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<document id="ADF5BCB21AFAA97F5026AAFBA92A6290" ID-CLB-Dataset="26414" ID-DOI="10.11646/zootaxa.4623.2.3" ID-GBIF-Dataset="c5c601d2-c2d1-4841-a206-5b6778989ad6" ID-ISSN="1175-5326" ID-Zenodo-Dep="3255421" ID-ZooBank="0ACA24AD-9249-4387-B1C2-8DF85FBCEDEA" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1561449449659" checkinUser="plazi" docAuthor="Bartholomay, Pedro R., Williams, Kevin A., Cambra, To. A. &amp; Oliveira, Márcio L." docDate="2019" docId="03E787F26750FFD560DEFBC4526FFDCC" docLanguage="en" docName="zootaxa.4623.2.3.pdf" docOrigin="Zootaxa 4623 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Quwitilla peruviana Bartholomay &amp; Williams &amp; Cambra &amp; Oliveira 2019" docType="treatment" docVersion="9" lastPageNumber="269" masterDocId="FFDEFF8A6758FFD96049FFA1565CFFD7" masterDocTitle="Does the genus Dasymutilla Ashmead occur in South America? The new genus Quwitilla, new combinations, and new distribution records from Neotropical velvet ants (Hymenoptera: Mutillidae)" masterLastPageNumber="282" masterPageNumber="261" pageNumber="269" updateTime="1698736201454" updateUser="ExternalLinkService">
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<mods:title id="4B9E3AEFCDB6ED3BAAB15D29DB6EBA18">Does the genus Dasymutilla Ashmead occur in South America? The new genus Quwitilla, new combinations, and new distribution records from Neotropical velvet ants (Hymenoptera: Mutillidae)</mods:title>
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<mods:name id="A0EC16A87D40D93E1D18E98F287548D8" type="personal">
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<mods:namePart id="D2F0007322A8AB39FD661FC5771631D6">Bartholomay, Pedro R.</mods:namePart>
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<mods:namePart id="92BCEC4D5A5E4BA034821DF63FF123D4">Williams, Kevin A.</mods:namePart>
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<mods:namePart id="06164E71ACFCD171D57EA22A9DC349F9">Cambra, To. A.</mods:namePart>
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<mods:namePart id="ACCE81AA1DE1B88F7DA3D2A62D487DD5">Oliveira, Márcio L.</mods:namePart>
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<mods:title id="419BB78056E1697010CA198AD5117570">Zootaxa</mods:title>
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<mods:date id="943D5F794FFC6791229AF3E2FF432860">2019</mods:date>
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<mods:number id="4D455D1BA87A6DCF152EB5D14F5E5123">2019-06-25</mods:number>
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<mods:number id="F085DBA362B2FA813C3E999CBDADDF8E">4623</mods:number>
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<treatment id="03E787F26750FFD560DEFBC4526FFDCC" ID-DOI="http://doi.org/10.5281/zenodo.5930948" ID-GBIF-Taxon="158380791" ID-Zenodo-Dep="5930948" LSID="urn:lsid:plazi:treatment:03E787F26750FFD560DEFBC4526FFDCC" httpUri="http://treatment.plazi.org/id/03E787F26750FFD560DEFBC4526FFDCC" lastPageId="12" lastPageNumber="269" pageId="8" pageNumber="269">
<subSubSection id="C354656F6750FFD160DEFBC4541FFBA8" box="[151,579,1124,1152]" pageId="8" pageNumber="269" type="nomenclature">
<paragraph id="8BF136E46750FFD160DEFBC4541FFBA8" blockId="8.[151,579,1124,1187]" box="[151,579,1124,1152]" pageId="8" pageNumber="269">
<heading id="D0B981886750FFD160DEFBC4541FFBA8" bold="true" box="[151,579,1124,1152]" fontSize="11" level="1" pageId="8" pageNumber="269" reason="1">
<taxonomicName id="4C4E4D676750FFD160DEFBC4541FFBA8" authority="(Suarez, 1970)" authorityName="Bartholomay &amp; Williams &amp; Cambra &amp; Oliveira" authorityYear="2019" baseAuthorityName="Suarez" baseAuthorityYear="1970" box="[151,579,1124,1152]" class="Insecta" family="Mutillidae" genus="Quwitilla" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="269" phylum="Arthropoda" rank="species" species="peruviana">
<emphasis id="B93AEAF66750FFD160DEFBC4541FFBA8" bold="true" box="[151,579,1124,1152]" pageId="8" pageNumber="269">
<emphasis id="B93AEAF66750FFD160DEFBC457D5FBA8" bold="true" box="[151,393,1125,1152]" italics="true" pageId="8" pageNumber="269">Quwitilla peruviana</emphasis>
(
<bibRefCitation id="EFDF4B156750FFD161D0FBC55467FBA8" author="Suarez, F. J." box="[409,571,1124,1151]" pageId="8" pageNumber="269" pagination="169 - 188" refId="ref10283" refString="Suarez, F. J. (1970) Datos sobre Mutilidos neotropicales VII. Algunas especies del Peru representadas en las colecciones del Museo de Historia Natural de Basilea (Suiza). Archivos del Instituto de Aclimataccion de Almeria, 15, 169 - 188." type="journal article" year="1970">Suárez, 1970</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C354656F6750FFD160DEFB28577BFB74" box="[151,295,1161,1187]" pageId="8" pageNumber="269" type="description">
<paragraph id="8BF136E46750FFD160DEFB28577BFB74" blockId="8.[151,579,1124,1187]" box="[151,295,1161,1187]" pageId="8" pageNumber="269">
(
<figureCitation id="13752A616750FFD160D6FB285743FB74" box="[159,287,1161,1187]" captionStart-0="FIGURES 2023" captionStart-1="FIGURES 2631" captionStartId-0="9.[151,264,1955,1980]" captionStartId-1="11.[151,264,1939,1964]" captionTargetBox-0="[151,1380,189,1928]" captionTargetBox-1="[151,1290,181,1920]" captionTargetId-0="figure@9.[151,1380,189,1928]" captionTargetId-1="figure@11.[151,1290,181,1920]" captionTargetPageId-0="9" captionTargetPageId-1="11" captionText-0="FIGURES 2023. Quwitilla peruviana (Suárez, 1970), ♀, holotype (of Dasymutilla homochroma Suárez), lines 2mm; 20. Dorsal view; 21. Lateral view; 22. Pygidium, posterodorsal view; 23. Type labels." captionText-1="FIGURES 2631. Quwitilla peruviana (Suárez, 1970), ♂, holotype; 26. Head, frontal view; 27. Pygidium (T7), posterodorsal view; 28. Mesosoma, dorsal view; 29. Hypopygium (S7), ventral view; 30. Type labels; 31. Genitalia (encased in resin)." httpUri-0="https://zenodo.org/record/3255427/files/figure.png" httpUri-1="https://zenodo.org/record/3255429/files/figure.png" pageId="8" pageNumber="269">Figs 2031</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C354656F6750FFD160DEFB6C546DFA96" pageId="8" pageNumber="269" type="reference_group">
<paragraph id="8BF136E46750FFD160DEFB6C5793FAD3" blockId="8.[151,1436,1229,1346]" pageId="8" pageNumber="269">
<taxonomicName id="4C4E4D676750FFD160DEFB6C5450FB32" authority="Suarez, 1970" authorityName="Suarez" authorityYear="1970" box="[151,524,1229,1253]" class="Insecta" family="Mutillidae" genus="Dasymutilla" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="269" phylum="Arthropoda" rank="species" species="peruviana">
<emphasis id="B93AEAF66750FFD160DEFB6C57DCFB32" box="[151,384,1229,1253]" italics="true" pageId="8" pageNumber="269">Dasymutilla peruviana</emphasis>
<materialsCitation id="3B263CB96750FFD161CCFB6C5450FB32" ID-GBIF-Occurrence="2269395014" box="[389,524,1229,1253]" collectionCode="NHMB" country="Peru" location="Archivos del Instituto de Aclimatacion Almeria" municipality="San Miguel" pageId="8" pageNumber="269" specimenCount="1" specimenCount-male="1" stateProvince="Lima" typeStatus="holotype">
<bibRefCitation id="EFDF4B156750FFD161CCFB6C5450FB32" author="Suarez, F. J." box="[389,524,1229,1253]" pageId="8" pageNumber="269" pagination="169 - 188" refId="ref10283" refString="Suarez, F. J. (1970) Datos sobre Mutilidos neotropicales VII. Algunas especies del Peru representadas en las colecciones del Museo de Historia Natural de Basilea (Suiza). Archivos del Instituto de Aclimataccion de Almeria, 15, 169 - 188." type="journal article" year="1970">Suárez, 1970</bibRefCitation>
</materialsCitation>
</taxonomicName>
.
<location id="8E91603F6750FFD1625AFB6C55B3FB31" LSID="urn:lsid:plazi:treatment:03E787F26750FFD560DEFBC4526FFDCC:8E91603F6750FFD1625AFB6C55B3FB31" box="[531,1007,1229,1254]" country="Peru" municipality="San Miguel" name="Archivos del Instituto de Aclimatacion Almeria" pageId="8" pageNumber="269" stateProvince="Lima">Archivos del Instituto de Aclimatación Almeria</location>
15: 177.
<typeStatus id="54F588466750FFD16405FB6C52F6FB31" box="[1100,1194,1229,1254]" pageId="8" pageNumber="269" type="holotype">Holotype</typeStatus>
<specimenCount id="9D48FD6D6750FFD164F9FB6F5299FB32" box="[1200,1221,1230,1253]" pageId="8" pageNumber="269" type="male"></specimenCount>
,
<collectingMunicipality id="6B95AC9E6750FFD16486FB6C531BFB31" box="[1231,1351,1229,1254]" pageId="8" pageNumber="269">San Miguel</collectingMunicipality>
, [
<collectingRegion id="498AF8066750FFD1651FFB6C53CEFB31" box="[1366,1426,1229,1254]" country="Peru" name="Lima" pageId="8" pageNumber="269">Lima</collectingRegion>
],
<collectingCountry id="F35976746750FFD1608EFB4D56A9FAD4" box="[199,245,1260,1283]" name="Peru" pageId="8" pageNumber="269">Peru</collectingCountry>
(NHMB), examined.
</paragraph>
<paragraph id="8BF136E46750FFD160DEFAAA546DFA96" blockId="8.[151,1436,1229,1346]" pageId="8" pageNumber="269">
<taxonomicName id="4C4E4D676750FFD160DEFAAA5479FAF4" authority="Suarez, 1970" authorityName="Suarez" authorityYear="1970" box="[151,549,1291,1315]" class="Insecta" family="Mutillidae" genus="Dasymutilla" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="269" phylum="Arthropoda" rank="species" species="homochroma">
<emphasis id="B93AEAF66750FFD160DEFAAA57C0FAF4" box="[151,412,1291,1315]" italics="true" pageId="8" pageNumber="269">Dasymutilla homochroma</emphasis>
<materialsCitation id="3B263CB96750FFD161E9FAAA5479FAF4" ID-GBIF-Occurrence="2269395018" box="[416,549,1291,1315]" collectionCode="NHMB" country="Peru" location="Archivos del Instituto de Aclimatacion Almeria" municipality="San Miguel" pageId="8" pageNumber="269" specimenCount="1" specimenCount-female="1" stateProvince="Lima" typeStatus="holotype">
<bibRefCitation id="EFDF4B156750FFD161E9FAAA5479FAF4" author="Suarez, F. J." box="[416,549,1291,1315]" pageId="8" pageNumber="269" pagination="169 - 188" refId="ref10283" refString="Suarez, F. J. (1970) Datos sobre Mutilidos neotropicales VII. Algunas especies del Peru representadas en las colecciones del Museo de Historia Natural de Basilea (Suiza). Archivos del Instituto de Aclimataccion de Almeria, 15, 169 - 188." type="journal article" year="1970">Suárez, 1970</bibRefCitation>
</materialsCitation>
</taxonomicName>
.
<location id="8E91603F6750FFD16263FAAB55A2FAF4" LSID="urn:lsid:plazi:treatment:03E787F26750FFD560DEFBC4526FFDCC:8E91603F6750FFD16263FAAB55A2FAF4" box="[554,1022,1290,1315]" country="Peru" municipality="San Miguel" name="Archivos del Instituto de Aclimatacion Almeria" pageId="8" pageNumber="269" stateProvince="Lima">Archivos del Instituto de Aclimatación Almeria</location>
15: 173.
<typeStatus id="54F588466750FFD1641EFAAB52E9FAF4" box="[1111,1205,1290,1315]" pageId="8" pageNumber="269" type="holotype">Holotype</typeStatus>
<specimenCount id="9D48FD6D6750FFD164F0FAAA5290FAF5" box="[1209,1228,1291,1314]" pageId="8" pageNumber="269" type="female"></specimenCount>
,
<collectingMunicipality id="6B95AC9E6750FFD1649FFAAA5312FAF4" box="[1238,1358,1290,1315]" pageId="8" pageNumber="269">San Miguel</collectingMunicipality>
,[
<collectingRegion id="498AF8066750FFD16511FAAB53CEFAF4" box="[1368,1426,1290,1315]" country="Peru" name="Lima" pageId="8" pageNumber="269">Lima</collectingRegion>
],
<collectingCountry id="F35976746750FFD1608EFA8B56A9FA96" box="[199,245,1322,1345]" name="Peru" pageId="8" pageNumber="269">Peru</collectingCountry>
(NHMB, examined).
<emphasis id="B93AEAF66750FFD1619CFA88546DFA96" bold="true" box="[469,561,1321,1346]" pageId="8" pageNumber="269">Syn. nov</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C354656F6750FFD160DEFAD15204F919" pageId="8" pageNumber="269" type="diagnosis">
<paragraph id="8BF136E46750FFD160DEFAD15204F919" blockId="8.[151,1437,1392,2030]" pageId="8" pageNumber="269">
<emphasis id="B93AEAF66750FFD160DEFAD15724FA5D" box="[151,376,1392,1418]" italics="true" pageId="8" pageNumber="269">Extended diagnosis.</emphasis>
Female. In addition to the characters described by
<bibRefCitation id="EFDF4B156750FFD163E5FAD15214FA5D" author="Suarez, F. J." box="[940,1096,1392,1418]" pageId="8" pageNumber="269" pagination="169 - 188" refId="ref10283" refString="Suarez, F. J. (1970) Datos sobre Mutilidos neotropicales VII. Algunas especies del Peru representadas en las colecciones del Museo de Historia Natural de Basilea (Suiza). Archivos del Instituto de Aclimataccion de Almeria, 15, 169 - 188." type="journal article" year="1970">Suárez (1970)</bibRefCitation>
the females can be recognized by the antennal scrobe with well-defined dorsal carina connected laterally with well-defined lateral carina; length of thorax (pronotum+mesonotum) 0.9 × its width; ratios of width of humeral angles, pronotal spiracles, widest point of mesonotum, narrowest point of mesonotum and propodeum posterior to propodeal spiracles, 58:65:65:53:48; scutellar scale small, narrowly connected to anterolateral carina; anterolateral carina more than 3 × the width of scutellar scale; ratios of width of T1, width of T2 and length of T2, 30:73:68; T2 widest point posterior to midlength; S2 with anteromedial flattened area; anteromedial longitudinal crest-fold conspicuous; subapical transverse slope present. Male. In addition to the characters described by
<bibRefCitation id="EFDF4B156750FFD1635CF9CD55EFF951" author="Suarez, F. J." box="[789,947,1644,1670]" pageId="8" pageNumber="269" pagination="169 - 188" refId="ref10283" refString="Suarez, F. J. (1970) Datos sobre Mutilidos neotropicales VII. Algunas especies del Peru representadas en las colecciones del Museo de Historia Natural de Basilea (Suiza). Archivos del Instituto de Aclimataccion de Almeria, 15, 169 - 188." type="journal article" year="1970">Suárez (1970)</bibRefCitation>
the ocelli are small, OOD 5.1 × DLO, IOD virtually equal to DLO; gena ecarinate; epaulet present, poorly developed, low, rounded, virtually flat against anterior margin of pronotum; meso and metatibial spurs black, finely serrated on margins.
</paragraph>
</subSubSection>
<subSubSection id="C354656F6750FFD1608EF97957C5F925" box="[199,409,1752,1778]" pageId="8" pageNumber="269" type="distribution">
<paragraph id="8BF136E46750FFD1608EF97957C5F925" blockId="8.[151,1437,1392,2030]" box="[199,409,1752,1778]" pageId="8" pageNumber="269">
<emphasis id="B93AEAF66750FFD1608EF9795704F925" box="[199,344,1752,1778]" italics="true" pageId="8" pageNumber="269">Distribution.</emphasis>
<collectingCountry id="F35976746750FFD16117F97957C9F925" box="[350,405,1752,1778]" name="Peru" pageId="8" pageNumber="269">Peru</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C354656F6750FFD1608EF95D57EBF855" pageId="8" pageNumber="269" type="materials_examined">
<paragraph id="8BF136E46750FFD1608EF95D57EBF855" blockId="8.[151,1437,1392,2030]" pageId="8" pageNumber="269">
<emphasis id="B93AEAF66750FFD1608EF95D57F4F8C1" box="[199,424,1788,1814]" italics="true" pageId="8" pageNumber="269">Material examined.</emphasis>
[
<specimenCount id="9D48FD6D6750FFD161F5F95D57BCF8C1" box="[444,480,1788,1814]" pageId="8" pageNumber="269" type="female">2♀</specimenCount>
<specimenCount id="9D48FD6D6750FFD161A2F95A5453F8C1" box="[491,527,1787,1814]" pageId="8" pageNumber="269" type="male">1♂</specimenCount>
] Type material.
<materialsCitation id="3B263CB96750FFD1629CF95D543FF8ED" ID-GBIF-Occurrence="2269395011" collectingDate="1955-03-08" collectionCode="NHMB" collectorName="M. Markl" country="Peru" location="San Miguel" pageId="8" pageNumber="269" specimenCount="1" specimenCount-male="1" stateProvince="Lima" typeStatus="holotype">
<typeStatus id="54F588466750FFD1629CF95D5561F8C1" box="[725,829,1788,1814]" pageId="8" pageNumber="269" type="holotype">Holotype</typeStatus>
of
<taxonomicName id="4C4E4D676750FFD16325F95D522EF8C2" authorityName="Suarez" authorityYear="1970" box="[876,1138,1788,1814]" class="Insecta" family="Mutillidae" genus="Dasymutilla" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="269" phylum="Arthropoda" rank="species" species="peruviana">
<emphasis id="B93AEAF66750FFD16325F95D522EF8C2" box="[876,1138,1788,1814]" italics="true" pageId="8" pageNumber="269">Dasymutilla peruviana</emphasis>
</taxonomicName>
,
<specimenCount id="9D48FD6D6750FFD164CCF95D52C0F8C1" box="[1157,1180,1788,1814]" pageId="8" pageNumber="269" type="male"></specimenCount>
,
<collectingCountry id="F35976746750FFD164E5F95D52AAF8C1" box="[1196,1270,1788,1814]" name="Peru" pageId="8" pageNumber="269">PERU</collectingCountry>
, [
<collectingRegion id="498AF8066750FFD16559F95D5316F8C2" box="[1296,1354,1788,1813]" country="Peru" name="Lima" pageId="8" pageNumber="269">
<emphasis id="B93AEAF66750FFD16559F95D5316F8C2" box="[1296,1354,1788,1813]" italics="true" pageId="8" pageNumber="269">Lima</emphasis>
</collectingRegion>
?],
<location id="8E91603F6750FFD1653BF95D56B7F8ED" LSID="urn:lsid:plazi:treatment:03E787F26750FFD560DEFBC4526FFDCC:8E91603F6750FFD1653BF95D56B7F8ED" country="Peru" name="San Miguel" pageId="8" pageNumber="269" stateProvince="Lima">San Miguel</location>
,
<collectingDate id="EFB4E9CC6750FFD160BEF8BE5724F8ED" box="[247,376,1823,1850]" pageId="8" pageNumber="269" value="1955-03-08">
<date id="FFF010246750FFD160BEF8BE5724F8ED" box="[247,376,1823,1850]" pageId="8" pageNumber="269" value="1955-03-08">08.III.1955</date>
</collectingDate>
,
<collectorName id="26BB53326750FFD161CDF88157ACF8ED" box="[388,496,1824,1850]" pageId="8" pageNumber="269">M. Markl</collectorName>
(
<collectionCode id="ED5FAE216750FFD16249F8815406F8ED" box="[512,602,1824,1850]" country="0" httpUri="http://grbio.org/cool/1d5a-4efs" name="Natural History Museum Bucharest" pageId="8" pageNumber="269">NHMB</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B263CB96750FFD16227F88157A1F889" ID-GBIF-Occurrence="2269395010" collectingDate="1955-03-08" collectionCode="NHMB" collectorName="M. Markl" country="Peru" location="San Miguel" pageId="8" pageNumber="269" specimenCount="1" specimenCount-female="1" stateProvince="Lima" typeStatus="holotype">
<typeStatus id="54F588466750FFD16227F881548AF8ED" box="[622,726,1824,1850]" pageId="8" pageNumber="269" type="holotype">Holotype</typeStatus>
of
<taxonomicName id="4C4E4D676750FFD162B7F881527DF8ED" box="[766,1057,1824,1850]" class="Insecta" family="Mutillidae" genus="Dasymutilla" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="269" phylum="Arthropoda" rank="species" species="homochroma">
<emphasis id="B93AEAF66750FFD162B7F881527DF8ED" box="[766,1057,1824,1850]" italics="true" pageId="8" pageNumber="269">Dasymutilla homochroma</emphasis>
</taxonomicName>
,
<specimenCount id="9D48FD6D6750FFD16479F8815219F8ED" box="[1072,1093,1824,1850]" pageId="8" pageNumber="269" type="female"></specimenCount>
,
<collectingCountry id="F35976746750FFD1641DF88152C2F8ED" box="[1108,1182,1824,1850]" name="Peru" pageId="8" pageNumber="269">PERU</collectingCountry>
, [
<collectingRegion id="498AF8066750FFD164FDF88152B2F8EE" box="[1204,1262,1824,1849]" country="Peru" name="Lima" pageId="8" pageNumber="269">
<emphasis id="B93AEAF66750FFD164FDF88152B2F8EE" box="[1204,1262,1824,1849]" italics="true" pageId="8" pageNumber="269">Lima</emphasis>
</collectingRegion>
?],
<location id="8E91603F6750FFD1655BF88153C4F8ED" LSID="urn:lsid:plazi:treatment:03E787F26750FFD560DEFBC4526FFDCC:8E91603F6750FFD1655BF88153C4F8ED" box="[1298,1432,1824,1850]" country="Peru" name="San Miguel" pageId="8" pageNumber="269" stateProvince="Lima">San Miguel</location>
,
<collectingDate id="EFB4E9CC6750FFD160DEF8E25744F889" box="[151,280,1859,1886]" pageId="8" pageNumber="269" value="1955-03-08">
<date id="FFF010246750FFD160DEF8E25744F889" box="[151,280,1859,1886]" pageId="8" pageNumber="269" value="1955-03-08">08.III.1955</date>
</collectingDate>
,
<collectorName id="26BB53326750FFD1616BF8E557D0F889" box="[290,396,1860,1886]" pageId="8" pageNumber="269">M. Markl</collectorName>
(
<collectionCode id="ED5FAE216750FFD161D2F8E557A9F889" box="[411,501,1860,1886]" country="0" httpUri="http://grbio.org/cool/1d5a-4efs" name="Natural History Museum Bucharest" pageId="8" pageNumber="269">NHMB</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B263CB96750FFD1624FF8E557EBF855" ID-GBIF-Occurrence="2269395012" collectingDate="1971-05-04" collectionCode="MIUP" collectorName="J. &amp; B. Bechyne" country="Peru" location="Desierto" pageId="8" pageNumber="269" specimenCount="1" specimenCount-female="1" stateProvince="Tumbes">
Other material:
<collectingRegion id="498AF8066750FFD162FFF8E55550F889" box="[694,780,1860,1886]" country="Peru" name="Tumbes" pageId="8" pageNumber="269">
<emphasis id="B93AEAF66750FFD162FFF8E55550F889" box="[694,780,1860,1886]" italics="true" pageId="8" pageNumber="269">Tumbes</emphasis>
</collectingRegion>
,
<location id="8E91603F6750FFD16351F8E55526F889" LSID="urn:lsid:plazi:treatment:03E787F26750FFD560DEFBC4526FFDCC:8E91603F6750FFD16351F8E55526F889" box="[792,890,1860,1886]" country="Peru" name="Desierto" pageId="8" pageNumber="269" stateProvince="Tumbes">Desierto</location>
, 4 horas en carro antes de
<collectingRegion id="498AF8066750FFD164D3F8E552AAF889" box="[1178,1270,1860,1886]" country="Peru" name="Tumbes" pageId="8" pageNumber="269">Tumbes</collectingRegion>
,
<specimenCount id="9D48FD6D6750FFD16548F8E5537FF889" box="[1281,1315,1860,1886]" pageId="8" pageNumber="269" type="female">1♀</specimenCount>
,
<collectingDate id="EFB4E9CC6750FFD16578F8E553CBF889" box="[1329,1431,1860,1886]" pageId="8" pageNumber="269" value="1971-05-04">
<date id="FFF010246750FFD16578F8E553CBF889" box="[1329,1431,1860,1886]" pageId="8" pageNumber="269" value="1971-05-04">4.V.1971</date>
</collectingDate>
,
<collectorName id="26BB53326750FFD160DEF8C9570FF855" box="[151,339,1896,1922]" pageId="8" pageNumber="269">J. &amp; B. Bechyne</collectorName>
, (
<collectionCode id="ED5FAE216750FFD1612CF8C957F2F855" box="[357,430,1896,1922]" pageId="8" pageNumber="269">MIUP</collectionCode>
)
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C354656F6750FFD5608EF82D526FFDCC" lastPageId="12" lastPageNumber="273" pageId="8" pageNumber="269" type="discussion">
<paragraph id="8BF136E46750FFD5608EF82D526FFDCC" blockId="8.[151,1437,1392,2030]" lastBlockId="12.[151,1437,153,539]" lastPageId="12" lastPageNumber="273" pageId="8" pageNumber="269">
<emphasis id="B93AEAF66750FFD1608EF82D576DF871" box="[199,305,1932,1958]" italics="true" pageId="8" pageNumber="269">Remarks.</emphasis>
Although
<taxonomicName id="4C4E4D676750FFD161E1F82D5460F872" authorityName="Suarez" authorityYear="1970" box="[424,572,1932,1958]" class="Insecta" family="Mutillidae" genus="Dasymutilla" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="269" phylum="Arthropoda" rank="species" species="peruviana">
<emphasis id="B93AEAF66750FFD161E1F82D5460F872" box="[424,572,1932,1958]" italics="true" pageId="8" pageNumber="269">Q. peruviana</emphasis>
</taxonomicName>
females are consistent in most aspects with the general morphological trends of the
<taxonomicName id="4C4E4D676750FFD16095F8115725F81D" authorityName="Brothers &amp; Lelej" authorityYear="2017" box="[220,377,1968,1994]" class="Insecta" family="Mutillidae" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="269" phylum="Arthropoda" rank="tribe" tribe="Dasymutillini">Dasymutillini</taxonomicName>
, the clypeus and mandibles of the males are even more distorted than the other known
<taxonomicName id="4C4E4D676750FFD1657BF81153C0F81D" authority="Bartholomay &amp; Williams &amp; Cambra &amp; Oliveira, 2019" authorityName="Bartholomay &amp; Williams &amp; Cambra &amp; Oliveira" authorityYear="2019" box="[1330,1436,1968,1994]" class="Insecta" family="Mutillidae" genus="Quwitilla" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="275" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B93AEAF66750FFD1657BF81153C0F81D" box="[1330,1436,1968,1994]" italics="true" pageId="8" pageNumber="269">Quwitilla</emphasis>
</taxonomicName>
males,
<taxonomicName id="4C4E4D676750FFD160AEF87557D3F839" authorityName="Bartholomay &amp; Williams &amp; Cambra &amp; Oliveira" authorityYear="2019" baseAuthorityName="Bartholomay &amp; Williams &amp; Cambra &amp; Oliveira" baseAuthorityYear="2019" box="[231,399,2004,2030]" class="Insecta" family="Mutillidae" genus="Quwitilla" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="269" phylum="Arthropoda" rank="species" species="blattoserica">
<emphasis id="B93AEAF66750FFD160AEF87557D3F839" box="[231,399,2004,2030]" italics="true" pageId="8" pageNumber="269">Q. blattoserica</emphasis>
</taxonomicName>
. It is clear, however that these belong to the newly described genus based on their genitalia, which only differs from those of
<taxonomicName id="4C4E4D676754FFD561B4FF3854FDFF64" authorityName="Bartholomay &amp; Williams &amp; Cambra &amp; Oliveira" authorityYear="2019" baseAuthorityName="Bartholomay &amp; Williams &amp; Cambra &amp; Oliveira" baseAuthorityYear="2019" box="[509,673,153,179]" class="Insecta" family="Mutillidae" genus="Quwitilla" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="273" phylum="Arthropoda" rank="species" species="blattoserica">
<emphasis id="B93AEAF66754FFD561B4FF3854FDFF64" box="[509,673,153,179]" italics="true" pageId="12" pageNumber="273">Q. blattoserica</emphasis>
</taxonomicName>
on the distribution of the paramere ventral setae.
<bibRefCitation id="EFDF4B156754FFD564F2FF385309FF64" author="Suarez, F. J." box="[1211,1365,153,179]" pageId="12" pageNumber="273" pagination="169 - 188" refId="ref10283" refString="Suarez, F. J. (1970) Datos sobre Mutilidos neotropicales VII. Algunas especies del Peru representadas en las colecciones del Museo de Historia Natural de Basilea (Suiza). Archivos del Instituto de Aclimataccion de Almeria, 15, 169 - 188." type="journal article" year="1970">Suárez (1970)</bibRefCitation>
raised the possibility of
<taxonomicName id="4C4E4D676754FFD5611EFF1C57BBFF00" authorityName="Bartholomay &amp; Williams &amp; Cambra &amp; Oliveira" authorityYear="2019" baseAuthorityName="Suarez" baseAuthorityYear="1970" box="[343,487,189,215]" class="Insecta" family="Mutillidae" genus="Quwitilla" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="273" phylum="Arthropoda" rank="species" species="peruviana">
<emphasis id="B93AEAF66754FFD5611EFF1C57BBFF00" box="[343,487,189,215]" italics="true" pageId="12" pageNumber="273">Q. peruviana</emphasis>
</taxonomicName>
being the male of
<taxonomicName id="4C4E4D676754FFD562FFFF1C5534FF00" box="[694,872,189,215]" class="Insecta" family="Mutillidae" genus="Dasymutilla" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="273" phylum="Arthropoda" rank="species" species="homochroma">
<emphasis id="B93AEAF66754FFD562FFFF1C5534FF00" box="[694,872,189,215]" italics="true" pageId="12" pageNumber="273">Q. homochroma</emphasis>
</taxonomicName>
and discarded the hypothesis based on the fact that while both sexes of
<taxonomicName id="4C4E4D676754FFD56133FF40547CFF2C" authorityName="Bartholomay &amp; Williams &amp; Cambra &amp; Oliveira" authorityYear="2019" baseAuthorityName="Bartholomay &amp; Williams &amp; Cambra &amp; Oliveira" baseAuthorityYear="2019" box="[378,544,225,251]" class="Insecta" family="Mutillidae" genus="Quwitilla" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="273" phylum="Arthropoda" rank="species" species="blattoserica">
<emphasis id="B93AEAF66754FFD56133FF40547CFF2C" box="[378,544,225,251]" italics="true" pageId="12" pageNumber="273">Q. blattoserica</emphasis>
</taxonomicName>
had coloration similarities,
<taxonomicName id="4C4E4D676754FFD56310FF4055B0FF2C" authorityName="Bartholomay &amp; Williams &amp; Cambra &amp; Oliveira" authorityYear="2019" baseAuthorityName="Suarez" baseAuthorityYear="1970" box="[857,1004,225,251]" class="Insecta" family="Mutillidae" genus="Quwitilla" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="273" phylum="Arthropoda" rank="species" species="peruviana">
<emphasis id="B93AEAF66754FFD56310FF4055B0FF2C" box="[857,1004,225,251]" italics="true" pageId="12" pageNumber="273">Q. peruviana</emphasis>
</taxonomicName>
and
<taxonomicName id="4C4E4D676754FFD5646CFF405284FF2C" box="[1061,1240,225,251]" class="Insecta" family="Mutillidae" genus="Dasymutilla" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="273" phylum="Arthropoda" rank="species" species="homochroma">
<emphasis id="B93AEAF66754FFD5646CFF405284FF2C" box="[1061,1240,225,251]" italics="true" pageId="12" pageNumber="273">Q. homochroma</emphasis>
</taxonomicName>
were remarkably different. We consider that having both species collected in the same date and place in an area where diurnal species of velvet-ant are uncommon, and their clear similarities with
<taxonomicName id="4C4E4D676754FFD56345FE8855F3FE94" authorityName="Bartholomay &amp; Williams &amp; Cambra &amp; Oliveira" authorityYear="2019" baseAuthorityName="Bartholomay &amp; Williams &amp; Cambra &amp; Oliveira" baseAuthorityYear="2019" box="[780,943,297,323]" class="Insecta" family="Mutillidae" genus="Quwitilla" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="273" phylum="Arthropoda" rank="species" species="blattoserica">
<emphasis id="B93AEAF66754FFD56345FE8855F3FE94" box="[780,943,297,323]" italics="true" pageId="12" pageNumber="273">Q. blattoserica</emphasis>
</taxonomicName>
are sufficient evidence for the sex association proposed here. As previously mentioned, the female of
<taxonomicName id="4C4E4D676754FFD56349FEEC55CEFEB0" authorityName="Bartholomay &amp; Williams &amp; Cambra &amp; Oliveira" authorityYear="2019" baseAuthorityName="Suarez" baseAuthorityYear="1970" box="[768,914,333,359]" class="Insecta" family="Mutillidae" genus="Quwitilla" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="273" phylum="Arthropoda" rank="species" species="peruviana">
<emphasis id="B93AEAF66754FFD56349FEEC55CEFEB0" box="[768,914,333,359]" italics="true" pageId="12" pageNumber="273">Q. peruviana</emphasis>
</taxonomicName>
appears to belong to a distinct color syndrome with other species west of the Andes. The male, however, remarkably approaches the color patterns observed in two other
<taxonomicName id="4C4E4D676754FFD5609CFE34571BFE78" authorityName="Latreille" authorityYear="1802" box="[213,327,405,431]" class="Insecta" family="Mutillidae" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="273" phylum="Arthropoda" rank="family">Mutillidae</taxonomicName>
species occurring west of the Andes:
<taxonomicName id="4C4E4D676754FFD56292FE3452BAFE78" authority="(Gerstaecker, 1874)" authorityName="Gerstaecker" authorityYear="1874" baseAuthorityName="Gerstaecker" baseAuthorityYear="1874" box="[731,1254,405,431]" class="Insecta" family="Mutillidae" genus="Traumatomutilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="273" phylum="Arthropoda" rank="species" species="vitelligera">
<emphasis id="B93AEAF66754FFD56292FE345255FE78" box="[731,1033,405,431]" italics="true" pageId="12" pageNumber="273">Traumatomutilla vitelligera</emphasis>
(
<bibRefCitation id="EFDF4B156754FFD5645FFE345281FE78" author="Gerstaecker, A." box="[1046,1245,405,431]" pageId="12" pageNumber="273" pagination="41 - 77" refId="ref9443" refString="Gerstaecker, A. (1874) Mutillarum Americae meridionalis indigenarum synopsis systematica et synonymica. Archiv fur Naturgeschichte, 40, 41 - 77 + 299 - 328." type="journal article" year="1874">Gerstaecker, 1874</bibRefCitation>
)
</taxonomicName>
and
<taxonomicName id="4C4E4D676754FFD56551FE3657E6FE03" authority="(Andre, 1906)" authorityName="Andre" authorityYear="1906" baseAuthorityName="Andre" baseAuthorityYear="1906" class="Insecta" family="Mutillidae" genus="Hoplomutilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="273" phylum="Arthropoda" rank="species" species="maculifrons">
<emphasis id="B93AEAF66754FFD56551FE365745FE04" italics="true" pageId="12" pageNumber="273">Hoplomutilla maculifrons</emphasis>
(André, 1906)
</taxonomicName>
(
<figureCitation id="13752A616754FFD56181FE185414FE04" box="[456,584,441,467]" captionStart="FIGURES 3840" captionStartId="16.[151,264,1940,1965]" captionTargetBox="[271,1316,181,1920]" captionTargetId="figure@16.[271,1316,181,1920]" captionTargetPageId="16" captionText="FIGURES 3840. 38. Quwitilla peruviana (Suárez, 1970), ♂, holotype; 39. Hoplomutilla maculifrons (André, 1906); 40. Traumatomutilla vitelligera (Gerstaecker, 1874)." httpUri="https://zenodo.org/record/3255433/files/figure.png" pageId="12" pageNumber="273">Figs 3840</figureCitation>
). This could be an example of Dual Sex-limited Mimicry, which was first proposed by
<bibRefCitation id="EFDF4B156754FFD56148FE7C57D2FE20" author="Evans, H. E." box="[257,398,477,503]" pageId="12" pageNumber="273" pagination="1 - 22" refId="ref9354" refString="Evans, H. E. (1968) Studies on Neotropical Pompilidae (Hymenoptera), IV. Examples of Dual Sex-Limited Mimicry in Chirodamus. Psyche, 75, 1 - 22. https: // doi. org / 10.1155 / 1968 / 76089" type="journal article" year="1968">Evans (1968</bibRefCitation>
,
<bibRefCitation id="EFDF4B156754FFD561D3FE7C578EFE2F" author="Evans, H. E." box="[410,466,477,504]" pageId="12" pageNumber="273" pagination="18 - 28" refId="ref9401" refString="Evans, H. E. (1969) Studies on Neotropical Pompilidae (Hymenoptera), V. Austrochares Banks. Psyche, 76, 18 - 28. https: // doi. org / 10.1155 / 1969 / 50293" type="journal article" year="1969">1969</bibRefCitation>
) in Neotropical pompilids, but has also been observed in
<taxonomicName id="4C4E4D676754FFD56407FE7C53C5FE20" authority="(Williams, 2012)" authorityName="Williams" authorityYear="2012" baseAuthorityName="Williams" baseAuthorityYear="2012" box="[1102,1433,477,503]" class="Insecta" family="Mutillidae" genus="Dasymutilla" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="273" phylum="Arthropoda" rank="genus">
<emphasis id="B93AEAF66754FFD56407FE7C5288FE20" box="[1102,1236,477,503]" italics="true" pageId="12" pageNumber="273">Dasymutilla</emphasis>
(
<bibRefCitation id="EFDF4B156754FFD564ABFE7C53CDFE20" author="Williams, K. A." box="[1250,1425,477,503]" pageId="12" pageNumber="273" refId="ref10406" refString="Williams, K. A. (2012) Systematics of Mutillidae (Hymenoptera) with special emphasis on Dasymutilla and their alles. All Graduate Theses and Dissertations. Paper 1200. Utah State Univeristy Press, Logan, Utah, 327 pp. Available from: http: // digitalcommons. usu. edu / etd / 1200 / (Las accessed November 2018)" type="book" year="2012">Williams, 2012</bibRefCitation>
)
</taxonomicName>
. Further studies into the mimetic syndromes of South American mutillids are needed.
</paragraph>
</subSubSection>
</treatment>
</document>