410 lines
49 KiB
XML
410 lines
49 KiB
XML
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<mods:title id="F8A75C396BDB900CDB9EFE8688CF18F5">Osteology of the Permian temnospondyl amphibian Glanochthon lellbachae and its relationships</mods:title>
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<mods:namePart id="4039D39CFD4AF0C2D0A775FAD39A4A25">Schoch Correspondence, Rainer R.</mods:namePart>
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<mods:title id="29C91197D8358DA534AFEF49AB44B7E4">Fossil Record</mods:title>
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<mods:date id="16EBE77EADC3117C07E09655E8999AA2">2021</mods:date>
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<mods:number id="BB1FBE1FBE1F0022749BF6A3FFF3F389">2021-03-23</mods:number>
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<treatment id="03E98784FFF6FFD5FF5841DDFAA6FC90" ID-DOI="http://doi.org/10.5281/zenodo.11618626" ID-GBIF-Taxon="232733555" ID-Zenodo-Dep="11618626" LSID="urn:lsid:plazi:treatment:03E98784FFF6FFD5FF5841DDFAA6FC90" httpUri="http://treatment.plazi.org/id/03E98784FFF6FFD5FF5841DDFAA6FC90" lastPageId="6" lastPageNumber="55" pageId="2" pageNumber="51">
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<subSubSection id="C35A6519FFF6FFD1FF5841DDFD14F972" box="[177,642,1664,1688]" pageId="2" pageNumber="51" type="nomenclature">
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<paragraph id="8BFF3692FFF6FFD1FF5841DDFD14F972" blockId="2.[177,642,1664,1688]" box="[177,642,1664,1688]" pageId="2" pageNumber="51">
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<heading id="D0B781FEFFF6FFD1FF5841DDFD14F972" box="[177,642,1664,1688]" fontSize="10" level="2" pageId="2" pageNumber="51" reason="7">
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<taxonomicName id="4C404D11FFF6FFD1FF5841DDFD14F972" authority="Kratschmer, 2006" authorityName="Kratschmer" authorityYear="2006" box="[177,642,1664,1688]" class="Bivalvia" family="Actinodontidae" genus="Cheliderpeton" higherTaxonomySource="GBIF" kingdom="Animalia" order="Actinodontida" pageId="2" pageNumber="51" phylum="Mollusca" rank="species" species="lellbachae">
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<emphasis id="B934EA80FFF6FFD1FF5841DDFE28F97D" box="[177,446,1664,1688]" italics="true" pageId="2" pageNumber="51">Cheliderpeton lellbachae</emphasis>
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<bibRefCitation id="EFD14B63FFF6FFD1FE2C41DDFD14F972" author="Kratschmer, K." box="[453,642,1664,1687]" pageId="2" pageNumber="51" pagination="3 - 46" refId="ref9586" refString="Kratschmer, K.: Neue temnospondyle Amphibien aus dem Rotliegend des sudwestdeutschen Saar-Nahe-Becken. Teil 1, Geowiss. Beitr. saarpfalz. Rotl., 4, 3 - 46, 2006." type="journal article" year="2006">Krätschmer, 2006</bibRefCitation>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C35A6519FFF6FFD1FF954189FCF8FDA3" pageId="2" pageNumber="51" type="materials_examined">
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<paragraph id="8BFF3692FFF6FFD1FF954189FF75F909" blockId="2.[124,227,1748,1772]" box="[124,227,1748,1772]" pageId="2" pageNumber="51">
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<emphasis id="B934EA80FFF6FFD1FF954189FF75F909" bold="true" box="[124,227,1748,1772]" pageId="2" pageNumber="51">
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<typeStatus id="54FB8830FFF6FFD1FF954189FF75F909" box="[124,227,1748,1772]" pageId="2" pageNumber="51" type="holotype">Holotype</typeStatus>
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</emphasis>
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</paragraph>
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<paragraph id="8BFF3692FFF6FFD1FF95404DFED8F8A3" blockId="2.[124,766,1808,1862]" pageId="2" pageNumber="51">
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<materialsCitation id="3B283CCFFFF6FFD1FF95404DFED8F8A3" ID-GBIF-Occurrence="4876372301" collectionCode="NHMM" pageId="2" pageNumber="51" specimenCode="NHMM 2006" specimenCount="1" typeStatus="holotype">
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<specimenCode id="DBE69EE9FFF6FFD1FF95404DFE85F8C2" box="[124,275,1808,1831]" collectionCode="NHMM" country="Netherlands" httpUri="http://grbio.org/cool/6t80-4rpg" name="Natuurhistorische Museum Maastricht" pageId="2" pageNumber="51">NHMM 2006</specimenCode>
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/
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<quantity id="4CB89B77FFF6FFD1FEF2404DFE00F8C2" box="[283,406,1808,1831]" metricMagnitude="-2" metricUnit="m" metricValue="1.4102" pageId="2" pageNumber="51" unit="mm" value="14.102">14, 102 mm</quantity>
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SL, complete skeleton with skin impression (
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<figureCitation id="137B2A17FFF6FFD1FF164072FED7F8A3" box="[255,321,1839,1862]" captionStart="Figure 2" captionStartId="3.[124,191,1842,1864]" captionTargetBox="[235,1359,200,1796]" captionTargetId="figure-12@3.[226,1360,199,1810]" captionTargetPageId="3" captionText="Figure 2. Type specimen of Glanochthon lellbachae (Krätschmer, 2006) comb. nov. (NHMM 2006/14). (a, c) Close-up of skull roof and (b) complete skeleton." figureDoi="http://doi.org/10.5281/zenodo.11359845" httpUri="https://zenodo.org/record/11359845/files/figure.png" pageId="2" pageNumber="51">Fig. 2</figureCitation>
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).
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</materialsCitation>
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</paragraph>
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<paragraph id="8BFF3692FFF6FFD1FCDA47F2FBA9FF22" blockId="2.[819,1087,175,199]" box="[819,1087,175,199]" pageId="2" pageNumber="51">
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<emphasis id="B934EA80FFF6FFD1FCDA47F2FBA9FF22" bold="true" box="[819,1087,175,199]" pageId="2" pageNumber="51">Type of locality and age</emphasis>
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</paragraph>
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<paragraph id="8BFF3692FFF6FFD1FCDA47ADFA21FE83" blockId="2.[819,1463,240,358]" pageId="2" pageNumber="51">
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The Klauswald southwest of Odernheim am Glan,
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<collectingRegion id="4984F870FFF6FFD1FCDA4652FB82FEC3" box="[819,1044,271,294]" country="Germany" name="Rheinland-Pfalz" pageId="2" pageNumber="51">Rhineland-Palatinate</collectingRegion>
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,
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<collectingCountry id="F3577602FFF6FFD1FBC24652FB1BFEC3" box="[1067,1165,271,294]" name="Germany" pageId="2" pageNumber="51">Germany</collectingCountry>
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(
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<figureCitation id="137B2A17FFF6FFD1FB404652FB65FEC3" box="[1193,1267,271,294]" captionStart="Figure 1" captionStartId="1.[124,191,1522,1544]" captionTargetBox="[358,1227,177,1491]" captionTargetId="figure-12@1.[358,1227,177,1491]" captionTargetPageId="1" captionText="Figure 1. Localities yielding Sclerocephalus and Glanochthon material and stratigraphy of the Rotliegend sequence with emphasis on the Meisenheim Formation (Autunian, Lower Permian) in the Saar–Nahe Basin of Germany." figureDoi="http://doi.org/10.5281/zenodo.11359843" httpUri="https://zenodo.org/record/11359843/files/figure.png" pageId="2" pageNumber="51">Fig. 1</figureCitation>
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). M9 sequence, Klauswald facies, Odernheim Subformation, Meisenheim Formation, lower Rotliegend, Autunian, lowermost Permian.
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</paragraph>
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<paragraph id="8BFF3692FFF6FFD1FCDA46D2FC69FE42" blockId="2.[819,1023,399,423]" box="[819,1023,399,423]" pageId="2" pageNumber="51">
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<emphasis id="B934EA80FFF6FFD1FCDA46D2FC69FE42" bold="true" box="[819,1023,399,423]" pageId="2" pageNumber="51">Referred material</emphasis>
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</paragraph>
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<paragraph id="8BFF3692FFF6FFD1FCDA4692FCF8FDA3" blockId="2.[819,1462,463,582]" pageId="2" pageNumber="51">
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Altogether, five additional specimens are referred to as
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<taxonomicName id="4C404D11FFF6FFD1FCDA46B3FBAFFDE3" authorityName="Schoch Correspondence" authorityYear="2021" baseAuthorityName="Kratschmer" baseAuthorityYear="2006" box="[819,1081,494,518]" class="Amphibia" genus="Glanochthon" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="2" pageNumber="51" phylum="Chordata" rank="species" species="lellbachae">
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<emphasis id="B934EA80FFF6FFD1FCDA46B3FBAFFDE3" box="[819,1081,494,518]" italics="true" pageId="2" pageNumber="51">Glanochthon lellbachae</emphasis>
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</taxonomicName>
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(see material section). There remains a substantial number of specimens in private collections.
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</paragraph>
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</subSubSection>
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<subSubSection id="C35A6519FFF6FFD5FCDA4533FA20FF67" lastPageId="6" lastPageNumber="55" pageId="2" pageNumber="51" type="diagnosis">
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<paragraph id="8BFF3692FFF6FFD1FCDA4533FC36FD63" blockId="2.[819,928,622,646]" box="[819,928,622,646]" pageId="2" pageNumber="51">
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<emphasis id="B934EA80FFF6FFD1FCDA4533FC36FD63" bold="true" box="[819,928,622,646]" pageId="2" pageNumber="51">Diagnosis</emphasis>
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</paragraph>
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<paragraph id="8BFF3692FFF6FFD1FCDA45F3FA17FC01" blockId="2.[819,1462,686,997]" pageId="2" pageNumber="51">
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Autapomorphies: (1) preorbital region in adults 1.8– 2.0 times as long as postorbital skull table, (2) dermal ornament with continuous and relatively tall radial ridges in the snout, frontals and cheek (contrasting the more polygonal arrangement in
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<taxonomicName id="4C404D11FFF6FFD1FBEE4470FBFDFCA0" baseAuthorityName="Schoch and Witzmann" baseAuthorityYear="2009" box="[1031,1131,813,837]" class="Magnoliopsida" family="Caryophyllaceae" genus="Sclerocephalus" kingdom="Plantae" order="Caryophyllales" pageId="2" pageNumber="51" phylum="Tracheophyta" rank="species" species="nobilis">
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<emphasis id="B934EA80FFF6FFD1FBEE4470FBFDFCA0" box="[1031,1131,813,837]" italics="true" pageId="2" pageNumber="51">S. nobilis</emphasis>
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</taxonomicName>
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), (3) prefrontal anterolaterally expanded to form a more equant pentagon, (4) squamosal posteriorly only half as wide as quadratojugal, (5) phalanges of manus and pes slightly longer and more gracile than in
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<taxonomicName id="4C404D11FFF6FFD1FA4B44D0FCEDFC20" baseAuthorityName="Schoch and Witzmann" baseAuthorityYear="2009" class="Magnoliopsida" family="Caryophyllaceae" genus="Sclerocephalus" kingdom="Plantae" order="Caryophyllales" pageId="2" pageNumber="51" phylum="Tracheophyta" rank="species" species="nobilis">
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<emphasis id="B934EA80FFF6FFD1FA4B44D0FCEDFC20" italics="true" pageId="2" pageNumber="51">S. nobilis</emphasis>
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</taxonomicName>
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and (6) tail substantially longer than skull and trunk combined (shorter than that measurement in
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<taxonomicName id="4C404D11FFF6FFD1FAE64490FAE7FC00" baseAuthorityName="Schoch and Witzmann" baseAuthorityYear="2009" box="[1295,1393,973,997]" class="Magnoliopsida" family="Caryophyllaceae" genus="Sclerocephalus" kingdom="Plantae" order="Caryophyllales" pageId="2" pageNumber="51" phylum="Tracheophyta" rank="species" species="nobilis">
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<emphasis id="B934EA80FFF6FFD1FAE64490FAE7FC00" box="[1295,1393,973,997]" italics="true" pageId="2" pageNumber="51">S. nobilis</emphasis>
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</taxonomicName>
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).
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</paragraph>
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<paragraph id="8BFF3692FFF6FFD1FCDA4350FBA0FBC0" blockId="2.[819,1078,1037,1061]" box="[819,1078,1037,1061]" pageId="2" pageNumber="51">
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<emphasis id="B934EA80FFF6FFD1FCDA4350FBA0FBC0" bold="true" box="[819,1078,1037,1061]" pageId="2" pageNumber="51">Taxonomic assignment</emphasis>
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</paragraph>
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<paragraph id="8BFF3692FFF6FFD1FCDA4310FAD3FA46" blockId="2.[819,1463,1100,1921]" pageId="2" pageNumber="51">
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This taxon was originally erected as
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<taxonomicName id="4C404D11FFF6FFD1FB0C4311FCE9FB61" authorityName="Kratschmer" authorityYear="2006" class="Bivalvia" family="Actinodontidae" genus="Cheliderpeton" higherTaxonomySource="GBIF" kingdom="Animalia" order="Actinodontida" pageId="2" pageNumber="51" phylum="Mollusca" rank="species" species="lellbachae">
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<emphasis id="B934EA80FFF6FFD1FB0C4311FCE9FB61" italics="true" pageId="2" pageNumber="51">Cheliderpeton lellbachae</emphasis>
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</taxonomicName>
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by
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<bibRefCitation id="EFD14B63FFF6FFD1FC414330FBE2FB61" author="Kratschmer, K." box="[936,1140,1133,1156]" pageId="2" pageNumber="51" pagination="3 - 46" refId="ref9586" refString="Kratschmer, K.: Neue temnospondyle Amphibien aus dem Rotliegend des sudwestdeutschen Saar-Nahe-Becken. Teil 1, Geowiss. Beitr. saarpfalz. Rotl., 4, 3 - 46, 2006." type="journal article" year="2006">Krätschmer (2006)</bibRefCitation>
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, who made reference to its resemblance to
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<taxonomicName id="4C404D11FFF6FFD1FC2543D1FB59FB41" baseAuthorityName="Schoch and Witzmann" baseAuthorityYear="2009" box="[972,1231,1164,1188]" class="Bivalvia" family="Actinodontidae" genus="Cheliderpeton" higherTaxonomySource="GBIF" kingdom="Animalia" order="Actinodontida" pageId="2" pageNumber="51" phylum="Mollusca" rank="species" species="latirostre">
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<emphasis id="B934EA80FFF6FFD1FC2543D1FB59FB41" box="[972,1231,1164,1188]" italics="true" pageId="2" pageNumber="51">Cheliderpeton latirostre</emphasis>
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</taxonomicName>
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as described and referred to by
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<bibRefCitation id="EFD14B63FFF6FFD1FC5D43F0FBA6FB21" author="Boy, J. A." box="[948,1072,1197,1220]" pageId="2" pageNumber="51" pagination="123 - 143" refId="ref9171" refString="Boy, J. A.: Uber einige Vertreter der Eryopoidea (Amphibia: Temnospondyli). 4. Cheliderpeton latirostre, Palaontol. Z., 67, 123 - 143, 1993." type="journal article" year="1993">Boy (1993)</bibRefCitation>
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.
