274 lines
57 KiB
XML
274 lines
57 KiB
XML
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<mods:title id="7F08F4593B2EAC51AB3C479E94F006C6">New species and first record of Helotrephes from India, and a check-list of Indian Helotrephidae (Hemiptera: Heteroptera)</mods:title>
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<mods:namePart id="E4FBCB53D8AFE4D618FE5D7400A12FFF">Jehamalar, Eyarin</mods:namePart>
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<mods:namePart id="A54934F2D8A71341740EC90631483E04">Chandra, Kailash</mods:namePart>
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<mods:namePart id="18DD554DD767B20AEF185B017D878275">Zettel, Herbert</mods:namePart>
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<treatment id="03E787D23857FFBD11988EAF1696CF94" ID-DOI="http://doi.org/10.5281/zenodo.3705454" ID-GBIF-Taxon="162462613" ID-Zenodo-Dep="3705454" LSID="urn:lsid:plazi:treatment:03E787D23857FFBD11988EAF1696CF94" httpUri="http://treatment.plazi.org/id/03E787D23857FFBD11988EAF1696CF94" lastPageNumber="245" pageId="2" pageNumber="245">
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<subSubSection id="C354654F3857FFBD11988EAF11F2CDB2" pageId="2" pageNumber="245" type="nomenclature">
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<paragraph id="8BF136C43857FFBD11988EAF1117CCE5" blockId="2.[182,729,685,740]" pageId="2" pageNumber="245">
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<heading id="D0B981A83857FFBD11988EAF1117CCE5" centered="true" fontSize="10" level="2" pageId="2" pageNumber="245" reason="3">
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<emphasis id="B93AEAD63857FFBD11988EAF11D3CCC5" bold="true" box="[182,729,685,708]" pageId="2" pageNumber="245">
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Key to the species of
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<taxonomicName id="4C4E4D473857FFBD108A8EAF1185CCC5" ID-CoL="6LMS7" authorityName="Zettel & Polhemus" authorityYear="1998" box="[420,655,685,708]" class="Insecta" family="Helotrephidae" genus="Helotrephes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="245" phylum="Arthropoda" rank="species" species="australis">
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<emphasis id="B93AEAD63857FFBD108A8EAF1185CCC5" bold="true" box="[420,655,685,708]" italics="true" pageId="2" pageNumber="245">Helotrephes australis</emphasis>
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</taxonomicName>
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group
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</emphasis>
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(only for males)
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</heading>
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</paragraph>
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<paragraph id="8BF136C43857FFBD11B98EFE11F2CDB2" blockId="2.[151,760,764,947]" pageId="2" pageNumber="245">
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<emphasis id="B93AEAD63857FFBD11B98EFE13DECD12" bold="true" box="[151,212,764,787]" pageId="2" pageNumber="245">Note.</emphasis>
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<emphasis id="B93AEAD63857FFBD11F78EFE12EBCD12" box="[217,481,764,787]" italics="true" pageId="2" pageNumber="245">
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<taxonomicName id="4C4E4D473857FFBD11F78EFE1256CD12" authorityName="Stal" authorityYear="1860" box="[217,348,764,787]" class="Insecta" family="Helotrephidae" genus="Helotrephes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="245" phylum="Arthropoda" rank="genus">Helotrephes</taxonomicName>
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kantonensis
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<bibRefCitation id="EFDF4B353857FFBD10C88EFE116DCD12" author="Zettel" box="[486,615,764,787]" firstAuthor="Zettel" pageId="2" pageNumber="245" pagination="397 - 409" refId="ref4584" refString="ZETTEL H. 2004 a: Helotrephidae (Insecta: Heteroptera) aus den chinesischen Provinzen Guangdong, Yunnan und Guizhou, mit Beschreibungen neuer Arten der Gattungen Helotrephes und Distotrephes. Annalen des Naturhistorischen Museums in Wien 105 B: 397 - 409." type="journal article" year="2004">Zettel, 2004</bibRefCitation>
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is not includ- ed in the key applicable only for male adults, since it is known only from a single female. It can be identified by a deep and wide incision of the pronotal plate, an angular anterior projection of the prosternal carina, and slightly angular carinae of metasternum and abdominal sternum 2.
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</paragraph>
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</subSubSection>
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<subSubSection id="C354654F3857FFBD11B98FC91696CF94" pageId="2" pageNumber="245" type="key">
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<keyStep id="30BA2E613857FFBD11B98FC911FDCA63" pageId="2" pageNumber="245">
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<paragraph id="8BF136C43857FFBD11B98FC911FDCA23" blockId="2.[151,760,971,2082]" pageId="2" pageNumber="245">
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<keyLead id="30BF95F13857FFBD11B98FC911FDCA23" pageId="2" pageNumber="245">
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1. Apex of aedeagus hook-shaped and facing backward (
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<figureCitation id="13752A413857FFBD11EB8FE91217CA03" box="[197,285,1003,1026]" captionStart="Fig" captionStartId="1.[151,182,1930,1949]" captionTargetBox="[299,1290,638,1908]" captionTargetId="figure@1.[298,1290,635,1911]" captionTargetPageId="1" captionText="Fig. 1. Key characters of species belonging to the Helotrephes australis species-group.A–E – aedeagi in right side view:A – H. recurvatus Zettel, 2000; B – H. australis Zettel & Polhemus, 1998; C – H. nieserianus Zettel & Polhemus, 1998; D – H. incisus Zettel & Polhemus, 1998; E – H. vietnamensis Zettel, 2005. F–I – pronotal and genal plate: F – H. recurvatus; G – H. globulus Zettel, 2004; H – H. australis; I – H. incisus. J – right paramere of H. australis. K – left paramere of H. australis. L–O – medioventral carinae: L – H. globulus; M – H. australis; N – H. recurvatus; O – H. incisus. Abbreviations: pr – prosternum; ms – mesosternum, mt – metasternum, 2–6 – carinae of abdominal sterna 2–6. Figures not on scale. Redrawn (with slight modifications) from ZETTEL & POLHEMUS (1998): B–D, H–K, M, O; ZETTEL (2000a): A, F, N; ZETTEL (2004a): G, L; and ZETTEL (2005): E." figureDoi="http://doi.org/10.5281/zenodo.3699242" httpUri="https://zenodo.org/record/3699242/files/figure.png" pageId="2" pageNumber="245">Fig. 1A</figureCitation>
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). Incision of pronotal plate deep and wide (
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<figureCitation id="13752A413857FFBD11EB8809121ECA23" box="[197,276,1035,1058]" captionStart="Fig" captionStartId="1.[151,182,1930,1949]" captionTargetBox="[299,1290,638,1908]" captionTargetId="figure@1.[298,1290,635,1911]" captionTargetPageId="1" captionText="Fig. 1. Key characters of species belonging to the Helotrephes australis species-group.A–E – aedeagi in right side view:A – H. recurvatus Zettel, 2000; B – H. australis Zettel & Polhemus, 1998; C – H. nieserianus Zettel & Polhemus, 1998; D – H. incisus Zettel & Polhemus, 1998; E – H. vietnamensis Zettel, 2005. F–I – pronotal and genal plate: F – H. recurvatus; G – H. globulus Zettel, 2004; H – H. australis; I – H. incisus. J – right paramere of H. australis. K – left paramere of H. australis. L–O – medioventral carinae: L – H. globulus; M – H. australis; N – H. recurvatus; O – H. incisus. Abbreviations: pr – prosternum; ms – mesosternum, mt – metasternum, 2–6 – carinae of abdominal sterna 2–6. Figures not on scale. Redrawn (with slight modifications) from ZETTEL & POLHEMUS (1998): B–D, H–K, M, O; ZETTEL (2000a): A, F, N; ZETTEL (2004a): G, L; and ZETTEL (2005): E." figureDoi="http://doi.org/10.5281/zenodo.3699242" httpUri="https://zenodo.org/record/3699242/files/figure.png" pageId="2" pageNumber="245">Fig. 1F</figureCitation>
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), or narrow and angular (
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<figureCitation id="13752A413857FFBD130A88091173CA23" box="[548,633,1035,1058]" captionStart="Fig" captionStartId="1.[151,182,1930,1949]" captionTargetBox="[299,1290,638,1908]" captionTargetId="figure@1.[298,1290,635,1911]" captionTargetPageId="1" captionText="Fig. 1. Key characters of species belonging to the Helotrephes australis species-group.A–E – aedeagi in right side view:A – H. recurvatus Zettel, 2000; B – H. australis Zettel & Polhemus, 1998; C – H. nieserianus Zettel & Polhemus, 1998; D – H. incisus Zettel & Polhemus, 1998; E – H. vietnamensis Zettel, 2005. F–I – pronotal and genal plate: F – H. recurvatus; G – H. globulus Zettel, 2004; H – H. australis; I – H. incisus. J – right paramere of H. australis. K – left paramere of H. australis. L–O – medioventral carinae: L – H. globulus; M – H. australis; N – H. recurvatus; O – H. incisus. Abbreviations: pr – prosternum; ms – mesosternum, mt – metasternum, 2–6 – carinae of abdominal sterna 2–6. Figures not on scale. Redrawn (with slight modifications) from ZETTEL & POLHEMUS (1998): B–D, H–K, M, O; ZETTEL (2000a): A, F, N; ZETTEL (2004a): G, L; and ZETTEL (2005): E." figureDoi="http://doi.org/10.5281/zenodo.3699242" httpUri="https://zenodo.org/record/3699242/files/figure.png" pageId="2" pageNumber="245">Fig. 1G</figureCitation>
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). ............ 2
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</keyLead>
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</paragraph>
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<paragraph id="8BF136C43857FFBD11B9882911FDCA63" blockId="2.[151,760,971,2082]" pageId="2" pageNumber="245">
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<keyLead id="30BF95F13857FFBD11B9882911FDCA63" pageId="2" pageNumber="245">
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– Apex of aedeagus facing forward (
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<figureCitation id="13752A413857FFBD136F882911B2CA43" box="[577,696,1067,1090]" captionStart="Fig" captionStartId="1.[151,182,1930,1949]" captionTargetBox="[299,1290,638,1908]" captionTargetId="figure@1.[298,1290,635,1911]" captionTargetPageId="1" captionText="Fig. 1. Key characters of species belonging to the Helotrephes australis species-group.A–E – aedeagi in right side view:A – H. recurvatus Zettel, 2000; B – H. australis Zettel & Polhemus, 1998; C – H. nieserianus Zettel & Polhemus, 1998; D – H. incisus Zettel & Polhemus, 1998; E – H. vietnamensis Zettel, 2005. F–I – pronotal and genal plate: F – H. recurvatus; G – H. globulus Zettel, 2004; H – H. australis; I – H. incisus. J – right paramere of H. australis. K – left paramere of H. australis. L–O – medioventral carinae: L – H. globulus; M – H. australis; N – H. recurvatus; O – H. incisus. Abbreviations: pr – prosternum; ms – mesosternum, mt – metasternum, 2–6 – carinae of abdominal sterna 2–6. Figures not on scale. Redrawn (with slight modifications) from ZETTEL & POLHEMUS (1998): B–D, H–K, M, O; ZETTEL (2000a): A, F, N; ZETTEL (2004a): G, L; and ZETTEL (2005): E." figureDoi="http://doi.org/10.5281/zenodo.3699242" httpUri="https://zenodo.org/record/3699242/files/figure.png" pageId="2" pageNumber="245">Figs 1B–E</figureCitation>
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,
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<figureCitation id="13752A413857FFBD13E6882911E6CA43" box="[712,748,1067,1090]" pageId="2" pageNumber="245">2G</figureCitation>
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). Incision on pronotal plate round (
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<figureCitation id="13752A413857FFBD13368849118FCA63" box="[536,645,1099,1122]" captionStart="Fig" captionStartId="1.[151,182,1930,1949]" captionTargetBox="[299,1290,638,1908]" captionTargetId="figure@1.[298,1290,635,1911]" captionTargetPageId="1" captionText="Fig. 1. Key characters of species belonging to the Helotrephes australis species-group.A–E – aedeagi in right side view:A – H. recurvatus Zettel, 2000; B – H. australis Zettel & Polhemus, 1998; C – H. nieserianus Zettel & Polhemus, 1998; D – H. incisus Zettel & Polhemus, 1998; E – H. vietnamensis Zettel, 2005. F–I – pronotal and genal plate: F – H. recurvatus; G – H. globulus Zettel, 2004; H – H. australis; I – H. incisus. J – right paramere of H. australis. K – left paramere of H. australis. L–O – medioventral carinae: L – H. globulus; M – H. australis; N – H. recurvatus; O – H. incisus. Abbreviations: pr – prosternum; ms – mesosternum, mt – metasternum, 2–6 – carinae of abdominal sterna 2–6. Figures not on scale. Redrawn (with slight modifications) from ZETTEL & POLHEMUS (1998): B–D, H–K, M, O; ZETTEL (2000a): A, F, N; ZETTEL (2004a): G, L; and ZETTEL (2005): E." figureDoi="http://doi.org/10.5281/zenodo.3699242" httpUri="https://zenodo.org/record/3699242/files/figure.png" pageId="2" pageNumber="245">Figs 1H–I</figureCitation>
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,
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<figureCitation id="13752A413857FFBD13A1884911BBCA63" box="[655,689,1099,1122]" pageId="2" pageNumber="245">2C</figureCitation>
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). .... 3
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</keyLead>
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</paragraph>
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</keyStep>
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<keyStep id="30BA2E613857FFBD11B9886911FDCB43" pageId="2" pageNumber="245">
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<paragraph id="8BF136C43857FFBD11B9886911FDCAE3" blockId="2.[151,760,971,2082]" pageId="2" pageNumber="245">
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<keyLead id="30BF95F13857FFBD11B9886911FDCAE3" pageId="2" pageNumber="245">
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2. Incision of pronotal plate narrow and angular (
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<figureCitation id="13752A413857FFBD13E2886913EACAA3" captionStart="Fig" captionStartId="1.[151,182,1930,1949]" captionTargetBox="[299,1290,638,1908]" captionTargetId="figure@1.[298,1290,635,1911]" captionTargetPageId="1" captionText="Fig. 1. Key characters of species belonging to the Helotrephes australis species-group.A–E – aedeagi in right side view:A – H. recurvatus Zettel, 2000; B – H. australis Zettel & Polhemus, 1998; C – H. nieserianus Zettel & Polhemus, 1998; D – H. incisus Zettel & Polhemus, 1998; E – H. vietnamensis Zettel, 2005. F–I – pronotal and genal plate: F – H. recurvatus; G – H. globulus Zettel, 2004; H – H. australis; I – H. incisus. J – right paramere of H. australis. K – left paramere of H. australis. L–O – medioventral carinae: L – H. globulus; M – H. australis; N – H. recurvatus; O – H. incisus. Abbreviations: pr – prosternum; ms – mesosternum, mt – metasternum, 2–6 – carinae of abdominal sterna 2–6. Figures not on scale. Redrawn (with slight modifications) from ZETTEL & POLHEMUS (1998): B–D, H–K, M, O; ZETTEL (2000a): A, F, N; ZETTEL (2004a): G, L; and ZETTEL (2005): E." figureDoi="http://doi.org/10.5281/zenodo.3699242" httpUri="https://zenodo.org/record/3699242/files/figure.png" pageId="2" pageNumber="245">Fig. 1G</figureCitation>
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). Anterior projection of prosternal carina blunt (
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<figureCitation id="13752A413857FFBD11EB88A9121CCAC3" box="[197,278,1195,1218]" captionStart="Fig" captionStartId="1.[151,182,1930,1949]" captionTargetBox="[299,1290,638,1908]" captionTargetId="figure@1.[298,1290,635,1911]" captionTargetPageId="1" captionText="Fig. 1. Key characters of species belonging to the Helotrephes australis species-group.A–E – aedeagi in right side view:A – H. recurvatus Zettel, 2000; B – H. australis Zettel & Polhemus, 1998; C – H. nieserianus Zettel & Polhemus, 1998; D – H. incisus Zettel & Polhemus, 1998; E – H. vietnamensis Zettel, 2005. F–I – pronotal and genal plate: F – H. recurvatus; G – H. globulus Zettel, 2004; H – H. australis; I – H. incisus. J – right paramere of H. australis. K – left paramere of H. australis. L–O – medioventral carinae: L – H. globulus; M – H. australis; N – H. recurvatus; O – H. incisus. Abbreviations: pr – prosternum; ms – mesosternum, mt – metasternum, 2–6 – carinae of abdominal sterna 2–6. Figures not on scale. Redrawn (with slight modifications) from ZETTEL & POLHEMUS (1998): B–D, H–K, M, O; ZETTEL (2000a): A, F, N; ZETTEL (2004a): G, L; and ZETTEL (2005): E." figureDoi="http://doi.org/10.5281/zenodo.3699242" httpUri="https://zenodo.org/record/3699242/files/figure.png" pageId="2" pageNumber="245">Fig. 1L</figureCitation>
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).
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<collectingCountry id="F35976543857FFBD100588A91261CAC3" box="[299,363,1195,1218]" name="China" pageId="2" pageNumber="245">China</collectingCountry>
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(
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<collectingRegion id="498AF8263857FFBD105688A912F1CAC3" box="[376,507,1195,1218]" country="China" name="Guangdong" pageId="2" pageNumber="245">Guangdong</collectingRegion>
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||
). ................................... ............................................
|
||
<taxonomicName id="4C4E4D473857FFBD10C188CE11FDCAE3" ID-CoL="6LN3V" authority="Zettel, 2004" authorityName="Zettel" authorityYear="2004" box="[495,759,1227,1251]" class="Insecta" family="Helotrephidae" genus="Helotrephes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="245" phylum="Arthropoda" rank="species" species="globulus">
|
||
<emphasis id="B93AEAD63857FFBD10C188CE117ACAE3" bold="true" box="[495,624,1227,1251]" italics="true" pageId="2" pageNumber="245">H. globulus</emphasis>
|
||
<bibRefCitation id="EFDF4B353857FFBD135988C911FDCAE3" author="Zettel" box="[631,759,1227,1250]" firstAuthor="Zettel" pageId="2" pageNumber="245" pagination="397 - 409" refId="ref4584" refString="ZETTEL H. 2004 a: Helotrephidae (Insecta: Heteroptera) aus den chinesischen Provinzen Guangdong, Yunnan und Guizhou, mit Beschreibungen neuer Arten der Gattungen Helotrephes und Distotrephes. Annalen des Naturhistorischen Museums in Wien 105 B: 397 - 409." type="journal article" year="2004">Zettel, 2004</bibRefCitation>
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||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BF136C43857FFBD11B988E911FDCB43" blockId="2.[151,760,971,2082]" pageId="2" pageNumber="245">
|
||
<keyLead id="30BF95F13857FFBD11B988E911FDCB43" pageId="2" pageNumber="245">
|
||
– Incision of pronotal plate deep and wide (
|
||
<figureCitation id="13752A413857FFBD13B888E911E0CB03" box="[662,746,1259,1282]" captionStart="Fig" captionStartId="1.[151,182,1930,1949]" captionTargetBox="[299,1290,638,1908]" captionTargetId="figure@1.[298,1290,635,1911]" captionTargetPageId="1" captionText="Fig. 1. Key characters of species belonging to the Helotrephes australis species-group.A–E – aedeagi in right side view:A – H. recurvatus Zettel, 2000; B – H. australis Zettel & Polhemus, 1998; C – H. nieserianus Zettel & Polhemus, 1998; D – H. incisus Zettel & Polhemus, 1998; E – H. vietnamensis Zettel, 2005. F–I – pronotal and genal plate: F – H. recurvatus; G – H. globulus Zettel, 2004; H – H. australis; I – H. incisus. J – right paramere of H. australis. K – left paramere of H. australis. L–O – medioventral carinae: L – H. globulus; M – H. australis; N – H. recurvatus; O – H. incisus. Abbreviations: pr – prosternum; ms – mesosternum, mt – metasternum, 2–6 – carinae of abdominal sterna 2–6. Figures not on scale. Redrawn (with slight modifications) from ZETTEL & POLHEMUS (1998): B–D, H–K, M, O; ZETTEL (2000a): A, F, N; ZETTEL (2004a): G, L; and ZETTEL (2005): E." figureDoi="http://doi.org/10.5281/zenodo.3699242" httpUri="https://zenodo.org/record/3699242/files/figure.png" pageId="2" pageNumber="245">Fig. 1F</figureCitation>
|
||
). Anterior projection of prosternal carina angulated (
|
||
<figureCitation id="13752A413857FFBD11EB89291210CB43" box="[197,282,1323,1346]" captionStart="Fig" captionStartId="1.[151,182,1930,1949]" captionTargetBox="[299,1290,638,1908]" captionTargetId="figure@1.[298,1290,635,1911]" captionTargetPageId="1" captionText="Fig. 1. Key characters of species belonging to the Helotrephes australis species-group.A–E – aedeagi in right side view:A – H. recurvatus Zettel, 2000; B – H. australis Zettel & Polhemus, 1998; C – H. nieserianus Zettel & Polhemus, 1998; D – H. incisus Zettel & Polhemus, 1998; E – H. vietnamensis Zettel, 2005. F–I – pronotal and genal plate: F – H. recurvatus; G – H. globulus Zettel, 2004; H – H. australis; I – H. incisus. J – right paramere of H. australis. K – left paramere of H. australis. L–O – medioventral carinae: L – H. globulus; M – H. australis; N – H. recurvatus; O – H. incisus. Abbreviations: pr – prosternum; ms – mesosternum, mt – metasternum, 2–6 – carinae of abdominal sterna 2–6. Figures not on scale. Redrawn (with slight modifications) from ZETTEL & POLHEMUS (1998): B–D, H–K, M, O; ZETTEL (2000a): A, F, N; ZETTEL (2004a): G, L; and ZETTEL (2005): E." figureDoi="http://doi.org/10.5281/zenodo.3699242" httpUri="https://zenodo.org/record/3699242/files/figure.png" pageId="2" pageNumber="245">Fig. 1N</figureCitation>
|
||
). Borneo. ..........
|
||
<taxonomicName id="4C4E4D473857FFBD10F7892E11FDCB43" ID-CoL="3KJTW" authority="Zettel, 2000" authorityName="Zettel" authorityYear="2000" box="[473,759,1323,1347]" class="Insecta" family="Helotrephidae" genus="Helotrephes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="245" phylum="Arthropoda" rank="species" species="recurvatus">
|
||
<emphasis id="B93AEAD63857FFBD10F7892E117ACB42" bold="true" box="[473,624,1324,1347]" italics="true" pageId="2" pageNumber="245">H. recurvatus</emphasis>
|
||
<bibRefCitation id="EFDF4B353857FFBD1359892911FDCB43" author="Zettel" box="[631,759,1323,1346]" firstAuthor="Zettel" pageId="2" pageNumber="245" pagination="1 - 22" refId="ref4497" refString="ZETTEL H. 2000 a: The Helotrephidae (Heteroptera) of Borneo. Entomological Problems 31: 1 - 22." type="journal article" year="2000">Zettel, 2000</bibRefCitation>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30BA2E613857FFBD11B9894911FDC823" pageId="2" pageNumber="245">
|
||
<paragraph id="8BF136C43857FFBD11B9894911FDCBA3" blockId="2.[151,760,971,2082]" pageId="2" pageNumber="245">
|
||
<keyLead id="30BF95F13857FFBD11B9894911FDCBA3" pageId="2" pageNumber="245">
|
||
3. Incision of pronotal plate constricted at mesal margin of plate (
|
||
<figureCitation id="13752A413857FFBD103789691267CB83" box="[281,365,1387,1410]" captionStart="Fig" captionStartId="1.[151,182,1930,1949]" captionTargetBox="[299,1290,638,1908]" captionTargetId="figure@1.[298,1290,635,1911]" captionTargetPageId="1" captionText="Fig. 1. Key characters of species belonging to the Helotrephes australis species-group.A–E – aedeagi in right side view:A – H. recurvatus Zettel, 2000; B – H. australis Zettel & Polhemus, 1998; C – H. nieserianus Zettel & Polhemus, 1998; D – H. incisus Zettel & Polhemus, 1998; E – H. vietnamensis Zettel, 2005. F–I – pronotal and genal plate: F – H. recurvatus; G – H. globulus Zettel, 2004; H – H. australis; I – H. incisus. J – right paramere of H. australis. K – left paramere of H. australis. L–O – medioventral carinae: L – H. globulus; M – H. australis; N – H. recurvatus; O – H. incisus. Abbreviations: pr – prosternum; ms – mesosternum, mt – metasternum, 2–6 – carinae of abdominal sterna 2–6. Figures not on scale. Redrawn (with slight modifications) from ZETTEL & POLHEMUS (1998): B–D, H–K, M, O; ZETTEL (2000a): A, F, N; ZETTEL (2004a): G, L; and ZETTEL (2005): E." figureDoi="http://doi.org/10.5281/zenodo.3699242" httpUri="https://zenodo.org/record/3699242/files/figure.png" pageId="2" pageNumber="245">Fig. 1H</figureCitation>
|
||
). In apical view, aedeagus with apical lamina (
|
||
<figureCitation id="13752A413857FFBD103A89891281CBA3" box="[276,395,1419,1442]" captionStart="Fig" captionStartId="1.[151,182,1930,1949]" captionTargetBox="[299,1290,638,1908]" captionTargetId="figure@1.[298,1290,635,1911]" captionTargetPageId="1" captionText="Fig. 1. Key characters of species belonging to the Helotrephes australis species-group.A–E – aedeagi in right side view:A – H. recurvatus Zettel, 2000; B – H. australis Zettel & Polhemus, 1998; C – H. nieserianus Zettel & Polhemus, 1998; D – H. incisus Zettel & Polhemus, 1998; E – H. vietnamensis Zettel, 2005. F–I – pronotal and genal plate: F – H. recurvatus; G – H. globulus Zettel, 2004; H – H. australis; I – H. incisus. J – right paramere of H. australis. K – left paramere of H. australis. L–O – medioventral carinae: L – H. globulus; M – H. australis; N – H. recurvatus; O – H. incisus. Abbreviations: pr – prosternum; ms – mesosternum, mt – metasternum, 2–6 – carinae of abdominal sterna 2–6. Figures not on scale. Redrawn (with slight modifications) from ZETTEL & POLHEMUS (1998): B–D, H–K, M, O; ZETTEL (2000a): A, F, N; ZETTEL (2004a): G, L; and ZETTEL (2005): E." figureDoi="http://doi.org/10.5281/zenodo.3699242" httpUri="https://zenodo.org/record/3699242/files/figure.png" pageId="2" pageNumber="245">Figs 1B–C</figureCitation>
|
||
, inserts). .................................... 4
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BF136C43857FFBD11B989A911FDC823" blockId="2.[151,760,971,2082]" pageId="2" pageNumber="245">
|
||
<keyLead id="30BF95F13857FFBD11B989A911FDC823" pageId="2" pageNumber="245">
|
||
– Incision on the pronotal plate not constricted at mesal margin of plate (
|
||
<figureCitation id="13752A413857FFBD104489C912BDCBE3" box="[362,439,1483,1506]" captionStart="Fig" captionStartId="1.[151,182,1930,1949]" captionTargetBox="[299,1290,638,1908]" captionTargetId="figure@1.[298,1290,635,1911]" captionTargetPageId="1" captionText="Fig. 1. Key characters of species belonging to the Helotrephes australis species-group.A–E – aedeagi in right side view:A – H. recurvatus Zettel, 2000; B – H. australis Zettel & Polhemus, 1998; C – H. nieserianus Zettel & Polhemus, 1998; D – H. incisus Zettel & Polhemus, 1998; E – H. vietnamensis Zettel, 2005. F–I – pronotal and genal plate: F – H. recurvatus; G – H. globulus Zettel, 2004; H – H. australis; I – H. incisus. J – right paramere of H. australis. K – left paramere of H. australis. L–O – medioventral carinae: L – H. globulus; M – H. australis; N – H. recurvatus; O – H. incisus. Abbreviations: pr – prosternum; ms – mesosternum, mt – metasternum, 2–6 – carinae of abdominal sterna 2–6. Figures not on scale. Redrawn (with slight modifications) from ZETTEL & POLHEMUS (1998): B–D, H–K, M, O; ZETTEL (2000a): A, F, N; ZETTEL (2004a): G, L; and ZETTEL (2005): E." figureDoi="http://doi.org/10.5281/zenodo.3699242" httpUri="https://zenodo.org/record/3699242/files/figure.png" pageId="2" pageNumber="245">Figs 1I</figureCitation>
|
||
,
|
||
<figureCitation id="13752A413857FFBD10EC89C912EFCBE3" box="[450,485,1483,1506]" captionStart="Fig" captionStartId="3.[151,182,1558,1577]" captionTargetBox="[255,1334,191,1536]" captionTargetId="figure@3.[254,1334,189,1538]" captionTargetPageId="3" captionText="Fig. 2. Helotrephes nainamalaii sp. nov. (holotype, brachypterous male).A – habitus, dorsal view; B – head, frontal view; C – pronotal and genal plate, ventral view (arrows indicate incision of pronotal plate and sub-basal tubercle of genal plate); D – ventromedian carinae, lateral view (arrows indicate strongly protruded prosternal carina and blunt posterior corner of metasternal carina); E – genital capsule, lateral view; F – apex of right paramere, dorsolateral view; G – aedeagus, lateral view; H–, right paramere, lateral view; I – left paramere, lateral view." figureDoi="http://doi.org/10.5281/zenodo.3699244" httpUri="https://zenodo.org/record/3699244/files/figure.png" pageId="2" pageNumber="245">2C</figureCitation>
|
||
). In apical view, aedeagus sharp, without apical lamina (
|
||
<figureCitation id="13752A413857FFBD133489E91192C803" box="[538,664,1515,1538]" captionStart="Fig" captionStartId="1.[151,182,1930,1949]" captionTargetBox="[299,1290,638,1908]" captionTargetId="figure@1.[298,1290,635,1911]" captionTargetPageId="1" captionText="Fig. 1. Key characters of species belonging to the Helotrephes australis species-group.A–E – aedeagi in right side view:A – H. recurvatus Zettel, 2000; B – H. australis Zettel & Polhemus, 1998; C – H. nieserianus Zettel & Polhemus, 1998; D – H. incisus Zettel & Polhemus, 1998; E – H. vietnamensis Zettel, 2005. F–I – pronotal and genal plate: F – H. recurvatus; G – H. globulus Zettel, 2004; H – H. australis; I – H. incisus. J – right paramere of H. australis. K – left paramere of H. australis. L–O – medioventral carinae: L – H. globulus; M – H. australis; N – H. recurvatus; O – H. incisus. Abbreviations: pr – prosternum; ms – mesosternum, mt – metasternum, 2–6 – carinae of abdominal sterna 2–6. Figures not on scale. Redrawn (with slight modifications) from ZETTEL & POLHEMUS (1998): B–D, H–K, M, O; ZETTEL (2000a): A, F, N; ZETTEL (2004a): G, L; and ZETTEL (2005): E." figureDoi="http://doi.org/10.5281/zenodo.3699242" httpUri="https://zenodo.org/record/3699242/files/figure.png" pageId="2" pageNumber="245">Figs 1D–E</figureCitation>
|
||
, inserts, 2G). .......................................................................... 5
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30BA2E613857FFBD11B98A2911FDC923" pageId="2" pageNumber="245">
|
||
<paragraph id="8BF136C43857FFBD11B98A2911FDC8A3" blockId="2.[151,760,971,2082]" pageId="2" pageNumber="245">
|
||
<keyLead id="30BF95F13857FFBD11B98A2911FDC8A3" pageId="2" pageNumber="245">
|
||
4. Anterior projection of prosternal carina spinous, pointed (
|
||
<figureCitation id="13752A413857FFBD100F8A49128AC863" box="[289,384,1611,1634]" captionStart="Fig" captionStartId="1.[151,182,1930,1949]" captionTargetBox="[299,1290,638,1908]" captionTargetId="figure@1.[298,1290,635,1911]" captionTargetPageId="1" captionText="Fig. 1. Key characters of species belonging to the Helotrephes australis species-group.A–E – aedeagi in right side view:A – H. recurvatus Zettel, 2000; B – H. australis Zettel & Polhemus, 1998; C – H. nieserianus Zettel & Polhemus, 1998; D – H. incisus Zettel & Polhemus, 1998; E – H. vietnamensis Zettel, 2005. F–I – pronotal and genal plate: F – H. recurvatus; G – H. globulus Zettel, 2004; H – H. australis; I – H. incisus. J – right paramere of H. australis. K – left paramere of H. australis. L–O – medioventral carinae: L – H. globulus; M – H. australis; N – H. recurvatus; O – H. incisus. Abbreviations: pr – prosternum; ms – mesosternum, mt – metasternum, 2–6 – carinae of abdominal sterna 2–6. Figures not on scale. Redrawn (with slight modifications) from ZETTEL & POLHEMUS (1998): B–D, H–K, M, O; ZETTEL (2000a): A, F, N; ZETTEL (2004a): G, L; and ZETTEL (2005): E." figureDoi="http://doi.org/10.5281/zenodo.3699242" httpUri="https://zenodo.org/record/3699242/files/figure.png" pageId="2" pageNumber="245">Fig. 1M</figureCitation>
|
||
). Aedeagus as in
|
||
<figureCitation id="13752A413857FFBD137E8A4911ADC863" box="[592,679,1611,1634]" captionStart="Fig" captionStartId="1.[151,182,1930,1949]" captionTargetBox="[299,1290,638,1908]" captionTargetId="figure@1.[298,1290,635,1911]" captionTargetPageId="1" captionText="Fig. 1. Key characters of species belonging to the Helotrephes australis species-group.A–E – aedeagi in right side view:A – H. recurvatus Zettel, 2000; B – H. australis Zettel & Polhemus, 1998; C – H. nieserianus Zettel & Polhemus, 1998; D – H. incisus Zettel & Polhemus, 1998; E – H. vietnamensis Zettel, 2005. F–I – pronotal and genal plate: F – H. recurvatus; G – H. globulus Zettel, 2004; H – H. australis; I – H. incisus. J – right paramere of H. australis. K – left paramere of H. australis. L–O – medioventral carinae: L – H. globulus; M – H. australis; N – H. recurvatus; O – H. incisus. Abbreviations: pr – prosternum; ms – mesosternum, mt – metasternum, 2–6 – carinae of abdominal sterna 2–6. Figures not on scale. Redrawn (with slight modifications) from ZETTEL & POLHEMUS (1998): B–D, H–K, M, O; ZETTEL (2000a): A, F, N; ZETTEL (2004a): G, L; and ZETTEL (2005): E." figureDoi="http://doi.org/10.5281/zenodo.3699242" httpUri="https://zenodo.org/record/3699242/files/figure.png" pageId="2" pageNumber="245">Fig. 1B</figureCitation>
|
||
.
|
||
<collectingCountry id="F35976543857FFBD13968A4911F2C863" box="[696,760,1611,1634]" name="China" pageId="2" pageNumber="245">China</collectingCountry>
|
||
(
|
||
<collectingRegion id="498AF8263857FFBD11EA8A691211C883" box="[196,283,1643,1666]" country="China" name="Yunnan" pageId="2" pageNumber="245">Yunnan</collectingRegion>
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||
),
|
||
<collectingCountry id="F35976543857FFBD10038A691285C883" box="[301,399,1643,1666]" name="Thailand" pageId="2" pageNumber="245">Thailand</collectingCountry>
|
||
,
|
||
<collectingCountry id="F35976543857FFBD10B78A6912C5C883" box="[409,463,1643,1666]" name="Laos" pageId="2" pageNumber="245">Laos</collectingCountry>
|
||
, West
|
||
<collectingCountry id="F35976543857FFBD133B8A691170C883" box="[533,634,1643,1666]" name="Malaysia" pageId="2" pageNumber="245">Malaysia</collectingCountry>
|
||
. ................. .......................
|
||
<taxonomicName id="4C4E4D473857FFBD104A8A8E11FDC8A3" ID-CoL="6LMS7" authority="Zettel & Polhemus, 1998" authorityName="Zettel & Polhemus" authorityYear="1998" box="[356,759,1675,1699]" class="Insecta" family="Helotrephidae" genus="Helotrephes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="245" phylum="Arthropoda" rank="species" species="australis">
|
||
<emphasis id="B93AEAD63857FFBD104A8A8E12EFC8A3" bold="true" box="[356,485,1675,1699]" italics="true" pageId="2" pageNumber="245">H. australis</emphasis>
|
||
<bibRefCitation id="EFDF4B353857FFBD10C58A8911FDC8A3" author="Zettel & Polhemus" box="[491,759,1675,1698]" firstAuthor="Zettel" pageId="2" pageNumber="245" pagination="99 - 136" refId="ref4780" refString="ZETTEL H. & POLHEMUS J. T. 1998: A revision of the genus Helotrephes Stal, 1860 (Insecta: Heteroptera: Helotrephidae) with descriptions of twelve new taxa from the Oriental Realm. Annalen des Naturhistorischen Museums in Wien 100 B: 99 - 136." type="journal article" year="1998">Zettel & Polhemus, 1998</bibRefCitation>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BF136C43857FFBD11B98AA911FDC923" blockId="2.[151,760,971,2082]" pageId="2" pageNumber="245">
|
||
<keyLead id="30BF95F13857FFBD11B98AA911FDC923" pageId="2" pageNumber="245">
|
||
– Anterior projection of prosternal carina weakly acute, not pointed (cf.
|
||
<figureCitation id="13752A413857FFBD10588AC912C4C8E3" box="[374,462,1739,1762]" captionStart="Fig" captionStartId="1.[151,182,1930,1949]" captionTargetBox="[299,1290,638,1908]" captionTargetId="figure@1.[298,1290,635,1911]" captionTargetPageId="1" captionText="Fig. 1. Key characters of species belonging to the Helotrephes australis species-group.A–E – aedeagi in right side view:A – H. recurvatus Zettel, 2000; B – H. australis Zettel & Polhemus, 1998; C – H. nieserianus Zettel & Polhemus, 1998; D – H. incisus Zettel & Polhemus, 1998; E – H. vietnamensis Zettel, 2005. F–I – pronotal and genal plate: F – H. recurvatus; G – H. globulus Zettel, 2004; H – H. australis; I – H. incisus. J – right paramere of H. australis. K – left paramere of H. australis. L–O – medioventral carinae: L – H. globulus; M – H. australis; N – H. recurvatus; O – H. incisus. Abbreviations: pr – prosternum; ms – mesosternum, mt – metasternum, 2–6 – carinae of abdominal sterna 2–6. Figures not on scale. Redrawn (with slight modifications) from ZETTEL & POLHEMUS (1998): B–D, H–K, M, O; ZETTEL (2000a): A, F, N; ZETTEL (2004a): G, L; and ZETTEL (2005): E." figureDoi="http://doi.org/10.5281/zenodo.3699242" httpUri="https://zenodo.org/record/3699242/files/figure.png" pageId="2" pageNumber="245">Fig. 1O</figureCitation>
|
||
). Aedeagus as in
|
||
<figureCitation id="13752A413857FFBD13B28AC911FEC8E3" box="[668,756,1739,1762]" captionStart="Fig" captionStartId="1.[151,182,1930,1949]" captionTargetBox="[299,1290,638,1908]" captionTargetId="figure@1.[298,1290,635,1911]" captionTargetPageId="1" captionText="Fig. 1. Key characters of species belonging to the Helotrephes australis species-group.A–E – aedeagi in right side view:A – H. recurvatus Zettel, 2000; B – H. australis Zettel & Polhemus, 1998; C – H. nieserianus Zettel & Polhemus, 1998; D – H. incisus Zettel & Polhemus, 1998; E – H. vietnamensis Zettel, 2005. F–I – pronotal and genal plate: F – H. recurvatus; G – H. globulus Zettel, 2004; H – H. australis; I – H. incisus. J – right paramere of H. australis. K – left paramere of H. australis. L–O – medioventral carinae: L – H. globulus; M – H. australis; N – H. recurvatus; O – H. incisus. Abbreviations: pr – prosternum; ms – mesosternum, mt – metasternum, 2–6 – carinae of abdominal sterna 2–6. Figures not on scale. Redrawn (with slight modifications) from ZETTEL & POLHEMUS (1998): B–D, H–K, M, O; ZETTEL (2000a): A, F, N; ZETTEL (2004a): G, L; and ZETTEL (2005): E." figureDoi="http://doi.org/10.5281/zenodo.3699242" httpUri="https://zenodo.org/record/3699242/files/figure.png" pageId="2" pageNumber="245">Fig. 1C</figureCitation>
|
||
.
|
||
<collectingCountry id="F35976543857FFBD11938AE913F7C903" box="[189,253,1771,1794]" name="China" pageId="2" pageNumber="245">China</collectingCountry>
|
||
(
|
||
<collectingRegion id="498AF8263857FFBD10258AE91268C903" box="[267,354,1771,1794]" country="China" name="Yunnan" pageId="2" pageNumber="245">Yunnan</collectingRegion>
|
||
),
|
||
<collectingCountry id="F35976543857FFBD105D8AE912DFC903" box="[371,469,1771,1794]" name="Thailand" pageId="2" pageNumber="245">Thailand</collectingCountry>
|
||
,
|
||
<collectingCountry id="F35976543857FFBD10F18AE9111CC903" box="[479,534,1771,1794]" name="Laos" pageId="2" pageNumber="245">Laos</collectingCountry>
|
||
. ................................ ..................
|
||
<taxonomicName id="4C4E4D473857FFBD106D8B0E11FDC923" ID-CoL="3KJTS" authority="Zettel & Polhemus, 1998" authorityName="Zettel & Polhemus" authorityYear="1998" box="[323,759,1803,1827]" class="Insecta" family="Helotrephidae" genus="Helotrephes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="245" phylum="Arthropoda" rank="species" species="nieserianus">
|
||
<emphasis id="B93AEAD63857FFBD106D8B0E12EEC923" bold="true" box="[323,484,1803,1827]" italics="true" pageId="2" pageNumber="245">H. nieserianus</emphasis>
|
||
<bibRefCitation id="EFDF4B353857FFBD10C58B0911FDC923" author="Zettel & Polhemus" box="[491,759,1803,1826]" firstAuthor="Zettel" pageId="2" pageNumber="245" pagination="99 - 136" refId="ref4780" refString="ZETTEL H. & POLHEMUS J. T. 1998: A revision of the genus Helotrephes Stal, 1860 (Insecta: Heteroptera: Helotrephidae) with descriptions of twelve new taxa from the Oriental Realm. Annalen des Naturhistorischen Museums in Wien 100 B: 99 - 136." type="journal article" year="1998">Zettel & Polhemus, 1998</bibRefCitation>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30BA2E613857FFBD11B98B2911FDC623" pageId="2" pageNumber="245">
|
||
<paragraph id="8BF136C43857FFBD11B98B2911F2C9A3" blockId="2.[151,760,971,2082]" pageId="2" pageNumber="245">
|
||
<keyLead id="30BF95F13857FFBD11B98B2911F2C9A3" pageId="2" pageNumber="245">
|
||
5. Apex of aedeagus hook-shaped (
|
||
<figureCitation id="13752A413857FFBD13038B291182C943" box="[557,648,1835,1858]" captionStart="Fig" captionStartId="3.[151,182,1558,1577]" captionTargetBox="[255,1334,191,1536]" captionTargetId="figure@3.[254,1334,189,1538]" captionTargetPageId="3" captionText="Fig. 2. Helotrephes nainamalaii sp. nov. (holotype, brachypterous male).A – habitus, dorsal view; B – head, frontal view; C – pronotal and genal plate, ventral view (arrows indicate incision of pronotal plate and sub-basal tubercle of genal plate); D – ventromedian carinae, lateral view (arrows indicate strongly protruded prosternal carina and blunt posterior corner of metasternal carina); E – genital capsule, lateral view; F – apex of right paramere, dorsolateral view; G – aedeagus, lateral view; H–, right paramere, lateral view; I – left paramere, lateral view." figureDoi="http://doi.org/10.5281/zenodo.3699244" httpUri="https://zenodo.org/record/3699244/files/figure.png" pageId="2" pageNumber="245">Fig. 2G</figureCitation>
|
||
). Anterior corner of prosternal carina strongly protruding (
|
||
<figureCitation id="13752A413857FFBD13E28B4913EBC983" captionStart="Fig" captionStartId="3.[151,182,1558,1577]" captionTargetBox="[255,1334,191,1536]" captionTargetId="figure@3.[254,1334,189,1538]" captionTargetPageId="3" captionText="Fig. 2. Helotrephes nainamalaii sp. nov. (holotype, brachypterous male).A – habitus, dorsal view; B – head, frontal view; C – pronotal and genal plate, ventral view (arrows indicate incision of pronotal plate and sub-basal tubercle of genal plate); D – ventromedian carinae, lateral view (arrows indicate strongly protruded prosternal carina and blunt posterior corner of metasternal carina); E – genital capsule, lateral view; F – apex of right paramere, dorsolateral view; G – aedeagus, lateral view; H–, right paramere, lateral view; I – left paramere, lateral view." figureDoi="http://doi.org/10.5281/zenodo.3699244" httpUri="https://zenodo.org/record/3699244/files/figure.png" pageId="2" pageNumber="245">Fig. 2D</figureCitation>
|
||
). Metasternal carina with blunt apex (
|
||
<figureCitation id="13752A413857FFBD13BF8B6911E6C983" box="[657,748,1899,1922]" captionStart="Fig" captionStartId="3.[151,182,1558,1577]" captionTargetBox="[255,1334,191,1536]" captionTargetId="figure@3.[254,1334,189,1538]" captionTargetPageId="3" captionText="Fig. 2. Helotrephes nainamalaii sp. nov. (holotype, brachypterous male).A – habitus, dorsal view; B – head, frontal view; C – pronotal and genal plate, ventral view (arrows indicate incision of pronotal plate and sub-basal tubercle of genal plate); D – ventromedian carinae, lateral view (arrows indicate strongly protruded prosternal carina and blunt posterior corner of metasternal carina); E – genital capsule, lateral view; F – apex of right paramere, dorsolateral view; G – aedeagus, lateral view; H–, right paramere, lateral view; I – left paramere, lateral view." figureDoi="http://doi.org/10.5281/zenodo.3699244" httpUri="https://zenodo.org/record/3699244/files/figure.png" pageId="2" pageNumber="245">Fig. 2D</figureCitation>
|
||
).
|
||
<collectingCountry id="F35976543857FFBD11938B8913FEC9A3" box="[189,244,1931,1954]" name="India" pageId="2" pageNumber="245">India</collectingCountry>
|
||
(
|
||
<collectingRegion id="498AF8263857FFBD102C8B891275C9A3" box="[258,383,1931,1954]" country="India" name="Meghalaya" pageId="2" pageNumber="245">Meghalaya</collectingRegion>
|
||
). ..............
|
||
<emphasis id="B93AEAD63857FFBD10DA8B8E11F2C9A3" bold="true" box="[500,760,1931,1955]" pageId="2" pageNumber="245">
|
||
<taxonomicName id="4C4E4D473857FFBD10DA8B8E1191C9A3" authority="Jehamalar & Chandra & Zettel, 2018" authorityName="Jehamalar & Chandra & Zettel" authorityYear="2018" box="[500,667,1931,1955]" class="Insecta" family="Helotrephidae" genus="Helotrephes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="245" phylum="Arthropoda" rank="species" species="nainamalaii" status="sp. nov.">
|
||
<emphasis id="B93AEAD63857FFBD10DA8B8E1191C9A3" bold="true" box="[500,667,1931,1955]" italics="true" pageId="2" pageNumber="245">H. nainamalaii</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A20957AD3857FFBD138C8B8911F2C9A3" box="[674,760,1931,1954]" pageId="2" pageNumber="245" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BF136C43857FFBD11B98BA911FDC623" blockId="2.[151,760,971,2082]" pageId="2" pageNumber="245">
|
||
<keyLead id="30BF95F13857FFBD11B98BA911FDC623" pageId="2" pageNumber="245">
|
||
– Apex of aedeagus beak-shaped (
|
||
<figureCitation id="13752A413857FFBD133B8BA91187C9C3" box="[533,653,1963,1986]" captionStart="Fig" captionStartId="1.[151,182,1930,1949]" captionTargetBox="[299,1290,638,1908]" captionTargetId="figure@1.[298,1290,635,1911]" captionTargetPageId="1" captionText="Fig. 1. Key characters of species belonging to the Helotrephes australis species-group.A–E – aedeagi in right side view:A – H. recurvatus Zettel, 2000; B – H. australis Zettel & Polhemus, 1998; C – H. nieserianus Zettel & Polhemus, 1998; D – H. incisus Zettel & Polhemus, 1998; E – H. vietnamensis Zettel, 2005. F–I – pronotal and genal plate: F – H. recurvatus; G – H. globulus Zettel, 2004; H – H. australis; I – H. incisus. J – right paramere of H. australis. K – left paramere of H. australis. L–O – medioventral carinae: L – H. globulus; M – H. australis; N – H. recurvatus; O – H. incisus. Abbreviations: pr – prosternum; ms – mesosternum, mt – metasternum, 2–6 – carinae of abdominal sterna 2–6. Figures not on scale. Redrawn (with slight modifications) from ZETTEL & POLHEMUS (1998): B–D, H–K, M, O; ZETTEL (2000a): A, F, N; ZETTEL (2004a): G, L; and ZETTEL (2005): E." figureDoi="http://doi.org/10.5281/zenodo.3699242" httpUri="https://zenodo.org/record/3699242/files/figure.png" pageId="2" pageNumber="245">Figs 1D–E</figureCitation>
|
||
). Anterior corner of prosternal carina weakly protruding (
|
||
<figureCitation id="13752A413857FFBD13E28BC913D5C603" captionStart="Fig" captionStartId="1.[151,182,1930,1949]" captionTargetBox="[299,1290,638,1908]" captionTargetId="figure@1.[298,1290,635,1911]" captionTargetPageId="1" captionText="Fig. 1. Key characters of species belonging to the Helotrephes australis species-group.A–E – aedeagi in right side view:A – H. recurvatus Zettel, 2000; B – H. australis Zettel & Polhemus, 1998; C – H. nieserianus Zettel & Polhemus, 1998; D – H. incisus Zettel & Polhemus, 1998; E – H. vietnamensis Zettel, 2005. F–I – pronotal and genal plate: F – H. recurvatus; G – H. globulus Zettel, 2004; H – H. australis; I – H. incisus. J – right paramere of H. australis. K – left paramere of H. australis. L–O – medioventral carinae: L – H. globulus; M – H. australis; N – H. recurvatus; O – H. incisus. Abbreviations: pr – prosternum; ms – mesosternum, mt – metasternum, 2–6 – carinae of abdominal sterna 2–6. Figures not on scale. Redrawn (with slight modifications) from ZETTEL & POLHEMUS (1998): B–D, H–K, M, O; ZETTEL (2000a): A, F, N; ZETTEL (2004a): G, L; and ZETTEL (2005): E." figureDoi="http://doi.org/10.5281/zenodo.3699242" httpUri="https://zenodo.org/record/3699242/files/figure.png" pageId="2" pageNumber="245">Fig. 1O</figureCitation>
|
||
). Metasternal carina with pointed apex (
|
||
<figureCitation id="13752A413857FFBD13BB8BE911E0C603" box="[661,746,2027,2050]" captionStart="Fig" captionStartId="1.[151,182,1930,1949]" captionTargetBox="[299,1290,638,1908]" captionTargetId="figure@1.[298,1290,635,1911]" captionTargetPageId="1" captionText="Fig. 1. Key characters of species belonging to the Helotrephes australis species-group.A–E – aedeagi in right side view:A – H. recurvatus Zettel, 2000; B – H. australis Zettel & Polhemus, 1998; C – H. nieserianus Zettel & Polhemus, 1998; D – H. incisus Zettel & Polhemus, 1998; E – H. vietnamensis Zettel, 2005. F–I – pronotal and genal plate: F – H. recurvatus; G – H. globulus Zettel, 2004; H – H. australis; I – H. incisus. J – right paramere of H. australis. K – left paramere of H. australis. L–O – medioventral carinae: L – H. globulus; M – H. australis; N – H. recurvatus; O – H. incisus. Abbreviations: pr – prosternum; ms – mesosternum, mt – metasternum, 2–6 – carinae of abdominal sterna 2–6. Figures not on scale. Redrawn (with slight modifications) from ZETTEL & POLHEMUS (1998): B–D, H–K, M, O; ZETTEL (2000a): A, F, N; ZETTEL (2004a): G, L; and ZETTEL (2005): E." figureDoi="http://doi.org/10.5281/zenodo.3699242" httpUri="https://zenodo.org/record/3699242/files/figure.png" pageId="2" pageNumber="245">Fig. 1O</figureCitation>
|
||
). ................................................................................. 6
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30BA2E613857FFBD12128CBC1696CF94" pageId="2" pageNumber="245">
|
||
<paragraph id="8BF136C43857FFBD12128CBC1696CF34" blockId="2.[828,1437,190,406]" pageId="2" pageNumber="245">
|
||
<keyLead id="30BF95F13857FFBD12128CBC1696CF34" pageId="2" pageNumber="245">
|
||
6. Carina of sternum 2 with small apical incision (
|
||
<figureCitation id="13752A413857FFBD145F8CBC108ECEF4" captionStart="Fig" captionStartId="1.[151,182,1930,1949]" captionTargetBox="[299,1290,638,1908]" captionTargetId="figure@1.[298,1290,635,1911]" captionTargetPageId="1" captionText="Fig. 1. Key characters of species belonging to the Helotrephes australis species-group.A–E – aedeagi in right side view:A – H. recurvatus Zettel, 2000; B – H. australis Zettel & Polhemus, 1998; C – H. nieserianus Zettel & Polhemus, 1998; D – H. incisus Zettel & Polhemus, 1998; E – H. vietnamensis Zettel, 2005. F–I – pronotal and genal plate: F – H. recurvatus; G – H. globulus Zettel, 2004; H – H. australis; I – H. incisus. J – right paramere of H. australis. K – left paramere of H. australis. L–O – medioventral carinae: L – H. globulus; M – H. australis; N – H. recurvatus; O – H. incisus. Abbreviations: pr – prosternum; ms – mesosternum, mt – metasternum, 2–6 – carinae of abdominal sterna 2–6. Figures not on scale. Redrawn (with slight modifications) from ZETTEL & POLHEMUS (1998): B–D, H–K, M, O; ZETTEL (2000a): A, F, N; ZETTEL (2004a): G, L; and ZETTEL (2005): E." figureDoi="http://doi.org/10.5281/zenodo.3699242" httpUri="https://zenodo.org/record/3699242/files/figure.png" pageId="2" pageNumber="245">Fig. 1O</figureCitation>
|
||
). Carina of sternum 3 with long setae (
|
||
<figureCitation id="13752A413857FFBD141A8CDC1684CEF4" box="[1332,1422,222,245]" captionStart="Fig" captionStartId="1.[151,182,1930,1949]" captionTargetBox="[299,1290,638,1908]" captionTargetId="figure@1.[298,1290,635,1911]" captionTargetPageId="1" captionText="Fig. 1. Key characters of species belonging to the Helotrephes australis species-group.A–E – aedeagi in right side view:A – H. recurvatus Zettel, 2000; B – H. australis Zettel & Polhemus, 1998; C – H. nieserianus Zettel & Polhemus, 1998; D – H. incisus Zettel & Polhemus, 1998; E – H. vietnamensis Zettel, 2005. F–I – pronotal and genal plate: F – H. recurvatus; G – H. globulus Zettel, 2004; H – H. australis; I – H. incisus. J – right paramere of H. australis. K – left paramere of H. australis. L–O – medioventral carinae: L – H. globulus; M – H. australis; N – H. recurvatus; O – H. incisus. Abbreviations: pr – prosternum; ms – mesosternum, mt – metasternum, 2–6 – carinae of abdominal sterna 2–6. Figures not on scale. Redrawn (with slight modifications) from ZETTEL & POLHEMUS (1998): B–D, H–K, M, O; ZETTEL (2000a): A, F, N; ZETTEL (2004a): G, L; and ZETTEL (2005): E." figureDoi="http://doi.org/10.5281/zenodo.3699242" httpUri="https://zenodo.org/record/3699242/files/figure.png" pageId="2" pageNumber="245">Figs 1O</figureCitation>
|
||
).
|
||
<collectingCountry id="F35976543857FFBD124F8CFC10ABCF14" box="[865,929,254,277]" name="China" pageId="2" pageNumber="245">China</collectingCountry>
|
||
(
|
||
<collectingRegion id="498AF8263857FFBD12818CFC170CCF14" box="[943,1030,254,277]" country="China" name="Yunnan" pageId="2" pageNumber="245">Yunnan</collectingRegion>
|
||
),
|
||
<collectingCountry id="F35976543857FFBD15368CFC1770CF14" box="[1048,1146,254,277]" name="Thailand" pageId="2" pageNumber="245">Thailand</collectingCountry>
|
||
. ......................................... ..........................
|
||
<taxonomicName id="4C4E4D473857FFBD15348D1D1696CF34" ID-CoL="3KJTG" authority="Zettel & Polhemus, 1998" authorityName="Zettel & Polhemus" authorityYear="1998" box="[1050,1436,286,310]" class="Insecta" family="Helotrephidae" genus="Helotrephes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="245" phylum="Arthropoda" rank="species" species="incisus">
|
||
<emphasis id="B93AEAD63857FFBD15348D1D1783CF34" bold="true" box="[1050,1161,286,310]" italics="true" pageId="2" pageNumber="245">H. incisus</emphasis>
|
||
<bibRefCitation id="EFDF4B353857FFBD15BE8D1C1696CF34" author="Zettel & Polhemus" box="[1168,1436,286,309]" firstAuthor="Zettel" pageId="2" pageNumber="245" pagination="99 - 136" refId="ref4780" refString="ZETTEL H. & POLHEMUS J. T. 1998: A revision of the genus Helotrephes Stal, 1860 (Insecta: Heteroptera: Helotrephidae) with descriptions of twelve new taxa from the Oriental Realm. Annalen des Naturhistorischen Museums in Wien 100 B: 99 - 136." type="journal article" year="1998">Zettel & Polhemus, 1998</bibRefCitation>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BF136C43857FFBD12128D3C1696CF94" blockId="2.[828,1437,190,406]" pageId="2" pageNumber="245">
|
||
<keyLead id="30BF95F13857FFBD12128D3C1696CF94" pageId="2" pageNumber="245">
|
||
– Carina of sternum 2 without apical incision. Carina of sternum 3 with short setae (cf.
|
||
<figureCitation id="13752A413857FFBD15858D5C160ECF74" box="[1195,1284,350,373]" captionStart="Fig" captionStartId="1.[151,182,1930,1949]" captionTargetBox="[299,1290,638,1908]" captionTargetId="figure@1.[298,1290,635,1911]" captionTargetPageId="1" captionText="Fig. 1. Key characters of species belonging to the Helotrephes australis species-group.A–E – aedeagi in right side view:A – H. recurvatus Zettel, 2000; B – H. australis Zettel & Polhemus, 1998; C – H. nieserianus Zettel & Polhemus, 1998; D – H. incisus Zettel & Polhemus, 1998; E – H. vietnamensis Zettel, 2005. F–I – pronotal and genal plate: F – H. recurvatus; G – H. globulus Zettel, 2004; H – H. australis; I – H. incisus. J – right paramere of H. australis. K – left paramere of H. australis. L–O – medioventral carinae: L – H. globulus; M – H. australis; N – H. recurvatus; O – H. incisus. Abbreviations: pr – prosternum; ms – mesosternum, mt – metasternum, 2–6 – carinae of abdominal sterna 2–6. Figures not on scale. Redrawn (with slight modifications) from ZETTEL & POLHEMUS (1998): B–D, H–K, M, O; ZETTEL (2000a): A, F, N; ZETTEL (2004a): G, L; and ZETTEL (2005): E." figureDoi="http://doi.org/10.5281/zenodo.3699242" httpUri="https://zenodo.org/record/3699242/files/figure.png" pageId="2" pageNumber="245">Fig. 1M</figureCitation>
|
||
).
|
||
<collectingCountry id="F35976543857FFBD14398D5C167FCF74" box="[1303,1397,350,373]" name="Vietnam" pageId="2" pageNumber="245">Vietnam</collectingCountry>
|
||
. .... .....................................
|
||
<taxonomicName id="4C4E4D473857FFBD154A8D7D1696CF94" ID-CoL="6LN42" authority="Zettel, 2005" authorityName="Zettel" authorityYear="2005" box="[1124,1436,382,406]" class="Insecta" family="Helotrephidae" genus="Helotrephes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="245" phylum="Arthropoda" rank="species" species="vietnamensis">
|
||
<emphasis id="B93AEAD63857FFBD154A8D7D161ECF94" bold="true" box="[1124,1300,382,406]" italics="true" pageId="2" pageNumber="245">H. vietnamensis</emphasis>
|
||
<bibRefCitation id="EFDF4B353857FFBD14358D7C1696CF94" author="Zettel" box="[1307,1436,382,405]" firstAuthor="Zettel" pageId="2" pageNumber="245" pagination="67 - 80" refId="ref4665" refString="ZETTEL H. 2005: Notes on the Helotrephini (Insecta: Heteroptera: Helotrephidae) from Thailand and Vietnam, with descriptions of three new species. Annalen des Naturhistorischen Museums in Wien 106 B: 67 - 80." type="journal article" year="2005">Zettel, 2005</bibRefCitation>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |