216 lines
26 KiB
XML
216 lines
26 KiB
XML
<document id="27021E41AA694144F1FA2B275A7A3E16" ID-CLB-Dataset="4613" ID-DOI="10.5281/zenodo.5332860" ID-GBIF-Dataset="cc461ce1-8e44-4862-9ff2-9c9ef7f73cd9" ID-ISSN="0253-116X" ID-Zenodo-Dep="5332860" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630172711352" checkinUser="carolina" docAuthor="Guéorguiev, B." docDate="2010" docId="03E887AB59593153FF3D6CADFCFCEF5F" docLanguage="en" docName="LinzerbiolBeitr.2010.42.1.659-669.pdf" docOrigin="Linzer biologische Beiträge 42 (1)" docStyle="DocumentStyle:24E5EB772D355DE2EE23B1E8D07853E0.9:LinzerbiolBeitr.2006-.journal_article" docStyleId="24E5EB772D355DE2EE23B1E8D07853E0" docStyleName="LinzerbiolBeitr.2006-.journal_article" docStyleVersion="9" docTitle="Pterostichus (Oreoplatysma) colchicus" docType="treatment" docVersion="7" lastPageNumber="661" masterDocId="FFD1FFD3595B3151FF8F6C75FF94E951" masterDocTitle="The " colchicus " group of Pterostichus (Coleoptera, Carabidae)" masterLastPageNumber="669" masterPageNumber="659" pageNumber="661" updateTime="1699187604789" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="9E299435160F7010171FA9342F703358">The " colchicus " group of Pterostichus (Coleoptera, Carabidae)</mods:title>
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<mods:namePart id="1481BBDACA57D72D613B38DA37049B76">Guéorguiev, B.</mods:namePart>
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<mods:title id="78735A5B9DF446481954667A6D95B9B9">Linzer biologische Beiträge</mods:title>
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<mods:date id="AF1327E8C8EC97325AAE6965E24CBC4B">2010</mods:date>
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<mods:number id="5CABD4A8FFC472A53F3C83BE474E5AF5">42</mods:number>
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<treatment id="03E887AB59593153FF3D6CADFCFCEF5F" ID-DOI="http://doi.org/10.5281/zenodo.5411001" ID-GBIF-Taxon="185227617" ID-Zenodo-Dep="5411001" LSID="urn:lsid:plazi:treatment:03E887AB59593153FF3D6CADFCFCEF5F" httpUri="http://treatment.plazi.org/id/03E887AB59593153FF3D6CADFCFCEF5F" lastPageNumber="661" pageId="2" pageNumber="661">
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<subSubSection id="C35B653659593153FF3D6CADFDB5E845" pageId="2" pageNumber="661" type="nomenclature">
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<paragraph id="8BFE36BD59593153FF3D6CADFCE7E9BF" blockId="2.[178,883,216,238]" box="[178,883,216,238]" pageId="2" pageNumber="661">
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<heading id="D0B681D159593153FF3D6CADFCE7E9BF" box="[178,883,216,238]" fontSize="8" level="2" pageId="2" pageNumber="661" reason="2">
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<emphasis id="B935EAAF59593153FF3D6CADFCE7E9BF" bold="true" box="[178,883,216,238]" pageId="2" pageNumber="661">
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<taxonomicName id="4C414D3E59593153FF3D6CADFC93E9BF" ID-CoL="4Q452" authority="(CHAUDOIR 1850)" baseAuthorityName="CHAUDOIR" baseAuthorityYear="1850" box="[178,775,216,238]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="661" phylum="Arthropoda" rank="species" species="colchicus" subGenus="Oreoplatysma">
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<emphasis id="B935EAAF59593153FF3D6CADFEA7E9BF" bold="true" box="[178,307,216,238]" italics="true" pageId="2" pageNumber="661">Pterostichus</emphasis>
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(
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<emphasis id="B935EAAF59593153FECD6CADFE46E9BF" bold="true" box="[322,466,216,238]" italics="true" pageId="2" pageNumber="661">Oreoplatysma</emphasis>
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)
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<emphasis id="B935EAAF59593153FE6E6CADFDD7E9BF" bold="true" box="[481,579,216,238]" italics="true" pageId="2" pageNumber="661">colchicus</emphasis>
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(
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<bibRefCitation id="EFD04B4C59593153FDDD6CADFD52E9BF" author="CHAUDOIR M" box="[594,710,216,238]" pageId="2" pageNumber="661" pagination="62 - 206" refId="ref4221" refString="CHAUDOIR M. de (1850): Supplement a la faune des carabiques de la Russie. - Bulletin de la Societe Imperiale des Naturalistes de Moscou 23 (3): 62 - 206." type="journal article" year="1850">CHAUDOIR</bibRefCitation>
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1850)
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</taxonomicName>
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(
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<figureCitation id="137A2A3859593153FC9B6CADFCFFE9BF" box="[788,875,216,238]" captionStart="Figs 1-5" captionStartId="8.[178,218,911,931]" captionTargetBox="[188,1074,230,887]" captionTargetId="figure-2@8.[178,1085,220,900]" captionTargetPageId="8" captionText="Figs 1-5: Feronia colchica CHAUDOIR, lectotype: Fig. (1) aedeagus, left lateral view; Fig. (2) aedeagus, right lateral view; Fig. (3) apex of aedeagus, dorsal view; Fig. (4) right paramere, internal view; Fig. (5) right paramere, external view. Scale bar 0.5 mm (1-5)." figureDoi="http://doi.org/10.5281/zenodo.5332862" httpUri="https://zenodo.org/record/5332862/files/figure.png" pageId="2" pageNumber="661">Figs 1-5</figureCitation>
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)
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BFE36BD59593153FF3D6D75FDB5E845" blockId="2.[178,545,256,276]" box="[178,545,256,276]" pageId="2" pageNumber="661">
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<emphasis id="B935EAAF59593153FF3D6D75FEDAE845" box="[178,334,256,276]" italics="true" pageId="2" pageNumber="661">Feronia colchica</emphasis>
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<bibRefCitation id="EFD04B4C59593153FEDC6D75FD89E845" author="CHAUDOIR M" box="[339,541,256,276]" pageId="2" pageNumber="661" pagination="62 - 206" refId="ref4221" refString="CHAUDOIR M. de (1850): Supplement a la faune des carabiques de la Russie. - Bulletin de la Societe Imperiale des Naturalistes de Moscou 23 (3): 62 - 206." type="journal article" year="1850">CHAUDOIR 1850: 137</bibRefCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C35B653659593153FF3D6D57FCFCEF5F" pageId="2" pageNumber="661" type="description">
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<paragraph id="8BFE36BD59593153FF3D6D57FCF4E897" blockId="2.[178,1086,290,654]" pageId="2" pageNumber="661">
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T y p e m a t e r i a l
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<specimenCount id="9D47FD3459593153FE236D57FD83E867" box="[428,535,290,310]" pageId="2" pageNumber="661" type="generic" typeStatus="syntype">1 syntype</specimenCount>
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(
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<typeStatus id="54FA881F59593153FDAA6D57FDE2E867" box="[549,630,290,310]" pageId="2" pageNumber="661" type="lectotype">lectotype</typeStatus>
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by present designation), relatively well preserved (only segments 6-11 of left antenna and segments 2-11 of right antenna missing), and without attached label under. Close to the specimen, the following handwritten label is pinned on the bottom of box: "
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<emphasis id="B935EAAF59593153FEFA6D1FFE2AE82F" box="[373,446,362,382]" italics="true" pageId="2" pageNumber="661">colchica</emphasis>
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Chaud. Monts Taurus 48. Nordman." (MNHN, "Collection Chaudoir", box no 216 "
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<emphasis id="B935EAAF59593153FED66DF7FE09E8C7" box="[345,413,386,406]" italics="true" pageId="2" pageNumber="661">Argutor</emphasis>
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"). As the original description does not implies that there are one or more
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<typeStatus id="54FA881F59593153FF506DEFFEB9E8FF" box="[223,301,410,430]" pageId="2" pageNumber="661" type="syntype">syntypes</typeStatus>
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, a
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<typeStatus id="54FA881F59593153FEC96DEFFE01E8FF" box="[326,405,410,430]" pageId="2" pageNumber="661" type="lectotype">lectotype</typeStatus>
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by inference of "
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<typeStatus id="54FA881F59593153FDCF6DEDFD0DE8FF" box="[576,665,408,430]" pageId="2" pageNumber="661" type="holotype">holotype</typeStatus>
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" for this species is designated following Article 74.6. of the International Code of Zoological Nomenclature (
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<bibRefCitation id="EFD04B4C59593153FD626DC7FCC2E897" author="ICZN" box="[749,854,434,454]" pageId="2" pageNumber="661" refId="ref4517" refString="ICZN [International Commission on Zoological Nomenclature] (1999): International Code on Zoological Nomenclature. Fourth Edition, adopted by the International Union of Biological Sciences. - London: Internaternational Trust for Zoological Nomenclature, online version: http: // www. iczn. org / iczn." type="url" year="1999">ICZN 1999</bibRefCitation>
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).
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</paragraph>
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<paragraph id="8BFE36BD59593153FF3D6DA7FEFFEBDF" blockId="2.[178,1086,290,654]" pageId="2" pageNumber="661">
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T y p e l o c a l i t y. "
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<collectingRegion id="4985F85F59593153FE256DA7FE79E8B7" box="[426,493,466,486]" country="Georgia" name="Ajaria" pageId="2" pageNumber="661">Adjara</collectingRegion>
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"(
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<bibRefCitation id="EFD04B4C59593153FE716DA7FD2FE8B7" author="CHAUDOIR M" box="[510,699,466,486]" pageId="2" pageNumber="661" pagination="62 - 206" refId="ref4221" refString="CHAUDOIR M. de (1850): Supplement a la faune des carabiques de la Russie. - Bulletin de la Societe Imperiale des Naturalistes de Moscou 23 (3): 62 - 206." type="journal article" year="1850">CHAUDOIR 1850: 78</bibRefCitation>
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). As far as the choice of type locality was based on a presumption of Chaudoir, we have doubt about the real place of finding the
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<typeStatus id="54FA881F59593153FC7E6D9FFBA9E8AF" box="[1009,1085,490,510]" pageId="2" pageNumber="661" type="holotype">holotype</typeStatus>
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and thus about the actual distribution of this species. In the last paragraph of the description,
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<bibRefCitation id="EFD04B4C59593153FF506E6FFE3FEB7F" author="CHAUDOIR M" box="[223,427,538,558]" pageId="2" pageNumber="661" pagination="62 - 206" refId="ref4221" refString="CHAUDOIR M. de (1850): Supplement a la faune des carabiques de la Russie. - Bulletin de la Societe Imperiale des Naturalistes de Moscou 23 (3): 62 - 206." type="journal article" year="1850">CHAUDOIR (1850: 78)</bibRefCitation>
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write: "M. de Normann l’a rapportée de son voyage en Abkhasie, en Mingrélie et dans le Gouriel; je presume qu’il l’a trouvé sur le plateau de l’Adjara, car évidemment un insecte alpin.". The supposition for inaccurate type locality is also supported by the label data of the
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<typeStatus id="54FA881F59593153FE8F6E17FED0EB27" box="[256,324,610,630]" pageId="2" pageNumber="661" type="syntype">syntype</typeStatus>
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(e.g. "Monts Taurus…") and by data of the morphology and distribution of the related taxa; see below.
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</paragraph>
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<paragraph id="8BFE36BD59593153FF3D6EE8FCA8EA33" blockId="2.[178,1086,668,866]" pageId="2" pageNumber="661">
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M a l e g e n i t a l i a. Median lobe of aedeagus in lateral view (
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<figureCitation id="137A2A3859593153FC106EE9FC6EEBE3" box="[927,1018,668,690]" captionStart="Figs 1-5" captionStartId="8.[178,218,911,931]" captionTargetBox="[188,1074,230,887]" captionTargetId="figure-2@8.[178,1085,220,900]" captionTargetPageId="8" captionText="Figs 1-5: Feronia colchica CHAUDOIR, lectotype: Fig. (1) aedeagus, left lateral view; Fig. (2) aedeagus, right lateral view; Fig. (3) apex of aedeagus, dorsal view; Fig. (4) right paramere, internal view; Fig. (5) right paramere, external view. Scale bar 0.5 mm (1-5)." figureDoi="http://doi.org/10.5281/zenodo.5332862" httpUri="https://zenodo.org/record/5332862/files/figure.png" pageId="2" pageNumber="661">Figs 1-2</figureCitation>
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) with ostium deflected to left; apical and basal parts equal in length, apical one with slanting dorsal and straight ventral margin, so that apex became pointed at tip; basal part with strongly developed and prominent ventral process; internal sac in apical position inside, with single strongly chitinised piece (visible in left lateral position). Apex of median lobe dorsally (
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<figureCitation id="137A2A3859593153FE9C6F5AFEC6EA14" box="[275,338,815,837]" captionStart="Figs 1-5" captionStartId="8.[178,218,911,931]" captionTargetBox="[188,1074,230,887]" captionTargetId="figure-2@8.[178,1085,220,900]" captionTargetPageId="8" captionText="Figs 1-5: Feronia colchica CHAUDOIR, lectotype: Fig. (1) aedeagus, left lateral view; Fig. (2) aedeagus, right lateral view; Fig. (3) apex of aedeagus, dorsal view; Fig. (4) right paramere, internal view; Fig. (5) right paramere, external view. Scale bar 0.5 mm (1-5)." figureDoi="http://doi.org/10.5281/zenodo.5332862" httpUri="https://zenodo.org/record/5332862/files/figure.png" pageId="2" pageNumber="661">Fig. 3</figureCitation>
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) with tip deflected to left. Left paramere (apical part broken) (
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<figureCitation id="137A2A3859593153FC516F5AFBA2EA14" box="[990,1078,815,837]" captionStart="Figs 1-5" captionStartId="8.[178,218,911,931]" captionTargetBox="[188,1074,230,887]" captionTargetId="figure-2@8.[178,1085,220,900]" captionTargetPageId="8" captionText="Figs 1-5: Feronia colchica CHAUDOIR, lectotype: Fig. (1) aedeagus, left lateral view; Fig. (2) aedeagus, right lateral view; Fig. (3) apex of aedeagus, dorsal view; Fig. (4) right paramere, internal view; Fig. (5) right paramere, external view. Scale bar 0.5 mm (1-5)." figureDoi="http://doi.org/10.5281/zenodo.5332862" httpUri="https://zenodo.org/record/5332862/files/figure.png" pageId="2" pageNumber="661">Figs 4-5</figureCitation>
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) with apical and basal parts bent each other almost at right angle.
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</paragraph>
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<paragraph id="8BFE36BD59593153FF3D6F04FEFAEAF4" blockId="2.[178,1085,881,933]" pageId="2" pageNumber="661">After drawing, the genitalia were placed in Euparal on the same label on which the specimen is glued.</paragraph>
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<paragraph id="8BFE36BD59593153FF3D6FC0FC4EEDC6" blockId="2.[178,1086,948,1175]" pageId="2" pageNumber="661">
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T a x o n o m i c n o t e s a n d r e l a t i o n s h i p s The presence of scanty pubescence on the distal part of left antennomere 3, dense and continuous punctuation on the basal part of pronotum (which also prolongs forward along the lateral margins and reaches the anterior part of pronotum), absence of medial setiferous punctures on the hind coxae, glabrous ventral side of onychium, and presumable sickle-shaped left paramere demonstrate that this species belongs to
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<taxonomicName id="4C414D3E59593153FD3E6833FCABED0D" authorityName="JACOBSON" authorityYear="1907" box="[689,831,1094,1116]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="666" phylum="Arthropoda" rank="subGenus" subGenus="Oreoplatysma">
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<emphasis id="B935EAAF59593153FD3E6833FCABED0D" box="[689,831,1094,1116]" italics="true" pageId="2" pageNumber="661">Oreoplatysma</emphasis>
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</taxonomicName>
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, and do not to
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<taxonomicName id="4C414D3E59593153FC546833FBA3ED0D" authorityName="CHAUDOIR" authorityYear="1838" box="[987,1079,1094,1116]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="666" phylum="Arthropoda" rank="subGenus" subGenus="Cryobius">
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<emphasis id="B935EAAF59593153FC546833FBA3ED0D" box="[987,1079,1094,1116]" italics="true" pageId="2" pageNumber="661">Cryobius</emphasis>
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</taxonomicName>
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. The following transfer is proposed:
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<taxonomicName id="4C414D3E59593153FDA26811FF7BEDC6" authority="(CHAUDOIR 1850)" baseAuthorityName="CHAUDOIR" baseAuthorityYear="1850" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="661" phylum="Arthropoda" rank="species" species="colchicus" subGenus="Oreoplatysma">
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<emphasis id="B935EAAF59593153FDA26811FD3FED2B" box="[557,683,1124,1146]" italics="true" pageId="2" pageNumber="661">Pterostichus</emphasis>
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(
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<emphasis id="B935EAAF59593153FD316811FCD9ED2B" box="[702,845,1124,1146]" italics="true" pageId="2" pageNumber="661">Oreoplatysma</emphasis>
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)
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<emphasis id="B935EAAF59593153FCD06811FC2AED2B" box="[863,958,1124,1146]" italics="true" pageId="2" pageNumber="661">colchicus</emphasis>
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(
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<bibRefCitation id="EFD04B4C59593153FC406811FF7CEDC6" author="CHAUDOIR M" pageId="2" pageNumber="661" pagination="62 - 206" refId="ref4221" refString="CHAUDOIR M. de (1850): Supplement a la faune des carabiques de la Russie. - Bulletin de la Societe Imperiale des Naturalistes de Moscou 23 (3): 62 - 206." type="journal article" year="1850">CHAUDOIR 1850</bibRefCitation>
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)
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</taxonomicName>
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, new assignment of
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<taxonomicName id="4C414D3E59593153FE3168F4FC41EDC6" authority="(CHAUDOIR 1850)" baseAuthorityName="CHAUDOIR" baseAuthorityYear="1850" box="[446,981,1153,1175]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="661" phylum="Arthropoda" rank="species" species="colchicus" subGenus="Cryobius">
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<emphasis id="B935EAAF59593153FE3168F4FDA8EDC6" box="[446,572,1153,1175]" italics="true" pageId="2" pageNumber="661">Pterostichus</emphasis>
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||
(
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<emphasis id="B935EAAF59593153FDC468F4FD33EDC6" box="[587,679,1153,1175]" italics="true" pageId="2" pageNumber="661">Cryobius</emphasis>
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)
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<emphasis id="B935EAAF59593153FD3968F4FC81EDC6" box="[694,789,1153,1175]" italics="true" pageId="2" pageNumber="661">colchicus</emphasis>
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(
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<bibRefCitation id="EFD04B4C59593153FCAC68F4FC5AEDC6" author="CHAUDOIR M" box="[803,974,1153,1175]" pageId="2" pageNumber="661" pagination="62 - 206" refId="ref4221" refString="CHAUDOIR M. de (1850): Supplement a la faune des carabiques de la Russie. - Bulletin de la Societe Imperiale des Naturalistes de Moscou 23 (3): 62 - 206." type="journal article" year="1850">CHAUDOIR 1850</bibRefCitation>
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)
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</taxonomicName>
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.
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</paragraph>
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<paragraph id="8BFE36BD59593153FF3D68D3FCB4ECD8" blockId="2.[178,1085,1190,1417]" pageId="2" pageNumber="661">
|
||
Details in the external morphology and morphology of both the aedeagus and of the left paramere suppose that this species is allied to several representatives of the group "
|
||
<emphasis id="B935EAAF59593153FF336894FE92EDA6" box="[188,262,1249,1271]" italics="true" pageId="2" pageNumber="661">koenigi</emphasis>
|
||
", like
|
||
<taxonomicName id="4C414D3E59593153FEC26897FD00EDA6" authority="(JAKOBSON 1907)" baseAuthorityName="JAKOBSON" baseAuthorityYear="1907" box="[333,660,1249,1271]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="661" phylum="Arthropoda" rank="species" species="cecchiniae">
|
||
<emphasis id="B935EAAF59593153FEC26897FECFEDA6" box="[333,347,1250,1271]" italics="true" pageId="2" pageNumber="661">P</emphasis>
|
||
.
|
||
<emphasis id="B935EAAF59593153FEE66894FE41EDA6" box="[361,469,1249,1271]" italics="true" pageId="2" pageNumber="661">cecchiniae</emphasis>
|
||
(JAKOBSON 1907)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C414D3E59593153FD2F6897FB99EDA6" authority="KIRSCHENHOFER 1987" authorityName="KIRSCHENHOFER" authorityYear="1987" box="[672,1037,1249,1271]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="661" phylum="Arthropoda" rank="species" species="rousianus">
|
||
<emphasis id="B935EAAF59593153FD2F6897FD3AEDA6" box="[672,686,1250,1271]" italics="true" pageId="2" pageNumber="661">P</emphasis>
|
||
.
|
||
<emphasis id="B935EAAF59593153FD326897FCB5EDA6" box="[701,801,1250,1271]" italics="true" pageId="2" pageNumber="661">rousianus</emphasis>
|
||
<bibRefCitation id="EFD04B4C59593153FCA76897FB99EDA6" author="KIRSCHENHOFER E" box="[808,1037,1249,1271]" pageId="2" pageNumber="661" pagination="45 - 85" refId="ref4479" refString="KIRSCHENHOFER E. (1987): Untersuchungen uber Pterostichus - Arten aus Kleinasien, dem Kaukasus und Zentralasien (Col., Carabidae, Pterostichinae). - Folia Entomologica Hungarica 48: 45 - 85." type="journal article" year="1987">KIRSCHENHOFER 1987</bibRefCitation>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="4C414D3E59593153FF3D688AFE76EC45" authority="REITTER 1898" authorityName="REITTER" authorityYear="1898" box="[178,482,1278,1300]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="661" phylum="Arthropoda" rank="species" species="strasseri">
|
||
<emphasis id="B935EAAF59593153FF3D688AFF55EC45" box="[178,193,1279,1300]" italics="true" pageId="2" pageNumber="661">P</emphasis>
|
||
.
|
||
<emphasis id="B935EAAF59593153FF54688AFEA6EC45" box="[219,306,1279,1300]" italics="true" pageId="2" pageNumber="661">strasseri</emphasis>
|
||
REITTER 1898
|
||
</taxonomicName>
|
||
. But, according to the best of our knowledge, no
|
||
<taxonomicName id="4C414D3E59593153FF3D696EFED5EC60" authorityName="JACOBSON" authorityYear="1907" box="[178,321,1307,1329]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="666" phylum="Arthropoda" rank="subGenus" subGenus="Oreoplatysma">
|
||
<emphasis id="B935EAAF59593153FF3D696EFED5EC60" box="[178,321,1307,1329]" italics="true" pageId="2" pageNumber="661">Oreoplatysma</emphasis>
|
||
</taxonomicName>
|
||
species with pubescent distal part of antennomeres 3 lives in the territories south of the Caucasus Major (except
|
||
<taxonomicName id="4C414D3E59593153FDCF694FFD2BEC1E" baseAuthorityName="CHAUDOIR" baseAuthorityYear="1850" box="[576,703,1337,1359]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="661" phylum="Arthropoda" rank="species" species="colchicus">
|
||
<emphasis id="B935EAAF59593153FDCF694FFDDBEC1E" box="[576,591,1338,1359]" italics="true" pageId="2" pageNumber="661">P</emphasis>
|
||
.
|
||
<emphasis id="B935EAAF59593153FDEF694CFD2BEC1E" box="[608,703,1337,1359]" italics="true" pageId="2" pageNumber="661">colchicus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C414D3E59593153FD78694FFB97EC1E" authority="DVO AK 1995" authorityName="DVO AK" authorityYear="1995" box="[759,1027,1337,1359]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="661" phylum="Arthropoda" rank="species" species="kadleci">
|
||
<emphasis id="B935EAAF59593153FD78694FFC92EC1E" box="[759,774,1338,1359]" italics="true" pageId="2" pageNumber="661">P</emphasis>
|
||
.
|
||
<emphasis id="B935EAAF59593153FC98694CFCF4EC1E" box="[791,864,1337,1359]" italics="true" pageId="2" pageNumber="661">kadleci</emphasis>
|
||
<bibRefCitation id="EFD04B4C59593153FCE5694CFB97EC1E" author="DVO AK M" box="[874,1027,1337,1359]" pageId="2" pageNumber="661" pagination="381 - 392" refId="ref4332" refString="DVO AK M. (1995): Neue Arten den neue Arten der Gattung Pterostichus Subgenus Oreoplatysma aus dem Kaukasus und der Turkei (Coleoptera, Carabidae, Pterostichini). - Entomofauna 16 (18): 381 - 392." type="journal article" year="1995">DVO ÁK 1995</bibRefCitation>
|
||
</taxonomicName>
|
||
), and especially in the area of Adzaria.
|
||
<taxonomicName id="4C414D3E59593153FD9D6922FDE3EC3D" authorityName="DVO AK" authorityYear="1995" box="[530,631,1366,1388]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="661" phylum="Arthropoda" rank="species" species="kadleci">
|
||
<emphasis id="B935EAAF59593153FD9D6922FDB4EC3D" box="[530,544,1367,1388]" italics="true" pageId="2" pageNumber="661">P</emphasis>
|
||
.
|
||
<emphasis id="B935EAAF59593153FDA16923FDE3EC3D" box="[558,631,1366,1388]" italics="true" pageId="2" pageNumber="661">kadleci</emphasis>
|
||
</taxonomicName>
|
||
from the Kop Da i (NE
|
||
<collectingCountry id="F356762D59593153FCF06923FC5EEC3D" box="[895,970,1366,1388]" name="Turkey" pageId="2" pageNumber="661">Turkey</collectingCountry>
|
||
) is distinct from
|
||
<taxonomicName id="4C414D3E59593153FF666901FEF1ECD8" baseAuthorityName="CHAUDOIR" baseAuthorityYear="1850" box="[233,357,1395,1417]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="661" phylum="Arthropoda" rank="species" species="colchicus">
|
||
<emphasis id="B935EAAF59593153FF666901FF6CECD8" box="[233,248,1396,1417]" italics="true" pageId="2" pageNumber="661">P</emphasis>
|
||
.
|
||
<emphasis id="B935EAAF59593153FE896906FEF1ECD8" box="[262,357,1395,1417]" italics="true" pageId="2" pageNumber="661">colchicus</emphasis>
|
||
</taxonomicName>
|
||
in the very thin and long apex of aedeagus.
|
||
</paragraph>
|
||
<paragraph id="8BFE36BD59593153FF3D69ECFECBECB8" blockId="2.[178,1085,1432,1513]" pageId="2" pageNumber="661">
|
||
<taxonomicName id="4C414D3E59593153FF3D69ECFEA0ECFF" baseAuthorityName="CHAUDOIR" baseAuthorityYear="1850" box="[178,308,1432,1454]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="661" phylum="Arthropoda" rank="species" species="colchicus">
|
||
<emphasis id="B935EAAF59593153FF3D69ECFF55ECFF" box="[178,193,1433,1454]" italics="true" pageId="2" pageNumber="661">P</emphasis>
|
||
.
|
||
<emphasis id="B935EAAF59593153FF5A69EDFEA0ECFF" box="[213,308,1432,1454]" italics="true" pageId="2" pageNumber="661">colchicus</emphasis>
|
||
</taxonomicName>
|
||
should be valid species, but further investigation looking for eventual synonymy between it and subsequently described species and for the true area of distribution is needed.
|
||
</paragraph>
|
||
<paragraph id="8BFE36BD59593153FF3D698CFCFCEF5F" blockId="2.[178,872,1528,1550]" box="[178,872,1528,1550]" pageId="2" pageNumber="661">
|
||
D i s t r i b u t i o n.?
|
||
<collectingRegion id="4985F85F59593153FE14698DFE6AEF5F" box="[411,510,1528,1550]" country="Georgia" name="Ajaria" pageId="2" pageNumber="661">Adzaria</collectingRegion>
|
||
(KRYZHANOVSKIJ et al. 1995: 102).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |