treatments-xml/data/03/E8/87/03E887D8CE4DFF9034F81A40D5A37651.xml

<document id="5F703D7DB22A284CB0D52177FAABAEFF" ID-DOI="10.26107/RBZ-2022-0009" ID-ISSN="2345-7600" ID-Zenodo-Dep="7174234" ID-ZooBank="BE6164AE-1C24-4E01-8B7B-D80764F147B3" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1665290677448" checkinUser="felipe" docAuthor="Ng, Peter K. L. &amp; Ahyong, Shane T." docDate="2022" docId="03E887D8CE4DFF9034F81A40D5A37651" docLanguage="en" docName="RafflesBZool.70.134–248.pdf" docOrigin="Raffles Bulletin of Zoology 70" docStyle="DocumentStyle:AF09A0581917EACD12E7F0991C3E7C26.5:RafflesBZool.2019-.journal_article" docStyleId="AF09A0581917EACD12E7F0991C3E7C26" docStyleName="RafflesBZool.2019-.journal_article" docStyleVersion="5" docTitle="Arcotheres rayi Ahyong &amp; Ng 2007" docType="treatment" docVersion="3" lastPageNumber="199" masterDocId="FFD1FFA0CE71FFD137601D0FD0617E6B" masterDocTitle="The pea crab genus Arcotheres Manning, 1993 (Crustacea: Brachyura: Pinnotheridae) from Singapore and Peninsular Malaysia, with a reappraisal of diagnostic characters and descriptions of two new genera" masterLastPageNumber="248" masterPageNumber="134" pageNumber="194" updateTime="1699379806661" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
<mods:mods id="0E9985C9476E967451CCC6DF1A4E4978" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="1CA064EDC2D4ED88F05114BAA04737B0">
<mods:title id="12591B92C0F996845D640F7D99208548">The pea crab genus Arcotheres Manning, 1993 (Crustacea: Brachyura: Pinnotheridae) from Singapore and Peninsular Malaysia, with a reappraisal of diagnostic characters and descriptions of two new genera</mods:title>
</mods:titleInfo>
<mods:name id="5AA2C26D598F0049745A585607D7A410" type="personal">
<mods:role id="59512DCF4508E31D03EEFC485BC5C82C">
<mods:roleTerm id="6E2F5BA9E8F3F83F16F17BDF30EE9782">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="CBB2659FED9789B8FE71C4BC547B6925">Ng, Peter K. L.</mods:namePart>
</mods:name>
<mods:name id="7F7ECA0CFF97D9CE19870371CB3860D6" type="personal">
<mods:role id="08F4A00428C624BDF91FC969A46469DE">
<mods:roleTerm id="6041F3BCBB68EAB72869FFD0B72D73CD">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="5EC319E82378433894CE85BD059A5AEB">Ahyong, Shane T.</mods:namePart>
</mods:name>
<mods:typeOfResource id="0FBB41EC11D2362D4A8B251463CF9559">text</mods:typeOfResource>
<mods:relatedItem id="8E8BEFFE5D397A8EFD58BEDA2ED51542" type="host">
<mods:titleInfo id="BE085187A5A6855D9EB82CF1526BCF8B">
<mods:title id="52A88F703E63872415DA5C849983FDE4">Raffles Bulletin of Zoology</mods:title>
</mods:titleInfo>
<mods:part id="5F795B2323C1D1730728918A495C8238">
<mods:date id="D440F463C98D20F47BE299275378C8BE">2022</mods:date>
<mods:detail id="2123CF575F7D740A247AEDE1FED02551" type="pubDate">
<mods:number id="82BBD356C13D691FE27675E6C50DEFB6">2022-04-01</mods:number>
</mods:detail>
<mods:detail id="4F83E45DA7E29B12411390D5449791F3" type="volume">
<mods:number id="FDE10A44DAD8156986019FCC94D552F7">70</mods:number>
</mods:detail>
<mods:extent id="805912040E9FC5300DD80FB5FDEEE720" unit="page">
<mods:start id="039EF6B0D7E2BEB4BA2B330411CDB766">134</mods:start>
<mods:end id="1091B8FF99C44B87515BE56A68585F39">248</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="92638D739E70F766D28C092FCD0F1856">journal article</mods:classification>
<mods:identifier id="47627284993D3E02EA5F6B318074AB25" type="DOI">10.26107/RBZ-2022-0009</mods:identifier>
<mods:identifier id="B1365165A9F50EC5E74B7FC54C0E0166" type="ISSN">2345-7600</mods:identifier>
<mods:identifier id="AF88C53C991B1706C6C7B7EB8BD5A82E" type="Zenodo-Dep">7174234</mods:identifier>
<mods:identifier id="8E5E85E06D3BA64D3697E395FC546598" type="ZooBank">BE6164AE-1C24-4E01-8B7B-D80764F147B3</mods:identifier>
</mods:mods>
<treatment id="03E887D8CE4DFF9034F81A40D5A37651" LSID="urn:lsid:plazi:treatment:03E887D8CE4DFF9034F81A40D5A37651" httpUri="http://treatment.plazi.org/id/03E887D8CE4DFF9034F81A40D5A37651" lastPageId="65" lastPageNumber="199" pageId="60" pageNumber="194">
<subSubSection id="C35B6545CE4DFFED34F81A40D523790C" box="[920,1346,1871,1895]" pageId="60" pageNumber="194" type="nomenclature">
<paragraph id="8BFE36CECE4DFFED34F81A40D523790C" blockId="60.[920,1346,1871,1927]" box="[920,1346,1871,1895]" pageId="60" pageNumber="194">
<heading id="D0B681A2CE4DFFED34F81A40D523790C" bold="true" box="[920,1346,1871,1895]" centered="true" fontSize="10" level="2" pageId="60" pageNumber="194" reason="2">
<taxonomicName id="4C414D4DCE4DFFED34F81A40D523790C" ID-CoL="67PY8" authority="Ahyong &amp; Ng, 2007 b" authorityName="Ahyong &amp; Ng" authorityYear="2007" box="[920,1346,1871,1895]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" kingdom="Animalia" order="Decapoda" pageId="60" pageNumber="194" phylum="Arthropoda" rank="species" species="rayi">
<emphasis id="B935EADCCE4DFFED34F81A40D523790C" bold="true" box="[920,1346,1871,1895]" pageId="60" pageNumber="194">
<emphasis id="B935EADCCE4DFFED34F81A40D422790C" bold="true" box="[920,1091,1871,1895]" italics="true" pageId="60" pageNumber="194">Arcotheres rayi</emphasis>
<bibRefCitation id="EFD04B3FCE4DFFED332E1A40D523790C" author="Ahyong ST &amp; Ng PKL" box="[1102,1346,1871,1895]" pageId="60" pageNumber="194" pagination="191 - 226" refId="ref62272" refString="Ahyong ST &amp; Ng PKL (2007 b) The pinnotherid type material of Semper (1880), Nauck (1880) and Burger (1895) (Crustacea: Decapoda: Brachyura). Raffles Bulletin of Zoology, Supplement 16: 191 - 226." type="journal article" year="2007">Ahyong &amp; Ng, 2007b</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C35B6545CE4DFFED34AC1A60D56C79EC" box="[972,1293,1903,1927]" pageId="60" pageNumber="194" type="description">
<paragraph id="8BFE36CECE4DFFED34AC1A60D56C79EC" blockId="60.[920,1346,1871,1927]" box="[972,1293,1903,1927]" pageId="60" pageNumber="194">
(
<figureCitation id="137A2A4BCE4DFFED34B41A60D43679EC" box="[980,1111,1903,1927]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStart-3="Fig" captionStartId-0="61.[136,171,1192,1214]" captionStartId-1="62.[113,148,1974,1996]" captionStartId-2="63.[135,170,1761,1783]" captionStartId-3="64.[113,148,1597,1619]" captionTargetBox-0="[136,1472,189,1177]" captionTargetBox-1="[113,1451,279,1958]" captionTargetBox-2="[135,1473,190,1746]" captionTargetBox-3="[117,1455,192,1580]" captionTargetId-0="figure-7@61.[136,1472,189,1177]" captionTargetId-1="figure-14@62.[113,1451,279,1958]" captionTargetId-2="figure-7@63.[135,1473,190,1746]" captionTargetId-3="figure-14@64.[107,1458,184,1592]" captionTargetPageId-0="61" captionTargetPageId-1="62" captionTargetPageId-2="63" captionTargetPageId-3="64" captionText-0="Fig. 46. Arcotheres rayi Ahyong &amp; Ng, 2007, holotype female (11.3 × 8.7 mm) (SMF-ZMG 952), Philippines. A, overall dorsal view; B, ventral view of cephalothorax; C, outer view of left chela." captionText-1="Fig. 47. Arcotheres rayi Ahyong &amp; Ng, 2007, female (6.9 × 5.2 mm) (ZRC 2013.1416), Peninsular Malaysia. A, overall dorsal view; B, ventral view of cephalothorax; C, dorsal view of carapace; D, frontal view of cephalothorax; E, outer view of left chela." captionText-2="Fig. 48. Arcotheres rayi Ahyong &amp; Ng, 2007. A, D, ovigerous female (7.6 × 5.8 mm) (ZRC 2013.1416), Peninsular Malaysia; B, C, E, female (6.2 × 4.4 mm) (ZRC 2013.1416), Peninsular Malaysia. A, B, overall dorsal view; C, dorsal view of carapace; D, E, right P5." captionText-3="Fig. 49. Arcotheres rayi Ahyong &amp; Ng, 2007, ovigerous female (7.6 × 5.8 mm) (ZRC 2013.1416), Peninsular Malaysia. A, overall dorsal view; B, right MXP3; C, frontal view of cephalothorax; D, anterior thoracic sternum; E, right chela; F–I, left P2–P5; J–M, right P2–P5. Scale = A, 2.0 mm; B, 0.5 mm; C–M, 1.0 mm." figureDoi-0="http://doi.org/10.5281/zenodo.7174331" figureDoi-1="http://doi.org/10.5281/zenodo.7174333" figureDoi-2="http://doi.org/10.5281/zenodo.7174335" figureDoi-3="http://doi.org/10.5281/zenodo.7174339" httpUri-0="https://zenodo.org/record/7174331/files/figure.png" httpUri-1="https://zenodo.org/record/7174333/files/figure.png" httpUri-2="https://zenodo.org/record/7174335/files/figure.png" httpUri-3="https://zenodo.org/record/7174339/files/figure.png" pageId="60" pageNumber="194">Figs. 46–49</figureCitation>
, 
<figureCitation id="137A2A4BCE4DFFED33041A60D4D379EC" box="[1124,1202,1903,1927]" captionStart="Fig" captionStartId="65.[135,170,1541,1563]" captionTargetBox="[137,1477,189,1531]" captionTargetId="figure-7@65.[132,1485,182,1535]" captionTargetPageId="65" captionText="Fig. 50. Right female P5. A–G, Arcotheres exiguus (Bürger, 1895), Peninsular Malaysia: A, juvenile (3.8 × 3.6 mm) (ZRC 2018.772); B, ovigerous (4.5 × 4.1 mm) (ZRC 2018.775); C, non-ovigerous (4.6 × 4.2 mm) (ZRC 2018.772); D, spent (5.4 × 4.6 mm) (ZRC 2018.775); E, ovigerous (6.8 × 6.1 mm) (ZRC 2019.516); F, ovigerous (7.7 × 6.7 mm) (ZRC 2018.772); G, ovigerous (8.9 × 7.6 mm) (ZRC 2018.775). H–L, A. rayi Ahyong &amp; Ng, 2007, Peninsular Malaysia: H, juvenile (5.4 × 5.8 mm) (ZRC 2013.1416); I, non-ovigerous (5.6 × 4.3 mm) (ZRC 2013.1416); J, non-ovigerous (8.1 × 5.2 mm) (ZRC 2013.1416); K, spent (6.9 × 5.2 mm) (ZRC 2013.1416); L, ovigerous (8.4 × 7.0 mm) (ZRC 2013.1416). Scale = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.7174341" httpUri="https://zenodo.org/record/7174341/files/figure.png" pageId="60" pageNumber="194">50H–L</figureCitation>
, 
<figureCitation id="137A2A4BCE4DFFED33DF1A60D4BD79EC" box="[1215,1244,1903,1927]" captionStart="Fig" captionStartId="66.[113,148,1045,1067]" captionTargetBox="[113,1451,189,1030]" captionTargetId="figure-14@66.[113,1451,189,1030]" captionTargetPageId="66" captionText="Fig. 51. Arcotheres rayi Ahyong &amp; Ng, 2007, male (4.2 × 3.8 mm) (ZRC 2013.1416), Peninsular Malaysia. A, overall dorsal view; B, frontal view of cephalothorax; C, outer view of right chela." figureDoi="http://doi.org/10.5281/zenodo.7174343" httpUri="https://zenodo.org/record/7174343/files/figure.png" pageId="60" pageNumber="194">51</figureCitation>
, 
<figureCitation id="137A2A4BCE4DFFED33891A60D56479EC" box="[1257,1285,1903,1927]" captionStart="Fig" captionStartId="67.[136,171,2034,2056]" captionTargetBox="[143,1467,191,2017]" captionTargetId="figure-8@67.[134,1472,183,2020]" captionTargetPageId="67" captionText="Fig. 52. Arcotheres rayi Ahyong &amp; Ng, 2007, male (4.2 × 3.8 mm) (ZRC 2013.1416), Peninsular Malaysia. A, overall dorsal view; B, right MXP3; C, frontal view of cephalothorax; D, right chela; E–H, left P2–P5; I–L, right P2–P5; M, pleon; N, O, right G1 (dorsal and ventral views, respectively); P, right G2. Scale = A, 2.0 mm; B, N–P, 0.5 mm; C–M, 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.7174345" httpUri="https://zenodo.org/record/7174345/files/figure.png" pageId="60" pageNumber="194">52</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C35B6545CE4DFFED344D1AA1D441765D" pageId="60" pageNumber="194" type="reference_group">
<paragraph id="8BFE36CECE4DFFED344D1AA1D441765D" blockId="60.[813,1454,1966,2103]" pageId="60" pageNumber="194">
<treatmentCitationGroup id="AB5111E0CE4DFFED344D1AA1D441765D" pageId="60" pageNumber="194">
<treatmentCitation id="0AE010DFCE4DFFED344D1AA1D4B279A8" author="Ahyong ST &amp; Ng PKL" box="[813,1235,1966,1988]" page="203" pageId="60" pageNumber="194" year="2007">
<taxonomicName id="4C414D4DCE4DFFED344D1AA1D4B279A8" ID-CoL="67PY8" authority="Ahyong &amp; Ng, 2007 b: 203" authorityName="Ahyong &amp; Ng" authorityPageNumber="203" authorityYear="2007" box="[813,1235,1966,1988]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" kingdom="Animalia" order="Decapoda" pageId="60" pageNumber="194" phylum="Arthropoda" rank="species" species="rayi">
<emphasis id="B935EADCCE4DFFED344D1AA1D3A579A8" box="[813,964,1966,1987]" italics="true" pageId="60" pageNumber="194">Arcotheres rayi</emphasis>
<bibRefCitation id="EFD04B3FCE4DFFED34AB1AA1D4B279A8" author="Ahyong ST &amp; Ng PKL" box="[971,1235,1966,1988]" pageId="60" pageNumber="194" pagination="191 - 226" refId="ref62272" refString="Ahyong ST &amp; Ng PKL (2007 b) The pinnotherid type material of Semper (1880), Nauck (1880) and Burger (1895) (Crustacea: Decapoda: Brachyura). Raffles Bulletin of Zoology, Supplement 16: 191 - 226." type="journal article" year="2007">Ahyong &amp; Ng, 2007b: 203</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, fig. 11 [
<typeStatus id="54FA886CCE4DFFED324A1AA1D53779A8" box="[1322,1366,1966,1987]" pageId="60" pageNumber="194">type</typeStatus>
locality: Mariveles or 
<collectingRegion id="4985F82CCE4DFFED34B61AC5D472798B" box="[982,1043,1994,2016]" country="Philippines" name="Bohol" pageId="60" pageNumber="194">Bohol</collectingRegion>
, 
<collectingCountry id="F356765ECE4DFFED337F1AC5D4F0798B" box="[1055,1169,1994,2016]" name="Philippines" pageId="60" pageNumber="194">Philippines</collectingCountry>
]; 
<treatmentCitation id="0AE010DFCE4DFFED33C11AC4D50B798B" author="Ng PKL &amp; Guinot D &amp; Davie PJF" box="[1185,1386,1994,2016]" page="248" pageId="60" pageNumber="194" year="2008">
<bibRefCitation id="EFD04B3FCE4DFFED33C11AC4D50B798B" author="Ng PKL &amp; Guinot D &amp; Davie PJF" box="[1185,1386,1994,2016]" pageId="60" pageNumber="194" pagination="1 - 286" refId="ref67385" refString="Ng PKL, Guinot D &amp; Davie PJF (2008) Systema Brachyurorum: Part I. An annotated checklist of extant brachyuran crabs of the world. Raffles Bulletin of Zoology, Supplement 17: 1 - 286." type="journal article" year="2008">Ng et al., 2008: 248</bibRefCitation>
</treatmentCitation>
; 
<treatmentCitation id="0AE010DFCE4DFFED32161AC5D4517996" author="Clark PF &amp; Schram FR" page="450" pageId="60" pageNumber="194" year="2009">
<bibRefCitation id="EFD04B3FCE4DFFED32161AC5D4517996" author="Clark PF &amp; Schram FR" pageId="60" pageNumber="194" pagination="431 - 457" refId="ref63654" refString="Clark PF &amp; Schram FR (2009) Raymond B. Manning: An appreciation. Journal of Crustacean Biology, 29: 431 - 457." type="journal article" year="2009">Clark &amp; Schram, 2009: 450</bibRefCitation>
</treatmentCitation>
; 
<treatmentCitation id="0AE010DFCE4DFFED335D1AE7D5797997" author="Ng PKL &amp; Clark PF &amp; Mitra S &amp; Kumar AB" box="[1085,1304,2023,2045]" page="1094" pageId="60" pageNumber="194" year="2017">
<bibRefCitation id="EFD04B3FCE4DFFED335D1AE7D5797997" author="Ng PKL &amp; Clark PF &amp; Mitra S &amp; Kumar AB" box="[1085,1304,2023,2045]" pageId="60" pageNumber="194" pagination="1079 - 1097" refId="ref67243" refString="Ng PKL, Clark PF, Mitra S &amp; Kumar AB (2017) Arcotheres borradailei (Nobili, 1906) and Pinnotheres ridgewayi Southwell, 1911: A reassessment of characters and generic assignment of species to Arcotheres Manning, 1993 (Decapoda, Brachyura, Pinnotheridae). Crustaceana, 90 (7 - 10): 1079 - 1097." type="journal article" year="2017">Ng et al., 2017: 1094</bibRefCitation>
</treatmentCitation>
; 
<treatmentCitation id="0AE010DFCE4DFFED32451AE8D3D97671" author="Trivedi JN &amp; Campos E &amp; Vachhrajani KD" page="54" pageId="60" pageNumber="194" year="2018">
<bibRefCitation id="EFD04B3FCE4DFFED32451AE8D3D97671" author="Trivedi JN &amp; Campos E &amp; Vachhrajani KD" pageId="60" pageNumber="194" pagination="195 - 200" refId="ref69878" refString="Trivedi JN, Campos E &amp; Vachhrajani KD (2018 a) A new pea crab species of the genus Arcotheres Manning, 1993 (Crustacea: Decapoda: Brachyura: Pinnotheridae) from India. Zootaxa, 4433 (1): 195 - 200." type="journal article" year="2018">Trivedi et al., 2018a: 54</bibRefCitation>
</treatmentCitation>
, 56; 
<treatmentCitation id="0AE010DFCE4DFFED348E150BD4887671" author="Trivedi JN &amp; Gosavi S &amp; Vachhrajani KD &amp; Ng PKL" box="[1006,1257,2052,2074]" page="590" pageId="60" pageNumber="194" year="2019">
<bibRefCitation id="EFD04B3FCE4DFFED348E150BD4887671" author="Trivedi JN &amp; Gosavi S &amp; Vachhrajani KD &amp; Ng PKL" box="[1006,1257,2052,2074]" pageId="60" pageNumber="194" pagination="587 - 593" refId="ref69992" refString="Trivedi JN, Gosavi S, Vachhrajani KD &amp; Ng PKL (2019) Redescription of Arcotheres pernicola (Burger, 1895) (Crustacea: Decapoda: Brachyura: Pinnotheridae) from the oyster Magallana gryphoides (Schlotheim, 1820) in India. Zootaxa, 4706 (4): 587 - 593." type="journal article" year="2019">Trivedi et al., 2019: 590</bibRefCitation>
</treatmentCitation>
, tab. 1; 
<bibRefCitation id="EFD04B3FCE4DFFED325F150AD3B9765D" author="De Gier W &amp; Becker C" pageId="60" pageNumber="194" pagination="431" refId="ref64297" refString="De Gier W &amp; Becker C (2020) A review of the ecomorphology of pinnotherine pea crabs (Brachyura: Pinnotheridae), with an updated list of symbiont-host associations. Diversity, 12: 431." type="journal article" year="2020">De Gier &amp; Becker, 2020</bibRefCitation>
: tab. 1.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<caption id="DF3E6646CE4CFFEC37E819A7D2857AB0" ID-DOI="http://doi.org/10.5281/zenodo.7174331" ID-Zenodo-Dep="7174331" httpUri="https://zenodo.org/record/7174331/files/figure.png" pageId="61" pageNumber="195" startId="61.[136,171,1192,1214]" targetBox="[136,1472,189,1177]" targetPageId="61">
<paragraph id="8BFE36CECE4CFFEC37E819A7D2857AB0" blockId="61.[136,1472,1192,1243]" pageId="61" pageNumber="195">
Fig. 46. 
<taxonomicName id="4C414D4DCE4CFFEC37BD19A7D22D7AD6" authority="Ahyong &amp; Ng, 2007" authorityName="Ahyong &amp; Ng" authorityYear="2007" box="[221,588,1192,1214]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" kingdom="Animalia" order="Decapoda" pageId="61" pageNumber="195" phylum="Arthropoda" rank="species" species="rayi">
<emphasis id="B935EADCCE4CFFEC37BD19A7D1147AD5" box="[221,373,1192,1214]" italics="true" pageId="61" pageNumber="195">Arcotheres rayi</emphasis>
Ahyong &amp; Ng, 2007
</taxonomicName>
, holotype female (11.3 × 8.7 mm) (SMF-ZMG 952), Philippines. A, overall dorsal view; B, ventral view of cephalothorax; C, outer view of left chela.
</paragraph>
</caption>
<subSubSection id="C35B6545CE4CFFEC37E8181DD1E87821" pageId="61" pageNumber="195" type="materials_examined">
<paragraph id="8BFE36CECE4CFFEC37E8181DD2987B02" blockId="61.[136,775,1297,1386]" pageId="61" pageNumber="195">
<emphasis id="B935EADCCE4CFFEC37E8181DD1537B42" bold="true" box="[136,306,1297,1322]" pageId="61" pageNumber="195">Type material.</emphasis>
<materialsCitation id="3B293C93CE4CFFEC365B181ED2987B02" collectionCode="SMFZMG" collectorName="Semper" country="Philippines" location="Bohol" pageId="61" pageNumber="195" specimenCode="SMFZMG952" specimenCount="2" stateProvince="Bohol" typeStatus="holotype">
<typeStatus id="54FA886CCE4CFFEC365B181ED1C37B42" box="[315,418,1297,1321]" pageId="61" pageNumber="195" type="holotype">Holotype</typeStatus>
: female (11.3 × 
<quantity id="4CB99B2BCE4CFFEC353A181DD2CE7B41" box="[602,687,1298,1322]" metricMagnitude="-3" metricUnit="m" metricValue="8.7" pageId="61" pageNumber="195" unit="mm" value="8.7">8.7 mm</quantity>
) (
<specimenCode id="DBE79EB5CE4CFFEC35A7181DD0917B22" collectionCode="SMFZMG" pageId="61" pageNumber="195">SMFZMG952</specimenCode>
), Mariveles or 
<collectingRegion id="4985F82CCE4CFFEC36FD183ED1837B22" box="[413,482,1329,1353]" country="Philippines" name="Bohol" pageId="61" pageNumber="195">Bohol</collectingRegion>
, 
<collectingCountry id="F356765ECE4CFFEC3690183ED20C7B22" box="[496,621,1329,1353]" name="Philippines" pageId="61" pageNumber="195">Philippines</collectingCountry>
, coll. 
<collectorName id="26B45318CE4CFFEC35D5183DD3667B21" box="[693,775,1330,1354]" pageId="61" pageNumber="195">Semper</collectorName>
(
<typeStatus id="54FA886CCE4CFFEC37EF185ED1407B02" box="[143,289,1361,1385]" pageId="61" pageNumber="195" type="paralectotype">paralectotype</typeStatus>
of 
<taxonomicName id="4C414D4DCE4CFFEC362A185ED28D7B02" authority="Burger, 1895" authorityName="Burger" authorityYear="1895" box="[330,748,1361,1386]" class="Malacostraca" family="Xenophthalmidae" genus="Xenophthalmus" kingdom="Animalia" order="Decapoda" pageId="61" pageNumber="195" phylum="Arthropoda" rank="species" species="latifrons">
<emphasis id="B935EADCCE4CFFEC362A185ED2327B02" box="[330,595,1361,1385]" italics="true" pageId="61" pageNumber="195">Xenophthalmus latifrons</emphasis>
<bibRefCitation id="EFD04B3FCE4CFFEC353A185DD28D7B02" author="Burger O" box="[602,748,1361,1386]" pageId="61" pageNumber="195" pagination="361 - 390" refId="ref62795" refString="Burger O (1895) Ein Beitrag zur Kenntnis der Pinnotherinen. Zoologische Jahrbucher, Abtheilung fur Systematik, Geographie und Biologie der Thiere, 8: 361 - 390, pls. 9, 10." type="book chapter" year="1895">Bürger, 1895</bibRefCitation>
</taxonomicName>
).
</materialsCitation>
</paragraph>
<paragraph id="8BFE36CECE4CFFEC37E8189ED1E87821" blockId="61.[135,776,1425,1610]" pageId="61" pageNumber="195">
<emphasis id="B935EADCCE4CFFEC37E8189ED2527BC2" bold="true" box="[136,563,1425,1449]" pageId="61" pageNumber="195">Other material examined. Peninsular</emphasis>
<materialsCitation id="3B293C93CE4CFFEC355B189ED2BF7B81" country="Malaysia" location="Malaysia" pageId="61" pageNumber="195" specimenCount="3" specimenCount-male="3">
<emphasis id="B935EADCCE4CFFEC355B189ED2CD7BC2" bold="true" box="[571,684,1425,1449]" pageId="61" pageNumber="195">
<collectingCountry id="F356765ECE4CFFEC355B189ED2C67BC2" box="[571,679,1425,1449]" name="Malaysia" pageId="61" pageNumber="195">Malaysia</collectingCountry>
:
</emphasis>
<specimenCount id="9D47FD47CE4CFFEC35D3189DD3667BC2" box="[691,775,1425,1450]" pageId="61" pageNumber="195" type="male">3 males</specimenCount>
(4.2 × 3.8–4.4 × 4.0 mm), 8 ovigerous females (5.4 × 5.1–8.4 × 7.0 mm), 5 spent females (5.6 × 4.0–6.9 × 
<quantity id="4CB99B2BCE4CFFEC351E18DED2B47B81" box="[638,725,1489,1514]" metricMagnitude="-3" metricUnit="m" metricValue="5.2" pageId="61" pageNumber="195" unit="mm" value="5.2">5.2 mm</quantity>
)
</materialsCitation>
, 
<materialsCitation id="3B293C93CE4CFFEC358B18DDD1E47821" collectingDate="2010-05-10" collectionCode="ZRC" collectorName="Bentong &amp; M. Low" country="Malaysia" location="Pahang" pageId="61" pageNumber="195" specimenCode="ZRC 2013.1416" specimenCount="10" specimenCount-female="10" stateProvince="Pahang">
<specimenCount id="9D47FD47CE4CFFEC358B18DDD0BC7862" pageId="61" pageNumber="195" type="female">10 females</specimenCount>
with bopyrids (5.4 × 3.8–10.4 × 
<quantity id="4CB99B2BCE4CFFEC353F18FED2D97861" box="[607,696,1521,1546]" metricMagnitude="-3" metricUnit="m" metricValue="6.2" pageId="61" pageNumber="195" unit="mm" value="6.2">6.2 mm</quantity>
) (
<specimenCode id="DBE79EB5CE4CFFEC35B018FDD09F7842" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="61" pageNumber="195">ZRC 2013.1416</specimenCode>
), 
<collectorName id="26B45318CE4CFFEC36731B1DD10F7841" box="[275,366,1554,1578]" pageId="61" pageNumber="195">Bentong</collectorName>
market, 
<collectingRegion id="4985F82CCE4CFFEC36B31B1ED2457842" box="[467,548,1553,1577]" country="Malaysia" name="Pahang" pageId="61" pageNumber="195">Pahang</collectingRegion>
, in 
<taxonomicName id="4C414D4DCE4CFFEC35311B1ED2A87842" box="[593,713,1553,1577]" class="Bivalvia" family="Veneridae" genus="Paphia" kingdom="Animalia" order="Venerida" pageId="61" pageNumber="195" phylum="Mollusca" rank="species" species="undetermined">
<emphasis id="B935EADCCE4CFFEC35311B1ED2FF7842" box="[593,670,1553,1577]" italics="true" pageId="61" pageNumber="195">Paphia</emphasis>
sp.
</taxonomicName>
, coll. 
<collectorName id="26B45318CE4CFFEC37E81B3DD0837822" box="[136,226,1586,1609]" pageId="61" pageNumber="195">M. Low</collectorName>
, 
<date id="FFFF100ECE4CFFEC378F1B3DD1E47821" box="[239,389,1586,1610]" pageId="61" pageNumber="195" value="2010-05-10">
<collectingDate id="EFBBE9E6CE4CFFEC378F1B3DD1E47821" box="[239,389,1586,1610]" pageId="61" pageNumber="195" value="2010-05-10">10 May 2010</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C35B6545CE4CFF9037E81B7ED2CC7991" lastPageId="65" lastPageNumber="199" pageId="61" pageNumber="195" type="description">
<paragraph id="8BFE36CECE4CFFEC37E81B7ED1CD7661" blockId="61.[135,775,1649,2058]" pageId="61" pageNumber="195">
<emphasis id="B935EADCCE4CFFEC37E81B7ED17C78E2" bold="true" box="[136,285,1649,1673]" pageId="61" pageNumber="195">Description.</emphasis>
Female: Carapace and pereopods poorly chitinised, soft. Carapace subcircular to subhexagonal, wider than long; dorsal and lateral surfaces smooth, glabrous; appearing domed in frontal view; front in line with (larger specimens) or slightly produced anteriorly (smaller specimens) beyond transverse anterolateral margins, margin gently convex; anterior margin with (larger specimens) or without (smaller specimens) shallow notch above orbit flanking front (
<figureCitation id="137A2A4BCE4CFFEC364E1A7ED1FC79E2" box="[302,413,1905,1929]" captionStart="Fig" captionStartId="61.[136,171,1192,1214]" captionTargetBox="[136,1472,189,1177]" captionTargetId="figure-7@61.[136,1472,189,1177]" captionTargetPageId="61" captionText="Fig. 46. Arcotheres rayi Ahyong &amp; Ng, 2007, holotype female (11.3 × 8.7 mm) (SMF-ZMG 952), Philippines. A, overall dorsal view; B, ventral view of cephalothorax; C, outer view of left chela." figureDoi="http://doi.org/10.5281/zenodo.7174331" httpUri="https://zenodo.org/record/7174331/files/figure.png" pageId="61" pageNumber="195">Figs. 46A</figureCitation>
, 
<figureCitation id="137A2A4BCE4CFFEC36CB1A7DD24179E2" box="[427,544,1906,1930]" captionStart="Fig" captionStartId="62.[113,148,1974,1996]" captionTargetBox="[113,1451,279,1958]" captionTargetId="figure-14@62.[113,1451,279,1958]" captionTargetPageId="62" captionText="Fig. 47. Arcotheres rayi Ahyong &amp; Ng, 2007, female (6.9 × 5.2 mm) (ZRC 2013.1416), Peninsular Malaysia. A, overall dorsal view; B, ventral view of cephalothorax; C, dorsal view of carapace; D, frontal view of cephalothorax; E, outer view of left chela." figureDoi="http://doi.org/10.5281/zenodo.7174333" httpUri="https://zenodo.org/record/7174333/files/figure.png" pageId="61" pageNumber="195">47A, C, D</figureCitation>
, 
<figureCitation id="137A2A4BCE4CFFEC354F1A7DD21E79E1" box="[559,639,1906,1930]" captionStart="Fig" captionStartId="63.[135,170,1761,1783]" captionTargetBox="[135,1473,190,1746]" captionTargetId="figure-7@63.[135,1473,190,1746]" captionTargetPageId="63" captionText="Fig. 48. Arcotheres rayi Ahyong &amp; Ng, 2007. A, D, ovigerous female (7.6 × 5.8 mm) (ZRC 2013.1416), Peninsular Malaysia; B, C, E, female (6.2 × 4.4 mm) (ZRC 2013.1416), Peninsular Malaysia. A, B, overall dorsal view; C, dorsal view of carapace; D, E, right P5." figureDoi="http://doi.org/10.5281/zenodo.7174335" httpUri="https://zenodo.org/record/7174335/files/figure.png" pageId="61" pageNumber="195">48A–C</figureCitation>
, 
<figureCitation id="137A2A4BCE4CFFEC35EC1A7DD2DF79E1" box="[652,702,1906,1930]" captionStart="Fig" captionStartId="64.[113,148,1597,1619]" captionTargetBox="[117,1455,192,1580]" captionTargetId="figure-14@64.[107,1458,184,1592]" captionTargetPageId="64" captionText="Fig. 49. Arcotheres rayi Ahyong &amp; Ng, 2007, ovigerous female (7.6 × 5.8 mm) (ZRC 2013.1416), Peninsular Malaysia. A, overall dorsal view; B, right MXP3; C, frontal view of cephalothorax; D, anterior thoracic sternum; E, right chela; F–I, left P2–P5; J–M, right P2–P5. Scale = A, 2.0 mm; B, 0.5 mm; C–M, 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.7174339" httpUri="https://zenodo.org/record/7174339/files/figure.png" pageId="61" pageNumber="195">49A</figureCitation>
). Eyes small, not or just visible in dorsal view in adults; mobile, completely filling orbit (
<figureCitation id="137A2A4BCE4CFFEC36EA1ABED19779A2" box="[394,502,1969,1993]" captionStart="Fig" captionStartId="61.[136,171,1192,1214]" captionTargetBox="[136,1472,189,1177]" captionTargetId="figure-7@61.[136,1472,189,1177]" captionTargetPageId="61" captionText="Fig. 46. Arcotheres rayi Ahyong &amp; Ng, 2007, holotype female (11.3 × 8.7 mm) (SMF-ZMG 952), Philippines. A, overall dorsal view; B, ventral view of cephalothorax; C, outer view of left chela." figureDoi="http://doi.org/10.5281/zenodo.7174331" httpUri="https://zenodo.org/record/7174331/files/figure.png" pageId="61" pageNumber="195">Figs. 46A</figureCitation>
, 
<figureCitation id="137A2A4BCE4CFFEC35601ABDD20F79A2" box="[512,622,1970,1994]" captionStart="Fig" captionStartId="62.[113,148,1974,1996]" captionTargetBox="[113,1451,279,1958]" captionTargetId="figure-14@62.[113,1451,279,1958]" captionTargetPageId="62" captionText="Fig. 47. Arcotheres rayi Ahyong &amp; Ng, 2007, female (6.9 × 5.2 mm) (ZRC 2013.1416), Peninsular Malaysia. A, overall dorsal view; B, ventral view of cephalothorax; C, dorsal view of carapace; D, frontal view of cephalothorax; E, outer view of left chela." figureDoi="http://doi.org/10.5281/zenodo.7174333" httpUri="https://zenodo.org/record/7174333/files/figure.png" pageId="61" pageNumber="195">47A, C, D</figureCitation>
, 
<figureCitation id="137A2A4BCE4CFFEC35191ABDD2A879A1" box="[633,713,1970,1994]" captionStart="Fig" captionStartId="63.[135,170,1761,1783]" captionTargetBox="[135,1473,190,1746]" captionTargetId="figure-7@63.[135,1473,190,1746]" captionTargetPageId="63" captionText="Fig. 48. Arcotheres rayi Ahyong &amp; Ng, 2007. A, D, ovigerous female (7.6 × 5.8 mm) (ZRC 2013.1416), Peninsular Malaysia; B, C, E, female (6.2 × 4.4 mm) (ZRC 2013.1416), Peninsular Malaysia. A, B, overall dorsal view; C, dorsal view of carapace; D, E, right P5." figureDoi="http://doi.org/10.5281/zenodo.7174335" httpUri="https://zenodo.org/record/7174335/files/figure.png" pageId="61" pageNumber="195">48A–C</figureCitation>
, 
<figureCitation id="137A2A4BCE4CFFEC35B21ABDD0FC7981" captionStart="Fig" captionStartId="64.[113,148,1597,1619]" captionTargetBox="[117,1455,192,1580]" captionTargetId="figure-14@64.[107,1458,184,1592]" captionTargetPageId="64" captionText="Fig. 49. Arcotheres rayi Ahyong &amp; Ng, 2007, ovigerous female (7.6 × 5.8 mm) (ZRC 2013.1416), Peninsular Malaysia. A, overall dorsal view; B, right MXP3; C, frontal view of cephalothorax; D, anterior thoracic sternum; E, right chela; F–I, left P2–P5; J–M, right P2–P5. Scale = A, 2.0 mm; B, 0.5 mm; C–M, 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.7174339" httpUri="https://zenodo.org/record/7174339/files/figure.png" pageId="61" pageNumber="195">49A, C</figureCitation>
). Epistome with median part triangular, lateral margins gently concave (
<figureCitation id="137A2A4BCE4CFFEC365B1AFED1C17661" box="[315,416,2033,2058]" captionStart="Fig" captionStartId="64.[113,148,1597,1619]" captionTargetBox="[117,1455,192,1580]" captionTargetId="figure-14@64.[107,1458,184,1592]" captionTargetPageId="64" captionText="Fig. 49. Arcotheres rayi Ahyong &amp; Ng, 2007, ovigerous female (7.6 × 5.8 mm) (ZRC 2013.1416), Peninsular Malaysia. A, overall dorsal view; B, right MXP3; C, frontal view of cephalothorax; D, anterior thoracic sternum; E, right chela; F–I, left P2–P5; J–M, right P2–P5. Scale = A, 2.0 mm; B, 0.5 mm; C–M, 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.7174339" httpUri="https://zenodo.org/record/7174339/files/figure.png" pageId="61" pageNumber="195">Fig. 49C</figureCitation>
).
</paragraph>
<paragraph id="8BFE36CECE4CFFEC3423181DD5D57861" blockId="61.[834,1474,1298,1546]" pageId="61" pageNumber="195">
MXP3 outer surface with scattered short setae; ischiomerus completely fused, subrhomboidal, inner margin usually rounded, at widest point; carpus short; propodus about 3 times as long as high, subspatulate, distinctly longer than carpus, tip rounded to subtruncate; dactylus slender, inserted medially on propodus, tip underreaching or just reaching propodal apex; exopod relatively slender, about two-thirds length of ischiomerus, flagellum 2-segmented (
<figureCitation id="137A2A4BCE4CFFEC322418FDD5C97861" box="[1348,1448,1522,1546]" captionStart="Fig" captionStartId="64.[113,148,1597,1619]" captionTargetBox="[117,1455,192,1580]" captionTargetId="figure-14@64.[107,1458,184,1592]" captionTargetPageId="64" captionText="Fig. 49. Arcotheres rayi Ahyong &amp; Ng, 2007, ovigerous female (7.6 × 5.8 mm) (ZRC 2013.1416), Peninsular Malaysia. A, overall dorsal view; B, right MXP3; C, frontal view of cephalothorax; D, anterior thoracic sternum; E, right chela; F–I, left P2–P5; J–M, right P2–P5. Scale = A, 2.0 mm; B, 0.5 mm; C–M, 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.7174339" httpUri="https://zenodo.org/record/7174339/files/figure.png" pageId="61" pageNumber="195">Fig. 49B</figureCitation>
).
</paragraph>
<paragraph id="8BFE36CECE4CFFEC34231B3DD54F7921" blockId="61.[834,1475,1586,1866]" pageId="61" pageNumber="195">
Chela not prominently elongate, dactylus half to two-thirds palm length; palm relatively slender, proximally narrower than distally; outer surfaces of palm, fingers (except for distal part) almost glabrous, with only scattered short setae; ventral margin of palm gently convex to sinuous; dactylus occlusal margin with distinct subproximal tooth; pollex occlusal margin usually with 1 low proximal tooth (sometimes obsolete), 1 submedian tooth, and minute denticles; tips of fingers sharp, hooked (
<figureCitation id="137A2A4BCE4CFFEC335F1A3DD4CD7921" box="[1087,1196,1842,1866]" captionStart="Fig" captionStartId="61.[136,171,1192,1214]" captionTargetBox="[136,1472,189,1177]" captionTargetId="figure-7@61.[136,1472,189,1177]" captionTargetPageId="61" captionText="Fig. 46. Arcotheres rayi Ahyong &amp; Ng, 2007, holotype female (11.3 × 8.7 mm) (SMF-ZMG 952), Philippines. A, overall dorsal view; B, ventral view of cephalothorax; C, outer view of left chela." figureDoi="http://doi.org/10.5281/zenodo.7174331" httpUri="https://zenodo.org/record/7174331/files/figure.png" pageId="61" pageNumber="195">Figs. 46C</figureCitation>
, 
<figureCitation id="137A2A4BCE4CFFEC33D91A3DD4877921" box="[1209,1254,1842,1866]" captionStart="Fig" captionStartId="62.[113,148,1974,1996]" captionTargetBox="[113,1451,279,1958]" captionTargetId="figure-14@62.[113,1451,279,1958]" captionTargetPageId="62" captionText="Fig. 47. Arcotheres rayi Ahyong &amp; Ng, 2007, female (6.9 × 5.2 mm) (ZRC 2013.1416), Peninsular Malaysia. A, overall dorsal view; B, ventral view of cephalothorax; C, dorsal view of carapace; D, frontal view of cephalothorax; E, outer view of left chela." figureDoi="http://doi.org/10.5281/zenodo.7174333" httpUri="https://zenodo.org/record/7174333/files/figure.png" pageId="61" pageNumber="195">47E</figureCitation>
, 
<figureCitation id="137A2A4BCE4CFFEC33931A3DD5437921" box="[1267,1314,1842,1866]" captionStart="Fig" captionStartId="64.[113,148,1597,1619]" captionTargetBox="[117,1455,192,1580]" captionTargetId="figure-14@64.[107,1458,184,1592]" captionTargetPageId="64" captionText="Fig. 49. Arcotheres rayi Ahyong &amp; Ng, 2007, ovigerous female (7.6 × 5.8 mm) (ZRC 2013.1416), Peninsular Malaysia. A, overall dorsal view; B, right MXP3; C, frontal view of cephalothorax; D, anterior thoracic sternum; E, right chela; F–I, left P2–P5; J–M, right P2–P5. Scale = A, 2.0 mm; B, 0.5 mm; C–M, 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.7174339" httpUri="https://zenodo.org/record/7174339/files/figure.png" pageId="61" pageNumber="195">49E</figureCitation>
).
</paragraph>
<paragraph id="8BFE36CECE4CFFEE34231A7DD2667648" blockId="61.[834,1475,1906,2090]" lastBlockId="63.[136,775,1867,2083]" lastPageId="63" lastPageNumber="197" pageId="61" pageNumber="195">
P2–P5 dorsally, ventrally unarmed; outer surface covered with scattered, very short setae or glabrous; ventral margins of propodus and dactylus slightly more setose; merus relatively longer, more slender, relative lengths of meri P4&gt;P3&gt;P5&gt;P2; left (sometimes right) P4 distinctly the longer; P2 and P3 dactyli short, subequal, tip gently hooked; P4 longer dactylus elongate, weakly falciform, distinctly longer than half propodus length, longer than P5 dactylus; P5 merus 3.9–4.5 times longer than wide; P5 dactylus shorter than propodus extensor margin, longer than P2 and P3 dactyli, length/height &lt;5.0, margins lined with short and long setae, denser on ventral margin, distoflexor margin without rows of spinules (
<figureCitation id="137A2A4BCE4EFFEE366F1504D1FF7648" box="[271,414,2059,2083]" captionStart="Fig" captionStartId="64.[113,148,1597,1619]" captionTargetBox="[117,1455,192,1580]" captionTargetId="figure-14@64.[107,1458,184,1592]" captionTargetPageId="64" captionText="Fig. 49. Arcotheres rayi Ahyong &amp; Ng, 2007, ovigerous female (7.6 × 5.8 mm) (ZRC 2013.1416), Peninsular Malaysia. A, overall dorsal view; B, right MXP3; C, frontal view of cephalothorax; D, anterior thoracic sternum; E, right chela; F–I, left P2–P5; J–M, right P2–P5. Scale = A, 2.0 mm; B, 0.5 mm; C–M, 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.7174339" httpUri="https://zenodo.org/record/7174339/files/figure.png" pageId="63" pageNumber="197">Figs. 49F–M</figureCitation>
, 
<figureCitation id="137A2A4BCE4EFFEE36CB1504D19A7648" box="[427,507,2059,2083]" captionStart="Fig" captionStartId="65.[135,170,1541,1563]" captionTargetBox="[137,1477,189,1531]" captionTargetId="figure-7@65.[132,1485,182,1535]" captionTargetPageId="65" captionText="Fig. 50. Right female P5. A–G, Arcotheres exiguus (Bürger, 1895), Peninsular Malaysia: A, juvenile (3.8 × 3.6 mm) (ZRC 2018.772); B, ovigerous (4.5 × 4.1 mm) (ZRC 2018.775); C, non-ovigerous (4.6 × 4.2 mm) (ZRC 2018.772); D, spent (5.4 × 4.6 mm) (ZRC 2018.775); E, ovigerous (6.8 × 6.1 mm) (ZRC 2019.516); F, ovigerous (7.7 × 6.7 mm) (ZRC 2018.772); G, ovigerous (8.9 × 7.6 mm) (ZRC 2018.775). H–L, A. rayi Ahyong &amp; Ng, 2007, Peninsular Malaysia: H, juvenile (5.4 × 5.8 mm) (ZRC 2013.1416); I, non-ovigerous (5.6 × 4.3 mm) (ZRC 2013.1416); J, non-ovigerous (8.1 × 5.2 mm) (ZRC 2013.1416); K, spent (6.9 × 5.2 mm) (ZRC 2013.1416); L, ovigerous (8.4 × 7.0 mm) (ZRC 2013.1416). Scale = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.7174341" httpUri="https://zenodo.org/record/7174341/files/figure.png" pageId="63" pageNumber="197">50H–L</figureCitation>
).
</paragraph>
<caption id="DF3E6646CE4FFFEF37111AB9D5727982" ID-DOI="http://doi.org/10.5281/zenodo.7174333" ID-Zenodo-Dep="7174333" httpUri="https://zenodo.org/record/7174333/files/figure.png" pageId="62" pageNumber="196" startId="62.[113,148,1974,1996]" targetBox="[113,1451,279,1958]" targetPageId="62">
<paragraph id="8BFE36CECE4FFFEF37111AB9D5727982" blockId="62.[113,1451,1974,2025]" pageId="62" pageNumber="196">
Fig. 47. 
<taxonomicName id="4C414D4DCE4FFFEF37A61AB9D25079A0" authority="Ahyong &amp; Ng, 2007" authorityName="Ahyong &amp; Ng" authorityYear="2007" box="[198,561,1974,1996]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" kingdom="Animalia" order="Decapoda" pageId="62" pageNumber="196" phylum="Arthropoda" rank="species" species="rayi">
<emphasis id="B935EADCCE4FFFEF37A61AB9D13F79A0" box="[198,350,1974,1995]" italics="true" pageId="62" pageNumber="196">Arcotheres rayi</emphasis>
Ahyong &amp; Ng, 2007
</taxonomicName>
, female (6.9 × 5.2 mm) (ZRC 2013.1416), Peninsular Malaysia. A, overall dorsal view; B, ventral view of cephalothorax; C, dorsal view of carapace; D, frontal view of cephalothorax; E, outer view of left chela.
</paragraph>
</caption>
<caption id="DF3E6646CE4EFFEE37E71BEED5C9797F" ID-DOI="http://doi.org/10.5281/zenodo.7174335" ID-Zenodo-Dep="7174335" httpUri="https://zenodo.org/record/7174335/files/figure.png" pageId="63" pageNumber="197" startId="63.[135,170,1761,1783]" targetBox="[135,1473,190,1746]" targetPageId="63">
<paragraph id="8BFE36CECE4EFFEE37E71BEED5C9797F" blockId="63.[135,1474,1761,1812]" pageId="63" pageNumber="197">
Fig. 48. 
<taxonomicName id="4C414D4DCE4EFFEE37BB1BEED22B789D" authority="Ahyong &amp; Ng, 2007" authorityName="Ahyong &amp; Ng" authorityYear="2007" box="[219,586,1761,1783]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" kingdom="Animalia" order="Decapoda" pageId="63" pageNumber="197" phylum="Arthropoda" rank="species" species="rayi">
<emphasis id="B935EADCCE4EFFEE37BB1BEED115789C" box="[219,372,1761,1783]" italics="true" pageId="63" pageNumber="197">Arcotheres rayi</emphasis>
Ahyong &amp; Ng, 2007
</taxonomicName>
. A, D, ovigerous female (7.6 × 5.8 mm) (ZRC 2013.1416), Peninsular Malaysia; B, C, E, female (6.2 × 4.4 mm) (ZRC 2013.1416), Peninsular Malaysia. A, B, overall dorsal view; C, dorsal view of carapace; D, E, right P5.
</paragraph>
</caption>
<paragraph id="8BFE36CECE4EFFEE34231A44D43779C8" blockId="63.[835,1474,1867,1955]" pageId="63" pageNumber="197">
Pleon extending to buccal region, covering bases of P2–P5; telson gently recessed into concave distal margin of somite 6 (
<figureCitation id="137A2A4BCE4EFFEE34021A84D3B179C8" box="[866,976,1931,1955]" captionStart="Fig" captionStartId="61.[136,171,1192,1214]" captionTargetBox="[136,1472,189,1177]" captionTargetId="figure-7@61.[136,1472,189,1177]" captionTargetPageId="61" captionText="Fig. 46. Arcotheres rayi Ahyong &amp; Ng, 2007, holotype female (11.3 × 8.7 mm) (SMF-ZMG 952), Philippines. A, overall dorsal view; B, ventral view of cephalothorax; C, outer view of left chela." figureDoi="http://doi.org/10.5281/zenodo.7174331" httpUri="https://zenodo.org/record/7174331/files/figure.png" pageId="63" pageNumber="197">Figs. 46B</figureCitation>
, 
<figureCitation id="137A2A4BCE4EFFEE34BC1A84D46A79C8" box="[988,1035,1931,1955]" captionStart="Fig" captionStartId="62.[113,148,1974,1996]" captionTargetBox="[113,1451,279,1958]" captionTargetId="figure-14@62.[113,1451,279,1958]" captionTargetPageId="62" captionText="Fig. 47. Arcotheres rayi Ahyong &amp; Ng, 2007, female (6.9 × 5.2 mm) (ZRC 2013.1416), Peninsular Malaysia. A, overall dorsal view; B, ventral view of cephalothorax; C, dorsal view of carapace; D, frontal view of cephalothorax; E, outer view of left chela." figureDoi="http://doi.org/10.5281/zenodo.7174333" httpUri="https://zenodo.org/record/7174333/files/figure.png" pageId="63" pageNumber="197">47B</figureCitation>
, 
<figureCitation id="137A2A4BCE4EFFEE33781A84D42A79C8" box="[1048,1099,1931,1955]" captionStart="Fig" captionStartId="64.[113,148,1597,1619]" captionTargetBox="[117,1455,192,1580]" captionTargetId="figure-14@64.[107,1458,184,1592]" captionTargetPageId="64" captionText="Fig. 49. Arcotheres rayi Ahyong &amp; Ng, 2007, ovigerous female (7.6 × 5.8 mm) (ZRC 2013.1416), Peninsular Malaysia. A, overall dorsal view; B, right MXP3; C, frontal view of cephalothorax; D, anterior thoracic sternum; E, right chela; F–I, left P2–P5; J–M, right P2–P5. Scale = A, 2.0 mm; B, 0.5 mm; C–M, 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.7174339" httpUri="https://zenodo.org/record/7174339/files/figure.png" pageId="63" pageNumber="197">49D</figureCitation>
).
</paragraph>
<paragraph id="8BFE36CECE4EFF9134231AC4D4A879D7" blockId="63.[835,1475,1995,2083]" lastBlockId="64.[812,1452,1732,1980]" lastPageId="64" lastPageNumber="198" pageId="63" pageNumber="197">
<collectingRegion id="4985F82CCE4EFFEE34231AC4D31C7988" box="[835,893,1995,2019]" country="Maldives" name="Maale" pageId="63" pageNumber="197">Male</collectingRegion>
: Carapace and pereopods well chitinised, firm. Carapace almost circular, slightly wider than long; dorsal surface almost smooth, not prominently inflated, lateral surfaces with setae; front projecting anteriorly, margin gently sinuous to almost straight (
<figureCitation id="137A2A4BCE31FF9136451BEBD1F57897" box="[293,404,1764,1788]" captionStart="Fig" captionStartId="66.[113,148,1045,1067]" captionTargetBox="[113,1451,189,1030]" captionTargetId="figure-14@66.[113,1451,189,1030]" captionTargetPageId="66" captionText="Fig. 51. Arcotheres rayi Ahyong &amp; Ng, 2007, male (4.2 × 3.8 mm) (ZRC 2013.1416), Peninsular Malaysia. A, overall dorsal view; B, frontal view of cephalothorax; C, outer view of right chela." figureDoi="http://doi.org/10.5281/zenodo.7174343" httpUri="https://zenodo.org/record/7174343/files/figure.png" pageId="64" pageNumber="198">Figs. 51A</figureCitation>
, 
<figureCitation id="137A2A4BCE31FF9136C31BEBD1B47897" box="[419,469,1764,1788]" captionStart="Fig" captionStartId="67.[136,171,2034,2056]" captionTargetBox="[143,1467,191,2017]" captionTargetId="figure-8@67.[134,1472,183,2020]" captionTargetPageId="67" captionText="Fig. 52. Arcotheres rayi Ahyong &amp; Ng, 2007, male (4.2 × 3.8 mm) (ZRC 2013.1416), Peninsular Malaysia. A, overall dorsal view; B, right MXP3; C, frontal view of cephalothorax; D, right chela; E–H, left P2–P5; I–L, right P2–P5; M, pleon; N, O, right G1 (dorsal and ventral views, respectively); P, right G2. Scale = A, 2.0 mm; B, N–P, 0.5 mm; C–M, 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.7174345" httpUri="https://zenodo.org/record/7174345/files/figure.png" pageId="64" pageNumber="198">52A</figureCitation>
). MXP3 as in female but dactylus shorter (
<figureCitation id="137A2A4BCE31FF9136491A0BD1EB7977" box="[297,394,1796,1820]" captionStart="Fig" captionStartId="67.[136,171,2034,2056]" captionTargetBox="[143,1467,191,2017]" captionTargetId="figure-8@67.[134,1472,183,2020]" captionTargetPageId="67" captionText="Fig. 52. Arcotheres rayi Ahyong &amp; Ng, 2007, male (4.2 × 3.8 mm) (ZRC 2013.1416), Peninsular Malaysia. A, overall dorsal view; B, right MXP3; C, frontal view of cephalothorax; D, right chela; E–H, left P2–P5; I–L, right P2–P5; M, pleon; N, O, right G1 (dorsal and ventral views, respectively); P, right G2. Scale = A, 2.0 mm; B, N–P, 0.5 mm; C–M, 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.7174345" httpUri="https://zenodo.org/record/7174345/files/figure.png" pageId="64" pageNumber="198">Fig. 52B</figureCitation>
). Anterior thoracic sternum wide, sternites 1, 2 fused, partially sunken into buccal cavity; suture between sternites 2 and 3 shallow; sternites 3, 4 completely fused, separated only by shallow grooves. Chela relatively stout, shorter than in female (
<figureCitation id="137A2A4BCE31FF9135521A8BD2C279F7" box="[562,675,1924,1948]" captionStart="Fig" captionStartId="66.[113,148,1045,1067]" captionTargetBox="[113,1451,189,1030]" captionTargetId="figure-14@66.[113,1451,189,1030]" captionTargetPageId="66" captionText="Fig. 51. Arcotheres rayi Ahyong &amp; Ng, 2007, male (4.2 × 3.8 mm) (ZRC 2013.1416), Peninsular Malaysia. A, overall dorsal view; B, frontal view of cephalothorax; C, outer view of right chela." figureDoi="http://doi.org/10.5281/zenodo.7174343" httpUri="https://zenodo.org/record/7174343/files/figure.png" pageId="64" pageNumber="198">Figs. 51C</figureCitation>
, 
<figureCitation id="137A2A4BCE31FF9135D21A8BD28579F7" box="[690,740,1924,1948]" captionStart="Fig" captionStartId="67.[136,171,2034,2056]" captionTargetBox="[143,1467,191,2017]" captionTargetId="figure-8@67.[134,1472,183,2020]" captionTargetPageId="67" captionText="Fig. 52. Arcotheres rayi Ahyong &amp; Ng, 2007, male (4.2 × 3.8 mm) (ZRC 2013.1416), Peninsular Malaysia. A, overall dorsal view; B, right MXP3; C, frontal view of cephalothorax; D, right chela; E–H, left P2–P5; I–L, right P2–P5; M, pleon; N, O, right G1 (dorsal and ventral views, respectively); P, right G2. Scale = A, 2.0 mm; B, N–P, 0.5 mm; C–M, 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.7174345" httpUri="https://zenodo.org/record/7174345/files/figure.png" pageId="64" pageNumber="198">52D</figureCitation>
). P2–P5 dorsally, ventrally unarmed; outer surface covered with short setae; P3 and P4 carpus and propodus with long natatory setae; left and right meri equal, relative lengths of meri P4&gt;P3&gt;P2&gt;P5; dactyli of P2–P4 progressively longer; right P4 dactylus slightly longer, more slender than left side; P4 and P5 dactylus subequal to that of P3, covered with short setae (
<figureCitation id="137A2A4BCE31FF9134D31BEBD4547897" box="[947,1077,1764,1788]" captionStart="Fig" captionStartId="67.[136,171,2034,2056]" captionTargetBox="[143,1467,191,2017]" captionTargetId="figure-8@67.[134,1472,183,2020]" captionTargetPageId="67" captionText="Fig. 52. Arcotheres rayi Ahyong &amp; Ng, 2007, male (4.2 × 3.8 mm) (ZRC 2013.1416), Peninsular Malaysia. A, overall dorsal view; B, right MXP3; C, frontal view of cephalothorax; D, right chela; E–H, left P2–P5; I–L, right P2–P5; M, pleon; N, O, right G1 (dorsal and ventral views, respectively); P, right G2. Scale = A, 2.0 mm; B, N–P, 0.5 mm; C–M, 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.7174345" httpUri="https://zenodo.org/record/7174345/files/figure.png" pageId="64" pageNumber="198">Fig. 52E–L</figureCitation>
). Pleon slender, triangular, widest at somite 3, lateral margins of somite 4 gently concave; somite 6 trapezoidal; telson semicircular, wider than long (
<figureCitation id="137A2A4BCE31FF9134551A4BD3C17937" box="[821,928,1860,1884]" captionStart="Fig" captionStartId="67.[136,171,2034,2056]" captionTargetBox="[143,1467,191,2017]" captionTargetId="figure-8@67.[134,1472,183,2020]" captionTargetPageId="67" captionText="Fig. 52. Arcotheres rayi Ahyong &amp; Ng, 2007, male (4.2 × 3.8 mm) (ZRC 2013.1416), Peninsular Malaysia. A, overall dorsal view; B, right MXP3; C, frontal view of cephalothorax; D, right chela; E–H, left P2–P5; I–L, right P2–P5; M, pleon; N, O, right G1 (dorsal and ventral views, respectively); P, right G2. Scale = A, 2.0 mm; B, N–P, 0.5 mm; C–M, 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.7174345" httpUri="https://zenodo.org/record/7174345/files/figure.png" pageId="64" pageNumber="198">Fig. 52M</figureCitation>
). G1 relatively stout, arcuate, curved outwards, with short triangular subdistal dorsal projection, tip rounded (
<figureCitation id="137A2A4BCE31FF9134541A8BD3D979F7" box="[820,952,1924,1948]" captionStart="Fig" captionStartId="67.[136,171,2034,2056]" captionTargetBox="[143,1467,191,2017]" captionTargetId="figure-8@67.[134,1472,183,2020]" captionTargetPageId="67" captionText="Fig. 52. Arcotheres rayi Ahyong &amp; Ng, 2007, male (4.2 × 3.8 mm) (ZRC 2013.1416), Peninsular Malaysia. A, overall dorsal view; B, right MXP3; C, frontal view of cephalothorax; D, right chela; E–H, left P2–P5; I–L, right P2–P5; M, pleon; N, O, right G1 (dorsal and ventral views, respectively); P, right G2. Scale = A, 2.0 mm; B, N–P, 0.5 mm; C–M, 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.7174345" httpUri="https://zenodo.org/record/7174345/files/figure.png" pageId="64" pageNumber="198">Fig. 52N, O</figureCitation>
). G2 short, with spatuliform tip; exopod about two-thirds endopod length (
<figureCitation id="137A2A4BCE31FF91333B1AABD4DD79D7" box="[1115,1212,1956,1980]" captionStart="Fig" captionStartId="67.[136,171,2034,2056]" captionTargetBox="[143,1467,191,2017]" captionTargetId="figure-8@67.[134,1472,183,2020]" captionTargetPageId="67" captionText="Fig. 52. Arcotheres rayi Ahyong &amp; Ng, 2007, male (4.2 × 3.8 mm) (ZRC 2013.1416), Peninsular Malaysia. A, overall dorsal view; B, right MXP3; C, frontal view of cephalothorax; D, right chela; E–H, left P2–P5; I–L, right P2–P5; M, pleon; N, O, right G1 (dorsal and ventral views, respectively); P, right G2. Scale = A, 2.0 mm; B, N–P, 0.5 mm; C–M, 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.7174345" httpUri="https://zenodo.org/record/7174345/files/figure.png" pageId="64" pageNumber="198">Fig. 52P</figureCitation>
).
</paragraph>
<caption id="DF3E6646CE31FF9137111B32D22778E7" ID-DOI="http://doi.org/10.5281/zenodo.7174339" ID-Zenodo-Dep="7174339" httpUri="https://zenodo.org/record/7174339/files/figure.png" pageId="64" pageNumber="198" startId="64.[113,148,1597,1619]" targetBox="[117,1455,192,1580]" targetPageId="64">
<paragraph id="8BFE36CECE31FF9137111B32D22778E7" blockId="64.[113,1453,1597,1677]" pageId="64" pageNumber="198">
Fig. 49. 
<taxonomicName id="4C414D4DCE31FF9137A41B32D24D7839" authority="Ahyong &amp; Ng, 2007" authorityName="Ahyong &amp; Ng" authorityYear="2007" box="[196,556,1597,1619]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" kingdom="Animalia" order="Decapoda" pageId="64" pageNumber="198" phylum="Arthropoda" rank="species" species="rayi">
<emphasis id="B935EADCCE31FF9137A41B32D13A7838" box="[196,347,1597,1619]" italics="true" pageId="64" pageNumber="198">Arcotheres rayi</emphasis>
Ahyong &amp; Ng, 2007
</taxonomicName>
, ovigerous female (7.6 × 5.8 mm) (ZRC 2013.1416), Peninsular Malaysia. A, overall dorsal view; B, right MXP3; C, frontal view of cephalothorax; D, anterior thoracic sternum; E, right chela; F–I, left P2–P5; J–M, right P2–P5. Scale = A, 2.0 mm; B, 0.5 mm; C–M, 1.0 mm.
</paragraph>
</caption>
<paragraph id="8BFE36CECE31FF90344D1AEBD2CC7991" blockId="64.[811,1452,2020,2108]" lastBlockId="65.[135,777,1763,2043]" lastPageId="65" lastPageNumber="199" pageId="64" pageNumber="198">
<emphasis id="B935EADCCE31FF91344D1AEBD3C07997" bold="true" box="[813,929,2020,2044]" pageId="64" pageNumber="198">Variation.</emphasis>
The carapace shape varies somewhat, the most important being the prominence of the front, whose variation is broadly size-related. In the 
<typeStatus id="54FA886CCE31FF913307152BD4A57657" box="[1127,1220,2084,2108]" pageId="64" pageNumber="198" type="holotype">holotype</typeStatus>
of 
<taxonomicName id="4C414D4DCE31FF913388152BD5517650" authorityName="Ahyong &amp; Ng" authorityYear="2007" box="[1256,1328,2084,2107]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" kingdom="Animalia" order="Decapoda" pageId="64" pageNumber="198" phylum="Arthropoda" rank="species" species="rayi">
<emphasis id="B935EADCCE31FF913388152BD5517650" box="[1256,1328,2084,2107]" italics="true" pageId="64" pageNumber="198">A. rayi</emphasis>
</taxonomicName>
, the largest known specimen of the species, the front does not protrude in advance of the general anterior margin, but is demarcated on either side by a shallow notch above each orbit (
<figureCitation id="137A2A4BCE30FF9035BA1A2CD0C67931" captionStart="Fig" captionStartId="61.[136,171,1192,1214]" captionTargetBox="[136,1472,189,1177]" captionTargetId="figure-7@61.[136,1472,189,1177]" captionTargetPageId="61" captionText="Fig. 46. Arcotheres rayi Ahyong &amp; Ng, 2007, holotype female (11.3 × 8.7 mm) (SMF-ZMG 952), Philippines. A, overall dorsal view; B, ventral view of cephalothorax; C, outer view of left chela." figureDoi="http://doi.org/10.5281/zenodo.7174331" httpUri="https://zenodo.org/record/7174331/files/figure.png" pageId="65" pageNumber="199">Fig. 46</figureCitation>
). Other specimens, however, have a front that protrudes anteriorly beyond the general anterior margin, some retaining the notch on either side of the front (
<figureCitation id="137A2A4BCE30FF9035721A8CD2F279F0" box="[530,659,1923,1947]" captionStart="Fig" captionStartId="63.[135,170,1761,1783]" captionTargetBox="[135,1473,190,1746]" captionTargetId="figure-7@63.[135,1473,190,1746]" captionTargetPageId="63" captionText="Fig. 48. Arcotheres rayi Ahyong &amp; Ng, 2007. A, D, ovigerous female (7.6 × 5.8 mm) (ZRC 2013.1416), Peninsular Malaysia; B, C, E, female (6.2 × 4.4 mm) (ZRC 2013.1416), Peninsular Malaysia. A, B, overall dorsal view; C, dorsal view of carapace; D, E, right P5." figureDoi="http://doi.org/10.5281/zenodo.7174335" httpUri="https://zenodo.org/record/7174335/files/figure.png" pageId="65" pageNumber="199">Fig. 48B, C</figureCitation>
), but most others without (
<figureCitation id="137A2A4BCE30FF9036521AACD1EA79D0" box="[306,395,1955,1979]" captionStart="Fig" captionStartId="62.[113,148,1974,1996]" captionTargetBox="[113,1451,279,1958]" captionTargetId="figure-14@62.[113,1451,279,1958]" captionTargetPageId="62" captionText="Fig. 47. Arcotheres rayi Ahyong &amp; Ng, 2007, female (6.9 × 5.2 mm) (ZRC 2013.1416), Peninsular Malaysia. A, overall dorsal view; B, ventral view of cephalothorax; C, dorsal view of carapace; D, frontal view of cephalothorax; E, outer view of left chela." figureDoi="http://doi.org/10.5281/zenodo.7174333" httpUri="https://zenodo.org/record/7174333/files/figure.png" pageId="65" pageNumber="199">Figs. 47</figureCitation>
, 
<figureCitation id="137A2A4BCE30FF9036F71AACD1A979D0" box="[407,456,1955,1979]" captionStart="Fig" captionStartId="63.[135,170,1761,1783]" captionTargetBox="[135,1473,190,1746]" captionTargetId="figure-7@63.[135,1473,190,1746]" captionTargetPageId="63" captionText="Fig. 48. Arcotheres rayi Ahyong &amp; Ng, 2007. A, D, ovigerous female (7.6 × 5.8 mm) (ZRC 2013.1416), Peninsular Malaysia; B, C, E, female (6.2 × 4.4 mm) (ZRC 2013.1416), Peninsular Malaysia. A, B, overall dorsal view; C, dorsal view of carapace; D, E, right P5." figureDoi="http://doi.org/10.5281/zenodo.7174335" httpUri="https://zenodo.org/record/7174335/files/figure.png" pageId="65" pageNumber="199">48A</figureCitation>
, 
<figureCitation id="137A2A4BCE30FF9036B31AACD19079D0" box="[467,497,1955,1979]" captionStart="Fig" captionStartId="64.[113,148,1597,1619]" captionTargetBox="[117,1455,192,1580]" captionTargetId="figure-14@64.[107,1458,184,1592]" captionTargetPageId="64" captionText="Fig. 49. Arcotheres rayi Ahyong &amp; Ng, 2007, ovigerous female (7.6 × 5.8 mm) (ZRC 2013.1416), Peninsular Malaysia. A, overall dorsal view; B, right MXP3; C, frontal view of cephalothorax; D, anterior thoracic sternum; E, right chela; F–I, left P2–P5; J–M, right P2–P5. Scale = A, 2.0 mm; B, 0.5 mm; C–M, 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.7174339" httpUri="https://zenodo.org/record/7174339/files/figure.png" pageId="65" pageNumber="199">49</figureCitation>
). The length of the longer P5 dactylus varies allometrically, being proportionately shorter in small specimens (
<tableCitation id="C6C30375CE30FF9036DA1AECD2707990" box="[442,529,2019,2043]" captionStart="Table 2" captionStartId="66.[814,869,1151,1173]" captionText="Table 2. Length/height proportions of P5 dactylus of Arcotheres exiguus and A. rayi." pageId="65" pageNumber="199">Table 2</tableCitation>
; 
<figureCitation id="137A2A4BCE30FF90357E1AECD2C07991" box="[542,673,2018,2043]" captionStart="Fig" captionStartId="65.[135,170,1541,1563]" captionTargetBox="[137,1477,189,1531]" captionTargetId="figure-7@65.[132,1485,182,1535]" captionTargetPageId="65" captionText="Fig. 50. Right female P5. A–G, Arcotheres exiguus (Bürger, 1895), Peninsular Malaysia: A, juvenile (3.8 × 3.6 mm) (ZRC 2018.772); B, ovigerous (4.5 × 4.1 mm) (ZRC 2018.775); C, non-ovigerous (4.6 × 4.2 mm) (ZRC 2018.772); D, spent (5.4 × 4.6 mm) (ZRC 2018.775); E, ovigerous (6.8 × 6.1 mm) (ZRC 2019.516); F, ovigerous (7.7 × 6.7 mm) (ZRC 2018.772); G, ovigerous (8.9 × 7.6 mm) (ZRC 2018.775). H–L, A. rayi Ahyong &amp; Ng, 2007, Peninsular Malaysia: H, juvenile (5.4 × 5.8 mm) (ZRC 2013.1416); I, non-ovigerous (5.6 × 4.3 mm) (ZRC 2013.1416); J, non-ovigerous (8.1 × 5.2 mm) (ZRC 2013.1416); K, spent (6.9 × 5.2 mm) (ZRC 2013.1416); L, ovigerous (8.4 × 7.0 mm) (ZRC 2013.1416). Scale = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.7174341" httpUri="https://zenodo.org/record/7174341/files/figure.png" pageId="65" pageNumber="199">Fig. 50H–L</figureCitation>
).
</paragraph>
</subSubSection>
<caption id="DF3E6646CE30FF9037E71B0AD22178C0" ID-DOI="http://doi.org/10.5281/zenodo.7174341" ID-Zenodo-Dep="7174341" httpUri="https://zenodo.org/record/7174341/files/figure.png" pageId="65" pageNumber="199" startId="65.[135,170,1541,1563]" targetBox="[137,1477,189,1531]" targetPageId="65">
<paragraph id="8BFE36CECE30FF9037E71B0AD22178C0" blockId="65.[135,1476,1541,1707]" pageId="65" pageNumber="199">
Fig. 50. Right female P5. A–G, 
<taxonomicName id="4C414D4DCE30FF9036A11B0AD3727870" authority="(Burger, 1895)" baseAuthorityName="Burger" baseAuthorityYear="1895" box="[449,787,1541,1563]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" kingdom="Animalia" order="Decapoda" pageId="65" pageNumber="199" phylum="Arthropoda" rank="species" species="exiguus">
<emphasis id="B935EADCCE30FF9036A11B0AD21A7870" box="[449,635,1541,1563]" italics="true" pageId="65" pageNumber="199">Arcotheres exiguus</emphasis>
(
<bibRefCitation id="EFD04B3FCE30FF9035E81B09D36D7870" author="Burger O" box="[648,780,1541,1563]" pageId="65" pageNumber="199" pagination="361 - 390" refId="ref62795" refString="Burger O (1895) Ein Beitrag zur Kenntnis der Pinnotherinen. Zoologische Jahrbucher, Abtheilung fur Systematik, Geographie und Biologie der Thiere, 8: 361 - 390, pls. 9, 10." type="book chapter" year="1895">Bürger, 1895</bibRefCitation>
)
</taxonomicName>
, Peninsular Malaysia: A, juvenile (3.8 × 3.6 mm) (ZRC 2018.772); B, ovigerous (4.5 × 4.1 mm) (ZRC 2018.775); C, non-ovigerous (4.6 × 4.2 mm) (ZRC 2018.772); D, spent (5.4 × 4.6 mm) (ZRC 2018.775); E, ovigerous (6.8 × 6.1 mm) (ZRC 2019.516); F, ovigerous (7.7 × 6.7 mm) (ZRC 2018.772); G, ovigerous (8.9 × 7.6 mm) (ZRC 2018.775). H–L, 
<taxonomicName id="4C414D4DCE30FF9037A11B53D1B9781A" authority="Ahyong &amp; Ng, 2007" authorityName="Ahyong &amp; Ng" authorityYear="2007" box="[193,472,1628,1650]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" kingdom="Animalia" order="Decapoda" pageId="65" pageNumber="199" phylum="Arthropoda" rank="species" species="rayi">
<emphasis id="B935EADCCE30FF9037A11B53D165781A" box="[193,260,1628,1649]" italics="true" pageId="65" pageNumber="199">A. rayi</emphasis>
Ahyong &amp; Ng, 2007
</taxonomicName>
, Peninsular Malaysia: H, juvenile (5.4 × 5.8 mm) (ZRC 2013.1416); I, non-ovigerous (5.6 × 4.3 mm) (ZRC 2013.1416); J, non-ovigerous (8.1 × 5.2 mm) (ZRC 2013.1416); K, spent (6.9 × 5.2 mm) (ZRC 2013.1416); L, ovigerous (8.4 × 7.0 mm) (ZRC 2013.1416). Scale = 1.0 mm.
</paragraph>
</caption>
<subSubSection id="C35B6545CE30FF9034231BECD3E87971" pageId="65" pageNumber="199" type="biology_ecology">
<paragraph id="8BFE36CECE30FF9034231BECD3E87971" blockId="65.[835,1474,1763,1818]" pageId="65" pageNumber="199">
<emphasis id="B935EADCCE30FF9034231BECD3E17890" bold="true" box="[835,896,1763,1787]" pageId="65" pageNumber="199">Host.</emphasis>
Known only from 
<taxonomicName id="4C414D4DCE30FF90332E1BECD4A17891" box="[1102,1216,1763,1787]" class="Bivalvia" family="Veneridae" genus="Paphia" kingdom="Animalia" order="Venerida" pageId="65" pageNumber="199" phylum="Mollusca" rank="species" species="undetermined">
<emphasis id="B935EADCCE30FF90332E1BECD4FA7890" box="[1102,1179,1763,1787]" italics="true" pageId="65" pageNumber="199">Paphia</emphasis>
sp.
</taxonomicName>
(
<taxonomicName id="4C414D4DCE30FF9033AE1BECD31D7971" authority="Rafinesque, 1815" authorityName="Rafinesque" authorityYear="1815" class="Bivalvia" family="Veneridae" kingdom="Animalia" order="Venerida" pageId="65" pageNumber="199" phylum="Mollusca" rank="family">Veneridae Rafinesque, 1815</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="C35B6545CE30FF9034231A4CD5A37651" pageId="65" pageNumber="199" type="discussion">
<paragraph id="8BFE36CECE30FF9034231A4CD5A37651" blockId="65.[835,1476,1859,2107]" pageId="65" pageNumber="199">
<emphasis id="B935EADCCE30FF9034231A4CD3D37930" bold="true" box="[835,946,1859,1883]" pageId="65" pageNumber="199">Remarks.</emphasis>
<taxonomicName id="4C414D4DCE30FF9034D91A4CD4017931" authorityName="Ahyong &amp; Ng" authorityYear="2007" box="[953,1120,1859,1883]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" kingdom="Animalia" order="Decapoda" pageId="65" pageNumber="199" phylum="Arthropoda" rank="species" species="rayi">
<emphasis id="B935EADCCE30FF9034D91A4CD4017931" box="[953,1120,1859,1883]" italics="true" pageId="65" pageNumber="199">Arcotheres rayi</emphasis>
</taxonomicName>
is morphologically very close to 
<taxonomicName id="4C414D4DCE30FF9034241A6CD3D57911" baseAuthorityName="Burger" baseAuthorityYear="1895" box="[836,948,1891,1914]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" kingdom="Animalia" order="Decapoda" pageId="65" pageNumber="199" phylum="Arthropoda" rank="species" species="exiguus">
<emphasis id="B935EADCCE30FF9034241A6CD3D57911" box="[836,948,1891,1914]" italics="true" pageId="65" pageNumber="199">A. exiguus</emphasis>
</taxonomicName>
, but the good series of specimens has permitted evaluation of intraspecific variability, enabling more reliable recognition of interspecific differences. Females of 
<taxonomicName id="4C414D4DCE30FF9032111AACD5DA79D1" authorityName="Ahyong &amp; Ng" authorityYear="2007" box="[1393,1467,1955,1978]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" kingdom="Animalia" order="Decapoda" pageId="65" pageNumber="199" phylum="Arthropoda" rank="species" species="rayi">
<emphasis id="B935EADCCE30FF9032111AACD5DA79D1" box="[1393,1467,1955,1978]" italics="true" pageId="65" pageNumber="199">A. rayi</emphasis>
</taxonomicName>
, in which the carapace front is demarcated by a notch in the anterior margin above each orbit (
<figureCitation id="137A2A4BCE30FF9033BF1AECD55B7991" box="[1247,1338,2018,2043]" captionStart="Fig" captionStartId="61.[136,171,1192,1214]" captionTargetBox="[136,1472,189,1177]" captionTargetId="figure-7@61.[136,1472,189,1177]" captionTargetPageId="61" captionText="Fig. 46. Arcotheres rayi Ahyong &amp; Ng, 2007, holotype female (11.3 × 8.7 mm) (SMF-ZMG 952), Philippines. A, overall dorsal view; B, ventral view of cephalothorax; C, outer view of left chela." figureDoi="http://doi.org/10.5281/zenodo.7174331" httpUri="https://zenodo.org/record/7174331/files/figure.png" pageId="65" pageNumber="199">Figs. 46</figureCitation>
, 
<figureCitation id="137A2A4BCE30FF9032261AECD5D47990" box="[1350,1461,2019,2043]" captionStart="Fig" captionStartId="63.[135,170,1761,1783]" captionTargetBox="[135,1473,190,1746]" captionTargetId="figure-7@63.[135,1473,190,1746]" captionTargetPageId="63" captionText="Fig. 48. Arcotheres rayi Ahyong &amp; Ng, 2007. A, D, ovigerous female (7.6 × 5.8 mm) (ZRC 2013.1416), Peninsular Malaysia; B, C, E, female (6.2 × 4.4 mm) (ZRC 2013.1416), Peninsular Malaysia. A, B, overall dorsal view; C, dorsal view of carapace; D, E, right P5." figureDoi="http://doi.org/10.5281/zenodo.7174335" httpUri="https://zenodo.org/record/7174335/files/figure.png" pageId="65" pageNumber="199">48B, C, E</figureCitation>
), are readily distinguished from 
<taxonomicName id="4C414D4DCE30FF9033F7150CD5687671" baseAuthorityName="Burger" baseAuthorityYear="1895" box="[1175,1289,2051,2074]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" kingdom="Animalia" order="Decapoda" pageId="65" pageNumber="199" phylum="Arthropoda" rank="species" species="exiguus">
<emphasis id="B935EADCCE30FF9033F7150CD5687671" box="[1175,1289,2051,2074]" italics="true" pageId="65" pageNumber="199">A. exiguus</emphasis>
</taxonomicName>
, which does not have the anterior carapace notches (
<figureCitation id="137A2A4BCE30FF9033B0152CD5017650" box="[1232,1376,2083,2107]" captionStart="Fig" captionStartId="46.[113,148,1474,1496]" captionTargetBox="[113,1451,191,1457]" captionTargetId="figure-15@46.[103,1466,176,1470]" captionTargetPageId="46" captionText="Fig. 33. Arcotheres similis (Bürger, 1895). A, B, ovigerous female (6.7 × 5.5 mm) (NHM 1936.6.19.2), Singapore (holotype of Pinnotheres spinidactylus Gordon, 1936); C, holotype female (8.3 × 6.4 mm) (SMF-ZMG 956 Go 950a), Philippines (holotype of Pinnotheres similis Bürger, 1895); D, female (7.3 × 6.0 mm; soft) (ZRC 1965.11.24.47), Sarawak; E, F, male (3.3 × 3.3 mm) (NHM 1936.6.19.3), Singapore; G–I, male (4.8 × 5.0 mm) (NHM 1936.6.19.14), Singapore; J, K, male (5.7 × 5.6 mm) (ZRC 2019.1026), Lombok, Indonesia; L, male (3.3 × 3.3 mm) (NHM 1936.6.19.3), Singapore (paratype of P. spinidactylus Gordon, 1936). A, left MXP3 (inner view); B, palp of left MXP3 (outer view); C, D, right MXP3 (outer view); E, G, J, left G1 (ventral view); F, distal part of left G1 (ventral view); H, K, distal part of left G1 (dorsal view); I, left G2; L, male pleonal somites 5, 6 and telson. A, B, E, F, after Gordon (1936: figs. 1c, 2a); C, after Ahyong &amp; Ng (2007: fig. 14C). Scales = D, G–K, 0.5 mm; L, 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.7174303" httpUri="https://zenodo.org/record/7174303/files/figure.png" pageId="65" pageNumber="199">Figs. 33B, C</figureCitation>
, 
<figureCitation id="137A2A4BCE30FF90320E152DD5DF7651" box="[1390,1470,2082,2106]" captionStart="Fig" captionStartId="50.[113,148,1511,1533]" captionTargetBox="[113,1451,189,1496]" captionTargetId="figure-14@50.[113,1451,189,1496]" captionTargetPageId="50" captionText="Fig. 35. Arcotheres exiguus (Bürger, 1895), colour in life. A, B, ovigerous female (7.9 × 7.3 mm) (ZRC 2019.516), in situ in Marcia recens (Holten, 1802) (Veneridae), Peninsular Malaysia; C, male (4.3 × 4.2 mm) (ZRC 2019.516), Peninsular Malaysia; D, male (4.5 × 4.2 mm) (ZRC 2018.768), India; E, F, ovigerous female, from Meretrix aurora Hornell, 1917 (Veneridae), India (specimens not examined). Photographs: A–C, Paul Y. C. Ng; D–F, R. Ravinesh." figureDoi="http://doi.org/10.5281/zenodo.7174307" httpUri="https://zenodo.org/record/7174307/files/figure.png" pageId="65" pageNumber="199">35B, E</figureCitation>
,
</paragraph>
</subSubSection>
</treatment>
</document>