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<document id="574351DE5759F60EB76B8B62032ED4A0" ID-DOI="10.1111/j.1096-3642.2012.00863.x" ID-ISSN="0024-4082" ID-Zenodo-Dep="5409181" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630350074602" checkinUser="marcus" docAuthor="Criscione, Francesco, Law, Margot Louisa &amp; Köhler, Frank" docDate="2012" docId="03E9878DFFC9FF8772CAA1795FCFFA15" docLanguage="en" docName="zoj863.pdf" docOrigin="Zoological Journal of the Linnean Society (Zool. J. Linn. Soc.) 166 (4)" docSource="http://dx.doi.org/10.1111/j.1096-3642.2012.00863.x" docStyle="DocumentStyle:FBF0BF18BB2B256F1F8008F27D80BF37.3:ZoolJLinnSoc.2010-2016.journal_article" docStyleId="FBF0BF18BB2B256F1F8008F27D80BF37" docStyleName="ZoolJLinnSoc.2010-2016.journal_article" docStyleVersion="3" docTitle="Exiligada pallida Criscione &amp; Law &amp; Köhler 2012, SP. NOV." docType="treatment" docUuid="8C6E1292-4DEF-4FBC-B790-9EF4111D347D" docUuidSource="ZooBank" docVersion="5" lastPageNumber="708" masterDocId="FFD0FFF5FFDAFF93710AA04F5F3FFFCD" masterDocTitle="Land snail diversity in the monsoon tropics of Northern Australia: revision of the genus Exiligada Iredale, 1939 (Mollusca: Pulmonata: Camaenidae), with description of 13 new species" masterLastPageNumber="722" masterPageNumber="689" pageNumber="708" updateTime="1699241497941" updateUser="plazi">
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<mods:title id="937CD529A2C608A382602CE7E7724E34">Land snail diversity in the monsoon tropics of Northern Australia: revision of the genus Exiligada Iredale, 1939 (Mollusca: Pulmonata: Camaenidae), with description of 13 new species</mods:title>
</mods:titleInfo>
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<mods:namePart id="475EECEE0991DCD91FBD791E57D7643F">Criscione, Francesco</mods:namePart>
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<mods:namePart id="FD52D46EF69A70DD2D52FA810B6A989D">Law, Margot Louisa</mods:namePart>
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<mods:namePart id="AF6DF02C0AFCE44AC39DBCB85C995C1D">Köhler, Frank</mods:namePart>
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<mods:title id="9BE1702E68FA5EA6DB0BD4F146B4CC26">Zoological Journal of the Linnean Society</mods:title>
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<mods:date id="4F6F569C14EBB3F2F72D71CCFA97FEFA">2012</mods:date>
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<mods:title id="0A7D3457CA473C53A796F1DA4F60843A">Zool. J. Linn. Soc.</mods:title>
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<mods:number id="0FC60E1DE003C6187E974D7FD4B5CAAE">2012-11-26</mods:number>
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<mods:number id="FB2EE5BD2763F8716D6C709C1101FC87">166</mods:number>
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<mods:classification id="7134ADC7D3D438CA916F919036417006">journal article</mods:classification>
<mods:identifier id="4DE6BEB3ADD57D36E37BD439364F783C" type="DOI">10.1111/j.1096-3642.2012.00863.x</mods:identifier>
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<mods:identifier id="AB36DED97908A3945314F7B98484112D" type="Zenodo-Dep">5409181</mods:identifier>
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<treatment id="03E9878DFFC9FF8772CAA1795FCFFA15" LSID="urn:lsid:zoobank.org:act:8C6E1292-4DEF-4FBC-B790-9EF4111D347D" httpUri="http://treatment.plazi.org/id/03E9878DFFC9FF8772CAA1795FCFFA15" lastPageId="20" lastPageNumber="708" pageId="19" pageNumber="708">
<subSubSection id="C35A6510FFC9FF8072CAA1795A24FE87" box="[960,1307,309,331]" pageId="19" pageNumber="708" type="nomenclature">
<paragraph id="8BFF369BFFC9FF8072CAA1795A24FE87" blockId="19.[960,1307,309,331]" box="[960,1307,309,331]" pageId="19" pageNumber="708">
<heading id="D0B781F7FFC9FF8072CAA1795A24FE87" box="[960,1307,309,331]" centered="true" fontSize="9" level="2" pageId="19" pageNumber="708" reason="4">
<taxonomicName id="4C404D18FFC9FF8072CAA1795B93FE87" ID-CoL="3DL85" authority="Criscione &amp; Law &amp; Köhler, 2012" authorityName="Criscione &amp; Law &amp; Köhler" authorityYear="2012" box="[960,1196,310,331]" class="Gastropoda" family="Camaenidae" genus="Exiligada" kingdom="Animalia" order="Stylommatophora" pageId="19" pageNumber="708" phylum="Mollusca" rank="species" species="pallida" status="sp. nov.">EXILIGADA PALLIDA</taxonomicName>
<emphasis id="B934EA89FFC9FF8075BCA1765A24FE87" bold="true" box="[1206,1307,309,330]" pageId="19" pageNumber="708">
<taxonomicNameLabel id="A20757F2FFC9FF8075BCA1765A24FE87" box="[1206,1307,309,330]" pageId="19" pageNumber="708" rank="species">SP. NOV.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C35A6510FFC9FF8772D2A1135FCFFA15" lastPageId="20" lastPageNumber="709" pageId="19" pageNumber="708" type="description">
<paragraph id="8BFF369BFFC9FF8072D2A1135A3CFEBF" blockId="19.[984,1283,348,370]" box="[984,1283,348,370]" pageId="19" pageNumber="708">
(
<figureCitation id="137B2A1EFFC9FF8072EAA1135B00FEBC" box="[992,1087,348,369]" captionStart="Figure 4" captionStartId="8.[144,223,954,973]" captionTargetBox="[144,1425,196,924]" captionTargetId="figure-472@8.[144,1425,196,924]" captionTargetPageId="8" captionText="Figure 4. Photographs of living specimens of Exiligada species (courtesy Vince Kessner, Adelaide River). A, E. negriensis Iredale, 1939. B, E. pallida sp. nov. C, E. nodulicaudata sp. nov. D, E. punctata sp. nov. E, E. qualis Iredale, 1939. F, E. monochroma sp. nov. G, E. unistriata sp. nov. H, E. floraevallis sp. nov. I, E. rivifontis sp. nov." figureDoi="http://doi.org/10.5281/zenodo.5409193" httpUri="https://zenodo.org/record/5409193/files/figure.png" pageId="19" pageNumber="708">FIGS 4B</figureCitation>
,
<figureCitation id="137B2A1EFFC9FF807544A1135B4FFEBF" box="[1102,1136,348,370]" captionStart="Figure 6" captionStartId="9.[826,905,1233,1252]" captionTargetBox="[834,1435,197,1200]" captionTargetId="figure-294@9.[830,1439,194,1203]" captionTargetPageId="9" captionText="Figure 6. Shells of Exiligada species. A, E. montejinni sp. nov. (paratype AM C.469982). B, E. nodulicauda sp. nov. (paratype WAM S49160). C, E. pallida sp. nov. (paratype WAM S49215). D, E. punctata sp. nov. (paratype WAM S49221). E, E. qualis Iredale, 1939 (holotype AM C.64866). F, E. rivifontis sp. nov. (WAM S49225). G, E. unistriata sp. nov. (paratype AM C.470460). Scale bar = 2 mm. Note that foot produces from some shells." figureDoi="http://doi.org/10.5281/zenodo.5409195" httpUri="https://zenodo.org/record/5409195/files/figure.png" pageId="19" pageNumber="708">6C</figureCitation>
,
<figureCitation id="137B2A1EFFC9FF807575A1135BEEFEBC" box="[1151,1233,348,369]" captionStart="Figure 20" captionStartId="22.[144,223,1220,1239]" captionTargetBox="[303,1262,196,1190]" captionTargetId="figure-272@22.[303,1263,196,1190]" captionTargetPageId="22" captionText="Figure 20. SEM micrographs of Exiligada shells.A-C, E. pallida sp. nov. (paratype WAM S49150). A, Apical view showing protoconch and first whorl. B, Close-up from above of sculpture across entire shell. C, Microsculpture on teleoconch whorl. D-F, E. punctata sp. nov. (paratype WAMS49139). D, Apical view showing protoconch and first whorl. E, Close-up from above of sculpture across entire shell. F, Microsculpture on teleoconch whorl. Scale bars: A-B, D-E = 1 mm, C, F = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5409227" httpUri="https://zenodo.org/record/5409227/files/figure.png" pageId="19" pageNumber="708">20A–C</figureCitation>
,
<figureCitation id="137B2A1EFFC9FF8075D5A1135BC4FEBF" box="[1247,1275,348,370]" captionStart="Figure 21" captionStartId="23.[164,243,781,800]" captionTargetBox="[233,711,199,750]" captionTargetId="figure-476@23.[232,712,198,751]" captionTargetPageId="23" captionText="Figure 21. Penial anatomy of E. pallida sp. nov. (holotype WAM S83174). Scale bar = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.5409229" httpUri="https://zenodo.org/record/5409229/files/figure.png" pageId="19" pageNumber="708">21</figureCitation>
)
</paragraph>
<paragraph id="8BFF369BFFC9FF807230A1CD5A6BFDFF" blockId="19.[826,1441,386,562]" pageId="19" pageNumber="708">
<materialsCitation id="3B283CC6FFC9FF807230A1CD5A6BFDFF" accessionNumber="S83174" collectingDate="2009-08-30" collectionCode="NT, VRD, WAM" collectorName="Spring Creek Station H &amp; V. Kessner &amp; Kessner, R" country="Australia" elevation="18" latitude="-17.012222" location="Spring Creek Station" longLatPrecision="21" longitude="129.12639" pageId="19" pageNumber="708" specimenCode="WA-66" specimenCount="1" typeStatus="holotype">
<emphasis id="B934EA89FFC9FF807230A1CD5C9BFE5A" box="[826,932,386,407]" italics="true" pageId="19" pageNumber="708">
<typeStatus id="54FB8839FFC9FF807230A1CD5CA1FE5A" box="[826,926,386,407]" pageId="19" pageNumber="708" type="holotype">Holotype</typeStatus>
:
</emphasis>
<collectingCountry id="F357760BFFC9FF8072BEA1CC5B72FE55" box="[948,1101,387,408]" name="Australia" pageId="19" pageNumber="708">AUSTRALIA</collectingCountry>
,
<collectionCode id="ED51AE5EFFC9FF807555A1CC5BBBFE55" box="[1119,1156,387,408]" pageId="19" pageNumber="708">NT</collectionCode>
,
<collectionCode id="ED51AE5EFFC9FF80759DA1CC5BEEFE55" box="[1175,1233,387,408]" pageId="19" pageNumber="708">VRD</collectionCode>
,
<quantity id="4CB89B7EFFC9FF8075EEA1CC5A00FE55" box="[1252,1343,386,408]" metricMagnitude="4" metricUnit="m" metricValue="2.82" pageId="19" pageNumber="708" unit="km" value="28.2">28.2 km</quantity>
east of
<collectorName id="26B5534DFFC9FF807230A1EE5B5FFE7B" box="[826,1120,417,439]" pageId="19" pageNumber="708">
<location id="8E9F6040FFC9FF807230A1EE5B04FE7A" LSID="urn:lsid:plazi:treatment:03E9878DFFC9FF8772CAA1795FCFFA15:8E9F6040FFC9FF807230A1EE5B04FE7A" box="[826,1083,417,439]" country="Australia" latitude="-17.012222" longLatPrecision="21" longitude="129.12639" name="Spring Creek Station" pageId="19" pageNumber="708">Spring Creek Station</location>
H
</collectorName>
/S, base of limestone cliffs along a seasonal stream bed,
<quantity id="4CB89B7EFFC9FF807592A18F5BC6FE18" box="[1176,1273,448,470]" metricMagnitude="1" metricUnit="m" metricValue="1.75" metricValueMax="2.0" metricValueMin="1.5" pageId="19" pageNumber="708" unit="m" value="17.5" valueMax="20.0" valueMin="15.0">
<elevation id="006DD1A8FFC9FF807592A18F5BC6FE18" box="[1176,1273,448,470]" metricMagnitude="1" metricUnit="m" metricValue="1.75" metricValueMax="2.0" metricValueMin="1.5" pageId="19" pageNumber="708" unit="m" value="17.5" valueMax="20.0" valueMin="15.0">15–20 m</elevation>
</quantity>
or more high, large overhangs, talus and boulders, under rocks,
<geoCoordinate id="EE74505CFFC9FF807230A1B25C86FDDF" box="[826,953,508,531]" degrees="17" direction="south" minutes="00" orientation="latitude" pageId="19" pageNumber="708" precision="15" seconds="44" value="-17.012222">
17°00
<emphasis id="B934EA89FFC9FF807271A1B35CBEFDDE" box="[891,897,508,531]" italics="true" pageId="19" pageNumber="708">′</emphasis>
44
<emphasis id="B934EA89FFC9FF807296A1B35C97FDDE" box="[924,936,508,531]" italics="true" pageId="19" pageNumber="708">′′</emphasis>
S
</geoCoordinate>
,
<geoCoordinate id="EE74505CFFC9FF8072C7A1B25B64FDDF" box="[973,1115,508,531]" degrees="129" direction="east" minutes="07" orientation="longitude" pageId="19" pageNumber="708" precision="15" seconds="35" value="129.12639">
129°07
<emphasis id="B934EA89FFC9FF807516A1B35B1DFDDE" box="[1052,1058,508,531]" italics="true" pageId="19" pageNumber="708">′</emphasis>
35
<emphasis id="B934EA89FFC9FF807537A1B35B76FDDE" box="[1085,1097,508,531]" italics="true" pageId="19" pageNumber="708">′′</emphasis>
E
</geoCoordinate>
(coll.
<collectorName id="26B5534DFFC9FF8075A4A1B25A10FDDF" box="[1198,1327,509,530]" pageId="19" pageNumber="708">V. Kessner</collectorName>
, R. Hokkanen,
<date id="FFFE105BFFC9FF80729AA2545B28FDFC" box="[912,1047,539,561]" pageId="19" pageNumber="708" value="2009-08-30">
<collectingDate id="EFBAE9B3FFC9FF80729AA2545B28FDFC" box="[912,1047,539,561]" pageId="19" pageNumber="708" value="2009-08-30">30.viii.2009</collectingDate>
</date>
;
<specimenCode id="DBE69EE0FFC9FF807529A2545B4EFDFC" box="[1059,1137,539,561]" collectionCode="WA-" pageId="19" pageNumber="708">WA-66</specimenCode>
/09) (
<collectionCode id="ED51AE5EFFC9FF8075A2A2535BD6FDFC" box="[1192,1257,540,561]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" lsid="urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="19" pageNumber="708" type="Museum">WAM</collectionCode>
<accessionNumber id="9413AB78FFC9FF8075FAA2535A77FDFF" box="[1264,1352,540,562]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/S83174" pageId="19" pageNumber="708" type="EnaNcbi">S83174</accessionNumber>
).
</materialsCitation>
</paragraph>
<paragraph id="8BFF369BFFC9FF807230A22F5CA1FD7F" blockId="19.[826,1441,607,690]" pageId="19" pageNumber="708">
<materialsCitation id="3B283CC6FFC9FF807230A22F5BF0FDB8" box="[826,1231,607,629]" collectionCode="NT, VRD" country="Australia" location="AUSTRALIA" pageId="19" pageNumber="708" specimenCount="1" typeStatus="paratype">
<emphasis id="B934EA89FFC9FF807230A22F5C8DFDB8" box="[826,946,608,629]" italics="true" pageId="19" pageNumber="708">
<typeStatus id="54FB8839FFC9FF807230A22F5C94FDB8" box="[826,939,608,629]" pageId="19" pageNumber="708" type="paratype">Paratypes</typeStatus>
:
</emphasis>
<collectingCountry id="F357760BFFC9FF8072B6A2105B6AFDB9" box="[956,1109,607,628]" name="Australia" pageId="19" pageNumber="708">AUSTRALIA</collectingCountry>
,
<collectionCode id="ED51AE5EFFC9FF807568A22F5BB8FDB8" box="[1122,1159,608,629]" pageId="19" pageNumber="708">NT</collectionCode>
,
<collectionCode id="ED51AE5EFFC9FF80759FA22F5BF0FDB8" box="[1173,1231,608,629]" pageId="19" pageNumber="708">VRD</collectionCode>
</materialsCitation>
,
<materialsCitation id="3B283CC6FFC9FF8075D6A22F5CA5FD7F" accessionNumber="S49215, S49150" collectingDate="2009-08-30" collectionCode="WAM" collectorName="Spring Creek Station H &amp; V. Kessner &amp; Kessner, R" country="Australia" elevation="18" latitude="-17.012222" location="Spring Creek Station" longLatPrecision="21" longitude="129.12639" pageId="19" pageNumber="708" specimenCount="1" typeStatus="paratype">
same data as for holotype, 22 wet (
<collectionCode id="ED51AE5EFFC9FF80752BA2315B5DFD5E" box="[1057,1122,638,659]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" lsid="urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="19" pageNumber="708" type="Museum">WAM</collectionCode>
S49215), 106 dry (
<collectionCode id="ED51AE5EFFC9FF80746AA2315A9EFD5E" box="[1376,1441,638,659]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" lsid="urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="19" pageNumber="708" type="Museum">WAM</collectionCode>
<accessionNumber id="9413AB78FFC9FF807230A2D35CADFD7F" box="[826,914,668,690]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/S49150" pageId="19" pageNumber="708" type="EnaNcbi">S49150</accessionNumber>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BFF369BFFC9FF807230A2AF5B25FCFE" blockId="19.[826,1441,736,819]" pageId="19" pageNumber="708">
<emphasis id="B934EA89FFC9FF807230A2AF5C87FD38" box="[826,952,736,757]" italics="true" pageId="19" pageNumber="708">Etymology:</emphasis>
In reference to the pale shell background colour, derived from ‘pallidus’ (Latin = pale); adjective of feminine gender.
</paragraph>
<caption id="DF3F6613FFC9FF8071AEA3F05D4DFC3D" ID-DOI="http://doi.org/10.5281/zenodo.5409217" ID-Zenodo-Dep="5409217" httpUri="https://zenodo.org/record/5409217/files/figure.png" pageId="19" pageNumber="708" startId="19.[164,243,959,978]" targetBox="[200,743,195,929]" targetPageId="19">
<paragraph id="8BFF369BFFC9FF8071AEA3F05D4DFC3D" blockId="19.[164,779,959,1008]" pageId="19" pageNumber="708">
<emphasis id="B934EA89FFC9FF8071AEA3F05E26FC1F" bold="true" box="[164,281,959,978]" pageId="19" pageNumber="708">Figure 16.</emphasis>
Penial anatomy of
<emphasis id="B934EA89FFC9FF8070FBA38F5C34FC1F" bold="true" box="[497,779,959,979]" pageId="19" pageNumber="708">
<taxonomicName id="4C404D18FFC9FF8070FBA38F5D95FC1F" authority="Criscione &amp; Law &amp; Köhler, 2012" authorityName="Criscione &amp; Law &amp; Köhler" authorityYear="2012" box="[497,682,959,979]" class="Gastropoda" family="Camaenidae" genus="Exiligada" kingdom="Animalia" order="Stylommatophora" pageId="19" pageNumber="708" phylum="Mollusca" rank="species" species="monochroma" status="sp. nov.">
<emphasis id="B934EA89FFC9FF8070FBA38F5D95FC1F" bold="true" box="[497,682,959,979]" italics="true" pageId="19" pageNumber="708">E. monochroma</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20757F2FFC9FF8073B8A3F05C34FC1F" box="[690,779,959,978]" pageId="19" pageNumber="708" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(holotype AM C.475768). Scale bar = 5 mm.
</paragraph>
</caption>
<paragraph id="8BFF369BFFC9FF8071B6A4605DE0FAF7" blockId="19.[164,779,1071,1338]" pageId="19" pageNumber="708">
Genitalia (
<figureCitation id="137B2A1EFFC9FF807034A4605EB0FB88" box="[318,399,1071,1093]" captionStart="Figure 19" captionStartId="21.[164,243,1789,1808]" captionTargetBox="[267,679,993,1757]" captionTargetId="figure-418@21.[264,681,991,1758]" captionTargetPageId="21" captionText="Figure 19. Penial anatomy of E. nodulicauda sp. nov. (holotype WAM S83173). Scale bar = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.5409225" httpUri="https://zenodo.org/record/5409225/files/figure.png" pageId="19" pageNumber="708">Fig. 19</figureCitation>
). Epiphallus length two thirds that of penis. Vas deferens entering penial sheath from one third up. Penis seven times longer than wide, coiled inside sheath, with posterior part of inner wall supporting three regularly nodulose, longitudinal pilasters fading into three irregularly shaped folds not reaching the genital pore. Vagina as long as penis, two times longer than free oviduct, bursa copulatrix with long, rounded well-differentiated end.
</paragraph>
<paragraph id="8BFF369BFFC9FF8071AEA5205A9EFB80" blockId="19.[164,779,1391,1903]" lastBlockId="19.[826,1441,865,1437]" pageId="19" pageNumber="708">
<emphasis id="B934EA89FFC9FF8071AEA5205E30FA49" box="[164,271,1391,1412]" italics="true" pageId="19" pageNumber="708">Remarks:</emphasis>
Together with
<taxonomicName id="4C404D18FFC9FF8070C3A53F5D11FA49" authorityName="Iredale" authorityYear="1939" box="[457,558,1391,1413]" class="Gastropoda" family="Camaenidae" genus="Exiligada" kingdom="Animalia" order="Stylommatophora" pageId="19" pageNumber="708" phylum="Mollusca" rank="species" species="qualis">
<emphasis id="B934EA89FFC9FF8070C3A53F5D11FA49" box="[457,558,1391,1413]" italics="true" pageId="19" pageNumber="708">E. qualis</emphasis>
</taxonomicName>
narrowest umbilicus of genus. Shell colour pattern very similar to shells of four congeners; to be distinguished from
<taxonomicName id="4C404D18FFC9FF8071AEA5835E7EFA2D" authorityName="Criscione &amp; Law &amp; Köhler" authorityYear="2012" box="[164,321,1483,1505]" class="Gastropoda" family="Camaenidae" genus="Exiligada" kingdom="Animalia" order="Stylommatophora" pageId="19" pageNumber="708" phylum="Mollusca" rank="species" species="longicauda">
<emphasis id="B934EA89FFC9FF8071AEA5835E7EFA2D" box="[164,321,1483,1505]" italics="true" pageId="19" pageNumber="708">E. longicauda</emphasis>
</taxonomicName>
by larger number of spiral bands, from
<taxonomicName id="4C404D18FFC9FF8071AEA5A55E0AFA32" authorityName="Iredale" authorityYear="1939" box="[164,309,1514,1535]" class="Gastropoda" family="Camaenidae" genus="Exiligada" kingdom="Animalia" order="Stylommatophora" pageId="19" pageNumber="708" phylum="Mollusca" rank="species" species="negriensis">
<emphasis id="B934EA89FFC9FF8071AEA5A55E0AFA32" box="[164,309,1514,1535]" italics="true" pageId="19" pageNumber="708">E. negriensis</emphasis>
</taxonomicName>
by more regularly interrupted bands, from
<taxonomicName id="4C404D18FFC9FF8071E1A6465E62F9D0" authorityName="Criscione &amp; Law &amp; Köhler" authorityYear="2012" box="[235,349,1544,1566]" class="Gastropoda" family="Camaenidae" genus="Exiligada" kingdom="Animalia" order="Stylommatophora" pageId="19" pageNumber="708" phylum="Mollusca" rank="species" species="pallida">
<emphasis id="B934EA89FFC9FF8071E1A6465E62F9D0" box="[235,349,1544,1566]" italics="true" pageId="19" pageNumber="708">E. pallida</emphasis>
</taxonomicName>
by darker background, and from
<taxonomicName id="4C404D18FFC9FF8071AEA6685E17F9F1" authorityName="Criscione &amp; Law &amp; Köhler" authorityYear="2012" box="[164,296,1575,1596]" class="Gastropoda" family="Camaenidae" genus="Exiligada" kingdom="Animalia" order="Stylommatophora" pageId="19" pageNumber="708" phylum="Mollusca" rank="species" species="punctata">
<emphasis id="B934EA89FFC9FF8071AEA6685E17F9F1" box="[164,296,1575,1596]" italics="true" pageId="19" pageNumber="708">E. punctata</emphasis>
</taxonomicName>
by absence of maculations. Along with
<taxonomicName id="4C404D18FFC9FF8071AEA6095E7EF997" authorityName="Criscione &amp; Law &amp; Köhler" authorityYear="2012" box="[164,321,1605,1627]" class="Gastropoda" family="Camaenidae" genus="Exiligada" kingdom="Animalia" order="Stylommatophora" pageId="19" pageNumber="708" phylum="Mollusca" rank="species" species="longicauda">
<emphasis id="B934EA89FFC9FF8071AEA6095E7EF997" box="[164,321,1605,1627]" italics="true" pageId="19" pageNumber="708">E. longicauda</emphasis>
</taxonomicName>
, only species in genus with a coiled penis. Penis length relative to width (pl/pw) larger than average and smaller than
<taxonomicName id="4C404D18FFC9FF80736CA6CC5C3CF955" authorityName="Criscione &amp; Law &amp; Köhler" authorityYear="2012" box="[614,771,1667,1688]" class="Gastropoda" family="Camaenidae" genus="Exiligada" kingdom="Animalia" order="Stylommatophora" pageId="19" pageNumber="708" phylum="Mollusca" rank="species" species="longicauda">
<emphasis id="B934EA89FFC9FF80736CA6CC5C3CF955" box="[614,771,1667,1688]" italics="true" pageId="19" pageNumber="708">E. longicauda</emphasis>
</taxonomicName>
; epiphallus length relative to penis (el/pl) smaller than average and two times that of
<taxonomicName id="4C404D18FFC9FF807311A68F5D87F918" authorityName="Criscione &amp; Law &amp; Köhler" authorityYear="2012" box="[539,696,1728,1749]" class="Gastropoda" family="Camaenidae" genus="Exiligada" kingdom="Animalia" order="Stylommatophora" pageId="19" pageNumber="708" phylum="Mollusca" rank="species" species="longicauda">
<emphasis id="B934EA89FFC9FF807311A68F5D87F918" box="[539,696,1728,1749]" italics="true" pageId="19" pageNumber="708">E. longicauda</emphasis>
</taxonomicName>
. Inner penial wall sculpture unique, in consisting of longitudinal pilasters sculptured by regularly spaced and shaped nodules. Vagina length relative to penis (vl/pl) larger than average and almost as large as in
<taxonomicName id="4C404D18FFC9FF8073B5A7745FC0F8A3" authorityName="Criscione &amp; Law &amp; Köhler" authorityYear="2012" class="Gastropoda" family="Camaenidae" genus="Exiligada" kingdom="Animalia" order="Stylommatophora" pageId="19" pageNumber="708" phylum="Mollusca" rank="species" species="longicauda">
<emphasis id="B934EA89FFC9FF8073B5A7745FC0F8A3" italics="true" pageId="19" pageNumber="708">E. longicauda</emphasis>
</taxonomicName>
; vagina length relative to free oviduct (vl/ol)
<emphasis id="B934EA89FFC9FF807230A32E5CFDFCBB" box="[826,962,865,886]" italics="true" pageId="19" pageNumber="708">Description:</emphasis>
Shell (
<figureCitation id="137B2A1EFFC9FF807512A32E5B4DFCBB" box="[1048,1138,865,887]" captionStart="Figure 4" captionStartId="8.[144,223,954,973]" captionTargetBox="[144,1425,196,924]" captionTargetId="figure-472@8.[144,1425,196,924]" captionTargetPageId="8" captionText="Figure 4. Photographs of living specimens of Exiligada species (courtesy Vince Kessner, Adelaide River). A, E. negriensis Iredale, 1939. B, E. pallida sp. nov. C, E. nodulicaudata sp. nov. D, E. punctata sp. nov. E, E. qualis Iredale, 1939. F, E. monochroma sp. nov. G, E. unistriata sp. nov. H, E. floraevallis sp. nov. I, E. rivifontis sp. nov." figureDoi="http://doi.org/10.5281/zenodo.5409193" httpUri="https://zenodo.org/record/5409193/files/figure.png" pageId="19" pageNumber="708">Figs 4B</figureCitation>
,
<figureCitation id="137B2A1EFFC9FF80758AA32E5B9DFCBA" box="[1152,1186,865,887]" captionStart="Figure 6" captionStartId="9.[826,905,1233,1252]" captionTargetBox="[834,1435,197,1200]" captionTargetId="figure-294@9.[830,1439,194,1203]" captionTargetPageId="9" captionText="Figure 6. Shells of Exiligada species. A, E. montejinni sp. nov. (paratype AM C.469982). B, E. nodulicauda sp. nov. (paratype WAM S49160). C, E. pallida sp. nov. (paratype WAM S49215). D, E. punctata sp. nov. (paratype WAM S49221). E, E. qualis Iredale, 1939 (holotype AM C.64866). F, E. rivifontis sp. nov. (WAM S49225). G, E. unistriata sp. nov. (paratype AM C.470460). Scale bar = 2 mm. Note that foot produces from some shells." figureDoi="http://doi.org/10.5281/zenodo.5409195" httpUri="https://zenodo.org/record/5409195/files/figure.png" pageId="19" pageNumber="708">6C</figureCitation>
,
<figureCitation id="137B2A1EFFC9FF8075BAA32E5A3EFCBA" box="[1200,1281,865,887]" captionStart="Figure 20" captionStartId="22.[144,223,1220,1239]" captionTargetBox="[303,1262,196,1190]" captionTargetId="figure-272@22.[303,1263,196,1190]" captionTargetPageId="22" captionText="Figure 20. SEM micrographs of Exiligada shells.A-C, E. pallida sp. nov. (paratype WAM S49150). A, Apical view showing protoconch and first whorl. B, Close-up from above of sculpture across entire shell. C, Microsculpture on teleoconch whorl. D-F, E. punctata sp. nov. (paratype WAMS49139). D, Apical view showing protoconch and first whorl. E, Close-up from above of sculpture across entire shell. F, Microsculpture on teleoconch whorl. Scale bars: A-B, D-E = 1 mm, C, F = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5409227" httpUri="https://zenodo.org/record/5409227/files/figure.png" pageId="19" pageNumber="708">20A–C</figureCitation>
). Large, moderately elevated, moderately thick; micropustulations present. Shell background colour pale horn fading to whitish towards shell base, with several conspicuous to faint brown-reddish spiral bands, equally conspicuous on entire shell, often present on shell base, often regularly interrupted radially and giving the appearance of a regular radial bandage; maculations absent.
</paragraph>
<paragraph id="8BFF369BFFC9FF807259A41A5C54FA50" blockId="19.[826,1441,865,1437]" pageId="19" pageNumber="708">
Genitalia (
<figureCitation id="137B2A1EFFC9FF8072DFA41A5B19FBA1" box="[981,1062,1109,1132]" captionStart="Figure 21" captionStartId="23.[164,243,781,800]" captionTargetBox="[233,711,199,750]" captionTargetId="figure-476@23.[232,712,198,751]" captionTargetPageId="23" captionText="Figure 21. Penial anatomy of E. pallida sp. nov. (holotype WAM S83174). Scale bar = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.5409229" httpUri="https://zenodo.org/record/5409229/files/figure.png" pageId="19" pageNumber="708">Fig. 21</figureCitation>
). Epiphallus as long as penis. Vas deferens entering penial sheath from halfway up. Penis six times longer than wide, not coiled inside sheath, with inner wall supporting a posterior coneshaped pad and five longitudinal pilasters, two arising close to posterior part of pad and reaching genital pore, three from posterior end of penis, only two of which reaching genital pore. Vagina one third shorter than penis length and as long as free oviduct, bursa copulatrix with long, pointed well-differentiated end.
</paragraph>
<paragraph id="8BFF369BFFC9FF807230A5845A9DF9CD" blockId="19.[826,1442,1483,1903]" pageId="19" pageNumber="708">
<emphasis id="B934EA89FFC9FF807230A5845C99FA2D" box="[826,934,1483,1504]" italics="true" pageId="19" pageNumber="708">Remarks:</emphasis>
Teleoconch microsculpture differing from that of
<taxonomicName id="4C404D18FFC9FF807286A5A55B21FA32" authorityName="Iredale" authorityYear="1939" box="[908,1054,1514,1535]" class="Gastropoda" family="Camaenidae" genus="Exiligada" kingdom="Animalia" order="Stylommatophora" pageId="19" pageNumber="708" phylum="Mollusca" rank="species" species="negriensis">
<emphasis id="B934EA89FFC9FF807286A5A55B21FA32" box="[908,1054,1514,1535]" italics="true" pageId="19" pageNumber="708">E. negriensis</emphasis>
</taxonomicName>
by presence of micropustulations.
</paragraph>
<paragraph id="8BFF369BFFC9FF807259A6475A9DF9B7" blockId="19.[826,1442,1483,1903]" pageId="19" pageNumber="708">
Shell colour pattern similar to that of four other
<taxonomicName id="4C404D18FFC9FF807230A6685B38F9F0" box="[826,1031,1575,1597]" class="Gastropoda" family="Camaenidae" genus="Exiligada" kingdom="Animalia" order="Stylommatophora" pageId="19" pageNumber="708" phylum="Mollusca" rank="species" species="undetermined">
<emphasis id="B934EA89FFC9FF807230A6685C96F9F1" box="[826,937,1575,1596]" italics="true" pageId="19" pageNumber="708">Exiligada</emphasis>
species
</taxonomicName>
; to be distinguished from
<taxonomicName id="4C404D18FFC9FF80744AA6685C40F997" authorityName="Criscione &amp; Law &amp; Köhler" authorityYear="2012" class="Gastropoda" family="Camaenidae" genus="Exiligada" kingdom="Animalia" order="Stylommatophora" pageId="19" pageNumber="708" phylum="Mollusca" rank="species" species="longicauda">
<emphasis id="B934EA89FFC9FF80744AA6685C40F997" italics="true" pageId="19" pageNumber="708">E. longicauda</emphasis>
</taxonomicName>
and
<taxonomicName id="4C404D18FFC9FF8072B5A6095B51F997" authorityName="Criscione &amp; Law &amp; Köhler" authorityYear="2012" box="[959,1134,1605,1627]" class="Gastropoda" family="Camaenidae" genus="Exiligada" kingdom="Animalia" order="Stylommatophora" pageId="19" pageNumber="708" phylum="Mollusca" rank="species" species="nodulicauda">
<emphasis id="B934EA89FFC9FF8072B5A6095B51F997" box="[959,1134,1605,1627]" italics="true" pageId="19" pageNumber="708">E. nodulicauda</emphasis>
</taxonomicName>
by its lighter background colour and from
<taxonomicName id="4C404D18FFC9FF8072E4A62A5B4CF9B7" authorityName="Criscione &amp; Law &amp; Köhler" authorityYear="2012" box="[1006,1139,1637,1658]" class="Gastropoda" family="Camaenidae" genus="Exiligada" kingdom="Animalia" order="Stylommatophora" pageId="19" pageNumber="708" phylum="Mollusca" rank="species" species="punctata">
<emphasis id="B934EA89FFC9FF8072E4A62A5B4CF9B7" box="[1006,1139,1637,1658]" italics="true" pageId="19" pageNumber="708">E. punctata</emphasis>
</taxonomicName>
by absence of maculations.
</paragraph>
<paragraph id="8BFF369BFFC9FF877259A6CC5FCFFA15" blockId="19.[826,1442,1483,1903]" lastBlockId="20.[144,759,1413,1496]" lastPageId="20" lastPageNumber="709" pageId="19" pageNumber="708">
Penis length relative to width (pl/pw) average and larger than in sister species
<taxonomicName id="4C404D18FFC9FF807402A6ED5AA5F97A" authorityName="Iredale" authorityYear="1939" box="[1288,1434,1698,1719]" class="Gastropoda" family="Camaenidae" genus="Exiligada" kingdom="Animalia" order="Stylommatophora" pageId="19" pageNumber="708" phylum="Mollusca" rank="species" species="negriensis">
<emphasis id="B934EA89FFC9FF807402A6ED5AA5F97A" box="[1288,1434,1698,1719]" italics="true" pageId="19" pageNumber="708">E. negriensis</emphasis>
</taxonomicName>
; epiphallus length relative to penis (el/pl) as long as average and as
<taxonomicName id="4C404D18FFC9FF807522A6905B85F939" authorityName="Iredale" authorityYear="1939" box="[1064,1210,1759,1780]" class="Gastropoda" family="Camaenidae" genus="Exiligada" kingdom="Animalia" order="Stylommatophora" pageId="19" pageNumber="708" phylum="Mollusca" rank="species" species="negriensis">
<emphasis id="B934EA89FFC9FF807522A6905B85F939" box="[1064,1210,1759,1780]" italics="true" pageId="19" pageNumber="708">E. negriensis</emphasis>
</taxonomicName>
. Inner penial wall sculpture similar to
<taxonomicName id="4C404D18FFC9FF807531A6B15BF3F8DE" authorityName="Iredale" authorityYear="1939" box="[1083,1228,1790,1811]" class="Gastropoda" family="Camaenidae" genus="Exiligada" kingdom="Animalia" order="Stylommatophora" pageId="19" pageNumber="708" phylum="Mollusca" rank="species" species="negriensis">
<emphasis id="B934EA89FFC9FF807531A6B15BF3F8DE" box="[1083,1228,1790,1811]" italics="true" pageId="19" pageNumber="708">E. negriensis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C404D18FFC9FF80741CA6B15AA5F8DE" authorityName="Criscione &amp; Law &amp; Köhler" authorityYear="2012" box="[1302,1434,1790,1811]" class="Gastropoda" family="Camaenidae" genus="Exiligada" kingdom="Animalia" order="Stylommatophora" pageId="19" pageNumber="708" phylum="Mollusca" rank="species" species="punctata">
<emphasis id="B934EA89FFC9FF80741CA6B15AA5F8DE" box="[1302,1434,1790,1811]" italics="true" pageId="19" pageNumber="708">E. punctata</emphasis>
</taxonomicName>
, to be distinguished from these species by narrower posterior pad and longer longitudinal pilasters, always reaching the posterior part of the wall, and by noncorrugated posterior pad. Vagina length relative to penis (vl/pl) and relative to free oviduct (vl/ol) average.
</paragraph>
</subSubSection>
</treatment>
</document>
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