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<document id="A7DEC5C48384CF0D4F299D2A39E6CCFB" ID-CLB-Dataset="291528" ID-DOI="10.11646/zootaxa.5428.2.5" ID-GBIF-Dataset="b0a98796-768e-43e2-be6b-1f79c24b9227" ID-ISSN="1175-5326" ID-Zenodo-Dep="10847635" ID-ZooBank="135D8285-BEC0-467E-BCD5-4DA116C6DFBC" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="valdenar" IM.materialsCitations_approvedBy="valdenar" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="valdenar" checkinTime="1711009984219" checkinUser="plazi" docAuthor="Janák, Jiří &amp; Baňař, Petr" docDate="2024" docId="03E8307A6655027F9992FF64C472F8E3" docLanguage="en" docName="zootaxa.5428.2.5.pdf" docOrigin="Zootaxa 5428 (2)" docSource="http://dx.doi.org/10.11646/zootaxa.5428.2.5" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Paulianidia vohimana Janak &amp; Banar 2024, sp. nov." docType="treatment" docVersion="5" lastPageNumber="260" masterDocId="FFD14802665202789905FFF3C74BFFCA" masterDocTitle="Four new species and new records of the genus Paulianidia of Madagascar (Coleoptera: Staphylinidae: Paederinae)" masterLastPageNumber="268" masterPageNumber="253" pageNumber="260" updateTime="1711133149338" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="C3EFE1B88C89D0DAAA9F06B91A879282" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="6921E27593707737C2D5EF42EE153E5E">
<mods:title id="F1C9406B063080AE96476A646299510F">Four new species and new records of the genus Paulianidia of Madagascar (Coleoptera: Staphylinidae: Paederinae)</mods:title>
</mods:titleInfo>
<mods:name id="F5E2DEBD7F3348ED997354D71EACC53B" type="personal">
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<mods:roleTerm id="DDBE3FBD31A6D01853F8E5A37258BF36">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="E61C9539057E6EEE6613E0BBD3EA1DD7">Janák, Jiří</mods:namePart>
<mods:affiliation id="E67762BFCC308C94D32BCB723016BB5C">CZ- 417 62 Rtyně nad Bílinou, č. 4, Czech Republic</mods:affiliation>
</mods:name>
<mods:name id="A88F5C83FED0964B142681D7318E74FF" type="personal">
<mods:role id="EA5760F7C14308D9CEB4FBD690F4E444">
<mods:roleTerm id="3B7113EA64C55543D125C4762E515BF4">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="5696AD02B86E236E686A12D951C27641">Baňař, Petr</mods:namePart>
<mods:affiliation id="A790214E94B22885DA281F870E80AD93">Moravian Museum, Department of Entomology, Hviezdoslavova 29 a, Brno, CZ- 627 00, Czech Republic</mods:affiliation>
</mods:name>
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<mods:title id="C160FF53FC8C40A525AF792094B62BCA">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="50BA89943B28B18966CE97A54000A0E7">
<mods:date id="9042A9781CD0637619495F32C6745F0A">2024</mods:date>
<mods:detail id="5A698DAB4C5B35FF9E36A1A663C92742" type="pubDate">
<mods:number id="2DF57E762D485B4C9E17570419BFA694">2024-03-21</mods:number>
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<mods:number id="DE6901641BB351E0031A99E2CC235D14">5428</mods:number>
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<mods:number id="9D937A9EFA25D3CE441B4300C1A9C510">2</mods:number>
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<mods:start id="D0CB7F66B9549FA641D4D0AA6191080B">253</mods:start>
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<mods:location id="5CBEDDB545009649A2E698F1536B9848">
<mods:url id="846DA8B88A9DF58C960793D352C7231D">http://dx.doi.org/10.11646/zootaxa.5428.2.5</mods:url>
</mods:location>
<mods:classification id="48D66142D385C9A6168EF6C0FD9DC220">journal article</mods:classification>
<mods:identifier id="E0C592B47548F94A6674D69826242C63" type="CLB-Dataset">291528</mods:identifier>
<mods:identifier id="592C26BB7991E6BFF2233473E64279E6" type="DOI">10.11646/zootaxa.5428.2.5</mods:identifier>
<mods:identifier id="F2F6A76F1A16DABEC67F0D3A3827330A" type="GBIF-Dataset">b0a98796-768e-43e2-be6b-1f79c24b9227</mods:identifier>
<mods:identifier id="4801723452F6780F624D78D1577AB365" type="ISSN">1175-5326</mods:identifier>
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</mods:mods>
<treatment id="03E8307A6655027F9992FF64C472F8E3" ID-DOI="http://doi.org/10.5281/zenodo.10847577" ID-Zenodo-Dep="10847577" LSID="urn:lsid:plazi:treatment:03E8307A6655027F9992FF64C472F8E3" httpUri="http://treatment.plazi.org/id/03E8307A6655027F9992FF64C472F8E3" lastPageNumber="260" pageId="7" pageNumber="260">
<subSubSection id="C35BD2E76655027F9992FF64C591FF7B" box="[151,730,150,178]" pageId="7" pageNumber="260" type="nomenclature">
<paragraph id="8BFE816C6655027F9992FF64C591FF7B" blockId="7.[151,730,150,213]" box="[151,730,150,178]" pageId="7" pageNumber="260">
<heading id="D0B636006655027F9992FF64C591FF7B" bold="true" box="[151,730,150,178]" fontSize="11" level="1" pageId="7" pageNumber="260" reason="1">
<emphasis id="B9355D7E6655027F9992FF64C591FF7B" bold="true" box="[151,730,150,178]" pageId="7" pageNumber="260">
<taxonomicName id="4C41FAEF6655027F9992FF64C53BFF7B" authority="Janak &amp; Banar" authorityName="Janak &amp; Banar" authorityYear="2024" box="[151,624,150,178]" class="Insecta" family="Staphylinidae" genus="Paulianidia" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="260" phylum="Arthropoda" rank="species" species="vohimana" status="sp. nov.">
<emphasis id="B9355D7E6655027F9992FF64C6EEFF78" bold="true" box="[151,421,151,178]" italics="true" pageId="7" pageNumber="260">Paulianidia vohimana</emphasis>
Janák &amp; Baňař
</taxonomicName>
,
<taxonomicNameLabel id="A206E0056655027F9B78FF64C591FF7B" box="[637,730,151,177]" pageId="7" pageNumber="260" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C35BD2E76655027F9992FF48C62CFF1F" box="[151,359,187,213]" pageId="7" pageNumber="260" type="description">
<paragraph id="8BFE816C6655027F9992FF48C62CFF1F" blockId="7.[151,730,150,213]" box="[151,359,187,213]" pageId="7" pageNumber="260">
(
<figureCitation id="137A9DE96655027F999AFF48C663FF1F" box="[159,296,187,213]" captionStart="FIGURES 31–44" captionStartId="6.[151,264,1784,1808]" captionTargetBox="[185,1404,196,1741]" captionTargetId="figure-34@6.[171,1416,181,1760]" captionTargetPageId="6" captionText="FIGURES 31–44. 31–37: Paulianidia valohoaka sp. nov., holotype: 31, habitus; 32, forebody; 33–35, aedeagus (33, ventral; 34, 1/3 lateral; 35, lateral); 36, male sternite VII; 37, male sternite VIII. 38–44: P. vohimana sp. nov., holotype: 38, habitus; 39, forebody; 40–42, aedeagus (40, ventral; 41, 1/3 lateral; 42, lateral); 43, male sternite VII; 44, male sternite VIII. Scales 1 mm: 31–32, 38–39; 0.5 mm: 33–35, 36–37, 40–42, 43–44." figureDoi="http://doi.org/10.5281/zenodo.10847644" httpUri="https://zenodo.org/record/10847644/files/figure.png" pageId="7" pageNumber="260">Figs. 38–44</figureCitation>
,
<figureCitation id="137A9DE96655027F9836FF48C615FF1F" box="[307,350,187,213]" captionStart="FIGURES 116–122" captionStartId="12.[151,264,1780,1804]" captionTargetBox="[180,1408,194,1743]" captionTargetId="figure-34@12.[167,1421,181,1756]" captionText="FIGURES 116–122. 116–120: Paulianidia steineri Lecoq, Andasibe: 116–118, aedeagus (116, ventral; 117, 1/3 lateral; 118, lateral); 119, male sternite VII; 120, male sternite VIII. Scale 0.5 mm: 116–120. 121: Distribution of P. densepunctata sp. nov., (rhombus), P. valohoaka sp. nov. (star), P. tanala sp. nov. and P. vohimana sp. nov. (rectangle). 122: Distribution of Paulianidia species in Madagascar." figureDoi="http://doi.org/10.5281/zenodo.10854991" httpUri="https://zenodo.org/record/10854991/files/figure.png" pageId="7" pageNumber="260">121</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C35BD2E76655027F9992FEF0C336FDAB" pageId="7" pageNumber="260" type="materials_examined">
<paragraph id="8BFE816C6655027F9992FEF0C403FED4" blockId="7.[151,1437,259,1834]" box="[151,840,259,286]" pageId="7" pageNumber="260">
<materialsCitation id="3B298B316655027F9992FEF0C40FFED4" box="[151,836,259,286]" country="Madagascar" location="Vohimana Experimental Reserve" pageId="7" pageNumber="260" specimenCount="1" typeStatus="holotype">
<emphasis id="B9355D7E6655027F9992FEF0C67DFED7" bold="true" box="[151,310,259,285]" pageId="7" pageNumber="260">
<typeStatus id="54FA3FCE6655027F9992FEF0C799FED7" box="[151,210,259,285]" pageId="7" pageNumber="260">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="F356C1FC6655027F9839FEF0C68EFED7" box="[316,453,259,285]" name="Madagascar" pageId="7" pageNumber="260">Madagascar</collectingCountry>
,
<location id="8E9ED7B76655027F98D5FEF0C40FFED4" LSID="urn:lsid:plazi:treatment:03E8307A6655027F9992FF64C472F8E3:8E9ED7B76655027F98D5FEF0C40FFED4" box="[464,836,259,286]" country="Madagascar" name="Vohimana Experimental Reserve" pageId="7" pageNumber="260">Vohimana Experimental Reserve</location>
</materialsCitation>
.
</paragraph>
<paragraph id="8BFE816C6655027F99C2FED4C336FDAB" blockId="7.[151,1437,259,1834]" pageId="7" pageNumber="260">
<emphasis id="B9355D7E6655027F99C2FED4C638FE8B" bold="true" box="[199,371,295,321]" pageId="7" pageNumber="260">Type material.</emphasis>
<materialsCitation id="3B298B316655027F987DFED4C6C0FE67" collectingDate="2012-08" collectingDateMax="2012-08-22" collectingDateMin="2012-08" collectionCode="MMBC, JJRC" collectorName="L. S. Rahanitriniaina &amp; E. M. Rabotoson" country="Madagascar" determinerName="Banar" elevation="891" latitude="-18.92886" location="Circuit" longLatPrecision="1" longitude="48.499947" pageId="7" pageNumber="260" specimenCount="1" specimenCount-female="4" specimenCount-male="1" typeStatus="holotype">
<emphasis id="B9355D7E6655027F987DFED4C6AEFE8B" bold="true" box="[376,485,295,321]" pageId="7" pageNumber="260">
<typeStatus id="54FA3FCE6655027F987DFED4C6AEFE8B" box="[376,485,295,321]" pageId="7" pageNumber="260" type="holotype">Holotype</typeStatus>
</emphasis>
<specimenCount id="9D474AE56655027F98EFFEDBC549FE88" box="[490,514,296,322]" count="1" pageId="7" pageNumber="260" type="male">♂</specimenCount>
:
<emphasis id="B9355D7E6655027F9B0EFED4C593FE8B" bold="true" box="[523,728,295,321]" pageId="7" pageNumber="260">
<collectingCountry id="F356C1FC6655027F9B0EFED4C593FE8B" box="[523,728,295,321]" name="Madagascar" pageId="7" pageNumber="260">MADAGASCAR</collectingCountry>
</emphasis>
: “VOH/
<date id="FFFFA7AC6655027F9A32FED4C4EEFE8B" box="[823,933,295,321]" pageId="7" pageNumber="260" value="2012-08">
<collectingDate id="EFBB5E446655027F9A32FED4C4EEFE8B" box="[823,933,295,321]" pageId="7" pageNumber="260" value="2012-08">Aug.2012</collectingDate>
</date>
/09 MADAGASCAR, Réserv. Expérimentale de VOHIMANA, „
<location id="8E9ED7B76655027F984EFEB8C6D7FEAF" LSID="urn:lsid:plazi:treatment:03E8307A6655027F9992FF64C472F8E3:8E9ED7B76655027F984EFEB8C6D7FEAF" box="[331,412,331,357]" country="Madagascar" latitude="-18.92886" longLatPrecision="1" longitude="48.499947" name="Circuit" pageId="7" pageNumber="260">Circuit</location>
5, PK 15“,
<date id="FFFFA7AC6655027F9B24FEB8C5E1FEAF" box="[545,682,331,357]" pageId="7" pageNumber="260" value="2012-08-22">
<collectingDate id="EFBB5E446655027F9B24FEB8C5E1FEAF" box="[545,682,331,357]" pageId="7" pageNumber="260" value="2012-08-22">22.viii.2012</collectingDate>
</date>
” // “
<geoCoordinate id="EE75E7AB6655027F9BDBFEB8C432FEAC" box="[734,889,331,358]" degrees="18" direction="south" minutes="55" orientation="latitude" pageId="7" pageNumber="260" precision="1" seconds="43.9" value="-18.92886">S18°55’43.9”</geoCoordinate>
<geoCoordinate id="EE75E7AB6655027F9A85FEBFC354FEAC" box="[896,1055,331,358]" degrees="48" direction="east" minutes="29" orientation="longitude" pageId="7" pageNumber="260" precision="1" seconds="59.8" value="48.499947">E48°29’59.8”</geoCoordinate>
; 891m, sifting litter; Winkler app. extr.;
<collectorName id="26B4E4BA6655027F99D3FE9CC6F3FE43" box="[214,440,367,393]" pageId="7" pageNumber="260">L.S. Rahanitriniaina</collectorName>
&amp;
<collectorName id="26B4E4BA6655027F98DFFE83C5DBFE43" box="[474,656,367,394]" pageId="7" pageNumber="260">E.M. Rabotoson</collectorName>
lgt.” // “HOLOTYPUS
<taxonomicName id="4C41FAEF6655027F9A99FE9CC3C5FE43" authority="Janák &amp; Baňař, 2024" authorityName="Janák &amp; Baňař" authorityYear="2024" box="[924,1166,367,393]" class="Insecta" family="Staphylinidae" genus="Paulianidia" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="260" phylum="Arthropoda" rank="species" species="vohimana" status="sp. nov.">Paulianidia vohimana</taxonomicName>
<taxonomicNameLabel id="A206E0056655027F9D90FE83C3A6FE40" box="[1173,1261,368,394]" pageId="7" pageNumber="260" rank="species">sp. nov.</taxonomicNameLabel>
, Janák &amp; Baňař det. 2023” (
<collectionCode id="ED5019A96655027F9824FE60C634FE67" box="[289,383,403,429]" country="Czech Republic" httpUri="http://grbio.org/cool/w50r-ge1v" name="Moravske Muzeum [Moravian Museum]" pageId="7" pageNumber="260">MMBC</collectionCode>
).
</materialsCitation>
<materialsCitation id="3B298B316655027F9890FE60C409FE3F" collectingDate="2012-08" collectingDateMax="2012-08-22" collectingDateMin="2012-08" collectionCode="MMBC, JJRC" country="Madagascar" determinerName="Banar" elevation="965" latitude="-18.917555" longLatPrecision="1" longitude="48.49361" pageId="7" pageNumber="260" specimenCount="4" specimenCount-female="4" typeStatus="holotype">
<emphasis id="B9355D7E6655027F9890FE60C546FE67" bold="true" box="[405,525,403,429]" pageId="7" pageNumber="260">
<typeStatus id="54FA3FCE6655027F9890FE60C546FE67" box="[405,525,403,429]" pageId="7" pageNumber="260" type="paratype">Paratypes</typeStatus>
</emphasis>
:
<specimenCount id="9D474AE56655027F9B1AFE60C505FE64" box="[543,590,403,430]" count="1" pageId="7" pageNumber="260" type="female">1 ♀</specimenCount>
: “
<specimenCount id="9D474AE56655027F9B69FE60C535FE67" box="[620,638,403,429]" count="1" pageId="7" pageNumber="260" type="female">♀</specimenCount>
” // “VOH/
<date id="FFFFA7AC6655027F9A07FE60C43AFE67" box="[770,881,403,429]" pageId="7" pageNumber="260" value="2012-08">
<collectingDate id="EFBB5E446655027F9A07FE60C43AFE67" box="[770,881,403,429]" pageId="7" pageNumber="260" value="2012-08">Aug.2012</collectingDate>
</date>
/01 MADAGASCAR, Réserv. Expérimentale de VOHIMANA, „Circuit 2, borne 18“,
<date id="FFFFA7AC6655027F9B1BFE44C5EDFE1B" box="[542,678,439,465]" pageId="7" pageNumber="260" value="2012-08-18">
<collectingDate id="EFBB5E446655027F9B1BFE44C5EDFE1B" box="[542,678,439,465]" pageId="7" pageNumber="260" value="2012-08-18">18.viii.2012</collectingDate>
</date>
” // “
<geoCoordinate id="EE75E7AB6655027F9BD9FE44C43DFE18" box="[732,886,439,466]" degrees="18" direction="south" minutes="55" orientation="latitude" pageId="7" pageNumber="260" precision="1" seconds="03.2" value="-18.917555">S18°55’03.2”</geoCoordinate>
<geoCoordinate id="EE75E7AB6655027F9A7BFE4BC356FE1B" box="[894,1053,439,466]" degrees="48" direction="east" minutes="29" orientation="longitude" pageId="7" pageNumber="260" precision="1" seconds="37.0" value="48.49361">E48°29’37.0”</geoCoordinate>
; 965m, sifting litter; Winkler app. extr.;
<collectorName id="26B4E4BA6655027F99DEFE28C68AFE3F" box="[219,449,475,501]" pageId="7" pageNumber="260">L.S. Rahanitriniaina</collectorName>
&amp;
<collectorName id="26B4E4BA6655027F98E9FE2FC5EDFE3F" box="[492,678,475,502]" pageId="7" pageNumber="260">E.M. Rabotoson</collectorName>
lgt.” (
<collectionCode id="ED5019A96655027F9BF1FE28C47EFE3F" box="[756,821,475,501]" pageId="7" pageNumber="260">JJRC</collectionCode>
);
</materialsCitation>
<materialsCitation id="3B298B316655027F9A48FE28C336FDAB" collectingDate="2012-08" collectingDateMax="2012-08-22" collectingDateMin="2012-08" collectionCode="MMBC, JJRC" country="Madagascar" determinerName="Banar" elevation="965" latitude="-18.917555" longLatPrecision="1" longitude="48.49361" pageId="7" pageNumber="260" specimenCount="4" specimenCount-female="4" typeStatus="holotype">
<specimenCount id="9D474AE56655027F9A48FE28C437FE3C" box="[845,892,475,502]" count="1" pageId="7" pageNumber="260" type="female">1 ♀</specimenCount>
: “
<specimenCount id="9D474AE56655027F9A9EFE28C4E6FE3F" box="[923,941,475,501]" count="1" pageId="7" pageNumber="260" type="female">♀</specimenCount>
” // “VOH/
<date id="FFFFA7AC6655027F9D34FE28C3D4FE3F" box="[1073,1183,475,501]" pageId="7" pageNumber="260" value="2012-08">
<collectingDate id="EFBB5E446655027F9D34FE28C3D4FE3F" box="[1073,1183,475,501]" pageId="7" pageNumber="260" value="2012-08">Aug.2012</collectingDate>
</date>
/08 MADAGASCAR, Réserv. Expérimentale de VOHIMANA, „Circuit 2, PK 12“,
<date id="FFFFA7AC6655027F9A3AFE0CC482FDD3" box="[831,969,511,537]" pageId="7" pageNumber="260" value="2012-08-22">
<collectingDate id="EFBB5E446655027F9A3AFE0CC482FDD3" box="[831,969,511,537]" pageId="7" pageNumber="260" value="2012-08-22">22.viii.2012</collectingDate>
</date>
” // “
<geoCoordinate id="EE75E7AB6655027F9D06FE0CC3D6FDD0" box="[1027,1181,511,538]" degrees="18" direction="south" minutes="55" orientation="latitude" pageId="7" pageNumber="260" precision="1" seconds="46.4" value="-18.929554">S18°55’46.4”</geoCoordinate>
<geoCoordinate id="EE75E7AB6655027F9DADFDF3C20CFDD3" box="[1192,1351,511,538]" degrees="48" direction="east" minutes="30" orientation="longitude" pageId="7" pageNumber="260" precision="1" seconds="02.4" value="48.500668">E48°30’02.4”</geoCoordinate>
; 916m, sifting litter; Winkler app. extr.;
<collectorName id="26B4E4BA6655027F9B11FDD0C5B1FDF7" box="[532,762,547,573]" pageId="7" pageNumber="260">L.S. Rahanitriniaina</collectorName>
&amp;
<collectorName id="26B4E4BA6655027F9A20FDD7C4ABFDF7" box="[805,992,547,574]" pageId="7" pageNumber="260">E.M. Rabotoson</collectorName>
lgt.” (
<collectionCode id="ED5019A96655027F9D2BFDD0C3C7FDF7" box="[1070,1164,547,573]" country="Czech Republic" httpUri="http://grbio.org/cool/w50r-ge1v" name="Moravske Muzeum [Moravian Museum]" pageId="7" pageNumber="260">MMBC</collectionCode>
). All paratypes with an additional label “PARATYPUS
<taxonomicName id="4C41FAEF6655027F98F8FDB4C5BBFDAB" authority="Janák &amp; Baňař, 2024" authorityName="Janák &amp; Baňař" authorityYear="2024" box="[509,752,583,609]" class="Insecta" family="Staphylinidae" genus="Paulianidia" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="260" phylum="Arthropoda" rank="species" species="vohimana" status="sp. nov.">Paulianidia vohimana</taxonomicName>
<taxonomicNameLabel id="A206E0056655027F9BF2FDBBC41BFDA8" box="[759,848,584,610]" pageId="7" pageNumber="260" rank="species">sp. nov.</taxonomicNameLabel>
, Janák &amp;
<determinerName id="7E226DA86655027F9ABAFDB4C349FDAB" box="[959,1026,583,609]" pageId="7" pageNumber="260">Baňař</determinerName>
det. 2023”
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C35BD2E76655027F99C2FD98C5B8FA2F" pageId="7" pageNumber="260" type="description">
<paragraph id="8BFE816C6655027F99C2FD98C6E6FD07" blockId="7.[151,1437,259,1834]" pageId="7" pageNumber="260">
<emphasis id="B9355D7E6655027F99C2FD98C61AFD4F" bold="true" box="[199,337,619,645]" pageId="7" pageNumber="260">Description</emphasis>
: Body length 5.7–6.0 mm (M =
<quantity id="4CB92C896655027F9BCEFD98C46EFD4C" box="[715,805,619,646]" metricMagnitude="-3" metricUnit="m" metricValue="5.8" pageId="7" pageNumber="260" unit="mm" value="5.8">5.8 mm</quantity>
, HT = 6.0 mm), forebody length 3.0–
<quantity id="4CB92C896655027F9DDDFD98C266FD4C" box="[1240,1325,619,646]" metricMagnitude="-3" metricUnit="m" metricValue="3.1" pageId="7" pageNumber="260" unit="mm" value="3.1">3.1 mm</quantity>
(M =
<quantity id="4CB92C896655027F9C7CFD98C78EFD60" metricMagnitude="-3" metricUnit="m" metricValue="3.1" pageId="7" pageNumber="260" unit="mm" value="3.1">3.1 mm</quantity>
, HT = 3.0 mm). Dark brown, abdomen reddish brown, mouthparts and antennomeres dark reddish brown, legs reddish yellow (
<figureCitation id="137A9DE96655027F9849FD40C6EAFD07" box="[332,417,691,717]" captionStart="FIGURES 31–44" captionStartId="6.[151,264,1784,1808]" captionTargetBox="[185,1404,196,1741]" captionTargetId="figure-34@6.[171,1416,181,1760]" captionTargetPageId="6" captionText="FIGURES 31–44. 31–37: Paulianidia valohoaka sp. nov., holotype: 31, habitus; 32, forebody; 33–35, aedeagus (33, ventral; 34, 1/3 lateral; 35, lateral); 36, male sternite VII; 37, male sternite VIII. 38–44: P. vohimana sp. nov., holotype: 38, habitus; 39, forebody; 40–42, aedeagus (40, ventral; 41, 1/3 lateral; 42, lateral); 43, male sternite VII; 44, male sternite VIII. Scales 1 mm: 31–32, 38–39; 0.5 mm: 33–35, 36–37, 40–42, 43–44." figureDoi="http://doi.org/10.5281/zenodo.10847644" httpUri="https://zenodo.org/record/10847644/files/figure.png" pageId="7" pageNumber="260">Fig. 38</figureCitation>
).
</paragraph>
<paragraph id="8BFE816C6655027F99C2FD24C388FC6F" blockId="7.[151,1437,259,1834]" pageId="7" pageNumber="260">
Head (
<figureCitation id="137A9DE96655027F9817FD24C62FFD3B" box="[274,356,727,753]" captionStart="FIGURES 31–44" captionStartId="6.[151,264,1784,1808]" captionTargetBox="[185,1404,196,1741]" captionTargetId="figure-34@6.[171,1416,181,1760]" captionTargetPageId="6" captionText="FIGURES 31–44. 31–37: Paulianidia valohoaka sp. nov., holotype: 31, habitus; 32, forebody; 33–35, aedeagus (33, ventral; 34, 1/3 lateral; 35, lateral); 36, male sternite VII; 37, male sternite VIII. 38–44: P. vohimana sp. nov., holotype: 38, habitus; 39, forebody; 40–42, aedeagus (40, ventral; 41, 1/3 lateral; 42, lateral); 43, male sternite VII; 44, male sternite VIII. Scales 1 mm: 31–32, 38–39; 0.5 mm: 33–35, 36–37, 40–42, 43–44." figureDoi="http://doi.org/10.5281/zenodo.10847644" httpUri="https://zenodo.org/record/10847644/files/figure.png" pageId="7" pageNumber="260">Fig. 39</figureCitation>
) moderately convex, about 1.1 times as long as wide (L/W = 1.10–1.14, M = 1.12, HT = 1.11). Surface between eyes with coarse and dense umbilicate punctures, centre of disc with punctation markedly sparser and finer and here and there with interstices larger than diameter of punctures. Eyes small, temples on average about 1.7 times as long as eyes (R = 1.48–1.84, M = 1.69, HT = 1.48). Temples markedly roundly narrowed posteriorly. Antennae long and slender, fifth antennomere more than one and half as long as wide (L/W = 1.64–1.88, M = 1.77, HT = 1.88), tenth antennomere about as long as wide (L/W = 0.93–1.07, M = 0.98, HT = 1.07).
</paragraph>
<paragraph id="8BFE816C6655027F99C2FC43C332FB93" blockId="7.[151,1437,259,1834]" pageId="7" pageNumber="260">
Pronotum (
<figureCitation id="137A9DE96655027F9842FC5CC6D1FC03" box="[327,410,943,969]" captionStart="FIGURES 31–44" captionStartId="6.[151,264,1784,1808]" captionTargetBox="[185,1404,196,1741]" captionTargetId="figure-34@6.[171,1416,181,1760]" captionTargetPageId="6" captionText="FIGURES 31–44. 31–37: Paulianidia valohoaka sp. nov., holotype: 31, habitus; 32, forebody; 33–35, aedeagus (33, ventral; 34, 1/3 lateral; 35, lateral); 36, male sternite VII; 37, male sternite VIII. 38–44: P. vohimana sp. nov., holotype: 38, habitus; 39, forebody; 40–42, aedeagus (40, ventral; 41, 1/3 lateral; 42, lateral); 43, male sternite VII; 44, male sternite VIII. Scales 1 mm: 31–32, 38–39; 0.5 mm: 33–35, 36–37, 40–42, 43–44." figureDoi="http://doi.org/10.5281/zenodo.10847644" httpUri="https://zenodo.org/record/10847644/files/figure.png" pageId="7" pageNumber="260">Fig. 39</figureCitation>
) egg-shaped, moderately convex, with shallow basal impressions, about 1.1 times as long as wide (L/W = 1.15–1.17, M = 1.16, HT = 1.15) and slightly narrower than head (PW/HW = 0.96). Sides slightly roundly widened posteriorly. Surface with moderately coarse and sparse punctures and with distinct interstices between punctures, punctation coarse and less dense than that on head and less dense along moderately narrow unpunctured midline; midline in anterior part of pronotum more than one puncture wide.
</paragraph>
<paragraph id="8BFE816C6655027F99C2FB90C54FFB23" blockId="7.[151,1437,259,1834]" pageId="7" pageNumber="260">
Elytra (
<figureCitation id="137A9DE96655027F981CFB90C623FBB7" box="[281,360,1123,1149]" captionStart="FIGURES 31–44" captionStartId="6.[151,264,1784,1808]" captionTargetBox="[185,1404,196,1741]" captionTargetId="figure-34@6.[171,1416,181,1760]" captionTargetPageId="6" captionText="FIGURES 31–44. 31–37: Paulianidia valohoaka sp. nov., holotype: 31, habitus; 32, forebody; 33–35, aedeagus (33, ventral; 34, 1/3 lateral; 35, lateral); 36, male sternite VII; 37, male sternite VIII. 38–44: P. vohimana sp. nov., holotype: 38, habitus; 39, forebody; 40–42, aedeagus (40, ventral; 41, 1/3 lateral; 42, lateral); 43, male sternite VII; 44, male sternite VIII. Scales 1 mm: 31–32, 38–39; 0.5 mm: 33–35, 36–37, 40–42, 43–44." figureDoi="http://doi.org/10.5281/zenodo.10847644" httpUri="https://zenodo.org/record/10847644/files/figure.png" pageId="7" pageNumber="260">Fig. 39</figureCitation>
) elongate with rounded sides and distinct depression along suture in first quarter behind scutellum, slightly flattened along suture, about 1.1 times longer than wide (R = 1.13–1.14, M = 1.14, HT = 1.13). Surface moderately coarsely and densely punctured. Interstices mostly slightly larger than diameter of punctures, surface shining, without microsculpture.
</paragraph>
<paragraph id="8BFE816C6655027F99C2FB00C4E9FAFB" blockId="7.[151,1437,259,1834]" pageId="7" pageNumber="260">
Abdomen (
<figureCitation id="137A9DE96655027F9843FB00C6D3FAC7" box="[326,408,1267,1293]" captionStart="FIGURES 31–44" captionStartId="6.[151,264,1784,1808]" captionTargetBox="[185,1404,196,1741]" captionTargetId="figure-34@6.[171,1416,181,1760]" captionTargetPageId="6" captionText="FIGURES 31–44. 31–37: Paulianidia valohoaka sp. nov., holotype: 31, habitus; 32, forebody; 33–35, aedeagus (33, ventral; 34, 1/3 lateral; 35, lateral); 36, male sternite VII; 37, male sternite VIII. 38–44: P. vohimana sp. nov., holotype: 38, habitus; 39, forebody; 40–42, aedeagus (40, ventral; 41, 1/3 lateral; 42, lateral); 43, male sternite VII; 44, male sternite VIII. Scales 1 mm: 31–32, 38–39; 0.5 mm: 33–35, 36–37, 40–42, 43–44." figureDoi="http://doi.org/10.5281/zenodo.10847644" httpUri="https://zenodo.org/record/10847644/files/figure.png" pageId="7" pageNumber="260">Fig. 38</figureCitation>
) slightly widened to segment V, finely and sparsely punctured. Interstices shining, without microsculpture. Tergite VII with distinct membranous fringe apically.
</paragraph>
<paragraph id="8BFE816C6655027F99C2FAC8C585FA08" blockId="7.[151,1437,259,1834]" pageId="7" pageNumber="260">
Male. Posterior margin of sternite VII with short and wide triangular emargination (
<figureCitation id="137A9DE96655027F9D77FAC8C38DFA9F" box="[1138,1222,1339,1365]" captionStart="FIGURES 31–44" captionStartId="6.[151,264,1784,1808]" captionTargetBox="[185,1404,196,1741]" captionTargetId="figure-34@6.[171,1416,181,1760]" captionTargetPageId="6" captionText="FIGURES 31–44. 31–37: Paulianidia valohoaka sp. nov., holotype: 31, habitus; 32, forebody; 33–35, aedeagus (33, ventral; 34, 1/3 lateral; 35, lateral); 36, male sternite VII; 37, male sternite VIII. 38–44: P. vohimana sp. nov., holotype: 38, habitus; 39, forebody; 40–42, aedeagus (40, ventral; 41, 1/3 lateral; 42, lateral); 43, male sternite VII; 44, male sternite VIII. Scales 1 mm: 31–32, 38–39; 0.5 mm: 33–35, 36–37, 40–42, 43–44." figureDoi="http://doi.org/10.5281/zenodo.10847644" httpUri="https://zenodo.org/record/10847644/files/figure.png" pageId="7" pageNumber="260">Fig. 43</figureCitation>
), sternite VIII with short and moderately wide triangular emargination in posterior tenth (
<figureCitation id="137A9DE96655027F9AA0FAACC4B2FAB3" box="[933,1017,1375,1401]" captionStart="FIGURES 31–44" captionStartId="6.[151,264,1784,1808]" captionTargetBox="[185,1404,196,1741]" captionTargetId="figure-34@6.[171,1416,181,1760]" captionTargetPageId="6" captionText="FIGURES 31–44. 31–37: Paulianidia valohoaka sp. nov., holotype: 31, habitus; 32, forebody; 33–35, aedeagus (33, ventral; 34, 1/3 lateral; 35, lateral); 36, male sternite VII; 37, male sternite VIII. 38–44: P. vohimana sp. nov., holotype: 38, habitus; 39, forebody; 40–42, aedeagus (40, ventral; 41, 1/3 lateral; 42, lateral); 43, male sternite VII; 44, male sternite VIII. Scales 1 mm: 31–32, 38–39; 0.5 mm: 33–35, 36–37, 40–42, 43–44." figureDoi="http://doi.org/10.5281/zenodo.10847644" httpUri="https://zenodo.org/record/10847644/files/figure.png" pageId="7" pageNumber="260">Fig. 44</figureCitation>
). Aedeagus (
<figureCitation id="137A9DE96655027F9D8CFAACC244FAB3" box="[1161,1295,1375,1401]" captionStart="FIGURES 31–44" captionStartId="6.[151,264,1784,1808]" captionTargetBox="[185,1404,196,1741]" captionTargetId="figure-34@6.[171,1416,181,1760]" captionTargetPageId="6" captionText="FIGURES 31–44. 31–37: Paulianidia valohoaka sp. nov., holotype: 31, habitus; 32, forebody; 33–35, aedeagus (33, ventral; 34, 1/3 lateral; 35, lateral); 36, male sternite VII; 37, male sternite VIII. 38–44: P. vohimana sp. nov., holotype: 38, habitus; 39, forebody; 40–42, aedeagus (40, ventral; 41, 1/3 lateral; 42, lateral); 43, male sternite VII; 44, male sternite VIII. Scales 1 mm: 31–32, 38–39; 0.5 mm: 33–35, 36–37, 40–42, 43–44." figureDoi="http://doi.org/10.5281/zenodo.10847644" httpUri="https://zenodo.org/record/10847644/files/figure.png" pageId="7" pageNumber="260">Figs. 40–42</figureCitation>
) moderately long and slender, about
<quantity id="4CB92C896655027F98A3FA70C6B6FA54" box="[422,509,1411,1438]" metricMagnitude="-4" metricUnit="m" metricValue="7.0" pageId="7" pageNumber="260" unit="mm" value="0.7">0.7 mm</quantity>
long (AL =
<quantity id="4CB92C896655027F9B89FA70C5BEFA57" box="[652,757,1411,1437]" metricMagnitude="-4" metricUnit="m" metricValue="6.8999999999999995" pageId="7" pageNumber="260" unit="mm" value="0.69">0.69 mm</quantity>
). Median lobe in ventral view with moderately wide middle part, narrowed before slightly widened apical part.
</paragraph>
<paragraph id="8BFE816C6655027F99C2FA38C5B8FA2F" blockId="7.[151,1437,259,1834]" box="[199,755,1483,1510]" pageId="7" pageNumber="260">Female. Sternite VIII largely rounded posteriorly.</paragraph>
</subSubSection>
<subSubSection id="C35BD2E76655027F99C2FA1CC439F950" pageId="7" pageNumber="260" type="diagnosis">
<paragraph id="8BFE816C6655027F99C2FA1CC439F950" blockId="7.[151,1437,259,1834]" pageId="7" pageNumber="260">
<emphasis id="B9355D7E6655027F99C2FA1CC686F9C3" bold="true" box="[199,461,1519,1545]" pageId="7" pageNumber="260">Differential diagnosis.</emphasis>
The new species is arranged in a group of species with normally (depressed) punctured elytra, impunctate or sparsely punctate disc of the head and integument of tergites smooth between punctures together with
<taxonomicName id="4C41FAEF6655027F9836F9CBC550F99B" authority="Lecoq, 2008" authorityName="Lecoq" authorityYear="2008" box="[307,539,1591,1618]" class="Insecta" family="Staphylinidae" genus="Paulianidia" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="260" phylum="Arthropoda" rank="species" species="hova">
<emphasis id="B9355D7E6655027F9836F9CBC6CEF99B" box="[307,389,1591,1617]" italics="true" pageId="7" pageNumber="260">P. hova</emphasis>
<bibRefCitation id="EFD0FC9D6655027F9889F9CBC550F99B" author="Lecoq, J. C." box="[396,539,1591,1618]" pageId="7" pageNumber="260" pagination="579 - 603" refId="ref12275" refString="Lecoq, J. C. (2008) Revision du genre Paulianidia Jarrige, 1978 de Madagascar (Insecta, Coleoptera, Staphylinidae, Paederinae). Zoosystema, 30 (3), 579 - 603." type="journal article" year="2008">Lecoq, 2008</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C41FAEF6655027F9B23F9CBC467F99B" authority="Lecoq, 2008" authorityName="Lecoq" authorityYear="2008" box="[550,812,1591,1618]" class="Insecta" family="Staphylinidae" genus="Paulianidia" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="260" phylum="Arthropoda" rank="species" species="minima">
<emphasis id="B9355D7E6655027F9B23F9CBC5DDF99B" box="[550,662,1592,1617]" italics="true" pageId="7" pageNumber="260">P. minima</emphasis>
<bibRefCitation id="EFD0FC9D6655027F9B98F9CBC467F99B" author="Lecoq, J. C." box="[669,812,1591,1618]" pageId="7" pageNumber="260" pagination="579 - 603" refId="ref12275" refString="Lecoq, J. C. (2008) Revision du genre Paulianidia Jarrige, 1978 de Madagascar (Insecta, Coleoptera, Staphylinidae, Paederinae). Zoosystema, 30 (3), 579 - 603." type="journal article" year="2008">Lecoq, 2008</bibRefCitation>
</taxonomicName>
, and
<taxonomicName id="4C41FAEF6655027F9A62F9CBC330F99B" authority="Janak &amp; Banar" authorityName="Janak &amp; Banar" authorityYear="2024" box="[871,1147,1591,1617]" class="Insecta" family="Staphylinidae" genus="Paulianidia" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="260" phylum="Arthropoda" rank="species" species="tanala" status="sp. nov.">
<emphasis id="B9355D7E6655027F9A62F9CBC481F99B" box="[871,970,1591,1617]" italics="true" pageId="7" pageNumber="260">P. tanala</emphasis>
Janák &amp; Baňař
</taxonomicName>
,
<emphasis id="B9355D7E6655027F9D83F9C4C3ABF99B" bold="true" box="[1158,1248,1591,1617]" pageId="7" pageNumber="260">
<taxonomicNameLabel id="A206E0056655027F9D83F9C4C3ABF99B" box="[1158,1248,1591,1617]" pageId="7" pageNumber="260" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
and differs from all these species by combination of fine and sparse punctation of tergites and head markedly narrowed behind eyes and in males also by very distinct shape of sternites VII and VIII.
</paragraph>
</subSubSection>
<subSubSection id="C35BD2E76655027F99C2F950C445F977" box="[199,782,1699,1726]" pageId="7" pageNumber="260" type="etymology">
<paragraph id="8BFE816C6655027F99C2F950C445F977" blockId="7.[151,1437,259,1834]" box="[199,782,1699,1726]" pageId="7" pageNumber="260">
<emphasis id="B9355D7E6655027F99C2F950C605F977" bold="true" box="[199,334,1699,1725]" pageId="7" pageNumber="260">Etymology.</emphasis>
The name is related to the
<typeStatus id="54FA3FCE6655027F9B78F957C5E6F974" box="[637,685,1700,1726]" pageId="7" pageNumber="260">type</typeStatus>
locality.
</paragraph>
</subSubSection>
<subSubSection id="C35BD2E76655027F99C2F934C472F8E3" pageId="7" pageNumber="260" type="distribution">
<paragraph id="8BFE816C6655027F99C2F934C414F8CF" blockId="7.[151,1437,259,1834]" pageId="7" pageNumber="260">
<emphasis id="B9355D7E6655027F99C2F934C62AF92B" bold="true" box="[199,353,1735,1761]" pageId="7" pageNumber="260">Distribution.</emphasis>
<taxonomicName id="4C41FAEF6655027F9862F934C45BF92B" authority="Janak &amp; Banar" authorityName="Janak &amp; Banar" authorityYear="2024" box="[359,784,1735,1761]" class="Insecta" family="Staphylinidae" genus="Paulianidia" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="260" phylum="Arthropoda" rank="species" species="vohimana" status="sp. nov.">
<emphasis id="B9355D7E6655027F9862F934C52BF92B" box="[359,608,1735,1761]" italics="true" pageId="7" pageNumber="260">Paulianidia vohimana</emphasis>
Janák &amp; Baňař
</taxonomicName>
,
<emphasis id="B9355D7E6655027F9A1FF934C438F92B" bold="true" box="[794,883,1735,1761]" pageId="7" pageNumber="260">
<taxonomicNameLabel id="A206E0056655027F9A1FF934C438F92B" box="[794,883,1735,1761]" pageId="7" pageNumber="260" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is currently recorded only from the Experimental Reserve Vohimana in the central part of
<collectingCountry id="F356C1FC6655027F9B5FF918C5AAF8CF" box="[602,737,1771,1797]" name="Madagascar" pageId="7" pageNumber="260">Madagascar</collectingCountry>
(
<figureCitation id="137A9DE96655027F9BF4F918C419F8CF" box="[753,850,1771,1797]" captionStart="FIGURES 116–122" captionStartId="12.[151,264,1780,1804]" captionTargetBox="[180,1408,194,1743]" captionTargetId="figure-34@12.[167,1421,181,1756]" captionText="FIGURES 116–122. 116–120: Paulianidia steineri Lecoq, Andasibe: 116–118, aedeagus (116, ventral; 117, 1/3 lateral; 118, lateral); 119, male sternite VII; 120, male sternite VIII. Scale 0.5 mm: 116–120. 121: Distribution of P. densepunctata sp. nov., (rhombus), P. valohoaka sp. nov. (star), P. tanala sp. nov. and P. vohimana sp. nov. (rectangle). 122: Distribution of Paulianidia species in Madagascar." figureDoi="http://doi.org/10.5281/zenodo.10854991" httpUri="https://zenodo.org/record/10854991/files/figure.png" pageId="7" pageNumber="260">Fig. 121</figureCitation>
).
</paragraph>
<paragraph id="8BFE816C6655027F99C2F8FCC472F8E3" blockId="7.[151,1437,259,1834]" box="[199,825,1807,1834]" pageId="7" pageNumber="260">
<emphasis id="B9355D7E6655027F99C2F8FCC600F8E3" bold="true" box="[199,331,1807,1833]" pageId="7" pageNumber="260">Bionomics.</emphasis>
The
<typeStatus id="54FA3FCE6655027F9881F8E3C6FFF8E0" box="[388,436,1808,1834]" pageId="7" pageNumber="260">type</typeStatus>
series was sifted from forest litter.
</paragraph>
</subSubSection>
</treatment>
</document>
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