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<document id="AB17CCC45A44E0FA8E07F1484178A368" ID-DOI="10.5252/adansonia2023v45a16" ID-ISSN="1639-4798" ID-Zenodo-Dep="8109269" IM.bibliography_approvedBy="juliana" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="juliana" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="juliana" IM.treatments_approvedBy="juliana" checkinTime="1688391199371" checkinUser="felipe" docAuthor="Calvo, Joel &amp; Callmander, Martin W." docDate="2023" docId="03E8878B4D37FF9AFCC214B11A008DA4" docLanguage="en" docName="Adansonia.45.16.279-283.pdf" docOrigin="Adansonia (3) 45 (16)" docStyle="DocumentStyle:2FE19AC740987067EC2A6C89A85F9648.8:Adansonia.2018-.journal_article" docStyleId="2FE19AC740987067EC2A6C89A85F9648" docStyleName="Adansonia.2018-.journal_article" docStyleVersion="8" docTitle="Senecio marinae J. Calvo &amp; Callm. 2023, sp. nov." docType="treatment" docVersion="3" lastPageNumber="283" masterDocId="FFD1FFF34D34FF9CFFBD15261B63896A" masterDocTitle="Novelties from the Northern Mountains Complex of Madagascar VII: A new species of Senecio L. (Compositae)" masterLastPageNumber="283" masterPageNumber="279" pageNumber="280" updateTime="1688405467860" updateUser="juliana" zenodo-license-document="CC-BY-4.0">
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<mods:title id="89C25557C71A6EB4942A2DAD58BCEAE9">Novelties from the Northern Mountains Complex of Madagascar VII: A new species of Senecio L. (Compositae)</mods:title>
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<mods:namePart id="1E878CC4FB1B50FF7FA6A06D642E565A">Calvo, Joel</mods:namePart>
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<mods:namePart id="C3E84A647894AF1BD1AD35DE6B5F893A">Callmander, Martin W.</mods:namePart>
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<mods:titleInfo id="91941CE4E72E8D602A120B26BE686267">
<mods:title id="3FB276B904D33A086A315BAFC7E56376">Adansonia</mods:title>
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<mods:part id="A19480B809B933431B5E5A46257461B6">
<mods:date id="B6CCBB4D82884930B4341C55D6A2357C">2023</mods:date>
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<mods:title id="D6158B247B565407A4F21F761FD4C977">3</mods:title>
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<mods:number id="E7BEF8532C4B79EF8E176FCED362DAEB">2023-07-03</mods:number>
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<mods:number id="7F37C079CDF3DE780EB8745FFE8B2E24">45</mods:number>
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<mods:identifier id="0543F9C6D1C9F2B3E334E40198241828" type="DOI">10.5252/adansonia2023v45a16</mods:identifier>
<mods:identifier id="02DC9CDA4A9DC81A59EC836847F762A8" type="ISSN">1639-4798</mods:identifier>
<mods:identifier id="D5C17F8A41ADD4CAC4E680AC3754DF55" type="Zenodo-Dep">8109269</mods:identifier>
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<treatment id="03E8878B4D37FF9AFCC214B11A008DA4" ID-DOI="http://doi.org/10.5281/zenodo.8109271" ID-Zenodo-Dep="8109271" LSID="urn:lsid:plazi:treatment:03E8878B4D37FF9AFCC214B11A008DA4" httpUri="http://treatment.plazi.org/id/03E8878B4D37FF9AFCC214B11A008DA4" lastPageId="6" lastPageNumber="283" pageId="3" pageNumber="280">
<subSubSection id="C35B65164D37FF9FFCC214B11E3D88D8" box="[895,1374,407,434]" pageId="3" pageNumber="280" type="nomenclature">
<paragraph id="8BFE369D4D37FF9FFCC214B11E3D88D8" blockId="3.[895,1374,407,465]" box="[895,1374,407,434]" pageId="3" pageNumber="280">
<heading id="D0B681F14D37FF9FFCC214B11E3D88D8" box="[895,1374,407,434]" centered="true" fontSize="11" level="2" pageId="3" pageNumber="280" reason="7">
<taxonomicName id="4C414D1E4D37FF9FFCC214B11E6388DB" authority="J. Calvo &amp; Callm." authorityName="J. Calvo &amp; Callm." authorityYear="2023" box="[895,1280,407,434]" class="Magnoliopsida" family="Asteraceae" genus="Senecio" kingdom="Plantae" order="Asterales" pageId="3" pageNumber="280" phylum="Tracheophyta" rank="species" species="marinae" status="sp. nov.">
<emphasis id="B935EA8F4D37FF9FFCC214B11F5288DB" bold="true" box="[895,1073,407,433]" italics="true" pageId="3" pageNumber="280">Senecio marinae</emphasis>
J. Calvo &amp; Callm.
</taxonomicName>
,
<taxonomicNameLabel id="A20657F44D37FF9FFAB614BE1E3D88D8" box="[1291,1374,408,434]" pageId="3" pageNumber="280" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C35B65164D37FF9FFB8A14911FC788BB" box="[1079,1188,439,465]" pageId="3" pageNumber="280" type="description">
<paragraph id="8BFE369D4D37FF9FFB8A14911FC788BB" blockId="3.[895,1374,407,465]" box="[1079,1188,439,465]" pageId="3" pageNumber="280">
(
<figureCitation id="137A2A184D37FF9FFB8314911FE388BB" box="[1086,1152,439,465]" captionStart="FIG" captionStartId="4.[133,144,1593,1611]" captionTargetBox="[132,1455,215,1582]" captionTargetId="figure-188@4.[132,1455,215,1582]" captionTargetPageId="4" captionText="FIG. 1. — Senecio marinae J. Calvo &amp; Callm., sp. nov.: A, flowering stem; B, capitulum at an early stage; C, capitulum at an advanced stage;D, achene with pap- pus; E, floret (ovary and pappus removed); F, anther (notice the caudate base). A, C, D, Antilahimena et al. 653; B, E, F, Wohlhauser et al. 785. Drawing: Roger Lala Andriamiarisoa. Scale bars: A, 1 cm; B, C, 6 mm; D, 2 mm, E, 1.5 mm, F, 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.8109273" httpUri="https://zenodo.org/record/8109273/files/figure.png" pageId="3" pageNumber="280">Figs 1</figureCitation>
;
<figureCitation id="137A2A184D37FF9FFB3314911FFE88BB" box="[1166,1181,439,465]" captionStart="FIG" captionStartId="5.[132,143,1985,2003]" captionTargetBox="[132,1456,216,1932]" captionTargetId="figure-59@5.[132,1454,989,1979]" captionTargetPageId="5" captionText="FIG. 2. — Field pictures of Senecio marinae J. Calvo &amp; Callm., sp. nov.: A, habit; B, synflorescence; C, detail of the capitulum; A-C, Wohlhauser et al. 785. Photographs: S. Wohlhauser." figureDoi="http://doi.org/10.5281/zenodo.8109275" httpUri="https://zenodo.org/record/8109275/files/figure.png" pageId="3" pageNumber="280">2</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C35B65164D37FF9FFC9014D21F688BC1" pageId="3" pageNumber="280" type="diagnosis">
<paragraph id="8BFE369D4D37FF9FFC9014D21F688BC1" blockId="3.[811,1458,500,683]" pageId="3" pageNumber="280">
<taxonomicName id="4C414D1E4D37FF9FFC9014D218B38B66" authority="Calvo &amp; Callmander, 2023" authorityName="Calvo &amp; Callmander" authorityYear="2023" box="[813,976,500,524]" class="Magnoliopsida" family="Asteraceae" genus="Senecio" kingdom="Plantae" order="Asterales" pageId="3" pageNumber="280" phylum="Tracheophyta" rank="species" species="marinae" status="sp. nov.">Senecio marinae</taxonomicName>
<emphasis id="B935EA8F4D37FF9FFC6514D318AD8BFA" italics="true" pageId="3" pageNumber="280">
<taxonomicNameLabel id="A20657F44D37FF9FFC6514D31F428B66" box="[984,1057,501,524]" pageId="3" pageNumber="280" rank="species">sp. nov.</taxonomicNameLabel>
can be distinguished by its scrambling or ascending habit with stems
<quantity id="4CB99B784D37FF9FFB9117291F068B4C" box="[1068,1125,527,551]" metricMagnitude="0" metricUnit="m" metricValue="2.5" metricValueMax="4.0" metricValueMin="1.0" pageId="3" pageNumber="280" unit="m" value="2.5" valueMax="4.0" valueMin="1.0">1-4 m</quantity>
long, the oblanceolate to obtrullate, fleshy, glabrous leaves, usually somewhat tricuspidate apically, the synflorescences with subumbelliform divisions, the discoid capitula with 10-11 involucral bracts, and the glabrous achenes. It mostly differs from the similar species
</emphasis>
<taxonomicName id="4C414D1E4D37FF9FFC6E175E1FC28BFA" authority="Humbert" authorityName="Humbert" box="[979,1185,632,656]" class="Magnoliopsida" family="Asteraceae" genus="Senecio" kingdom="Plantae" order="Asterales" pageId="3" pageNumber="280" phylum="Tracheophyta" rank="species" species="francoisii">
S. francoisii
<emphasis id="B935EA8F4D37FF9FFBF6175E1FC28BFA" box="[1099,1185,632,656]" italics="true" pageId="3" pageNumber="280">Humbert</emphasis>
</taxonomicName>
<emphasis id="B935EA8F4D37FF9FFB1A175E1FA88BFA" box="[1191,1227,632,656]" italics="true" pageId="3" pageNumber="280">and</emphasis>
S. handiensis
<emphasis id="B935EA8F4D37FF9FFAE8175E1F688BC1" italics="true" pageId="3" pageNumber="280">Forssk. in having discoid capitula.</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C35B65164D37FF9FFC9017E11E4E8A23" pageId="3" pageNumber="280" type="materials_examined">
<paragraph id="8BFE369D4D37FF9FFC9017E11E4E8A23" blockId="3.[811,1456,711,841]" pageId="3" pageNumber="280">
<materialsCitation id="3B293CC04D37FF9FFC9017E11E1F8A44" collectingDate="2005-10-20" collectionCode="G" collectorName="Wohlhauser &amp; Callmander &amp; Buerki" country="Madagascar" county="Boeny Region" elevation="1980" latitude="-14.403333" location="Mangindrano" longLatPrecision="21" longitude="49.065556" municipality="Mahajanga" pageId="3" pageNumber="280" specimenCode="G00398286" specimenCount="1" stateProvince="Boeny Region" typeStatus="holotype">
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. —
<emphasis id="B935EA8F4D37FF9FFC3817E1189A8BB5" bold="true" box="[901,1017,711,735]" pageId="3" pageNumber="280">
<collectingCountry id="F356760D4D37FF9FFC3817E1189A8BB5" box="[901,1017,711,735]" name="Madagascar" pageId="3" pageNumber="280">Madagascar</collectingCountry>
</emphasis>
.
<collectingRegion id="4985F87F4D37FF9FFBB917E11FE98BB5" box="[1028,1162,711,735]" pageId="3" pageNumber="280">Boeny Region</collectingRegion>
[Prov.
<collectingMunicipality id="6B9AACE74D37FF9FFB7617E11E588BB5" box="[1227,1339,711,735]" pageId="3" pageNumber="280">Mahajanga</collectingMunicipality>
]:
<collectingMunicipality id="6B9AACE74D37FF9FFAF517E11ECF8BB5" box="[1352,1452,711,735]" pageId="3" pageNumber="280">Bealanana</collectingMunicipality>
,
<location id="8E9E60464D37FF9FFC9617C718D38B93" LSID="urn:lsid:plazi:treatment:03E8878B4D37FF9AFCC214B11A008DA4:8E9E60464D37FF9FFC9617C718D38B93" box="[811,944,737,761]" country="Madagascar" county="Boeny Region" latitude="-14.403333" longLatPrecision="21" longitude="49.065556" municipality="Mahajanga" name="Mangindrano" pageId="3" pageNumber="280" stateProvince="Boeny Region">Mangindrano</location>
,
<location id="8E9E60464D37FF9FFC0A17C71F128B93" LSID="urn:lsid:plazi:treatment:03E8878B4D37FF9AFCC214B11A008DA4:8E9E60464D37FF9FFC0A17C71F128B93" box="[951,1137,737,761]" country="Madagascar" county="Boeny Region" latitude="-14.403333" longLatPrecision="21" longitude="49.065556" municipality="Mahajanga" name="Ambohimirahavavy" pageId="3" pageNumber="280" stateProvince="Boeny Region">Ambohimirahavavy</location>
,
<location id="8E9E60464D37FF9FFBC417C71FB98B93" LSID="urn:lsid:plazi:treatment:03E8878B4D37FF9AFCC214B11A008DA4:8E9E60464D37FF9FFBC417C71FB98B93" box="[1145,1242,737,761]" country="Madagascar" county="Boeny Region" latitude="-14.403333" longLatPrecision="21" longitude="49.065556" municipality="Mahajanga" name="Antsahivo" pageId="3" pageNumber="280" stateProvince="Boeny Region">Antsahivo</location>
,
<locationDeviation id="9A4C2FBF4D37FF9FFB5F17C41ECA8B93" box="[1250,1449,737,761]" pageId="3" pageNumber="280">W du campement 01</locationDeviation>
,
<location id="8E9E60464D37FF9FFC9617DD18F78A79" LSID="urn:lsid:plazi:treatment:03E8878B4D37FF9AFCC214B11A008DA4:8E9E60464D37FF9FFC9617DD18F78A79" box="[811,916,763,787]" country="Madagascar" county="Boeny Region" latitude="-14.403333" longLatPrecision="21" longitude="49.065556" municipality="Mahajanga" name="Matsabory" pageId="3" pageNumber="280" stateProvince="Boeny Region">Matsabory</location>
,
<geoCoordinate id="EE75505A4D37FF9FFC1C17DA1F778A7E" box="[929,1044,764,788]" degrees="14" direction="south" minutes="24" orientation="latitude" pageId="3" pageNumber="280" precision="15" seconds="12" value="-14.403333">14°24’12”S</geoCoordinate>
,
<geoCoordinate id="EE75505A4D37FF9FFB9D17DD1FF08A79" box="[1056,1171,763,787]" degrees="49" direction="east" minutes="03" orientation="longitude" pageId="3" pageNumber="280" precision="15" seconds="56" value="49.065556">49°03’56”E</geoCoordinate>
,
<quantity id="4CB99B784D37FF9FFB1C17DA1F918A79" box="[1185,1266,764,788]" metricMagnitude="3" metricUnit="m" metricValue="1.98" pageId="3" pageNumber="280" unit="m" value="1980.0">
<elevation id="006CD1AE4D37FF9FFB1C17DA1F918A79" box="[1185,1266,764,788]" metricMagnitude="3" metricUnit="m" metricValue="1.98" pageId="3" pageNumber="280" unit="m" value="1980.0">1980 m</elevation>
</quantity>
,
<date id="FFFF105D4D37FF9FFB4317DD1E098A79" box="[1278,1386,763,787]" pageId="3" pageNumber="280" value="2005-10-20">
<collectingDate id="EFBBE9B54D37FF9FFB4317DD1E098A79" box="[1278,1386,763,787]" pageId="3" pageNumber="280" value="2005-10-20">20.X.2005</collectingDate>
</date>
,
<emphasis id="B935EA8F4D37FF9FFACB17DD1F1E8A44" italics="true" pageId="3" pageNumber="280">
<collectorName id="26B4534B4D37FF9FFACB17DD180F8A44" pageId="3" pageNumber="280">Wohlhauser</collectorName>
,
<collectorName id="26B4534B4D37FF9FFCCA1630188A8A44" box="[887,1001,790,814]" pageId="3" pageNumber="280">Callmander</collectorName>
&amp;
<collectorName id="26B4534B4D37FF9FFBB316301F2D8A44" box="[1038,1102,790,814]" pageId="3" pageNumber="280">Buerki</collectorName>
785
</emphasis>
(
<typeStatus id="54FA883F4D37FF9FFB3116301FA68A44" box="[1164,1221,790,814]" pageId="3" pageNumber="280">holo-</typeStatus>
,
<collectionCode id="ED50AE584D37FF9FFB6C16301F858A44" box="[1233,1254,790,814]" country="Switzerland" lsid="urn:lsid:biocol.org:col:15706" name="Conservatoire et Jardin botaniques de la Ville de Genève" pageId="3" pageNumber="280" type="Herbarium">G</collectionCode>
[
<specimenCode id="DBE79EE64D37FF9FFB5016301E0B8A44" box="[1261,1384,790,814]" collectionCode="G" country="Switzerland" lsid="urn:lsid:biocol.org:col:15706" name="Conservatoire et Jardin botaniques de la Ville de Genève" pageId="3" pageNumber="280" type="Herbarium">G00398286</specimenCode>
]!;
</materialsCitation>
<materialsCitation id="3B293CC04D37FF9FFA3916301E4A8A23" collectingDate="2005-10-20" collectionCode="MO, P, TAN" collectorName="Wohlhauser &amp; Callmander &amp; Buerki" country="Madagascar" county="Boeny Region" elevation="1980" latitude="-14.403333" location="Mangindrano" longLatPrecision="21" longitude="49.065556" municipality="Mahajanga" pageId="3" pageNumber="280" specimenCode="MO-3401237, P02473236" specimenCount="2" stateProvince="Boeny Region" typeStatus="isotype">
<typeStatus id="54FA883F4D37FF9FFA3916301EC98A44" box="[1412,1450,790,814]" pageId="3" pageNumber="280">iso-</typeStatus>
,
<collectionCode id="ED50AE584D37FF9FFC931617183A8A23" box="[814,857,817,841]" country="USA" lsid="urn:lsid:biocol.org:col:15859" name="Missouri Botanical Garden" pageId="3" pageNumber="280" type="Herbarium">MO</collectionCode>
[
<specimenCode id="DBE79EE64D37FF9FFCE2161718938A23" box="[863,1008,817,841]" pageId="3" pageNumber="280">MO-3401237</specimenCode>
] image!;
<collectionCode id="ED50AE584D37FF9FFBF516171F3B8A22" box="[1096,1112,817,840]" country="France" lsid="urn:lsid:biocol.org:col:15763" name="Museum National d' Histoire Naturelle, Paris (MNHN) - Vascular Plants" pageId="3" pageNumber="280" type="Herbarium">P</collectionCode>
[
<specimenCode id="DBE79EE64D37FF9FFBE316161FB18A22" box="[1118,1234,816,840]" collectionCode="P" country="France" lsid="urn:lsid:biocol.org:col:15763" name="Museum National d' Histoire Naturelle, Paris (MNHN) - Vascular Plants" pageId="3" pageNumber="280" type="Herbarium">P02473236</specimenCode>
]!;
<collectionCode id="ED50AE584D37FF9FFB5616171E418A23" box="[1259,1314,817,841]" country="Madagascar" httpUri="http://biocol.org/urn:lsid:biocol.org:col:13842" lsid="urn:lsid:biocol.org:col:13842" name="Parc de Tsimbazaza" pageId="3" pageNumber="280" type="Herbarium">TAN</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C35B65164D37FF9FFC9016431F7F8AFD" pageId="3" pageNumber="280" type="biology_ecology">
<paragraph id="8BFE369D4D37FF9FFC9016431F7F8AFD" blockId="3.[813,1454,868,919]" pageId="3" pageNumber="280">
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. — The new species has been found in bloom in October and November.
</paragraph>
</subSubSection>
<subSubSection id="C35B65164D37FF9FFC9016951E198D24" pageId="3" pageNumber="280" type="distribution">
<paragraph id="8BFE369D4D37FF9FFC9016951E198D24" blockId="3.[811,1456,946,1103]" pageId="3" pageNumber="280">
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<smallCapsWord id="8D18A0414D37FF9FFC56169E1F278AA3" baselines="965" box="[1003,1092,952,969]" lowerCaseFontSize="7" mainFontSize="10" normCase="lower" normString="ecology" pageId="3" pageNumber="280">ECOLOGY</smallCapsWord>
. —
<taxonomicName id="4C414D1E4D37FF9FFBD016951E618AA0" authority="Calvo &amp; Callmander, 2023" authorityName="Calvo &amp; Callmander" authorityYear="2023" box="[1133,1282,947,970]" class="Magnoliopsida" family="Asteraceae" genus="Senecio" kingdom="Plantae" order="Asterales" pageId="3" pageNumber="280" phylum="Tracheophyta" rank="species" species="marinae" status="sp. nov.">
<emphasis id="B935EA8F4D37FF9FFBD016951E618AA0" box="[1133,1282,947,970]" italics="true" pageId="3" pageNumber="280">Senecio marinae</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A20657F44D37FF9FFAB016951E358AA0" box="[1293,1366,947,970]" pageId="3" pageNumber="280" rank="species">sp. nov.</taxonomicNameLabel>
is known at elevations of
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in Ambohimiravavy, Biempoko, Sorata, and Tsaratanana massifs. The new species grows on plateaux and ridges in open forests (chablis) rich in Bamboo in medium altitude moist evergreen forest and at the transition between these forests and the ericoid mountain thicket (
<emphasis id="B935EA8F4D37FF9FFBC311111FCE8D24" box="[1150,1197,1079,1102]" italics="true" pageId="3" pageNumber="280">sensu</emphasis>
<bibRefCitation id="EFD04B6C4D37FF9FFB0911111E0E8D25" author="GAUTIER L. &amp; TAHINARIVONY J. A. &amp; RANIRISON P. &amp; WOHLHAUSER S." box="[1204,1389,1079,1103]" pageId="3" pageNumber="280" pagination="207 - 242" refId="ref2698" refString="GAUTIER L., TAHINARIVONY J. A., RANIRISON P. &amp; WOHLHAUSER S. 2018. - Vegetation, in GOODMAN S. M., RAHERILALAO J. &amp; WOHLHAUSER S. (eds), The Terrestrial Protected Areas of Madagascar: Their History, Description, and Biota. Association Vahatra, Antananarivo: 207 - 242." type="book chapter" year="2018">
Gautier
<emphasis id="B935EA8F4D37FF9FFAB811111E578D25" box="[1285,1332,1079,1103]" italics="true" pageId="3" pageNumber="280">et al.</emphasis>
2018
</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C35B65164D37FF9FFC90114D1EC88C6D" pageId="3" pageNumber="280" type="etymology">
<paragraph id="8BFE369D4D37FF9FFC90114D1EC88C6D" blockId="3.[812,1457,1130,1287]" pageId="3" pageNumber="280">
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. — The epithet
<taxonomicName id="4C414D1E4D37FF9FFBFB114D1FF78DE8" authorityName="Calvo &amp; Callmander" authorityYear="2023" box="[1094,1172,1131,1154]" class="Magnoliopsida" family="Asteraceae" genus="Senecio" kingdom="Plantae" order="Asterales" pageId="3" pageNumber="280" phylum="Tracheophyta" rank="species" species="marinae">
<emphasis id="B935EA8F4D37FF9FFBFB114D1FF78DE8" box="[1094,1172,1131,1154]" italics="true" pageId="3" pageNumber="280">marinae</emphasis>
</taxonomicName>
honors
<taxonomicName id="4C414D1E4D37FF9FFB59114C18358DF7" authority="Rabarimanarivo" authorityName="Rabarimanarivo" class="Magnoliopsida" family="Fabaceae" genus="Marina" higherTaxonomySource="GBIF" kingdom="Plantae" order="Fabales" pageId="3" pageNumber="280" phylum="Tracheophyta" rank="genus">Marina Rabarimanarivo</taxonomicName>
, botanist at the Missouri Botanical Garden in
<collectingRegion id="4985F87F4D37FF9FFA9711A31ECC8DF7" box="[1322,1455,1157,1181]" country="Madagascar" name="Antananarivo" pageId="3" pageNumber="280">Antananarivo</collectingRegion>
who contributes immensely to the
<collectingCountry id="F356760D4D37FF9FFB3511B91F988DDD" box="[1160,1275,1183,1207]" name="Madagascar" pageId="3" pageNumber="280">Madagascar</collectingCountry>
Catalogue project (
<bibRefCitation id="EFD04B6C4D37FF9FFC88119C1F298DB8" author="MADAGASCAR CATALOGUE" box="[821,1098,1210,1234]" pageId="3" pageNumber="280" refId="ref2983" refString="MADAGASCAR CATALOGUE 2023. - Catalogue of the Plants of Madagascar. Missouri Botanical Garden, St. Louis and Antananarivo. [http: // www. tropicos. org / Project / Madagascar]" type="url" year="2023">
<collectingCountry id="F356760D4D37FF9FFC88119C18CB8DB8" box="[821,936,1210,1234]" name="Madagascar" pageId="3" pageNumber="280">Madagascar</collectingCountry>
Catalogue 2023
</bibRefCitation>
) and understood that the collections from high altitudes in Northern
<collectingCountry id="F356760D4D37FF9FFBDF11F21FB78D86" box="[1122,1236,1236,1260]" name="Madagascar" pageId="3" pageNumber="280">Madagascar</collectingCountry>
represented a new species when she evaluated the genus
<taxonomicName id="4C414D1E4D37FF9FFBC111C91FA38C6C" box="[1148,1216,1263,1286]" class="Magnoliopsida" family="Asteraceae" genus="Senecio" kingdom="Plantae" order="Asterales" pageId="3" pageNumber="280" phylum="Tracheophyta" rank="genus">
<emphasis id="B935EA8F4D37FF9FFBC111C91FA38C6C" box="[1148,1216,1263,1286]" italics="true" pageId="3" pageNumber="280">Senecio</emphasis>
</taxonomicName>
for the project in 2009.
</paragraph>
</subSubSection>
<subSubSection id="C35B65164D37FF9FFC9010051E098F43" pageId="3" pageNumber="280" type="conservation">
<paragraph id="8BFE369D4D37FF9FFC9010051E098F43" blockId="3.[811,1457,1315,1577]" pageId="3" pageNumber="280">
<smallCapsWord id="8D18A0414D37FF9FFC90100518A58C53" baselines="1333,1333" box="[813,966,1315,1339]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Conservation" pageId="3" pageNumber="280">CONSERVATION</smallCapsWord>
<smallCapsWord id="8D18A0414D37FF9FFC72100E1F768C53" baselines="1333" box="[975,1045,1320,1337]" lowerCaseFontSize="7" mainFontSize="10" normCase="lower" normString="status" pageId="3" pageNumber="280">STATUS</smallCapsWord>
. —
<taxonomicName id="4C414D1E4D37FF9FFBF910051FBD8C50" authority="Calvo &amp; Callmander, 2023" authorityName="Calvo &amp; Callmander" authorityYear="2023" box="[1092,1246,1315,1338]" class="Magnoliopsida" family="Asteraceae" genus="Senecio" kingdom="Plantae" order="Asterales" pageId="3" pageNumber="280" phylum="Tracheophyta" rank="species" species="marinae" status="sp. nov.">
<emphasis id="B935EA8F4D37FF9FFBF910051FBD8C50" box="[1092,1246,1315,1338]" italics="true" pageId="3" pageNumber="280">Senecio marinae</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A20657F44D37FF9FFB5110051E5A8C50" box="[1260,1337,1315,1338]" pageId="3" pageNumber="280" rank="species">sp. nov.</taxonomicNameLabel>
has an estimated EOO of
<emphasis id="B935EA8F4D37FF9FFC7E101818B08C3F" box="[963,979,1342,1365]" italics="true" pageId="3" pageNumber="280">c.</emphasis>
<quantity id="4CB99B784D37FF9FFC6A101B1F518C32" box="[983,1074,1341,1368]" metricMagnitude="6" metricUnit="m" metricValue="1.319" pageId="3" pageNumber="280" unit="km" value="1319.0">1319 km</quantity>
², an AOO of
<quantity id="4CB99B784D37FF9FFB08101B1F958C32" box="[1205,1270,1341,1368]" metricMagnitude="4" metricUnit="m" metricValue="2.4" pageId="3" pageNumber="280" unit="km" value="24.0">24 km</quantity>
², and five locations encompassed within the COMATSA Nord and Tsaratanana protected areas. Fires are known to occasionally enter pristine forests in those protected areas and are also known along pastures and summit zones (
<bibRefCitation id="EFD04B6C4D37FF9FFCCD10811F298CD5" author="GOODMAN S. M. &amp; RAHERILALAO J. &amp; WOHLHAUSER S." box="[880,1098,1447,1471]" pageId="3" pageNumber="280" pagination="597 - 611" refId="ref2762" refString="GOODMAN S. M., RAHERILALAO J. &amp; WOHLHAUSER S. 2018. - Site 11. COMATSA Nord, in GOODMAN S. M., RAHERILALAO J. &amp; WOHLHAUSER S. (eds), The Terrestrial Protected Areas of Madagascar: Their History, Description, and Biota. Association Vahatra, Antananarivo: 597 - 611." type="book chapter" year="2018">
Goodman
<emphasis id="B935EA8F4D37FF9FFC63108E1F6C8CD5" box="[990,1039,1447,1471]" italics="true" pageId="3" pageNumber="280">et al.</emphasis>
2018
</bibRefCitation>
). Due to the plausible threats to its habitat for slash-and-burn agriculture and fires caused by lightning,
<taxonomicName id="4C414D1E4D37FF9FFC9610FA18F18C99" authority="Calvo &amp; Callmander, 2023" authorityName="Calvo &amp; Callmander" authorityYear="2023" box="[811,914,1500,1523]" class="Magnoliopsida" family="Asteraceae" genus="Senecio" kingdom="Plantae" order="Asterales" pageId="3" pageNumber="280" phylum="Tracheophyta" rank="species" species="marinae" status="sp. nov.">
<emphasis id="B935EA8F4D37FF9FFC9610FA18F18C99" box="[811,914,1500,1523]" italics="true" pageId="3" pageNumber="280">S. marinae</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A20657F44D37FF9FFC1D10FA188F8C99" box="[928,1004,1500,1523]" pageId="3" pageNumber="280" rank="species">sp. nov.</taxonomicNameLabel>
is assigned to a preliminary risk of extinction status of “Vulnerable” [VU B1ab(i,ii,iii,iv,v)+2ab(i,ii,iii,iv,v)] using the IUCN Red List Categories and Criteria (
<bibRefCitation id="EFD04B6C4D37FF9FFB5F13371E3E8F43" author="IUCN" box="[1250,1373,1553,1577]" pageId="3" pageNumber="280" refId="ref2897" refString="IUCN 2012. - IUCN Red List Categories and Criteria: Version 3.1. Ed. 2. IUCN Species Survival Commission, IUCN, Gland and Cambridge." type="book" year="2012">IUCN 2012</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C35B65164D37FF9FFC9013631EC68E83" pageId="3" pageNumber="280" type="materials_examined">
<paragraph id="8BFE369D4D37FF9FFC9013631EC68E83" blockId="3.[811,1456,1604,2025]" pageId="3" pageNumber="280">
<smallCapsWord id="8D18A0414D37FF9FFC90136318CA8F31" baselines="1623,1623" box="[813,937,1605,1629]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Additional" pageId="3" pageNumber="280">ADDITIONAL</smallCapsWord>
<smallCapsWord id="8D18A0414D37FF9FFC10136C1F768F31" baselines="1623" box="[941,1045,1610,1627]" lowerCaseFontSize="7" mainFontSize="10" normCase="lower" normString="specimens" pageId="3" pageNumber="280">SPECIMENS</smallCapsWord>
<smallCapsWord id="8D18A0414D37FF9FFBA7136C1F1D8F31" baselines="1623" box="[1050,1150,1610,1627]" lowerCaseFontSize="7" mainFontSize="10" normCase="lower" normString="examined" pageId="3" pageNumber="280">EXAMINED</smallCapsWord>
. —
<materialsCitation id="3B293CC04D37FF9FFB15136318D48FAC" collectingDate="2000-11-05" collectingDateMax="2007-10-25" collectingDateMin="1966-11" collectionCode="G, K, MO, P, TAN" collectorName="Antilahimena" country="Madagascar" county="Diana Region" elevation="2288" latitude="-14.228055" location="Ambanja" longLatPrecision="21" longitude="49.137222" municipality="Antsiranana" pageId="3" pageNumber="280" specimenCode="MO798854, P02473228" specimenCount="1" stateProvince="Diana Region">
<emphasis id="B935EA8F4D37FF9FFB1513631E478F37" bold="true" box="[1192,1316,1605,1629]" pageId="3" pageNumber="280">
<collectingCountry id="F356760D4D37FF9FFB1513631E438F37" box="[1192,1312,1605,1629]" name="Madagascar" pageId="3" pageNumber="280">Madagascar</collectingCountry>
.
</emphasis>
<collectingRegion id="4985F87F4D37FF9FFA9513621ECC8F36" box="[1320,1455,1604,1628]" pageId="3" pageNumber="280">Diana Region</collectingRegion>
[Prov.
<collectingMunicipality id="6B9AACE74D37FF9FFCD01379188F8F1D" box="[877,1004,1631,1655]" pageId="3" pageNumber="280">Antsiranana</collectingMunicipality>
]:
<collectingMunicipality id="6B9AACE74D37FF9FFC4013791F168F1D" box="[1021,1141,1631,1655]" pageId="3" pageNumber="280">Marotolana</collectingMunicipality>
,
<location id="8E9E60464D37FF9FFB3C13791FBC8F1D" LSID="urn:lsid:plazi:treatment:03E8878B4D37FF9AFCC214B11A008DA4:8E9E60464D37FF9FFB3C13791FBC8F1D" box="[1153,1247,1631,1655]" country="Madagascar" county="Diana Region" latitude="-14.228055" longLatPrecision="21" longitude="49.137222" municipality="Antsiranana" name="Ambanja" pageId="3" pageNumber="280" stateProvince="Diana Region">Ambanja</location>
,
<location id="8E9E60464D37FF9FFB5413791E028F1C" LSID="urn:lsid:plazi:treatment:03E8878B4D37FF9AFCC214B11A008DA4:8E9E60464D37FF9FFB5413791E028F1C" box="[1257,1377,1631,1654]" country="Madagascar" county="Diana Region" latitude="-14.228055" longLatPrecision="21" longitude="49.137222" municipality="Antsiranana" name="Tsaratanana" pageId="3" pageNumber="280" stateProvince="Diana Region">Tsaratanana</location>
massif, S slopes,
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,
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,
<quantity id="4CB99B784D37FF9FFBCE135F1F9B8FFB" box="[1139,1272,1657,1681]" metricMagnitude="3" metricUnit="m" metricValue="2.288" metricValueMax="2.3649999999999998" metricValueMin="2.211" pageId="3" pageNumber="280" unit="m" value="2288.0" valueMax="2365.0" valueMin="2211.0">
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</quantity>
,
<date id="FFFF105D4D37FF9FFABD135F1E058FFB" box="[1280,1382,1657,1681]" pageId="3" pageNumber="280" value="2000-11-05">
<collectingDate id="EFBBE9B54D37FF9FFABD135F1E058FFB" box="[1280,1382,1657,1681]" pageId="3" pageNumber="280" value="2000-11-05">5.XI.2000</collectingDate>
</date>
,
<emphasis id="B935EA8F4D37FF9FFAD3135F18AF8FC6" italics="true" pageId="3" pageNumber="280">
<collectorName id="26B4534B4D37FF9FFAD3135F18118FC6" pageId="3" pageNumber="280">Antilahimena</collectorName>
et al. 653
</emphasis>
(
<collectionCode id="ED50AE584D37FF9FFC6513B2188D8FC6" box="[984,1006,1684,1708]" country="Switzerland" lsid="urn:lsid:biocol.org:col:15706" name="Conservatoire et Jardin botaniques de la Ville de Genève" pageId="3" pageNumber="280" type="Herbarium">G</collectionCode>
!,
<collectionCode id="ED50AE584D37FF9FFC4613B21F6D8FC1" box="[1019,1038,1684,1707]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15867" name="Royal Botanic Gardens" pageId="3" pageNumber="280" type="Herbarium">K</collectionCode>
,
<collectionCode id="ED50AE584D37FF9FFBAB13B21F238FC6" box="[1046,1088,1684,1708]" country="USA" lsid="urn:lsid:biocol.org:col:15859" name="Missouri Botanical Garden" pageId="3" pageNumber="280" type="Herbarium">MO</collectionCode>
[
<specimenCode id="DBE79EE64D37FF9FFBFB13B21FA18FC6" box="[1094,1218,1684,1708]" collectionCode="MO" country="USA" lsid="urn:lsid:biocol.org:col:15859" name="Missouri Botanical Garden" pageId="3" pageNumber="280" type="Herbarium">MO798854</specimenCode>
] image!,
<collectionCode id="ED50AE584D37FF9FFAA813B21E478FC1" box="[1301,1316,1684,1707]" country="France" lsid="urn:lsid:biocol.org:col:15763" name="Museum National d' Histoire Naturelle, Paris (MNHN) - Vascular Plants" pageId="3" pageNumber="280" type="Herbarium">P</collectionCode>
[
<specimenCode id="DBE79EE64D37FF9FFA9713B21EFF8FC6" box="[1322,1436,1684,1708]" collectionCode="P" country="France" lsid="urn:lsid:biocol.org:col:15763" name="Museum National d' Histoire Naturelle, Paris (MNHN) - Vascular Plants" pageId="3" pageNumber="280" type="Herbarium">P02473228</specimenCode>
]!,
<collectionCode id="ED50AE584D37FF9FFC90138818028FAC" box="[813,865,1710,1734]" country="Madagascar" httpUri="http://biocol.org/urn:lsid:biocol.org:col:13842" lsid="urn:lsid:biocol.org:col:13842" name="Parc de Tsimbazaza" pageId="3" pageNumber="280" type="Herbarium">TAN</collectionCode>
image!);
</materialsCitation>
<materialsCitation id="3B293CC04D37FF9FFC0113891ECC8FAC" box="[956,1455,1710,1734]" collectingDate="1966-11" collectingDateMax="2007-10-25" collectingDateMin="1966-11" collectionCode="TAN, MO, P" collectorName="Morat &amp; Randriambololomamonjy" country="Madagascar" county="Diana Region" elevation="1559" latitude="-14.228055" location="Antsahavalany" longLatPrecision="21" longitude="49.137222" municipality="Marotolana" pageId="3" pageNumber="280" specimenCode="MO3401222, P04276661" specimenCount="1" stateProvince="Tsaratanana">
<collectingRegion id="4985F87F4D37FF9FFC0113891F558FAC" box="[956,1078,1711,1734]" pageId="3" pageNumber="280">Tsaratanana</collectingRegion>
,
<date id="FFFF105D4D37FF9FFBFD13881FF68FAC" box="[1088,1173,1710,1734]" pageId="3" pageNumber="280" value="1966-11">
<collectingDate id="EFBBE9B54D37FF9FFBFD13881FF68FAC" box="[1088,1173,1710,1734]" pageId="3" pageNumber="280" value="1966-11">XI.1966</collectingDate>
</date>
,
<emphasis id="B935EA8F4D37FF9FFB1D13891E758FAC" box="[1184,1302,1710,1734]" italics="true" pageId="3" pageNumber="280">
<collectorName id="26B4534B4D37FF9FFB1D13891FB88FAC" box="[1184,1243,1711,1734]" pageId="3" pageNumber="280">Morat</collectorName>
2256
</emphasis>
(
<collectionCode id="ED50AE584D37FF9FFA9E13881E3A8FAC" box="[1315,1369,1710,1734]" country="Madagascar" httpUri="http://biocol.org/urn:lsid:biocol.org:col:13842" lsid="urn:lsid:biocol.org:col:13842" name="Parc de Tsimbazaza" pageId="3" pageNumber="280" type="Herbarium">TAN</collectionCode>
image!);
</materialsCitation>
<materialsCitation id="3B293CC04D37FF9FFC9013EF18138E5A" collectingDate="1966-11" collectingDateMax="2007-10-25" collectingDateMin="1966-11" collectionCode="TAN, MO, P" collectorName="Morat &amp; Randriambololomamonjy" country="Madagascar" county="Diana Region" elevation="1559" latitude="-14.228055" location="Antsahavalany" longLatPrecision="21" longitude="49.137222" municipality="Marotolana" pageId="3" pageNumber="280" specimenCode="MO3401222, P04276661" specimenCount="1" stateProvince="Tsaratanana">
<collectingRegion id="4985F87F4D37FF9FFC9013EF18C48F8B" box="[813,935,1737,1761]" pageId="3" pageNumber="280">Manambato</collectingRegion>
, à
<quantity id="4CB99B784D37FF9FFC0313EF18988F8B" box="[958,1019,1737,1761]" metricMagnitude="4" metricUnit="m" metricValue="1.0" pageId="3" pageNumber="280" unit="km" value="10.0">
<locationDeviation id="9A4C2FBF4D37FF9FFC0313EF18988F8B" box="[958,1019,1737,1761]" metricMagnitude="4" metricUnit="m" metricValue="1.0" pageId="3" pageNumber="280" unit="km" value="10.0">10 km</locationDeviation>
</quantity>
à vol d’oiseau du fkt.
<location id="8E9E60464D37FF9FFB7513EF1E368F8B" LSID="urn:lsid:plazi:treatment:03E8878B4D37FF9AFCC214B11A008DA4:8E9E60464D37FF9FFB7513EF1E368F8B" box="[1224,1365,1737,1761]" country="Madagascar" county="Diana Region" latitude="-14.228055" longLatPrecision="21" longitude="49.137222" municipality="Marotolana" name="Antsahavalany" pageId="3" pageNumber="280" stateProvince="Tsaratanana">Antsahavalany</location>
, [Sorata],
<geoCoordinate id="EE75505A4D37FF9FFC9613C518C28F91" box="[811,929,1763,1787]" degrees="13" direction="south" minutes="43" orientation="latitude" pageId="3" pageNumber="280" precision="15" seconds="28" value="-13.724444">13°43’28”S</geoCoordinate>
,
<geoCoordinate id="EE75505A4D37FF9FFC1013C51F408F91" box="[941,1059,1763,1787]" degrees="49" direction="east" minutes="22" orientation="longitude" pageId="3" pageNumber="280" precision="15" seconds="43" value="49.37861">49°22’43”E</geoCoordinate>
,
<quantity id="4CB99B784D37FF9FFB8C13C51FE78F91" box="[1073,1156,1763,1787]" metricMagnitude="3" metricUnit="m" metricValue="1.559" pageId="3" pageNumber="280" unit="m" value="1559.0">
<elevation id="006CD1AE4D37FF9FFB8C13C51FE78F91" box="[1073,1156,1763,1787]" metricMagnitude="3" metricUnit="m" metricValue="1.559" pageId="3" pageNumber="280" unit="m" value="1559.0">1559 m</elevation>
</quantity>
,
<date id="FFFF105D4D37FF9FFB2D13C51F9D8F91" box="[1168,1278,1763,1787]" pageId="3" pageNumber="280" value="2007-10-25">
<collectingDate id="EFBBE9B54D37FF9FFB2D13C51F9D8F91" box="[1168,1278,1763,1787]" pageId="3" pageNumber="280" value="2007-10-25">25.X.2007</collectingDate>
</date>
,
<emphasis id="B935EA8F4D37FF9FFAB713C5188A8E7C" italics="true" pageId="3" pageNumber="280">
<collectorName id="26B4534B4D37FF9FFAB713C518E58E7F" pageId="3" pageNumber="280">Randriambololomamonjy</collectorName>
et al. 103
</emphasis>
(
<collectionCode id="ED50AE584D37FF9FFC4B13D81F418E7C" box="[1014,1058,1790,1814]" country="USA" lsid="urn:lsid:biocol.org:col:15859" name="Missouri Botanical Garden" pageId="3" pageNumber="280" type="Herbarium">MO</collectionCode>
[
<specimenCode id="DBE79EE64D37FF9FFB9513D81FD68E7C" box="[1064,1205,1790,1814]" collectionCode="MO" country="USA" lsid="urn:lsid:biocol.org:col:15859" name="Missouri Botanical Garden" pageId="3" pageNumber="280" type="Herbarium">MO3401222</specimenCode>
] image!,
<collectionCode id="ED50AE584D37FF9FFAB213D81E7C8E7F" box="[1295,1311,1790,1813]" country="France" lsid="urn:lsid:biocol.org:col:15763" name="Museum National d' Histoire Naturelle, Paris (MNHN) - Vascular Plants" pageId="3" pageNumber="280" type="Herbarium">P</collectionCode>
[
<specimenCode id="DBE79EE64D37FF9FFA9813D81EF88E7C" box="[1317,1435,1790,1814]" collectionCode="P" country="France" lsid="urn:lsid:biocol.org:col:15763" name="Museum National d' Histoire Naturelle, Paris (MNHN) - Vascular Plants" pageId="3" pageNumber="280" type="Herbarium">P04276661</specimenCode>
]!,
<collectionCode id="ED50AE584D37FF9FFC90123E18068E5A" box="[813,869,1816,1840]" country="Madagascar" httpUri="http://biocol.org/urn:lsid:biocol.org:col:13842" lsid="urn:lsid:biocol.org:col:13842" name="Parc de Tsimbazaza" pageId="3" pageNumber="280" type="Herbarium">TAN</collectionCode>
).
</materialsCitation>
—
<materialsCitation id="3B293CC04D37FF9FFC29123E18098EEA" collectingDate="2005-11-17" collectingDateMax="2007-10-25" collectingDateMin="1966-11" collectionCode="MO, P, TAN" collectorName="Randrianarivelo" country="Madagascar" county="Diana Region" elevation="1991" latitude="-14.228055" location="l'E de Bealanana" longLatPrecision="21" longitude="49.137222" municipality="Antisranana" pageId="3" pageNumber="280" specimenCode="P02473238" specimenCount="1" stateProvince="Sava Region">
<collectingRegion id="4985F87F4D37FF9FFC29123E1F6E8E5A" box="[916,1037,1816,1840]" pageId="3" pageNumber="280">Sava Region</collectingRegion>
[Prov.
<collectingMunicipality id="6B9AACE74D37FF9FFBEF123E1FB38E5A" box="[1106,1232,1816,1840]" pageId="3" pageNumber="280">Antisranana</collectingMunicipality>
]:
<quantity id="4CB99B784D37FF9FFB63123E1E7C8E5A" box="[1246,1311,1816,1840]" metricMagnitude="4" metricUnit="m" metricValue="2.2" pageId="3" pageNumber="280" unit="km" value="22.0">
<locationDeviation id="9A4C2FBF4D37FF9FFB63123E1E7C8E5A" box="[1246,1311,1816,1840]" metricMagnitude="4" metricUnit="m" metricValue="2.2" pageId="3" pageNumber="280" unit="km" value="22.0">22 km</locationDeviation>
</quantity>
à
<location id="8E9E60464D37FF9FFA89123E183C8E20" LSID="urn:lsid:plazi:treatment:03E8878B4D37FF9AFCC214B11A008DA4:8E9E60464D37FF9FFA89123E183C8E20" country="Madagascar" county="Diana Region" latitude="-14.228055" longLatPrecision="21" longitude="49.137222" municipality="Antisranana" name="l'E de Bealanana" pageId="3" pageNumber="280" stateProvince="Sava Region">l’E de Bealanana</location>
, montagne de
<location id="8E9E60464D37FF9FFC5612151F3B8E21" LSID="urn:lsid:plazi:treatment:03E8878B4D37FF9AFCC214B11A008DA4:8E9E60464D37FF9FFC5612151F3B8E21" box="[1003,1112,1843,1867]" country="Madagascar" county="Diana Region" latitude="-14.228055" longLatPrecision="21" longitude="49.137222" municipality="Antisranana" name="Beampoko" pageId="3" pageNumber="280" stateProvince="Sava Region">Beampoko</location>
,
<geoCoordinate id="EE75505A4D37FF9FFBDC12151FB78E21" box="[1121,1236,1843,1867]" degrees="14" direction="south" minutes="13" orientation="latitude" pageId="3" pageNumber="280" precision="15" seconds="41" value="-14.228055">14°13’41”S</geoCoordinate>
,
<geoCoordinate id="EE75505A4D37FF9FFB6012151E328E21" box="[1245,1361,1843,1867]" degrees="49" direction="east" minutes="08" orientation="longitude" pageId="3" pageNumber="280" precision="15" seconds="14" value="49.137222">49°08’14”E</geoCoordinate>
,
<quantity id="4CB99B784D37FF9FFAE112151EC88E20" box="[1372,1451,1843,1867]" metricMagnitude="3" metricUnit="m" metricValue="1.991" pageId="3" pageNumber="280" unit="m" value="1991.0">
<elevation id="006CD1AE4D37FF9FFAE112151EC88E20" box="[1372,1451,1843,1867]" metricMagnitude="3" metricUnit="m" metricValue="1.991" pageId="3" pageNumber="280" unit="m" value="1991.0">1991 m</elevation>
</quantity>
,
<date id="FFFF105D4D37FF9FFC96126B18C78E0F" box="[811,932,1869,1893]" pageId="3" pageNumber="280" value="2005-11-17">
<collectingDate id="EFBBE9B54D37FF9FFC96126B18C78E0F" box="[811,932,1869,1893]" pageId="3" pageNumber="280" value="2005-11-17">17.XI.2005</collectingDate>
</date>
,
<emphasis id="B935EA8F4D37FF9FFC0E126B1FDE8E0F" box="[947,1213,1869,1893]" italics="true" pageId="3" pageNumber="280">
<collectorName id="26B4534B4D37FF9FFC0E126B1F308E0F" box="[947,1107,1869,1893]" pageId="3" pageNumber="280">Randrianarivelo</collectorName>
et al. 329
</emphasis>
(
<collectionCode id="ED50AE584D37FF9FFB73126B1F9D8E0F" box="[1230,1278,1869,1893]" country="USA" lsid="urn:lsid:biocol.org:col:15859" name="Missouri Botanical Garden" pageId="3" pageNumber="280" type="Herbarium">MO</collectionCode>
,
<collectionCode id="ED50AE584D37FF9FFAB612681E788E0F" box="[1291,1307,1870,1893]" country="France" lsid="urn:lsid:biocol.org:col:15763" name="Museum National d' Histoire Naturelle, Paris (MNHN) - Vascular Plants" pageId="3" pageNumber="280" type="Herbarium">P</collectionCode>
[
<specimenCode id="DBE79EE64D37FF9FFA9F126B1EF98E0F" box="[1314,1434,1869,1893]" collectionCode="P" country="France" lsid="urn:lsid:biocol.org:col:15763" name="Museum National d' Histoire Naturelle, Paris (MNHN) - Vascular Plants" pageId="3" pageNumber="280" type="Herbarium">P02473238</specimenCode>
]!,
<collectionCode id="ED50AE584D37FF9FFC90124E18008EEA" box="[813,867,1896,1920]" country="Madagascar" httpUri="http://biocol.org/urn:lsid:biocol.org:col:13842" lsid="urn:lsid:biocol.org:col:13842" name="Parc de Tsimbazaza" pageId="3" pageNumber="280" type="Herbarium">TAN</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B293CC04D37FF9FFCCF12411F5E8EDF" collectingDate="2005-11-09" collectionCode="G, MO, P, TAN" collectorName="Randrianarivony" country="Madagascar" elevation="2220" latitude="-14.201944" location="Beampoko" longLatPrecision="21" longitude="49.118053" pageId="3" pageNumber="280" specimenCode="P02473237" specimenCount="1" stateProvince="Antsiranana">
plateau entre
<location id="8E9E60464D37FF9FFC4C12411F3A8E15" LSID="urn:lsid:plazi:treatment:03E8878B4D37FF9AFCC214B11A008DA4:8E9E60464D37FF9FFC4C12411F3A8E15" box="[1009,1113,1895,1919]" country="Madagascar" latitude="-14.201944" longLatPrecision="21" longitude="49.118053" name="Beampoko" pageId="3" pageNumber="280" stateProvince="Antsiranana">Beampoko</location>
et
<location id="8E9E60464D37FF9FFBCE12411E528E15" LSID="urn:lsid:plazi:treatment:03E8878B4D37FF9AFCC214B11A008DA4:8E9E60464D37FF9FFBCE12411E528E15" box="[1139,1329,1895,1919]" country="Madagascar" latitude="-14.201944" longLatPrecision="21" longitude="49.118053" name="Ambohimirahavavy" pageId="3" pageNumber="280" stateProvince="Antsiranana">Ambohimirahavavy</location>
,
<geoCoordinate id="EE75505A4D37FF9FFA84124E1EC98EEA" box="[1337,1450,1896,1920]" degrees="14" direction="south" minutes="12" orientation="latitude" pageId="3" pageNumber="280" precision="15" seconds="07" value="-14.201944">14°12’07”S</geoCoordinate>
,
<geoCoordinate id="EE75505A4D37FF9FFC9012A418C18EF0" box="[813,930,1922,1946]" degrees="49" direction="east" minutes="07" orientation="longitude" pageId="3" pageNumber="280" precision="15" seconds="05" value="49.118053">49°07’05”E</geoCoordinate>
,
<quantity id="4CB99B784D37FF9FFC0D12A41F618EF3" box="[944,1026,1922,1946]" metricMagnitude="3" metricUnit="m" metricValue="2.2199999999999998" pageId="3" pageNumber="280" unit="m" value="2220.0">
<elevation id="006CD1AE4D37FF9FFC0D12A41F618EF3" box="[944,1026,1922,1946]" metricMagnitude="3" metricUnit="m" metricValue="2.2199999999999998" pageId="3" pageNumber="280" unit="m" value="2220.0">2220 m</elevation>
</quantity>
,
<date id="FFFF105D4D37FF9FFBB312A41F148EF0" box="[1038,1143,1922,1946]" pageId="3" pageNumber="280" value="2005-11-09">
<collectingDate id="EFBBE9B54D37FF9FFBB312A41F148EF0" box="[1038,1143,1922,1946]" pageId="3" pageNumber="280" value="2005-11-09">9.XI.2005</collectingDate>
</date>
,
<emphasis id="B935EA8F4D37FF9FFB3E12A41EE38EF0" box="[1155,1408,1922,1946]" italics="true" pageId="3" pageNumber="280">
<collectorName id="26B4534B4D37FF9FFB3E12A41E458EF0" box="[1155,1318,1922,1946]" pageId="3" pageNumber="280">Randrianarivony</collectorName>
et al. 74
</emphasis>
(
<collectionCode id="ED50AE584D37FF9FFA2D12A41EC48EF0" box="[1424,1447,1922,1946]" country="Switzerland" lsid="urn:lsid:biocol.org:col:15706" name="Conservatoire et Jardin botaniques de la Ville de Genève" pageId="3" pageNumber="280" type="Herbarium">G</collectionCode>
!,
<collectionCode id="ED50AE584D37FF9FFC9312BB18388EDF" box="[814,859,1949,1973]" country="USA" lsid="urn:lsid:biocol.org:col:15859" name="Missouri Botanical Garden" pageId="3" pageNumber="280" type="Herbarium">MO</collectionCode>
,
<collectionCode id="ED50AE584D37FF9FFCDE12BB18118EDE" box="[867,882,1949,1972]" country="France" lsid="urn:lsid:biocol.org:col:15763" name="Museum National d' Histoire Naturelle, Paris (MNHN) - Vascular Plants" pageId="3" pageNumber="280" type="Herbarium">P</collectionCode>
[
<specimenCode id="DBE79EE64D37FF9FFCC412BB188A8EDF" box="[889,1001,1949,1973]" collectionCode="P" country="France" lsid="urn:lsid:biocol.org:col:15763" name="Museum National d' Histoire Naturelle, Paris (MNHN) - Vascular Plants" pageId="3" pageNumber="280" type="Herbarium">P02473237</specimenCode>
]!,
<collectionCode id="ED50AE584D37FF9FFBBD12BB1F558EDF" box="[1024,1078,1949,1973]" country="Madagascar" httpUri="http://biocol.org/urn:lsid:biocol.org:col:13842" lsid="urn:lsid:biocol.org:col:13842" name="Parc de Tsimbazaza" pageId="3" pageNumber="280" type="Herbarium">TAN</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B293CC04D37FF9FFBF812BA1EC28E83" collectingDate="2007-10-01" collectionCode="MO, P, TAN" collectorName="Razakamalala" country="Madagascar" elevation="1364" latitude="-13.6825" location="Sorata" longLatPrecision="21" longitude="49.4425" municipality="Andrafainkona" pageId="3" pageNumber="280" specimenCode="MO3185014, P04276663" specimenCount="1" stateProvince="Antsiranana">
<collectingMunicipality id="6B9AACE74D37FF9FFBF812BA1FB48EDE" box="[1093,1239,1948,1972]" pageId="3" pageNumber="280">Andrafainkona</collectingMunicipality>
, forêt dense humide de
<location id="8E9E60464D37FF9FFC96129118088EA5" LSID="urn:lsid:plazi:treatment:03E8878B4D37FF9AFCC214B11A008DA4:8E9E60464D37FF9FFC96129118088EA5" box="[811,875,1975,1999]" country="Madagascar" latitude="-13.6825" longLatPrecision="21" longitude="49.4425" municipality="Andrafainkona" name="Sorata" pageId="3" pageNumber="280" stateProvince="Antsiranana">Sorata</location>
,
<date id="FFFF105D4D37FF9FFCCF129118B28EA5" box="[882,977,1975,1999]" pageId="3" pageNumber="280" value="2007-10-01">
<collectingDate id="EFBBE9B54D37FF9FFCCF129118B28EA5" box="[882,977,1975,1999]" pageId="3" pageNumber="280" value="2007-10-01">1.X.2007</collectingDate>
</date>
,
<geoCoordinate id="EE75505A4D37FF9FFC6412911F288EA5" box="[985,1099,1975,1999]" degrees="13" direction="south" minutes="40" orientation="latitude" pageId="3" pageNumber="280" precision="15" seconds="57" value="-13.6825">13°40’57”S</geoCoordinate>
,
<geoCoordinate id="EE75505A4D37FF9FFBE912911FA58EA5" box="[1108,1222,1975,1999]" degrees="49" direction="east" minutes="26" orientation="longitude" pageId="3" pageNumber="280" precision="15" seconds="33" value="49.4425">49°26’33”E</geoCoordinate>
,
<quantity id="4CB99B784D37FF9FFB6C12911E7D8EA4" box="[1233,1310,1975,1999]" metricMagnitude="3" metricUnit="m" metricValue="1.364" pageId="3" pageNumber="280" unit="m" value="1364.0">
<elevation id="006CD1AE4D37FF9FFB6C12911E7D8EA4" box="[1233,1310,1975,1999]" metricMagnitude="3" metricUnit="m" metricValue="1.364" pageId="3" pageNumber="280" unit="m" value="1364.0">1364 m</elevation>
</quantity>
,
<emphasis id="B935EA8F4D37FF9FFA9A129118F58E83" italics="true" pageId="3" pageNumber="280">
<collectorName id="26B4534B4D37FF9FFA9A12911ECD8EA5" box="[1319,1454,1975,1999]" pageId="3" pageNumber="280">Razakamalala</collectorName>
et al. 3634
</emphasis>
(
<collectionCode id="ED50AE584D37FF9FFC1912F718B08E83" box="[932,979,2001,2025]" country="USA" lsid="urn:lsid:biocol.org:col:15859" name="Missouri Botanical Garden" pageId="3" pageNumber="280" type="Herbarium">MO</collectionCode>
[
<specimenCode id="DBE79EE64D37FF9FFC6412F71F038E83" box="[985,1120,2001,2025]" collectionCode="MO" country="USA" lsid="urn:lsid:biocol.org:col:15859" name="Missouri Botanical Garden" pageId="3" pageNumber="280" type="Herbarium">MO3185014</specimenCode>
] image!,
<collectionCode id="ED50AE584D37FF9FFB0112F41FAF8E83" box="[1212,1228,2002,2025]" country="France" lsid="urn:lsid:biocol.org:col:15763" name="Museum National d' Histoire Naturelle, Paris (MNHN) - Vascular Plants" pageId="3" pageNumber="280" type="Herbarium">P</collectionCode>
[
<specimenCode id="DBE79EE64D37FF9FFB6F12F71E2B8E83" box="[1234,1352,2001,2025]" collectionCode="P" country="France" lsid="urn:lsid:biocol.org:col:15763" name="Museum National d' Histoire Naturelle, Paris (MNHN) - Vascular Plants" pageId="3" pageNumber="280" type="Herbarium">P04276663</specimenCode>
]!,
<collectionCode id="ED50AE584D37FF9FFADF12F71EF98E83" box="[1378,1434,2001,2025]" country="Madagascar" httpUri="http://biocol.org/urn:lsid:biocol.org:col:13842" lsid="urn:lsid:biocol.org:col:13842" name="Parc de Tsimbazaza" pageId="3" pageNumber="280" type="Herbarium">TAN</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<caption id="DF3E66154D30FF98FF38131F18328F13" ID-DOI="http://doi.org/10.5281/zenodo.8109273" ID-Zenodo-Dep="8109273" httpUri="https://zenodo.org/record/8109273/files/figure.png" pageId="4" pageNumber="281" startId="4.[133,144,1593,1611]" targetBox="[132,1455,215,1582]" targetPageId="4" targetType="figure">
<paragraph id="8BFE369D4D30FF98FF38131F18328F13" blockId="4.[133,1455,1593,1657]" pageId="4" pageNumber="281">
<smallCapsWord id="8D18A0414D30FF98FF38131F1BFE8F20" baselines="1606,1607" box="[133,157,1593,1611]" lowerCaseFontSize="5" mainFontSize="7" normCase="title" normString="Fig" pageId="4" pageNumber="281">FIG</smallCapsWord>
. 1. —
<taxonomicName id="4C414D1E4D30FF98FF6E131F1A978F21" authority="J. Calvo &amp; Callm." authorityName="J. Calvo &amp; Callm." authorityYear="2023" box="[211,500,1593,1611]" class="Magnoliopsida" family="Asteraceae" genus="Senecio" kingdom="Plantae" order="Asterales" pageId="4" pageNumber="281" phylum="Tracheophyta" rank="species" species="marinae" status="sp. nov.">
<emphasis id="B935EA8F4D30FF98FF6E131F1A3D8F21" box="[211,350,1593,1611]" italics="true" pageId="4" pageNumber="281">Senecio marinae</emphasis>
J. Calvo &amp; Callm.
</taxonomicName>
,
<taxonomicNameLabel id="A20657F44D30FF98FE44131F195D8F21" box="[505,574,1593,1611]" pageId="4" pageNumber="281" rank="species">sp. nov.</taxonomicNameLabel>
:
<emphasis id="B935EA8F4D30FF98FDFE131F19338F21" bold="true" box="[579,592,1593,1611]" pageId="4" pageNumber="281">A</emphasis>
, flowering stem;
<emphasis id="B935EA8F4D30FF98FD63131F19888F21" bold="true" box="[734,747,1593,1611]" pageId="4" pageNumber="281">B</emphasis>
, capitulum at an early stage;
<emphasis id="B935EA8F4D30FF98FC60131F18888F21" bold="true" box="[989,1003,1593,1611]" pageId="4" pageNumber="281">C</emphasis>
, capitulum at an advanced stage;
<emphasis id="B935EA8F4D30FF98FABA131F1E768F21" bold="true" box="[1287,1301,1593,1611]" pageId="4" pageNumber="281">D</emphasis>
, achene with pappus;
<emphasis id="B935EA8F4D30FF98FF1313761BD98F08" bold="true" box="[174,186,1616,1634]" pageId="4" pageNumber="281">E</emphasis>
, floret (ovary and pappus removed);
<emphasis id="B935EA8F4D30FF98FE5313761A9A8F08" bold="true" box="[494,505,1616,1634]" pageId="4" pageNumber="281">F</emphasis>
, anther (notice the caudate base).
<emphasis id="B935EA8F4D30FF98FCA61376184B8F08" bold="true" box="[795,808,1616,1634]" pageId="4" pageNumber="281">A</emphasis>
,
<emphasis id="B935EA8F4D30FF98FC8E137618228F08" bold="true" box="[819,833,1616,1634]" pageId="4" pageNumber="281">C</emphasis>
,
<emphasis id="B935EA8F4D30FF98FCF61376183A8F08" bold="true" box="[843,857,1616,1634]" pageId="4" pageNumber="281">D</emphasis>
,
<emphasis id="B935EA8F4D30FF98FCDE13761F428F08" box="[867,1057,1616,1634]" italics="true" pageId="4" pageNumber="281">Antilahimena et al. 653</emphasis>
;
<emphasis id="B935EA8F4D30FF98FB9613761F5B8F08" bold="true" box="[1067,1080,1616,1634]" pageId="4" pageNumber="281">B</emphasis>
,
<emphasis id="B935EA8F4D30FF98FBFE13761F2C8F08" bold="true" box="[1091,1103,1616,1634]" pageId="4" pageNumber="281">E</emphasis>
,
<emphasis id="B935EA8F4D30FF98FBE713761F068F08" bold="true" box="[1114,1125,1616,1634]" pageId="4" pageNumber="281">F</emphasis>
,
<emphasis id="B935EA8F4D30FF98FBD213761E408F08" box="[1135,1315,1616,1634]" italics="true" pageId="4" pageNumber="281">Wohlhauser et al. 785</emphasis>
. Drawing: Roger Lala Andriamiarisoa. Scale bars: A, 1 cm; B, C, 6 mm; D, 2 mm, E, 1.5 mm, F, 0.5 mm.
</paragraph>
</caption>
<subSubSection id="C35B65164D30FF9AFF3913951964883B" lastPageId="6" lastPageNumber="283" pageId="4" pageNumber="281" type="description">
<paragraph id="8BFE369D4D30FF98FF3913951A768FA1" blockId="4.[130,777,1715,2027]" box="[132,277,1715,1741]" pageId="4" pageNumber="281">
<smallCapsWord id="8D18A0414D30FF98FF3913951A768FA1" baselines="1734,1734" box="[132,277,1715,1741]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Description" pageId="4" pageNumber="281">DESCRIPTION</smallCapsWord>
</paragraph>
<paragraph id="8BFE369D4D30FF9AFF3913F41964883B" blockId="4.[130,777,1715,2027]" lastBlockId="6.[130,775,216,338]" lastPageId="6" lastPageNumber="283" pageId="4" pageNumber="281">
Suffrutescent subshrub, scrambling or ascending,
<quantity id="4CB99B784D30FF98FD2A13F519BB8F87" box="[663,728,1747,1773]" metricMagnitude="0" metricUnit="m" metricValue="2.5" metricValueMax="4.0" metricValueMin="1.0" pageId="4" pageNumber="281" unit="m" value="2.5" valueMax="4.0" valueMin="1.0">1-4 m</quantity>
tall. Stem terete, smooth, fistulous (at least upper parts), glabrous, with leaves only on upper part as plant ages. Leaves alternate, simple, pseudopetiolate; laminas oblanceolate to obtrullate, usually somewhat tricuspidate apically, 6-13 ×
<quantity id="4CB99B784D30FF98FD38127418618E07" box="[645,770,1874,1901]" metricMagnitude="-2" metricUnit="m" metricValue="3.9" metricValueMax="5.5" metricValueMin="2.3" pageId="4" pageNumber="281" unit="cm" value="3.9" valueMax="5.5" valueMin="2.3">2.3-5.5 cm</quantity>
, base attenuate, apex acute, margins entire, glabrous on both faces, with venation barely noticeable, fleshy, shiny in living plants; pseudopetioles
<quantity id="4CB99B784D30FF98FED712971AAB8EA6" box="[362,456,1969,1996]" metricMagnitude="-2" metricUnit="m" metricValue="1.75" metricValueMax="2.5" metricValueMin="1.0" pageId="4" pageNumber="281" unit="cm" value="1.75" valueMax="2.5" valueMin="1.0">1-2.5 cm</quantity>
long, glabrous.Synflorescences corymbiform in overall shape, with divisions subumbelliform; synflorescence bracts linear-subulate. Capitula discoid on peduncles
<quantity id="4CB99B784D30FF98FC7C13F41F448F87" box="[961,1063,1746,1773]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" metricValueMax="10.0" metricValueMin="6.0" pageId="4" pageNumber="281" unit="mm" value="8.0" valueMax="10.0" valueMin="6.0">6-10 mm</quantity>
long with 1-3 bracteoles. Involucres cylindrical, 8-9 ×
<quantity id="4CB99B784D30FF98FC5513D41F268E67" box="[1000,1093,1778,1805]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" metricValueMax="5.0" metricValueMin="4.0" pageId="4" pageNumber="281" unit="mm" value="4.5" valueMax="5.0" valueMin="4.0">4-5 mm</quantity>
, glabrous; receptables flat, somewhate fimbrillate; involucral bracts 10-11, linear-oblong, 5.8-7.4×
<quantity id="4CB99B784D30FF98FC2F12141F668E27" box="[914,1029,1842,1869]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" metricValueMax="10.0" metricValueMin="6.0" pageId="4" pageNumber="281" unit="mm" value="0.8" valueMax="1.0" valueMin="0.6">0.6-1 mm</quantity>
, glabrous; supplementary bracts (calyculus) 3-5, linear-subulate, 1.5-2.3 ×
<emphasis id="B935EA8F4D30FF98FB7512751FBA8E06" box="[1224,1241,1875,1900]" italics="true" pageId="4" pageNumber="281">c.</emphasis>
<quantity id="4CB99B784D30FF98FB5C12741E5E8E07" box="[1249,1341,1874,1901]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="4" pageNumber="281" unit="mm" value="0.3">0.3 mm</quantity>
, glabrous. Florets
<emphasis id="B935EA8F4D30FF98FCC0125518ED8EE6" box="[893,910,1907,1932]" italics="true" pageId="4" pageNumber="281">c.</emphasis>
22, hermaphrodite; corollas
<quantity id="4CB99B784D30FF98FB7A12541E298EE6" box="[1223,1354,1906,1933]" metricMagnitude="-3" metricUnit="m" metricValue="5.95" metricValueMax="6.2" metricValueMin="5.7" pageId="4" pageNumber="281" unit="mm" value="5.95" valueMax="6.2" valueMin="5.7">5.7-6.2 mm</quantity>
long, tubular, limbs
<emphasis id="B935EA8F4D30FF98FC0D12B518A28EC6" box="[944,961,1939,1964]" italics="true" pageId="4" pageNumber="281">c.</emphasis>
<quantity id="4CB99B784D30FF98FC7B12B41F798EC6" box="[966,1050,1938,1964]" metricMagnitude="-3" metricUnit="m" metricValue="2.7" pageId="4" pageNumber="281" unit="mm" value="2.7">2.7 mm</quantity>
long, 5-lobed, yellow; filament collars balustrifrom; anthers
<quantity id="4CB99B784D30FF98FBA512971F138EA6" box="[1048,1136,1969,1996]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="4" pageNumber="281" unit="mm" value="1.5">1.5 mm</quantity>
long (including appendage), bases caudate, a half as long as the filament collar, yellowish, appendages
<emphasis id="B935EA8F4D32FF9AFEBA15FF1A7B8998" box="[263,280,217,242]" italics="true" pageId="6" pageNumber="283">c.</emphasis>
0.3 ×
<quantity id="4CB99B784D32FF9AFEE215FE1ADA8998" box="[351,441,216,242]" metricMagnitude="-4" metricUnit="m" metricValue="2.0" pageId="6" pageNumber="283" unit="mm" value="0.2">0.2 mm</quantity>
; style branches truncate with a crown of sweeping trichomes, yellowish. Achenes cylindrical,
<emphasis id="B935EA8F4D32FF9AFF3F143E1BF0885B" box="[130,147,280,305]" italics="true" pageId="6" pageNumber="283">c.</emphasis>
1.7 ×
<quantity id="4CB99B784D32FF9AFF6A143E1A4C8858" box="[215,303,280,306]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="6" pageNumber="283" unit="mm" value="0.3">0.3 mm</quantity>
(immature),
<emphasis id="B935EA8F4D32FF9AFE78143E1AB5885B" box="[453,470,280,305]" italics="true" pageId="6" pageNumber="283">c.</emphasis>
9-ribbed, glabrous; pappus
<quantity id="4CB99B784D32FF9AFF3B14111A6B8838" box="[134,264,311,338]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" metricValueMax="6.2" metricValueMin="5.8" pageId="6" pageNumber="283" unit="mm" value="6.0" valueMax="6.2" valueMin="5.8">5.8-6.2 mm</quantity>
long, barbellate, white.
</paragraph>
</subSubSection>
<caption id="DF3E66154D31FF99FF3912E71A1B8E83" ID-DOI="http://doi.org/10.5281/zenodo.8109275" ID-Zenodo-Dep="8109275" httpUri="https://zenodo.org/record/8109275/files/figure.png" pageId="5" pageNumber="282" startId="5.[132,143,1985,2003]" targetBox="[132,1456,216,1932]" targetPageId="5" targetType="figure">
<paragraph id="8BFE369D4D31FF99FF3912E71A1B8E83" blockId="5.[132,1455,1984,2025]" pageId="5" pageNumber="282">
<smallCapsWord id="8D18A0414D31FF99FF3912E71BFE8EBB" baselines="1998,1998" box="[132,157,1985,2003]" lowerCaseFontSize="5" mainFontSize="7" normCase="title" normString="Fig" pageId="5" pageNumber="282">FIG</smallCapsWord>
. 2. — Field pictures of
<taxonomicName id="4C414D1E4D31FF99FED512E619F68EB8" authority="J. Calvo &amp; Callm." authorityName="J. Calvo &amp; Callm." authorityYear="2023" box="[360,661,1984,2003]" class="Magnoliopsida" family="Asteraceae" genus="Senecio" kingdom="Plantae" order="Asterales" pageId="5" pageNumber="282" phylum="Tracheophyta" rank="species" species="marinae" status="sp. nov.">
<emphasis id="B935EA8F4D31FF99FED512E61A968EB9" box="[360,501,1984,2003]" italics="true" pageId="5" pageNumber="282">Senecio marinae</emphasis>
J. Calvo &amp; Callm.
</taxonomicName>
,
<taxonomicNameLabel id="A20657F44D31FF99FD2112E719878EB9" box="[668,740,1985,2003]" pageId="5" pageNumber="282" rank="species">sp. nov.</taxonomicNameLabel>
:
<emphasis id="B935EA8F4D31FF99FD5612E7199B8EB9" bold="true" box="[747,760,1985,2003]" pageId="5" pageNumber="282">A</emphasis>
, habit;
<emphasis id="B935EA8F4D31FF99FC8712E718248EB9" bold="true" box="[826,839,1985,2003]" pageId="5" pageNumber="282">B</emphasis>
, synflorescence;
<emphasis id="B935EA8F4D31FF99FC6012E618888EB8" bold="true" box="[989,1003,1984,2002]" pageId="5" pageNumber="282">C</emphasis>
, detail of the capitulum;
<emphasis id="B935EA8F4D31FF99FB7E12E61F868EB8" bold="true" box="[1219,1253,1984,2002]" pageId="5" pageNumber="282">A-C</emphasis>
,
<emphasis id="B935EA8F4D31FF99FB4C12E71ECA8EB9" box="[1265,1449,1985,2003]" italics="true" pageId="5" pageNumber="282">Wohlhauser et al. 785</emphasis>
. Photographs: S. Wohlhauser.
</paragraph>
</caption>
<subSubSection id="C35B65164D32FF9AFF39145E1A008DA4" pageId="6" pageNumber="283" type="discussion">
<paragraph id="8BFE369D4D32FF9AFF39145E1BAC88FA" blockId="6.[130,777,376,1231]" box="[132,207,376,402]" pageId="6" pageNumber="283">
<smallCapsWord id="8D18A0414D32FF9AFF39145E1BAC88FA" baselines="395,395" box="[132,207,376,402]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Notes" pageId="6" pageNumber="283">NOTES</smallCapsWord>
</paragraph>
<paragraph id="8BFE369D4D32FF9AFF3914BE1AA38D45" blockId="6.[130,777,376,1231]" pageId="6" pageNumber="283">
In its habit and leaf morphology,
<taxonomicName id="4C414D1E4D32FF9AFE4514B119C788DB" authority="Calvo &amp; Callmander, 2023" authorityName="Calvo &amp; Callmander" authorityYear="2023" box="[504,676,407,433]" class="Magnoliopsida" family="Asteraceae" genus="Senecio" kingdom="Plantae" order="Asterales" pageId="6" pageNumber="283" phylum="Tracheophyta" rank="species" species="marinae" status="sp. nov.">
<emphasis id="B935EA8F4D32FF9AFE4514B119C788DB" box="[504,676,407,433]" italics="true" pageId="6" pageNumber="283">Senecio marinae</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A20657F44D32FF9AFD0E14BE186488D8" box="[691,775,408,434]" pageId="6" pageNumber="283" rank="species">sp. nov.</taxonomicNameLabel>
presents morphological affinities with both
<taxonomicName id="4C414D1E4D32FF9AFD3914911B93889B" authority="Humbert" authorityName="Humbert" class="Magnoliopsida" family="Asteraceae" genus="Senecio" kingdom="Plantae" order="Asterales" pageId="6" pageNumber="283" phylum="Tracheophyta" rank="species" species="francoisii">
<emphasis id="B935EA8F4D32FF9AFD391491186B88BB" box="[644,776,439,465]" italics="true" pageId="6" pageNumber="283">S. francoisii</emphasis>
Humbert
</taxonomicName>
and
<taxonomicName id="4C414D1E4D32FF9AFE9214F1196D889B" authority="Forssk." authorityName="Forssk." box="[303,526,471,497]" class="Magnoliopsida" family="Asteraceae" genus="Senecio" kingdom="Plantae" order="Asterales" pageId="6" pageNumber="283" phylum="Tracheophyta" rank="species" species="hadiensis">
<emphasis id="B935EA8F4D32FF9AFE9214F11AD1889B" box="[303,434,471,497]" italics="true" pageId="6" pageNumber="283">S. hadiensis</emphasis>
Forssk.
</taxonomicName>
[synonym of
<taxonomicName id="4C414D1E4D32FF9AFD0814F11A488B7B" authority="A. Rich." authorityName="A. Rich." class="Magnoliopsida" family="Asteraceae" genus="Senecio" kingdom="Plantae" order="Asterales" pageId="6" pageNumber="283" phylum="Tracheophyta" rank="species" species="petitianus">
<emphasis id="B935EA8F4D32FF9AFD0814F11BA68B7B" italics="true" pageId="6" pageNumber="283">S. petitianus</emphasis>
A. Rich.
</taxonomicName>
]. These species are characterized by having a scrambling or ascending habit, fleshy, pinnatinerved leaves, and radiate capitula.
<bibRefCitation id="EFD04B6C4D32FF9AFED21710194A8B3A" author="HUMBERT H." box="[367,553,566,592]" pageId="6" pageNumber="283" pagination="623 - 911" refId="ref2850" refString="HUMBERT H. 1963. - Flore de Madagascar et des Comores. 189 e Famille, Composees, tome III. Museum national d'Histoire naturelle, Paris: 623 - 911. https: // doi. org / 10.5962 / bhl. title. 6600" type="book chapter" year="1963">Humbert (1963)</bibRefCitation>
treated them as the only members of his informal
<taxonomicName id="4C414D1E4D32FF9AFE6F1771197D8B1B" box="[466,542,599,625]" class="Magnoliopsida" family="Asteraceae" genus="Senecio" kingdom="Plantae" order="Asterales" pageId="6" pageNumber="283" phylum="Tracheophyta" rank="genus">
<emphasis id="B935EA8F4D32FF9AFE6F1771197D8B1B" box="[466,542,599,625]" italics="true" pageId="6" pageNumber="283">Senecio</emphasis>
</taxonomicName>
group XIV. The new species, however, clearly differs in having discoid capitula. In addition,
<taxonomicName id="4C414D1E4D32FF9AFEA817B11AEF8BDA" authority="Calvo &amp; Callmander, 2023" authorityName="Calvo &amp; Callmander" authorityYear="2023" box="[277,396,663,689]" class="Magnoliopsida" family="Asteraceae" genus="Senecio" kingdom="Plantae" order="Asterales" pageId="6" pageNumber="283" phylum="Tracheophyta" rank="species" species="marinae" status="sp. nov.">
<emphasis id="B935EA8F4D32FF9AFEA817B11AEF8BDA" box="[277,396,663,689]" italics="true" pageId="6" pageNumber="283">S. marinae</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A20657F44D32FF9AFE2117B11A908BDB" box="[412,499,663,689]" pageId="6" pageNumber="283" rank="species">sp. nov.</taxonomicNameLabel>
differs from
<taxonomicName id="4C414D1E4D32FF9AFD3417B1186B8BDA" box="[649,776,662,689]" class="Magnoliopsida" family="Asteraceae" genus="Senecio" kingdom="Plantae" order="Asterales" pageId="6" pageNumber="283" phylum="Tracheophyta" rank="species" species="francoisii">
<emphasis id="B935EA8F4D32FF9AFD3417B1186B8BDA" box="[649,776,662,689]" italics="true" pageId="6" pageNumber="283">S. francoisii</emphasis>
</taxonomicName>
in stem length (
<quantity id="4CB99B784D32FF9AFE8117901AE18BBB" box="[316,386,694,721]" metricMagnitude="0" metricUnit="m" metricValue="2.5" metricValueMax="4.0" metricValueMin="1.0" pageId="6" pageNumber="283" unit="m" value="2.5" valueMax="4.0" valueMin="1.0">1-4 m</quantity>
vs
<quantity id="4CB99B784D32FF9AFE171790197E8BBB" box="[426,541,694,721]" metricMagnitude="-1" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="6" pageNumber="283" unit="m" value="0.25" valueMax="0.3" valueMin="0.2">0.2-0.3 m</quantity>
in
<taxonomicName id="4C414D1E4D32FF9AFDFB179019A58BBA" box="[582,710,694,720]" class="Magnoliopsida" family="Asteraceae" genus="Senecio" kingdom="Plantae" order="Asterales" pageId="6" pageNumber="283" phylum="Tracheophyta" rank="species" species="francoisii">
<emphasis id="B935EA8F4D32FF9AFDFB179019A58BBA" box="[582,710,694,720]" italics="true" pageId="6" pageNumber="283">S. francoisii</emphasis>
</taxonomicName>
), leaf size (6-13×
<quantity id="4CB99B784D32FF9AFEBB17F01AE38B9A" box="[262,384,726,753]" metricMagnitude="-2" metricUnit="m" metricValue="3.9" metricValueMax="5.5" metricValueMin="2.3" pageId="6" pageNumber="283" unit="cm" value="3.9" valueMax="5.5" valueMin="2.3">2.3-5.5 cm</quantity>
vs 3-8 ×
<quantity id="4CB99B784D32FF9AFE6017F019348B9A" box="[477,599,726,753]" metricMagnitude="-2" metricUnit="m" metricValue="1.85" metricValueMax="2.5" metricValueMin="1.2" pageId="6" pageNumber="283" unit="cm" value="1.85" valueMax="2.5" valueMin="1.2">1.2-2.5 cm</quantity>
in
<taxonomicName id="4C414D1E4D32FF9AFDC617F019948B9A" box="[635,759,726,752]" class="Magnoliopsida" family="Asteraceae" genus="Senecio" kingdom="Plantae" order="Asterales" pageId="6" pageNumber="283" phylum="Tracheophyta" rank="species" species="francoisii">
<emphasis id="B935EA8F4D32FF9AFDC617F019948B9A" box="[635,759,726,752]" italics="true" pageId="6" pageNumber="283">S. francoisii</emphasis>
</taxonomicName>
), and involucral bract length (
<quantity id="4CB99B784D32FF9AFE7B17D0192D8A7A" box="[454,590,758,785]" metricMagnitude="-3" metricUnit="m" metricValue="6.6" metricValueMax="7.4" metricValueMin="5.8" pageId="6" pageNumber="283" unit="mm" value="6.6" valueMax="7.4" valueMin="5.8">5.8-7.4 mm</quantity>
vs
<quantity id="4CB99B784D32FF9AFDCB17D019848A7A" box="[630,743,758,785]" metricMagnitude="-3" metricUnit="m" metricValue="5.2" metricValueMax="5.4" metricValueMin="5.0" pageId="6" pageNumber="283" unit="mm" value="5.2" valueMax="5.4" valueMin="5.0">5-5.4 mm</quantity>
in
<taxonomicName id="4C414D1E4D32FF9AFF3B16301A608A5A" box="[134,259,790,816]" class="Magnoliopsida" family="Asteraceae" genus="Senecio" kingdom="Plantae" order="Asterales" pageId="6" pageNumber="283" phylum="Tracheophyta" rank="species" species="francoisii">
<emphasis id="B935EA8F4D32FF9AFF3B16301A608A5A" box="[134,259,790,816]" italics="true" pageId="6" pageNumber="283">S. francoisii</emphasis>
</taxonomicName>
). With respect to
<taxonomicName id="4C414D1E4D32FF9AFE6C1630192D8A5A" box="[465,590,790,816]" class="Magnoliopsida" family="Asteraceae" genus="Senecio" kingdom="Plantae" order="Asterales" pageId="6" pageNumber="283" phylum="Tracheophyta" rank="species" species="hadiensis">
<emphasis id="B935EA8F4D32FF9AFE6C1630192D8A5A" box="[465,590,790,816]" italics="true" pageId="6" pageNumber="283">S. hadiensis</emphasis>
</taxonomicName>
, the new species further differs in the involucral bract length (
<quantity id="4CB99B784D32FF9AFD3C1613186B8A3A" box="[641,776,821,848]" metricMagnitude="-3" metricUnit="m" metricValue="6.6" metricValueMax="7.4" metricValueMin="5.8" pageId="6" pageNumber="283" unit="mm" value="6.6" valueMax="7.4" valueMin="5.8">5.8-7.4 mm</quantity>
vs
<quantity id="4CB99B784D32FF9AFF1F16701A458A1A" box="[162,294,854,880]" metricMagnitude="-3" metricUnit="m" metricValue="3.95" metricValueMax="4.2" metricValueMin="3.7" pageId="6" pageNumber="283" unit="mm" value="3.95" valueMax="4.2" valueMin="3.7">3.7-4.2 mm</quantity>
in
<taxonomicName id="4C414D1E4D32FF9AFEF716701AA78A05" box="[330,452,853,880]" class="Magnoliopsida" family="Asteraceae" genus="Senecio" kingdom="Plantae" order="Asterales" pageId="6" pageNumber="283" phylum="Tracheophyta" rank="species" species="hadiensis">
<emphasis id="B935EA8F4D32FF9AFEF716701AA78A05" box="[330,452,853,880]" italics="true" pageId="6" pageNumber="283">S. hadiensis</emphasis>
</taxonomicName>
). The
<taxonomicName id="4C414D1E4D32FF9AFDB6167319EE8A05" box="[523,653,853,879]" pageId="6" pageNumber="283" rank="variety" variety="with">variety with</taxonomicName>
discoid capitula described from
<collectingCountry id="F356760D4D32FF9AFED316501AB78AFA" box="[366,468,886,912]" name="Comoros" pageId="6" pageNumber="283">Comoros</collectingCountry>
under the name
<taxonomicName id="4C414D1E4D32FF9AFD3A16501AC88AC5" authority="Humbert" authorityName="Humbert" class="Magnoliopsida" family="Asteraceae" genus="Senecio" kingdom="Plantae" order="Asterales" pageId="6" pageNumber="283" phylum="Tracheophyta" rank="subVariety" species="petitianus" subVariety="discoideus">
<emphasis id="B935EA8F4D32FF9AFD3A165018648AE5" box="[647,775,886,912]" italics="true" pageId="6" pageNumber="283">S. petitianus</emphasis>
subvar.
<emphasis id="B935EA8F4D32FF9AFF6B16B31A5E8AC5" box="[214,317,917,943]" italics="true" pageId="6" pageNumber="283">discoideus</emphasis>
Humbert
</taxonomicName>
, can be also differentiated by the remarkable shorter involucral bracts; it has not been recorded in
<collectingCountry id="F356760D4D32FF9AFF2216F31A408A85" box="[159,291,981,1007]" name="Madagascar" pageId="6" pageNumber="283">Madagascar</collectingCountry>
. It is interesting to note that
<taxonomicName id="4C414D1E4D32FF9AFDE716F319B68A85" box="[602,725,981,1007]" class="Magnoliopsida" family="Asteraceae" genus="Senecio" kingdom="Plantae" order="Asterales" pageId="6" pageNumber="283" phylum="Tracheophyta" rank="species" species="francoisii">
<emphasis id="B935EA8F4D32FF9AFDE716F319B68A85" box="[602,725,981,1007]" italics="true" pageId="6" pageNumber="283">S. francoisii</emphasis>
</taxonomicName>
, the Malagasy populations of
<taxonomicName id="4C414D1E4D32FF9AFE2016D3197A8D65" box="[413,537,1013,1039]" class="Magnoliopsida" family="Asteraceae" genus="Senecio" kingdom="Plantae" order="Asterales" pageId="6" pageNumber="283" phylum="Tracheophyta" rank="species" species="hadiensis">
<emphasis id="B935EA8F4D32FF9AFE2016D3197A8D65" box="[413,537,1013,1039]" italics="true" pageId="6" pageNumber="283">S. hadiensis</emphasis>
</taxonomicName>
, and the new species display caudate anther bases.
</paragraph>
<paragraph id="8BFE369D4D32FF9AFF2611131A008DA4" blockId="6.[130,777,376,1231]" pageId="6" pageNumber="283">
The new species seems to be restricted to the northern mountains of
<collectingCountry id="F356760D4D32FF9AFEA611731AF88D05" box="[283,411,1109,1135]" name="Madagascar" pageId="6" pageNumber="283">Madagascar</collectingCountry>
, whereas
<taxonomicName id="4C414D1E4D32FF9AFE42117319148D05" box="[511,631,1109,1135]" class="Magnoliopsida" family="Asteraceae" genus="Senecio" kingdom="Plantae" order="Asterales" pageId="6" pageNumber="283" phylum="Tracheophyta" rank="species" species="francoisii">
<emphasis id="B935EA8F4D32FF9AFE42117319148D05" box="[511,631,1109,1135]" italics="true" pageId="6" pageNumber="283">S. francoisii</emphasis>
</taxonomicName>
is distributed through the central-east
<collectingCountry id="F356760D4D32FF9AFE321152196D8DE4" box="[399,526,1140,1166]" name="Madagascar" pageId="6" pageNumber="283">Madagascar</collectingCountry>
and
<taxonomicName id="4C414D1E4D32FF9AFDFB115319DC8DE4" box="[582,703,1140,1167]" class="Magnoliopsida" family="Asteraceae" genus="Senecio" kingdom="Plantae" order="Asterales" pageId="6" pageNumber="283" phylum="Tracheophyta" rank="species" species="hadiensis">
<emphasis id="B935EA8F4D32FF9AFDFB115319DC8DE4" box="[582,703,1140,1167]" italics="true" pageId="6" pageNumber="283">S. hadiensis</emphasis>
</taxonomicName>
in the centre and south-east part of the island. Their distributions areas do not overlap.
</paragraph>
</subSubSection>
</treatment>
</document>
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