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<bibRefCitation id="EFD14B63FFF6FFD1FBD343F0FA14FB21" author="Schoch, R. R. & Witzmann, F." box="[1082,1410,1197,1220]" pageId="2" pageNumber="51" pagination="121 - 136" refId="ref10146" refString="Schoch, R. R. and Witzmann, F.: The temnospondyl Glanochthon from the Lower Permian Meisenheim Formation of Germany, Spec. Pap. Palaeont., 81, 121 - 136, 2009 b." type="journal article" year="2009">Schoch and Witzmann (2009b)</bibRefCitation>
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suggested the new genus name
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<taxonomicName id="4C404D11FFF6FFD1FB8D4391FB78FB01" authorityName="Schoch and Witzmann" authorityYear="2009" box="[1124,1262,1228,1252]" class="Amphibia" genus="Glanochthon" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="2" pageNumber="51" phylum="Chordata" rank="genus">
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<emphasis id="B934EA80FFF6FFD1FB8D4391FB78FB01" box="[1124,1262,1228,1252]" italics="true" pageId="2" pageNumber="51">Glanochthon</emphasis>
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</taxonomicName>
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for
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<taxonomicName id="4C404D11FFF6FFD1FAF74391FC02FAE1" baseAuthorityName="Schoch and Witzmann" baseAuthorityYear="2009" class="Bivalvia" family="Actinodontidae" genus="Cheliderpeton" higherTaxonomySource="GBIF" kingdom="Animalia" order="Actinodontida" pageId="2" pageNumber="51" phylum="Mollusca" rank="species" species="latirostre">
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<emphasis id="B934EA80FFF6FFD1FAF74391FC02FAE1" italics="true" pageId="2" pageNumber="51">Cheliderpeton latirostre</emphasis>
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</taxonomicName>
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after the
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<typeStatus id="54FB8830FFF6FFD1FBE043B1FBA1FAE6" box="[1033,1079,1260,1283]" pageId="2" pageNumber="51">type</typeStatus>
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species
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<taxonomicName id="4C404D11FFF6FFD1FB7543B1FA15FAE1" box="[1180,1411,1260,1284]" class="Bivalvia" family="Actinodontidae" genus="Cheliderpeton" higherTaxonomySource="GBIF" kingdom="Animalia" order="Actinodontida" pageId="2" pageNumber="51" phylum="Mollusca" rank="species" species="vranyi">
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<emphasis id="B934EA80FFF6FFD1FB7543B1FA15FAE1" box="[1180,1411,1260,1284]" italics="true" pageId="2" pageNumber="51">Cheliderpeton vranyi</emphasis>
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</taxonomicName>
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had been redescribed by
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<bibRefCitation id="EFD14B63FFF6FFD1FBFC4251FAC2FAC6" author="Werneburg, R. & Steyer, J. S." box="[1045,1364,1292,1315]" pageId="2" pageNumber="51" pagination="149 - 162" refId="ref10546" refString="Werneburg, R. and Steyer, J. S.: Revision of Cheliderpeton vranyi Fritsch, 1877 (Amphibia, Temnospondyli) from the Lower Permian of Bohemia (Czech Republic), Palaontol. Z., 76, 149 - 162, 2002." type="journal article" year="2002">Werneburg and Steyer (2002)</bibRefCitation>
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and was found not to be closely related to
|
||
<taxonomicName id="4C404D11FFF6FFD1FB4C4271FB96FA86" authority="(Schoch and Witzmann, 2009 b)" baseAuthorityName="Schoch and Witzmann" baseAuthorityYear="2009" class="Bivalvia" family="Actinodontidae" genus="Cheliderpeton" higherTaxonomySource="GBIF" kingdom="Animalia" order="Actinodontida" pageId="2" pageNumber="51" phylum="Mollusca" rank="species" species="latirostre">
|
||
<emphasis id="B934EA80FFF6FFD1FB4C4271FABEFAA1" box="[1189,1320,1324,1348]" italics="true" pageId="2" pageNumber="51">G. latirostre</emphasis>
|
||
(
|
||
<bibRefCitation id="EFD14B63FFF6FFD1FADE4271FC6EFA86" author="Schoch, R. R. & Witzmann, F." pageId="2" pageNumber="51" pagination="121 - 136" refId="ref10146" refString="Schoch, R. R. and Witzmann, F.: The temnospondyl Glanochthon from the Lower Permian Meisenheim Formation of Germany, Spec. Pap. Palaeont., 81, 121 - 136, 2009 b." type="journal article" year="2009">Schoch and Witzmann, 2009b</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
. Based on its below-demonstrated close relationship to
|
||
<taxonomicName id="4C404D11FFF6FFD1FC3B4236FBC5FA66" baseAuthorityName="Schoch and Witzmann" baseAuthorityYear="2009" box="[978,1107,1387,1411]" class="Bivalvia" family="Actinodontidae" genus="Cheliderpeton" higherTaxonomySource="GBIF" kingdom="Animalia" order="Actinodontida" pageId="2" pageNumber="51" phylum="Mollusca" rank="species" species="latirostre">
|
||
<emphasis id="B934EA80FFF6FFD1FC3B4236FBC5FA66" box="[978,1107,1387,1411]" italics="true" pageId="2" pageNumber="51">G. latirostre</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C404D11FFF6FFD1FB894236FAFAFA66" authorityName="Kratschmer" authorityYear="2006" box="[1120,1388,1387,1411]" class="Bivalvia" family="Actinodontidae" genus="Cheliderpeton" higherTaxonomySource="GBIF" kingdom="Animalia" order="Actinodontida" pageId="2" pageNumber="51" phylum="Mollusca" rank="species" species="lellbachae">
|
||
<emphasis id="B934EA80FFF6FFD1FB894236FAFAFA66" box="[1120,1388,1387,1411]" italics="true" pageId="2" pageNumber="51">Cheliderpeton lellbachae</emphasis>
|
||
</taxonomicName>
|
||
is also referred to as
|
||
<taxonomicName id="4C404D11FFF6FFD1FC2F42D6FBC6FA46" authorityName="Schoch and Witzmann" authorityYear="2009" box="[966,1104,1419,1443]" class="Amphibia" genus="Glanochthon" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="2" pageNumber="51" phylum="Chordata" rank="genus">
|
||
<emphasis id="B934EA80FFF6FFD1FC2F42D6FBC6FA46" box="[966,1104,1419,1443]" italics="true" pageId="2" pageNumber="51">Glanochthon</emphasis>
|
||
</taxonomicName>
|
||
as a new combination.
|
||
</paragraph>
|
||
<paragraph id="8BFF3692FFF6FFD1FCA742F6FADCF864" blockId="2.[819,1463,1100,1921]" pageId="2" pageNumber="51">
|
||
<taxonomicName id="4C404D11FFF6FFD1FCA742F6FC4AFA26" authorityName="Kratschmer" authorityYear="2006" box="[846,988,1451,1475]" class="Bivalvia" family="Actinodontidae" genus="Cheliderpeton" higherTaxonomySource="GBIF" kingdom="Animalia" order="Actinodontida" pageId="2" pageNumber="51" phylum="Mollusca" rank="species" species="lellbachae">
|
||
<emphasis id="B934EA80FFF6FFD1FCA742F6FC4AFA26" box="[846,988,1451,1475]" italics="true" pageId="2" pageNumber="51">G. lellbachae</emphasis>
|
||
</taxonomicName>
|
||
co-occurs with
|
||
<taxonomicName id="4C404D11FFF6FFD1FB6C42F6FAE2FA26" baseAuthorityName="Schoch and Witzmann" baseAuthorityYear="2009" box="[1157,1396,1451,1475]" class="Magnoliopsida" family="Caryophyllaceae" genus="Sclerocephalus" kingdom="Plantae" order="Caryophyllales" pageId="2" pageNumber="51" phylum="Tracheophyta" rank="species" species="nobilis">
|
||
<emphasis id="B934EA80FFF6FFD1FB6C42F6FAE2FA26" box="[1157,1396,1451,1475]" italics="true" pageId="2" pageNumber="51">Sclerocephalus nobilis</emphasis>
|
||
</taxonomicName>
|
||
in the same locality and horizon (
|
||
<bibRefCitation id="EFD14B63FFF6FFD1FB804296FA27FA07" author="Kratschmer, K. & Resch, M." box="[1129,1457,1483,1506]" pageId="2" pageNumber="51" pagination="39 - 65" refId="ref9621" refString="Kratschmer, K. and Resch, M.: Klauswaldia nobile gen. et spec. nov. Ein aussergewohnlicher stereospondyler Stegocephale aus dem Rotliegend (Unter-Perm) von Odernheim (Rheinland-Pfalz, SW-Deutschland), Geowiss. Beitr. saarpfalz. Rotl., 3, 39 - 65, 2005." type="journal article" year="2005">Krätschmer and Resch, 2005</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFD14B63FFF6FFD1FCDA42B6FBE2F9E7" author="Schoch, R. R. & Witzmann, F." box="[819,1140,1515,1538]" pageId="2" pageNumber="51" pagination="135 - 168" refId="ref10104" refString="Schoch, R. R. and Witzmann, F.: Osteology and relationships of the temnospondyl genus Sclerocephalus, Zool. J. Linn. Soc. - Lond., 157, 135 - 168, 2009 a." type="journal article" year="2009">Schoch and Witzmann, 2009a</bibRefCitation>
|
||
). The combined list of autapomorphic characters 1–5 for the genus distinguish this taxon from
|
||
<taxonomicName id="4C404D11FFF6FFD1FC864176FC45F9A6" baseAuthorityName="Schoch and Witzmann" baseAuthorityYear="2009" box="[879,979,1579,1603]" class="Magnoliopsida" family="Caryophyllaceae" genus="Sclerocephalus" kingdom="Plantae" order="Caryophyllales" pageId="2" pageNumber="51" phylum="Tracheophyta" rank="species" species="nobilis">
|
||
<emphasis id="B934EA80FFF6FFD1FC864176FC45F9A6" box="[879,979,1579,1603]" italics="true" pageId="2" pageNumber="51">S. nobilis</emphasis>
|
||
</taxonomicName>
|
||
and all other species of
|
||
<taxonomicName id="4C404D11FFF6FFD1FB344176FAE8F9A6" authorityName="Goldfuss" authorityYear="1847" box="[1245,1406,1579,1603]" class="Magnoliopsida" family="Caryophyllaceae" genus="Sclerocephalus" kingdom="Plantae" order="Caryophyllales" pageId="2" pageNumber="51" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="B934EA80FFF6FFD1FB344176FAE8F9A6" box="[1245,1406,1579,1603]" italics="true" pageId="2" pageNumber="51">Sclerocephalus</emphasis>
|
||
</taxonomicName>
|
||
. Hypothetical larvae and small juveniles of
|
||
<taxonomicName id="4C404D11FFF6FFD1FB374117FAFBF987" authorityName="Kratschmer" authorityYear="2006" box="[1246,1389,1610,1634]" class="Bivalvia" family="Actinodontidae" genus="Cheliderpeton" higherTaxonomySource="GBIF" kingdom="Animalia" order="Actinodontida" pageId="2" pageNumber="51" phylum="Mollusca" rank="species" species="lellbachae">
|
||
<emphasis id="B934EA80FFF6FFD1FB374117FAFBF987" box="[1246,1389,1610,1634]" italics="true" pageId="2" pageNumber="51">G. lellbachae</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C404D11FFF6FFD1FA4B4117FCEDF967" baseAuthorityName="Schoch and Witzmann" baseAuthorityYear="2009" class="Magnoliopsida" family="Caryophyllaceae" genus="Sclerocephalus" kingdom="Plantae" order="Caryophyllales" pageId="2" pageNumber="51" phylum="Tracheophyta" rank="species" species="nobilis">
|
||
<emphasis id="B934EA80FFF6FFD1FA4B4117FCEDF967" italics="true" pageId="2" pageNumber="51">S. nobilis</emphasis>
|
||
</taxonomicName>
|
||
may not be distinguished on the basis of the mentioned features, but
|
||
<taxonomicName id="4C404D11FFF6FFD1FBFA41D7FBECF947" baseAuthorityName="Schoch and Witzmann" baseAuthorityYear="2009" box="[1043,1146,1674,1698]" class="Magnoliopsida" family="Caryophyllaceae" genus="Sclerocephalus" kingdom="Plantae" order="Caryophyllales" pageId="2" pageNumber="51" phylum="Tracheophyta" rank="species" species="nobilis">
|
||
<emphasis id="B934EA80FFF6FFD1FBFA41D7FBECF947" box="[1043,1146,1674,1698]" italics="true" pageId="2" pageNumber="51">S. nobilis</emphasis>
|
||
</taxonomicName>
|
||
generally has a wider jugal and more rounded orbit even at small stages. The presence of a third eryopiform temnospondyl at the Klauswald locality (suggested by
|
||
<bibRefCitation id="EFD14B63FFF6FFD1FC1241B7FB28F8E4" author="Kratschmer, K." box="[1019,1214,1770,1793]" pageId="2" pageNumber="51" pagination="3 - 46" refId="ref9586" refString="Kratschmer, K.: Neue temnospondyle Amphibien aus dem Rotliegend des sudwestdeutschen Saar-Nahe-Becken. Teil 1, Geowiss. Beitr. saarpfalz. Rotl., 4, 3 - 46, 2006." type="journal article" year="2006">Krätschmer, 2006</bibRefCitation>
|
||
) cannot be confirmed. With a maximum skull length of
|
||
<quantity id="4CB89B77FFF6FFD1FB794057FB59F8C4" box="[1168,1231,1802,1825]" metricMagnitude="-1" metricUnit="m" metricValue="1.7" pageId="2" pageNumber="51" unit="cm" value="17.0">17 cm</quantity>
|
||
(specimens in private collections),
|
||
<taxonomicName id="4C404D11FFF6FFD1FC554077FBDCF8A7" authorityName="Kratschmer" authorityYear="2006" box="[956,1098,1834,1858]" class="Bivalvia" family="Actinodontidae" genus="Cheliderpeton" higherTaxonomySource="GBIF" kingdom="Animalia" order="Actinodontida" pageId="2" pageNumber="51" phylum="Mollusca" rank="species" species="lellbachae">
|
||
<emphasis id="B934EA80FFF6FFD1FC554077FBDCF8A7" box="[956,1098,1834,1858]" italics="true" pageId="2" pageNumber="51">G. lellbachae</emphasis>
|
||
</taxonomicName>
|
||
appears to have been smaller than the more heavily ossified
|
||
<taxonomicName id="4C404D11FFF6FFD1FBA54017FB27F887" baseAuthorityName="Schoch and Witzmann" baseAuthorityYear="2009" box="[1100,1201,1866,1890]" class="Magnoliopsida" family="Caryophyllaceae" genus="Sclerocephalus" kingdom="Plantae" order="Caryophyllales" pageId="2" pageNumber="51" phylum="Tracheophyta" rank="species" species="nobilis">
|
||
<emphasis id="B934EA80FFF6FFD1FBA54017FB27F887" box="[1100,1201,1866,1890]" italics="true" pageId="2" pageNumber="51">S. nobilis</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<quantity id="4CB89B77FFF6FFD1FB284017FA90F884" box="[1217,1286,1866,1889]" metricMagnitude="-1" metricUnit="m" metricValue="2.4" pageId="2" pageNumber="51" unit="cm" value="24.0">24 cm</quantity>
|
||
), but admittedly the number of available specimens is very limited.
|
||
</paragraph>
|
||
<caption id="DF3F661AFFF7FFD0FF95406FFEC5F880" ID-DOI="http://doi.org/10.5281/zenodo.11359845" ID-Zenodo-Dep="11359845" httpUri="https://zenodo.org/record/11359845/files/figure.png" pageId="3" pageNumber="52" startId="3.[124,191,1842,1864]" targetBox="[235,1359,200,1796]" targetPageId="3" targetType="figure">
|
||
<paragraph id="8BFF3692FFF7FFD0FF95406FFEC5F880" blockId="3.[124,1462,1842,1893]" pageId="3" pageNumber="52">
|
||
<emphasis id="B934EA80FFF7FFD0FF95406FFF4FF8AD" bold="true" box="[124,217,1842,1864]" pageId="3" pageNumber="52">Figure 2.</emphasis>
|
||
Type specimen of
|
||
<taxonomicName id="4C404D11FFF7FFD0FE7C406FFCA9F8AD" authority="(Kratschmer, 2006)" authorityName="Schoch Correspondence" authorityYear="2021" baseAuthorityName="Kratschmer" baseAuthorityYear="2006" box="[405,831,1842,1864]" class="Amphibia" genus="Glanochthon" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="3" pageNumber="52" phylum="Chordata" rank="species" species="lellbachae" status="comb. nov.">
|
||
<emphasis id="B934EA80FFF7FFD0FE7C406FFDEAF8A2" box="[405,636,1842,1863]" italics="true" pageId="3" pageNumber="52">Glanochthon lellbachae</emphasis>
|
||
(
|
||
<bibRefCitation id="EFD14B63FFF7FFD0FD63406EFCAFF8AD" author="Kratschmer, K." box="[650,825,1843,1864]" pageId="3" pageNumber="52" pagination="3 - 46" refId="ref9586" refString="Kratschmer, K.: Neue temnospondyle Amphibien aus dem Rotliegend des sudwestdeutschen Saar-Nahe-Becken. Teil 1, Geowiss. Beitr. saarpfalz. Rotl., 4, 3 - 46, 2006." type="journal article" year="2006">Krätschmer, 2006</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A20757FBFFF7FFD0FCAE406EFC24F8AD" box="[839,946,1843,1864]" pageId="3" pageNumber="52" rank="species">comb. nov.</taxonomicNameLabel>
|
||
(NHMM 2006/14).
|
||
<emphasis id="B934EA80FFF7FFD0FB90406FFB3BF8AD" bold="true" box="[1145,1197,1842,1864]" pageId="3" pageNumber="52">(a, c)</emphasis>
|
||
Close-up of skull roof and
|
||
<emphasis id="B934EA80FFF7FFD0FF954012FF0FF880" bold="true" box="[124,153,1871,1893]" pageId="3" pageNumber="52">(b)</emphasis>
|
||
complete skeleton.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF3F661AFFF0FFD7FF95406EFEDEF883" ID-DOI="http://doi.org/10.5281/zenodo.11359847" ID-Zenodo-Dep="11359847" httpUri="https://zenodo.org/record/11359847/files/figure.png" pageId="4" pageNumber="53" startId="4.[124,191,1843,1865]" targetBox="[283,1303,199,1811]" targetPageId="4" targetType="figure">
|
||
<paragraph id="8BFF3692FFF0FFD7FF95406EFEDEF883" blockId="4.[124,1462,1843,1894]" pageId="4" pageNumber="53">
|
||
<emphasis id="B934EA80FFF0FFD7FF95406EFF4CF8AC" bold="true" box="[124,218,1843,1865]" pageId="4" pageNumber="53">Figure 3.</emphasis>
|
||
<taxonomicName id="4C404D11FFF0FFD7FF0A406EFD04F8AC" authority="(Kratschmer, 2006)" authorityName="Schoch Correspondence" authorityYear="2021" baseAuthorityName="Kratschmer" baseAuthorityYear="2006" box="[227,658,1843,1865]" class="Amphibia" genus="Glanochthon" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="4" pageNumber="53" phylum="Chordata" rank="species" species="lellbachae" status="comb. nov.">
|
||
<emphasis id="B934EA80FFF0FFD7FF0A406EFE5AF8AD" box="[227,460,1843,1864]" italics="true" pageId="4" pageNumber="53">Glanochthon lellbachae</emphasis>
|
||
(
|
||
<bibRefCitation id="EFD14B63FFF0FFD7FE324069FD1AF8AC" author="Kratschmer, K." box="[475,652,1844,1865]" pageId="4" pageNumber="53" pagination="3 - 46" refId="ref9586" refString="Kratschmer, K.: Neue temnospondyle Amphibien aus dem Rotliegend des sudwestdeutschen Saar-Nahe-Becken. Teil 1, Geowiss. Beitr. saarpfalz. Rotl., 4, 3 - 46, 2006." type="journal article" year="2006">Krätschmer, 2006</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A20757FBFFF0FFD7FD724069FC9DF8AC" box="[667,779,1844,1865]" pageId="4" pageNumber="53" rank="species">comb. nov.</taxonomicNameLabel>
|
||
:
|
||
<emphasis id="B934EA80FFF0FFD7FCFF406EFCA4F8AC" bold="true" box="[790,818,1843,1865]" pageId="4" pageNumber="53">(a)</emphasis>
|
||
SMNS 91281,
|
||
<emphasis id="B934EA80FFF0FFD7FC39406EFC7BF8AC" bold="true" box="[976,1005,1843,1865]" pageId="4" pageNumber="53">(b)</emphasis>
|
||
UGKU POL-F 1997/1,
|
||
<emphasis id="B934EA80FFF0FFD7FB08406EFB6DF8AC" bold="true" box="[1249,1275,1843,1865]" pageId="4" pageNumber="53">(c)</emphasis>
|
||
SMNS 90507 and
|
||
<emphasis id="B934EA80FFF0FFD7FF95400DFF0FF883" bold="true" box="[124,153,1872,1894]" pageId="4" pageNumber="53">(d)</emphasis>
|
||
NHMM 2006/14.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF3F661AFFF1FFD6FF95401EFF53F893" ID-DOI="http://doi.org/10.5281/zenodo.11359849" ID-Zenodo-Dep="11359849" httpUri="https://zenodo.org/record/11359849/files/figure.png" pageId="5" pageNumber="54" startId="5.[124,191,1859,1881]" targetBox="[283,1303,187,1828]" targetPageId="5" targetType="figure">
|
||
<paragraph id="8BFF3692FFF1FFD6FF95401EFF53F893" blockId="5.[124,1461,1859,1910]" pageId="5" pageNumber="54">
|
||
<emphasis id="B934EA80FFF1FFD6FF95401EFF4EF8BC" bold="true" box="[124,216,1859,1881]" pageId="5" pageNumber="54">Figure 4.</emphasis>
|
||
Palate and postcranium of
|
||
<taxonomicName id="4C404D11FFF1FFD6FE37401EFC12F8BC" authority="(Kratschmer, 2006)" authorityName="Schoch Correspondence" authorityYear="2021" baseAuthorityName="Kratschmer" baseAuthorityYear="2006" box="[478,900,1859,1881]" class="Amphibia" genus="Glanochthon" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="5" pageNumber="54" phylum="Chordata" rank="species" species="lellbachae" status="comb. nov.">
|
||
<emphasis id="B934EA80FFF1FFD6FE37401EFD55F8BD" box="[478,707,1859,1880]" italics="true" pageId="5" pageNumber="54">Glanochthon lellbachae</emphasis>
|
||
(
|
||
<bibRefCitation id="EFD14B63FFF1FFD6FD394019FCE8F8BC" author="Kratschmer, K." box="[720,894,1860,1881]" pageId="5" pageNumber="54" pagination="3 - 46" refId="ref9586" refString="Kratschmer, K.: Neue temnospondyle Amphibien aus dem Rotliegend des sudwestdeutschen Saar-Nahe-Becken. Teil 1, Geowiss. Beitr. saarpfalz. Rotl., 4, 3 - 46, 2006." type="journal article" year="2006">Krätschmer, 2006</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A20757FBFFF1FFD6FC624019FC61F8BC" box="[907,1015,1860,1881]" pageId="5" pageNumber="54" rank="species">comb. nov.</taxonomicNameLabel>
|
||
:
|
||
<emphasis id="B934EA80FFF1FFD6FBE9401EFBA3F8BC" bold="true" box="[1024,1077,1859,1881]" pageId="5" pageNumber="54">(a, b)</emphasis>
|
||
SMNS 90507 and
|
||
<emphasis id="B934EA80FFF1FFD6FB07401EFAB4F8BC" bold="true" box="[1262,1314,1859,1881]" pageId="5" pageNumber="54">(c, d)</emphasis>
|
||
UGKU POL-F 1997/1.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BFF3692FFF2FFD5FF954737FA20FF67" blockId="6.[124,1462,106,130]" box="[124,1462,106,130]" pageId="6" pageNumber="55">
|
||
<emphasis id="B934EA80FFF2FFD5FF954737FA20FF67" bold="true" box="[124,1462,106,130]" pageId="6" pageNumber="55">
|
||
<collectionCode id="ED51AE57FFF2FFD5FF954737FF07FF67" box="[124,145,106,130]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="6" pageNumber="55">R</collectionCode>
|
||
.
|
||
<collectionCode id="ED51AE57FFF2FFD5FF754737FF27FF67" box="[156,177,106,130]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="6" pageNumber="55">R</collectionCode>
|
||
. Schoch: Osteology of the Permian temnospondyl amphibian
|
||
<taxonomicName id="4C404D11FFF2FFD5FC834737FBE7FF67" authorityName="Schoch Correspondence" authorityYear="2021" baseAuthorityName="Kratschmer" baseAuthorityYear="2006" box="[874,1137,106,130]" class="Amphibia" genus="Glanochthon" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="6" pageNumber="55" phylum="Chordata" rank="species" species="lellbachae">
|
||
<emphasis id="B934EA80FFF2FFD5FC834737FBE7FF67" bold="true" box="[874,1137,106,130]" italics="true" pageId="6" pageNumber="55">Glanochthon lellbachae</emphasis>
|
||
</taxonomicName>
|
||
55
|
||
</emphasis>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF3F661AFFF2FFD5FF954595FF53FD1E" ID-DOI="http://doi.org/10.5281/zenodo.11359851" ID-Zenodo-Dep="11359851" httpUri="https://zenodo.org/record/11359851/files/figure.png" pageId="6" pageNumber="55" startId="6.[124,191,712,734]" targetBox="[188,1397,177,681]" targetPageId="6" targetType="figure">
|
||
<paragraph id="8BFF3692FFF2FFD5FF954595FF53FD1E" blockId="6.[124,1462,712,763]" pageId="6" pageNumber="55">
|
||
<emphasis id="B934EA80FFF2FFD5FF954595FF41FD3B" bold="true" box="[124,215,712,734]" pageId="6" pageNumber="55">Figure 5.</emphasis>
|
||
Comparison of
|
||
<taxonomicName id="4C404D11FFF2FFD5FE9B4595FDC1FD38" authorityName="Schoch Correspondence" authorityYear="2021" baseAuthorityName="Kratschmer" baseAuthorityYear="2006" box="[370,599,712,733]" class="Amphibia" genus="Glanochthon" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="6" pageNumber="55" phylum="Chordata" rank="species" species="lellbachae">
|
||
<emphasis id="B934EA80FFF2FFD5FE9B4595FDC1FD38" box="[370,599,712,733]" italics="true" pageId="6" pageNumber="55">Glanochthon lellbachae</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B934EA80FFF2FFD5FDB74595FDEFFD3B" bold="true" box="[606,633,712,734]" pageId="6" pageNumber="55">(a)</emphasis>
|
||
with
|
||
<taxonomicName id="4C404D11FFF2FFD5FD594595FC8FFD38" authorityName="Schoch & Witzmann" authorityYear="2009" box="[688,793,712,733]" class="Amphibia" genus="Glanochthon" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="6" pageNumber="55" phylum="Chordata" rank="species" species="angusta">
|
||
<emphasis id="B934EA80FFF2FFD5FD594595FC8FFD38" box="[688,793,712,733]" italics="true" pageId="6" pageNumber="55">G. angusta</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B934EA80FFF2FFD5FCF64595FCAAFD3B" bold="true" box="[799,828,712,734]" pageId="6" pageNumber="55">(b)</emphasis>
|
||
and
|
||
<taxonomicName id="4C404D11FFF2FFD5FC824595FC49FD38" baseAuthorityName="Schoch and Witzmann" baseAuthorityYear="2009" box="[875,991,712,733]" class="Bivalvia" family="Actinodontidae" genus="Cheliderpeton" higherTaxonomySource="GBIF" kingdom="Animalia" order="Actinodontida" pageId="6" pageNumber="55" phylum="Mollusca" rank="species" species="latirostre">
|
||
<emphasis id="B934EA80FFF2FFD5FC824595FC49FD38" box="[875,991,712,733]" italics="true" pageId="6" pageNumber="55">G. latirostre</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B934EA80FFF2FFD5FC0C4595FC69FD3B" bold="true" box="[997,1023,712,734]" pageId="6" pageNumber="55">(c)</emphasis>
|
||
. (Panels
|
||
<emphasis id="B934EA80FFF2FFD5FBB14595FBF3FD3B" bold="true" box="[1112,1125,712,734]" pageId="6" pageNumber="55">b</emphasis>
|
||
and
|
||
<emphasis id="B934EA80FFF2FFD5FB7A4595FB08FD3B" bold="true" box="[1171,1182,712,734]" pageId="6" pageNumber="55">c</emphasis>
|
||
from
|
||
<bibRefCitation id="EFD14B63FFF2FFD5FB314594FF2DFD1E" author="Schoch, R. R. & Witzmann, F." pageId="6" pageNumber="55" pagination="121 - 136" refId="ref10146" refString="Schoch, R. R. and Witzmann, F.: The temnospondyl Glanochthon from the Lower Permian Meisenheim Formation of Germany, Spec. Pap. Palaeont., 81, 121 - 136, 2009 b." type="journal article" year="2009">Schoch and Witzmann, 2009b</bibRefCitation>
|
||
.)
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C35A6519FFF2FFD5FF954460FE99FAF5" pageId="6" pageNumber="55" type="description">
|
||
<paragraph id="8BFF3692FFF2FFD5FF954460FE0CFCB0" blockId="6.[124,410,829,853]" box="[124,410,829,853]" pageId="6" pageNumber="55">
|
||
<emphasis id="B934EA80FFF2FFD5FF954460FE0CFCB0" bold="true" box="[124,410,829,853]" pageId="6" pageNumber="55">Phylogeny and taxonomy</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BFF3692FFF2FFD5FF954427FE99FAF5" blockId="6.[124,767,890,1296]" pageId="6" pageNumber="55">
|
||
In the analysis reported below,
|
||
<taxonomicName id="4C404D11FFF2FFD5FDFE4427FD1CFC77" baseAuthorityName="Schoch and Witzmann" baseAuthorityYear="2009" box="[535,650,890,914]" class="Magnoliopsida" family="Caryophyllaceae" genus="Sclerocephalus" kingdom="Plantae" order="Caryophyllales" pageId="6" pageNumber="55" phylum="Tracheophyta" rank="species" species="nobilis">
|
||
<emphasis id="B934EA80FFF2FFD5FDFE4427FD1CFC77" box="[535,650,890,914]" italics="true" pageId="6" pageNumber="55">S. nobilis</emphasis>
|
||
</taxonomicName>
|
||
and the
|
||
<taxonomicName id="4C404D11FFF2FFD5FF9544C7FE90FC57" authorityName="Schoch and Witzmann" authorityYear="2009" box="[124,262,922,946]" class="Amphibia" genus="Glanochthon" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="6" pageNumber="55" phylum="Chordata" rank="genus">
|
||
<emphasis id="B934EA80FFF2FFD5FF9544C7FE90FC57" box="[124,262,922,946]" italics="true" pageId="6" pageNumber="55">Glanochthon</emphasis>
|
||
</taxonomicName>
|
||
clade are found to be sister taxa. This recognized topological pattern leads to the phylogenetically problematic situation that
|
||
<taxonomicName id="4C404D11FFF2FFD5FE8C4484FD90FC14" authorityName="Goldfuss" authorityYear="1847" box="[357,518,985,1009]" class="Magnoliopsida" family="Caryophyllaceae" genus="Sclerocephalus" kingdom="Plantae" order="Caryophyllales" pageId="6" pageNumber="55" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="B934EA80FFF2FFD5FE8C4484FD90FC14" box="[357,518,985,1009]" italics="true" pageId="6" pageNumber="55">Sclerocephalus</emphasis>
|
||
</taxonomicName>
|
||
forms a grade towards
|
||
<taxonomicName id="4C404D11FFF2FFD5FF9544A4FE90FBF4" authorityName="Schoch and Witzmann" authorityYear="2009" box="[124,262,1017,1041]" class="Amphibia" genus="Glanochthon" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="6" pageNumber="55" phylum="Chordata" rank="genus">
|
||
<emphasis id="B934EA80FFF2FFD5FF9544A4FE90FBF4" box="[124,262,1017,1041]" italics="true" pageId="6" pageNumber="55">Glanochthon</emphasis>
|
||
</taxonomicName>
|
||
, which under strict application of cladistic principles would prompt erection of new genera at each node. An alternative option would be to define
|
||
<taxonomicName id="4C404D11FFF2FFD5FDB44364FD68FBB4" authorityName="Goldfuss" authorityYear="1847" box="[605,766,1081,1105]" class="Magnoliopsida" family="Caryophyllaceae" genus="Sclerocephalus" kingdom="Plantae" order="Caryophyllales" pageId="6" pageNumber="55" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="B934EA80FFF2FFD5FDB44364FD68FBB4" box="[605,766,1081,1105]" italics="true" pageId="6" pageNumber="55">Sclerocephalus</emphasis>
|
||
</taxonomicName>
|
||
more broadly, encompassing all of
|
||
<taxonomicName id="4C404D11FFF2FFD5FE1F4304FD3DFB95" authorityName="Jaekel" authorityYear="1909" box="[502,683,1113,1136]" class="Amphibia" family="Sclerocephalidae" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="6" pageNumber="55" phylum="Chordata" rank="family">Sclerocephalidae</taxonomicName>
|
||
including
|
||
<taxonomicName id="4C404D11FFF2FFD5FF4E4324FEA7FB74" authorityName="Schoch and Witzmann" authorityYear="2009" box="[167,305,1145,1169]" class="Amphibia" genus="Glanochthon" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="6" pageNumber="55" phylum="Chordata" rank="genus">
|
||
<emphasis id="B934EA80FFF2FFD5FF4E4324FEA7FB74" box="[167,305,1145,1169]" italics="true" pageId="6" pageNumber="55">Glanochthon</emphasis>
|
||
</taxonomicName>
|
||
. At the present stage, I consider any such step premature as long as morphological effects of microevolution cannot be distinguished from other effects, especially plasticity (ecophenotypes), which is beyond the scope of the present study.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C35A6519FFF2FFD5FF954268FE96F940" pageId="6" pageNumber="55" type="discussion">
|
||
<paragraph id="8BFF3692FFF2FFD5FF954268FF7AFAA8" blockId="6.[124,236,1333,1357]" box="[124,236,1333,1357]" pageId="6" pageNumber="55">
|
||
<emphasis id="B934EA80FFF2FFD5FF954268FF7AFAA8" bold="true" box="[124,236,1333,1357]" pageId="6" pageNumber="55">Comment</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BFF3692FFF2FFD5FF95422FFF68F98D" blockId="6.[124,767,1394,1640]" pageId="6" pageNumber="55">
|
||
Based on the current phylogenetic findings,
|
||
<taxonomicName id="4C404D11FFF2FFD5FDB3422FFD72FA6F" authorityName="Schoch and Witzmann" authorityYear="2009" box="[602,740,1394,1418]" class="Amphibia" genus="Glanochthon" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="6" pageNumber="55" phylum="Chordata" rank="genus">
|
||
<emphasis id="B934EA80FFF2FFD5FDB3422FFD72FA6F" box="[602,740,1394,1418]" italics="true" pageId="6" pageNumber="55">Glanochthon</emphasis>
|
||
</taxonomicName>
|
||
is referred to as the family
|
||
<taxonomicName id="4C404D11FFF2FFD5FE6F42CFFDA8FA4C" authorityName="Jaekel" authorityYear="1909" box="[390,574,1426,1449]" class="Amphibia" family="Sclerocephalidae" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="6" pageNumber="55" phylum="Chordata" rank="family">Sclerocephalidae</taxonomicName>
|
||
, which contains a monophyletic group including a long
|
||
<taxonomicName id="4C404D11FFF2FFD5FDF042ECFD2CFA2C" authorityName="Goldfuss" authorityYear="1847" box="[537,698,1457,1481]" class="Magnoliopsida" family="Caryophyllaceae" genus="Sclerocephalus" kingdom="Plantae" order="Caryophyllales" pageId="6" pageNumber="55" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="B934EA80FFF2FFD5FDF042ECFD2CFA2C" box="[537,698,1457,1481]" italics="true" pageId="6" pageNumber="55">Sclerocephalus</emphasis>
|
||
</taxonomicName>
|
||
grade and a terminal
|
||
<taxonomicName id="4C404D11FFF2FFD5FEC1428CFE24FA0C" authorityName="Schoch and Witzmann" authorityYear="2009" box="[296,434,1489,1513]" class="Amphibia" genus="Glanochthon" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="6" pageNumber="55" phylum="Chordata" rank="genus">
|
||
<emphasis id="B934EA80FFF2FFD5FEC1428CFE24FA0C" box="[296,434,1489,1513]" italics="true" pageId="6" pageNumber="55">Glanochthon</emphasis>
|
||
</taxonomicName>
|
||
clade. The former referral of
|
||
<taxonomicName id="4C404D11FFF2FFD5FF9542ACFE90F9EC" authorityName="Schoch and Witzmann" authorityYear="2009" box="[124,262,1521,1545]" class="Amphibia" genus="Glanochthon" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="6" pageNumber="55" phylum="Chordata" rank="genus">
|
||
<emphasis id="B934EA80FFF2FFD5FF9542ACFE90F9EC" box="[124,262,1521,1545]" italics="true" pageId="6" pageNumber="55">Glanochthon</emphasis>
|
||
</taxonomicName>
|
||
(as
|
||
<taxonomicName id="4C404D11FFF2FFD5FED242ACFDA9F9EC" baseAuthorityName="Schoch and Witzmann" baseAuthorityYear="2009" box="[315,575,1521,1545]" class="Bivalvia" family="Actinodontidae" genus="Cheliderpeton" higherTaxonomySource="GBIF" kingdom="Animalia" order="Actinodontida" pageId="6" pageNumber="55" phylum="Mollusca" rank="species" species="latirostre">
|
||
<emphasis id="B934EA80FFF2FFD5FED242ACFDA9F9EC" box="[315,575,1521,1545]" italics="true" pageId="6" pageNumber="55">Cheliderpeton latirostre</emphasis>
|
||
</taxonomicName>
|
||
) within the
|
||
<taxonomicName id="4C404D11FFF2FFD5FD2542AFFE69F9CD" authority="(Schoch and Milner, 2000)" baseAuthorityName="Schoch and Milner" baseAuthorityYear="2000" class="Amphibia" family="Intasuchidae" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="6" pageNumber="55" phylum="Chordata" rank="family">
|
||
Intasuchidae (
|
||
<bibRefCitation id="EFD14B63FFF2FFD5FF0E414CFE6EF9CD" author="Schoch, R. R. & Milner, A. R." box="[231,504,1553,1576]" pageId="6" pageNumber="55" refId="ref10016" refString="Schoch, R. R. and Milner, A. R.: Stereospondyli, in: Handbuch der Palaoherpetologie, Vol. 3 B " edited by: Wellnhofer, P., Friedrich Pfeil, Munich, 203 pp., 2000." type="book" year="2000">Schoch and Milner, 2000</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
was based on a suite of characters which appear to be convergent in the present light of evidence.
|
||
</paragraph>
|
||
<paragraph id="8BFF3692FFF2FFD5FF9541D0FE96F940" blockId="6.[124,256,1677,1701]" box="[124,256,1677,1701]" pageId="6" pageNumber="55">
|
||
<emphasis id="B934EA80FFF2FFD5FF9541D0FE96F940" bold="true" box="[124,256,1677,1701]" pageId="6" pageNumber="55">Occurrence</emphasis>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C35A6519FFF2FFD5FF954197FAA6FC90" pageId="6" pageNumber="55" type="description">
|
||
<paragraph id="8BFF3692FFF2FFD5FF954197FAA6FC90" blockId="6.[124,767,1738,1921]" lastBlockId="6.[819,1462,830,885]" pageId="6" pageNumber="55">
|
||
Although
|
||
<taxonomicName id="4C404D11FFF2FFD5FF184197FECBF907" baseAuthorityName="Schoch and Witzmann" baseAuthorityYear="2009" box="[241,349,1738,1762]" class="Magnoliopsida" family="Caryophyllaceae" genus="Sclerocephalus" kingdom="Plantae" order="Caryophyllales" pageId="6" pageNumber="55" phylum="Tracheophyta" rank="species" species="nobilis">
|
||
<emphasis id="B934EA80FFF2FFD5FF184197FECBF907" box="[241,349,1738,1762]" italics="true" pageId="6" pageNumber="55">S. nobilis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C404D11FFF2FFD5FE4D4197FDAAF907" authorityName="Schoch Correspondence" authorityYear="2021" baseAuthorityName="Kratschmer" baseAuthorityYear="2006" box="[420,572,1738,1762]" class="Amphibia" genus="Glanochthon" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="6" pageNumber="55" phylum="Chordata" rank="species" species="lellbachae">
|
||
<emphasis id="B934EA80FFF2FFD5FE4D4197FDAAF907" box="[420,572,1738,1762]" italics="true" pageId="6" pageNumber="55">G. lellbachae</emphasis>
|
||
</taxonomicName>
|
||
co-occur in the same locality and several successive horizons, they differ in sample size per horizon. Most notably,
|
||
<taxonomicName id="4C404D11FFF2FFD5FDB84057FD74F8C7" authorityName="Schoch Correspondence" authorityYear="2021" baseAuthorityName="Kratschmer" baseAuthorityYear="2006" box="[593,738,1802,1826]" class="Amphibia" genus="Glanochthon" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="6" pageNumber="55" phylum="Chordata" rank="species" species="lellbachae">
|
||
<emphasis id="B934EA80FFF2FFD5FDB84057FD74F8C7" box="[593,738,1802,1826]" italics="true" pageId="6" pageNumber="55">G. lellbachae</emphasis>
|
||
</taxonomicName>
|
||
is more common in the upper fish beds (Obere Fischschiefer, “kalkige Papierschiefer”, K5), whereas
|
||
<taxonomicName id="4C404D11FFF2FFD5FDD84017FD0EF887" baseAuthorityName="Schoch and Witzmann" baseAuthorityYear="2009" box="[561,664,1866,1890]" class="Magnoliopsida" family="Caryophyllaceae" genus="Sclerocephalus" kingdom="Plantae" order="Caryophyllales" pageId="6" pageNumber="55" phylum="Tracheophyta" rank="species" species="nobilis">
|
||
<emphasis id="B934EA80FFF2FFD5FDD84017FD0EF887" box="[561,664,1866,1890]" italics="true" pageId="6" pageNumber="55">S. nobilis</emphasis>
|
||
</taxonomicName>
|
||
peaks in the lower fish beds (Untere Fischschiefer, K2) as defined by
|
||
<bibRefCitation id="EFD14B63FFF2FFD5FCDA4463FB95FCB0" author="Kratschmer, K." box="[819,1027,830,853]" pageId="6" pageNumber="55" pagination="1 - 52" refId="ref9550" refString="Kratschmer, K.: Revision von " Sclerocephalus haeuseri " (Goldfuss) 1847 (Stem-Stereospondyli), Geowiss. Beitr. saarpfalz. Rotl., 2, 1 - 52, 2004." type="journal article" year="2004">Krätschmer (2004)</bibRefCitation>
|
||
. In the other beds (K3, 4), the two taxa co-occur, mostly represented by large juveniles.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |