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<mods:title id="E303EE04F96C99BC889C36C7CB7ACE79">Description of a new species of catshark, Atelomycterus marnkalha n. sp. (Carcharhiniformes: Scyliorhinidae) from north-east Australia</mods:title>
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<emphasis id="B935EA87FFA8FFACFF19FD83FE5DFDF8" bold="true" box="[151,464,597,623]" italics="true" pageId="5" pageNumber="24">Atelomycterus marnkalha</emphasis>
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<paragraph id="8BFE3695FFA8FFACFF19FDAEFE32FD05" blockId="5.[151,615,597,698]" box="[151,447,632,658]" pageId="5" pageNumber="24">Eastern Banded Catshark</paragraph>
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<figureCitation id="137A2A10FFA8FFACFF19FD76FE8DFD2D" box="[151,256,672,698]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStart-4="FIGURE 5" captionStart-5="FIGURE 6" captionStart-6="FIGURE 7" captionStartId-0="6.[151,255,1082,1106]" captionStartId-1="6.[151,255,1460,1484]" captionStartId-2="8.[151,255,1103,1127]" captionStartId-3="9.[151,255,1795,1819]" captionStartId-4="10.[151,255,1595,1619]" captionStartId-5="11.[151,255,937,961]" captionStartId-6="12.[151,255,1433,1457]" captionTargetBox-0="[270,1316,625,1035]" captionTargetBox-1="[268,1315,1250,1378]" captionTargetBox-2="[286,1322,411,1068]" captionTargetBox-3="[304,1297,710,1736]" captionTargetBox-4="[384,1203,194,1551]" captionTargetBox-5="[270,1322,202,898]" captionTargetBox-6="[264,1322,474,1405]" captionTargetId-0="figure@6.[264,1323,597,1055]" captionTargetId-1="figure@6.[264,1323,1194,1437]" captionTargetId-2="figure@8.[264,1323,394,1079]" captionTargetId-3="figure@9.[285,1303,674,1737]" captionTargetId-4="figure@10.[384,1203,194,1571]" captionTargetId-5="figure@11.[264,1323,194,913]" captionTargetId-6="figure@12.[264,1323,474,1409]" captionTargetPageId-0="6" captionTargetPageId-1="6" captionTargetPageId-2="8" captionTargetPageId-3="9" captionTargetPageId-4="10" captionTargetPageId-5="11" captionTargetPageId-6="12" captionText-0="FIGURE 1. Atelomycterus marnkalha n. sp.; fixed, holotype adult male, 386 mm TL (CSIRO H 6144 - 01). A: Lateral view; B Dorsal view. Photo by S. Riddoch." captionText-1="FIGURE 2. Lateral view of Atelomycterus marnkalha n. sp.; fresh specimen, paratype, adult female, 351 mm TL, (CSIRO H 6145 - 01). Photo by S. Riddoch." captionText-2="FIGURE 3. Ventral head view of Atelomycterus marnkalha n. sp.; fixed, holotype adult male, 386 mm TL (CSIRO H 6144 - 01). Photo by S. Riddoch." captionText-3="FIGURE 4. Atelomycterus marnkalha n. sp.; scanning electron micrographs, 459 mm TL male A – C upper teeth; D – H lower teeth. A – B upper anterolateral teeth from near symphysis in A, labial; and B, lingual views; C, posterior tooth in lingual view; D – F, lower anterolateral teeth from near symphysis in D, labial; E, lingual; and F, dorso-lateral views; G – H, posterior teeth in G, labial and H, lingual views. Photos by K. Sewell." captionText-4="FIGURE 5. Atelomycterus marnkalha n. sp.; scanning electron micrographs of lateral trunk denticles in three-quarter anteriorlateral view, 459 mm TL male at A, 100 Μm, and B, 50 Μm. Photo by K. Sewell." captionText-5="FIGURE 6. Claspers of Atelomycterus marnkalha n. sp.; 459 mm TL adult male (QM I 38089). A: Glans not dilated; B: Glans spread. Abbreviations: AP: Apopyle; CLD: Clasper denticles; CRH: Cover Rhipidion; ERH: Exorhipidion; FCG: Fused clasper groove; FRT: Free tab on cover rhipidion; P- 2: Pelvic fin; PSP: pseudopera; PSS: Pseudosiphon; RH: Rhipidion. Illustration by I. P. Jacobsen." captionText-6="FIGURE 7. Claspers of Atelomycterus marnkalha n. sp.; 459 mm TL adult male (QM I 38089). A: Ventral view; B: Dorsal vies; C: dorsal view of glans, terminal cartilage spread. Abbreviations: AX: Axial cartilage; β: Beta cartilage; B: Basypterygium; B 1: Intermediate segment; G: End style; RD: Dorsal marginal cartilage; RD 2: Accessory dorsal marginal cartilage; RV: Ventral marginal cartilage; T 3: accessory terminal cartilage; TD: Dorsal terminal cartilage; TD 2: dorsal terminal 2 cartilage; TV: ventral terminal cartilage; TV 2: Ventral terminal 2 cartilage; TV 3: ventral terminal 3 cartilage. Illustration by I. P. Jacobsen." httpUri-0="https://zenodo.org/record/177425/files/figure.png" httpUri-1="https://zenodo.org/record/177426/files/figure.png" httpUri-2="https://zenodo.org/record/177427/files/figure.png" httpUri-3="https://zenodo.org/record/177428/files/figure.png" httpUri-4="https://zenodo.org/record/177429/files/figure.png" httpUri-5="https://zenodo.org/record/177430/files/figure.png" httpUri-6="https://zenodo.org/record/177431/files/figure.png" pageId="5" pageNumber="24">Figs 1–7</figureCitation>
; 
<tableCitation id="C6C3032EFFA8FFACFE82FD76FEE3FD2D" box="[268,366,672,698]" captionStart="TABLE 1" captionStartId="1.[151,239,1066,1090]" captionText="TABLE 1. Morphometric data expressed as % of TL for the Atelomycterus marnkalha n. sp. holotype, paratype size ranges and A. fasciatus comparisons." pageId="5" pageNumber="24">Tables 1</tableCitation>
and 
<tableCitation id="C6C3032EFFA8FFACFE29FD76FE3BFD2D" box="[423,438,672,698]" captionStart="TABLE 2" captionStartId="4.[151,239,151,175]" captionTargetBox="[159,1402,242,820]" captionTargetPageId="4" captionText="TABLE 2. Vertebral count comparisons between Atelomycterus marnkalha n. sp. and A. fasciatus. * Vertebral counts and percentages compiled from Compagno and Stevens’ (1993) original description." httpUri="http://table.plazi.org/id/DF3E661DFFA9FFADFF19FF41FC62FF46" pageId="5" pageNumber="24" tableUuid="DF3E661DFFA9FFADFF19FF41FC62FF46">2</tableCitation>
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sp. A (Torres Strait form, not 
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sp. A = 
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<emphasis id="B935EA87FFA8FFACFD6DFD26FCEDFC90" box="[739,864,752,775]" italics="true" pageId="5" pageNumber="24">A. fasciatus</emphasis>
, 
<bibRefCitation id="EFD04B64FFA8FFACFCE3FD26FB13FC90" author="Compagno" box="[877,1182,752,775]" pageId="5" pageNumber="36" refString="Compagno, L. J. V. &amp; Stevens, J. D. (1993) Atelomycterus fasciatus n. sp., a new catshark (Chondrichthyes: Carcharhiniformes: Scyliorhinidae) from tropical Australia. Records of the Australian Museum, Vol. 45, 147 - 169." type="journal article" year="1993">Compagno &amp; Stevens, 1993</bibRefCitation>
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): 
<treatmentCitation id="0AE01084FFA8FFACFB3EFD26FF62FCB0" author="Last" page="189" pageId="5" pageNumber="24" year="1994">
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: 189
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(in part) (Plate 22, Figure 26.17b)
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Material studied. 
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:
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H 6144.01, mature male, 
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: 
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H 6148-01, adult male, 
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, Torres Strait, Queensland, 
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H 6147-01, adult male, 
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, Torres Strait, Queensland, 0 
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H 6145-01, female, 
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<collectionCode id="ED50AE50FFA8FFACFC9FFBF6FCEAFBAD" box="[785,871,1056,1082]" httpUri="http://grbio.org/cool/0k2v-y6wz" name="Australian National Fish Collection" pageId="5" pageNumber="24">CSIRO</collectionCode>
H 6149-02, immature male, 
<quantity id="4CB99B70FFA8FFACFB39FBF6FA95FBAD" box="[1207,1304,1056,1082]" metricMagnitude="-1" metricUnit="m" metricValue="3.22" pageId="5" pageNumber="24" unit="mm" value="322.0">322 mm</quantity>
<collectionCode id="ED50AE50FFA8FFACFA90FBF6FAC9FBAD" box="[1310,1348,1056,1082]" pageId="5" pageNumber="24">TL</collectionCode>
, 
<date id="FFFF1055FFA8FFACFADEFBF6FF59FBF5" pageId="5" pageNumber="24" value="2003-12-04">4 Dec. 2003</date>
, Whitsunday Passage, Queensland, 
<geoCoordinate id="EE755052FFA8FFACFD0FFB9EFD61FBF5" box="[641,748,1095,1122]" direction="south" orientation="latitude" pageId="5" pageNumber="24" precision="925" value="-20.283333">20°17´S</geoCoordinate>
, 
<geoCoordinate id="EE755052FFA8FFACFD75FB9EFCFBFBF5" box="[763,886,1095,1122]" direction="east" orientation="longitude" pageId="5" pageNumber="24" precision="925" value="148.88333">148°53´E</geoCoordinate>
, 
<quantity id="4CB99B70FFA8FFACFC0BFB9EFC48FBF5" box="[901,965,1096,1122]" metricMagnitude="1" metricUnit="m" metricValue="4.4" pageId="5" pageNumber="24" unit="m" value="44.0">44 m</quantity>
; 
<collectionCode id="ED50AE50FFA8FFACFC5DFB9EFBA4FBF5" box="[979,1065,1096,1122]" httpUri="http://grbio.org/cool/0k2v-y6wz" name="Australian National Fish Collection" pageId="5" pageNumber="24">CSIRO</collectionCode>
H 6146-01, female, 
<quantity id="4CB99B70FFA8FFACFAADFB9EFA11FBF5" box="[1315,1436,1096,1122]" metricMagnitude="-1" metricUnit="m" metricValue="2.935" pageId="5" pageNumber="24" unit="mm" value="293.5">293.5 mm</quantity>
<collectionCode id="ED50AE50FFA8FFACFF19FBA6FF33FB1D" box="[151,190,1136,1162]" pageId="5" pageNumber="24">TL</collectionCode>
, 
<date id="FFFF1055FFA8FFACFF47FBA6FEECFB1D" box="[201,353,1136,1162]" pageId="5" pageNumber="24" value="2004-01-22">22 Jan. 2004</date>
, Torres Strait, Queensland, 0 
<geoCoordinate id="EE755052FFA8FFACFD3CFBA6FC81FB1D" box="[690,780,1135,1162]" direction="south" orientation="latitude" pageId="5" pageNumber="24" precision="925" value="-9.6">9°36´S</geoCoordinate>
, 
<geoCoordinate id="EE755052FFA8FFACFC96FBA6FC1FFB1D" box="[792,914,1135,1162]" direction="east" orientation="longitude" pageId="5" pageNumber="24" precision="925" value="142.56667">142°34´E</geoCoordinate>
, 
<quantity id="4CB99B70FFA8FFACFC10FBA6FB8AFB1D" box="[926,1031,1136,1162]" metricMagnitude="1" metricUnit="m" metricValue="1.4" metricValueMax="1.7" metricValueMin="1.1" pageId="5" pageNumber="24" unit="m" value="14.0" valueMax="17.0" valueMin="11.0">11–17 m</quantity>
; QM I 34023, immature male, 
<quantity id="4CB99B70FFA8FFACFAFEFBA6FF48FB25" metricMagnitude="-1" metricUnit="m" metricValue="2.77" pageId="5" pageNumber="24" unit="mm" value="277.0">277 mm</quantity>
<collectionCode id="ED50AE50FFA8FFACFF40FB4EFF78FB25" box="[206,245,1176,1202]" pageId="5" pageNumber="24">TL</collectionCode>
, 
<date id="FFFF1055FFA8FFACFE8CFB4EFE2BFB25" box="[258,422,1176,1202]" pageId="5" pageNumber="24" value="2002-11-27">27 Nov. 2002</date>
, Polmaise Reef, Queensland, 
<geoCoordinate id="EE755052FFA8FFACFC85FB4EFCF8FB25" box="[779,885,1175,1202]" direction="south" orientation="latitude" pageId="5" pageNumber="24" precision="925" value="-23.633333">23°38´S</geoCoordinate>
, 
<geoCoordinate id="EE755052FFA8FFACFC0AFB4EFB8DFB25" box="[900,1024,1175,1202]" direction="east" orientation="longitude" pageId="5" pageNumber="24" precision="925" value="151.88333">151°53´E</geoCoordinate>
, 
<quantity id="4CB99B70FFA8FFACFB80FB4EFBC2FB25" box="[1038,1103,1176,1202]" metricMagnitude="1" metricUnit="m" metricValue="3.6" pageId="5" pageNumber="24" unit="m" value="36.0">36 m</quantity>
; QM I 34022, adult female, 
<quantity id="4CB99B70FFA8FFACFF19FB16FF75FB4D" box="[151,248,1216,1242]" metricMagnitude="-1" metricUnit="m" metricValue="4.66" pageId="5" pageNumber="24" unit="mm" value="466.0">466 mm</quantity>
<collectionCode id="ED50AE50FFA8FFACFF71FB16FEA8FB4D" box="[255,293,1216,1242]" pageId="5" pageNumber="24">TL</collectionCode>
, 
<date id="FFFF1055FFA8FFACFEBFFB16FE5FFB4D" box="[305,466,1216,1242]" pageId="5" pageNumber="24" value="2002-11-27">27 Nov. 2002</date>
, SE of Masthead Island, Queensland, 
<geoCoordinate id="EE755052FFA8FFACFC1EFB16FC74FB4D" box="[912,1017,1215,1242]" direction="south" orientation="latitude" pageId="5" pageNumber="24" precision="925" value="-23.683332">23°41´S</geoCoordinate>
, 
<geoCoordinate id="EE755052FFA8FFACFB88FB16FB0DFB4D" box="[1030,1152,1215,1242]" direction="east" orientation="longitude" pageId="5" pageNumber="24" precision="925" value="151.81667">151°49´E</geoCoordinate>
, 
<quantity id="4CB99B70FFA8FFACFB03FB16FB7AFB4D" box="[1165,1271,1216,1242]" metricMagnitude="1" metricUnit="m" metricValue="3.5" metricValueMax="3.6" metricValueMin="3.4" pageId="5" pageNumber="24" unit="m" value="35.0" valueMax="36.0" valueMin="34.0">34–36 m</quantity>
; QM I 38078, female, 
<quantity id="4CB99B70FFA8FFACFF79FB3EFED6FA95" box="[247,347,1256,1282]" metricMagnitude="-1" metricUnit="m" metricValue="3.2399999999999998" pageId="5" pageNumber="24" unit="mm" value="324.0">324 mm</quantity>
<collectionCode id="ED50AE50FFA8FFACFEEAFB3EFE06FA95" box="[356,395,1256,1282]" pageId="5" pageNumber="24">TL</collectionCode>
, 
<date id="FFFF1055FFA8FFACFE17FB3EFDB3FA95" box="[409,574,1256,1282]" pageId="5" pageNumber="24" value="2005-11-13">13 Nov. 2005</date>
, Queensland, 
<geoCoordinate id="EE755052FFA8FFACFD66FB3EFCDEFA95" box="[744,851,1255,1282]" direction="south" orientation="latitude" pageId="5" pageNumber="24" precision="925" value="-21.883333">21°53´S</geoCoordinate>
, 
<geoCoordinate id="EE755052FFA8FFACFCECFB3EFC52FA95" box="[866,991,1255,1282]" direction="east" orientation="longitude" pageId="5" pageNumber="24" precision="925" value="150.35">150°21´E</geoCoordinate>
, 
<quantity id="4CB99B70FFA8FFACFC63FB3EFBA3FA95" box="[1005,1070,1256,1282]" metricMagnitude="1" metricUnit="m" metricValue="3.9" pageId="5" pageNumber="24" unit="m" value="39.0">39 m</quantity>
; 
<collectionCode id="ED50AE50FFA8FFACFBB3FB3EFB1EFA95" box="[1085,1171,1256,1282]" httpUri="http://grbio.org/cool/0k2v-y6wz" name="Australian National Fish Collection" pageId="5" pageNumber="24">CSIRO</collectionCode>
H 1118-01, immature male, 
<quantity id="4CB99B70FFA8FFACFF6FFAC6FEC9FABD" box="[225,324,1296,1322]" metricMagnitude="-1" metricUnit="m" metricValue="3.0" pageId="5" pageNumber="24" unit="mm" value="300.0">300 mm</quantity>
<collectionCode id="ED50AE50FFA8FFACFEC0FAC6FEF8FABD" box="[334,373,1296,1322]" pageId="5" pageNumber="24">TL</collectionCode>
, 
<date id="FFFF1055FFA8FFACFE0CFAC6FD94FABD" box="[386,537,1296,1322]" pageId="5" pageNumber="24" value="1987-11-04">4 Nov. 1987</date>
, Arafura Sea, Queensland, 
<geoCoordinate id="EE755052FFA8FFACFCEDFAC6FC42FABD" box="[867,975,1295,1322]" direction="south" orientation="latitude" pageId="5" pageNumber="24" precision="925" value="-10.516666">10°31´S</geoCoordinate>
, 
<geoCoordinate id="EE755052FFA8FFACFC50FAC6FBD6FABD" box="[990,1115,1295,1322]" direction="east" orientation="longitude" pageId="5" pageNumber="24" precision="925" value="140.8">140°48´E</geoCoordinate>
, 
<quantity id="4CB99B70FFA8FFACFBE4FAC6FB27FABD" box="[1130,1194,1296,1322]" metricMagnitude="1" metricUnit="m" metricValue="4.3" pageId="5" pageNumber="24" unit="m" value="43.0">43 m</quantity>
; AMS I 15557-003, adult male, 
<quantity id="4CB99B70FFA8FFACFEABFAEEFE05FAC5" box="[293,392,1336,1362]" metricMagnitude="-1" metricUnit="m" metricValue="3.75" pageId="5" pageNumber="24" unit="mm" value="375.0">375 mm</quantity>
<collectionCode id="ED50AE50FFA8FFACFE1FFAEEFE35FAC5" box="[401,440,1336,1362]" pageId="5" pageNumber="24">TL</collectionCode>
, 1964, North Mornington, Queensland, 
<geoCoordinate id="EE755052FFA8FFACFC19FAEEFB8EFAC5" box="[919,1027,1335,1362]" direction="south" orientation="latitude" pageId="5" pageNumber="24" precision="925" value="-16.633333">16°38´S</geoCoordinate>
, 
<geoCoordinate id="EE755052FFA8FFACFB9CFAEEFB03FAC5" box="[1042,1166,1335,1362]" direction="east" orientation="longitude" pageId="5" pageNumber="24" precision="925" value="140.03334">140°02´E</geoCoordinate>
, 
<quantity id="4CB99B70FFA8FFACFB13FAEEFB53FAC5" box="[1181,1246,1336,1362]" metricMagnitude="1" metricUnit="m" metricValue="3.2" pageId="5" pageNumber="24" unit="m" value="32.0">32 m</quantity>
; AMS I 21842- 0 0 1, female, 
<quantity id="4CB99B70FFA8FFACFEBEFAB6FE1CFAED" box="[304,401,1376,1402]" metricMagnitude="-1" metricUnit="m" metricValue="2.7" pageId="5" pageNumber="24" unit="mm" value="270.0">270 mm</quantity>
<collectionCode id="ED50AE50FFA8FFACFE16FAB6FE32FAED" box="[408,447,1376,1402]" pageId="5" pageNumber="24">TL</collectionCode>
, 
<date id="FFFF1055FFA8FFACFE44FAB6FDE6FAED" box="[458,619,1376,1402]" pageId="5" pageNumber="24" value="1980-11-16">16 Nov. 1980</date>
, Arafura Sea, Queensland, 
<geoCoordinate id="EE755052FFA8FFACFC22FAB6FB98FAED" box="[940,1045,1375,1402]" direction="south" orientation="latitude" pageId="5" pageNumber="24" precision="925" value="-10.616667">10°37´S</geoCoordinate>
, 
<geoCoordinate id="EE755052FFA8FFACFBACFAB6FB11FAED" box="[1058,1180,1375,1402]" direction="east" orientation="longitude" pageId="5" pageNumber="24" precision="925" value="133.78334">133°47´E</geoCoordinate>
, 
<specimenCount id="9D47FD1CFFA8FFACFB26FAB6FB66FAED" box="[1192,1259,1376,1402]" pageId="5" pageNumber="24" type="generic">
<quantity id="4CB99B70FFA8FFACFB26FAB6FB6AFAED" box="[1192,1255,1376,1402]" metricMagnitude="1" metricUnit="m" metricValue="6.0" pageId="5" pageNumber="24" unit="m" value="60.0">60 m</quantity>
.
</specimenCount>
</paragraph>
<paragraph id="8BFE3695FFA8FFACFF48FA51FDE2F89D" blockId="5.[151,1436,855,2002]" pageId="5" pageNumber="24">
<emphasis id="B935EA87FFA8FFACFF48FA51FE39FA36" bold="true" box="[198,436,1415,1441]" pageId="5" pageNumber="24">Non-type material:</emphasis>
<specimenCount id="9D47FD1CFFA8FFACFE32FA5EFDD0FA35" box="[444,605,1416,1442]" pageId="5" pageNumber="24" type="generic">11 specimens</specimenCount>
: 
<collectionCode id="ED50AE50FFA8FFACFDE6FA5EFD33FA35" box="[616,702,1416,1442]" httpUri="http://grbio.org/cool/0k2v-y6wz" name="Australian National Fish Collection" pageId="5" pageNumber="24">CSIRO</collectionCode>
H 6150-01, 2 immature females, 270 and 
<quantity id="4CB99B70FFA8FFACFB21FA5EFA9DFA35" box="[1199,1296,1416,1442]" metricMagnitude="-1" metricUnit="m" metricValue="2.65" pageId="5" pageNumber="24" unit="mm" value="265.0">265 mm</quantity>
<collectionCode id="ED50AE50FFA8FFACFA99FA5EFAB0FA35" box="[1303,1341,1416,1442]" pageId="5" pageNumber="24">TL</collectionCode>
, 
<date id="FFFF1055FFA8FFACFAC7FA5EFF58FA5D" pageId="5" pageNumber="24" value="2004-01-16">16 Jan. 2004</date>
, Torres Strait, 
<geoCoordinate id="EE755052FFA8FFACFEF3FA66FE6BFA5D" box="[381,486,1455,1482]" direction="south" orientation="latitude" pageId="5" pageNumber="24" precision="925" value="-10.666667">10°40´S</geoCoordinate>
, 
<geoCoordinate id="EE755052FFA8FFACFE7DFA66FDE1FA5D" box="[499,620,1455,1482]" direction="east" orientation="longitude" pageId="5" pageNumber="24" precision="925" value="141.55">141°33´E</geoCoordinate>
, 
<quantity id="4CB99B70FFA8FFACFDF6FA66FD3BFA5D" box="[632,694,1456,1482]" metricMagnitude="1" metricUnit="m" metricValue="1.9" pageId="5" pageNumber="24" unit="m" value="19.0">19 m</quantity>
; 
<collectionCode id="ED50AE50FFA8FFACFD4CFA66FC95FA5D" box="[706,792,1456,1482]" httpUri="http://grbio.org/cool/0k2v-y6wz" name="Australian National Fish Collection" pageId="5" pageNumber="24">CSIRO</collectionCode>
H 6146-02, immature male, 
<quantity id="4CB99B70FFA8FFACFBE6FA66FB44FA5D" box="[1128,1225,1456,1482]" metricMagnitude="-1" metricUnit="m" metricValue="1.9" pageId="5" pageNumber="24" unit="mm" value="190.0">190 mm</quantity>
<collectionCode id="ED50AE50FFA8FFACFB41FA66FB7BFA5D" box="[1231,1270,1456,1482]" pageId="5" pageNumber="24">TL</collectionCode>
, 
<date id="FFFF1055FFA8FFACFA8FFA66FA15FA5D" box="[1281,1432,1456,1482]" pageId="5" pageNumber="24" value="2004-01-22">22 Jan. 2004</date>
, Torres Strait, Queensland, 0 
<geoCoordinate id="EE755052FFA8FFACFE6EFA0EFDB6FA65" box="[480,571,1495,1522]" direction="south" orientation="latitude" pageId="5" pageNumber="24" precision="925" value="-9.6">9°36´S</geoCoordinate>
, 
<geoCoordinate id="EE755052FFA8FFACFDC6FA0EFD4EFA65" box="[584,707,1495,1522]" direction="east" orientation="longitude" pageId="5" pageNumber="24" precision="925" value="142.56667">142°34´E</geoCoordinate>
, 
<quantity id="4CB99B70FFA8FFACFD5EFA0EFCB8FA65" box="[720,821,1496,1522]" metricMagnitude="1" metricUnit="m" metricValue="1.4" metricValueMax="1.7" metricValueMin="1.1" pageId="5" pageNumber="24" unit="m" value="14.0" valueMax="17.0" valueMin="11.0">11-17 m</quantity>
; 
<collectionCode id="ED50AE50FFA8FFACFCCCFA0EFC15FA65" box="[834,920,1496,1522]" httpUri="http://grbio.org/cool/0k2v-y6wz" name="Australian National Fish Collection" pageId="5" pageNumber="24">CSIRO</collectionCode>
H 6149-01, 2 immature females, 284.5 and 
<quantity id="4CB99B70FFA8FFACFF19F9D6FE9FF98D" box="[151,274,1536,1562]" metricMagnitude="-1" metricUnit="m" metricValue="2.535" pageId="5" pageNumber="24" unit="mm" value="253.5">253.5 mm</quantity>
<collectionCode id="ED50AE50FFA8FFACFE93F9D6FEC9F98D" box="[285,324,1536,1562]" pageId="5" pageNumber="24">TL</collectionCode>
, 1 immature male, 
<quantity id="4CB99B70FFA8FFACFDBBF9D6FD17F98D" box="[565,666,1536,1562]" metricMagnitude="-1" metricUnit="m" metricValue="3.0" pageId="5" pageNumber="24" unit="mm" value="300.0">300 mm</quantity>
<collectionCode id="ED50AE50FFA8FFACFD2BF9D6FD41F98D" box="[677,716,1536,1562]" pageId="5" pageNumber="24">TL</collectionCode>
, 
<date id="FFFF1055FFA8FFACFD55F9D6FCF9F98D" box="[731,884,1536,1562]" pageId="5" pageNumber="24" value="2003-12-04">4 Dec. 2003</date>
, Whitsunday Passage, Queensland, 
<geoCoordinate id="EE755052FFA8FFACFAA7F9D6FA1BF98D" box="[1321,1430,1535,1562]" direction="south" orientation="latitude" pageId="5" pageNumber="24" precision="925" value="-20.283333">20°17´S</geoCoordinate>
, 
<geoCoordinate id="EE755052FFA8FFACFF19F9FEFE9EF9D5" box="[151,275,1575,1602]" direction="east" orientation="longitude" pageId="5" pageNumber="24" precision="925" value="148.88333">148°53´E</geoCoordinate>
, 
<quantity id="4CB99B70FFA8FFACFEACF9FEFEEFF9D5" box="[290,354,1576,1602]" metricMagnitude="1" metricUnit="m" metricValue="4.4" pageId="5" pageNumber="24" unit="m" value="44.0">44 m</quantity>
; 
<collectionCode id="ED50AE50FFA8FFACFEFEF9FEFE4BF9D5" box="[368,454,1576,1602]" httpUri="http://grbio.org/cool/0k2v-y6wz" name="Australian National Fish Collection" pageId="5" pageNumber="24">CSIRO</collectionCode>
H 6151-01, female, 
<quantity id="4CB99B70FFA8FFACFD4CF9FEFCA8F9D5" box="[706,805,1576,1602]" metricMagnitude="-1" metricUnit="m" metricValue="3.3" pageId="5" pageNumber="24" unit="mm" value="330.0">330 mm</quantity>
<collectionCode id="ED50AE50FFA8FFACFCA0F9FEFCD8F9D5" box="[814,853,1576,1602]" pageId="5" pageNumber="24">TL</collectionCode>
, 
<date id="FFFF1055FFA8FFACFCECF9FEFC7DF9D5" box="[866,1008,1576,1602]" pageId="5" pageNumber="24" value="2004-01-09">9 Jan. 2004</date>
, Torres Strait, Queensland, 
<geoCoordinate id="EE755052FFA8FFACFAB2F9FEFA11F9D5" box="[1340,1436,1575,1602]" direction="south" orientation="latitude" pageId="5" pageNumber="24" precision="925" value="-10.6">10°36´S</geoCoordinate>
<geoCoordinate id="EE755052FFA8FFACFF19F986FE87F9FD" box="[151,266,1615,1642]" direction="east" orientation="longitude" pageId="5" pageNumber="24" precision="925" value="143.15">143°09´E</geoCoordinate>
, 
<quantity id="4CB99B70FFA8FFACFE96F986FED4F9FD" box="[280,345,1616,1642]" metricMagnitude="1" metricUnit="m" metricValue="2.6" pageId="5" pageNumber="24" unit="m" value="26.0">26 m</quantity>
; 
<collectionCode id="ED50AE50FFA8FFACFEE6F986FE33F9FD" box="[360,446,1616,1642]" httpUri="http://grbio.org/cool/0k2v-y6wz" name="Australian National Fish Collection" pageId="5" pageNumber="24">CSIRO</collectionCode>
H 6152-01, immature male, 
<quantity id="4CB99B70FFA8FFACFC90F986FC0CF9FD" box="[798,897,1616,1642]" metricMagnitude="-1" metricUnit="m" metricValue="3.1" pageId="5" pageNumber="24" unit="mm" value="310.0">310 mm</quantity>
<collectionCode id="ED50AE50FFA8FFACFC05F986FC3FF9FD" box="[907,946,1616,1642]" pageId="5" pageNumber="24">TL</collectionCode>
, 
<date id="FFFF1055FFA8FFACFC4EF986FBD8F9FD" box="[960,1109,1616,1642]" pageId="5" pageNumber="24" value="2003-12-07">7 Dec. 2003</date>
, Swain Reefs, Queensland, 
<geoCoordinate id="EE755052FFA8FFACFF19F9AEFE8EF905" box="[151,259,1655,1682]" direction="south" orientation="latitude" pageId="5" pageNumber="24" precision="925" value="-21.516666">21°31´S</geoCoordinate>
, 
<geoCoordinate id="EE755052FFA8FFACFE9CF9AEFE03F905" box="[274,398,1655,1682]" direction="east" orientation="longitude" pageId="5" pageNumber="24" precision="925" value="151.46666">151°28´E</geoCoordinate>
, 
<quantity id="4CB99B70FFA8FFACFE13F9AEFE53F905" box="[413,478,1656,1682]" metricMagnitude="1" metricUnit="m" metricValue="5.7" pageId="5" pageNumber="24" unit="m" value="57.0">57 m</quantity>
; QM I 38079, adult female, 
<quantity id="4CB99B70FFA8FFACFCB6F9AEFC16F905" box="[824,923,1656,1682]" metricMagnitude="-1" metricUnit="m" metricValue="4.59" pageId="5" pageNumber="24" unit="mm" value="459.0">459 mm</quantity>
<collectionCode id="ED50AE50FFA8FFACFC2BF9AEFC41F905" box="[933,972,1656,1682]" pageId="5" pageNumber="24">TL</collectionCode>
, 
<date id="FFFF1055FFA8FFACFC54F9AEFBF2F905" box="[986,1151,1656,1682]" pageId="5" pageNumber="24" value="2005-11-14">14 Nov. 2005</date>
, Broad Sound Channel, Queensland, 
<geoCoordinate id="EE755052FFA8FFACFEBFF976FE10F92D" box="[305,413,1695,1722]" direction="south" orientation="latitude" pageId="5" pageNumber="24" precision="925" value="-22.116667">22°07´S</geoCoordinate>
, 
<geoCoordinate id="EE755052FFA8FFACFE25F976FDAAF92D" box="[427,551,1695,1722]" direction="east" orientation="longitude" pageId="5" pageNumber="24" precision="925" value="150.31667">150°19´E</geoCoordinate>
, 
<quantity id="4CB99B70FFA8FFACFDBBF976FDF8F92D" box="[565,629,1696,1722]" metricMagnitude="1" metricUnit="m" metricValue="2.5" pageId="5" pageNumber="24" unit="m" value="25.0">25 m</quantity>
; QM I 38080, immature male, 
<quantity id="4CB99B70FFA8FFACFC65F976FBC0F92D" box="[1003,1101,1696,1722]" metricMagnitude="-1" metricUnit="m" metricValue="3.08" pageId="5" pageNumber="24" unit="mm" value="308.0">308 mm</quantity>
<collectionCode id="ED50AE50FFA8FFACFBD8F976FBF0F92D" box="[1110,1149,1696,1722]" pageId="5" pageNumber="24">TL</collectionCode>
, 
<date id="FFFF1055FFA8FFACFB04F976FA96F92D" box="[1162,1307,1696,1722]" pageId="5" pageNumber="24" value="2004-10-06">6 Oct. 2004</date>
, Lads Passage, Queensland, 
<geoCoordinate id="EE755052FFA8FFACFEFCF91EFE56F975" box="[370,475,1735,1762]" direction="south" orientation="latitude" pageId="5" pageNumber="24" precision="925" value="-13.85">13°51´S</geoCoordinate>
, 
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, 
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; QM I 38081, female, 
<quantity id="4CB99B70FFA8FFACFC38F91EFB9AF975" box="[950,1047,1736,1762]" metricMagnitude="-1" metricUnit="m" metricValue="3.54" pageId="5" pageNumber="24" unit="mm" value="354.0">354 mm</quantity>
<collectionCode id="ED50AE50FFA8FFACFB90F91EFBC8F975" box="[1054,1093,1736,1762]" pageId="5" pageNumber="24">TL</collectionCode>
, 
<date id="FFFF1055FFA8FFACFBDEF91EFB60F975" box="[1104,1261,1736,1762]" pageId="5" pageNumber="24" value="2006-02-18">18 Feb. 2006</date>
, Wyborn Reef, Queensland, 
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, 
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, 
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<quantity id="4CB99B70FFA8FFACFDA3F926FDE6F89D" box="[557,619,1776,1802]" metricMagnitude="1" metricUnit="m" metricValue="2.5" pageId="5" pageNumber="24" unit="m" value="25.0">25 m</quantity>
.
</specimenCount>
</paragraph>
</subSubSection>
<subSubSection id="C35B651EFFA8FFAFFF48F8C1FF63FD8D" lastPageId="6" lastPageNumber="25" pageId="5" pageNumber="24" type="diagnosis">
<paragraph id="8BFE3695FFA8FFAFFF48F8C1FF63FD8D" blockId="5.[151,1436,855,2002]" lastBlockId="6.[151,1437,152,538]" lastPageId="6" lastPageNumber="25" pageId="5" pageNumber="24">
<emphasis id="B935EA87FFA8FFACFF48F8C1FECBF8A6" bold="true" box="[198,326,1815,1841]" pageId="5" pageNumber="24">Diagnosis.</emphasis>
A comparatively small 
<taxonomicName id="4C414D16FFA8FFACFDEEF8CEFC85F8A6" box="[608,776,1816,1841]" class="Elasmobranchii" family="Scyliorhinidae" genus="Atelomycterus" kingdom="Animalia" order="Carcharhiniformes" pageId="5" pageNumber="24" phylum="Chordata" rank="genus">
<emphasis id="B935EA87FFA8FFACFDEEF8CEFC85F8A6" box="[608,776,1816,1841]" italics="true" pageId="5" pageNumber="24">Atelomycterus</emphasis>
</taxonomicName>
species (TL up to 
<quantity id="4CB99B70FFA8FFACFC6DF8CEFBB3F8A5" box="[995,1086,1816,1842]" metricMagnitude="-1" metricUnit="m" metricValue="4.9" pageId="5" pageNumber="24" unit="mm" value="490.0">490mm</quantity>
) with the following combination of characters: a relatively long snout, preoral length 4.7–5.1% TL; head length 19.3–19.9% TL, precaudal length 79.7–82.6% TL; short interdorsal space 12.2–13.9% TL; anal fin height 2.5–3.0% TL; anal fin length and anal fin base-length comparatively long 9.7–10.8 and 8.2–9.1% TL respectively. Adult claspers elongate, base broad, tapering moderately from base to tip. In adult males, clasper outer length 7.2–9.0% TL, clasper base width 20.2–22.5% of clasper outer length; clasper glans covering more than half length of entire clasper; enlarged tab on cover rhipidion. Rhipidion moderately large, concealed predominantly by large cover rhipidion and exorhipidion; psuedosiphon over half length of cover rhipidion; pseudopera large; clasper tip narrow and of moderate width, bluntly pointed. Total vertebral counts 147–153 (N=16); precaudal counts 95–99 (n=16). Colour pattern dominated by broad brownish saddles and transverse bands on tan/light brown background. Prominent dark brown to black spots and white spots on dorsal and lateral surfaces. Dark brown to black spots concentrated predominantly on saddle margins and as irregular transverse lines in inter-saddle spaces; decreasing in number post second dorsal fin. Significant numbers of small to medium sized white spots found along entire length of animal; white spots increasing in concentration towards posterior end of animal.
</paragraph>
</subSubSection>
<caption id="DF3E661DFFABFFAFFF19FBECFDEFFBE4" httpUri="https://zenodo.org/record/177425/files/figure.png" pageId="6" pageNumber="25" targetBox="[270,1316,625,1035]" targetPageId="6">
<paragraph id="8BFE3695FFABFFAFFF19FBECFDEFFBE4" blockId="6.[151,1435,1082,1140]" pageId="6" pageNumber="25">
<emphasis id="B935EA87FFABFFAFFF19FBECFE97FBC5" bold="true" box="[151,282,1082,1106]" pageId="6" pageNumber="25">FIGURE 1.</emphasis>
<taxonomicName id="4C414D16FFABFFAFFEADFBECFDBAFBC6" box="[291,567,1082,1105]" class="Elasmobranchii" family="Scyliorhinidae" genus="Atelomycterus" kingdom="Animalia" order="Carcharhiniformes" pageId="6" pageNumber="25" phylum="Chordata" rank="species" species="marnkalha" status="sp. nov.">
<emphasis id="B935EA87FFABFFAFFEADFBECFDBAFBC6" box="[291,567,1082,1105]" italics="true" pageId="6" pageNumber="25">Atelomycterus marnkalha</emphasis>
</taxonomicName>
<emphasis id="B935EA87FFABFFAFFDB1FBECFDF1FBC5" bold="true" box="[575,636,1082,1106]" pageId="6" pageNumber="25">
<taxonomicNameLabel id="A20657FCFFABFFAFFDB1FBECFDF1FBC5" box="[575,636,1082,1106]" pageId="6" pageNumber="25" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
; fixed, holotype adult male, 386 mm TL (CSIRO H 6144-01). 
<emphasis id="B935EA87FFABFFAFFAA3FBECFACDFBC5" bold="true" box="[1325,1344,1082,1106]" pageId="6" pageNumber="25">A</emphasis>
: Lateral view; 
<emphasis id="B935EA87FFABFFAFFF57FB8AFF66FBE3" bold="true" box="[217,235,1116,1140]" pageId="6" pageNumber="25">B</emphasis>
Dorsal view. Photo by S. Riddoch.
</paragraph>
</caption>
<caption id="DF3E661DFFABFFAFFF19FA62FDEFFA79" httpUri="https://zenodo.org/record/177426/files/figure.png" pageId="6" pageNumber="25" targetBox="[268,1315,1250,1378]" targetPageId="6">
<paragraph id="8BFE3695FFABFFAFFF19FA62FDEFFA79" blockId="6.[151,1436,1460,1518]" pageId="6" pageNumber="25">
<emphasis id="B935EA87FFABFFAFFF19FA62FE90FA5B" bold="true" box="[151,285,1460,1484]" pageId="6" pageNumber="25">FIGURE 2.</emphasis>
Lateral view of 
<taxonomicName id="4C414D16FFABFFAFFE53FA63FD79FA5B" box="[477,756,1461,1484]" class="Elasmobranchii" family="Scyliorhinidae" genus="Atelomycterus" kingdom="Animalia" order="Carcharhiniformes" pageId="6" pageNumber="25" phylum="Chordata" rank="species" species="marnkalha" status="sp. nov.">
<emphasis id="B935EA87FFABFFAFFE53FA63FD79FA5B" box="[477,756,1461,1484]" italics="true" pageId="6" pageNumber="25">Atelomycterus marnkalha</emphasis>
</taxonomicName>
<emphasis id="B935EA87FFABFFAFFD71FA62FCB2FA5B" bold="true" box="[767,831,1460,1484]" pageId="6" pageNumber="25">
<taxonomicNameLabel id="A20657FCFFABFFAFFD71FA62FCB2FA5B" box="[767,831,1460,1484]" pageId="6" pageNumber="25" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
; fresh specimen, paratype, adult female, 351 mm TL, (CSIRO H 6145-01). Photo by S. Riddoch.
</paragraph>
</caption>
<subSubSection id="C35B651EFFABFFA4FF48F9F7FE9FFD2D" lastPageId="13" lastPageNumber="32" pageId="6" pageNumber="25" type="description">
<paragraph id="8BFE3695FFABFFAFFF48F9F7FE9EF9F3" blockId="6.[151,1437,1569,2036]" pageId="6" pageNumber="25">
<emphasis id="B935EA87FFABFFAFFF48F9F7FED3F9AC" bold="true" box="[198,350,1569,1595]" pageId="6" pageNumber="25">Description.</emphasis>
Proportions as percentages of total length for the 
<typeStatus id="54FA8837FFABFFAFFC20F9F4FB98F9AB" box="[942,1045,1570,1596]" pageId="6" pageNumber="25" type="holotype">holotype</typeStatus>
and 
<typeStatus id="54FA8837FFABFFAFFBDEF9F4FB4DF9AB" box="[1104,1216,1570,1596]" pageId="6" pageNumber="25" type="paratype">paratypes</typeStatus>
(11) are presented in 
<tableCitation id="C6C3032EFFABFFAFFF3BF99CFE80F9F3" box="[181,269,1610,1636]" captionStart="TABLE 1" captionStartId="1.[151,239,1066,1090]" captionText="TABLE 1. Morphometric data expressed as % of TL for the Atelomycterus marnkalha n. sp. holotype, paratype size ranges and A. fasciatus comparisons." pageId="6" pageNumber="25">Table 1</tableCitation>
.
</paragraph>
<paragraph id="8BFE3695FFABFFAFFF48F9A4FE29F8C3" blockId="6.[151,1437,1569,2036]" pageId="6" pageNumber="25">Head short, length approximately equal 0.93 (0.96–1.13) to pectoral-pelvic space. Head narrow and moderately depressed; roughly trapezoidal in cross section at anterior most point of eyes. Outline of head in lateral view dorsally convex, with minor concavity in front of eyes; head with narrow parabolic outline anterior to gill openings in dorsoventral view. Preoral/snout length short 0.76 (0.67–0.84) times mouth width; broadly rounded in dorsoventral view; not or very slightly indented anterior to nostrils; snout bulbous and bluntlypointed in lateral view.</paragraph>
<paragraph id="8BFE3695FFABFFAEFF48F8B4FC6BFF25" blockId="6.[151,1437,1569,2036]" lastBlockId="7.[151,1437,152,1138]" lastPageId="7" lastPageNumber="26" pageId="6" pageNumber="25">
Eyes small and spindle shaped, eye length 6.75 (6.12–7.55) in head length and 3.11 (2.92–4.91) times eye height. Eyes dorsolateral on head, with lower edges well medial to horizontal head rim in dorsal view; subocular ridges strong but narrow; external eye openings with prominent anterior and posterior eye notches. Posterior notch not connected to spiracle. Nictitating lower eyelids of rudimentary 
<typeStatus id="54FA8837FFABFFAFFBD0F80CFB1EF863" box="[1118,1171,2010,2036]" pageId="6" pageNumber="25">type</typeStatus>
, with shallow, scaled subocular pouches and secondary lower eyelids free from upper eyelids.
</paragraph>
<paragraph id="8BFE3695FFAAFFAEFF48FF16FCAAFE35" blockId="7.[151,1437,152,1138]" pageId="7" pageNumber="26">Spiracles small, length 4.36 (3.28–7.54) in eye length, 0.28 (0.24–0.42) eye lengths behind and below posterior eye notch. First four gill openings substantially higher than fifth, height of fifth 0.84 (0.64–0.84) of first and 0.78 (0.58–0.81) of third; first gill slit height 11.53 (8.31–12.09) in head length and 0.58 (0.58–0.76) of eye length. Gill openings straight, undulated or slightly concave, gill filaments not visible from outside. Upper ends of gill openings about opposite lower edges of eyes, gill openings not elevated on dorsolateral surface of head. Gill-raker papillae absent from gill arches.</paragraph>
<paragraph id="8BFE3695FFAAFFAEFF48FE66FCA8FD05" blockId="7.[151,1437,152,1138]" pageId="7" pageNumber="26">Nostrils with small incurrent apertures lacking posterolateral keels, broadly angular nasal flaps with narrowly rounded tips, mesonarial flaps very small and positioned well laterally on anterior nasal flaps, large excurrent apertures, no posterior nasal flap. Nostrils reaching mouth, with anterior nasal flaps partially covering upper symphysis. Anterior nasal flaps very large, meeting at midline of mouth, covering excurrent apertures. Nostril width 0.97 (0.72–0.96) in internarial space, 0.78 (0.56–0.87) in eye length, 1.32 (0.85–1.33) in first gill height and 1.23 (0.78–1.24) in third gill height.</paragraph>
<paragraph id="8BFE3695FFAAFFAEFF48FD76FCA9FC3D" blockId="7.[151,1437,152,1138]" pageId="7" pageNumber="26">Mouth broadly angular, small, short, mouth width 3.17 (2.73–3.40) in head length; mouth length 2.20 (2.23–2.56) in mouth width. Lower symphysis nearly reaching upper symphysis, teeth adjacent upper symphysis exposed in ventral view. Tongue moderate-sized, flat and rounded, filling most of floor of mouth. Maxillary valve narrow, not highly papillose. No large buccal papillae in mouth, palate and floor of mouth covered with buccopharyngeal denticles, except just in front of tongue. Upper labial furrows long, reaching upper symphysis or stopping just prior to upper symphysis. Lower furrows slightly longer than upper furrows; 1.09 (1.04–1.34) times upper furrows, labial cartilages large.</paragraph>
<paragraph id="8BFE3695FFAAFFAEFF48FC6EFECEFBE5" blockId="7.[151,1437,152,1138]" pageId="7" pageNumber="26">Teeth in 67–78/64–70 rows; 3–5/4–5 series functional, with more series functional in adults than in young and in posterior tooth rows than anterolaterals. Lateroposterior teeth not arranged in diagonal files, no toothless spaces at symphysis. Teeth not strongly differentiated in upper and lower jaws; tooth row groups along upper and lower jaws including medials (M), anterolaterals (AL) and posteriors (P) weakly defined. Tooth formula (N=4) is:</paragraph>
<paragraph id="8BFE3695FFAAFFAEFE4AFB60FBE8FB45" blockId="7.[151,1436,1206,2032]" box="[452,1125,1206,1234]" pageId="7" pageNumber="26">Left P13-20 AL18-20 M3 AL17-20 P14-20 right</paragraph>
<paragraph id="8BFE3695FFAAFFAEFD83FB0DFB90FB60" blockId="7.[151,1436,1206,2032]" box="[525,1053,1243,1271]" pageId="7" pageNumber="26">P14-19 AL14-18 M1 AL14-20 P14-20</paragraph>
<paragraph id="8BFE3695FFAAFFAEFC8AFAD6FC92FA8B" blockId="7.[151,1436,1206,2032]" box="[772,799,1280,1308]" pageId="7" pageNumber="26">or</paragraph>
<paragraph id="8BFE3695FFAAFFAEFD3CFAF2FCFAFAD7" blockId="7.[151,1436,1206,2032]" box="[690,887,1316,1344]" pageId="7" pageNumber="26">
<date id="FFFF1055FFAAFFAEFD3CFAF2FC90FAD7" box="[690,797,1316,1344]" pageId="7" pageNumber="26" value="1938-03-31">31-38 3</date>
33-38
</paragraph>
<paragraph id="8BFE3695FFAAFFAEFD3CFA9FFCF5FAF2" blockId="7.[151,1436,1206,2032]" box="[690,888,1353,1381]" pageId="7" pageNumber="26">
31- 
<date id="FFFF1055FFAAFFAEFD55FA9FFD70FAF2" box="[731,765,1353,1381]" pageId="7" pageNumber="26" value="1934-01-31">34</date>
1 31-35
</paragraph>
<paragraph id="8BFE3695FFAAFFAEFF48FAA8FC1CF9F7" blockId="7.[151,1436,1206,2032]" pageId="7" pageNumber="26">
Tooth formation tricuspid (
<figureCitation id="137A2A10FFAAFFAEFD89FAA8FDDCFA0F" box="[519,593,1406,1432]" captionStart="FIGURE 4" captionStartId="9.[151,255,1795,1819]" captionTargetBox="[304,1297,710,1736]" captionTargetId="figure@9.[285,1303,674,1737]" captionTargetPageId="9" captionText="FIGURE 4. Atelomycterus marnkalha n. sp.; scanning electron micrographs, 459 mm TL male A – C upper teeth; D – H lower teeth. A – B upper anterolateral teeth from near symphysis in A, labial; and B, lingual views; C, posterior tooth in lingual view; D – F, lower anterolateral teeth from near symphysis in D, labial; E, lingual; and F, dorso-lateral views; G – H, posterior teeth in G, labial and H, lingual views. Photos by K. Sewell." httpUri="https://zenodo.org/record/177428/files/figure.png" pageId="7" pageNumber="26">Fig. 4</figureCitation>
); central cusp strong, well-developed and substantially longer than lateral cusps. Cusps strongly erect to semi-oblique. Upper anterolateral teeth smaller than those adjacent; cusplets of upper anterolateral teeth with well-developed transverse ridges; longer than lower. Medials smaller and weak differentiated in comparison with anterolaterals. Posterior teeth forming several rows in both jaws; distinctly smaller than anterolateral and median teeth; with low weak cusps. Sexual heterodonty absent; tooth morphology not particularly enlarged or modified in adult males.
</paragraph>
<paragraph id="8BFE3695FFAAFFA1FF48F9B8FC9FFF25" blockId="7.[151,1436,1206,2032]" lastBlockId="8.[151,1436,152,338]" lastPageId="8" lastPageNumber="27" pageId="7" pageNumber="26">
Lateral trunk denticles below first dorsal fin moderately elongate, with weak to moderately tricuspid teardrop-shaped crowns approximately 1.5 times as long as wide (
<figureCitation id="137A2A10FFAAFFAEFC1FF940FC51F927" box="[913,988,1686,1712]" captionStart="FIGURE 5" captionStartId="10.[151,255,1595,1619]" captionTargetBox="[384,1203,194,1551]" captionTargetId="figure@10.[384,1203,194,1571]" captionTargetPageId="10" captionText="FIGURE 5. Atelomycterus marnkalha n. sp.; scanning electron micrographs of lateral trunk denticles in three-quarter anteriorlateral view, 459 mm TL male at A, 100 Μm, and B, 50 Μm. Photo by K. Sewell." httpUri="https://zenodo.org/record/177429/files/figure.png" pageId="7" pageNumber="26">Fig. 5</figureCitation>
). Denticle crown with pair of strong medial ridges extending along almost entire length of crown and onto a strong, medial cusp; lateral cusps short and obtuse when compared to medial cusp; numerous shallow depressions present proximally, covering approximately one third to half of denticle crown, occasionally extending onto lateral cusps; shallow depressions absent between medial ridges and on medial cusp. Denticle crowns closely spaced and well-imbricated. Pectoral fins broad and triangular in shape with rounded apices, not falcate, with broadly convex anterior margins 1.08 (1.00–1.16) times its length; weakly convex to broadly rounded posterior margins, free rear tips inner margins and narrow bases. Total area of pectoral fins less than twice the area of first dorsal fin. Origins of pectoral fins under interspace between third and forth gill openings. Apex of pectoral fin slightly anterior to its free tip when fin is elevated and appressed to body.
</paragraph>
<paragraph id="8BFE3695FFA5FFA1FF48FF16FC10FEC5" blockId="8.[151,1436,152,338]" pageId="8" pageNumber="27">Pelvic fins broadly triangular; pelvic fin anterior margins nearly straight or very weakly convex; 0.74 (0.63–0.78) pectoral fin anterior margins; rounded apically, with posterior margins convex or nearly straight, free rear tips narrowly rounded and not attenuated, inner margins straight or slightly convex, not fused over claspers of adult males. Pelvic area nearly 2–3 times anal fin area.</paragraph>
<caption id="DF3E661DFFA5FFA1FF19FB99FE67FB1E" httpUri="https://zenodo.org/record/177427/files/figure.png" pageId="8" pageNumber="27" targetBox="[286,1322,411,1068]" targetPageId="8">
<paragraph id="8BFE3695FFA5FFA1FF19FB99FE67FB1E" blockId="8.[151,1436,1103,1161]" pageId="8" pageNumber="27">
<emphasis id="B935EA87FFA5FFA1FF19FB99FE97FBF0" bold="true" box="[151,282,1103,1127]" pageId="8" pageNumber="27">FIGURE 3.</emphasis>
Ventral head view of 
<taxonomicName id="4C414D16FFA5FFA1FD82FB99FCADFBF1" box="[524,800,1103,1126]" class="Elasmobranchii" family="Scyliorhinidae" genus="Atelomycterus" kingdom="Animalia" order="Carcharhiniformes" pageId="8" pageNumber="27" phylum="Chordata" rank="species" species="marnkalha" status="sp. nov.">
<emphasis id="B935EA87FFA5FFA1FD82FB99FCADFBF1" box="[524,800,1103,1126]" italics="true" pageId="8" pageNumber="27">Atelomycterus marnkalha</emphasis>
</taxonomicName>
<emphasis id="B935EA87FFA5FFA1FCA6FB99FCE8FBF0" bold="true" box="[808,869,1103,1127]" pageId="8" pageNumber="27">
<taxonomicNameLabel id="A20657FCFFA5FFA1FCA6FB99FCE8FBF0" box="[808,869,1103,1127]" pageId="8" pageNumber="27" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
; fixed, holotype adult male, 386 mm TL (CSIRO H 6144-01). Photo by S. Riddoch.
</paragraph>
</caption>
<paragraph id="8BFE3695FFA5FFA1FF48FB6AFA9AF8C1" blockId="8.[151,1437,1212,1998]" pageId="8" pageNumber="27">
Claspers relatively long, slightly convex and tapering on lateral edge, with a slightly undulating clasper glan (
<figureCitation id="137A2A10FFA5FFA1FF55FB32FEA9FB69" box="[219,292,1252,1278]" captionStart="FIGURE 6" captionStartId="11.[151,255,937,961]" captionTargetBox="[270,1322,202,898]" captionTargetId="figure@11.[264,1323,194,913]" captionTargetPageId="11" captionText="FIGURE 6. Claspers of Atelomycterus marnkalha n. sp.; 459 mm TL adult male (QM I 38089). A: Glans not dilated; B: Glans spread. Abbreviations: AP: Apopyle; CLD: Clasper denticles; CRH: Cover Rhipidion; ERH: Exorhipidion; FCG: Fused clasper groove; FRT: Free tab on cover rhipidion; P- 2: Pelvic fin; PSP: pseudopera; PSS: Pseudosiphon; RH: Rhipidion. Illustration by I. P. Jacobsen." httpUri="https://zenodo.org/record/177430/files/figure.png" pageId="8" pageNumber="27">Fig. 6</figureCitation>
). Claspers of adult males extending well behind pelvic fin free rear tips, distance approximately 1.7 times pelvic fin inner margin, posterior most point of distal apex falling in front of anal fin origin by 0.77– 0.93 times anal fin base. Clasper glans moderately long, length approximately half to two thirds of outer clasper margin; distal apex blunt and without specialised clasper hooks. Dorsolateral and ventral surfaces covered with small clasper denticles, dorsomedial and posteromedial surfaces of glans (including rhipidion) and lateral strip adjacent to clasper groove naked; clasper denticles much longer than broad with anteriorly directed cusps; narrow band of low semi-upright tricuspidate denticles near tip of glans. Exorhipidion strongly differentiated, originating opposite the last third of cover rhipidion. Pseudopera present, situated below anterior end of exorhipidion and about opposite posterior end of cover rhipidion. Rhipidion present and greatly enlarged, forming a flat convex-edged blade-like structure extending over most of clasper glan length; posterior end below apex of exorhipidion. Cover rhipidion very large, formed as a distally tapering wedge with large, lobate anterior tab, extending from apopyle to apex of exorhipidion; angle between free tab and base of cover rhipidion acute; posterior end proximal to exorhipidion and rear end of the rhipidion. Anterior margin of cover rhipidion slightly convex, without obvious notch (
<figureCitation id="137A2A10FFA5FFA1FCC1F93AFC11F891" box="[847,924,1772,1798]" captionStart="FIGURE 6" captionStartId="11.[151,255,937,961]" captionTargetBox="[270,1322,202,898]" captionTargetId="figure@11.[264,1323,194,913]" captionTargetPageId="11" captionText="FIGURE 6. Claspers of Atelomycterus marnkalha n. sp.; 459 mm TL adult male (QM I 38089). A: Glans not dilated; B: Glans spread. Abbreviations: AP: Apopyle; CLD: Clasper denticles; CRH: Cover Rhipidion; ERH: Exorhipidion; FCG: Fused clasper groove; FRT: Free tab on cover rhipidion; P- 2: Pelvic fin; PSP: pseudopera; PSS: Pseudosiphon; RH: Rhipidion. Illustration by I. P. Jacobsen." httpUri="https://zenodo.org/record/177430/files/figure.png" pageId="8" pageNumber="27">Fig. 6</figureCitation>
). Pseudosiphon long, narrow and slitlike, extending opposite most of base of cover rhipidion; posterior end terminating anterior to exorhipidion apex. Apopyle and hypopyle connected by long clasper groove, dorsal margins fused over clasper canal.
</paragraph>
<paragraph id="8BFE3695FFA5FFA0FF48F8B2FAACFDFD" blockId="8.[151,1437,1212,1998]" lastBlockId="9.[151,1437,152,618]" lastPageId="9" lastPageNumber="28" pageId="8" pageNumber="27">
Clasper skeleton with all elements present (
<figureCitation id="137A2A10FFA5FFA1FD47F8B2FC9CF8E9" box="[713,785,1892,1918]" captionStart="FIGURE 7" captionStartId="12.[151,255,1433,1457]" captionTargetBox="[264,1322,474,1405]" captionTargetId="figure@12.[264,1323,474,1409]" captionTargetPageId="12" captionText="FIGURE 7. Claspers of Atelomycterus marnkalha n. sp.; 459 mm TL adult male (QM I 38089). A: Ventral view; B: Dorsal vies; C: dorsal view of glans, terminal cartilage spread. Abbreviations: AX: Axial cartilage; β: Beta cartilage; B: Basypterygium; B 1: Intermediate segment; G: End style; RD: Dorsal marginal cartilage; RD 2: Accessory dorsal marginal cartilage; RV: Ventral marginal cartilage; T 3: accessory terminal cartilage; TD: Dorsal terminal cartilage; TD 2: dorsal terminal 2 cartilage; TV: ventral terminal cartilage; TV 2: Ventral terminal 2 cartilage; TV 3: ventral terminal 3 cartilage. Illustration by I. P. Jacobsen." httpUri="https://zenodo.org/record/177431/files/figure.png" pageId="8" pageNumber="27">Fig. 7</figureCitation>
). Axial cartilage connected proximally to pelvic basipterygium by single, short intermediate segment (B1) and moderately long, posteriorly tapering, dorsal beta cartilage (β). Beta cartilage with rounded anterior and posterior margins and possessing a small, central, proximal protuberance. Clasper shaft formed from axial cartilage and tightly rolled dorsal and ventral marginal cartilages. Marginal cartilages anteriorly slender and tapering; expanding posteriorly along axial cartilage. Dorsal terminal (TD) of clasper glan large, narrow and curved; ventral terminal (TV) distally expanded and spear-shaped; separated along their proximomesial by narrow, cylindrical end-style (terminal extension of axial cartilage; short free posterior ends of terminal cartilages separated by narrow gap. Accessory dorsal marginal cartilage (RD2) long, wedge shaped, tapering distally, with separate, short distal segment (RD3). Dorsal terminal 2 cartilage (TD2) large, elongate, tapering distally, extending inside rhipidion, along almost entire clasper glans from dorsal margin to end-style tip. Cartilages of exorhipidion include an elongate, anterior tapered ventral terminal 2 (TV2) cartilage and a ventral terminal 3 (TV3) cartilage tapering posteriorly; TV2 cartilage articulating with anterolateral end of TV3 cartilage and extending anteriorly to partially cover distal end of ventral margin. Short, accessory terminal cartilage (T3) present under anterior half of TD2 cartilage, partially covered by terminal extension of ventral margin; T3 well anterior to TV3-TV articulation.
</paragraph>
<caption id="DF3E661DFFA4FFA0FF19F8D5FC10F814" httpUri="https://zenodo.org/record/177428/files/figure.png" pageId="9" pageNumber="28" targetBox="[304,1297,710,1736]" targetPageId="9">
<paragraph id="8BFE3695FFA4FFA0FF19F8D5FC10F814" blockId="9.[151,1436,1795,1923]" pageId="9" pageNumber="28">
<emphasis id="B935EA87FFA4FFA0FF19F8D5FE97F88C" bold="true" box="[151,282,1795,1819]" pageId="9" pageNumber="28">FIGURE 4.</emphasis>
<taxonomicName id="4C414D16FFA4FFA0FEACF8D2FDB8F88C" box="[290,565,1796,1819]" class="Elasmobranchii" family="Scyliorhinidae" genus="Atelomycterus" kingdom="Animalia" order="Carcharhiniformes" pageId="9" pageNumber="28" phylum="Chordata" rank="species" species="marnkalha" status="sp. nov.">
<emphasis id="B935EA87FFA4FFA0FEACF8D2FDB8F88C" box="[290,565,1796,1819]" italics="true" pageId="9" pageNumber="28">Atelomycterus marnkalha</emphasis>
</taxonomicName>
<emphasis id="B935EA87FFA4FFA0FDB3F8D5FDF7F88C" bold="true" box="[573,634,1795,1819]" pageId="9" pageNumber="28">
<taxonomicNameLabel id="A20657FCFFA4FFA0FDB3F8D5FDF7F88C" box="[573,634,1795,1819]" pageId="9" pageNumber="28" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
; scanning electron micrographs, 459 mm TL male A–C upper teeth; D–H lower teeth. A–B upper anterolateral teeth from near symphysis in A, labial; and B, lingual views; C, posterior tooth in lingual view; D–F, lower anterolateral teeth from near symphysis in D, labial; E, lingual; and F, dorso-lateral views; G– H, posterior teeth in G, labial and H, lingual views. Photos by K. Sewell.
</paragraph>
</caption>
<caption id="DF3E661DFFA7FFA3FF19F9EDFBCEF9E2" httpUri="https://zenodo.org/record/177429/files/figure.png" pageId="10" pageNumber="29" targetBox="[384,1203,194,1551]" targetPageId="10">
<paragraph id="8BFE3695FFA7FFA3FF19F9EDFBCEF9E2" blockId="10.[151,1436,1595,1656]" pageId="10" pageNumber="29">
<emphasis id="B935EA87FFA7FFA3FF19F9EDFE91F9C4" bold="true" box="[151,284,1595,1619]" pageId="10" pageNumber="29">FIGURE 5.</emphasis>
<taxonomicName id="4C414D16FFA7FFA3FEAAF9EAFDB5F9C4" box="[292,568,1596,1619]" class="Elasmobranchii" family="Scyliorhinidae" genus="Atelomycterus" kingdom="Animalia" order="Carcharhiniformes" pageId="10" pageNumber="29" phylum="Chordata" rank="species" species="marnkalha" status="sp. nov.">
<emphasis id="B935EA87FFA7FFA3FEAAF9EAFDB5F9C4" box="[292,568,1596,1619]" italics="true" pageId="10" pageNumber="29">Atelomycterus marnkalha</emphasis>
</taxonomicName>
<emphasis id="B935EA87FFA7FFA3FDCFF9EDFDF2F9C4" bold="true" box="[577,639,1595,1619]" pageId="10" pageNumber="29">
<taxonomicNameLabel id="A20657FCFFA7FFA3FDCFF9EDFDF2F9C4" box="[577,639,1595,1619]" pageId="10" pageNumber="29" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
; scanning electron micrographs of lateral trunk denticles in three-quarter anteriorlateral view, 459 mm TL male at A, 100 Μm, and B, 50 Μm. Photo by K. Sewell.
</paragraph>
</caption>
<paragraph id="8BFE3695FFA7FFA3FF48F97FFE29F84C" blockId="10.[151,1437,1705,2011]" pageId="10" pageNumber="29">First dorsal fin high, apically narrow and not falcate, with nearly straight or weakly convex anterior margin, narrowly rounded apex, weakly concave posterior margin, angular free rear tip, inner margin straight. First dorsal-fin origin at midpoint approximate of pelvic-fin bases; midpoint of base slightly anterior or opposite pelvic-fin free rear tips, insertion closer to pelvic-fin insertions than anal-fin origin; free rear tip 1.67 (1.68–2.48) times length of inner margin anterior to anal-fin origin. Fin insertion anterior to fin apex; first dorsal-fin base 1.58 (1.32–1.62) in interdorsal space, 2.09 (1.88–2.35) in dorsal caudal-fin margin, first dorsal-fin height 1.58 (1.31–1.86) in its base, first dorsal-fin inner margin 1.66 (1.64–2.06) in first dorsal-fin height, 2.63 (2.70–3.40) in its base.</paragraph>
<caption id="DF3E661DFFA6FFA2FF19FC7FFD91FBBF" httpUri="https://zenodo.org/record/177430/files/figure.png" pageId="11" pageNumber="30" targetBox="[270,1322,202,898]" targetPageId="11">
<paragraph id="8BFE3695FFA6FFA2FF19FC7FFD91FBBF" blockId="11.[151,1437,937,1064]" pageId="11" pageNumber="30">
<emphasis id="B935EA87FFA6FFA2FF19FC7FFE94FC56" bold="true" box="[151,281,937,961]" pageId="11" pageNumber="30">FIGURE 6.</emphasis>
Claspers of 
<taxonomicName id="4C414D16FFA6FFA2FE10FC7FFD3DFC57" box="[414,688,937,960]" class="Elasmobranchii" family="Scyliorhinidae" genus="Atelomycterus" kingdom="Animalia" order="Carcharhiniformes" pageId="11" pageNumber="30" phylum="Chordata" rank="species" species="marnkalha" status="sp. nov.">
<emphasis id="B935EA87FFA6FFA2FE10FC7FFD3DFC57" box="[414,688,937,960]" italics="true" pageId="11" pageNumber="30">Atelomycterus marnkalha</emphasis>
</taxonomicName>
<emphasis id="B935EA87FFA6FFA2FD39FC7FFD7EFC56" bold="true" box="[695,755,937,961]" pageId="11" pageNumber="30">
<taxonomicNameLabel id="A20657FCFFA6FFA2FD39FC7FFD7EFC56" box="[695,755,937,961]" pageId="11" pageNumber="30" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
; 459 mm TL adult male (QM I 38089). A: Glans not dilated; B: Glans spread. Abbreviations: AP: Apopyle; CLD: Clasper denticles; CRH: Cover Rhipidion; ERH: Exorhipidion; FCG: Fused clasper groove; FRT: Free tab on cover rhipidion; P-2: Pelvic fin; PSP: pseudopera; PSS: Pseudosiphon; RH: Rhipidion. Illustration by I. P. Jacobsen.
</paragraph>
</caption>
<paragraph id="8BFE3695FFA6FFA2FF48FB8AFEC4FA49" blockId="11.[151,1437,1116,2022]" pageId="11" pageNumber="30">Second dorsal fin similar to first dorsal fin. Second dorsal fin high, apically narrow, not falcate, equal to first dorsal fin area or marginally subequal; second dorsal-fin height 0.96 (0.91–1.21) of first dorsal-fin height, base 0.95 (0.85–1.08) of first dorsal-fin base. Second dorsal fin with slightly convex anterior margin, narrowly rounded apex, nearly straight posterior margin, bluntly pointed free rear tip, and weakly concave or straight inner margin. Second dorsal-fin origin opposite or slightly in front of anal fin midbase, insertion well behind anal-fin free rear tip; free rear tip in front of upper caudal-fin origin 0.99 (0.54–2.00) times the inner margin. Posterior margin slanting posteroventrally from apex, insertion below or slightly in front of dorsal apex. Second dorsal-fin base 0.77 (0.65–0.91) in dorsal-caudal space, second dorsal-fin height 1.74 (1.49– 4.98) in its base, second dorsal-fin inner margin 1.74 (1.85–2.03) in its height and 3.04 (2.91–3.89) in seconddorsal fin base.</paragraph>
<paragraph id="8BFE3695FFA6FFA2FF48FA3AFB23F889" blockId="11.[151,1437,1116,2022]" pageId="11" pageNumber="30">Anal fin low apically broad and semifalcate, much smaller than second dorsal fin, anal-fin height 0.49 (0.52–0.62) in second dorsal-fin height and anal-fin base 0.91 (0.88–1.03) times second dorsal-fin base. Analfin anterior margin moderately convex to nearly straight, apex broadly rounded, posterior margin nearly straight, inner margin nearly straight. Anal-fin base without preanal ridges, anal-fin origin 1.80 (1.44–1.83) times its base length behind pelvic-fin insertions, free rear tips 3.72 (3.05–6.02) times anal fin inner margin length anterior to lower caudal fin origin. Anal-fin posterior margin slanting posterodorsally, anal-fin insertion posterior to apex. Anal-fin base 1.14 (0.86–1.16) in anal-caudal space, anal-fin height 3.26 (2.72–3.25) in its base; anal-fin inner margin 1.11 (1.33–2.03) in its height and 3.62 (3.69–6.07) in its base.</paragraph>
<paragraph id="8BFE3695FFA6FFA2FF48F8FAFA10F871" blockId="11.[151,1437,1116,2022]" pageId="11" pageNumber="30">Caudal fin narrow, elongate and asymmetrical, with large terminal lobe, ventral lobe not developed; dorsal margin slightly undulating, 4.63 (4.29–4.67) in precaudal length. Preventral caudal-fin margin 2.34 (2.00– 2.54) in dorsal caudal-fin margin, terminal lobe 4.43 (3.48–5.11) in dorsal caudal-fin margin, subterminal margin 1.45 (1.05–1.67) in terminal margin. Dorsal caudal-fin margin without lateral undulations; proximally and distally straight, basally concave and apically straight, tip of ventral caudal fin lobe bluntly rounded.</paragraph>
<paragraph id="8BFE3695FFA1FFA5FF19FF4EFD18FEBD" blockId="12.[151,1436,152,418]" pageId="12" pageNumber="31">Postventral caudal-fin margin not differentiated into upper and lower parts, margin straight to slightly concave. Subterminal notch a narrow, deep slot, subterminal margin straight to convex; terminal margin concave and sometimes notched, edges of lobe bluntly angular, tip of tail broadly rounded; terminal margin length 3.81 (3.66–5.47) times dorsal caudal-fin margin.</paragraph>
<paragraph id="8BFE3695FFA1FFA5FF48FEEEFAF4FE35" blockId="12.[151,1436,152,418]" pageId="12" pageNumber="31">
Vertebral counts, ratios and statistics are given in 
<tableCitation id="C6C3032EFFA1FFA5FC80FEEEFCE5FEC5" box="[782,872,312,338]" captionStart="TABLE 2" captionStartId="4.[151,239,151,175]" captionTargetBox="[159,1402,242,820]" captionTargetPageId="4" captionText="TABLE 2. Vertebral count comparisons between Atelomycterus marnkalha n. sp. and A. fasciatus. * Vertebral counts and percentages compiled from Compagno and Stevens’ (1993) original description." httpUri="http://table.plazi.org/id/DF3E661DFFA9FFADFF19FF41FC62FF46" pageId="12" pageNumber="31" tableUuid="DF3E661DFFA9FFADFF19FF41FC62FF46">Table 2</tableCitation>
. Transition between monospondylous (MP) and diplospondylous (DP) centra on average 6 (5–7) centra behind front of pelvic girdle. Monospondylous – Diplospondylous transition hardly enlarged, not forming ‘stutter zone’ of alternating long and short centra.
</paragraph>
<caption id="DF3E661DFFA1FFA5FF19FA4FFDB9F9C9" httpUri="https://zenodo.org/record/177431/files/figure.png" pageId="12" pageNumber="31" targetBox="[264,1322,474,1405]" targetPageId="12">
<paragraph id="8BFE3695FFA1FFA5FF19FA4FFDB9F9C9" blockId="12.[151,1436,1433,1630]" pageId="12" pageNumber="31">
<emphasis id="B935EA87FFA1FFA5FF19FA4FFE96FA26" bold="true" box="[151,283,1433,1457]" pageId="12" pageNumber="31">FIGURE 7.</emphasis>
Claspers of 
<taxonomicName id="4C414D16FFA1FFA5FE26FA4CFD31FA26" box="[424,700,1434,1457]" class="Elasmobranchii" family="Scyliorhinidae" genus="Atelomycterus" kingdom="Animalia" order="Carcharhiniformes" pageId="12" pageNumber="31" phylum="Chordata" rank="species" species="marnkalha" status="sp. nov.">
<emphasis id="B935EA87FFA1FFA5FE26FA4CFD31FA26" box="[424,700,1434,1457]" italics="true" pageId="12" pageNumber="31">Atelomycterus marnkalha</emphasis>
</taxonomicName>
<emphasis id="B935EA87FFA1FFA5FD48FA4FFC89FA26" bold="true" box="[710,772,1433,1457]" pageId="12" pageNumber="31">
<taxonomicNameLabel id="A20657FCFFA1FFA5FD48FA4FFC89FA26" box="[710,772,1433,1457]" pageId="12" pageNumber="31" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
; 459 mm TL adult male (QM I 38089). A: Ventral view; B: Dorsal vies; C: dorsal view of glans, terminal cartilage spread. Abbreviations: AX: Axial cartilage; β: Beta cartilage; B: Basypterygium; B1: Intermediate segment; G: End style; RD: Dorsal marginal cartilage; RD2: Accessory dorsal marginal cartilage; RV: Ventral marginal cartilage; T3: accessory terminal cartilage; TD: Dorsal terminal cartilage; TD2: dorsal terminal 2 cartilage; TV: ventral terminal cartilage; TV2: Ventral terminal 2 cartilage; TV3: ventral terminal 3 cartilage. Illustration by I. P. Jacobsen.
</paragraph>
</caption>
<paragraph id="8BFE3695FFA1FFA5FF48F947FA10F87B" blockId="12.[151,1437,1681,2028]" pageId="12" pageNumber="31">
<emphasis id="B935EA87FFA1FFA5FF48F947FEAEF93C" bold="true" box="[198,291,1681,1707]" pageId="12" pageNumber="31">Colour.</emphasis>
In alcohol, colour light brownish above, light brown to tan colouration on ventral surfaces of head, trunk, precaudal tail and fins, bands and saddles dark brown. Broad, brown saddle-marks on head above eyes, extending ventrally to include eyes; over gills, extending ventrally to incorporate first three gill slits; over, but not including, pectoral-fin free rear tips; on abdomen well anterior of pelvic-fin origins; over pelvicfin insertions with anterior margin of saddle posterior to first dorsal-fin origin, including section of first dorsal-fin anterior margin; between dorsal-fin bases; over and including second dorsal fin, extending ventrally to include inner margin of anal fins (
<figureCitation id="137A2A10FFA1FFA5FDBAF854FD0AF80B" box="[564,647,1922,1948]" captionStart="FIGURE 1" captionStartId="6.[151,255,1082,1106]" captionTargetBox="[270,1316,625,1035]" captionTargetId="figure@6.[264,1323,597,1055]" captionTargetPageId="6" captionText="FIGURE 1. Atelomycterus marnkalha n. sp.; fixed, holotype adult male, 386 mm TL (CSIRO H 6144 - 01). A: Lateral view; B Dorsal view. Photo by S. Riddoch." httpUri="https://zenodo.org/record/177425/files/figure.png" pageId="12" pageNumber="31">Figs. 1</figureCitation>
and 
<figureCitation id="137A2A10FFA1FFA5FD4BF854FD5BF80B" box="[709,726,1922,1948]" captionStart="FIGURE 2" captionStartId="6.[151,255,1460,1484]" captionTargetBox="[268,1315,1250,1378]" captionTargetId="figure@6.[264,1323,1194,1437]" captionTargetPageId="6" captionText="FIGURE 2. Lateral view of Atelomycterus marnkalha n. sp.; fresh specimen, paratype, adult female, 351 mm TL, (CSIRO H 6145 - 01). Photo by S. Riddoch." httpUri="https://zenodo.org/record/177426/files/figure.png" pageId="12" pageNumber="31">2</figureCitation>
). Thinner saddle marks present in dorsal-caudal space/anal caudal space; continuing as two bands on caudal fin and one on caudal fin terminal lobe. Ventral limits of abdomen saddle-marks approximately 75% of trunk depth; caudal fin saddles continuing to ventral limits.
</paragraph>
<paragraph id="8BFE3695FFA0FFA4FF19FF4EFECCFE95" blockId="13.[151,1437,152,698]" pageId="13" pageNumber="32">Secondary, thinner saddles of light brown colouration may occur over first dorsal-fin origin extending anteriorly and over second dorsal-fin origin extending anteriorly. Development and prominence of secondary saddles may vary.</paragraph>
<paragraph id="8BFE3695FFA0FFA4FF48FEC6FE9FFD2D" blockId="13.[151,1437,152,698]" pageId="13" pageNumber="32">Small dark brown to black spots the size of eye width or smaller, loosely arranged in lines around saddlemark margins and centre of inter-saddle spaces (strongly demarcated in dorsal view) of head and trunk, irregularly arranged on pectoral fins and dorsal fins. Brown/black spots on trunk region decreasing in number approaching second dorsal fin, black spots absent from body surfaces post-second dorsal fin free rear tip. First and second dorsal fin apices with white tips, posterior tip of caudal fin terminal lobe with small white tip, occasionally extending along terminal margin. Small white spots absent on head anterior to third gill arch, pectoral fins, pelvic fins and on dorsal surface; excluding caudal fin dorsal margin. Small white spots present on lateral surfaces post-third gill arch, irregularly arranged on the trunk, dorsal fins and caudal fin on saddlemarks and inter-saddle spaces. White spots increasing in intensity and number on posterior surfaces of animal; most pronounced on caudal-tail. Overall, white spots slightly smaller than brown/black spots and more numerous.</paragraph>
</subSubSection>
<caption id="DF3E661DFFA0FFA4FF19F939FE08F8F1" httpUri="https://zenodo.org/record/177432/files/figure.png" pageId="13" pageNumber="32" targetBox="[196,1255,749,1758]" targetPageId="13">
<paragraph id="8BFE3695FFA0FFA4FF19F939FE08F8F1" blockId="13.[151,1436,1775,1895]" pageId="13" pageNumber="32">
<emphasis id="B935EA87FFA0FFA4FF19F939FE94F890" bold="true" box="[151,281,1775,1799]" pageId="13" pageNumber="32">FIGURE 8.</emphasis>
<taxonomicName id="4C414D16FFA0FFA4FEAFF926FDB9F890" box="[289,564,1776,1799]" class="Elasmobranchii" family="Scyliorhinidae" genus="Atelomycterus" kingdom="Animalia" order="Carcharhiniformes" pageId="13" pageNumber="32" phylum="Chordata" rank="species" species="marnkalha" status="sp. nov.">
<emphasis id="B935EA87FFA0FFA4FEAFF926FDB9F890" box="[289,564,1776,1799]" italics="true" pageId="13" pageNumber="32">Atelomycterus marnkalha</emphasis>
</taxonomicName>
<emphasis id="B935EA87FFA0FFA4FDB2F939FDF5F890" bold="true" box="[572,632,1775,1799]" pageId="13" pageNumber="32">
<taxonomicNameLabel id="A20657FCFFA0FFA4FDB2F939FDF5F890" box="[572,632,1775,1799]" pageId="13" pageNumber="32" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
distribution. Inset: sample range of 
<taxonomicName id="4C414D16FFA0FFA4FC73F926FB03F890" box="[1021,1166,1776,1799]" class="Elasmobranchii" family="Scyliorhinidae" genus="Atelomycterus" kingdom="Animalia" order="Carcharhiniformes" pageId="13" pageNumber="32" phylum="Chordata" rank="species" species="marnkalha">
<emphasis id="B935EA87FFA0FFA4FC73F926FB03F890" box="[1021,1166,1776,1799]" italics="true" pageId="13" pageNumber="32">A. marnkalha</emphasis>
</taxonomicName>
collected from the Great Barrier Reef and Torres Strait region circumscribed by square. Black dots identified with A, B, C represent easternmost specimens of 
<taxonomicName id="4C414D16FFA0FFA4FEA7F8E6FE34F8D0" box="[297,441,1840,1863]" class="Elasmobranchii" family="Scyliorhinidae" genus="Atelomycterus" kingdom="Animalia" order="Carcharhiniformes" pageId="13" pageNumber="32" phylum="Chordata" rank="species" species="marnkalha">
<emphasis id="B935EA87FFA0FFA4FEA7F8E6FE34F8D0" box="[297,441,1840,1863]" italics="true" pageId="13" pageNumber="32">A. marnkalha</emphasis>
</taxonomicName>
which were originally identified as 
<taxonomicName id="4C414D16FFA0FFA4FCB6F8E6FC38F8D0" box="[824,949,1840,1863]" class="Elasmobranchii" family="Scyliorhinidae" genus="Atelomycterus" kingdom="Animalia" order="Carcharhiniformes" pageId="13" pageNumber="32" phylum="Chordata" rank="species" species="fasciatus">
<emphasis id="B935EA87FFA0FFA4FCB6F8E6FC38F8D0" box="[824,949,1840,1863]" italics="true" pageId="13" pageNumber="32">A. fasciatus</emphasis>
</taxonomicName>
; 
<emphasis id="B935EA87FFA0FFA4FC4CF8F9FC51F8D0" bold="true" box="[962,988,1839,1863]" pageId="13" pageNumber="32">A.</emphasis>
AMS I 21842-001; 
<emphasis id="B935EA87FFA0FFA4FB3DF8F9FB46F8D0" bold="true" box="[1203,1227,1839,1863]" pageId="13" pageNumber="32">B.</emphasis>
AMS I 15557-003; 
<emphasis id="B935EA87FFA0FFA4FF19F899FF3DF8F0" bold="true" box="[151,176,1871,1895]" pageId="13" pageNumber="32">C.</emphasis>
CSIRO H 1118-01.
</paragraph>
</caption>
<subSubSection id="C35B651EFFA0FFA7FF48F864FB5CFCBC" lastPageId="14" lastPageNumber="33" pageId="13" pageNumber="32" type="distribution">
<paragraph id="8BFE3695FFA0FFA7FF48F864FE6EFEBF" blockId="13.[151,1436,1970,2036]" lastBlockId="14.[151,1437,151,2036]" lastPageId="14" lastPageNumber="33" pageId="13" pageNumber="32">
<emphasis id="B935EA87FFA0FFA4FF48F864FE73F85B" bold="true" box="[198,510,1970,1996]" pageId="13" pageNumber="32">Distribution and habitat.</emphasis>
The distribution of 
<taxonomicName id="4C414D16FFA0FFA4FD66F864FC05F85C" box="[744,904,1970,1995]" class="Elasmobranchii" family="Scyliorhinidae" genus="Atelomycterus" kingdom="Animalia" order="Carcharhiniformes" pageId="13" pageNumber="32" phylum="Chordata" rank="species" species="marnkalha">
<emphasis id="B935EA87FFA0FFA4FD66F864FC05F85C" box="[744,904,1970,1995]" italics="true" pageId="13" pageNumber="32">A. marnkalha</emphasis>
</taxonomicName>
specimens collected from the present survey, extends from Gladstone in Central East Queensland (22°S, 150°E) to the southern reaches of 
<collectingCountry id="F3567605FFA0FFA7FA9CF80CFF61FF25" lastPageId="14" lastPageNumber="33" name="Papua New Guinea" pageId="13" pageNumber="32">Papua New Guinea</collectingCountry>
(9° S, 143° E) (
<figureCitation id="137A2A10FFA3FFA7FE2BFF4EFE66FF25" box="[421,491,152,178]" captionStart="FIGURE 8" captionStartId="13.[151,255,1775,1799]" captionTargetBox="[196,1255,749,1758]" captionTargetId="figure@13.[195,1393,746,1758]" captionTargetPageId="13" captionText="FIGURE 8. Atelomycterus marnkalha n. sp. distribution. Inset: sample range of A. marnkalha collected from the Great Barrier Reef and Torres Strait region circumscribed by square. Black dots identified with A, B, C represent easternmost specimens of A. marnkalha which were originally identified as A. fasciatus; A. AMS I 21842 - 001; B. AMS I 15557 - 003; C. CSIRO H 1118 - 01." httpUri="https://zenodo.org/record/177432/files/figure.png" pageId="14" pageNumber="33">Fig. 8</figureCitation>
). 
<taxonomicName id="4C414D16FFA3FFA7FD8CFF4EFCA2FF26" box="[514,815,152,177]" class="Elasmobranchii" family="Scyliorhinidae" genus="Atelomycterus" kingdom="Animalia" order="Carcharhiniformes" pageId="14" pageNumber="33" phylum="Chordata" rank="species" species="marnkalha">
<emphasis id="B935EA87FFA3FFA7FD8CFF4EFCA2FF26" box="[514,815,152,177]" italics="true" pageId="14" pageNumber="33">Atelomycterus marnkalha</emphasis>
</taxonomicName>
specimens collected from the Great Barrier Reef and the Torres Strait were caught in a variety of environmental substrates ranging from sandy to course rubble (Great Barrier Reef Seabed Biodiversity Project). Depths of capture were from 
<quantity id="4CB99B70FFA3FFA7FBE3FF31FA9EFE96" box="[1133,1299,231,257]" metricMagnitude="1" metricUnit="m" metricValue="4.2299999999999995" metricValueMax="7.4" metricValueMin="1.06" pageId="14" pageNumber="33" unit="m" value="42.3" valueMax="74.0" valueMin="10.6">10.6 to 74 m</quantity>
, with most records shallower than 
<specimenCount id="9D47FD1CFFA3FFA7FE29FED8FE6EFEBF" box="[423,483,270,296]" pageId="14" pageNumber="33" type="generic">
<quantity id="4CB99B70FFA3FFA7FE29FED8FE52FEBF" box="[423,479,270,296]" metricMagnitude="1" metricUnit="m" metricValue="5.0" pageId="14" pageNumber="33" unit="m" value="50.0">50m</quantity>
.
</specimenCount>
</paragraph>
<paragraph id="8BFE3695FFA3FFA7FF48FEE0FE15FD81" blockId="14.[151,1437,151,2036]" pageId="14" pageNumber="33">
<emphasis id="B935EA87FFA3FFA7FF48FEE0FDA9FEC7" bold="true" box="[198,548,310,336]" pageId="14" pageNumber="33">Size, reproduction and diet.</emphasis>
The total length of males and females of 
<taxonomicName id="4C414D16FFA3FFA7FB92FEE0FB33FED8" box="[1052,1214,310,335]" class="Elasmobranchii" family="Scyliorhinidae" genus="Atelomycterus" kingdom="Animalia" order="Carcharhiniformes" pageId="14" pageNumber="33" phylum="Chordata" rank="species" species="marnkalha">
<emphasis id="B935EA87FFA3FFA7FB92FEE0FB33FED8" box="[1052,1214,310,335]" italics="true" pageId="14" pageNumber="33">A. marnkalha</emphasis>
</taxonomicName>
ranged from 190– 470 and 
<quantity id="4CB99B70FFA3FFA7FF75FE88FE1AFEEF" box="[251,407,350,376]" metricMagnitude="-1" metricUnit="m" metricValue="3.31" metricValueMax="4.84" metricValueMin="1.78" pageId="14" pageNumber="33" unit="mm" value="331.0" valueMax="484.0" valueMin="178.0">178–484 mm</quantity>
TL respectively. The average mass of adult males was 92.5% of the adult females, with considerable overlap between large males and small females. The mass of adult males ranged from 182.5– 286.0 g (n=5, mean = 203.8, sd = 43.6); adult females mass range 
<quantity id="4CB99B70FFA3FFA7FC48FE7BFBD1FE50" box="[966,1116,429,455]" metricMagnitude="-1" metricUnit="kg" metricValue="2.6814999999999998" metricValueMax="3.863" metricValueMin="1.5" pageId="14" pageNumber="33" unit="g" value="268.15" valueMax="386.3" valueMin="150.0">150–386.3 g</quantity>
(n=4, mean = 220.3, sd = 111.2). There was a considerable overlap in the mass of juveniles and adults for both males and females in the 
<taxonomicName id="4C414D16FFA3FFA7FF19FE2AFEBBFD82" box="[151,310,508,533]" class="Elasmobranchii" family="Scyliorhinidae" genus="Atelomycterus" kingdom="Animalia" order="Carcharhiniformes" pageId="14" pageNumber="33" phylum="Chordata" rank="species" species="marnkalha">
<emphasis id="B935EA87FFA3FFA7FF19FE2AFEBBFD82" box="[151,310,508,533]" italics="true" pageId="14" pageNumber="33">A. marnkalha</emphasis>
</taxonomicName>
sample.
</paragraph>
<paragraph id="8BFE3695FFA3FFA7FF48FDF2FC91FD23" blockId="14.[151,1437,151,2036]" pageId="14" pageNumber="33">
The smallest sexually mature female was 
<quantity id="4CB99B70FFA3FFA7FD22FDF2FC8BFDA9" box="[684,774,548,574]" metricMagnitude="-1" metricUnit="m" metricValue="3.54" pageId="14" pageNumber="33" unit="mm" value="354.0">354mm</quantity>
TL; although size at first maturity is possibly lower. The smallest sexually mature male was recorded at 
<quantity id="4CB99B70FFA3FFA7FD5FFD9DFCA6FDF2" box="[721,811,587,613]" metricMagnitude="-1" metricUnit="m" metricValue="3.45" pageId="14" pageNumber="33" unit="mm" value="345.0">345mm</quantity>
TL. Reproduction, as with the other atelomycterine species is expected to be oviparous (McKay, 1965; 
<bibRefCitation id="EFD04B64FFA3FFA7FD63FDA5FBB4FD1A" author="Compagno" box="[749,1081,627,653]" pageId="14" pageNumber="36" refString="Compagno, L. J. V. &amp; Stevens, J. D. (1993) Atelomycterus fasciatus n. sp., a new catshark (Chondrichthyes: Carcharhiniformes: Scyliorhinidae) from tropical Australia. Records of the Australian Museum, Vol. 45, 147 - 169." type="journal article" year="1993">Compagno &amp; Stevens, 1993</bibRefCitation>
; 
<bibRefCitation id="EFD04B64FFA3FFA7FBCAFDA5FB73FD1A" author="Bor" box="[1092,1278,627,653]" pageId="14" pageNumber="36" refString="Bor, P. H. F., van Oijen, M. J. P. &amp; Magenta, C. (2003) The egg capsule of the coral cat shark, Atelomycterus marmoratus (Bennett 1830) (Chondrichthyes: Scyliorhinidae). Zoologische Mededelingen, Vol. 77, 87 - 92." type="journal article" year="2003">
Bor 
<emphasis id="B935EA87FFA3FFA7FBF9FDA5FB3CFD1B" box="[1143,1201,627,652]" italics="true" pageId="14" pageNumber="33">et al.</emphasis>
, 2003
</bibRefCitation>
), although no egg casings were recorded from the sample specimens.
</paragraph>
<paragraph id="8BFE3695FFA3FFA7FF48FD14FB5CFCBC" blockId="14.[151,1437,151,2036]" pageId="14" pageNumber="33">
<taxonomicName id="4C414D16FFA3FFA7FF48FD14FE75FD4C" box="[198,504,706,731]" class="Elasmobranchii" family="Scyliorhinidae" genus="Atelomycterus" kingdom="Animalia" order="Carcharhiniformes" pageId="14" pageNumber="33" phylum="Chordata" rank="species" species="marnkalha">
<emphasis id="B935EA87FFA3FFA7FF48FD14FE75FD4C" box="[198,504,706,731]" italics="true" pageId="14" pageNumber="33">Atelomycterus marnkalha</emphasis>
</taxonomicName>
feeds predominately on benthic invertebrates and small teleost species. Prey items found in the stomach content of 
<taxonomicName id="4C414D16FFA3FFA7FDD9FD3CFD7BFC94" box="[599,758,746,771]" class="Elasmobranchii" family="Scyliorhinidae" genus="Atelomycterus" kingdom="Animalia" order="Carcharhiniformes" pageId="14" pageNumber="33" phylum="Chordata" rank="species" species="marnkalha">
<emphasis id="B935EA87FFA3FFA7FDD9FD3CFD7BFC94" box="[599,758,746,771]" italics="true" pageId="14" pageNumber="33">A. marnkalha</emphasis>
</taxonomicName>
(n=4) included penaeid prawns, brachyuran crabs, cephalopods (Order Teuthida) and teleost species from the families 
<taxonomicName id="4C414D16FFA3FFA7FCE5FCC7FC5BFCBC" box="[875,982,785,811]" class="Actinopterygii" family="Gobiidae" kingdom="Animalia" order="Perciformes" pageId="14" pageNumber="33" phylum="Chordata" rank="family">Gobiidae</taxonomicName>
and 
<taxonomicName id="4C414D16FFA3FFA7FB81FCC7FB40FCBC" box="[1039,1229,785,811]" class="Actinopterygii" family="Platycephalidae" kingdom="Animalia" order="Scorpaeniformes" pageId="14" pageNumber="33" phylum="Chordata" rank="family">Platycephalidae</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C35B651EFFA3FFA7FF48FCEEFEC1FA5C" pageId="14" pageNumber="33" type="etymology">
<paragraph id="8BFE3695FFA3FFA7FF48FCEEFDFDFB07" blockId="14.[151,1437,151,2036]" pageId="14" pageNumber="33">
<emphasis id="B935EA87FFA3FFA7FF48FCEEFEDEFCC5" bold="true" box="[198,339,824,850]" pageId="14" pageNumber="33">Etymology.</emphasis>
The term 
<taxonomicName id="4C414D16FFA3FFA7FE45FCEFFDC7FCC5" box="[459,586,825,850]" class="Elasmobranchii" family="Scyliorhinidae" genus="Atelomycterus" kingdom="Animalia" order="Carcharhiniformes" pageId="14" pageNumber="33" phylum="Chordata" rank="species" species="marnkalha">
<emphasis id="B935EA87FFA3FFA7FE45FCEFFDC7FCC5" box="[459,586,825,850]" italics="true" pageId="14" pageNumber="33">marnkalha</emphasis>
</taxonomicName>
originates from an indigenous Australian dialect and is the word used by the Rrumburriya clan to describe regional catshark species in the Gulf of Carpentaria. The Rrumburriya clan belong to the Yanyuwa peoples and Yanyuwa country, an expansive area encompassing the south-western coastline of the Gulf of Carpentaria, the tidal reaches of McArthur and Wearyan Rivers, and the Sir Edward Pellew Islands (
<bibRefCitation id="EFD04B64FFA3FFA7FEDAFC01FDC4FC66" author="Bradley" box="[340,585,983,1009]" pageId="14" pageNumber="36" refString="Bradley, J., Homes, M., Norman, D., Isaac, A., Miller, J &amp; Ninganga, I. (2006) Yumbulyumbulmantha ki-Awawawu: All Kinds of Things from Country: Yanyuwa ethnobiological classification. Aboriginal and Torres Strait Islander Studies Unit Research Report Series Vol. 6. Aboriginal and Torres Strait Islander Studies Unit, University of Queensland, Brisbane, 174 pp." type="book" year="2006">
Bradley 
<emphasis id="B935EA87FFA3FFA7FE32FC0EFE75FC66" box="[444,504,984,1009]" italics="true" pageId="14" pageNumber="33">et al.</emphasis>
, 2006
</bibRefCitation>
). The Yanyuwa peoples have a close affinity to several marine animals including dugongs, turtles and elasmobranchs, which is reflected in their cultural and subsistence patterns. Yanyuwa country is where one of the first 
<taxonomicName id="4C414D16FFA3FFA7FD28FBF1FCC4FBD7" box="[678,841,1063,1088]" class="Elasmobranchii" family="Scyliorhinidae" genus="Atelomycterus" kingdom="Animalia" order="Carcharhiniformes" pageId="14" pageNumber="33" phylum="Chordata" rank="species" species="marnkalha">
<emphasis id="B935EA87FFA3FFA7FD28FBF1FCC4FBD7" box="[678,841,1063,1088]" italics="true" pageId="14" pageNumber="33">A. marnkalha</emphasis>
</taxonomicName>
specimens was recorded (Compagno &amp; 
<bibRefCitation id="EFD04B64FFA3FFA7FAB4FBF0FF58FBFF" author="Stevens" pageId="14" pageNumber="36" refString="Last, P. R. &amp; Stevens, J. D. (1994) Sharks and Rays of Australia, CSIRO: Australia, 513 pp." type="book" year="1994">Stevens, 1994</bibRefCitation>
; 
<bibRefCitation id="EFD04B64FFA3FFA7FF6CFB98FE5FFBFF" author="Bradley" box="[226,466,1102,1128]" pageId="14" pageNumber="36" refString="Bradley, J., Homes, M., Norman, D., Isaac, A., Miller, J &amp; Ninganga, I. (2006) Yumbulyumbulmantha ki-Awawawu: All Kinds of Things from Country: Yanyuwa ethnobiological classification. Aboriginal and Torres Strait Islander Studies Unit Research Report Series Vol. 6. Aboriginal and Torres Strait Islander Studies Unit, University of Queensland, Brisbane, 174 pp." type="book" year="2006">
Bradley 
<emphasis id="B935EA87FFA3FFA7FEC9FB98FE0EFBF0" box="[327,387,1102,1127]" italics="true" pageId="14" pageNumber="33">et al.</emphasis>
, 2006
</bibRefCitation>
). The common name Eastern Banded Catshark is in reference to the species’ geographical occurrence and colour pattern.
</paragraph>
<paragraph id="8BFE3695FFA3FFA7FF48FB4BFEC1FA5C" blockId="14.[151,1437,151,2036]" pageId="14" pageNumber="33">
<emphasis id="B935EA87FFA3FFA7FF48FB4BFDDBFB20" bold="true" box="[198,598,1181,1207]" pageId="14" pageNumber="33">Comparison with other species.</emphasis>
<taxonomicName id="4C414D16FFA3FFA7FDEEFB48FC1CFB20" box="[608,913,1182,1207]" class="Elasmobranchii" family="Scyliorhinidae" genus="Atelomycterus" kingdom="Animalia" order="Carcharhiniformes" pageId="14" pageNumber="33" phylum="Chordata" rank="species" species="marnkalha" status="sp. nov.">
<emphasis id="B935EA87FFA3FFA7FDEEFB48FC1CFB20" box="[608,913,1182,1207]" italics="true" pageId="14" pageNumber="33">Atelomycterus marnkalha</emphasis>
</taxonomicName>
<emphasis id="B935EA87FFA3FFA7FC14FB4BFC52FB20" bold="true" box="[922,991,1181,1207]" pageId="14" pageNumber="33">
<taxonomicNameLabel id="A20657FCFFA3FFA7FC14FB4BFC52FB20" box="[922,991,1181,1207]" pageId="14" pageNumber="33" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
is readily distinguishable from other atelomycterine species in external morphology and morphometrics, clasper structure, colouration and vertebral counts. It is most similar to the West Australian species 
<taxonomicName id="4C414D16FFA3FFA7FCEDFB3AFC61FA92" box="[867,1004,1260,1285]" class="Elasmobranchii" family="Scyliorhinidae" genus="Atelomycterus" kingdom="Animalia" order="Carcharhiniformes" pageId="14" pageNumber="33" phylum="Chordata" rank="species" species="fasciatus">
<emphasis id="B935EA87FFA3FFA7FCEDFB3AFC61FA92" box="[867,1004,1260,1285]" italics="true" pageId="14" pageNumber="33">A. fasciatus</emphasis>
</taxonomicName>
with the main morphological differences seen in the size of the anal fin (
<tableCitation id="C6C3032EFFA3FFA7FDECFAC2FD30FAB9" box="[610,701,1300,1326]" captionStart="TABLE 1" captionStartId="1.[151,239,1066,1090]" captionText="TABLE 1. Morphometric data expressed as % of TL for the Atelomycterus marnkalha n. sp. holotype, paratype size ranges and A. fasciatus comparisons." pageId="14" pageNumber="33">Table 1</tableCitation>
). In addition, 
<taxonomicName id="4C414D16FFA3FFA7FCE2FAC2FB83FABA" box="[876,1038,1300,1325]" class="Elasmobranchii" family="Scyliorhinidae" genus="Atelomycterus" kingdom="Animalia" order="Carcharhiniformes" pageId="14" pageNumber="33" phylum="Chordata" rank="species" species="marnkalha">
<emphasis id="B935EA87FFA3FFA7FCE2FAC2FB83FABA" box="[876,1038,1300,1325]" italics="true" pageId="14" pageNumber="33">A. marnkalha</emphasis>
</taxonomicName>
has a higher pelvic-anal to analcaudal ratio (1.65–2.38 
<emphasis id="B935EA87FFA3FFA7FE3AFAEDFE41FAC3" box="[436,460,1339,1364]" italics="true" pageId="14" pageNumber="33">vs</emphasis>
. 1.33–1.59 times in 
<taxonomicName id="4C414D16FFA3FFA7FD4AFAEDFCC2FAC3" box="[708,847,1339,1364]" class="Elasmobranchii" family="Scyliorhinidae" genus="Atelomycterus" kingdom="Animalia" order="Carcharhiniformes" pageId="14" pageNumber="33" phylum="Chordata" rank="species" species="fasciatus">
<emphasis id="B935EA87FFA3FFA7FD4AFAEDFCC2FAC3" box="[708,847,1339,1364]" italics="true" pageId="14" pageNumber="33">A. fasciatus</emphasis>
</taxonomicName>
) and a larger dorsal fin-caudal fin margin (17.5– 18.7 
<emphasis id="B935EA87FFA3FFA7FF5FFAB5FF64FAEB" box="[209,233,1379,1404]" italics="true" pageId="14" pageNumber="33">vs</emphasis>
16.3–17.3) (
<tableCitation id="C6C3032EFFA3FFA7FEF3FAB4FE58FAEB" box="[381,469,1378,1404]" captionStart="TABLE 1" captionStartId="1.[151,239,1066,1090]" captionText="TABLE 1. Morphometric data expressed as % of TL for the Atelomycterus marnkalha n. sp. holotype, paratype size ranges and A. fasciatus comparisons." pageId="14" pageNumber="33">Table 1</tableCitation>
). The lateral trunk denticles of 
<taxonomicName id="4C414D16FFA3FFA7FCCDFAB5FC6FFAEB" box="[835,994,1379,1404]" class="Elasmobranchii" family="Scyliorhinidae" genus="Atelomycterus" kingdom="Animalia" order="Carcharhiniformes" pageId="14" pageNumber="33" phylum="Chordata" rank="species" species="marnkalha">
<emphasis id="B935EA87FFA3FFA7FCCDFAB5FC6FFAEB" box="[835,994,1379,1404]" italics="true" pageId="14" pageNumber="33">A. marnkalha</emphasis>
</taxonomicName>
lack the prominent lateral ridges seen in 
<taxonomicName id="4C414D16FFA3FFA7FF36FA5CFEC8FA34" box="[184,325,1418,1443]" class="Elasmobranchii" family="Scyliorhinidae" genus="Atelomycterus" kingdom="Animalia" order="Carcharhiniformes" pageId="14" pageNumber="33" phylum="Chordata" rank="species" species="fasciatus">
<emphasis id="B935EA87FFA3FFA7FF36FA5CFEC8FA34" box="[184,325,1418,1443]" italics="true" pageId="14" pageNumber="33">A. fasciatus</emphasis>
</taxonomicName>
and have clearly visible shallow depressions (
<figureCitation id="137A2A10FFA3FFA7FCF2FA5CFC4BFA33" box="[892,966,1418,1444]" captionStart="FIGURE 5" captionStartId="10.[151,255,1595,1619]" captionTargetBox="[384,1203,194,1551]" captionTargetId="figure@10.[384,1203,194,1571]" captionTargetPageId="10" captionText="FIGURE 5. Atelomycterus marnkalha n. sp.; scanning electron micrographs of lateral trunk denticles in three-quarter anteriorlateral view, 459 mm TL male at A, 100 Μm, and B, 50 Μm. Photo by K. Sewell." httpUri="https://zenodo.org/record/177429/files/figure.png" pageId="14" pageNumber="33">Fig. 5</figureCitation>
); absent in 
<taxonomicName id="4C414D16FFA3FFA7FBD7FA5CFEC5FA5C" authority="Compagno &amp; Stevens, 1993" authorityName="Compagno &amp; Stevens" authorityYear="1993" class="Elasmobranchii" family="Scyliorhinidae" genus="Atelomycterus" kingdom="Animalia" order="Carcharhiniformes" pageId="14" pageNumber="33" phylum="Chordata" rank="species" species="fasciatus">
<emphasis id="B935EA87FFA3FFA7FBD7FA5CFB68FA34" box="[1113,1253,1418,1443]" italics="true" pageId="14" pageNumber="33">A. fasciatus</emphasis>
(
<bibRefCitation id="EFD04B64FFA3FFA7FB76FA5CFEB2FA5C" author="Compagno" pageId="14" pageNumber="36" refString="Compagno, L. J. V. &amp; Stevens, J. D. (1993) Atelomycterus fasciatus n. sp., a new catshark (Chondrichthyes: Carcharhiniformes: Scyliorhinidae) from tropical Australia. Records of the Australian Museum, Vol. 45, 147 - 169." type="journal article" year="1993">Compagno &amp; Stevens, 1993</bibRefCitation>
)
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C35B651EFFA3FFA7FF48FA0FFF6EF863" pageId="14" pageNumber="33" type="reference_group">
<paragraph id="8BFE3695FFA3FFA7FF48FA0FFE15F890" blockId="14.[151,1437,151,2036]" pageId="14" pageNumber="33">
<taxonomicName id="4C414D16FFA3FFA7FF48FA0FFE79FA65" ID-CoL="J8V8" box="[198,500,1497,1522]" class="Elasmobranchii" family="Scyliorhinidae" genus="Atelomycterus" kingdom="Animalia" order="Carcharhiniformes" pageId="14" pageNumber="33" phylum="Chordata" rank="species" species="marnkalha">
<emphasis id="B935EA87FFA3FFA7FF48FA0FFE79FA65" box="[198,500,1497,1522]" italics="true" pageId="14" pageNumber="33">Atelomycterus marnkalha</emphasis>
</taxonomicName>
, as with 
<taxonomicName id="4C414D16FFA3FFA7FDD3FA0FFBC0FA64" ID-CoL="J8V4" authority="Compagno &amp; Stevens, 1993" authorityName="Compagno &amp; Stevens" authorityYear="1993" box="[605,1101,1497,1523]" class="Elasmobranchii" family="Scyliorhinidae" genus="Atelomycterus" kingdom="Animalia" order="Carcharhiniformes" pageId="14" pageNumber="33" phylum="Chordata" rank="species" species="fasciatus">
<emphasis id="B935EA87FFA3FFA7FDD3FA0FFD6BFA65" box="[605,742,1497,1522]" italics="true" pageId="14" pageNumber="33">A. fasciatus</emphasis>
(
<bibRefCitation id="EFD04B64FFA3FFA7FD7BFA0FFBC9FA64" author="Compagno" box="[757,1092,1497,1523]" pageId="14" pageNumber="36" refString="Compagno, L. J. V. &amp; Stevens, J. D. (1993) Atelomycterus fasciatus n. sp., a new catshark (Chondrichthyes: Carcharhiniformes: Scyliorhinidae) from tropical Australia. Records of the Australian Museum, Vol. 45, 147 - 169." type="journal article" year="1993">Compagno &amp; Stevens, 1993</bibRefCitation>
)
</taxonomicName>
, differs from 
<taxonomicName id="4C414D16FFA3FFA7FB62FA0FFA11FA65" ID-CoL="J8V6" box="[1260,1436,1497,1522]" class="Elasmobranchii" family="Scyliorhinidae" genus="Atelomycterus" kingdom="Animalia" order="Carcharhiniformes" pageId="14" pageNumber="33" phylum="Chordata" rank="species" species="marmoratus">
<emphasis id="B935EA87FFA3FFA7FB62FA0FFA11FA65" box="[1260,1436,1497,1522]" italics="true" pageId="14" pageNumber="33">A. marmoratus</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C414D16FFA3FFA7FF47F9D7FED9F98D" ID-CoL="J8V5" box="[201,340,1537,1562]" class="Elasmobranchii" family="Scyliorhinidae" genus="Atelomycterus" kingdom="Animalia" order="Carcharhiniformes" pageId="14" pageNumber="33" phylum="Chordata" rank="species" species="macleayi">
<emphasis id="B935EA87FFA3FFA7FF47F9D7FED9F98D" box="[201,340,1537,1562]" italics="true" pageId="14" pageNumber="33">A. macleayi</emphasis>
</taxonomicName>
in having nearly straight to weakly concave dorsal fin posterior margins, slanting posterventrally from the apex. This difference is present though less pronounced when comparing 
<taxonomicName id="4C414D16FFA3FFA7FB37F9FEFAD7F9D6" ID-CoL="J8V8" box="[1209,1370,1576,1601]" class="Elasmobranchii" family="Scyliorhinidae" genus="Atelomycterus" kingdom="Animalia" order="Carcharhiniformes" pageId="14" pageNumber="33" phylum="Chordata" rank="species" species="marnkalha">
<emphasis id="B935EA87FFA3FFA7FB37F9FEFAD7F9D6" box="[1209,1370,1576,1601]" italics="true" pageId="14" pageNumber="33">A. marnkalha</emphasis>
</taxonomicName>
to 
<taxonomicName id="4C414D16FFA3FFA7FA0DF9FEFF72F9FE" ID-CoL="J8V2" class="Elasmobranchii" family="Scyliorhinidae" genus="Atelomycterus" kingdom="Animalia" order="Carcharhiniformes" pageId="14" pageNumber="33" phylum="Chordata" rank="species" species="baliensis">
<emphasis id="B935EA87FFA3FFA7FA0DF9FEFF72F9FE" italics="true" pageId="14" pageNumber="33">A. baliensis</emphasis>
</taxonomicName>
, which has weakly falcate dorsal fins (
<bibRefCitation id="EFD04B64FFA3FFA7FD47F999FC2EF9FE" author="White" box="[713,931,1615,1641]" pageId="14" pageNumber="36" refString="White, W. T., Last, P. R. &amp; Dharmadi. (2005) Description of a new species of catshark Atelomycterus baliensis (Carcharhiniformes: scyliorhinidae) from eastern Indonesia. Cybium, Vol. 29 (1), 33 - 40." type="journal article" year="2005">
White 
<emphasis id="B935EA87FFA3FFA7FC94F986FCD8F9FE" box="[794,853,1616,1641]" italics="true" pageId="14" pageNumber="33">et al.</emphasis>
, 2005
</bibRefCitation>
). In addition, 
<taxonomicName id="4C414D16FFA3FFA7FBC9F986FB5DF9FE" ID-CoL="J8V2" box="[1095,1232,1616,1641]" class="Elasmobranchii" family="Scyliorhinidae" genus="Atelomycterus" kingdom="Animalia" order="Carcharhiniformes" pageId="14" pageNumber="33" phylum="Chordata" rank="species" species="baliensis">
<emphasis id="B935EA87FFA3FFA7FBC9F986FB5DF9FE" box="[1095,1232,1616,1641]" italics="true" pageId="14" pageNumber="33">A. baliensis</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C414D16FFA3FFA7FB51F986FAE7F9FE" ID-CoL="J8V5" box="[1247,1386,1616,1641]" class="Elasmobranchii" family="Scyliorhinidae" genus="Atelomycterus" kingdom="Animalia" order="Carcharhiniformes" pageId="14" pageNumber="33" phylum="Chordata" rank="species" species="macleayi">
<emphasis id="B935EA87FFA3FFA7FB51F986FAE7F9FE" box="[1247,1386,1616,1641]" italics="true" pageId="14" pageNumber="33">A. macleayi</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C414D16FFA3FFA7FF19F9A1FEC5F907" ID-CoL="J8V6" box="[151,328,1655,1680]" class="Elasmobranchii" family="Scyliorhinidae" genus="Atelomycterus" kingdom="Animalia" order="Carcharhiniformes" pageId="14" pageNumber="33" phylum="Chordata" rank="species" species="marmoratus">
<emphasis id="B935EA87FFA3FFA7FF19F9A1FEC5F907" box="[151,328,1655,1680]" italics="true" pageId="14" pageNumber="33">A. marmoratus</emphasis>
</taxonomicName>
have relatively short preoral (POR) and prenarial (PRN) snout lengths when compared to 
<taxonomicName id="4C414D16FFA3FFA7FA0DF9A1FE9BF92F" ID-CoL="J8V8" class="Elasmobranchii" family="Scyliorhinidae" genus="Atelomycterus" kingdom="Animalia" order="Carcharhiniformes" pageId="14" pageNumber="33" phylum="Chordata" rank="species" species="marnkalha">
<emphasis id="B935EA87FFA3FFA7FA0DF9A1FE9BF92F" italics="true" pageId="14" pageNumber="33">A. marnkalha</emphasis>
</taxonomicName>
; POR: 
<taxonomicName id="4C414D16FFA3FFA7FEE2F949FE78F92F" ID-CoL="J8V2" box="[364,501,1695,1720]" class="Elasmobranchii" family="Scyliorhinidae" genus="Atelomycterus" kingdom="Animalia" order="Carcharhiniformes" pageId="14" pageNumber="33" phylum="Chordata" rank="species" species="baliensis">
<emphasis id="B935EA87FFA3FFA7FEE2F949FE78F92F" box="[364,501,1695,1720]" italics="true" pageId="14" pageNumber="33">A. baliensis</emphasis>
</taxonomicName>
4.14–4.51, 
<taxonomicName id="4C414D16FFA3FFA7FDF0F949FC84F92F" ID-CoL="J8V5" box="[638,777,1695,1720]" class="Elasmobranchii" family="Scyliorhinidae" genus="Atelomycterus" kingdom="Animalia" order="Carcharhiniformes" pageId="14" pageNumber="33" phylum="Chordata" rank="species" species="macleayi">
<emphasis id="B935EA87FFA3FFA7FDF0F949FC84F92F" box="[638,777,1695,1720]" italics="true" pageId="14" pageNumber="33">A. macleayi</emphasis>
</taxonomicName>
3.25–3.39, 
<taxonomicName id="4C414D16FFA3FFA7FC1CF949FBCCF92F" ID-CoL="J8V6" box="[914,1089,1695,1720]" class="Elasmobranchii" family="Scyliorhinidae" genus="Atelomycterus" kingdom="Animalia" order="Carcharhiniformes" pageId="14" pageNumber="33" phylum="Chordata" rank="species" species="marmoratus">
<emphasis id="B935EA87FFA3FFA7FC1CF949FBCCF92F" box="[914,1089,1695,1720]" italics="true" pageId="14" pageNumber="33">A. marmoratus</emphasis>
</taxonomicName>
4.06–4.47; PRN: 
<taxonomicName id="4C414D16FFA3FFA7FA9AF949FA11F92F" ID-CoL="J8V2" box="[1300,1436,1695,1720]" class="Elasmobranchii" family="Scyliorhinidae" genus="Atelomycterus" kingdom="Animalia" order="Carcharhiniformes" pageId="14" pageNumber="33" phylum="Chordata" rank="species" species="baliensis">
<emphasis id="B935EA87FFA3FFA7FA9AF949FA11F92F" box="[1300,1436,1695,1720]" italics="true" pageId="14" pageNumber="33">A. baliensis</emphasis>
</taxonomicName>
3.42–3.60, 
<taxonomicName id="4C414D16FFA3FFA7FE94F910FE29F948" ID-CoL="J8V5" box="[282,420,1734,1759]" class="Elasmobranchii" family="Scyliorhinidae" genus="Atelomycterus" kingdom="Animalia" order="Carcharhiniformes" pageId="14" pageNumber="33" phylum="Chordata" rank="species" species="macleayi">
<emphasis id="B935EA87FFA3FFA7FE94F910FE29F948" box="[282,420,1734,1759]" italics="true" pageId="14" pageNumber="33">A. macleayi</emphasis>
</taxonomicName>
2.46–2.55, 
<taxonomicName id="4C414D16FFA3FFA7FDA0F910FD50F948" ID-CoL="J8V6" box="[558,733,1734,1759]" class="Elasmobranchii" family="Scyliorhinidae" genus="Atelomycterus" kingdom="Animalia" order="Carcharhiniformes" pageId="14" pageNumber="33" phylum="Chordata" rank="species" species="marmoratus">
<emphasis id="B935EA87FFA3FFA7FDA0F910FD50F948" box="[558,733,1734,1759]" italics="true" pageId="14" pageNumber="33">A. marmoratus</emphasis>
</taxonomicName>
3.18–3.63. Some overlap occurs between 
<taxonomicName id="4C414D16FFA3FFA7FB42F910FAE6F948" ID-CoL="J8V8" box="[1228,1387,1734,1759]" class="Elasmobranchii" family="Scyliorhinidae" genus="Atelomycterus" kingdom="Animalia" order="Carcharhiniformes" pageId="14" pageNumber="33" phylum="Chordata" rank="species" species="marnkalha">
<emphasis id="B935EA87FFA3FFA7FB42F910FAE6F948" box="[1228,1387,1734,1759]" italics="true" pageId="14" pageNumber="33">A. marnkalha</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C414D16FFA3FFA7FF19F938FE92F890" ID-CoL="J8V4" box="[151,287,1774,1799]" class="Elasmobranchii" family="Scyliorhinidae" genus="Atelomycterus" kingdom="Animalia" order="Carcharhiniformes" pageId="14" pageNumber="33" phylum="Chordata" rank="species" species="fasciatus">
<emphasis id="B935EA87FFA3FFA7FF19F938FE92F890" box="[151,287,1774,1799]" italics="true" pageId="14" pageNumber="33">A. fasciatus</emphasis>
</taxonomicName>
(
<tableCitation id="C6C3032EFFA3FFA7FEA0F93BFE0BF890" box="[302,390,1773,1799]" captionStart="TABLE 1" captionStartId="1.[151,239,1066,1090]" captionText="TABLE 1. Morphometric data expressed as % of TL for the Atelomycterus marnkalha n. sp. holotype, paratype size ranges and A. fasciatus comparisons." pageId="14" pageNumber="33">Table 1</tableCitation>
).
</paragraph>
<paragraph id="8BFE3695FFA3FFA7FF48F8C3FF6EF863" blockId="14.[151,1437,151,2036]" pageId="14" pageNumber="33">
The five 
<taxonomicName id="4C414D16FFA3FFA7FEBAF8C3FE51F8B9" ID-CoL="34VW" box="[308,476,1813,1838]" class="Elasmobranchii" family="Scyliorhinidae" genus="Atelomycterus" kingdom="Animalia" order="Carcharhiniformes" pageId="14" pageNumber="33" phylum="Chordata" rank="genus">
<emphasis id="B935EA87FFA3FFA7FEBAF8C3FE51F8B9" box="[308,476,1813,1838]" italics="true" pageId="14" pageNumber="33">Atelomycterus</emphasis>
</taxonomicName>
species are clearly differentiated by the external morphology and dimensions of their claspers. Adult males of 
<taxonomicName id="4C414D16FFA3FFA7FE71F8EAFD3DF8C2" ID-CoL="J8V6" box="[511,688,1852,1877]" class="Elasmobranchii" family="Scyliorhinidae" genus="Atelomycterus" kingdom="Animalia" order="Carcharhiniformes" pageId="14" pageNumber="33" phylum="Chordata" rank="species" species="marmoratus">
<emphasis id="B935EA87FFA3FFA7FE71F8EAFD3DF8C2" box="[511,688,1852,1877]" italics="true" pageId="14" pageNumber="33">A. marmoratus</emphasis>
</taxonomicName>
have very elongate and attenuated claspers when compared to the other four species (
<figureCitation id="137A2A10FFA3FFA7FE2BF8B2FE63F8E9" box="[421,494,1892,1918]" captionStart="FIGURE 5" captionStartId="10.[151,255,1595,1619]" captionTargetBox="[384,1203,194,1551]" captionTargetId="figure@10.[384,1203,194,1571]" captionTargetPageId="10" captionText="FIGURE 5. Atelomycterus marnkalha n. sp.; scanning electron micrographs of lateral trunk denticles in three-quarter anteriorlateral view, 459 mm TL male at A, 100 Μm, and B, 50 Μm. Photo by K. Sewell." httpUri="https://zenodo.org/record/177429/files/figure.png" pageId="14" pageNumber="33">Fig. 5</figureCitation>
; 
<bibRefCitation id="EFD04B64FFA3FFA7FE72F8B2FCC1F8E9" author="Compagno" box="[508,844,1892,1918]" pageId="14" pageNumber="36" refString="Compagno, L. J. V. &amp; Stevens, J. D. (1993) Atelomycterus fasciatus n. sp., a new catshark (Chondrichthyes: Carcharhiniformes: Scyliorhinidae) from tropical Australia. Records of the Australian Museum, Vol. 45, 147 - 169." type="journal article" year="1993">Compagno &amp; Stevens, 1993</bibRefCitation>
). Mature 
<taxonomicName id="4C414D16FFA3FFA7FC30F8B2FBC4F8EA" ID-CoL="J8V5" box="[958,1097,1892,1917]" class="Elasmobranchii" family="Scyliorhinidae" genus="Atelomycterus" kingdom="Animalia" order="Carcharhiniformes" pageId="14" pageNumber="33" phylum="Chordata" rank="species" species="macleayi">
<emphasis id="B935EA87FFA3FFA7FC30F8B2FBC4F8EA" box="[958,1097,1892,1917]" italics="true" pageId="14" pageNumber="33">A. macleayi</emphasis>
</taxonomicName>
males have short, very stout claspers, with the clasper structure of mature 
<taxonomicName id="4C414D16FFA3FFA7FD20F85DFCC3F833" ID-CoL="J8V8" box="[686,846,1931,1956]" class="Elasmobranchii" family="Scyliorhinidae" genus="Atelomycterus" kingdom="Animalia" order="Carcharhiniformes" pageId="14" pageNumber="33" phylum="Chordata" rank="species" species="marnkalha">
<emphasis id="B935EA87FFA3FFA7FD20F85DFCC3F833" box="[686,846,1931,1956]" italics="true" pageId="14" pageNumber="33">A. marnkalha</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C414D16FFA3FFA7FCD3F85DFC6AF833" ID-CoL="J8V4" box="[861,999,1931,1956]" class="Elasmobranchii" family="Scyliorhinidae" genus="Atelomycterus" kingdom="Animalia" order="Carcharhiniformes" pageId="14" pageNumber="33" phylum="Chordata" rank="species" species="fasciatus">
<emphasis id="B935EA87FFA3FFA7FCD3F85DFC6AF833" box="[861,999,1931,1956]" italics="true" pageId="14" pageNumber="33">A. fasciatus</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C414D16FFA3FFA7FBAFF85DFB27F833" ID-CoL="J8V2" box="[1057,1194,1931,1956]" class="Elasmobranchii" family="Scyliorhinidae" genus="Atelomycterus" kingdom="Animalia" order="Carcharhiniformes" pageId="14" pageNumber="33" phylum="Chordata" rank="species" species="baliensis">
<emphasis id="B935EA87FFA3FFA7FBAFF85DFB27F833" box="[1057,1194,1931,1956]" italics="true" pageId="14" pageNumber="33">A. baliensis</emphasis>
</taxonomicName>
having intermediate characteristics; moderately stout, elongate claspers (
<figureCitation id="137A2A10FFA3FFA7FD70F864FCCAF85B" box="[766,839,1970,1996]" captionStart="FIGURE 5" captionStartId="10.[151,255,1595,1619]" captionTargetBox="[384,1203,194,1551]" captionTargetId="figure@10.[384,1203,194,1571]" captionTargetPageId="10" captionText="FIGURE 5. Atelomycterus marnkalha n. sp.; scanning electron micrographs of lateral trunk denticles in three-quarter anteriorlateral view, 459 mm TL male at A, 100 Μm, and B, 50 Μm. Photo by K. Sewell." httpUri="https://zenodo.org/record/177429/files/figure.png" pageId="14" pageNumber="33">Fig. 5</figureCitation>
; 
<bibRefCitation id="EFD04B64FFA3FFA7FCDBF864FB2BF85B" author="Compagno" box="[853,1190,1970,1996]" pageId="14" pageNumber="36" refString="Compagno, L. J. V. &amp; Stevens, J. D. (1993) Atelomycterus fasciatus n. sp., a new catshark (Chondrichthyes: Carcharhiniformes: Scyliorhinidae) from tropical Australia. Records of the Australian Museum, Vol. 45, 147 - 169." type="journal article" year="1993">Compagno &amp; Stevens, 1993</bibRefCitation>
; 
<figureCitation id="137A2A10FFA3FFA7FB3DF864FB76F85B" box="[1203,1275,1970,1996]" captionStart="FIGURE 6" captionStartId="11.[151,255,937,961]" captionTargetBox="[270,1322,202,898]" captionTargetId="figure@11.[264,1323,194,913]" captionTargetPageId="11" captionText="FIGURE 6. Claspers of Atelomycterus marnkalha n. sp.; 459 mm TL adult male (QM I 38089). A: Glans not dilated; B: Glans spread. Abbreviations: AP: Apopyle; CLD: Clasper denticles; CRH: Cover Rhipidion; ERH: Exorhipidion; FCG: Fused clasper groove; FRT: Free tab on cover rhipidion; P- 2: Pelvic fin; PSP: pseudopera; PSS: Pseudosiphon; RH: Rhipidion. Illustration by I. P. Jacobsen." httpUri="https://zenodo.org/record/177430/files/figure.png" pageId="14" pageNumber="33">Fig. 6</figureCitation>
; 
<bibRefCitation id="EFD04B64FFA3FFA7FA87F864FF5AF863" author="White" pageId="14" pageNumber="36" refString="White, W. T., Last, P. R. &amp; Dharmadi. (2005) Description of a new species of catshark Atelomycterus baliensis (Carcharhiniformes: scyliorhinidae) from eastern Indonesia. Cybium, Vol. 29 (1), 33 - 40." type="journal article" year="2005">
White 
<emphasis id="B935EA87FFA3FFA7FAD7F865FA18F85B" box="[1369,1429,1971,1996]" italics="true" pageId="14" pageNumber="33">et al.</emphasis>
, 2005
</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C35B651EFFA2FFA6FF48FF4EFD84FC79" pageId="15" pageNumber="34" type="description">
<paragraph id="8BFE3695FFA2FFA6FF48FF4EFAFBFE09" blockId="15.[151,1437,152,2033]" pageId="15" pageNumber="34">
Claspers of mature 
<taxonomicName id="4C414D16FFA2FFA6FE3EFF4EFDDFFF26" box="[432,594,152,177]" class="Elasmobranchii" family="Scyliorhinidae" genus="Atelomycterus" kingdom="Animalia" order="Carcharhiniformes" pageId="15" pageNumber="34" phylum="Chordata" rank="species" species="marnkalha">
<emphasis id="B935EA87FFA2FFA6FE3EFF4EFDDFFF26" box="[432,594,152,177]" italics="true" pageId="15" pageNumber="34">A. marnkalha</emphasis>
</taxonomicName>
males are less tapered than 
<taxonomicName id="4C414D16FFA2FFA6FC2BFF4EFBBDFF26" box="[933,1072,152,177]" class="Elasmobranchii" family="Scyliorhinidae" genus="Atelomycterus" kingdom="Animalia" order="Carcharhiniformes" pageId="15" pageNumber="34" phylum="Chordata" rank="species" species="fasciatus">
<emphasis id="B935EA87FFA2FFA6FC2BFF4EFBBDFF26" box="[933,1072,152,177]" italics="true" pageId="15" pageNumber="34">A. fasciatus</emphasis>
</taxonomicName>
, and stouter than in mature 
<taxonomicName id="4C414D16FFA2FFA6FA0DFF4EFF72FF4F" class="Elasmobranchii" family="Scyliorhinidae" genus="Atelomycterus" kingdom="Animalia" order="Carcharhiniformes" pageId="15" pageNumber="34" phylum="Chordata" rank="species" species="fasciatus">
<emphasis id="B935EA87FFA2FFA6FA0DFF4EFF72FF4F" italics="true" pageId="15" pageNumber="34">A. fasciatus</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C414D16FFA2FFA6FECDFF69FE5CFF4F" box="[323,465,191,216]" class="Elasmobranchii" family="Scyliorhinidae" genus="Atelomycterus" kingdom="Animalia" order="Carcharhiniformes" pageId="15" pageNumber="34" phylum="Chordata" rank="species" species="baliensis">
<emphasis id="B935EA87FFA2FFA6FECDFF69FE5CFF4F" box="[323,465,191,216]" italics="true" pageId="15" pageNumber="34">A. baliensis</emphasis>
</taxonomicName>
; clasper base width (CLB) as a percentage of clasper outer length (CLO), 
<taxonomicName id="4C414D16FFA2FFA6FA0DFF69FE9BFE97" class="Elasmobranchii" family="Scyliorhinidae" genus="Atelomycterus" kingdom="Animalia" order="Carcharhiniformes" pageId="15" pageNumber="34" phylum="Chordata" rank="species" species="marnkalha">
<emphasis id="B935EA87FFA2FFA6FA0DFF69FE9BFE97" italics="true" pageId="15" pageNumber="34">A. marnkalha</emphasis>
</taxonomicName>
22.0%, 
<taxonomicName id="4C414D16FFA2FFA6FEF4FF31FD89FE97" box="[378,516,231,256]" class="Elasmobranchii" family="Scyliorhinidae" genus="Atelomycterus" kingdom="Animalia" order="Carcharhiniformes" pageId="15" pageNumber="34" phylum="Chordata" rank="species" species="fasciatus">
<emphasis id="B935EA87FFA2FFA6FEF4FF31FD89FE97" box="[378,516,231,256]" italics="true" pageId="15" pageNumber="34">A. fasciatus</emphasis>
</taxonomicName>
17.1%, 
<taxonomicName id="4C414D16FFA2FFA6FDE7FF31FD7EFE97" box="[617,755,231,256]" class="Elasmobranchii" family="Scyliorhinidae" genus="Atelomycterus" kingdom="Animalia" order="Carcharhiniformes" pageId="15" pageNumber="34" phylum="Chordata" rank="species" species="baliensis">
<emphasis id="B935EA87FFA2FFA6FDE7FF31FD7EFE97" box="[617,755,231,256]" italics="true" pageId="15" pageNumber="34">A. baliensis</emphasis>
</taxonomicName>
15.4%. The claspers of 
<taxonomicName id="4C414D16FFA2FFA6FB98FF31FB3AFE97" box="[1046,1207,231,256]" class="Elasmobranchii" family="Scyliorhinidae" genus="Atelomycterus" kingdom="Animalia" order="Carcharhiniformes" pageId="15" pageNumber="34" phylum="Chordata" rank="species" species="marnkalha">
<emphasis id="B935EA87FFA2FFA6FB98FF31FB3AFE97" box="[1046,1207,231,256]" italics="true" pageId="15" pageNumber="34">A. marnkalha</emphasis>
</taxonomicName>
also differ from 
<taxonomicName id="4C414D16FFA2FFA6FA0DFF31FF72FEB0" class="Elasmobranchii" family="Scyliorhinidae" genus="Atelomycterus" kingdom="Animalia" order="Carcharhiniformes" pageId="15" pageNumber="34" phylum="Chordata" rank="species" species="fasciatus">
<emphasis id="B935EA87FFA2FFA6FA0DFF31FF72FEB0" italics="true" pageId="15" pageNumber="34">A. fasciatus</emphasis>
</taxonomicName>
in having a cover rhipidion with a straight to slightly concave anterior margin. The cover rhipidion in 
<taxonomicName id="4C414D16FFA2FFA6FF38FEE0FEB2FED8" box="[182,319,310,335]" class="Elasmobranchii" family="Scyliorhinidae" genus="Atelomycterus" kingdom="Animalia" order="Carcharhiniformes" pageId="15" pageNumber="34" phylum="Chordata" rank="species" species="fasciatus">
<emphasis id="B935EA87FFA2FFA6FF38FEE0FEB2FED8" box="[182,319,310,335]" italics="true" pageId="15" pageNumber="34">A. fasciatus</emphasis>
</taxonomicName>
has a distinct cover-rhipidion notch originating approximately opposite the anteriormost point of the pseudosiphon (
<figureCitation id="137A2A10FFA2FFA6FE17FE8BFE52FEE0" box="[409,479,349,375]" captionStart="FIGURE 5" captionStartId="10.[151,255,1595,1619]" captionTargetBox="[384,1203,194,1551]" captionTargetId="figure@10.[384,1203,194,1571]" captionTargetPageId="10" captionText="FIGURE 5. Atelomycterus marnkalha n. sp.; scanning electron micrographs of lateral trunk denticles in three-quarter anteriorlateral view, 459 mm TL male at A, 100 Μm, and B, 50 Μm. Photo by K. Sewell." httpUri="https://zenodo.org/record/177429/files/figure.png" pageId="15" pageNumber="34">Fig. 5</figureCitation>
A; 
<bibRefCitation id="EFD04B64FFA2FFA6FD88FE8BFCD6FEE0" author="Compagno" box="[518,859,349,375]" pageId="15" pageNumber="36" refString="Compagno, L. J. V. &amp; Stevens, J. D. (1993) Atelomycterus fasciatus n. sp., a new catshark (Chondrichthyes: Carcharhiniformes: Scyliorhinidae) from tropical Australia. Records of the Australian Museum, Vol. 45, 147 - 169." type="journal article" year="1993">Compagno &amp; Stevens, 1993</bibRefCitation>
). The clasper structure of 
<taxonomicName id="4C414D16FFA2FFA6FB1BFE8BFACBFEE1" box="[1173,1350,349,374]" class="Elasmobranchii" family="Scyliorhinidae" genus="Atelomycterus" kingdom="Animalia" order="Carcharhiniformes" pageId="15" pageNumber="34" phylum="Chordata" rank="species" species="marmoratus">
<emphasis id="B935EA87FFA2FFA6FB1BFE8BFACBFEE1" box="[1173,1350,349,374]" italics="true" pageId="15" pageNumber="34">A. marmoratus</emphasis>
</taxonomicName>
differs from all other atelomycterine species in having glans less than half the length of the claspers outer margin.
</paragraph>
<paragraph id="8BFE3695FFA2FFA6FF48FE7AFD42FD24" blockId="15.[151,1437,152,2033]" pageId="15" pageNumber="34">
The colouration of 
<taxonomicName id="4C414D16FFA2FFA6FE26FE7AFDCAFE52" box="[424,583,428,453]" class="Elasmobranchii" family="Scyliorhinidae" genus="Atelomycterus" kingdom="Animalia" order="Carcharhiniformes" pageId="15" pageNumber="34" phylum="Chordata" rank="species" species="marnkalha">
<emphasis id="B935EA87FFA2FFA6FE26FE7AFDCAFE52" box="[424,583,428,453]" italics="true" pageId="15" pageNumber="34">A. marnkalha</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C414D16FFA2FFA6FD0EFE7AFC84FE52" box="[640,777,428,453]" class="Elasmobranchii" family="Scyliorhinidae" genus="Atelomycterus" kingdom="Animalia" order="Carcharhiniformes" pageId="15" pageNumber="34" phylum="Chordata" rank="species" species="fasciatus">
<emphasis id="B935EA87FFA2FFA6FD0EFE7AFC84FE52" box="[640,777,428,453]" italics="true" pageId="15" pageNumber="34">A. fasciatus</emphasis>
</taxonomicName>
differs from other 
<taxonomicName id="4C414D16FFA2FFA6FC66FE7AFB1DFE52" box="[1000,1168,428,453]" class="Elasmobranchii" family="Scyliorhinidae" genus="Atelomycterus" kingdom="Animalia" order="Carcharhiniformes" pageId="15" pageNumber="34" phylum="Chordata" rank="genus">
<emphasis id="B935EA87FFA2FFA6FC66FE7AFB1DFE52" box="[1000,1168,428,453]" italics="true" pageId="15" pageNumber="34">Atelomycterus</emphasis>
</taxonomicName>
species in having dark brown-grey saddles and bands on a light grey background. Saddles may be present though less well-defined in 
<taxonomicName id="4C414D16FFA2FFA6FF19FE2DFE92FD83" box="[151,287,507,532]" class="Elasmobranchii" family="Scyliorhinidae" genus="Atelomycterus" kingdom="Animalia" order="Carcharhiniformes" pageId="15" pageNumber="34" phylum="Chordata" rank="species" species="baliensis">
<emphasis id="B935EA87FFA2FFA6FF19FE2DFE92FD83" box="[151,287,507,532]" italics="true" pageId="15" pageNumber="34">A. baliensis</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C414D16FFA2FFA6FED7FE2DFE69FD83" box="[345,484,507,532]" class="Elasmobranchii" family="Scyliorhinidae" genus="Atelomycterus" kingdom="Animalia" order="Carcharhiniformes" pageId="15" pageNumber="34" phylum="Chordata" rank="species" species="macleayi">
<emphasis id="B935EA87FFA2FFA6FED7FE2DFE69FD83" box="[345,484,507,532]" italics="true" pageId="15" pageNumber="34">A. macleayi</emphasis>
</taxonomicName>
adults and of equal width (
<bibRefCitation id="EFD04B64FFA2FFA6FCADFE2DFBFEFD82" author="Compagno" box="[803,1139,507,533]" pageId="15" pageNumber="36" refString="Compagno, L. J. V. &amp; Stevens, J. D. (1993) Atelomycterus fasciatus n. sp., a new catshark (Chondrichthyes: Carcharhiniformes: Scyliorhinidae) from tropical Australia. Records of the Australian Museum, Vol. 45, 147 - 169." type="journal article" year="1993">Compagno &amp; Stevens, 1993</bibRefCitation>
; 
<bibRefCitation id="EFD04B64FFA2FFA6FBF1FE2DFADAFD82" author="White" box="[1151,1367,507,533]" pageId="15" pageNumber="36" refString="White, W. T., Last, P. R. &amp; Dharmadi. (2005) Description of a new species of catshark Atelomycterus baliensis (Carcharhiniformes: scyliorhinidae) from eastern Indonesia. Cybium, Vol. 29 (1), 33 - 40." type="journal article" year="2005">
White 
<emphasis id="B935EA87FFA2FFA6FB40FE2DFA84FD83" box="[1230,1289,507,532]" italics="true" pageId="15" pageNumber="34">et al.</emphasis>
, 2005
</bibRefCitation>
). 
<emphasis id="B935EA87FFA2FFA6FAE5FE2DFDCAFDAC" italics="true" pageId="15" pageNumber="34">
<taxonomicName id="4C414D16FFA2FFA6FAE5FE2DFE01FDAC" class="Elasmobranchii" family="Scyliorhinidae" genus="Atelomycterus" kingdom="Animalia" order="Carcharhiniformes" pageId="15" pageNumber="34" phylum="Chordata" rank="species" species="macleayi">Atelomycterus macleayi</taxonomicName>
, 
<taxonomicName id="4C414D16FFA2FFA6FE19FDF4FDCAFDAC" box="[407,583,546,571]" class="Elasmobranchii" family="Scyliorhinidae" genus="Atelomycterus" kingdom="Animalia" order="Carcharhiniformes" pageId="15" pageNumber="34" phylum="Chordata" rank="species" species="marmoratus">A. marmoratus</taxonomicName>
</emphasis>
and 
<taxonomicName id="4C414D16FFA2FFA6FD0EFDF4FC84FDAC" box="[640,777,546,571]" class="Elasmobranchii" family="Scyliorhinidae" genus="Atelomycterus" kingdom="Animalia" order="Carcharhiniformes" pageId="15" pageNumber="34" phylum="Chordata" rank="species" species="baliensis">
<emphasis id="B935EA87FFA2FFA6FD0EFDF4FC84FDAC" box="[640,777,546,571]" italics="true" pageId="15" pageNumber="34">A. baliensis</emphasis>
</taxonomicName>
all have numerous, large and small dark brown to black spots covering the entire length of the dorsal surface. Spots of 
<taxonomicName id="4C414D16FFA2FFA6FC2CFD9CFBDBFDF4" box="[930,1110,586,611]" class="Elasmobranchii" family="Scyliorhinidae" genus="Atelomycterus" kingdom="Animalia" order="Carcharhiniformes" pageId="15" pageNumber="34" phylum="Chordata" rank="species" species="marmoratus">
<emphasis id="B935EA87FFA2FFA6FC2CFD9CFBDBFDF4" box="[930,1110,586,611]" italics="true" pageId="15" pageNumber="34">A. marmoratus</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C414D16FFA2FFA6FB16FD9CFAA8FDF4" box="[1176,1317,586,611]" class="Elasmobranchii" family="Scyliorhinidae" genus="Atelomycterus" kingdom="Animalia" order="Carcharhiniformes" pageId="15" pageNumber="34" phylum="Chordata" rank="species" species="baliensis">
<emphasis id="B935EA87FFA2FFA6FB16FD9CFAA8FDF4" box="[1176,1317,586,611]" italics="true" pageId="15" pageNumber="34">A. baliensis</emphasis>
</taxonomicName>
are often enlarged and merging when compared to 
<emphasis id="B935EA87FFA2FFA6FD0DFDA7FC9BFD1D" box="[643,790,625,650]" italics="true" pageId="15" pageNumber="34">
<taxonomicName id="4C414D16FFA2FFA6FD0DFDA7FC9CFD1D" box="[643,785,625,650]" class="Elasmobranchii" family="Scyliorhinidae" genus="Atelomycterus" kingdom="Animalia" order="Carcharhiniformes" pageId="15" pageNumber="34" phylum="Chordata" rank="species" species="macleayi">A. macleayi</taxonomicName>
.
</emphasis>
Black spots are less numerous and more defined in 
<taxonomicName id="4C414D16FFA2FFA6FA0DFDA7FE9BFD25" class="Elasmobranchii" family="Scyliorhinidae" genus="Atelomycterus" kingdom="Animalia" order="Carcharhiniformes" pageId="15" pageNumber="34" phylum="Chordata" rank="species" species="marnkalha">
<emphasis id="B935EA87FFA2FFA6FA0DFDA7FE9BFD25" italics="true" pageId="15" pageNumber="34">A. marnkalha</emphasis>
</taxonomicName>
, and least represented in 
<taxonomicName id="4C414D16FFA2FFA6FDB1FD4FFD45FD25" box="[575,712,665,690]" class="Elasmobranchii" family="Scyliorhinidae" genus="Atelomycterus" kingdom="Animalia" order="Carcharhiniformes" pageId="15" pageNumber="34" phylum="Chordata" rank="species" species="fasciatus">
<emphasis id="B935EA87FFA2FFA6FDB1FD4FFD45FD25" box="[575,712,665,690]" italics="true" pageId="15" pageNumber="34">A. fasciatus</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BFE3695FFA2FFA6FF48FD17FD84FC79" blockId="15.[151,1437,152,2033]" pageId="15" pageNumber="34">
<taxonomicName id="4C414D16FFA2FFA6FF48FD17FE7AFD4D" box="[198,503,705,730]" class="Elasmobranchii" family="Scyliorhinidae" genus="Atelomycterus" kingdom="Animalia" order="Carcharhiniformes" pageId="15" pageNumber="34" phylum="Chordata" rank="species" species="marnkalha">
<emphasis id="B935EA87FFA2FFA6FF48FD17FE7AFD4D" box="[198,503,705,730]" italics="true" pageId="15" pageNumber="34">Atelomycterus marnkalha</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C414D16FFA2FFA6FDBBFD17FD6AFD4D" box="[565,743,705,730]" class="Elasmobranchii" family="Scyliorhinidae" genus="Atelomycterus" kingdom="Animalia" order="Carcharhiniformes" pageId="15" pageNumber="34" phylum="Chordata" rank="species" species="marmoratus">
<emphasis id="B935EA87FFA2FFA6FDBBFD17FD6AFD4D" box="[565,743,705,730]" italics="true" pageId="15" pageNumber="34">A. marmoratus</emphasis>
</taxonomicName>
can be further differentiated from the other three species based on the presence or absence of white spots. 
<taxonomicName id="4C414D16FFA2FFA6FD5CFD3EFC72FC96" box="[722,1023,744,769]" class="Elasmobranchii" family="Scyliorhinidae" genus="Atelomycterus" kingdom="Animalia" order="Carcharhiniformes" pageId="15" pageNumber="34" phylum="Chordata" rank="species" species="marnkalha">
<emphasis id="B935EA87FFA2FFA6FD5CFD3EFC72FC96" box="[722,1023,744,769]" italics="true" pageId="15" pageNumber="34">Atelomycterus marnkalha</emphasis>
</taxonomicName>
has an irregular, though numerous, scattering of white spots across the dorsal and lateral surfaces, which intensify towards the posterior end of the animal. In comparison, the pale dorsal surfaces of 
<taxonomicName id="4C414D16FFA2FFA6FD72FCE1FC23FCC7" box="[764,942,823,848]" class="Elasmobranchii" family="Scyliorhinidae" genus="Atelomycterus" kingdom="Animalia" order="Carcharhiniformes" pageId="15" pageNumber="34" phylum="Chordata" rank="species" species="marmoratus">
<emphasis id="B935EA87FFA2FFA6FD72FCE1FC23FCC7" box="[764,942,823,848]" italics="true" pageId="15" pageNumber="34">A. marmoratus</emphasis>
</taxonomicName>
are dramatically reduced and form large irregular white spots on the lateral and dorsal surfaces (
<bibRefCitation id="EFD04B64FFA2FFA6FCB8FC88FB03FCEF" author="Compagno" box="[822,1166,862,888]" pageId="15" pageNumber="36" refString="Compagno, L. J. V. &amp; Stevens, J. D. (1993) Atelomycterus fasciatus n. sp., a new catshark (Chondrichthyes: Carcharhiniformes: Scyliorhinidae) from tropical Australia. Records of the Australian Museum, Vol. 45, 147 - 169." type="journal article" year="1993">Compagno &amp; Stevens, 1993</bibRefCitation>
). A sequence of white spots situated below the eye in 
<taxonomicName id="4C414D16FFA2FFA6FD83FC50FD30FC08" box="[525,701,902,927]" class="Elasmobranchii" family="Scyliorhinidae" genus="Atelomycterus" kingdom="Animalia" order="Carcharhiniformes" pageId="15" pageNumber="34" phylum="Chordata" rank="species" species="marmoratus">
<emphasis id="B935EA87FFA2FFA6FD83FC50FD30FC08" box="[525,701,902,927]" italics="true" pageId="15" pageNumber="34">A. marmoratus</emphasis>
</taxonomicName>
have also merged, forming a distinct white stripe through the gill slits. White spots are absent in 
<taxonomicName id="4C414D16FFA2FFA6FDB0FC7BFD46FC51" box="[574,715,941,966]" class="Elasmobranchii" family="Scyliorhinidae" genus="Atelomycterus" kingdom="Animalia" order="Carcharhiniformes" pageId="15" pageNumber="34" phylum="Chordata" rank="species" species="macleayi">
<emphasis id="B935EA87FFA2FFA6FDB0FC7BFD46FC51" box="[574,715,941,966]" italics="true" pageId="15" pageNumber="34">A. macleayi</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C414D16FFA2FFA6FC86FC7BFC1FFC51" box="[776,914,941,966]" class="Elasmobranchii" family="Scyliorhinidae" genus="Atelomycterus" kingdom="Animalia" order="Carcharhiniformes" pageId="15" pageNumber="34" phylum="Chordata" rank="species" species="baliensis">
<emphasis id="B935EA87FFA2FFA6FC86FC7BFC1FFC51" box="[776,914,941,966]" italics="true" pageId="15" pageNumber="34">A. baliensis</emphasis>
</taxonomicName>
; white spots are absent or fewer in number than black spots in 
<taxonomicName id="4C414D16FFA2FFA6FEF7FC03FD8FFC79" box="[377,514,981,1006]" class="Elasmobranchii" family="Scyliorhinidae" genus="Atelomycterus" kingdom="Animalia" order="Carcharhiniformes" pageId="15" pageNumber="34" phylum="Chordata" rank="species" species="fasciatus">
<emphasis id="B935EA87FFA2FFA6FEF7FC03FD8FFC79" box="[377,514,981,1006]" italics="true" pageId="15" pageNumber="34">A. fasciatus</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C35B651EFFA2FFA6FF48FC2AFAFFF94B" pageId="15" pageNumber="34" type="reference_group">
<paragraph id="8BFE3695FFA2FFA6FF48FC2AFDB0FB23" blockId="15.[151,1437,152,2033]" pageId="15" pageNumber="34">
<taxonomicName id="4C414D16FFA2FFA6FF48FC2AFE52FB82" ID-CoL="J8V5" box="[198,479,1020,1045]" class="Elasmobranchii" family="Scyliorhinidae" genus="Atelomycterus" kingdom="Animalia" order="Carcharhiniformes" pageId="15" pageNumber="34" phylum="Chordata" rank="species" species="macleayi">
<emphasis id="B935EA87FFA2FFA6FF48FC2AFE52FB82" box="[198,479,1020,1045]" italics="true" pageId="15" pageNumber="34">Atelomycterus macleayi</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C414D16FFA2FFA6FD96FC2AFD45FB82" ID-CoL="J8V6" box="[536,712,1020,1045]" class="Elasmobranchii" family="Scyliorhinidae" genus="Atelomycterus" kingdom="Animalia" order="Carcharhiniformes" pageId="15" pageNumber="34" phylum="Chordata" rank="species" species="marmoratus">
<emphasis id="B935EA87FFA2FFA6FD96FC2AFD45FB82" box="[536,712,1020,1045]" italics="true" pageId="15" pageNumber="34">A. marmoratus</emphasis>
</taxonomicName>
have higher total centra (TC), precaudal centra (PC) and precaudal diplospondylous centra (DP) count ranges than 
<emphasis id="B935EA87FFA2FFA6FC98FBF2FBC1FBAA" box="[790,1100,1060,1085]" italics="true" pageId="15" pageNumber="34">
<taxonomicName id="4C414D16FFA2FFA6FC98FBF2FC35FBAA" ID-CoL="J8V8" box="[790,952,1060,1085]" class="Elasmobranchii" family="Scyliorhinidae" genus="Atelomycterus" kingdom="Animalia" order="Carcharhiniformes" pageId="15" pageNumber="34" phylum="Chordata" rank="species" species="marnkalha">A. marnkalha</taxonomicName>
, 
<taxonomicName id="4C414D16FFA2FFA6FC4AFBF2FBC1FBAA" ID-CoL="J8V4" box="[964,1100,1060,1085]" class="Elasmobranchii" family="Scyliorhinidae" genus="Atelomycterus" kingdom="Animalia" order="Carcharhiniformes" pageId="15" pageNumber="34" phylum="Chordata" rank="species" species="fasciatus">A. fasciatus</taxonomicName>
</emphasis>
and 
<taxonomicName id="4C414D16FFA2FFA6FB0AFBF2FA81FBAA" ID-CoL="J8V2" box="[1156,1292,1060,1085]" class="Elasmobranchii" family="Scyliorhinidae" genus="Atelomycterus" kingdom="Animalia" order="Carcharhiniformes" pageId="15" pageNumber="34" phylum="Chordata" rank="species" species="baliensis">
<emphasis id="B935EA87FFA2FFA6FB0AFBF2FA81FBAA" box="[1156,1292,1060,1085]" italics="true" pageId="15" pageNumber="34">A. baliensis</emphasis>
</taxonomicName>
(TC = 161– 183 
<emphasis id="B935EA87FFA2FFA6FF42FB9DFF66FBF3" box="[204,235,1099,1124]" italics="true" pageId="15" pageNumber="34">vs.</emphasis>
147–163; PC = 106–132 
<emphasis id="B935EA87FFA2FFA6FDAEFB9DFDCDFBF3" box="[544,576,1099,1124]" italics="true" pageId="15" pageNumber="34">vs.</emphasis>
95–110; DP = 63–85 
<emphasis id="B935EA87FFA2FFA6FCC4FB9DFCE4FBF3" box="[842,873,1099,1124]" italics="true" pageId="15" pageNumber="34">vs.</emphasis>
53–69) (
<bibRefCitation id="EFD04B64FFA2FFA6FC5BFB9DFAA4FBF2" author="Compagno" box="[981,1321,1099,1125]" pageId="15" pageNumber="36" refString="Compagno, L. J. V. &amp; Stevens, J. D. (1993) Atelomycterus fasciatus n. sp., a new catshark (Chondrichthyes: Carcharhiniformes: Scyliorhinidae) from tropical Australia. Records of the Australian Museum, Vol. 45, 147 - 169." type="journal article" year="1993">Compagno &amp; Stevens, 1993</bibRefCitation>
; 
<bibRefCitation id="EFD04B64FFA2FFA6FAB8FB9DFE8BFB1B" author="White" pageId="15" pageNumber="36" refString="White, W. T., Last, P. R. &amp; Dharmadi. (2005) Description of a new species of catshark Atelomycterus baliensis (Carcharhiniformes: scyliorhinidae) from eastern Indonesia. Cybium, Vol. 29 (1), 33 - 40." type="journal article" year="2005">
White 
<emphasis id="B935EA87FFA2FFA6FA09FB9DFF38FB1B" italics="true" pageId="15" pageNumber="34">et al.</emphasis>
, 2005
</bibRefCitation>
). 
<taxonomicName id="4C414D16FFA2FFA6FE93FBA5FDC2FB1B" ID-CoL="J8V8" box="[285,591,1139,1164]" class="Elasmobranchii" family="Scyliorhinidae" genus="Atelomycterus" kingdom="Animalia" order="Carcharhiniformes" pageId="15" pageNumber="34" phylum="Chordata" rank="species" species="marnkalha">
<emphasis id="B935EA87FFA2FFA6FE93FBA5FDC2FB1B" box="[285,591,1139,1164]" italics="true" pageId="15" pageNumber="34">Atelomycterus marnkalha</emphasis>
</taxonomicName>
has the lowest PC count range (95–99) and the lowest minimum TC count (147–153) of the five species.
</paragraph>
<paragraph id="8BFE3695FFA2FFA6FF48FB17FACCFAC5" blockId="15.[151,1437,152,2033]" pageId="15" pageNumber="34">
<taxonomicName id="4C414D16FFA2FFA6FF48FB17FE6EFB4D" ID-CoL="J8V5" box="[198,483,1217,1242]" class="Elasmobranchii" family="Scyliorhinidae" genus="Atelomycterus" kingdom="Animalia" order="Carcharhiniformes" pageId="15" pageNumber="34" phylum="Chordata" rank="species" species="macleayi">
<emphasis id="B935EA87FFA2FFA6FF48FB17FE6EFB4D" box="[198,483,1217,1242]" italics="true" pageId="15" pageNumber="34">Atelomycterus macleayi</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C414D16FFA2FFA6FDADFB17FD5BFB4D" ID-CoL="J8V6" box="[547,726,1217,1242]" class="Elasmobranchii" family="Scyliorhinidae" genus="Atelomycterus" kingdom="Animalia" order="Carcharhiniformes" pageId="15" pageNumber="34" phylum="Chordata" rank="species" species="marmoratus">
<emphasis id="B935EA87FFA2FFA6FDADFB17FD5BFB4D" box="[547,726,1217,1242]" italics="true" pageId="15" pageNumber="34">A. marmoratus</emphasis>
</taxonomicName>
apparently attain a larger size (mature males 
<quantity id="4CB99B70FFA2FFA6FA89FB17FA11FB4C" box="[1287,1436,1217,1243]" metricMagnitude="-1" metricUnit="m" metricValue="5.5" metricValueMax="6.1899999999999995" metricValueMin="4.81" pageId="15" pageNumber="34" unit="mm" value="550.0" valueMax="619.0" valueMin="481.0">481–619mm</quantity>
TL) than 
<taxonomicName id="4C414D16FFA2FFA6FE86FB3FFE25FA95" ID-CoL="J8V8" box="[264,424,1257,1282]" class="Elasmobranchii" family="Scyliorhinidae" genus="Atelomycterus" kingdom="Animalia" order="Carcharhiniformes" pageId="15" pageNumber="34" phylum="Chordata" rank="species" species="marnkalha">
<emphasis id="B935EA87FFA2FFA6FE86FB3FFE25FA95" box="[264,424,1257,1282]" italics="true" pageId="15" pageNumber="34">A. marnkalha</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C414D16FFA2FFA6FE36FB3FFDCFFA95" ID-CoL="J8V4" box="[440,578,1257,1282]" class="Elasmobranchii" family="Scyliorhinidae" genus="Atelomycterus" kingdom="Animalia" order="Carcharhiniformes" pageId="15" pageNumber="34" phylum="Chordata" rank="species" species="fasciatus">
<emphasis id="B935EA87FFA2FFA6FE36FB3FFDCFFA95" box="[440,578,1257,1282]" italics="true" pageId="15" pageNumber="34">A. fasciatus</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C414D16FFA2FFA6FDF0FB3FFC8AFA95" ID-CoL="J8V2" box="[638,775,1257,1282]" class="Elasmobranchii" family="Scyliorhinidae" genus="Atelomycterus" kingdom="Animalia" order="Carcharhiniformes" pageId="15" pageNumber="34" phylum="Chordata" rank="species" species="baliensis">
<emphasis id="B935EA87FFA2FFA6FDF0FB3FFC8AFA95" box="[638,775,1257,1282]" italics="true" pageId="15" pageNumber="34">A. baliensis</emphasis>
</taxonomicName>
: mature males 
<quantity id="4CB99B70FFA2FFA6FC31FB3FFBD9FA94" box="[959,1108,1257,1283]" metricMagnitude="-1" metricUnit="m" metricValue="4.02" metricValueMax="4.18" metricValueMin="3.86" pageId="15" pageNumber="34" unit="mm" value="402.0" valueMax="418.0" valueMin="386.0">386–418mm</quantity>
TL; 
<quantity id="4CB99B70FFA2FFA6FB1EFB3FFAA9FA94" box="[1168,1316,1257,1283]" metricMagnitude="-1" metricUnit="m" metricValue="3.655" metricValueMax="4.02" metricValueMin="3.29" pageId="15" pageNumber="34" unit="mm" value="365.5" valueMax="402.0" valueMin="329.0">329–402mm</quantity>
TL; 
<quantity id="4CB99B70FFA2FFA6FAEFFB3FFF7DFABD" metricMagnitude="-1" metricUnit="m" metricValue="4.535" metricValueMax="4.74" metricValueMin="4.33" pageId="15" pageNumber="34" unit="mm" value="453.5" valueMax="474.0" valueMin="433.0">433– 474mm</quantity>
TL respectively (
<bibRefCitation id="EFD04B64FFA2FFA6FE4DFAC6FC98FABD" author="Compagno" box="[451,789,1296,1322]" pageId="15" pageNumber="36" refString="Compagno, L. J. V. &amp; Stevens, J. D. (1993) Atelomycterus fasciatus n. sp., a new catshark (Chondrichthyes: Carcharhiniformes: Scyliorhinidae) from tropical Australia. Records of the Australian Museum, Vol. 45, 147 - 169." type="journal article" year="1993">Compagno &amp; Stevens, 1993</bibRefCitation>
; 
<bibRefCitation id="EFD04B64FFA2FFA6FCACFAC6FC73FABD" author="White" box="[802,1022,1296,1322]" pageId="15" pageNumber="36" refString="White, W. T., Last, P. R. &amp; Dharmadi. (2005) Description of a new species of catshark Atelomycterus baliensis (Carcharhiniformes: scyliorhinidae) from eastern Indonesia. Cybium, Vol. 29 (1), 33 - 40." type="journal article" year="2005">
White 
<emphasis id="B935EA87FFA2FFA6FCFCFAC7FC2AFABD" box="[882,935,1297,1322]" italics="true" pageId="15" pageNumber="34">et al</emphasis>
., 2005
</bibRefCitation>
). Females of 
<taxonomicName id="4C414D16FFA2FFA6FB12FAC7FAB0FABD" ID-CoL="J8V8" box="[1180,1341,1297,1322]" class="Elasmobranchii" family="Scyliorhinidae" genus="Atelomycterus" kingdom="Animalia" order="Carcharhiniformes" pageId="15" pageNumber="34" phylum="Chordata" rank="species" species="marnkalha">
<emphasis id="B935EA87FFA2FFA6FB12FAC7FAB0FABD" box="[1180,1341,1297,1322]" italics="true" pageId="15" pageNumber="34">A. marnkalha</emphasis>
</taxonomicName>
attain a similar size to those of 
<taxonomicName id="4C414D16FFA2FFA6FE29FAEEFDBDFAC6" ID-CoL="J8V4" box="[423,560,1336,1361]" class="Elasmobranchii" family="Scyliorhinidae" genus="Atelomycterus" kingdom="Animalia" order="Carcharhiniformes" pageId="15" pageNumber="34" phylum="Chordata" rank="species" species="fasciatus">
<emphasis id="B935EA87FFA2FFA6FE29FAEEFDBDFAC6" box="[423,560,1336,1361]" italics="true" pageId="15" pageNumber="34">A. fasciatus</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C414D16FFA2FFA6FDE7FAEEFD7FFAC6" ID-CoL="J8V2" box="[617,754,1336,1361]" class="Elasmobranchii" family="Scyliorhinidae" genus="Atelomycterus" kingdom="Animalia" order="Carcharhiniformes" pageId="15" pageNumber="34" phylum="Chordata" rank="species" species="baliensis">
<emphasis id="B935EA87FFA2FFA6FDE7FAEEFD7FFAC6" box="[617,754,1336,1361]" italics="true" pageId="15" pageNumber="34">A. baliensis</emphasis>
</taxonomicName>
(
<bibRefCitation id="EFD04B64FFA2FFA6FC8FFAEEFBDDFAC5" author="Compagno" box="[769,1104,1336,1362]" pageId="15" pageNumber="36" refString="Compagno, L. J. V. &amp; Stevens, J. D. (1993) Atelomycterus fasciatus n. sp., a new catshark (Chondrichthyes: Carcharhiniformes: Scyliorhinidae) from tropical Australia. Records of the Australian Museum, Vol. 45, 147 - 169." type="journal article" year="1993">Compagno &amp; Stevens, 1993</bibRefCitation>
; 
<bibRefCitation id="EFD04B64FFA2FFA6FBD2FAEEFAB8FAC5" author="White" box="[1116,1333,1336,1362]" pageId="15" pageNumber="36" refString="White, W. T., Last, P. R. &amp; Dharmadi. (2005) Description of a new species of catshark Atelomycterus baliensis (Carcharhiniformes: scyliorhinidae) from eastern Indonesia. Cybium, Vol. 29 (1), 33 - 40." type="journal article" year="2005">
White 
<emphasis id="B935EA87FFA2FFA6FB25FAEEFB52FAC6" box="[1195,1247,1336,1361]" italics="true" pageId="15" pageNumber="34">et al</emphasis>
., 2005
</bibRefCitation>
).
</paragraph>
<paragraph id="8BFE3695FFA2FFA6FF48FAB6FDC0FA67" blockId="15.[151,1437,152,2033]" pageId="15" pageNumber="34">
<taxonomicName id="4C414D16FFA2FFA6FF48FAB6FE79FAEE" ID-CoL="J8V8" box="[198,500,1376,1401]" class="Elasmobranchii" family="Scyliorhinidae" genus="Atelomycterus" kingdom="Animalia" order="Carcharhiniformes" pageId="15" pageNumber="34" phylum="Chordata" rank="species" species="marnkalha">
<emphasis id="B935EA87FFA2FFA6FF48FAB6FE79FAEE" box="[198,500,1376,1401]" italics="true" pageId="15" pageNumber="34">Atelomycterus marnkalha</emphasis>
</taxonomicName>
is found further south than 
<taxonomicName id="4C414D16FFA2FFA6FCB6FAB6FC4DFAEE" ID-CoL="J8V4" box="[824,960,1376,1401]" class="Elasmobranchii" family="Scyliorhinidae" genus="Atelomycterus" kingdom="Animalia" order="Carcharhiniformes" pageId="15" pageNumber="34" phylum="Chordata" rank="species" species="fasciatus">
<emphasis id="B935EA87FFA2FFA6FCB6FAB6FC4DFAEE" box="[824,960,1376,1401]" italics="true" pageId="15" pageNumber="34">A. fasciatus</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C414D16FFA2FFA6FC77FAB6FB09FAEE" ID-CoL="J8V5" box="[1017,1156,1376,1401]" class="Elasmobranchii" family="Scyliorhinidae" genus="Atelomycterus" kingdom="Animalia" order="Carcharhiniformes" pageId="15" pageNumber="34" phylum="Chordata" rank="species" species="macleayi">
<emphasis id="B935EA87FFA2FFA6FC77FAB6FB09FAEE" box="[1017,1156,1376,1401]" italics="true" pageId="15" pageNumber="34">A. macleayi</emphasis>
</taxonomicName>
(
<bibRefCitation id="EFD04B64FFA2FFA6FB1CFA89FF58FA36" author="Compagno" pageId="15" pageNumber="36" refString="Compagno, L. J. V. &amp; Stevens, J. D. (1993) Atelomycterus fasciatus n. sp., a new catshark (Chondrichthyes: Carcharhiniformes: Scyliorhinidae) from tropical Australia. Records of the Australian Museum, Vol. 45, 147 - 169." type="journal article" year="1993">Compagno &amp; Stevens, 1993</bibRefCitation>
; 
<bibRefCitation id="EFD04B64FFA2FFA6FF6CFA51FE68FA36" author="Last" box="[226,485,1415,1441]" pageId="15" pageNumber="36" refString="Last, P. R. &amp; Stevens, J. D. (1994) Sharks and Rays of Australia, CSIRO: Australia, 513 pp." type="book" year="1994">Last &amp; Stevens, 1994</bibRefCitation>
), but may overlap with 
<taxonomicName id="4C414D16FFA2FFA6FC8CFA51FC03FA37" ID-CoL="J8V5" box="[770,910,1415,1440]" class="Elasmobranchii" family="Scyliorhinidae" genus="Atelomycterus" kingdom="Animalia" order="Carcharhiniformes" pageId="15" pageNumber="34" phylum="Chordata" rank="species" species="macleayi">
<emphasis id="B935EA87FFA2FFA6FC8CFA51FC03FA37" box="[770,910,1415,1440]" italics="true" pageId="15" pageNumber="34">A. macleayi</emphasis>
</taxonomicName>
in the Gulf of Carpentaria and in the Torres Strait. 
<taxonomicName id="4C414D16FFA2FFA6FF6AFA78FDACFA50" ID-CoL="J8V6" box="[228,545,1454,1479]" class="Elasmobranchii" family="Scyliorhinidae" genus="Atelomycterus" kingdom="Animalia" order="Carcharhiniformes" pageId="15" pageNumber="34" phylum="Chordata" rank="species" species="marmoratus">
<emphasis id="B935EA87FFA2FFA6FF6AFA78FDACFA50" box="[228,545,1454,1479]" italics="true" pageId="15" pageNumber="34">Atelomycterus marmoratus</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C414D16FFA2FFA6FDD7FA78FD6CFA50" ID-CoL="J8V2" box="[601,737,1454,1479]" class="Elasmobranchii" family="Scyliorhinidae" genus="Atelomycterus" kingdom="Animalia" order="Carcharhiniformes" pageId="15" pageNumber="34" phylum="Chordata" rank="species" species="baliensis">
<emphasis id="B935EA87FFA2FFA6FDD7FA78FD6CFA50" box="[601,737,1454,1479]" italics="true" pageId="15" pageNumber="34">A. baliensis</emphasis>
</taxonomicName>
have not been recorded from Australian waters (
<bibRefCitation id="EFD04B64FFA2FFA6FA97FA78FED1FA67" author="Compagno" pageId="15" pageNumber="36" refString="Compagno, L. J. V. &amp; Stevens, J. D. (1993) Atelomycterus fasciatus n. sp., a new catshark (Chondrichthyes: Carcharhiniformes: Scyliorhinidae) from tropical Australia. Records of the Australian Museum, Vol. 45, 147 - 169." type="journal article" year="1993">Compagno &amp; Stevens, 1993</bibRefCitation>
; 
<bibRefCitation id="EFD04B64FFA2FFA6FEE6FA00FDCDFA67" author="White" box="[360,576,1494,1520]" pageId="15" pageNumber="36" refString="White, W. T., Last, P. R. &amp; Dharmadi. (2005) Description of a new species of catshark Atelomycterus baliensis (Carcharhiniformes: scyliorhinidae) from eastern Indonesia. Cybium, Vol. 29 (1), 33 - 40." type="journal article" year="2005">
White 
<emphasis id="B935EA87FFA2FFA6FE39FA00FE7FFA78" box="[439,498,1494,1519]" italics="true" pageId="15" pageNumber="34">et al.</emphasis>
, 2005
</bibRefCitation>
).
</paragraph>
<paragraph id="8BFE3695FFA2FFA6FF48FA2BFAFFF94B" blockId="15.[151,1437,152,2033]" pageId="15" pageNumber="34">
<emphasis id="B935EA87FFA2FFA6FF48FA2BFE17F980" bold="true" box="[198,410,1533,1559]" pageId="15" pageNumber="34">MDS ordination.</emphasis>
Within the MDS ordination, 
<taxonomicName id="4C414D16FFA2FFA6FD7BFA2BFC18F981" ID-CoL="J8V8" box="[757,917,1533,1558]" class="Elasmobranchii" family="Scyliorhinidae" genus="Atelomycterus" kingdom="Animalia" order="Carcharhiniformes" pageId="15" pageNumber="34" phylum="Chordata" rank="species" species="marnkalha">
<emphasis id="B935EA87FFA2FFA6FD7BFA2BFC18F981" box="[757,917,1533,1558]" italics="true" pageId="15" pageNumber="34">A. marnkalha</emphasis>
</taxonomicName>
specimens were distributed around the central-left side of the plot, with 
<taxonomicName id="4C414D16FFA2FFA6FE7AF9F3FDF0F9A9" ID-CoL="J8V4" box="[500,637,1573,1598]" class="Elasmobranchii" family="Scyliorhinidae" genus="Atelomycterus" kingdom="Animalia" order="Carcharhiniformes" pageId="15" pageNumber="34" phylum="Chordata" rank="species" species="fasciatus">
<emphasis id="B935EA87FFA2FFA6FE7AF9F3FDF0F9A9" box="[500,637,1573,1598]" italics="true" pageId="15" pageNumber="34">A. fasciatus</emphasis>
</taxonomicName>
specimens, producing a relatively tight group at the top-left corner. As expected, the 
<taxonomicName id="4C414D16FFA2FFA6FEEEF99AFD8DF9F2" ID-CoL="J8V8" box="[352,512,1612,1637]" class="Elasmobranchii" family="Scyliorhinidae" genus="Atelomycterus" kingdom="Animalia" order="Carcharhiniformes" pageId="15" pageNumber="34" phylum="Chordata" rank="species" species="marnkalha">
<emphasis id="B935EA87FFA2FFA6FEEEF99AFD8DF9F2" box="[352,512,1612,1637]" italics="true" pageId="15" pageNumber="34">A. marnkalha</emphasis>
</taxonomicName>
sample plot was closer to 
<taxonomicName id="4C414D16FFA2FFA6FCB8F99AFC32F9F2" ID-CoL="J8V4" box="[822,959,1612,1637]" class="Elasmobranchii" family="Scyliorhinidae" genus="Atelomycterus" kingdom="Animalia" order="Carcharhiniformes" pageId="15" pageNumber="34" phylum="Chordata" rank="species" species="fasciatus">
<emphasis id="B935EA87FFA2FFA6FCB8F99AFC32F9F2" box="[822,959,1612,1637]" italics="true" pageId="15" pageNumber="34">A. fasciatus</emphasis>
</taxonomicName>
than the other three species (
<figureCitation id="137A2A10FFA2FFA6FA9AF99AFAD8F9F1" box="[1300,1365,1612,1638]" captionStart="FIGURE 9" captionStartId="16.[151,255,745,769]" captionTargetBox="[460,1131,370,717]" captionTargetId="figure@16.[454,1135,359,721]" captionTargetPageId="16" captionText="FIGURE 9. Non-metric multidimensional scaling ordination of atelomycterine morphometric percentages." httpUri="https://zenodo.org/record/177433/files/figure.png" pageId="15" pageNumber="34">Fig 9</figureCitation>
). 
<taxonomicName id="4C414D16FFA2FFA6FAE5F99AFE04F91A" ID-CoL="J8V2" class="Elasmobranchii" family="Scyliorhinidae" genus="Atelomycterus" kingdom="Animalia" order="Carcharhiniformes" pageId="15" pageNumber="34" phylum="Chordata" rank="species" species="baliensis">
<emphasis id="B935EA87FFA2FFA6FAE5F99AFE04F91A" italics="true" pageId="15" pageNumber="34">Atelomycterus baliensis</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C414D16FFA2FFA6FE46F9A2FDF7F91A" ID-CoL="J8V6" box="[456,634,1652,1677]" class="Elasmobranchii" family="Scyliorhinidae" genus="Atelomycterus" kingdom="Animalia" order="Carcharhiniformes" pageId="15" pageNumber="34" phylum="Chordata" rank="species" species="marmoratus">
<emphasis id="B935EA87FFA2FFA6FE46F9A2FDF7F91A" box="[456,634,1652,1677]" italics="true" pageId="15" pageNumber="34">A. marmoratus</emphasis>
</taxonomicName>
were located in a relatively confined area in the lower half of the plot, indicating the two species are morphologically similar. This is consistent with results obtained by 
<bibRefCitation id="EFD04B64FFA2FFA6FADAF94DFEA6F94B" pageId="15" pageNumber="36" refString="White, W. T., Last, P. R. &amp; Dharmadi. (2005) Description of a new species of catshark Atelomycterus baliensis (Carcharhiniformes: scyliorhinidae) from eastern Indonesia. Cybium, Vol. 29 (1), 33 - 40." type="journal article">
White 
<emphasis id="B935EA87FFA2FFA6FF19F915FF5CF94B" box="[151,209,1731,1756]" italics="true" pageId="15" pageNumber="34">et al.</emphasis>
(2005)
</bibRefCitation>
. The two 
<taxonomicName id="4C414D16FFA2FFA6FE13F915FDA5F94B" ID-CoL="J8V5" box="[413,552,1731,1756]" class="Elasmobranchii" family="Scyliorhinidae" genus="Atelomycterus" kingdom="Animalia" order="Carcharhiniformes" pageId="15" pageNumber="34" phylum="Chordata" rank="species" species="macleayi">
<emphasis id="B935EA87FFA2FFA6FE13F915FDA5F94B" box="[413,552,1731,1756]" italics="true" pageId="15" pageNumber="34">A. macleayi</emphasis>
</taxonomicName>
specimens were located to the right of the plot on the outer extremities.
</paragraph>
</subSubSection>
<subSubSection id="C35B651EFFA2FFB9FF48F93CFD09FEBD" lastPageId="16" lastPageNumber="35" pageId="15" pageNumber="34" type="description">
<paragraph id="8BFE3695FFA2FFA6FF48F93CFC86F866" blockId="15.[151,1437,152,2033]" pageId="15" pageNumber="34">
The resulting ANOSIM demonstrated that all five species were significantly different overall (P&lt;0.01; global R statistic = 0.826). Individual pairwise comparisons between 
<taxonomicName id="4C414D16FFA2FFA6FC5FF8C7FBFFF8BD" box="[977,1138,1809,1834]" class="Elasmobranchii" family="Scyliorhinidae" genus="Atelomycterus" kingdom="Animalia" order="Carcharhiniformes" pageId="15" pageNumber="34" phylum="Chordata" rank="species" species="marnkalha">
<emphasis id="B935EA87FFA2FFA6FC5FF8C7FBFFF8BD" box="[977,1138,1809,1834]" italics="true" pageId="15" pageNumber="34">A. marnkalha</emphasis>
</taxonomicName>
and the four other atelomycterine species all showed significant differences; 
<emphasis id="B935EA87FFA2FFA6FC90F8EFFBFDF8C5" box="[798,1136,1849,1874]" italics="true" pageId="15" pageNumber="34">
<taxonomicName id="4C414D16FFA2FFA6FC90F8EFFC23F8C5" box="[798,942,1849,1874]" class="Elasmobranchii" family="Scyliorhinidae" genus="Atelomycterus" kingdom="Animalia" order="Carcharhiniformes" pageId="15" pageNumber="34" phylum="Chordata" rank="species" species="fasciatus">A. fasciatus</taxonomicName>
, 
<taxonomicName id="4C414D16FFA2FFA6FC33F8EFFBFDF8C5" box="[957,1136,1849,1874]" class="Elasmobranchii" family="Scyliorhinidae" genus="Atelomycterus" kingdom="Animalia" order="Carcharhiniformes" pageId="15" pageNumber="34" phylum="Chordata" rank="species" species="marmoratus">A. marmoratus</taxonomicName>
</emphasis>
P&lt;0.01; 
<emphasis id="B935EA87FFA2FFA6FB6DF8EFFF72F8EE" italics="true" pageId="15" pageNumber="34">
<taxonomicName id="4C414D16FFA2FFA6FB6DF8EFFAF9F8C5" box="[1251,1396,1849,1874]" class="Elasmobranchii" family="Scyliorhinidae" genus="Atelomycterus" kingdom="Animalia" order="Carcharhiniformes" pageId="15" pageNumber="34" phylum="Chordata" rank="species" species="macleayi">A. macleayi</taxonomicName>
, 
<taxonomicName id="4C414D16FFA2FFA6FA0DF8EFFF72F8EE" class="Elasmobranchii" family="Scyliorhinidae" genus="Atelomycterus" kingdom="Animalia" order="Carcharhiniformes" pageId="15" pageNumber="34" phylum="Chordata" rank="species" species="baliensis">A. baliensis</taxonomicName>
</emphasis>
P&lt;0.05. The morphometric measurements identified by SIMPER as most responsible for the differences seen between 
<taxonomicName id="4C414D16FFA2FFA6FE1FF85EFDB8F836" box="[401,565,1928,1953]" class="Elasmobranchii" family="Scyliorhinidae" genus="Atelomycterus" kingdom="Animalia" order="Carcharhiniformes" pageId="15" pageNumber="34" phylum="Chordata" rank="species" species="marnkalha">
<emphasis id="B935EA87FFA2FFA6FE1FF85EFDB8F836" box="[401,565,1928,1953]" italics="true" pageId="15" pageNumber="34">A. marnkalha</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C414D16FFA2FFA6FDF8F85EFC8EF836" box="[630,771,1928,1953]" class="Elasmobranchii" family="Scyliorhinidae" genus="Atelomycterus" kingdom="Animalia" order="Carcharhiniformes" pageId="15" pageNumber="34" phylum="Chordata" rank="species" species="fasciatus">
<emphasis id="B935EA87FFA2FFA6FDF8F85EFC8EF836" box="[630,771,1928,1953]" italics="true" pageId="15" pageNumber="34">A. fasciatus</emphasis>
</taxonomicName>
were the anal-caudal space (ACS) length and preanal (PAL) length. The corresponding morphometric results showed that both the ACS length and PAL length for 
<taxonomicName id="4C414D16FFA2FFA6FF19F801FEBBF867" box="[151,310,2007,2032]" class="Elasmobranchii" family="Scyliorhinidae" genus="Atelomycterus" kingdom="Animalia" order="Carcharhiniformes" pageId="15" pageNumber="34" phylum="Chordata" rank="species" species="marnkalha">
<emphasis id="B935EA87FFA2FFA6FF19F801FEBBF867" box="[151,310,2007,2032]" italics="true" pageId="15" pageNumber="34">A. marnkalha</emphasis>
</taxonomicName>
was smaller than 
<taxonomicName id="4C414D16FFA2FFA6FD84F801FD1FF867" box="[522,658,2007,2032]" class="Elasmobranchii" family="Scyliorhinidae" genus="Atelomycterus" kingdom="Animalia" order="Carcharhiniformes" pageId="15" pageNumber="34" phylum="Chordata" rank="species" species="fasciatus">
<emphasis id="B935EA87FFA2FFA6FD84F801FD1FF867" box="[522,658,2007,2032]" italics="true" pageId="15" pageNumber="34">A. fasciatus</emphasis>
</taxonomicName>
(
<tableCitation id="C6C3032EFFA2FFA6FD2CF801FD74F866" box="[674,761,2007,2033]" captionStart="TABLE 1" captionStartId="1.[151,239,1066,1090]" captionText="TABLE 1. Morphometric data expressed as % of TL for the Atelomycterus marnkalha n. sp. holotype, paratype size ranges and A. fasciatus comparisons." pageId="15" pageNumber="34">Table 1</tableCitation>
).
</paragraph>
<paragraph id="8BFE3695FFBDFFB9FF48FF4EFD09FEBD" blockId="16.[151,1437,152,298]" pageId="16" pageNumber="35">
The results obtained from the MDS ordination, ANOSIM, and SIMPER analysis further supports the morphometric results and the reidentification of AMS I.15557-003, CSIRO H1118-01 and AMS I.21842-001, 10°S37´S 
<geoCoordinate id="EE755052FFBDFFB9FE80FF3EFE0DFE95" box="[270,384,231,258]" direction="east" orientation="longitude" pageId="16" pageNumber="35" precision="925" value="140.03334">140°02´E</geoCoordinate>
) as 
<taxonomicName id="4C414D16FFBDFFB9FE21FF3EFDC3FE96" box="[431,590,232,257]" class="Elasmobranchii" family="Scyliorhinidae" genus="Atelomycterus" kingdom="Animalia" order="Carcharhiniformes" pageId="16" pageNumber="35" phylum="Chordata" rank="species" species="marnkalha">
<emphasis id="B935EA87FFBDFFB9FE21FF3EFDC3FE96" box="[431,590,232,257]" italics="true" pageId="16" pageNumber="35">A. marnkalha</emphasis>
</taxonomicName>
. Given these samples are some of the earliest records of the species they have also been included in the 
<typeStatus id="54FA8837FFBDFFB9FD8EFEC6FDBFFEBD" box="[512,562,272,298]" pageId="16" pageNumber="35">type</typeStatus>
series.
</paragraph>
</subSubSection>
</treatment>
</document>