FF243316FF7D" blockId="9.[813,1451,188,246]" pageId="9" pageNumber="186">
Island,
<geoCoordinate id="EE745044FF9E1806FC93FF253220FF5D" box="[889,1027,188,213]" degrees="01" direction="north" minutes="13.043" orientation="latitude" pageId="9" pageNumber="186" precision="1" value="1.2173834">01°13.043′N</geoCoordinate>
,
<geoCoordinate id="EE745044FF9E1806FBE6FF253283FF5D" box="[1036,1184,188,213]" degrees="103" direction="east" minutes="51.319" orientation="longitude" pageId="9" pageNumber="186" precision="1" value="103.855316">103°51.319′E</geoCoordinate>
, OTC0103 OTR356, trap
<quantity id="4CB89B66FF9E1806FCC7FF443558FF7D" box="[813,891,221,245]" metricMagnitude="1" metricUnit="m" metricValue="2.2399999999999998" pageId="9" pageNumber="186" unit="m" value="22.4">22.4 m</quantity>
, coll. H. Wong and party,
<date id="FFFE1043FF9E1806FB4BFF443312FF7D" box="[1185,1329,221,245]" pageId="9" pageNumber="186" value="2014-04-03">3 April 2014</date>
.
</paragraph>
<paragraph id="8BFF3683FF9E1806FCC7FE843214FDBD" blockId="9.[812,1454,285,566]" pageId="9" pageNumber="186">
<emphasis id="B934EA91FF9E1806FCC7FE843585FEBD" bold="true" box="[813,934,285,309]" pageId="9" pageNumber="186">
<typeStatus id="54FB8821FF9E1806FCC7FE843581FEBD" box="[813,930,285,309]" pageId="9" pageNumber="186" type="paratype">Paratypes</typeStatus>
.
</emphasis>
<specimenCount id="9D46FD0AFF9E1806FC5AFE843224FEBD" box="[944,1031,285,309]" pageId="9" pageNumber="186" type="male">4 males</specimenCount>
(6.0 [dissected], 6.3, 6.6,
<quantity id="4CB89B66FF9E1806FAC5FE8433AAFEBD" box="[1327,1417,285,309]" metricMagnitude="-3" metricUnit="m" metricValue="6.9" pageId="9" pageNumber="186" unit="mm" value="6.9">6.9 mm</quantity>
),
<materialsCitation id="3B283CDEFF9E1806FA74FE8435AEFEFD" collectionCode="ZRC" country="Singapore" location="St John's Island" pageId="9" pageNumber="186" specimenCode="ZRC 2018.0207" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<specimenCount id="9D46FD0AFF9E1806FA74FE843555FEDD" pageId="9" pageNumber="186" type="female">1 female</specimenCount>
(6.3 [dissected] mm) (
<specimenCode id="DBE69EF8FF9E1806FB8EFEA43336FEDD" box="[1124,1301,317,341]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="9" pageNumber="186">ZRC 2018.0207</specimenCode>
), same data as holotype
</materialsCitation>
;
<materialsCitation id="3B283CDEFF9E1806FC73FEC43207FE5D" collectingDate="2013-10-07" collectionCode="ZRC" collectorName="S. Cheng" country="Singapore" elevation="9" latitude="1.4403833" location="Pulau Ubin" longLatPrecision="1" longitude="104.04504" municipality="Beting Bronok" pageId="9" pageNumber="186" specimenCode="ZRC 2018.0208" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<specimenCount id="9D46FD0AFF9E1806FC73FEC435C0FEFD" box="[921,995,349,373]" pageId="9" pageNumber="186" type="male">1 male</specimenCount>
(
<quantity id="4CB89B66FF9E1806FC1EFEC43269FEFD" box="[1012,1098,349,373]" metricMagnitude="-3" metricUnit="m" metricValue="7.8" pageId="9" pageNumber="186" unit="mm" value="7.8">7.8 mm</quantity>
) (
<specimenCode id="DBE69EF8FF9E1806FB8BFEC43335FEFD" box="[1121,1302,349,373]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="9" pageNumber="186">ZRC 2018.0208</specimenCode>
),
<location id="8E9F6058FF9E1806FAC3FEC43384FEFD" LSID="urn:lsid:plazi:treatment:03E98795FF9E1801FEF9F80432A7FA1D:8E9F6058FF9E1806FAC3FEC43384FEFD" box="[1321,1447,349,373]" country="Singapore" latitude="1.4403833" longLatPrecision="1" longitude="104.04504" municipality="Beting Bronok" name="Pulau Ubin" pageId="9" pageNumber="186">Pulau Ubin</location>
,
<collectingMunicipality id="6B9BACF9FF9E1806FCC5FEE435F6FE1D" box="[815,981,381,405]" pageId="9" pageNumber="186">Beting Bronok</collectingMunicipality>
,
<geoCoordinate id="EE745044FF9E1806FC0EFEE53252FE1D" box="[996,1137,380,405]" degrees="01" direction="north" minutes="26.423" orientation="latitude" pageId="9" pageNumber="186" precision="1" value="1.4403833">01°26.423′N</geoCoordinate>
,
<geoCoordinate id="EE745044FF9E1806FB6AFEE53334FE1D" box="[1152,1303,380,405]" degrees="104" direction="east" minutes="02.702" orientation="longitude" pageId="9" pageNumber="186" precision="1" value="104.04504">104°02.702′E</geoCoordinate>
, CMBS SEA 1273 DR214, rectangular dredge
<quantity id="4CB89B66FF9E1806FB7DFE0432F4FE3D" box="[1175,1239,413,437]" metricMagnitude="0" metricUnit="m" metricValue="8.7" pageId="9" pageNumber="186" unit="m" value="8.7">
<elevation id="006DD1B0FF9E1806FB7DFE0432F4FE3D" box="[1175,1239,413,437]" metricMagnitude="0" metricUnit="m" metricValue="8.7" pageId="9" pageNumber="186" unit="m" value="8.7">8.7 m</elevation>
</quantity>
, coll.
<collectorName id="26B55355FF9E1806FAF0FE04335DFE3D" box="[1306,1406,413,437]" pageId="9" pageNumber="186">S. Cheng</collectorName>
and party,
<date id="FFFE1043FF9E1806FC9FFE273207FE5D" box="[885,1060,445,469]" pageId="9" pageNumber="186" value="2013-10-07">
<collectingDate id="EFBAE9ABFF9E1806FC9FFE273207FE5D" box="[885,1060,445,469]" pageId="9" pageNumber="186" value="2013-10-07">7 October 2013</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="3B283CDEFF9E1806FBDBFE243210FDBD" accessionNumber="W34884" collectingDate="2014-04-08" collectionCode="MTQ" collectorName="T. Tay" country="Singapore" elevation="19" latitude="1.3082" location="Changi Naval Base" longLatPrecision="1" longitude="104.04892" pageId="9" pageNumber="186" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<specimenCount id="9D46FD0AFF9E1806FBDBFE243258FE5D" box="[1073,1147,445,469]" pageId="9" pageNumber="186" type="male">1 male</specimenCount>
(
<quantity id="4CB89B66FF9E1806FB66FE2432C0FE5D" box="[1164,1251,445,469]" metricMagnitude="-3" metricUnit="m" metricValue="5.2" pageId="9" pageNumber="186" unit="mm" value="5.2">5.2 mm</quantity>
) (
<collectionCode id="ED51AE46FF9E1806FB11FE24331BFE5D" box="[1275,1336,445,469]" pageId="9" pageNumber="186">MTQ</collectionCode>
W34884), near
<location id="8E9F6058FF9E1806FCB5FE443208FE7D" LSID="urn:lsid:plazi:treatment:03E98795FF9E1801FEF9F80432A7FA1D:8E9F6058FF9E1806FCB5FE443208FE7D" box="[863,1067,477,501]" country="Singapore" latitude="1.3082" longLatPrecision="1" longitude="104.04892" name="Changi Naval Base" pageId="9" pageNumber="186">Changi Naval Base</location>
,
<geoCoordinate id="EE745044FF9E1806FBDEFE45329FFE7D" box="[1076,1212,476,501]" degrees="01" direction="north" minutes="18.492" orientation="latitude" pageId="9" pageNumber="186" precision="1" value="1.3082">01°18.492′N</geoCoordinate>
,
<geoCoordinate id="EE745044FF9E1806FB2CFE45337AFE7D" box="[1222,1369,476,501]" degrees="104" direction="east" minutes="02.935" orientation="longitude" pageId="9" pageNumber="186" precision="1" value="104.04892">104°02.935′E</geoCoordinate>
, CMBS SEA 6139 DR361, rectangular dredge
<quantity id="4CB89B66FF9E1806FB3BFE64333CFD9D" box="[1233,1311,509,533]" metricMagnitude="1" metricUnit="m" metricValue="1.92" pageId="9" pageNumber="186" unit="m" value="19.2">
<elevation id="006DD1B0FF9E1806FB3BFE64333CFD9D" box="[1233,1311,509,533]" metricMagnitude="1" metricUnit="m" metricValue="1.92" pageId="9" pageNumber="186" unit="m" value="19.2">19.2 m</elevation>
</quantity>
, coll.
<collectorName id="26B55355FF9E1806FA89FE673388FD9E" box="[1379,1451,510,534]" pageId="9" pageNumber="186">T. Tay</collectorName>
and party,
<date id="FFFE1043FF9E1806FC49FD843210FDBD" box="[931,1075,541,565]" pageId="9" pageNumber="186" value="2014-04-08">
<collectingDate id="EFBAE9ABFF9E1806FC49FD843210FDBD" box="[931,1075,541,565]" pageId="9" pageNumber="186" value="2014-04-08">8 April 2014</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C35A6508FF9E1801FCC7FDC436D8F83D" lastPageId="14" lastPageNumber="191" pageId="9" pageNumber="186" type="description">
<paragraph id="8BFF3683FF9E1806FCC7FDC433AEFABD" blockId="9.[811,1453,605,1333]" pageId="9" pageNumber="186">
<emphasis id="B934EA91FF9E1806FCC7FDC4359BFDFD" bold="true" box="[813,952,605,629]" pageId="9" pageNumber="186">Description.</emphasis>
Body (
<figureCitation id="137B2A06FF9E1806FBE6FDC43242FDFD" box="[1036,1121,605,629]" captionStart="Fig" captionStartId="10.[135,170,2035,2057]" captionTargetBox="[169,1442,191,2020]" captionTargetId="figure-8@10.[121,1442,187,2021]" captionTargetPageId="10" captionText="Fig. 5. Cirolana trulla n. sp. A–C, E–F, holotype male (7.4 mm) (ZRC 2018.0206), Singapore. D, G, H, paratype male (6.0 mm) (ZRC 2018.0207), Singapore. I, paratype female (6.3 mm) (ZRC 2018.0207), Singapore. A, dorsal view; B, lateral view; C, head; D, antennula; E, frons; F, pleon; G, antenna peduncle; H, uropod male; I, uropod female." figureDoi="http://doi.org/10.5281/zenodo.5358571" httpUri="https://zenodo.org/record/5358571/files/figure.png" pageId="9" pageNumber="186">Fig. 5A</figureCitation>
) 4.3 times as long as greatest width, dorsal surfaces punctate, widest at pereonite 4, lateral margins subparallel. Rostral point absent. Eyes (
<figureCitation id="137B2A06FF9E1806FAA6FD043381FD3D" box="[1356,1442,669,693]" captionStart="Fig" captionStartId="10.[135,170,2035,2057]" captionTargetBox="[169,1442,191,2020]" captionTargetId="figure-8@10.[121,1442,187,2021]" captionTargetPageId="10" captionText="Fig. 5. Cirolana trulla n. sp. A–C, E–F, holotype male (7.4 mm) (ZRC 2018.0206), Singapore. D, G, H, paratype male (6.0 mm) (ZRC 2018.0207), Singapore. I, paratype female (6.3 mm) (ZRC 2018.0207), Singapore. A, dorsal view; B, lateral view; C, head; D, antennula; E, frons; F, pleon; G, antenna peduncle; H, uropod male; I, uropod female." figureDoi="http://doi.org/10.5281/zenodo.5358571" httpUri="https://zenodo.org/record/5358571/files/figure.png" pageId="9" pageNumber="186">Fig. 5C</figureCitation>
) separated by about 79% width of head, eye colour black. Pereonite 1 and coxae 2–3 (
<figureCitation id="137B2A06FF9E1806FBA6FD4432BEFD7D" box="[1100,1181,733,757]" captionStart="Fig" captionStartId="10.[135,170,2035,2057]" captionTargetBox="[169,1442,191,2020]" captionTargetId="figure-8@10.[121,1442,187,2021]" captionTargetPageId="10" captionText="Fig. 5. Cirolana trulla n. sp. A–C, E–F, holotype male (7.4 mm) (ZRC 2018.0206), Singapore. D, G, H, paratype male (6.0 mm) (ZRC 2018.0207), Singapore. I, paratype female (6.3 mm) (ZRC 2018.0207), Singapore. A, dorsal view; B, lateral view; C, head; D, antennula; E, frons; F, pleon; G, antenna peduncle; H, uropod male; I, uropod female." figureDoi="http://doi.org/10.5281/zenodo.5358571" httpUri="https://zenodo.org/record/5358571/files/figure.png" pageId="9" pageNumber="186">Fig. 5B</figureCitation>
) each with posteroventral angle rounded; coxae 5–7 with entire oblique carina; posterior margins of pereonites 6–7 with irregular submarginal nodules. Pleon (
<figureCitation id="137B2A06FF9E1806FC93FCA435E8FCDD" box="[889,971,829,853]" captionStart="Fig" captionStartId="10.[135,170,2035,2057]" captionTargetBox="[169,1442,191,2020]" captionTargetId="figure-8@10.[121,1442,187,2021]" captionTargetPageId="10" captionText="Fig. 5. Cirolana trulla n. sp. A–C, E–F, holotype male (7.4 mm) (ZRC 2018.0206), Singapore. D, G, H, paratype male (6.0 mm) (ZRC 2018.0207), Singapore. I, paratype female (6.3 mm) (ZRC 2018.0207), Singapore. A, dorsal view; B, lateral view; C, head; D, antennula; E, frons; F, pleon; G, antenna peduncle; H, uropod male; I, uropod female." figureDoi="http://doi.org/10.5281/zenodo.5358571" httpUri="https://zenodo.org/record/5358571/files/figure.png" pageId="9" pageNumber="186">Fig. 5F</figureCitation>
) with pleonite 1 not visible in dorsal view; pleonites 3–5 posterior margin with row of small, regularly spaced nodules; posterolateral angles of pleonite 2 forming acute point, extending posteriorly to anterior of pleonite 3; pleonite 3 without median tubercle and 4 small sublateral tubercles on each side, posterolateral margins not extending to posterior margin of pleonite 4 or 5, acute; pleonite 4 without median tubercle and 4 sublateral tubercles on each side, posterolateral margin of pleonite 4 rounded, clearly extending beyond posterior margin of pleonite 5; pleonite 5 without median tubercle and 1 sublateral tubercle on each side and posterolateral angles overlapped by lateral margins of pleonite 4. Pleotelson (
<figureCitation id="137B2A06FF9E1806FBB3FB243293FB5D" box="[1113,1200,1213,1237]" captionStart="Fig" captionStartId="11.[113,148,1417,1439]" captionTargetBox="[114,1451,192,1399]" captionTargetId="figure-10@11.[114,1451,191,1400]" captionTargetPageId="11" captionText="Fig. 6. Cirolana trulla n. sp. E, holotype male (7.4 mm) (ZRC 2018.0206), Singapore. A–D, F, paratype male (6.0 mm) (ZRC 2018.0207), Singapore. A, maxilliped; B, maxilla; C, maxillule; D, right mandible; E, penial process; F, pleotelson." figureDoi="http://doi.org/10.5281/zenodo.5358573" httpUri="https://zenodo.org/record/5358573/files/figure.png" pageId="9" pageNumber="186">Fig. 6F</figureCitation>
) 0.9 times as long as anterior width; with 2 tubercles on posterior of dorsal surface; lateral margins straight, margins smooth, posterior margin sub-truncate, without median point, with 8 robust setae.
</paragraph>
<paragraph id="8BFF3683FF9E1806FCC7FAC4333CF9FD" blockId="9.[811,1452,1373,1653]" pageId="9" pageNumber="186">
Antennula (
<figureCitation id="137B2A06FF9E1806FC40FAC435DEFAFD" box="[938,1021,1373,1397]" captionStart="Fig" captionStartId="10.[135,170,2035,2057]" captionTargetBox="[169,1442,191,2020]" captionTargetId="figure-8@10.[121,1442,187,2021]" captionTargetPageId="10" captionText="Fig. 5. Cirolana trulla n. sp. A–C, E–F, holotype male (7.4 mm) (ZRC 2018.0206), Singapore. D, G, H, paratype male (6.0 mm) (ZRC 2018.0207), Singapore. I, paratype female (6.3 mm) (ZRC 2018.0207), Singapore. A, dorsal view; B, lateral view; C, head; D, antennula; E, frons; F, pleon; G, antenna peduncle; H, uropod male; I, uropod female." figureDoi="http://doi.org/10.5281/zenodo.5358571" httpUri="https://zenodo.org/record/5358571/files/figure.png" pageId="9" pageNumber="186">Fig. 5D</figureCitation>
) peduncle articles 1 and 2 fused; articles 3 and 4 0.8 times as long as combined lengths of articles 1 and 2, article 3 1.9 times as long as wide; flagellum with 9 articles, extending to anterior of pereonite 1. Antenna (
<figureCitation id="137B2A06FF9E1806FA6AFA24356DFA7D" captionStart="Fig" captionStartId="10.[135,170,2035,2057]" captionTargetBox="[169,1442,191,2020]" captionTargetId="figure-8@10.[121,1442,187,2021]" captionTargetPageId="10" captionText="Fig. 5. Cirolana trulla n. sp. A–C, E–F, holotype male (7.4 mm) (ZRC 2018.0206), Singapore. D, G, H, paratype male (6.0 mm) (ZRC 2018.0207), Singapore. I, paratype female (6.3 mm) (ZRC 2018.0207), Singapore. A, dorsal view; B, lateral view; C, head; D, antennula; E, frons; F, pleon; G, antenna peduncle; H, uropod male; I, uropod female." figureDoi="http://doi.org/10.5281/zenodo.5358571" httpUri="https://zenodo.org/record/5358571/files/figure.png" pageId="9" pageNumber="186">Fig. 5G</figureCitation>
) peduncle article 4 1.6 times as long as wide, 1.3 times as long as article 3, and 6 short simple setae (distal); article 5 1.3 as long as article 4, 2.5 times as long as wide, anterodistal angle with cluster of 1 short simple setae; flagellum with 19 articles, extending to posterior of pereonite 4.
</paragraph>
<paragraph id="8BFF3683FF9E1806FCC7F90432D2F97D" blockId="9.[812,1451,1693,1781]" pageId="9" pageNumber="186">
Frontal lamina (
<figureCitation id="137B2A06FF9E1806FC3CF9043206F93D" box="[982,1061,1693,1717]" captionStart="Fig" captionStartId="10.[135,170,2035,2057]" captionTargetBox="[169,1442,191,2020]" captionTargetId="figure-8@10.[121,1442,187,2021]" captionTargetPageId="10" captionText="Fig. 5. Cirolana trulla n. sp. A–C, E–F, holotype male (7.4 mm) (ZRC 2018.0206), Singapore. D, G, H, paratype male (6.0 mm) (ZRC 2018.0207), Singapore. I, paratype female (6.3 mm) (ZRC 2018.0207), Singapore. A, dorsal view; B, lateral view; C, head; D, antennula; E, frons; F, pleon; G, antenna peduncle; H, uropod male; I, uropod female." figureDoi="http://doi.org/10.5281/zenodo.5358571" httpUri="https://zenodo.org/record/5358571/files/figure.png" pageId="9" pageNumber="186">Fig. 5E</figureCitation>
) pentagonal, 1.9 longer than greatest width, lateral margins straight and parallel, anterior margin straight and acute, forming median point.
</paragraph>
<paragraph id="8BFF3683FF9E1804FCC7F884343CF942" blockId="9.[811,1453,1821,2101]" lastBlockId="11.[113,755,1523,1739]" lastPageId="11" lastPageNumber="188" pageId="9" pageNumber="186">
Mandible (
<figureCitation id="137B2A06FF9E1806FC49F88435DAF8BD" box="[931,1017,1821,1845]" captionStart="Fig" captionStartId="11.[113,148,1417,1439]" captionTargetBox="[114,1451,192,1399]" captionTargetId="figure-10@11.[114,1451,191,1400]" captionTargetPageId="11" captionText="Fig. 6. Cirolana trulla n. sp. E, holotype male (7.4 mm) (ZRC 2018.0206), Singapore. A–D, F, paratype male (6.0 mm) (ZRC 2018.0207), Singapore. A, maxilliped; B, maxilla; C, maxillule; D, right mandible; E, penial process; F, pleotelson." figureDoi="http://doi.org/10.5281/zenodo.5358573" httpUri="https://zenodo.org/record/5358573/files/figure.png" pageId="9" pageNumber="186">Fig. 6D</figureCitation>
) molar process anterior margin with 15 flat teeth; with proximal cluster of long simple setae; right mandible spine row composed of 4 spines; palp article 2 with 12 distolateral setae, palp article 3 with 4 robust biserrate and 11 plumose setae. Maxillule (
<figureCitation id="137B2A06FF9E1806FB72F80432C8F83D" box="[1176,1259,1949,1973]" captionStart="Fig" captionStartId="11.[113,148,1417,1439]" captionTargetBox="[114,1451,192,1399]" captionTargetId="figure-10@11.[114,1451,191,1400]" captionTargetPageId="11" captionText="Fig. 6. Cirolana trulla n. sp. E, holotype male (7.4 mm) (ZRC 2018.0206), Singapore. A–D, F, paratype male (6.0 mm) (ZRC 2018.0207), Singapore. A, maxilliped; B, maxilla; C, maxillule; D, right mandible; E, penial process; F, pleotelson." figureDoi="http://doi.org/10.5281/zenodo.5358573" httpUri="https://zenodo.org/record/5358573/files/figure.png" pageId="9" pageNumber="186">Fig. 6C</figureCitation>
) mesial lobe with 3 large and circumplumose RS; lateral lobe with 13 RS. Maxilla (
<figureCitation id="137B2A06FF9E1806FC7FF84435CEF87D" box="[917,1005,2013,2037]" captionStart="Fig" captionStartId="11.[113,148,1417,1439]" captionTargetBox="[114,1451,192,1399]" captionTargetId="figure-10@11.[114,1451,191,1400]" captionTargetPageId="11" captionText="Fig. 6. Cirolana trulla n. sp. E, holotype male (7.4 mm) (ZRC 2018.0206), Singapore. A–D, F, paratype male (6.0 mm) (ZRC 2018.0207), Singapore. A, maxilliped; B, maxilla; C, maxillule; D, right mandible; E, penial process; F, pleotelson." figureDoi="http://doi.org/10.5281/zenodo.5358573" httpUri="https://zenodo.org/record/5358573/files/figure.png" pageId="9" pageNumber="186">Fig. 6B</figureCitation>
) lateral lobe with 4 long simple setae; middle lobe with 8 long simple setae; mesial lobe with 4 distal simple setae, with 2 proximal simple and 4 plumose setae. Maxilliped (
<figureCitation id="137B2A06FF9C1804FEAEFA6A37BEF983" box="[324,413,1523,1547]" captionStart="Fig" captionStartId="11.[113,148,1417,1439]" captionTargetBox="[114,1451,192,1399]" captionTargetId="figure-10@11.[114,1451,191,1400]" captionTargetPageId="11" captionText="Fig. 6. Cirolana trulla n. sp. E, holotype male (7.4 mm) (ZRC 2018.0206), Singapore. A–D, F, paratype male (6.0 mm) (ZRC 2018.0207), Singapore. A, maxilliped; B, maxilla; C, maxillule; D, right mandible; E, penial process; F, pleotelson." figureDoi="http://doi.org/10.5281/zenodo.5358573" httpUri="https://zenodo.org/record/5358573/files/figure.png" pageId="11" pageNumber="188">Fig. 6A</figureCitation>
) palp article 2 mesial margin with 9 slender setae, lateral margin distally with 1 slender setae; article 3 mesial margin with 16 slender setae, lateral margin with 10 slender setae; article 4 mesial margin with 11 slender setae, lateral margin with 5 slender setae; article 5 distal margin 12 setae, lateral margin with 6 setae; endite with 4 long CPS, and 2 coupling setae.
</paragraph>
<caption id="DF3F660BFF9D1805FF6DF86A354FF7CB" ID-DOI="http://doi.org/10.5281/zenodo.5358571" ID-Zenodo-Dep="5358571" httpUri="https://zenodo.org/record/5358571/files/figure.png" pageId="10" pageNumber="187" startId="10.[135,170,2035,2057]" targetBox="[169,1442,191,2020]" targetPageId="10">
<paragraph id="8BFF3683FF9D1805FF6DF86A354FF7CB" blockId="10.[135,1476,2035,2115]" pageId="10" pageNumber="187">
Fig. 5.
<taxonomicName id="4C404D00FF9D1805FF3BF86A3746F780" authority="Sidabalok &amp; Bruce, 2018" authorityName="Sidabalok &amp; Bruce" authorityYear="2018" box="[209,357,2035,2056]" class="Malacostraca" family="Cirolanidae" genus="Cirolana" kingdom="Animalia" order="Isopoda" pageId="10" pageNumber="187" phylum="Arthropoda" rank="species" species="trulla" status="sp. nov.">
<emphasis id="B934EA91FF9D1805FF3BF86A3746F780" box="[209,357,2035,2056]" italics="true" pageId="10" pageNumber="187">Cirolana trulla</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20757EAFF9D1805FE87F86D3780F781" box="[365,419,2036,2057]" pageId="10" pageNumber="187" rank="species">n. sp.</taxonomicNameLabel>
A–C, E–F, holotype male (7.4 mm) (ZRC 2018.0206), Singapore. D, G, H, paratype male (6.0 mm) (ZRC 2018.0207), Singapore. I, paratype female (6.3 mm) (ZRC 2018.0207), Singapore. A, dorsal view; B, lateral view; C, head; D, antennula; E, frons; F, pleon; G, antenna peduncle; H, uropod male; I, uropod female.
</paragraph>
</caption>
<caption id="DF3F660BFF9C1804FF9BFA103245FA33" ID-DOI="http://doi.org/10.5281/zenodo.5358573" ID-Zenodo-Dep="5358573" httpUri="https://zenodo.org/record/5358573/files/figure.png" pageId="11" pageNumber="188" startId="11.[113,148,1417,1439]" targetBox="[114,1451,192,1399]" targetPageId="11">
<paragraph id="8BFF3683FF9C1804FF9BFA103245FA33" blockId="11.[113,1452,1417,1467]" pageId="11" pageNumber="188">
Fig. 6.
<taxonomicName id="4C404D00FF9C1804FF5CFA10376BFA16" authority="Sidabalok &amp; Bruce, 2018" authorityName="Sidabalok &amp; Bruce" authorityYear="2018" box="[182,328,1417,1438]" class="Malacostraca" family="Cirolanidae" genus="Cirolana" kingdom="Animalia" order="Isopoda" pageId="11" pageNumber="188" phylum="Arthropoda" rank="species" species="trulla" status="sp. nov.">
<emphasis id="B934EA91FF9C1804FF5CFA10376BFA16" box="[182,328,1417,1438]" italics="true" pageId="11" pageNumber="188">Cirolana trulla</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20757EAFF9C1804FEA5FA1037A0FA16" box="[335,387,1417,1438]" pageId="11" pageNumber="188" rank="species">n. sp.</taxonomicNameLabel>
E, holotype male (7.4 mm) (ZRC 2018.0206), Singapore. A–D, F, paratype male (6.0 mm) (ZRC 2018.0207), Singapore. A, maxilliped; B, maxilla; C, maxillule; D, right mandible; E, penial process; F, pleotelson.
</paragraph>
</caption>
<paragraph id="8BFF3683FF9C1801FF9BF96A3494FDBD" blockId="11.[113,754,1779,2091]" lastBlockId="14.[135,776,189,565]" lastPageId="14" lastPageNumber="191" pageId="11" pageNumber="188">
Pereopod 1 (
<figureCitation id="137B2A06FF9C1804FEE8F96A3778F883" box="[258,347,1779,1803]" captionStart="Fig" captionStartId="12.[135,170,2064,2086]" captionTargetBox="[168,1442,189,2047]" captionTargetId="figure-8@12.[168,1442,189,2047]" captionTargetPageId="12" captionText="Fig. 7. Cirolana trulla n. sp. A–D, paratype male (6.0 mm) (ZRC 2018.0207), Singapore. A, pereopod 1; B, pereopod 2; C, pereopod 6; D, pereopod 7." figureDoi="http://doi.org/10.5281/zenodo.5358575" httpUri="https://zenodo.org/record/5358575/files/figure.png" pageId="11" pageNumber="188">Fig. 7A</figureCitation>
) basis 2.2 times as long as greatest width, superior distal angle without cluster of acute setae, inferior distal angle with a cluster of 4 acute setae; ischium 0.6 times as long as basis, inferior margin with 2 setae, inferior distal margin with 2 RS, superior distal margin with 4 long RS; merus inferior margin with 6 molariform RS, set as 4 and 2, with 5 simple setae, superior distal angle with 3 RS; carpus inferior margin with 1 RS and 1 simple seta; propodus 1.7 times as long as wide, inferior margin with 2 RS, inferior distal margin with 1 large RS and 1 simple seta, superior distal with 2 simple setae; dactylus 0.9 as long as propodus. Pereopod 2 (
<figureCitation id="137B2A06FF9C1804FB7AF98A32C5F9A3" box="[1168,1254,1555,1579]" captionStart="Fig" captionStartId="12.[135,170,2064,2086]" captionTargetBox="[168,1442,189,2047]" captionTargetId="figure-8@12.[168,1442,189,2047]" captionTargetPageId="12" captionText="Fig. 7. Cirolana trulla n. sp. A–D, paratype male (6.0 mm) (ZRC 2018.0207), Singapore. A, pereopod 1; B, pereopod 2; C, pereopod 6; D, pereopod 7." figureDoi="http://doi.org/10.5281/zenodo.5358575" httpUri="https://zenodo.org/record/5358575/files/figure.png" pageId="11" pageNumber="188">Fig. 7B</figureCitation>
) ischium inferior margin with 1 long acute RS, inferior distal margin with 2 RS, 1 acute RS, 1 simple seta, superior distal margin with 2 long acute setae and 3 plumose setae; merus inferior margin with 4 stout RS and 2 simple setae, set as single row, inferior distal margin with 1 stout RS, 2 acute RS and 1 simple seta, superior distal margin with 1 long acute RS and 4 simple setae; carpus inferior distal margin with 2 acute RS; propodus 2.7 as long as wide, with 2 clusters of acute RS, inferior margin with 2 RS, inferior distal margin with 1 large RS and 1 simple seta; dactylus 0.7 as long as propodus. Pereopod 3 similar to pereopod 2. Pereopod 6 (
<figureCitation id="137B2A06FF9C1804FCDFF80A35A4F823" box="[821,903,1939,1963]" captionStart="Fig" captionStartId="12.[135,170,2064,2086]" captionTargetBox="[168,1442,189,2047]" captionTargetId="figure-8@12.[168,1442,189,2047]" captionTargetPageId="12" captionText="Fig. 7. Cirolana trulla n. sp. A–D, paratype male (6.0 mm) (ZRC 2018.0207), Singapore. A, pereopod 1; B, pereopod 2; C, pereopod 6; D, pereopod 7." figureDoi="http://doi.org/10.5281/zenodo.5358575" httpUri="https://zenodo.org/record/5358575/files/figure.png" pageId="11" pageNumber="188">Fig. 7C</figureCitation>
) similar to pereopod 7. Pereopod 7 (
<figureCitation id="137B2A06FF9C1804FAF9F80A3345F823" box="[1299,1382,1939,1963]" captionStart="Fig" captionStartId="12.[135,170,2064,2086]" captionTargetBox="[168,1442,189,2047]" captionTargetId="figure-8@12.[168,1442,189,2047]" captionTargetPageId="12" captionText="Fig. 7. Cirolana trulla n. sp. A–D, paratype male (6.0 mm) (ZRC 2018.0207), Singapore. A, pereopod 1; B, pereopod 2; C, pereopod 6; D, pereopod 7." figureDoi="http://doi.org/10.5281/zenodo.5358575" httpUri="https://zenodo.org/record/5358575/files/figure.png" pageId="11" pageNumber="188">Fig. 7D</figureCitation>
) basis 2.5 times as long as greatest width, superior margin straight, with 3 palmate setae; ischium 0.5 as long as basis, inferior margin with 4 long simple setae (set in group of 2 and 2), with 3 triangular processes, inferior distal angle with 2 long simple setae, superior distal angle with 5 long setae and 2 biserrate setae; merus 1.1 as long as ischium, 1.9 times as long as wide, inferior margin with 1 acute RS and 3 simple setae, superior distal angle with 11 RS (10 biserrate), inferior distal angle with 5 RS and 3 simple setae; carpus as long as ischium, 0.4 times as long as wide, inferior margin with 3 RS and 1 simple seta, superior distal angle with 8 biserrate RS, inferior distal angle with 9 RS (5 biserrate); propodus as long as ischium, 3.8 times as long as wide, inferior margin with 2 clusters of RS (set as 1 and 1), superior distal angle with 1 slender seta, 1plumose seta and 3 RS, inferior distal angle with 2 RS; dactylus 0.5 as long as propodus.
</paragraph>
<caption id="DF3F660BFF9B1803FF6DF7893738F7CA" ID-DOI="http://doi.org/10.5281/zenodo.5358575" ID-Zenodo-Dep="5358575" httpUri="https://zenodo.org/record/5358575/files/figure.png" pageId="12" pageNumber="189" startId="12.[135,170,2064,2086]" targetBox="[168,1442,189,2047]" targetPageId="12">
<paragraph id="8BFF3683FF9B1803FF6DF7893738F7CA" blockId="12.[135,1474,2064,2115]" pageId="12" pageNumber="189">
Fig. 7.
<taxonomicName id="4C404D00FF9B1803FF3AF7893740F7AD" authority="Sidabalok &amp; Bruce, 2018" authorityName="Sidabalok &amp; Bruce" authorityYear="2018" box="[208,355,2064,2085]" class="Malacostraca" family="Cirolanidae" genus="Cirolana" kingdom="Animalia" order="Isopoda" pageId="12" pageNumber="189" phylum="Arthropoda" rank="species" species="trulla" status="sp. nov.">
<emphasis id="B934EA91FF9B1803FF3AF7893740F7AD" box="[208,355,2064,2085]" italics="true" pageId="12" pageNumber="189">Cirolana trulla</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20757EAFF9B1803FE80F78937BCF7AD" box="[362,415,2064,2085]" pageId="12" pageNumber="189" rank="species">n. sp.</taxonomicNameLabel>
A–D, paratype male (6.0 mm) (ZRC 2018.0207), Singapore. A, pereopod 1; B, pereopod 2; C, pereopod 6; D, pereopod 7.
</paragraph>
</caption>
<caption id="DF3F660BFF9A1802FF9BF8143745F837" ID-DOI="http://doi.org/10.5281/zenodo.5358577" ID-Zenodo-Dep="5358577" httpUri="https://zenodo.org/record/5358577/files/figure.png" pageId="13" pageNumber="190" startId="13.[113,148,1933,1955]" targetBox="[114,1451,320,1917]" targetPageId="13">
<paragraph id="8BFF3683FF9A1802FF9BF8143745F837" blockId="13.[113,1453,1932,1983]" pageId="13" pageNumber="190">
Fig. 8.
<taxonomicName id="4C404D00FF9A1802FF51F814376DF82A" authority="Sidabalok &amp; Bruce, 2018" authorityName="Sidabalok &amp; Bruce" authorityYear="2018" box="[187,334,1933,1954]" class="Malacostraca" family="Cirolanidae" genus="Cirolana" kingdom="Animalia" order="Isopoda" pageId="13" pageNumber="190" phylum="Arthropoda" rank="species" species="trulla" status="sp. nov.">
<emphasis id="B934EA91FF9A1802FF51F814376DF82A" box="[187,334,1933,1954]" italics="true" pageId="13" pageNumber="190">Cirolana trulla</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20757EAFF9A1802FEBDF81437AEF82A" box="[343,397,1933,1954]" pageId="13" pageNumber="190" rank="species">n. sp.</taxonomicNameLabel>
A–E, paratype male (6.0 mm) (ZRC 2018.0207), Singapore. A, pleopod 1; B, pleopod 2; C, pleopod 3; D, pleopod 4; E, pleopod 5.
</paragraph>
</caption>
<paragraph id="8BFF3683FF991801FF62FDC73758FD1D" blockId="14.[136,775,605,661]" pageId="14" pageNumber="191">
Penes (
<figureCitation id="137B2A06FF991801FF3FFDC43705FDFD" box="[213,294,605,629]" captionStart="Fig" captionStartId="11.[113,148,1417,1439]" captionTargetBox="[114,1451,192,1399]" captionTargetId="figure-10@11.[114,1451,191,1400]" captionTargetPageId="11" captionText="Fig. 6. Cirolana trulla n. sp. E, holotype male (7.4 mm) (ZRC 2018.0206), Singapore. A–D, F, paratype male (6.0 mm) (ZRC 2018.0207), Singapore. A, maxilliped; B, maxilla; C, maxillule; D, right mandible; E, penial process; F, pleotelson." figureDoi="http://doi.org/10.5281/zenodo.5358573" httpUri="https://zenodo.org/record/5358573/files/figure.png" pageId="14" pageNumber="191">Fig. 6E</figureCitation>
) low tubercles, penial openings separated by 2.9% of sternal width.
</paragraph>
<paragraph id="8BFF3683FF991801FF62FD2437F9FB3D" blockId="14.[136,775,701,1205]" pageId="14" pageNumber="191">
Pleopod 1 (
<figureCitation id="137B2A06FF991801FEE4FD24374BFD5D" box="[270,360,701,725]" captionStart="Fig" captionStartId="13.[113,148,1933,1955]" captionTargetBox="[114,1451,320,1917]" captionTargetId="figure-10@13.[114,1451,320,1918]" captionTargetPageId="13" captionText="Fig. 8. Cirolana trulla n. sp. A–E, paratype male (6.0 mm) (ZRC 2018.0207), Singapore. A, pleopod 1; B, pleopod 2; C, pleopod 3; D, pleopod 4; E, pleopod 5." figureDoi="http://doi.org/10.5281/zenodo.5358577" httpUri="https://zenodo.org/record/5358577/files/figure.png" pageId="14" pageNumber="191">Fig. 8A</figureCitation>
) exopod 1.7 times as long as wide, lateral margin straight, distally narrowly rounded, medial margin weakly oblique, mesial margin strongly convex, with PMS from distal one-third, with ~38 PMS; endopod 2.4 times as long as wide, distally narrowly rounded, lateral margin straight, with PMS on distal margin only, mesial margin with PMS from distal one-third, endopod with ~13 PMS; peduncle 1.3 times as wide as long. Pleopod 2 (
<figureCitation id="137B2A06FF991801FF7AFC2436C7FC5D" box="[144,228,957,981]" captionStart="Fig" captionStartId="13.[113,148,1933,1955]" captionTargetBox="[114,1451,320,1917]" captionTargetId="figure-10@13.[114,1451,320,1918]" captionTargetPageId="13" captionText="Fig. 8. Cirolana trulla n. sp. A–E, paratype male (6.0 mm) (ZRC 2018.0207), Singapore. A, pleopod 1; B, pleopod 2; C, pleopod 3; D, pleopod 4; E, pleopod 5." figureDoi="http://doi.org/10.5281/zenodo.5358577" httpUri="https://zenodo.org/record/5358577/files/figure.png" pageId="14" pageNumber="191">Fig. 8B</figureCitation>
) exopod with ~40 PMS, endopod with ~16 PMS; appendix masculina with parallel margins, 0.9 times as long as endopod, distally acute. Pleopod 3 (
<figureCitation id="137B2A06FF991801FDC8FC643450FB9D" box="[546,627,1021,1045]" captionStart="Fig" captionStartId="13.[113,148,1933,1955]" captionTargetBox="[114,1451,320,1917]" captionTargetId="figure-10@13.[114,1451,320,1918]" captionTargetPageId="13" captionText="Fig. 8. Cirolana trulla n. sp. A–E, paratype male (6.0 mm) (ZRC 2018.0207), Singapore. A, pleopod 1; B, pleopod 2; C, pleopod 3; D, pleopod 4; E, pleopod 5." figureDoi="http://doi.org/10.5281/zenodo.5358577" httpUri="https://zenodo.org/record/5358577/files/figure.png" pageId="14" pageNumber="191">Fig. 8C</figureCitation>
) exopod with ~43 PMS, endopod with ~12 PMS. Pleopod 4 (
<figureCitation id="137B2A06FF991801FD4CFB8434DDFBBD" box="[678,766,1053,1077]" captionStart="Fig" captionStartId="13.[113,148,1933,1955]" captionTargetBox="[114,1451,320,1917]" captionTargetId="figure-10@13.[114,1451,320,1918]" captionTargetPageId="13" captionText="Fig. 8. Cirolana trulla n. sp. A–E, paratype male (6.0 mm) (ZRC 2018.0207), Singapore. A, pleopod 1; B, pleopod 2; C, pleopod 3; D, pleopod 4; E, pleopod 5." figureDoi="http://doi.org/10.5281/zenodo.5358577" httpUri="https://zenodo.org/record/5358577/files/figure.png" pageId="14" pageNumber="191">Fig. 8D</figureCitation>
) exopod with ~37 PMS, endopod with ~9 PMS. Pleopod 5 (
<figureCitation id="137B2A06FF991801FF7AFBC436C7FBFD" box="[144,228,1117,1141]" captionStart="Fig" captionStartId="13.[113,148,1933,1955]" captionTargetBox="[114,1451,320,1917]" captionTargetId="figure-10@13.[114,1451,320,1918]" captionTargetPageId="13" captionText="Fig. 8. Cirolana trulla n. sp. A–E, paratype male (6.0 mm) (ZRC 2018.0207), Singapore. A, pleopod 1; B, pleopod 2; C, pleopod 3; D, pleopod 4; E, pleopod 5." figureDoi="http://doi.org/10.5281/zenodo.5358577" httpUri="https://zenodo.org/record/5358577/files/figure.png" pageId="14" pageNumber="191">Fig. 8E</figureCitation>
) exopod with ~38 PMS. Pleopods 2–5 peduncle distolateral margin with prominent acute RS, 3–5 endopods with distomesial serrate scales.
</paragraph>
<paragraph id="8BFF3683FF991801FF62FB443524F9BD" blockId="14.[136,775,1245,1589]" pageId="14" pageNumber="191">
Uropod (
<figureCitation id="137B2A06FF991801FF04FB443748FB7D" box="[238,363,1245,1269]" captionStart="Fig" captionStartId="10.[135,170,2035,2057]" captionTargetBox="[169,1442,191,2020]" captionTargetId="figure-8@10.[121,1442,187,2021]" captionTargetPageId="10" captionText="Fig. 5. Cirolana trulla n. sp. A–C, E–F, holotype male (7.4 mm) (ZRC 2018.0206), Singapore. D, G, H, paratype male (6.0 mm) (ZRC 2018.0207), Singapore. I, paratype female (6.3 mm) (ZRC 2018.0207), Singapore. A, dorsal view; B, lateral view; C, head; D, antennula; E, frons; F, pleon; G, antenna peduncle; H, uropod male; I, uropod female." figureDoi="http://doi.org/10.5281/zenodo.5358571" httpUri="https://zenodo.org/record/5358571/files/figure.png" pageId="14" pageNumber="191">Fig. 5H–5I</figureCitation>
) peduncle ventrolateral margin with 1 RS, lateral margin with medial short acute RS, posterior lobe about 0.9 as long as endopod; rami extending beyond pleotelson, marginal setae dense, in two tiers, apices not bifid. Endopod lateral margin proximally convex, without prominent excision, proximal lateral margin with 1 RS, distal lateral margin with 1 RS; mesial margin strongly convex, with 9 RS. Exopod not extending to end of endopod, 1.8 times as long as greatest width; lateral margin weakly convex with setal fringe on the posterior, with 3 visible widely separated RS; mesial margin straight, proximally convex, with 4 RS.
</paragraph>
<paragraph id="8BFF3683FF991801FF62F9C437F9F91D" blockId="14.[136,775,1629,1685]" pageId="14" pageNumber="191">
<emphasis id="B934EA91FF991801FF62F9C436C1F9FD" bold="true" box="[136,226,1629,1653]" pageId="14" pageNumber="191">Female.</emphasis>
The uropodal exopod lacks the dense setal fringe on the posterior lateral margin.
</paragraph>
<paragraph id="8BFF3683FF991801FF62F9243479F97D" blockId="14.[135,775,1725,1782]" pageId="14" pageNumber="191">
<emphasis id="B934EA91FF991801FF62F924369EF95D" bold="true" box="[136,189,1725,1749]" pageId="14" pageNumber="191">Size.</emphasis>
Males
<quantity id="4CB89B66FF991801FEE2F92437AAF95D" box="[264,393,1725,1749]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" metricValueMax="7.8" metricValueMin="5.2" pageId="14" pageNumber="191" unit="mm" value="6.5" valueMax="7.8" valueMin="5.2">5.2–7.8 mm</quantity>
, mean
<quantity id="4CB89B66FF991801FE3BF9243403F95D" box="[465,544,1725,1749]" metricMagnitude="-3" metricUnit="m" metricValue="6.6" pageId="14" pageNumber="191" unit="mm" value="6.6">6.6 mm</quantity>
(n=7); non-ovigerous female
<quantity id="4CB89B66FF991801FF32F9443708F97D" box="[216,299,1757,1781]" metricMagnitude="-3" metricUnit="m" metricValue="6.3" pageId="14" pageNumber="191" unit="mm" value="6.3">6.3 mm</quantity>
(n=1); all from
<typeStatus id="54FB8821FF991801FE35F947342EF97E" box="[479,525,1758,1782]" pageId="14" pageNumber="191">type</typeStatus>
series.
</paragraph>
<paragraph id="8BFF3683FF991801FF62F88436D8F83D" blockId="14.[136,778,1821,1973]" pageId="14" pageNumber="191">
<emphasis id="B934EA91FF991801FF62F88436DEF8BD" bold="true" box="[136,253,1821,1845]" pageId="14" pageNumber="191">Variation.</emphasis>
RS count was measured from all
<typeStatus id="54FB8821FF991801FD9EF8873481F8BE" box="[628,674,1822,1846]" pageId="14" pageNumber="191">type</typeStatus>
series (n = 8). Uropod exopod lateral: 3 = 75%, 4 = 25%; mesial: 3 = 50%, 4 = 50%. Uropod endopod lateral: 2 = 87.5%, 3 = 12.5%; mesial: 7 = 12.5%, 8 = 62.5%, 9 = 25%. Pleotelson: 8 = 100%.
</paragraph>
</subSubSection>
<subSubSection id="C35A6508FF991801FF62F8443381FB7D" pageId="14" pageNumber="191" type="discussion">
<paragraph id="8BFF3683FF991801FF62F84432FCFE3D" blockId="14.[136,775,2013,2101]" lastBlockId="14.[834,1476,189,437]" pageId="14" pageNumber="191">
<emphasis id="B934EA91FF991801FF62F84436D4F87D" bold="true" box="[136,247,2013,2037]" pageId="14" pageNumber="191">Remarks.</emphasis>
<taxonomicName id="4C404D00FF991801FF14F8443782F87D" authority="Sidabalok &amp; Bruce, 2018" authorityName="Sidabalok &amp; Bruce" authorityYear="2018" box="[254,417,2013,2037]" class="Malacostraca" family="Cirolanidae" genus="Cirolana" kingdom="Animalia" order="Isopoda" pageId="14" pageNumber="191" phylum="Arthropoda" rank="species" species="trulla" status="sp. nov.">
<emphasis id="B934EA91FF991801FF14F8443782F87D" box="[254,417,2013,2037]" italics="true" pageId="14" pageNumber="191">Cirolana trulla</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A20757EAFF991801FE45F8473410F87D" box="[431,563,2013,2037]" pageId="14" pageNumber="191" rank="species">new species</taxonomicNameLabel>
, is characterised by having a more elongate body than most
<taxonomicName id="4C404D00FF991801FDA0F8643489F79D" authorityName="Leach" authorityYear="1818" box="[586,682,2045,2069]" class="Malacostraca" family="Cirolanidae" genus="Cirolana" kingdom="Animalia" order="Isopoda" pageId="14" pageNumber="191" phylum="Arthropoda" rank="genus">
<emphasis id="B934EA91FF991801FDA0F8643489F79D" box="[586,682,2045,2069]" italics="true" pageId="14" pageNumber="191">Cirolana</emphasis>
</taxonomicName>
(4.3 vs. ~3.0 times as long as wide), a triangular rostrum in dorsal view, the presence of three triangular processes (= broadbased spines) on the inferior margin of ischium of pereopods 6–7, low tubercle-like penial openings (not processes) and the presence of two small posteriorly positioned nodules on of pleotelson. The male uropodal rami are distinctive in shape, but especially distinctive in terms of their setation, with a two-tier and dense setal fringe on the posterior of lateral and mesial margins of exopod.
</paragraph>
<paragraph id="8BFF3683FF991801FCA9FE443381FB7D" blockId="14.[834,1476,477,1269]" pageId="14" pageNumber="191">
<taxonomicName id="4C404D00FF991801FCA9FE4435C4FE7D" authority="Sidabalok &amp; Bruce, 2018" authorityName="Sidabalok &amp; Bruce" authorityYear="2018" box="[835,999,477,501]" class="Malacostraca" family="Cirolanidae" genus="Cirolana" kingdom="Animalia" order="Isopoda" pageId="14" pageNumber="191" phylum="Arthropoda" rank="species" species="trulla" status="sp. nov.">
<emphasis id="B934EA91FF991801FCA9FE4435C4FE7D" box="[835,999,477,501]" italics="true" pageId="14" pageNumber="191">Cirolana trulla</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A20757EAFF991801FC1DFE47325FFE7D" box="[1015,1148,477,501]" pageId="14" pageNumber="191" rank="species">new species</taxonomicNameLabel>
, is most similar to
<taxonomicName id="4C404D00FF991801FAB9FE47326BFD9D" authority="Bruce &amp; Olesen, 2002" authorityName="Bruce &amp; Olesen" authorityYear="2002" class="Malacostraca" family="Cirolanidae" genus="Cirolana" kingdom="Animalia" order="Isopoda" pageId="14" pageNumber="191" phylum="Arthropoda" rank="species" species="bruscai">
<emphasis id="B934EA91FF991801FAB9FE4733E7FE7D" box="[1363,1476,477,501]" italics="true" pageId="14" pageNumber="191">C. bruscai</emphasis>
<bibRefCitation id="EFD14B72FF991801FCA9FE67326BFD9D" author="Bruce NL &amp; Olesen J" box="[835,1096,509,533]" pageId="14" pageNumber="191" pagination="109 - 131" refId="ref16210" refString="Bruce NL &amp; Olesen J (2002) Cirolanid isopods from the Andaman Sea off Phuket, Thailand, with description of two new species. Phuket Marine Biological Center Special Publication, 23: 109 - 131." type="journal article" year="2002">Bruce &amp; Olesen, 2002</bibRefCitation>
</taxonomicName>
, both species having a relatively elongate body, the head anteriorly produced and overriding the antennular bases, a few sublateral tubercles on pereonites 3–5, scattered nodules on posterior of pleotelson, long simple setae on superior distal angle of the ischium– merus of pereopods 1–2 and on the inferior margin of the ischium–merus of pereopods 6–7, the triangular processes on the ischium of pereopods 6–7, the two tiered-setae that form dense mass of setae on the posterior lateral margin of uropod exopod and endopod. Despite these similarities there are numerous differences between the two species:
<taxonomicName id="4C404D00FF991801FCA9FCC435CAFCFD" authorityName="Sidabalok &amp; Bruce" authorityYear="2018" box="[835,1001,861,885]" class="Malacostraca" family="Cirolanidae" genus="Cirolana" kingdom="Animalia" order="Isopoda" pageId="14" pageNumber="191" phylum="Arthropoda" rank="species" species="trulla">
<emphasis id="B934EA91FF991801FCA9FCC435CAFCFD" box="[835,1001,861,885]" italics="true" pageId="14" pageNumber="191">Cirolana trulla</emphasis>
</taxonomicName>
is 4.3 times as long as greatest width (vs.
<quantity id="4CB89B66FF991801FCA9FCE435A1FC1D" box="[835,898,893,917]" metricMagnitude="-2" metricUnit="m" metricValue="7.874" pageId="14" pageNumber="191" unit="in" value="3.1">3.1 in</quantity>
<taxonomicName id="4C404D00FF991801FC61FCE735D9FC1D" authorityName="Bruce &amp; Olesen" authorityYear="2002" box="[907,1018,893,917]" class="Malacostraca" family="Cirolanidae" genus="Cirolana" kingdom="Animalia" order="Isopoda" pageId="14" pageNumber="191" phylum="Arthropoda" rank="species" species="bruscai">
<emphasis id="B934EA91FF991801FC61FCE735D9FC1D" box="[907,1018,893,917]" italics="true" pageId="14" pageNumber="191">C. bruscai</emphasis>
</taxonomicName>
), has 8 robust setae on the pleotelson (vs. 6), the anterior of the head is strongly produced (vs. weakly produced), pleopod 1 endopod and exopod lateral margins are straight (vs. convex); uropodal exopod not extending to end of endopod (vs. beyond) with a narrow apex (vs. semi-truncate), the mesial margin is distinctly angled midlength (vs. evenly convex) and the second submarginal tier of setae (distal one-third vs. distal three-quarters), and two small submedian tubercles on pleotelson (vs. absent). The body length-to-width ratio (2.5) of
<taxonomicName id="4C404D00FF991801FB2FFB04335DFB3D" authorityName="Bruce &amp; Olesen" authorityYear="2002" box="[1221,1406,1181,1205]" class="Malacostraca" family="Cirolanidae" genus="Cirolana" kingdom="Animalia" order="Isopoda" pageId="14" pageNumber="191" phylum="Arthropoda" rank="species" species="bruscai">
<emphasis id="B934EA91FF991801FB2FFB04335DFB3D" box="[1221,1406,1181,1205]" italics="true" pageId="14" pageNumber="191">Cirolana bruscai</emphasis>
</taxonomicName>
in the original description is inaccurate owing to the curved state of the
<typeStatus id="54FB8821FF991801FC67FB4435CDFB7D" box="[909,1006,1245,1269]" pageId="14" pageNumber="191" type="holotype">holotype</typeStatus>
; we recalculated the ratio, which is 3.1.
</paragraph>
</subSubSection>
<subSubSection id="C35A6508FF991801FCA9FA8432DCFABD" box="[835,1279,1309,1333]" pageId="14" pageNumber="191" type="distribution">
<paragraph id="8BFF3683FF991801FCA9FA8432DCFABD" blockId="14.[835,1279,1309,1333]" box="[835,1279,1309,1333]" pageId="14" pageNumber="191">
<emphasis id="B934EA91FF991801FCA9FA8435F5FABD" bold="true" box="[835,982,1309,1333]" pageId="14" pageNumber="191">Distribution.</emphasis>
<collectingCountry id="F3577613FF991801FC34FA84326DFABD" box="[990,1102,1309,1333]" name="Singapore" pageId="14" pageNumber="191">Singapore</collectingCountry>
; 8 to 24 metres.
</paragraph>
</subSubSection>
<subSubSection id="C35A6508FF991801FCA9FAC432A7FA1D" pageId="14" pageNumber="191" type="etymology">
<paragraph id="8BFF3683FF991801FCA9FAC432A7FA1D" blockId="14.[834,1474,1373,1429]" pageId="14" pageNumber="191">
<emphasis id="B934EA91FF991801FCA9FAC435E7FAFD" bold="true" box="[835,964,1373,1397]" pageId="14" pageNumber="191">Etymology.</emphasis>
The epithet is the Latin word for shovel, alluding to the shovel-shaped rostrum.
</paragraph>
</subSubSection>
</treatment>
</document>
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