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<document id="44A91C149FD39E5C34E0D0B4E34DE267" ID-DOI="10.11646/zootaxa.5414.1.1" ID-ISSN="1175-5334" ID-Zenodo-Dep="10681263" ID-ZooBank="3B1DFADA-C20E-473B-A5E9-843548B54AD3" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1708377054888" checkinUser="plazi" docAuthor="Paredes-Munguía, Williams, Brescovit, Antonio D. &amp; Teixeira, Renato A." docDate="2024" docId="03E887D2BA13FE0EFDDBFD966687FDEF" docLanguage="en" docName="zootaxa.5414.1.1.pdf" docOrigin="Zootaxa 5414 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5414.1.1" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.18:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="18" docTitle="Arctosa pacaya Paredes-Munguía &amp; Brescovit &amp; Teixeira 2024, sp. nov." docType="treatment" docVersion="1" lastPageNumber="35" masterDocId="FFD1FFAABA0DFE2CFD4CFF8E6559FFD2" masterDocTitle="Revision of Neotropical wolf spider genus Arctosa C. L. Koch, 1847 (Araneae: Lycosidae), with description of seven new species" masterLastPageNumber="83" masterPageNumber="1" pageNumber="31" updateTime="1708377875290" updateUser="ExternalLinkService">
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<mods:title id="8202333EC5E684D2A76754B3E7A29DC2">Revision of Neotropical wolf spider genus Arctosa C. L. Koch, 1847 (Araneae: Lycosidae), with description of seven new species</mods:title>
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<mods:namePart id="0071BE6B8CB6110A94AE6CF6F647F87E">Paredes-Munguía, Williams</mods:namePart>
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<mods:affiliation id="7DD85767581AED48FA42D5AEA5F2116E">Pontifícia Universidade Católica do Rio Grande do Sul, Escola de Ciências da Saúde e da Vida, Museu de Ciências e Tecnologia, Laboratório de Aracnologia, Porto Alegre, Brazil. &amp; Museo de Historia Natural de la Universidad Nacional Mayor de San Marcos, Laboratorio de Entomología, Av. Arenales 1256, Lima 14, Peru. &amp; williams. paredes @ acad. pucrs. br; https: // orcid. org / 0000 - 0002 - 3320 - 4611</mods:affiliation>
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<mods:namePart id="63EF2BF50720D4614BA44230BDCDBD38">Brescovit, Antonio D.</mods:namePart>
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<mods:affiliation id="637DAD952A3CFF74B5293610582EC63D">Laboratório de Coleções Zoológicas, Av. Vital Brasil, 1500, Butantã, 05503 - 900, São Paulo, SP, Brazil. &amp; antonio. brescovit @ butantan. gov. br; https: // orcid. org / 0000 - 0002 - 1511 - 5324</mods:affiliation>
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<mods:namePart id="52F346411D206CF00DA0B737D9D55F94">Teixeira, Renato A.</mods:namePart>
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<mods:affiliation id="DB180BE1BB60C671D8232D106D76D9D1">Pontifícia Universidade Católica do Rio Grande do Sul, Escola de Ciências da Saúde e da Vida, Museu de Ciências e Tecnologia, Laboratório de Aracnologia, Porto Alegre, Brazil. &amp; Pontifícia Universidade Católica do Rio Grande do Sul, Escola de Ciências da Saúde e da Vida, Museu de Ciências e Tecnologia, Laboratório de Aracnologia, Porto Alegre, Brazil. &amp; renatoaug. tx @ gmail. com; https: // orcid. org / 0000 - 0002 - 1756 - 9821</mods:affiliation>
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<mods:date id="BCDC284AD4686C44979867F0EBF03F9D">2024</mods:date>
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<taxonomicName id="4C414D47BA13FE32FDDBFD966409FDE0" authority="Paredes-Munguía &amp; Brescovit &amp; Teixeira, 2024" authorityName="Paredes-Munguía &amp; Brescovit &amp; Teixeira" authorityYear="2024" box="[151,336,536,562]" class="Arachnida" family="Lycosidae" genus="Arctosa" kingdom="Animalia" order="Araneae" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="pacaya" status="sp. nov.">
<emphasis id="B935EAD6BA13FE32FDDBFD966409FDE0" bold="true" box="[151,336,536,562]" italics="true" pageId="30" pageNumber="31">Arctosa pacaya</emphasis>
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<taxonomicNameLabel id="A20657ADBA13FE32FC1BFD9664ECFDE0" box="[343,437,536,562]" pageId="30" pageNumber="31" rank="species">sp. nov.</taxonomicNameLabel>
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<subSubSection id="C35B654FBA13FE32FDDBFDB26467FD84" box="[151,318,572,598]" pageId="30" pageNumber="31" type="description">
<paragraph id="8BFE36C4BA13FE32FDDBFDB26467FD84" blockId="30.[151,437,536,598]" box="[151,318,572,598]" pageId="30" pageNumber="31">
<figureCitation id="137A2A41BA13FE32FDDBFDB2644EFD84" box="[151,279,572,598]" captionStart-0="FIGURE 24" captionStart-1="FIGURE 25" captionStart-2="FIGURE 26" captionStart-3="FIGURE 27" captionStartId-0="30.[151,250,1977,2001]" captionStartId-1="31.[151,250,1908,1932]" captionStartId-2="32.[151,250,1646,1670]" captionStartId-3="33.[151,250,1132,1156]" captionTargetBox-0="[174,1413,841,1944]" captionTargetBox-1="[188,1400,181,1884]" captionTargetBox-2="[151,1436,181,1621]" captionTargetBox-3="[151,1436,181,1108]" captionTargetId-0="figure-259@30.[164,1423,833,1954]" captionTargetId-1="figure-21@31.[188,1400,181,1884]" captionTargetId-2="figure-25@32.[151,1436,181,1621]" captionTargetId-3="figure-21@33.[151,1436,181,1108]" captionTargetPageId-0="30" captionTargetPageId-1="31" captionTargetPageId-2="32" captionTargetPageId-3="33" captionText-0="FIGURE 24. Arctosa pacaya sp. nov. Holotype male pedipalp (MUSM-ENT 505356) ac (a, ventral; b, retrolateral; c, anterior). Scale bars: ac, 0.25 mm. CMS cymbium macrosetae, Pa posterior arm, Ra retrolateral arm, TA terminal apophysis." captionText-1="FIGURE 25. Arctosa pacaya sp. nov. Male pedipalp (MUSM-ENT 505356) ac (a, retrolateral; b, ventral; c, prolateral). Pedipalp bulb df, h (d, retrolateral; e, ventral; f, prolateral; h, anterior). g, apical cymbium setae, i, protuberance on male chelicerae. Scale bars: ac, 0.2 mm, dh, 100 µm, i, 125 µm. C conductor, DMA depresion of median apophysis, E embolus, P palea, Pa posterior arm, SD seminal duct, ST subtegulum, T tegulum, TA terminal apophysis." captionText-2="FIGURE 26. Arctosa pacaya sp. nov. Male habitus (MUSM-ENT 505356) ac (a, dorsal; b, ventral; c, lateral). Pedipalp, expanded d, e (d, prolateral; e, anterior). Scale bars: ac, 1 mm, d, e, 200 µm. BH basal hematodocha, CMS cymbial macrosetae, DH distal hematodocha, E embolus, Pa posterior arm, SD seminal duct, ST subtegulum, T tegulum, TA terminal apophysis." captionText-3="FIGURE 27. Arctosa pacaya sp. nov. Male pedipalp SEM (MUSM-ENT 505356) ac (a, ventral; b, TA detail; c, Pa detail). Scale bars: a, 150 µm, b, 100 µm, c, 50 µm. C conductor, E embolus, MA median apophysis, P palea, Pa posterior arm, Ra retrolateral arm, ST subtegulum, T tegulum, TA terminal apophysis." figureDoi-0="http://doi.org/10.5281/zenodo.10681313" figureDoi-1="http://doi.org/10.5281/zenodo.10681315" figureDoi-2="http://doi.org/10.5281/zenodo.10681317" figureDoi-3="http://doi.org/10.5281/zenodo.10681319" httpUri-0="https://zenodo.org/record/10681313/files/figure.png" httpUri-1="https://zenodo.org/record/10681315/files/figure.png" httpUri-2="https://zenodo.org/record/10681317/files/figure.png" httpUri-3="https://zenodo.org/record/10681319/files/figure.png" pageId="30" pageNumber="31">Figs 2427</figureCitation>
,
<figureCitation id="137A2A41BA13FE32FC6EFDB26467FD84" box="[290,318,572,598]" captionStart="FIGURE 62" captionStartId="72.[151,250,1618,1642]" captionTargetBox="[171,1418,564,1590]" captionTargetId="figure-197@72.[164,1423,561,1595]" captionTargetPageId="72" captionText="FIGURE 62. Geographic distribution records of Arctosa ayaymama sp. nov. (blue dot), Arctosa pacaya sp. nov. (red dot), Hogna andina comb. nov. (orange dot)." figureDoi="http://doi.org/10.5281/zenodo.10681391" httpUri="https://zenodo.org/record/10681391/files/figure.png" pageId="30" pageNumber="31">62</figureCitation>
</paragraph>
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<emphasis id="B935EAD6BA13FE32FDDBFDF1641FFD4A" bold="true" box="[151,326,638,665]" pageId="30" pageNumber="31">Type Material.</emphasis>
<materialsCitation id="3B293C99BA13FE32FC07FDF167BFFD6F" collectingDate="1989-08-11" collectorName="D. Silva &amp; Silva D." country="Peru" elevation="100" latitude="-5.1" location="Loreto" longLatPrecision="1305" longitude="-74.833336" pageId="30" pageNumber="31" specimenCode="MUSM-ENT 505356" specimenCount="1" stateProvince="Loreto" typeStatus="holotype">
Male
<typeStatus id="54FA8866BA13FE32FCC5FDF164B2FD4B" box="[393,491,639,665]" pageId="30" pageNumber="31" type="holotype">holotype</typeStatus>
from Pithecia,
<geoCoordinate id="EE755003BA13FE32FFDDFDF06783FD4B" box="[657,730,638,665]" degrees="5" direction="south" minutes="06" orientation="latitude" pageId="30" pageNumber="31" precision="925" value="-5.1">5°06'S</geoCoordinate>
<geoCoordinate id="EE755003BA13FE32FF93FDF0661BFD4B" box="[735,834,638,665]" degrees="74" direction="west" minutes="50" orientation="longitude" pageId="30" pageNumber="31" precision="925" value="-74.833336">74°50'W</geoCoordinate>
,
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<elevation id="006CD1F7BA13FE32FE07FDF166CBFD4B" box="[843,914,639,665]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="30" pageNumber="31" unit="m" value="100.0">100 m</elevation>
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,
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<collectingDate id="EFBBE9ECBA13FE32FED0FDF1617DFD4B" box="[924,1060,639,665]" pageId="30" pageNumber="31" value="1989-08-11">11.viii.1989</collectingDate>
</date>
, Parque Nacional Pacaya Samiria,
<collectingRegion id="4985F826BA13FE32FDDBFD2D65BCFD6F" box="[151,229,675,701]" country="Peru" name="Loreto" pageId="30" pageNumber="31">Loreto</collectingRegion>
,
<collectingCountry id="F3567654BA13FE32FDBCFD2D647FFD6F" box="[240,294,675,701]" name="Peru" pageId="30" pageNumber="31">Peru</collectingCountry>
,
<collectorName id="26B45312BA13FE32FC7DFD2D64D4FD6F" box="[305,397,675,701]" pageId="30" pageNumber="31">D. Silva</collectorName>
D.
<emphasis id="B935EAD6BA13FE32FCFAFD2D6481FD6F" box="[438,472,675,701]" italics="true" pageId="30" pageNumber="31">leg</emphasis>
(
<specimenCode id="DBE79EBFBA13FE32FCABFD2D6784FD6F" box="[487,733,674,701]" collectionCode="MUSM-ENT" pageId="30" pageNumber="31">MUSM-ENT 505356</specimenCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C35B654FBA13FE32FD8BFD496113FCFB" pageId="30" pageNumber="31" type="etymology">
<paragraph id="8BFE36C4BA13FE32FD8BFD496113FCFB" blockId="30.[151,1437,638,809]" pageId="30" pageNumber="31">
<emphasis id="B935EAD6BA13FE32FD8BFD496417FD33" bold="true" box="[199,334,711,737]" pageId="30" pageNumber="31">Etymology.</emphasis>
The species epithet is dedicated to the Pacaya Samiria Natural Reserve (PSNR), a protected natural wetland area. PSNR is the biggest protected flooded forest in the Peruvian Amazon, the biggest RAMSAR site in
<collectingCountry id="F3567654BA13FE32FDDBFC816594FCFB" box="[151,205,783,809]" name="Peru" pageId="30" pageNumber="31">Peru</collectingCountry>
, and serves as habitat to 25% of the bird fauna reported from
<collectingCountry id="F3567654BA13FE32FECCFC8166EAFCFB" box="[896,947,783,809]" name="Peru" pageId="30" pageNumber="31">Peru</collectingCountry>
(
<bibRefCitation id="EFD04B35BA13FE32FE8EFC816167FCFB" author="Isola, S." box="[962,1086,783,809]" pageId="30" pageNumber="31" refId="ref49846" refString="Isola, S. (2007) Birds of Pacaya Samiria National Reserve. The Nature Conservancy (TNC) and United States Agency for International Development (USAID), Lima, 64 pp." type="book" year="2007">
<collectingCountry id="F3567654BA13FE32FE8EFC8166A3FCFB" box="[962,1018,783,809]" name="Iceland" pageId="30" pageNumber="31">Isola</collectingCountry>
2007
</bibRefCitation>
).
</paragraph>
</subSubSection>
<caption id="DF3E664CBA13FE32FDDBF8376062F824" ID-DOI="http://doi.org/10.5281/zenodo.10681313" ID-Zenodo-Dep="10681313" httpUri="https://zenodo.org/record/10681313/files/figure.png" pageId="30" pageNumber="31" startId="30.[151,250,1977,2001]" targetBox="[174,1413,841,1944]" targetPageId="30" targetType="figure">
<paragraph id="8BFE36C4BA13FE32FDDBF8376062F824" blockId="30.[151,1437,1977,2038]" pageId="30" pageNumber="31">
<emphasis id="B935EAD6BA13FE32FDDBF8376447F803" bold="true" box="[151,286,1977,2001]" pageId="30" pageNumber="31">FIGURE 24.</emphasis>
<taxonomicName id="4C414D47BA13FE32FC6FF83464E6F803" authority="Paredes-Munguía &amp; Brescovit &amp; Teixeira, 2024" authorityName="Paredes-Munguía &amp; Brescovit &amp; Teixeira" authorityYear="2024" box="[291,447,1978,2001]" class="Arachnida" family="Lycosidae" genus="Arctosa" kingdom="Animalia" order="Araneae" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="pacaya" status="sp. nov.">
<emphasis id="B935EAD6BA13FE32FC6FF83464E6F803" box="[291,447,1978,2001]" italics="true" pageId="30" pageNumber="31">Arctosa pacaya</emphasis>
</taxonomicName>
<emphasis id="B935EAD6BA13FE32FC88F834674AF800" bold="true" box="[452,531,1978,2002]" pageId="30" pageNumber="31">
<taxonomicNameLabel id="A20657ADBA13FE32FC88F834674AF800" box="[452,531,1978,2002]" pageId="30" pageNumber="31" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Holotype male pedipalp (MUSM-ENT 505356)
<emphasis id="B935EAD6BA13FE32FEB7F8346179F800" bold="true" box="[1019,1056,1978,2002]" pageId="30" pageNumber="31">ac</emphasis>
(a, ventral; b, retrolateral; c, anterior). Scale bars: ac, 0.25 mm.
<emphasis id="B935EAD6BA13FE32FCEFF8536482F827" bold="true" box="[419,475,2013,2037]" pageId="30" pageNumber="31">CMS</emphasis>
cymbium macrosetae,
<emphasis id="B935EAD6BA13FE32FF8AF85067BBF824" bold="true" box="[710,738,2014,2038]" pageId="30" pageNumber="31">Pa</emphasis>
posterior arm,
<emphasis id="B935EAD6BA13FE32FE31F85066C5F824" bold="true" box="[893,924,2014,2038]" pageId="30" pageNumber="31">Ra</emphasis>
retrolateral arm,
<emphasis id="B935EAD6BA13FE32F907F8536135F827" bold="true" box="[1099,1132,2013,2037]" pageId="30" pageNumber="31">TA</emphasis>
terminal apophysis.
</paragraph>
</caption>
<caption id="DF3E664CBA12FE33FDDBF8FA610AF82A" ID-DOI="http://doi.org/10.5281/zenodo.10681315" ID-Zenodo-Dep="10681315" httpUri="https://zenodo.org/record/10681315/files/figure.png" pageId="31" pageNumber="32" startId="31.[151,250,1908,1932]" targetBox="[188,1400,181,1884]" targetPageId="31" targetType="figure">
<paragraph id="8BFE36C4BA12FE33FDDBF8FA610AF82A" blockId="31.[151,1437,1908,2041]" pageId="31" pageNumber="32">
<emphasis id="B935EAD6BA12FE33FDDBF8FA647AF85E" bold="true" box="[151,291,1908,1932]" pageId="31" pageNumber="32">FIGURE 25.</emphasis>
<taxonomicName id="4C414D47BA12FE33FC61F8FB6494F85E" authority="Paredes-Munguía &amp; Brescovit &amp; Teixeira, 2024" authorityName="Paredes-Munguía &amp; Brescovit &amp; Teixeira" authorityYear="2024" box="[301,461,1909,1932]" class="Arachnida" family="Lycosidae" genus="Arctosa" kingdom="Animalia" order="Araneae" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="pacaya" status="sp. nov.">
<emphasis id="B935EAD6BA12FE33FC61F8FB6494F85E" box="[301,461,1909,1932]" italics="true" pageId="31" pageNumber="32">Arctosa pacaya</emphasis>
</taxonomicName>
<emphasis id="B935EAD6BA12FE33FC9BF8FB6772F85F" bold="true" box="[471,555,1909,1933]" pageId="31" pageNumber="32">
<taxonomicNameLabel id="A20657ADBA12FE33FC9BF8FB6772F85F" box="[471,555,1909,1933]" pageId="31" pageNumber="32" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Male pedipalp (MUSM-ENT 505356)
<emphasis id="B935EAD6BA12FE33FE87F8FB66A9F85F" bold="true" box="[971,1008,1909,1933]" pageId="31" pageNumber="32">ac</emphasis>
(a, retrolateral; b, ventral; c, prolateral). Pedipalp bulb
<emphasis id="B935EAD6BA12FE33FC63F8166436F863" bold="true" box="[303,367,1944,1969]" pageId="31" pageNumber="32">df, h</emphasis>
(d, retrolateral; e, ventral; f, prolateral; h, anterior). g, apical cymbium setae, i, protuberance on male chelicerae. Scale bars: ac, 0.2 mm, dh, 100 µm, i, 125 µm.
<emphasis id="B935EAD6BA12FE33FE4EF832664DF806" bold="true" box="[770,788,1980,2004]" pageId="31" pageNumber="32">C</emphasis>
conductor,
<emphasis id="B935EAD6BA12FE33FEC7F832669EF806" bold="true" box="[907,967,1980,2004]" pageId="31" pageNumber="32">DMA</emphasis>
depresion of median apophysis,
<emphasis id="B935EAD6BA12FE33F858F833607CF807" bold="true" box="[1300,1317,1981,2005]" pageId="31" pageNumber="32">E</emphasis>
embolus,
<emphasis id="B935EAD6BA12FE33F8C1F83360C5F807" bold="true" box="[1421,1436,1981,2005]" pageId="31" pageNumber="32">P</emphasis>
palea,
<emphasis id="B935EAD6BA12FE33FD95F86F65ACF82B" bold="true" box="[217,245,2017,2041]" pageId="31" pageNumber="32">Pa</emphasis>
posterior arm,
<emphasis id="B935EAD6BA12FE33FCDCF86E64E9F82A" bold="true" box="[400,432,2016,2040]" pageId="31" pageNumber="32">SD</emphasis>
seminal duct,
<emphasis id="B935EAD6BA12FE33FF08F86E673AF82A" bold="true" box="[580,611,2016,2040]" pageId="31" pageNumber="32">ST</emphasis>
subtegulum,
<emphasis id="B935EAD6BA12FE33FFA1F86F67A7F82B" bold="true" box="[749,766,2017,2041]" pageId="31" pageNumber="32">T</emphasis>
tegulum,
<emphasis id="B935EAD6BA12FE33FE2FF86E66DDF82A" bold="true" box="[867,900,2016,2040]" pageId="31" pageNumber="32">TA</emphasis>
terminal apophysis.
</paragraph>
</caption>
<caption id="DF3E664CBA2DFE0CFDDBF9E060DEF91C" ID-DOI="http://doi.org/10.5281/zenodo.10681317" ID-Zenodo-Dep="10681317" httpUri="https://zenodo.org/record/10681317/files/figure.png" pageId="32" pageNumber="33" startId="32.[151,250,1646,1670]" targetBox="[151,1436,181,1621]" targetPageId="32" targetType="figure">
<paragraph id="8BFE36C4BA2DFE0CFDDBF9E060DEF91C" blockId="32.[151,1437,1646,1742]" pageId="32" pageNumber="33">
<emphasis id="B935EAD6BA2DFE0CFDDBF9E0647AF954" bold="true" box="[151,291,1646,1670]" pageId="32" pageNumber="33">FIGURE 26.</emphasis>
<taxonomicName id="4C414D47BA2DFE0CFC60F9E06495F954" authority="Paredes-Munguía &amp; Brescovit &amp; Teixeira, 2024" authorityName="Paredes-Munguía &amp; Brescovit &amp; Teixeira" authorityYear="2024" box="[300,460,1646,1670]" class="Arachnida" family="Lycosidae" genus="Arctosa" kingdom="Animalia" order="Araneae" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="pacaya" status="sp. nov.">
<emphasis id="B935EAD6BA2DFE0CFC60F9E06495F954" box="[300,460,1646,1670]" italics="true" pageId="32" pageNumber="33">Arctosa pacaya</emphasis>
</taxonomicName>
<emphasis id="B935EAD6BA2DFE0CFC9AF9E06770F954" bold="true" box="[470,553,1646,1670]" pageId="32" pageNumber="33">
<taxonomicNameLabel id="A20657ADBA2DFE0CFC9AF9E06770F954" box="[470,553,1646,1670]" pageId="32" pageNumber="33" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Male habitus (MUSM-ENT 505356)
<emphasis id="B935EAD6BA2DFE0CFEF5F9E06684F954" bold="true" box="[953,989,1646,1670]" pageId="32" pageNumber="33">ac</emphasis>
(a, dorsal; b, ventral; c, lateral). Pedipalp, expanded
<emphasis id="B935EAD6BA2DFE0CFDB1F91C647BF978" bold="true" box="[253,290,1682,1706]" pageId="32" pageNumber="33">d, e</emphasis>
(d, prolateral; e, anterior). Scale bars: ac, 1 mm, d, e, 200 µm.
<emphasis id="B935EAD6BA2DFE0CFED7F91C6699F978" bold="true" box="[923,960,1682,1706]" pageId="32" pageNumber="33">BH</emphasis>
basal hematodocha,
<emphasis id="B935EAD6BA2DFE0CF9DCF91C6191F978" bold="true" box="[1168,1224,1682,1706]" pageId="32" pageNumber="33">CMS</emphasis>
cymbial macrosetae,
<emphasis id="B935EAD6BA2DFE0CFDDBF93865E4F91C" bold="true" box="[151,189,1718,1742]" pageId="32" pageNumber="33">DH</emphasis>
distal hematodocha,
<emphasis id="B935EAD6BA2DFE0CFCD8F93864FCF91C" bold="true" box="[404,421,1718,1742]" pageId="32" pageNumber="33">E</emphasis>
embolus,
<emphasis id="B935EAD6BA2DFE0CFF42F9386773F91C" bold="true" box="[526,554,1718,1742]" pageId="32" pageNumber="33">Pa</emphasis>
posterior arm,
<emphasis id="B935EAD6BA2DFE0CFF89F93867BCF91C" bold="true" box="[709,741,1718,1742]" pageId="32" pageNumber="33">SD</emphasis>
seminal duct,
<emphasis id="B935EAD6BA2DFE0CFE35F93866C1F91C" bold="true" box="[889,920,1718,1742]" pageId="32" pageNumber="33">ST</emphasis>
subtegulum,
<emphasis id="B935EAD6BA2DFE0CF96DF938616BF91C" bold="true" box="[1057,1074,1718,1742]" pageId="32" pageNumber="33">T</emphasis>
tegulum,
<emphasis id="B935EAD6BA2DFE0CF9D4F93861E0F91C" bold="true" box="[1176,1209,1718,1742]" pageId="32" pageNumber="33">TA</emphasis>
terminal apophysis.
</paragraph>
</caption>
<subSubSection id="C35B654FBA2DFE0CFD8BF8AE64FBF874" pageId="32" pageNumber="33" type="diagnosis">
<paragraph id="8BFE36C4BA2DFE0CFD8BF8AE64FBF874" blockId="32.[151,1437,1824,1958]" pageId="32" pageNumber="33">
<emphasis id="B935EAD6BA2DFE0CFD8BF8AE641BF8E8" bold="true" box="[199,322,1824,1850]" pageId="32" pageNumber="33">Diagnosis.</emphasis>
Males of
<taxonomicName id="4C414D47BA2DFE0CFC8EF8AE672CF8E8" authorityName="Paredes-Munguía &amp; Brescovit &amp; Teixeira" authorityYear="2024" box="[450,629,1824,1850]" class="Arachnida" family="Lycosidae" genus="Arctosa" kingdom="Animalia" order="Araneae" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="pacaya">
<emphasis id="B935EAD6BA2DFE0CFC8EF8AE672CF8E8" box="[450,629,1824,1850]" italics="true" pageId="32" pageNumber="33">Arctosa pacaya</emphasis>
</taxonomicName>
resemble
<taxonomicName id="4C414D47BA2DFE0CFFBEF8AE669FF8E8" authorityName="Silva &amp; Lise" authorityYear="2009" box="[754,966,1824,1850]" class="Arachnida" family="Lycosidae" genus="Arctosa" kingdom="Animalia" order="Araneae" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="sapiranga">
<emphasis id="B935EAD6BA2DFE0CFFBEF8AE669FF8E8" box="[754,966,1824,1850]" italics="true" pageId="32" pageNumber="33">Arctosa sapiranga</emphasis>
</taxonomicName>
in dorsal pattern of habitus but can be recognized by the wider carapace and the tibia/metatarsus I and II whitish (
<figureCitation id="137A2A41BA2DFE0CF945F8CA61D0F88C" box="[1033,1161,1860,1886]" captionStart="FIGURE 26" captionStartId="32.[151,250,1646,1670]" captionTargetBox="[151,1436,181,1621]" captionTargetId="figure-25@32.[151,1436,181,1621]" captionTargetPageId="32" captionText="FIGURE 26. Arctosa pacaya sp. nov. Male habitus (MUSM-ENT 505356) ac (a, dorsal; b, ventral; c, lateral). Pedipalp, expanded d, e (d, prolateral; e, anterior). Scale bars: ac, 1 mm, d, e, 200 µm. BH basal hematodocha, CMS cymbial macrosetae, DH distal hematodocha, E embolus, Pa posterior arm, SD seminal duct, ST subtegulum, T tegulum, TA terminal apophysis." figureDoi="http://doi.org/10.5281/zenodo.10681317" httpUri="https://zenodo.org/record/10681317/files/figure.png" pageId="32" pageNumber="33">Fig. 26a, c</figureCitation>
); the male palpus of
<taxonomicName id="4C414D47BA2DFE0CF8C8F8CA65B1F850" authorityName="Paredes-Munguía &amp; Brescovit &amp; Teixeira" authorityYear="2024" class="Arachnida" family="Lycosidae" genus="Arctosa" kingdom="Animalia" order="Araneae" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="pacaya">
<emphasis id="B935EAD6BA2DFE0CF8C8F8CA65B1F850" italics="true" pageId="32" pageNumber="33">A. pacaya</emphasis>
</taxonomicName>
have the embolus twisted once in the middle part (
<figureCitation id="137A2A41BA2DFE0CFE5BF8E6662EF851" box="[791,887,1896,1923]" captionStart="FIGURE 25" captionStartId="31.[151,250,1908,1932]" captionTargetBox="[188,1400,181,1884]" captionTargetId="figure-21@31.[188,1400,181,1884]" captionTargetPageId="31" captionText="FIGURE 25. Arctosa pacaya sp. nov. Male pedipalp (MUSM-ENT 505356) ac (a, retrolateral; b, ventral; c, prolateral). Pedipalp bulb df, h (d, retrolateral; e, ventral; f, prolateral; h, anterior). g, apical cymbium setae, i, protuberance on male chelicerae. Scale bars: ac, 0.2 mm, dh, 100 µm, i, 125 µm. C conductor, DMA depresion of median apophysis, E embolus, P palea, Pa posterior arm, SD seminal duct, ST subtegulum, T tegulum, TA terminal apophysis." figureDoi="http://doi.org/10.5281/zenodo.10681315" httpUri="https://zenodo.org/record/10681315/files/figure.png" pageId="32" pageNumber="33">Fig. 25b</figureCitation>
), and a T-shaped posterior arm (Pa) of the median apophysis (
<figureCitation id="137A2A41BA2DFE0CFC5AF80264CDF874" box="[278,404,1932,1958]" captionStart="FIGURE 24" captionStartId="30.[151,250,1977,2001]" captionTargetBox="[174,1413,841,1944]" captionTargetId="figure-259@30.[164,1423,833,1954]" captionTargetPageId="30" captionText="FIGURE 24. Arctosa pacaya sp. nov. Holotype male pedipalp (MUSM-ENT 505356) ac (a, ventral; b, retrolateral; c, anterior). Scale bars: ac, 0.25 mm. CMS cymbium macrosetae, Pa posterior arm, Ra retrolateral arm, TA terminal apophysis." figureDoi="http://doi.org/10.5281/zenodo.10681313" httpUri="https://zenodo.org/record/10681313/files/figure.png" pageId="32" pageNumber="33">Figs 24a, c</figureCitation>
).
</paragraph>
</subSubSection>
<caption id="DF3E664CBA2CFE0DFDDBFBE26616FB1E" ID-DOI="http://doi.org/10.5281/zenodo.10681319" ID-Zenodo-Dep="10681319" httpUri="https://zenodo.org/record/10681319/files/figure.png" pageId="33" pageNumber="34" startId="33.[151,250,1132,1156]" targetBox="[151,1436,181,1108]" targetPageId="33" targetType="figure">
<paragraph id="8BFE36C4BA2CFE0DFDDBFBE26616FB1E" blockId="33.[151,1437,1132,1228]" pageId="33" pageNumber="34">
<emphasis id="B935EAD6BA2CFE0DFDDBFBE26478FB56" bold="true" box="[151,289,1132,1156]" pageId="33" pageNumber="34">FIGURE 27.</emphasis>
<taxonomicName id="4C414D47BA2CFE0DFC65FBE2649EFB56" authority="Paredes-Munguía &amp; Brescovit &amp; Teixeira, 2024" authorityName="Paredes-Munguía &amp; Brescovit &amp; Teixeira" authorityYear="2024" box="[297,455,1132,1156]" class="Arachnida" family="Lycosidae" genus="Arctosa" kingdom="Animalia" order="Araneae" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="pacaya" status="sp. nov.">
<emphasis id="B935EAD6BA2CFE0DFC65FBE2649EFB56" box="[297,455,1132,1156]" italics="true" pageId="33" pageNumber="34">Arctosa pacaya</emphasis>
</taxonomicName>
<emphasis id="B935EAD6BA2CFE0DFC83FBE26778FB56" bold="true" box="[463,545,1132,1156]" pageId="33" pageNumber="34">
<taxonomicNameLabel id="A20657ADBA2CFE0DFC83FBE26778FB56" box="[463,545,1132,1156]" pageId="33" pageNumber="34" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Male pedipalp SEM (MUSM-ENT 505356)
<emphasis id="B935EAD6BA2CFE0DFEBDFBE2614FFB56" bold="true" box="[1009,1046,1132,1156]" pageId="33" pageNumber="34">ac</emphasis>
(a, ventral; b, TA detail; c, Pa detail). Scale bars: a, 150 µm, b, 100 µm, c, 50 µm.
<emphasis id="B935EAD6BA2CFE0DFF24FB1E6723FB7A" bold="true" box="[616,634,1168,1192]" pageId="33" pageNumber="34">C</emphasis>
conductor,
<emphasis id="B935EAD6BA2CFE0DFFB9FB1E665FFB7A" bold="true" box="[757,774,1168,1192]" pageId="33" pageNumber="34">E</emphasis>
embolus,
<emphasis id="B935EAD6BA2CFE0DFE3DFB1E66C2FB7A" bold="true" box="[881,923,1168,1192]" pageId="33" pageNumber="34">MA</emphasis>
median apophysis,
<emphasis id="B935EAD6BA2CFE0DF924FB1E612EFB7A" bold="true" box="[1128,1143,1168,1192]" pageId="33" pageNumber="34">P</emphasis>
palea,
<emphasis id="B935EAD6BA2CFE0DF98FFB1E6186FB7A" bold="true" box="[1219,1247,1168,1192]" pageId="33" pageNumber="34">Pa</emphasis>
posterior arm,
<emphasis id="B935EAD6BA2CFE0DF831FB1E60C5FB7A" bold="true" box="[1405,1436,1168,1192]" pageId="33" pageNumber="34">Ra</emphasis>
retrolateral arm,
<emphasis id="B935EAD6BA2CFE0DFC0DFB3A6439FB1E" bold="true" box="[321,352,1204,1228]" pageId="33" pageNumber="34">ST</emphasis>
subtegulum,
<emphasis id="B935EAD6BA2CFE0DFCA5FB3A64A3FB1E" bold="true" box="[489,506,1204,1228]" pageId="33" pageNumber="34">T</emphasis>
tegulum,
<emphasis id="B935EAD6BA2CFE0DFF2CFB3A67D8FB1E" bold="true" box="[608,641,1204,1228]" pageId="33" pageNumber="34">TA</emphasis>
terminal apophysis.
</paragraph>
</caption>
<subSubSection id="C35B654FBA2CFE0EFDDBFA906617FDCB" lastPageId="34" lastPageNumber="35" pageId="33" pageNumber="34" type="description">
<paragraph id="8BFE36C4BA2CFE0DFDDBFA906751FAEA" blockId="33.[151,1437,1310,2021]" box="[151,520,1310,1336]" pageId="33" pageNumber="34">
<emphasis id="B935EAD6BA2CFE0DFDDBFA906470FAEA" bold="true" box="[151,297,1310,1336]" pageId="33" pageNumber="34">Description.</emphasis>
<emphasis id="B935EAD6BA2CFE0DFC7CFA9064D2FAEA" box="[304,395,1310,1336]" italics="true" pageId="33" pageNumber="34">Female.</emphasis>
Unknown.
</paragraph>
<paragraph id="8BFE36C4BA2CFE0DFD8BFACC67E1F8DE" blockId="33.[151,1437,1310,2021]" pageId="33" pageNumber="34">
<emphasis id="B935EAD6BA2CFE0DFD8BFACC6727FA8E" box="[199,638,1346,1372]" italics="true" pageId="33" pageNumber="34">
Male.
<typeStatus id="54FA8866BA2CFE0DFC42FACC642CFA8E" box="[270,373,1346,1372]" pageId="33" pageNumber="34" type="holotype">Holotype</typeStatus>
(MUSM-ENT 505356).
</emphasis>
Dorsum of Prosoma light brown sparsely covered with tiny black setae and a pale yellow longitudinal median band tapering to fovea; non-recognizable submarginal bands (
<figureCitation id="137A2A41BA2CFE0DF9B6FAE86003FA52" box="[1274,1370,1382,1408]" captionStart="FIGURE 26" captionStartId="32.[151,250,1646,1670]" captionTargetBox="[151,1436,181,1621]" captionTargetId="figure-25@32.[151,1436,181,1621]" captionTargetPageId="32" captionText="FIGURE 26. Arctosa pacaya sp. nov. Male habitus (MUSM-ENT 505356) ac (a, dorsal; b, ventral; c, lateral). Pedipalp, expanded d, e (d, prolateral; e, anterior). Scale bars: ac, 1 mm, d, e, 200 µm. BH basal hematodocha, CMS cymbial macrosetae, DH distal hematodocha, E embolus, Pa posterior arm, SD seminal duct, ST subtegulum, T tegulum, TA terminal apophysis." figureDoi="http://doi.org/10.5281/zenodo.10681317" httpUri="https://zenodo.org/record/10681317/files/figure.png" pageId="33" pageNumber="34">Fig. 26a</figureCitation>
), four long black bristles on the clypeal condyle. Ocular area is the same color as the longitudinal median band, covered by a sparse white seta. First row of eyes procurved and shorter than the second row; AME slightly bigger than ALE; ALE resting on tubercles distally oriented; PME separated by more than half the length of its radii. anterior to AER. Sternum yellow sparsely covered with brown bristles (
<figureCitation id="137A2A41BA2CFE0DFE77FA7866C4F9C2" box="[827,925,1526,1552]" captionStart="FIGURE 26" captionStartId="32.[151,250,1646,1670]" captionTargetBox="[151,1436,181,1621]" captionTargetId="figure-25@32.[151,1436,181,1621]" captionTargetPageId="32" captionText="FIGURE 26. Arctosa pacaya sp. nov. Male habitus (MUSM-ENT 505356) ac (a, dorsal; b, ventral; c, lateral). Pedipalp, expanded d, e (d, prolateral; e, anterior). Scale bars: ac, 1 mm, d, e, 200 µm. BH basal hematodocha, CMS cymbial macrosetae, DH distal hematodocha, E embolus, Pa posterior arm, SD seminal duct, ST subtegulum, T tegulum, TA terminal apophysis." figureDoi="http://doi.org/10.5281/zenodo.10681317" httpUri="https://zenodo.org/record/10681317/files/figure.png" pageId="33" pageNumber="34">Fig. 26b</figureCitation>
). Labium yellow covered with brown bristles, border of the labium and endite clearer than the base; chelicerae dark brown, darker than the endites and labium. Condyle short and poorly notorious, pointed bulge in the outer margin of the cheliceral claw (
<figureCitation id="137A2A41BA2CFE0DF9EDF9B061A2F98B" box="[1185,1275,1598,1625]" captionStart="FIGURE 25" captionStartId="31.[151,250,1908,1932]" captionTargetBox="[188,1400,181,1884]" captionTargetId="figure-21@31.[188,1400,181,1884]" captionTargetPageId="31" captionText="FIGURE 25. Arctosa pacaya sp. nov. Male pedipalp (MUSM-ENT 505356) ac (a, retrolateral; b, ventral; c, prolateral). Pedipalp bulb df, h (d, retrolateral; e, ventral; f, prolateral; h, anterior). g, apical cymbium setae, i, protuberance on male chelicerae. Scale bars: ac, 0.2 mm, dh, 100 µm, i, 125 µm. C conductor, DMA depresion of median apophysis, E embolus, P palea, Pa posterior arm, SD seminal duct, ST subtegulum, T tegulum, TA terminal apophysis." figureDoi="http://doi.org/10.5281/zenodo.10681315" httpUri="https://zenodo.org/record/10681315/files/figure.png" pageId="33" pageNumber="34">Fig. 25i</figureCitation>
). Legs yellow, becoming lighter distally. All femora are yellow with dusky irregular patches; only the leg I white from tibiae to tarsi (
<figureCitation id="137A2A41BA2CFE0DFDD3F9086458F972" box="[159,257,1670,1696]" captionStart="FIGURE 26" captionStartId="32.[151,250,1646,1670]" captionTargetBox="[151,1436,181,1621]" captionTargetId="figure-25@32.[151,1436,181,1621]" captionTargetPageId="32" captionText="FIGURE 26. Arctosa pacaya sp. nov. Male habitus (MUSM-ENT 505356) ac (a, dorsal; b, ventral; c, lateral). Pedipalp, expanded d, e (d, prolateral; e, anterior). Scale bars: ac, 1 mm, d, e, 200 µm. BH basal hematodocha, CMS cymbial macrosetae, DH distal hematodocha, E embolus, Pa posterior arm, SD seminal duct, ST subtegulum, T tegulum, TA terminal apophysis." figureDoi="http://doi.org/10.5281/zenodo.10681317" httpUri="https://zenodo.org/record/10681317/files/figure.png" pageId="33" pageNumber="34">Fig. 26c</figureCitation>
). All tarsi with no scopula, spinules on tarsi III and IV; trichobothria in tibiae, tarsi and metatarsi distally increasing in size. Opisthosoma dorsal and lateral shield olive gray; a broad yellow longitudinal band runs along the spinnerets finishing in not defined chevron pattern stripes, long black bristles in the yellow band; venter yellow with some irregular dusky patches; spinnerets yellow.
</paragraph>
<paragraph id="8BFE36C4BA2CFE0DFD8BF89864B6F836" blockId="33.[151,1437,1310,2021]" pageId="33" pageNumber="34">
Pedipalp (
<figureCitation id="137A2A41BA2CFE0DFC75F89864E3F8E3" box="[313,442,1814,1841]" captionStart="FIGURE 25" captionStartId="31.[151,250,1908,1932]" captionTargetBox="[188,1400,181,1884]" captionTargetId="figure-21@31.[188,1400,181,1884]" captionTargetPageId="31" captionText="FIGURE 25. Arctosa pacaya sp. nov. Male pedipalp (MUSM-ENT 505356) ac (a, retrolateral; b, ventral; c, prolateral). Pedipalp bulb df, h (d, retrolateral; e, ventral; f, prolateral; h, anterior). g, apical cymbium setae, i, protuberance on male chelicerae. Scale bars: ac, 0.2 mm, dh, 100 µm, i, 125 µm. C conductor, DMA depresion of median apophysis, E embolus, P palea, Pa posterior arm, SD seminal duct, ST subtegulum, T tegulum, TA terminal apophysis." figureDoi="http://doi.org/10.5281/zenodo.10681315" httpUri="https://zenodo.org/record/10681315/files/figure.png" pageId="33" pageNumber="34">Figs 25ah</figureCitation>
,
<figureCitation id="137A2A41BA2CFE0DFC8AF8986755F8E3" box="[454,524,1814,1841]" captionStart="FIGURE 26" captionStartId="32.[151,250,1646,1670]" captionTargetBox="[151,1436,181,1621]" captionTargetId="figure-25@32.[151,1436,181,1621]" captionTargetPageId="32" captionText="FIGURE 26. Arctosa pacaya sp. nov. Male habitus (MUSM-ENT 505356) ac (a, dorsal; b, ventral; c, lateral). Pedipalp, expanded d, e (d, prolateral; e, anterior). Scale bars: ac, 1 mm, d, e, 200 µm. BH basal hematodocha, CMS cymbial macrosetae, DH distal hematodocha, E embolus, Pa posterior arm, SD seminal duct, ST subtegulum, T tegulum, TA terminal apophysis." figureDoi="http://doi.org/10.5281/zenodo.10681317" httpUri="https://zenodo.org/record/10681317/files/figure.png" pageId="33" pageNumber="34">26d, e</figureCitation>
,
<figureCitation id="137A2A41BA2CFE0DFF56F898676EF8E2" box="[538,567,1814,1840]" captionStart="FIGURE 27" captionStartId="33.[151,250,1132,1156]" captionTargetBox="[151,1436,181,1108]" captionTargetId="figure-21@33.[151,1436,181,1108]" captionTargetPageId="33" captionText="FIGURE 27. Arctosa pacaya sp. nov. Male pedipalp SEM (MUSM-ENT 505356) ac (a, ventral; b, TA detail; c, Pa detail). Scale bars: a, 150 µm, b, 100 µm, c, 50 µm. C conductor, E embolus, MA median apophysis, P palea, Pa posterior arm, Ra retrolateral arm, ST subtegulum, T tegulum, TA terminal apophysis." figureDoi="http://doi.org/10.5281/zenodo.10681319" httpUri="https://zenodo.org/record/10681319/files/figure.png" pageId="33" pageNumber="34">27</figureCitation>
) femora slightly curved in the dorsal view. Cymbium with a short but strong subterminal spine in the apical area (
<figureCitation id="137A2A41BA2CFE0DFF60F8B467D4F886" box="[556,653,1850,1876]" captionStart="FIGURE 26" captionStartId="32.[151,250,1646,1670]" captionTargetBox="[151,1436,181,1621]" captionTargetId="figure-25@32.[151,1436,181,1621]" captionTargetPageId="32" captionText="FIGURE 26. Arctosa pacaya sp. nov. Male habitus (MUSM-ENT 505356) ac (a, dorsal; b, ventral; c, lateral). Pedipalp, expanded d, e (d, prolateral; e, anterior). Scale bars: ac, 1 mm, d, e, 200 µm. BH basal hematodocha, CMS cymbial macrosetae, DH distal hematodocha, E embolus, Pa posterior arm, SD seminal duct, ST subtegulum, T tegulum, TA terminal apophysis." figureDoi="http://doi.org/10.5281/zenodo.10681317" httpUri="https://zenodo.org/record/10681317/files/figure.png" pageId="33" pageNumber="34">Fig. 26d</figureCitation>
). Sperm duct crossing the tegulum in a C-shaped pathway, subtegulum visible and bowl shaped (
<figureCitation id="137A2A41BA2CFE0DFCF4F8D06765F8AA" box="[440,572,1886,1913]" captionStart="FIGURE 25" captionStartId="31.[151,250,1908,1932]" captionTargetBox="[188,1400,181,1884]" captionTargetId="figure-21@31.[188,1400,181,1884]" captionTargetPageId="31" captionText="FIGURE 25. Arctosa pacaya sp. nov. Male pedipalp (MUSM-ENT 505356) ac (a, retrolateral; b, ventral; c, prolateral). Pedipalp bulb df, h (d, retrolateral; e, ventral; f, prolateral; h, anterior). g, apical cymbium setae, i, protuberance on male chelicerae. Scale bars: ac, 0.2 mm, dh, 100 µm, i, 125 µm. C conductor, DMA depresion of median apophysis, E embolus, P palea, Pa posterior arm, SD seminal duct, ST subtegulum, T tegulum, TA terminal apophysis." figureDoi="http://doi.org/10.5281/zenodo.10681315" httpUri="https://zenodo.org/record/10681315/files/figure.png" pageId="33" pageNumber="34">Figs 25b, d</figureCitation>
). In ventral view, Median apophysis with an evident posterior arm T-shaped ventrally projected (
<figureCitation id="137A2A41BA2CFE0DFC3BF80C64ACF84F" box="[375,501,1922,1949]" captionStart="FIGURE 25" captionStartId="31.[151,250,1908,1932]" captionTargetBox="[188,1400,181,1884]" captionTargetId="figure-21@31.[188,1400,181,1884]" captionTargetPageId="31" captionText="FIGURE 25. Arctosa pacaya sp. nov. Male pedipalp (MUSM-ENT 505356) ac (a, retrolateral; b, ventral; c, prolateral). Pedipalp bulb df, h (d, retrolateral; e, ventral; f, prolateral; h, anterior). g, apical cymbium setae, i, protuberance on male chelicerae. Scale bars: ac, 0.2 mm, dh, 100 µm, i, 125 µm. C conductor, DMA depresion of median apophysis, E embolus, P palea, Pa posterior arm, SD seminal duct, ST subtegulum, T tegulum, TA terminal apophysis." figureDoi="http://doi.org/10.5281/zenodo.10681315" httpUri="https://zenodo.org/record/10681315/files/figure.png" pageId="33" pageNumber="34">Figs 25b, e</figureCitation>
,
<figureCitation id="137A2A41BA2CFE0DFF4CF80C6774F84E" box="[512,557,1922,1948]" captionStart="FIGURE 27" captionStartId="33.[151,250,1132,1156]" captionTargetBox="[151,1436,181,1108]" captionTargetId="figure-21@33.[151,1436,181,1108]" captionTargetPageId="33" captionText="FIGURE 27. Arctosa pacaya sp. nov. Male pedipalp SEM (MUSM-ENT 505356) ac (a, ventral; b, TA detail; c, Pa detail). Scale bars: a, 150 µm, b, 100 µm, c, 50 µm. C conductor, E embolus, MA median apophysis, P palea, Pa posterior arm, Ra retrolateral arm, ST subtegulum, T tegulum, TA terminal apophysis." figureDoi="http://doi.org/10.5281/zenodo.10681319" httpUri="https://zenodo.org/record/10681319/files/figure.png" pageId="33" pageNumber="34">27c</figureCitation>
). The basal portion of the terminal apophysis is more sclerotized than the distal and ribbed with its distal part sharpened (
<figureCitation id="137A2A41BA2CFE0DFF21F8286788F812" box="[621,721,1958,1984]" captionStart="FIGURE 24" captionStartId="30.[151,250,1977,2001]" captionTargetBox="[174,1413,841,1944]" captionTargetId="figure-259@30.[164,1423,833,1954]" captionTargetPageId="30" captionText="FIGURE 24. Arctosa pacaya sp. nov. Holotype male pedipalp (MUSM-ENT 505356) ac (a, ventral; b, retrolateral; c, anterior). Scale bars: ac, 0.25 mm. CMS cymbium macrosetae, Pa posterior arm, Ra retrolateral arm, TA terminal apophysis." figureDoi="http://doi.org/10.5281/zenodo.10681313" httpUri="https://zenodo.org/record/10681313/files/figure.png" pageId="33" pageNumber="34">Figs 24a</figureCitation>
,
<figureCitation id="137A2A41BA2CFE0DFF92F8286655F813" box="[734,780,1958,1985]" captionStart="FIGURE 25" captionStartId="31.[151,250,1908,1932]" captionTargetBox="[188,1400,181,1884]" captionTargetId="figure-21@31.[188,1400,181,1884]" captionTargetPageId="31" captionText="FIGURE 25. Arctosa pacaya sp. nov. Male pedipalp (MUSM-ENT 505356) ac (a, retrolateral; b, ventral; c, prolateral). Pedipalp bulb df, h (d, retrolateral; e, ventral; f, prolateral; h, anterior). g, apical cymbium setae, i, protuberance on male chelicerae. Scale bars: ac, 0.2 mm, dh, 100 µm, i, 125 µm. C conductor, DMA depresion of median apophysis, E embolus, P palea, Pa posterior arm, SD seminal duct, ST subtegulum, T tegulum, TA terminal apophysis." figureDoi="http://doi.org/10.5281/zenodo.10681315" httpUri="https://zenodo.org/record/10681315/files/figure.png" pageId="33" pageNumber="34">25b</figureCitation>
). Embolus is fusiform, sclerotized, and twisted once in its median part (
<figureCitation id="137A2A41BA2CFE0DFC67F84464F3F837" box="[299,426,1994,2021]" captionStart="FIGURE 25" captionStartId="31.[151,250,1908,1932]" captionTargetBox="[188,1400,181,1884]" captionTargetId="figure-21@31.[188,1400,181,1884]" captionTargetPageId="31" captionText="FIGURE 25. Arctosa pacaya sp. nov. Male pedipalp (MUSM-ENT 505356) ac (a, retrolateral; b, ventral; c, prolateral). Pedipalp bulb df, h (d, retrolateral; e, ventral; f, prolateral; h, anterior). g, apical cymbium setae, i, protuberance on male chelicerae. Scale bars: ac, 0.2 mm, dh, 100 µm, i, 125 µm. C conductor, DMA depresion of median apophysis, E embolus, P palea, Pa posterior arm, SD seminal duct, ST subtegulum, T tegulum, TA terminal apophysis." figureDoi="http://doi.org/10.5281/zenodo.10681315" httpUri="https://zenodo.org/record/10681315/files/figure.png" pageId="33" pageNumber="34">Figs 25b, e</figureCitation>
,
<figureCitation id="137A2A41BA2CFE0DFCFAF84464BAF836" box="[438,483,1994,2020]" captionStart="FIGURE 27" captionStartId="33.[151,250,1132,1156]" captionTargetBox="[151,1436,181,1108]" captionTargetId="figure-21@33.[151,1436,181,1108]" captionTargetPageId="33" captionText="FIGURE 27. Arctosa pacaya sp. nov. Male pedipalp SEM (MUSM-ENT 505356) ac (a, ventral; b, TA detail; c, Pa detail). Scale bars: a, 150 µm, b, 100 µm, c, 50 µm. C conductor, E embolus, MA median apophysis, P palea, Pa posterior arm, Ra retrolateral arm, ST subtegulum, T tegulum, TA terminal apophysis." figureDoi="http://doi.org/10.5281/zenodo.10681319" httpUri="https://zenodo.org/record/10681319/files/figure.png" pageId="33" pageNumber="34">27a</figureCitation>
).
</paragraph>
<paragraph id="8BFE36C4BA2FFE0EFD8BFF1966F5FE93" blockId="34.[151,1437,151,573]" pageId="34" pageNumber="35">
Leg formula IV&gt;I&gt;II&gt;III. Spination pattern: femur
<emphasis id="B935EAD6BA2FFE0EFE4EFF19667DFF63" bold="true" box="[770,804,151,177]" pageId="34" pageNumber="35">I p</emphasis>
0-0-2
<emphasis id="B935EAD6BA2FFE0EFE2BFF19662EFF63" bold="true" box="[871,887,151,177]" pageId="34" pageNumber="35">d</emphasis>
1-1-1
<emphasis id="B935EAD6BA2FFE0EFEF5FF19669CFF63" bold="true" box="[953,965,151,177]" pageId="34" pageNumber="35">r</emphasis>
0-1-1,
<emphasis id="B935EAD6BA2FFE0EF95CFF196165FF63" bold="true" box="[1040,1084,151,177]" pageId="34" pageNumber="35">II p</emphasis>
0-0-2
<emphasis id="B935EAD6BA2FFE0EF933FF1961D6FF63" bold="true" box="[1151,1167,151,177]" pageId="34" pageNumber="35">d</emphasis>
1-1-1
<emphasis id="B935EAD6BA2FFE0EF99EFF196187FF63" bold="true" box="[1234,1246,151,177]" pageId="34" pageNumber="35">r</emphasis>
0-1-1,
<emphasis id="B935EAD6BA2FFE0EF865FF196039FF63" bold="true" box="[1321,1376,151,177]" pageId="34" pageNumber="35">III p</emphasis>
0-1-1
<emphasis id="B935EAD6BA2FFE0EFDDBFF3565FEFF07" bold="true" box="[151,167,187,213]" pageId="34" pageNumber="35">d</emphasis>
1-1-1
<emphasis id="B935EAD6BA2FFE0EFDA0FF3565A1FF07" bold="true" box="[236,248,187,213]" pageId="34" pageNumber="35">r</emphasis>
0-1-1,
<emphasis id="B935EAD6BA2FFE0EFC08FF356422FF07" bold="true" box="[324,379,187,213]" pageId="34" pageNumber="35">IV p</emphasis>
1-0-1
<emphasis id="B935EAD6BA2FFE0EFCF3FF356496FF07" bold="true" box="[447,463,187,213]" pageId="34" pageNumber="35">d</emphasis>
1-1-1
<emphasis id="B935EAD6BA2FFE0EFF5FFF356746FF07" bold="true" box="[531,543,187,213]" pageId="34" pageNumber="35">r</emphasis>
0-0-1; patella
<emphasis id="B935EAD6BA2FFE0EFFF3FF3567BBFF07" bold="true" box="[703,738,187,213]" pageId="34" pageNumber="35">I p</emphasis>
1
<emphasis id="B935EAD6BA2FFE0EFFB4FF356651FF07" bold="true" box="[760,776,187,213]" pageId="34" pageNumber="35">d</emphasis>
1bas-1ap
<emphasis id="B935EAD6BA2FFE0EFE39FF3566D8FF07" bold="true" box="[885,897,187,213]" pageId="34" pageNumber="35">r</emphasis>
1,
<emphasis id="B935EAD6BA2FFE0EFED2FF356695FF07" bold="true" box="[926,972,187,213]" pageId="34" pageNumber="35">II p</emphasis>
1
<emphasis id="B935EAD6BA2FFE0EFEAEFF3566ABFF07" bold="true" box="[994,1010,187,213]" pageId="34" pageNumber="35">d</emphasis>
1bas-1ap
<emphasis id="B935EAD6BA2FFE0EF913FF356132FF07" bold="true" box="[1119,1131,187,213]" pageId="34" pageNumber="35">r</emphasis>
1,
<emphasis id="B935EAD6BA2FFE0EF9C5FF356198FF07" bold="true" box="[1161,1217,187,213]" pageId="34" pageNumber="35">III p</emphasis>
1
<emphasis id="B935EAD6BA2FFE0EF99BFF3561BEFF07" bold="true" box="[1239,1255,187,213]" pageId="34" pageNumber="35">d</emphasis>
1bas-1ap
<emphasis id="B935EAD6BA2FFE0EF818FF356039FF07" bold="true" box="[1364,1376,187,213]" pageId="34" pageNumber="35">r</emphasis>
1,
<emphasis id="B935EAD6BA2FFE0EF832FF3565FEFF2B" bold="true" pageId="34" pageNumber="35">IV p</emphasis>
1
<emphasis id="B935EAD6BA2FFE0EFDF0FF516595FF2B" bold="true" box="[188,204,223,249]" pageId="34" pageNumber="35">d</emphasis>
1bas-1ap
<emphasis id="B935EAD6BA2FFE0EFC7BFF51641AFF2B" bold="true" box="[311,323,223,249]" pageId="34" pageNumber="35">r</emphasis>
1; tibia
<emphasis id="B935EAD6BA2FFE0EFCD5FF5164E3FF2B" bold="true" box="[409,442,223,249]" pageId="34" pageNumber="35">I p</emphasis>
1-1v
<emphasis id="B935EAD6BA2FFE0EFCB8FF51675BFF2B" bold="true" box="[500,514,223,249]" pageId="34" pageNumber="35">v</emphasis>
2-2-2ap
<emphasis id="B935EAD6BA2FFE0EFF2CFF516735FF2B" bold="true" box="[608,620,223,249]" pageId="34" pageNumber="35">r</emphasis>
1-1v,
<emphasis id="B935EAD6BA2FFE0EFFE0FF516781FF2B" bold="true" box="[684,728,223,249]" pageId="34" pageNumber="35">II p</emphasis>
1-1
<emphasis id="B935EAD6BA2FFE0EFE48FF51664DFF2B" bold="true" box="[772,788,223,249]" pageId="34" pageNumber="35">d</emphasis>
1-(1 bristle)
<emphasis id="B935EAD6BA2FFE0EFED3FF5166F2FF2B" bold="true" box="[927,939,223,249]" pageId="34" pageNumber="35">r</emphasis>
1-1
<emphasis id="B935EAD6BA2FFE0EFE94FF5166BFFF2B" bold="true" box="[984,998,223,249]" pageId="34" pageNumber="35">v</emphasis>
2-2-2ap
<emphasis id="B935EAD6BA2FFE0EF908FF51613CFF2B" bold="true" box="[1092,1125,223,249]" pageId="34" pageNumber="35">III</emphasis>
,
<emphasis id="B935EAD6BA2FFE0EF93EFF5161DBFF2B" bold="true" box="[1138,1154,223,249]" pageId="34" pageNumber="35">p</emphasis>
d1-1
<emphasis id="B935EAD6BA2FFE0EF9F0FF516195FF2B" bold="true" box="[1212,1228,223,249]" pageId="34" pageNumber="35">d</emphasis>
1-1
<emphasis id="B935EAD6BA2FFE0EF9B4FF51605DFF2B" bold="true" box="[1272,1284,223,249]" pageId="34" pageNumber="35">r</emphasis>
1-1
<emphasis id="B935EAD6BA2FFE0EF87CFF516067FF2B" bold="true" box="[1328,1342,223,249]" pageId="34" pageNumber="35">v</emphasis>
2
<emphasis id="B935EAD6BA2FFE0EF800FF51600CFF2B" bold="true" box="[1356,1365,223,249]" pageId="34" pageNumber="35">-</emphasis>
2-2ap,
<emphasis id="B935EAD6BA2FFE0EFDDBFE8D6595FECF" bold="true" box="[151,204,259,285]" pageId="34" pageNumber="35">IV p</emphasis>
1-1
<emphasis id="B935EAD6BA2FFE0EFDBBFE8D645EFECF" bold="true" box="[247,263,259,285]" pageId="34" pageNumber="35">d</emphasis>
1-1
<emphasis id="B935EAD6BA2FFE0EFC7EFE8D640DFECF" bold="true" box="[306,340,259,285]" pageId="34" pageNumber="35">r d</emphasis>
1-1
<emphasis id="B935EAD6BA2FFE0EFC33FE8D64D4FECF" bold="true" box="[383,397,259,285]" pageId="34" pageNumber="35">v</emphasis>
2-2-2ap; metatarsus
<emphasis id="B935EAD6BA2FFE0EFF3CFE8D67C8FECF" bold="true" box="[624,657,259,285]" pageId="34" pageNumber="35">I p</emphasis>
0-1-2ap
<emphasis id="B935EAD6BA2FFE0EFFA2FE8D67A3FECF" bold="true" box="[750,762,259,285]" pageId="34" pageNumber="35">r</emphasis>
0-12ap
<emphasis id="B935EAD6BA2FFE0EFE02FE8D6605FECF" bold="true" box="[846,860,259,285]" pageId="34" pageNumber="35">v</emphasis>
2-2-1ap,
<emphasis id="B935EAD6BA2FFE0EFE8CFE8D66B5FECF" bold="true" box="[960,1004,259,285]" pageId="34" pageNumber="35">II p</emphasis>
0-1-2ap
<emphasis id="B935EAD6BA2FFE0EF905FE8D610CFECF" bold="true" box="[1097,1109,259,285]" pageId="34" pageNumber="35">r</emphasis>
0-1-2ap
<emphasis id="B935EAD6BA2FFE0EF9FEFE8D6199FECF" bold="true" box="[1202,1216,259,285]" pageId="34" pageNumber="35">v</emphasis>
2-2-1ap,
<emphasis id="B935EAD6BA2FFE0EF869FE8D6002FECF" bold="true" box="[1317,1371,259,285]" pageId="34" pageNumber="35">III p</emphasis>
1d-1d
<emphasis id="B935EAD6BA2FFE0EFDDBFEA965FAFE93" bold="true" box="[151,163,295,321]" pageId="34" pageNumber="35">r</emphasis>
1d-1v-1d-0-1ap
<emphasis id="B935EAD6BA2FFE0EFC16FEA96431FE93" bold="true" box="[346,360,295,321]" pageId="34" pageNumber="35">v</emphasis>
2-2-1ap,
<emphasis id="B935EAD6BA2FFE0EFC81FEA9675AFE93" bold="true" box="[461,515,295,321]" pageId="34" pageNumber="35">IV p</emphasis>
1d-1d-2ap
<emphasis id="B935EAD6BA2FFE0EFF31FEA967D0FE93" bold="true" box="[637,649,295,321]" pageId="34" pageNumber="35">r</emphasis>
1d-1v-1d-0-2ap
<emphasis id="B935EAD6BA2FFE0EFE0CFEA96617FE93" bold="true" box="[832,846,295,321]" pageId="34" pageNumber="35">v</emphasis>
2-2-1ap.
</paragraph>
<paragraph id="8BFE36C4BA2FFE0EFD8BFEC56617FDCB" blockId="34.[151,1437,151,573]" pageId="34" pageNumber="35">
Measurements: TL 4.92, CL 2.83, CCW 2.14, CTW 1.22, CCH 1.10, CTH 1.16. Eyes: AME 0.50, ALE 0.33, PME 1.50, PLE 0.26, interdistances: AME-AME 0.50, AME-ALE 0.33; PME-PME 0.24, POQ length 0.58, POQ posterior width 0.90, POQ anterior width 0.56. Chelicerae: 0.92 length. Opisthosoma: length 2.17, width 1.33. Legs: length of segments femur + patella+tibia + metatarsus + tarsus = total length:
<emphasis id="B935EAD6BA2FFE0EFEBDFE386100FE02" bold="true" box="[1009,1113,438,464]" pageId="34" pageNumber="35">pedipalp</emphasis>
1.16 + 0.60 + - + 0.82 + 1.00 = 3.58,
<emphasis id="B935EAD6BA2FFE0EFDA1FE5565A1FE27" bold="true" box="[237,248,475,501]" pageId="34" pageNumber="35">I</emphasis>
2.00 + 0.80 + 1.70 + 1.64+1.08 = 7.22,
<emphasis id="B935EAD6BA2FFE0EFFF0FE55678BFE27" bold="true" box="[700,722,475,501]" pageId="34" pageNumber="35">II</emphasis>
1.80 + 0.82 + 1.52 + 1.56+1.02 = 6.72,
<emphasis id="B935EAD6BA2FFE0EF9D9FE5561EFFE27" bold="true" box="[1173,1206,475,501]" pageId="34" pageNumber="35">III</emphasis>
1.76
<emphasis id="B935EAD6BA2FFE0EF9B9FE55605CFE27" bold="true" box="[1269,1285,475,501]" pageId="34" pageNumber="35">+</emphasis>
0.66 +1.30 + 1.66+0.92 = 6.30,
<emphasis id="B935EAD6BA2FFE0EFC2AFE7164DCFDCB" bold="true" box="[358,389,511,537]" pageId="34" pageNumber="35">IV</emphasis>
2. 40 + 0.92 + 2. 10 + 2.74+1.30 = 9.46.
</paragraph>
</subSubSection>
<subSubSection id="C35B654FBA2FFE0EFD8BFDAC6687FDEF" box="[199,990,546,573]" pageId="34" pageNumber="35" type="distribution">
<paragraph id="8BFE36C4BA2FFE0EFD8BFDAC6687FDEF" blockId="34.[151,1437,151,573]" box="[199,990,546,573]" pageId="34" pageNumber="35">
<emphasis id="B935EAD6BA2FFE0EFD8BFDAC6438FDEE" bold="true" box="[199,353,546,572]" pageId="34" pageNumber="35">Distribution.</emphasis>
Known only for the
<typeStatus id="54FA8866BA2FFE0EFF07FDAD6722FDEF" box="[587,635,547,573]" pageId="34" pageNumber="35">type</typeStatus>
locality,
<collectingRegion id="4985F826BA2FFE0EFFAEFDAD6669FDEF" box="[738,816,547,573]" country="Peru" name="Loreto" pageId="34" pageNumber="35">Loreto</collectingRegion>
,
<collectingCountry id="F3567654BA2FFE0EFE77FDAD6637FDEF" box="[827,878,547,573]" name="Peru" pageId="34" pageNumber="35">Peru</collectingCountry>
(
<figureCitation id="137A2A41BA2FFE0EFE31FDAD668BFDEF" box="[893,978,547,573]" captionStart="FIGURE 62" captionStartId="72.[151,250,1618,1642]" captionTargetBox="[171,1418,564,1590]" captionTargetId="figure-197@72.[164,1423,561,1595]" captionTargetPageId="72" captionText="FIGURE 62. Geographic distribution records of Arctosa ayaymama sp. nov. (blue dot), Arctosa pacaya sp. nov. (red dot), Hogna andina comb. nov. (orange dot)." figureDoi="http://doi.org/10.5281/zenodo.10681391" httpUri="https://zenodo.org/record/10681391/files/figure.png" pageId="34" pageNumber="35">Fig. 62</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
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