314 lines
51 KiB
XML
314 lines
51 KiB
XML
<document id="87F17ED5FA68A46CE711C91DAFA4E832" ID-DOI="10.2478/aemnp-2018-0021" ID-GBIF-Dataset="c6c56876-6d8f-4c85-892a-37ea32bab386" ID-ISSN="1804-6487" ID-Zenodo-Dep="3699240" ID-ZooBank="66F5BF07-F271-4CE7-A40F-DFAFAF6076A2" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1583497418992" checkinUser="tatiana" docAuthor="Jehamalar, Eyarin, Chandra, Kailash & Zettel, Herbert" docDate="2018" docId="03E787D23857FFBB129B8DBC17D0CF74" docLanguage="en" docName="ActaEntMusNatPra.58.1.243-248.pdf.imf" docOrigin="Acta Entomologica Musei Nationalis Pragae 58 (1)" docStyle="DocumentStyle:BA2C5C8540539042E893A65A182CEA85.8:ActaEntMusNatPra.2018-.journal_article" docStyleId="BA2C5C8540539042E893A65A182CEA85" docStyleName="ActaEntMusNatPra.2018-.journal_article" docStyleVersion="8" docTitle="Helotrephes nainamalaii Jehamalar & Chandra & Zettel 2018, sp. nov." docType="treatment" docVersion="7" lastPageNumber="247" masterDocId="FFDEFFAA3855FFBF112E8C02130ACE01" masterDocTitle="New species and first record of Helotrephes from India, and a check-list of Indian Helotrephidae (Hemiptera: Heteroptera)" masterLastPageNumber="248" masterPageNumber="243" pageNumber="245" updateTime="1698725395183" updateUser="plazi" zenodo-license-document="CC-BY-NC-ND-3.0">
|
||
<mods:mods id="3506A4F500A15798F779893B28AD7901" xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo id="03FF4E599BB7EDF7C3ABDE3A5CF9558B">
|
||
<mods:title id="1602FCADF2BDC67F9E31D4E728F051CE">New species and first record of Helotrephes from India, and a check-list of Indian Helotrephidae (Hemiptera: Heteroptera)</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name id="95038D0FDEB9C98618A0CA7E4FC9B5E2" type="personal">
|
||
<mods:role id="DC05CB73B5F21749FA45E36D640F777B">
|
||
<mods:roleTerm id="30A9C0740D8F4B5A75B3CF7A573CB106">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="95006733EB8EE07768C6812B902BE6F7">Jehamalar, Eyarin</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="6BC5CB1C68307C5E99D1B45F01136C16" type="personal">
|
||
<mods:role id="0472E7C5AB756FB8B418DE5F373FBDE3">
|
||
<mods:roleTerm id="A7C70737884A82283CCF67EAC7D80257">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="68C4C54CB46B370BD92DE001CE54E8A4">Chandra, Kailash</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="D12D5103C13A138805602C6C6A660EBE" type="personal">
|
||
<mods:role id="7C0EFEEE86F512EFCB6D9B454C93B5B6">
|
||
<mods:roleTerm id="D3E2C215A916629220391F0069B8FF4F">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="754EAAF837CCB28E4C5AECBBDF232746">Zettel, Herbert</mods:namePart>
|
||
</mods:name>
|
||
<mods:typeOfResource id="2A931E8E37C9E5312D682AD7EA182B32">text</mods:typeOfResource>
|
||
<mods:relatedItem id="0F20116206E5AAF52BE25C1496DD1C0A" type="host">
|
||
<mods:titleInfo id="2DFA35322946A53C78E9B9B8749FE494">
|
||
<mods:title id="D8D7B1B6080FFE488669BD8C2CD8A869">Acta Entomologica Musei Nationalis Pragae</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part id="091BB3648FCE0112B364B2985604FF48">
|
||
<mods:date id="571ED5626A1F2001D58341FC3AC338DA">2018</mods:date>
|
||
<mods:detail id="77732E03C34705B1A023975997C5852F" type="pubDate">
|
||
<mods:number id="B8FD8F6629B5BF2CE869E6878416568E">2018-07-30</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="B220602F1AAFCB606D3ED2B0D0A49162" type="volume">
|
||
<mods:number id="26594A673C98DFAD191774F32D1A275D">58</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="30F9E0F01C997DCFAAFA872E49718B20" type="issue">
|
||
<mods:number id="ECEA462D829A0B86532985A1FCBE01F7">1</mods:number>
|
||
</mods:detail>
|
||
<mods:extent id="249ED0BF13C6F8C11758706B33303981" unit="page">
|
||
<mods:start id="10A2A57B091FA525C165342F0644AF55">243</mods:start>
|
||
<mods:end id="CE733FF834CD223DB2E1A754EB9068EE">248</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:classification id="0484ED566E2DA3B036FC77B457C5327A">journal article</mods:classification>
|
||
<mods:identifier id="1B9C4F2AC1D86B3BD3985AA38CB41B59" type="DOI">10.2478/aemnp-2018-0021</mods:identifier>
|
||
<mods:identifier id="90F916A37016425BA9804FAC8A832F63" type="GBIF-Dataset">c6c56876-6d8f-4c85-892a-37ea32bab386</mods:identifier>
|
||
<mods:identifier id="27E34A51D97B1FDD2B30CC7245547C3D" type="ISSN">1804-6487</mods:identifier>
|
||
<mods:identifier id="2C909335F6B06BC1508A367F52124A55" type="Zenodo-Dep">3699240</mods:identifier>
|
||
<mods:identifier id="8D953FEFDD917028F56C25FE362B1160" type="ZooBank">66F5BF07-F271-4CE7-A40F-DFAFAF6076A2</mods:identifier>
|
||
</mods:mods>
|
||
<treatment id="03E787D23857FFBB129B8DBC17D0CF74" ID-DOI="http://doi.org/10.5281/zenodo.3705450" ID-GBIF-Taxon="162462615" ID-Zenodo-Dep="3705450" LSID="urn:lsid:plazi:treatment:03E787D23857FFBB129B8DBC17D0CF74" httpUri="http://treatment.plazi.org/id/03E787D23857FFBB129B8DBC17D0CF74" lastPageId="4" lastPageNumber="247" pageId="2" pageNumber="245">
|
||
<subSubSection id="C354654F3857FFBD129B8DBC1629CFD4" box="[949,1315,446,469]" pageId="2" pageNumber="245" type="nomenclature">
|
||
<paragraph id="8BF136C43857FFBD129B8DBC1629CFD4" blockId="2.[828,1437,446,689]" box="[949,1315,446,469]" pageId="2" pageNumber="245">
|
||
<heading id="D0B981A83857FFBD129B8DBC1629CFD4" box="[949,1315,446,469]" centered="true" fontSize="10" level="2" pageId="2" pageNumber="245" reason="3">
|
||
<emphasis id="B93AEAD63857FFBD129B8DBC1629CFD4" bold="true" box="[949,1315,446,469]" pageId="2" pageNumber="245">
|
||
<taxonomicName id="4C4E4D473857FFBD129B8DBC17CCCFD4" authority="Jehamalar & Chandra & Zettel, 2018" authorityName="Jehamalar & Chandra & Zettel" authorityYear="2018" box="[949,1222,446,469]" class="Insecta" family="Helotrephidae" genus="Helotrephes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="245" phylum="Arthropoda" rank="species" species="nainamalaii" status="sp. nov.">
|
||
<emphasis id="B93AEAD63857FFBD129B8DBC17CCCFD4" bold="true" box="[949,1222,446,469]" italics="true" pageId="2" pageNumber="245">Helotrephes nainamalaii</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A20957AD3857FFBD15E38DBC1629CFD4" box="[1229,1315,446,469]" pageId="2" pageNumber="245" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C354654F3857FFBD15148DDE1794CFEE" box="[1082,1182,476,495]" pageId="2" pageNumber="245" type="description">
|
||
<paragraph id="8BF136C43857FFBD15148DDE1794CFEE" blockId="2.[828,1437,446,689]" box="[1082,1182,476,495]" pageId="2" pageNumber="245">
|
||
<heading id="D0B981A83857FFBD15148DDE1794CFEE" box="[1082,1182,476,495]" pageId="2" pageNumber="245">
|
||
(
|
||
<figureCitation id="13752A413857FFBD156E8DDE1792CFEE" box="[1088,1176,476,495]" captionStart="Fig" captionStartId="3.[151,182,1558,1577]" captionTargetBox="[255,1334,191,1536]" captionTargetId="figure@3.[254,1334,189,1538]" captionTargetPageId="3" captionText="Fig. 2. Helotrephes nainamalaii sp. nov. (holotype, brachypterous male).A – habitus, dorsal view; B – head, frontal view; C – pronotal and genal plate, ventral view (arrows indicate incision of pronotal plate and sub-basal tubercle of genal plate); D – ventromedian carinae, lateral view (arrows indicate strongly protruded prosternal carina and blunt posterior corner of metasternal carina); E – genital capsule, lateral view; F – apex of right paramere, dorsolateral view; G – aedeagus, lateral view; H–, right paramere, lateral view; I – left paramere, lateral view." figureDoi="http://doi.org/10.5281/zenodo.3699244" httpUri="https://zenodo.org/record/3699244/files/figure.png" pageId="2" pageNumber="245">Figs 2A–I</figureCitation>
|
||
)
|
||
</heading>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C354654F3857FFBD12128DFC167BCCB0" pageId="2" pageNumber="245" type="materials_examined">
|
||
<paragraph id="8BF136C43857FFBD12128DFC167BCCB0" blockId="2.[828,1437,446,689]" pageId="2" pageNumber="245">
|
||
<emphasis id="B93AEAD63857FFBD12128DFC10CBCC10" bold="true" box="[828,961,510,529]" pageId="2" pageNumber="245">Type material.</emphasis>
|
||
<materialsCitation id="3B263C993857FFBD12E88DFC166CCC46" ID-GBIF-Occurrence="2574906701" collectingDate="2016-03-04" collectionCode="PARA-, TYPE, CEL, ZSI" collectorName="Coll. E. E. Jehamalar. & Both & New Alipore & Kolkata & India & Reg. No." country="India" county="East Khasi Hills District" elevation="1042" latitude="25.25731" location="Phudjynniaw Stream" longLatPrecision="5" longitude="91.5745" municipality="Janiaw Village" pageId="2" pageNumber="245" specimenCount="3" stateProvince="Meghalaya" typeStatus="holotype">
|
||
<typeStatus id="54F588663857FFBD12E88DFC1710CC11" box="[966,1050,510,529]" pageId="2" pageNumber="245" type="holotype">
|
||
<subScript id="17CA34813857FFBD12E88DFC1710CC11" attach="left" box="[966,1050,510,529]" fontSize="6" pageId="2" pageNumber="245">HOLOTYPE</subScript>
|
||
</typeStatus>
|
||
(brachypterous male):
|
||
<emphasis id="B93AEAD63857FFBD15CC8DFC1696CC10" bold="true" box="[1250,1436,510,529]" pageId="2" pageNumber="245">
|
||
<collectingCountry id="F35976543857FFBD15CC8DFC162ECC10" box="[1250,1316,510,529]" name="India" pageId="2" pageNumber="245">INDIA</collectingCountry>
|
||
:
|
||
<collectingRegion id="498AF8263857FFBD14038DFC169FCC11" box="[1325,1429,510,529]" country="India" name="Meghalaya" pageId="2" pageNumber="245">
|
||
<subScript id="17CA34813857FFBD14038DFC169FCC11" attach="left" box="[1325,1429,510,529]" fontSize="6" pageId="2" pageNumber="245">MEGHALAYA</subScript>
|
||
</collectingRegion>
|
||
:
|
||
</emphasis>
|
||
<collectingCounty id="62904E483857FFBD12128E1B1704CC2D" box="[828,1038,537,556]" pageId="2" pageNumber="245">East Khasi Hills District</collectingCounty>
|
||
,
|
||
<collectingMunicipality id="6B95ACBE3857FFBD15388E1B179CCC2D" box="[1046,1174,537,556]" pageId="2" pageNumber="245">Janiaw Village</collectingMunicipality>
|
||
,
|
||
<location id="8E91601F3857FFBD15B18E1B165ECC2D" LSID="urn:lsid:plazi:treatment:03E787D23857FFBB129B8DBC17D0CF74:8E91601F3857FFBD15B18E1B165ECC2D" box="[1183,1364,537,556]" country="India" county="East Khasi Hills District" latitude="25.25731" longLatPrecision="5" longitude="91.5745" municipality="Janiaw Village" name="Phudjynniaw Stream" pageId="2" pageNumber="245" stateProvince="Meghalaya">Phudjynniaw Stream</location>
|
||
,
|
||
<elevation id="0063D1F73857FFBD14728E1B106FCC46" metricMagnitude="3" metricUnit="m" metricValue="1.042" pageId="2" pageNumber="245" unit="m" value="1042.0">
|
||
<quantity id="4CB69B213857FFBD14728E1B1697CC2D" box="[1372,1437,537,556]" metricMagnitude="3" metricUnit="m" metricValue="1.042" pageId="2" pageNumber="245" unit="m" value="1042.0">1042 m</quantity>
|
||
a.s.l.
|
||
</elevation>
|
||
,
|
||
<geoCoordinate id="EE7A50033857FFBD12408E3610D6CC46" box="[878,988,564,583]" degrees="25.25731" direction="north" orientation="latitude" pageId="2" pageNumber="245" precision="1" value="25.25731">25.25731° N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE7A50033857FFBD12CB8E36174DCC46" box="[997,1095,564,583]" degrees="91.5745" direction="east" orientation="longitude" pageId="2" pageNumber="245" precision="5" value="91.5745">91.5745° E</geoCoordinate>
|
||
,
|
||
<date id="FFF010043857FFBD157E8E3617AECC46" box="[1104,1188,564,583]" pageId="2" pageNumber="245" value="2016-03-04">
|
||
<collectingDate id="EFB4E9EC3857FFBD157E8E3617AECC46" box="[1104,1188,564,583]" pageId="2" pageNumber="245" value="2016-03-04">4.iii.2016</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="26BB53123857FFBD15828E36166CCC46" box="[1196,1382,564,583]" pageId="2" pageNumber="245">Coll. E. E. Jehamalar.</collectorName>
|
||
</materialsCitation>
|
||
<materialsCitation id="3B263C993857FFBD14448E36167BCCB0" ID-GBIF-Occurrence="2574906702" collectingDate="2016-03-04" collectionCode="PARA-, TYPE, CEL, ZSI" collectorName="Coll. E. E. Jehamalar. & Both & New Alipore & Kolkata & India & Reg. No." country="India" county="East Khasi Hills District" elevation="1042" latitude="25.25731" location="Phudjynniaw Stream" longLatPrecision="5" longitude="91.5745" municipality="Janiaw Village" pageId="2" pageNumber="245" specimenCount="3" stateProvince="Meghalaya" typeStatus="holotype">
|
||
<collectionCode id="ED5FAE013857FFBD14448E361696CC46" box="[1386,1436,564,583]" pageId="2" pageNumber="245">PARA-</collectionCode>
|
||
<subScript id="17CA34813857FFBD12128E51106BCC61" attach="none" box="[828,865,595,608]" fontSize="6" pageId="2" pageNumber="245">
|
||
<collectionCode id="ED5FAE013857FFBD12128E51106BCC61" box="[828,865,595,608]" pageId="2" pageNumber="245">TYPE</collectionCode>
|
||
</subScript>
|
||
: 1 brachypterous male, same collection data as for holotype.
|
||
<collectorName id="26BB53123857FFBD145D8E4C1696CC60" box="[1395,1436,590,609]" pageId="2" pageNumber="245">Both</collectorName>
|
||
specimens are deposited in the
|
||
<location id="8E91601F3857FFBD15128E6B1657CC7D" LSID="urn:lsid:plazi:treatment:03E787D23857FFBB129B8DBC17D0CF74:8E91601F3857FFBD15128E6B1657CC7D" box="[1084,1373,617,636]" country="India" county="East Khasi Hills District" latitude="25.25731" longLatPrecision="5" longitude="91.5745" municipality="Janiaw Village" name="Central Entomological Laboratory" pageId="2" pageNumber="245" stateProvince="Meghalaya">Central Entomological Laboratory</location>
|
||
(
|
||
<collectionCode id="ED5FAE013857FFBD14498E6B1699CC7D" box="[1383,1427,617,636]" country="USA" lsid="urn:lsid:biocol.org:col:14528" name="University of Illinois" pageId="2" pageNumber="245" type="Herbarium">CEL</collectionCode>
|
||
),
|
||
<location id="8E91601F3857FFBD12128E86172ECC96" LSID="urn:lsid:plazi:treatment:03E787D23857FFBB129B8DBC17D0CF74:8E91601F3857FFBD12128E86172ECC96" box="[828,1060,644,663]" country="India" county="East Khasi Hills District" latitude="25.25731" longLatPrecision="5" longitude="91.5745" municipality="Janiaw Village" name="Zoological Survey of India" pageId="2" pageNumber="245" stateProvince="Meghalaya">Zoological Survey of India</location>
|
||
(
|
||
<collectionCode id="ED5FAE013857FFBD151E8E86175ECC96" box="[1072,1108,644,663]" pageId="2" pageNumber="245">ZSI</collectionCode>
|
||
),
|
||
<collectorName id="26BB53123857FFBD154D8E8617D9CC96" box="[1123,1235,644,663]" pageId="2" pageNumber="245">New Alipore</collectorName>
|
||
,
|
||
<collectorName id="26BB53123857FFBD15F28E861628CC96" box="[1244,1314,644,663]" pageId="2" pageNumber="245">Kolkata</collectorName>
|
||
,
|
||
<collectingRegion id="498AF8263857FFBD14048E861693CC96" box="[1322,1433,644,663]" country="India" name="West Bengal" pageId="2" pageNumber="245">West Bengal</collectingRegion>
|
||
,
|
||
<collectorName id="26BB53123857FFBD12128E9C1061CCB0" box="[828,875,670,689]" pageId="2" pageNumber="245">India</collectorName>
|
||
:
|
||
<typeStatus id="54F588663857FFBD125D8E9C10B4CCB0" box="[883,958,670,689]" pageId="2" pageNumber="245" type="holotype">holotype</typeStatus>
|
||
<collectorName id="26BB53123857FFBD12ED8E9C171ACCB0" box="[963,1040,670,689]" pageId="2" pageNumber="245">Reg. No.</collectorName>
|
||
5887/H15,
|
||
<typeStatus id="54F588663857FFBD155B8E9C17B4CCB0" box="[1141,1214,670,689]" pageId="2" pageNumber="245" type="paratype">paratype</typeStatus>
|
||
<collectorName id="26BB53123857FFBD15ED8E9C161BCCB0" box="[1219,1297,670,689]" pageId="2" pageNumber="245">Reg. No.</collectorName>
|
||
5888/H15.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C354654F3857FFBD12128EC81633C980" pageId="2" pageNumber="245" type="description">
|
||
<paragraph id="8BF136C43857FFBD12128EC8170DCD40" blockId="2.[828,1437,714,2081]" pageId="2" pageNumber="245">
|
||
<emphasis id="B93AEAD63857FFBD12128EC817D8CCE0" bold="true" box="[828,1234,714,737]" pageId="2" pageNumber="245">Description of brachypterous male</emphasis>
|
||
(
|
||
<figureCitation id="13752A413857FFBD15CA8EC81653CCE0" box="[1252,1369,714,737]" captionStart="Fig" captionStartId="3.[151,182,1558,1577]" captionTargetBox="[255,1334,191,1536]" captionTargetId="figure@3.[254,1334,189,1538]" captionTargetPageId="3" captionText="Fig. 2. Helotrephes nainamalaii sp. nov. (holotype, brachypterous male).A – habitus, dorsal view; B – head, frontal view; C – pronotal and genal plate, ventral view (arrows indicate incision of pronotal plate and sub-basal tubercle of genal plate); D – ventromedian carinae, lateral view (arrows indicate strongly protruded prosternal carina and blunt posterior corner of metasternal carina); E – genital capsule, lateral view; F – apex of right paramere, dorsolateral view; G – aedeagus, lateral view; H–, right paramere, lateral view; I – left paramere, lateral view." figureDoi="http://doi.org/10.5281/zenodo.3699244" httpUri="https://zenodo.org/record/3699244/files/figure.png" pageId="2" pageNumber="245">Figs 2A–I</figureCitation>
|
||
). Size (
|
||
<typeStatus id="54F588663857FFBD126D8EE810A8CD00" box="[835,930,746,769]" pageId="2" pageNumber="245" type="holotype">holotype</typeStatus>
|
||
is a smaller specimen): Body length: 2.28 / 2.45. Maximum body width at posterolateral angle of cephalonotum: 1.71 / 1.81.
|
||
</paragraph>
|
||
<paragraph id="8BF136C43857FFBD12748F48173ACDC0" blockId="2.[828,1437,714,2081]" pageId="2" pageNumber="245">
|
||
<emphasis id="B93AEAD63857FFBD12748F4810A6CD60" bold="true" box="[858,940,842,865]" italics="true" pageId="2" pageNumber="245">Colour.</emphasis>
|
||
Dark brown to black. Anteromedian region of pronotum pale brown (
|
||
<figureCitation id="13752A413857FFBD15168F68179ACD80" box="[1080,1168,874,897]" captionStart="Fig" captionStartId="3.[151,182,1558,1577]" captionTargetBox="[255,1334,191,1536]" captionTargetId="figure@3.[254,1334,189,1538]" captionTargetPageId="3" captionText="Fig. 2. Helotrephes nainamalaii sp. nov. (holotype, brachypterous male).A – habitus, dorsal view; B – head, frontal view; C – pronotal and genal plate, ventral view (arrows indicate incision of pronotal plate and sub-basal tubercle of genal plate); D – ventromedian carinae, lateral view (arrows indicate strongly protruded prosternal carina and blunt posterior corner of metasternal carina); E – genital capsule, lateral view; F – apex of right paramere, dorsolateral view; G – aedeagus, lateral view; H–, right paramere, lateral view; I – left paramere, lateral view." figureDoi="http://doi.org/10.5281/zenodo.3699244" httpUri="https://zenodo.org/record/3699244/files/figure.png" pageId="2" pageNumber="245">Fig. 2A</figureCitation>
|
||
). Head yellowish brown with irregular black mark (
|
||
<figureCitation id="13752A413857FFBD15758F8817A5CDA0" box="[1115,1199,906,929]" captionStart="Fig" captionStartId="3.[151,182,1558,1577]" captionTargetBox="[255,1334,191,1536]" captionTargetId="figure@3.[254,1334,189,1538]" captionTargetPageId="3" captionText="Fig. 2. Helotrephes nainamalaii sp. nov. (holotype, brachypterous male).A – habitus, dorsal view; B – head, frontal view; C – pronotal and genal plate, ventral view (arrows indicate incision of pronotal plate and sub-basal tubercle of genal plate); D – ventromedian carinae, lateral view (arrows indicate strongly protruded prosternal carina and blunt posterior corner of metasternal carina); E – genital capsule, lateral view; F – apex of right paramere, dorsolateral view; G – aedeagus, lateral view; H–, right paramere, lateral view; I – left paramere, lateral view." figureDoi="http://doi.org/10.5281/zenodo.3699244" httpUri="https://zenodo.org/record/3699244/files/figure.png" pageId="2" pageNumber="245">Fig. 2B</figureCitation>
|
||
), near eyes yellowish. Legs yellowish brown.
|
||
</paragraph>
|
||
<paragraph id="8BF136C43857FFBD12748FC8161BCAA0" blockId="2.[828,1437,714,2081]" pageId="2" pageNumber="245">
|
||
<emphasis id="B93AEAD63857FFBD12748FC81752CDE0" bold="true" box="[858,1112,970,993]" italics="true" pageId="2" pageNumber="245">Structural characters.</emphasis>
|
||
Head: vertex near posterior margin of eye rugulose; anterior part without distinct punctures. Cephalonotal suture sinuate and distinct (
|
||
<figureCitation id="13752A413857FFBD145F88081054CA40" captionStart="Fig" captionStartId="3.[151,182,1558,1577]" captionTargetBox="[255,1334,191,1536]" captionTargetId="figure@3.[254,1334,189,1538]" captionTargetPageId="3" captionText="Fig. 2. Helotrephes nainamalaii sp. nov. (holotype, brachypterous male).A – habitus, dorsal view; B – head, frontal view; C – pronotal and genal plate, ventral view (arrows indicate incision of pronotal plate and sub-basal tubercle of genal plate); D – ventromedian carinae, lateral view (arrows indicate strongly protruded prosternal carina and blunt posterior corner of metasternal carina); E – genital capsule, lateral view; F – apex of right paramere, dorsolateral view; G – aedeagus, lateral view; H–, right paramere, lateral view; I – left paramere, lateral view." figureDoi="http://doi.org/10.5281/zenodo.3699244" httpUri="https://zenodo.org/record/3699244/files/figure.png" pageId="2" pageNumber="245">Fig. 2B</figureCitation>
|
||
). Eye length 0.59, width 0.28. Fourth rostral segment almost 3.0 times as long as third. Pronotal plate with a very large, almost squared incision (
|
||
<figureCitation id="13752A413857FFBD15928868161ACA80" box="[1212,1296,1130,1153]" captionStart="Fig" captionStartId="3.[151,182,1558,1577]" captionTargetBox="[255,1334,191,1536]" captionTargetId="figure@3.[254,1334,189,1538]" captionTargetPageId="3" captionText="Fig. 2. Helotrephes nainamalaii sp. nov. (holotype, brachypterous male).A – habitus, dorsal view; B – head, frontal view; C – pronotal and genal plate, ventral view (arrows indicate incision of pronotal plate and sub-basal tubercle of genal plate); D – ventromedian carinae, lateral view (arrows indicate strongly protruded prosternal carina and blunt posterior corner of metasternal carina); E – genital capsule, lateral view; F – apex of right paramere, dorsolateral view; G – aedeagus, lateral view; H–, right paramere, lateral view; I – left paramere, lateral view." figureDoi="http://doi.org/10.5281/zenodo.3699244" httpUri="https://zenodo.org/record/3699244/files/figure.png" pageId="2" pageNumber="245">Fig. 2C</figureCitation>
|
||
). Genal plate narrow, sub-basally with a small protrusion.
|
||
</paragraph>
|
||
<paragraph id="8BF136C43857FFBD127488A81725CB40" blockId="2.[828,1437,714,2081]" pageId="2" pageNumber="245">Punctures on entire dorsum bearing yellowish brown setae; pronotum with dense, small and shallow punctures; scutellum and hemelytra with sparse, large, deep punctures; corium sub-laterally reticulated. Mesoscutellum, length 1.09, basal width 0.94.</paragraph>
|
||
<paragraph id="8BF136C43857FFBD127489481760C860" blockId="2.[828,1437,714,2081]" pageId="2" pageNumber="245">
|
||
Ventral carinae (
|
||
<figureCitation id="13752A413857FFBD15398948177ECB60" box="[1047,1140,1354,1377]" captionStart="Fig" captionStartId="3.[151,182,1558,1577]" captionTargetBox="[255,1334,191,1536]" captionTargetId="figure@3.[254,1334,189,1538]" captionTargetPageId="3" captionText="Fig. 2. Helotrephes nainamalaii sp. nov. (holotype, brachypterous male).A – habitus, dorsal view; B – head, frontal view; C – pronotal and genal plate, ventral view (arrows indicate incision of pronotal plate and sub-basal tubercle of genal plate); D – ventromedian carinae, lateral view (arrows indicate strongly protruded prosternal carina and blunt posterior corner of metasternal carina); E – genital capsule, lateral view; F – apex of right paramere, dorsolateral view; G – aedeagus, lateral view; H–, right paramere, lateral view; I – left paramere, lateral view." figureDoi="http://doi.org/10.5281/zenodo.3699244" httpUri="https://zenodo.org/record/3699244/files/figure.png" pageId="2" pageNumber="245">Fig. 2D</figureCitation>
|
||
): Prosternal carina deeply notched posteriorly, strongly protruding anterior corner and blunt posterior corner. Mesosternal carina anteriorly knob-like, posteriorly rounded. Metasternal carina almost squared, anteriorly elevated, posteriorly with blunt corner. Carina of sternum 2 large and almost sigmoid. Carina of sternum 3 with apical part almost reniform, with sparse long thin setae. Carinae of abdominal sterna 4 to 6 small and basally with thick setae.
|
||
</paragraph>
|
||
<paragraph id="8BF136C43857FFBD12748A681633C980" blockId="2.[828,1437,714,2081]" pageId="2" pageNumber="245">
|
||
<emphasis id="B93AEAD63857FFBD12748A6810CCC880" bold="true" box="[858,966,1642,1665]" italics="true" pageId="2" pageNumber="245">Genitalia.</emphasis>
|
||
Genital capsule as in
|
||
<figureCitation id="13752A413857FFBD15888A6817FFC880" box="[1190,1269,1642,1665]" captionStart="Fig" captionStartId="3.[151,182,1558,1577]" captionTargetBox="[255,1334,191,1536]" captionTargetId="figure@3.[254,1334,189,1538]" captionTargetPageId="3" captionText="Fig. 2. Helotrephes nainamalaii sp. nov. (holotype, brachypterous male).A – habitus, dorsal view; B – head, frontal view; C – pronotal and genal plate, ventral view (arrows indicate incision of pronotal plate and sub-basal tubercle of genal plate); D – ventromedian carinae, lateral view (arrows indicate strongly protruded prosternal carina and blunt posterior corner of metasternal carina); E – genital capsule, lateral view; F – apex of right paramere, dorsolateral view; G – aedeagus, lateral view; H–, right paramere, lateral view; I – left paramere, lateral view." figureDoi="http://doi.org/10.5281/zenodo.3699244" httpUri="https://zenodo.org/record/3699244/files/figure.png" pageId="2" pageNumber="245">Fig. 2E</figureCitation>
|
||
. Aedeagus (
|
||
<figureCitation id="13752A413857FFBD145F8A681057C8A0" captionStart="Fig" captionStartId="3.[151,182,1558,1577]" captionTargetBox="[255,1334,191,1536]" captionTargetId="figure@3.[254,1334,189,1538]" captionTargetPageId="3" captionText="Fig. 2. Helotrephes nainamalaii sp. nov. (holotype, brachypterous male).A – habitus, dorsal view; B – head, frontal view; C – pronotal and genal plate, ventral view (arrows indicate incision of pronotal plate and sub-basal tubercle of genal plate); D – ventromedian carinae, lateral view (arrows indicate strongly protruded prosternal carina and blunt posterior corner of metasternal carina); E – genital capsule, lateral view; F – apex of right paramere, dorsolateral view; G – aedeagus, lateral view; H–, right paramere, lateral view; I – left paramere, lateral view." figureDoi="http://doi.org/10.5281/zenodo.3699244" httpUri="https://zenodo.org/record/3699244/files/figure.png" pageId="2" pageNumber="245">Fig. 2G</figureCitation>
|
||
) medium-sized, stout, with slightly convex posterior margin; apex of characteristic hook-like shape. Right paramere (
|
||
<figureCitation id="13752A413857FFBD129C8AC81700C8E0" box="[946,1034,1738,1761]" captionStart="Fig" captionStartId="3.[151,182,1558,1577]" captionTargetBox="[255,1334,191,1536]" captionTargetId="figure@3.[254,1334,189,1538]" captionTargetPageId="3" captionText="Fig. 2. Helotrephes nainamalaii sp. nov. (holotype, brachypterous male).A – habitus, dorsal view; B – head, frontal view; C – pronotal and genal plate, ventral view (arrows indicate incision of pronotal plate and sub-basal tubercle of genal plate); D – ventromedian carinae, lateral view (arrows indicate strongly protruded prosternal carina and blunt posterior corner of metasternal carina); E – genital capsule, lateral view; F – apex of right paramere, dorsolateral view; G – aedeagus, lateral view; H–, right paramere, lateral view; I – left paramere, lateral view." figureDoi="http://doi.org/10.5281/zenodo.3699244" httpUri="https://zenodo.org/record/3699244/files/figure.png" pageId="2" pageNumber="245">Fig. 2H</figureCitation>
|
||
) slightly shorter than left paramere, slender and curved, posteriorly with 13 long setae evenly distributed in distal two thirds; apex round, but in a more posterior view (
|
||
<figureCitation id="13752A413857FFBD12F38B281723C940" box="[989,1065,1834,1857]" captionStart="Fig" captionStartId="3.[151,182,1558,1577]" captionTargetBox="[255,1334,191,1536]" captionTargetId="figure@3.[254,1334,189,1538]" captionTargetPageId="3" captionText="Fig. 2. Helotrephes nainamalaii sp. nov. (holotype, brachypterous male).A – habitus, dorsal view; B – head, frontal view; C – pronotal and genal plate, ventral view (arrows indicate incision of pronotal plate and sub-basal tubercle of genal plate); D – ventromedian carinae, lateral view (arrows indicate strongly protruded prosternal carina and blunt posterior corner of metasternal carina); E – genital capsule, lateral view; F – apex of right paramere, dorsolateral view; G – aedeagus, lateral view; H–, right paramere, lateral view; I – left paramere, lateral view." figureDoi="http://doi.org/10.5281/zenodo.3699244" httpUri="https://zenodo.org/record/3699244/files/figure.png" pageId="2" pageNumber="245">Fig. 2F</figureCitation>
|
||
) angularly produced. Left paramere (
|
||
<figureCitation id="13752A413857FFBD126A8B481086C960" box="[836,908,1866,1889]" captionStart="Fig" captionStartId="3.[151,182,1558,1577]" captionTargetBox="[255,1334,191,1536]" captionTargetId="figure@3.[254,1334,189,1538]" captionTargetPageId="3" captionText="Fig. 2. Helotrephes nainamalaii sp. nov. (holotype, brachypterous male).A – habitus, dorsal view; B – head, frontal view; C – pronotal and genal plate, ventral view (arrows indicate incision of pronotal plate and sub-basal tubercle of genal plate); D – ventromedian carinae, lateral view (arrows indicate strongly protruded prosternal carina and blunt posterior corner of metasternal carina); E – genital capsule, lateral view; F – apex of right paramere, dorsolateral view; G – aedeagus, lateral view; H–, right paramere, lateral view; I – left paramere, lateral view." figureDoi="http://doi.org/10.5281/zenodo.3699244" httpUri="https://zenodo.org/record/3699244/files/figure.png" pageId="2" pageNumber="245">Fig. 2I</figureCitation>
|
||
) broad, abruptly narrowed distally, with blunt tip, with some setae posteriorly and on medial face.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C354654F3857FFBC12128B8810CBC911" lastPageId="3" lastPageNumber="246" pageId="2" pageNumber="245" type="diagnosis">
|
||
<paragraph id="8BF136C43857FFBC12128B8810CBC911" blockId="2.[828,1437,714,2081]" lastBlockId="3.[828,1437,1753,2096]" lastPageId="3" lastPageNumber="246" pageId="2" pageNumber="245">
|
||
<emphasis id="B93AEAD63857FFBD12128B88173CC9A0" bold="true" box="[828,1078,1930,1953]" pageId="2" pageNumber="245">Differential diagnosis.</emphasis>
|
||
<taxonomicName id="4C4E4D473857FFBD15128B88164EC9A0" authority="Jehamalar & Chandra & Zettel, 2018" authorityName="Jehamalar & Chandra & Zettel" authorityYear="2018" box="[1084,1348,1930,1953]" class="Insecta" family="Helotrephidae" genus="Helotrephes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="245" phylum="Arthropoda" rank="species" species="nainamalaii" status="sp. nov.">
|
||
<emphasis id="B93AEAD63857FFBD15128B88164EC9A0" box="[1084,1348,1930,1953]" italics="true" pageId="2" pageNumber="245">Helotrephes nainamalaii</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A20957AD3857FFBD14648B881697C9A0" box="[1354,1437,1930,1953]" pageId="2" pageNumber="245" rank="species">sp. nov.</taxonomicNameLabel>
|
||
belongs to the
|
||
<taxonomicName id="4C4E4D473857FFBD12FE8BA91743C9C0" authorityName="Zettel & Polhemus" authorityYear="1998" box="[976,1097,1962,1986]" class="Insecta" family="Helotrephidae" genus="Helotrephes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="245" phylum="Arthropoda" rank="species" species="australis">
|
||
<emphasis id="B93AEAD63857FFBD12FE8BA91743C9C0" box="[976,1097,1962,1986]" italics="true" pageId="2" pageNumber="245">H. australis</emphasis>
|
||
</taxonomicName>
|
||
group sensu
|
||
<bibRefCitation id="EFDF4B353857FFBD15E18BA8108CC9E0" author="ZETTEL & POLHEMUS" firstAuthor="ZETTEL" pageId="2" pageNumber="245" pagination="99 - 136" refId="ref4780" refString="ZETTEL H. & POLHEMUS J. T. 1998: A revision of the genus Helotrephes Stal, 1860 (Insecta: Heteroptera: Helotrephidae) with descriptions of twelve new taxa from the Oriental Realm. Annalen des Naturhistorischen Museums in Wien 100 B: 99 - 136." type="journal article" year="1998">ZETTEL & POLHEMUS (1998)</bibRefCitation>
|
||
. It can easily be distinguished from all the other known species of the
|
||
<taxonomicName id="4C4E4D473857FFBD15098BE917ACC600" authorityName="Zettel & Polhemus" authorityYear="1998" box="[1063,1190,2026,2050]" class="Insecta" family="Helotrephidae" genus="Helotrephes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="245" phylum="Arthropoda" rank="species" species="australis">
|
||
<emphasis id="B93AEAD63857FFBD15098BE91730C603" box="[1063,1082,2027,2050]" italics="true" pageId="2" pageNumber="245">H</emphasis>
|
||
.
|
||
<emphasis id="B93AEAD63857FFBD15678BE817ACC600" box="[1097,1190,2026,2049]" italics="true" pageId="2" pageNumber="245">australis</emphasis>
|
||
</taxonomicName>
|
||
group by the presence of a sub-basal tubercle on the genal plate (
|
||
<figureCitation id="13752A413857FFBD142B84081650C620" box="[1285,1370,2058,2081]" captionStart="Fig" captionStartId="3.[151,182,1558,1577]" captionTargetBox="[255,1334,191,1536]" captionTargetId="figure@3.[254,1334,189,1538]" captionTargetPageId="3" captionText="Fig. 2. Helotrephes nainamalaii sp. nov. (holotype, brachypterous male).A – habitus, dorsal view; B – head, frontal view; C – pronotal and genal plate, ventral view (arrows indicate incision of pronotal plate and sub-basal tubercle of genal plate); D – ventromedian carinae, lateral view (arrows indicate strongly protruded prosternal carina and blunt posterior corner of metasternal carina); E – genital capsule, lateral view; F – apex of right paramere, dorsolateral view; G – aedeagus, lateral view; H–, right paramere, lateral view; I – left paramere, lateral view." figureDoi="http://doi.org/10.5281/zenodo.3699244" httpUri="https://zenodo.org/record/3699244/files/figure.png" pageId="2" pageNumber="245">Fig. 2C</figureCitation>
|
||
), by a strongly protruding anterior corner of the prosternal carina (
|
||
<figureCitation id="13752A413856FFBC11B18AFB13FEC911" box="[159,244,1785,1808]" captionStart="Fig" captionStartId="3.[151,182,1558,1577]" captionTargetBox="[255,1334,191,1536]" captionTargetId="figure@3.[254,1334,189,1538]" captionTargetPageId="3" captionText="Fig. 2. Helotrephes nainamalaii sp. nov. (holotype, brachypterous male).A – habitus, dorsal view; B – head, frontal view; C – pronotal and genal plate, ventral view (arrows indicate incision of pronotal plate and sub-basal tubercle of genal plate); D – ventromedian carinae, lateral view (arrows indicate strongly protruded prosternal carina and blunt posterior corner of metasternal carina); E – genital capsule, lateral view; F – apex of right paramere, dorsolateral view; G – aedeagus, lateral view; H–, right paramere, lateral view; I – left paramere, lateral view." figureDoi="http://doi.org/10.5281/zenodo.3699244" httpUri="https://zenodo.org/record/3699244/files/figure.png" pageId="3" pageNumber="246">Fig. 2D</figureCitation>
|
||
), and by the apical hook of the aedeagus basally bent backward and then forward (
|
||
<figureCitation id="13752A413856FFBC132D8B1B1151C931" box="[515,603,1817,1840]" captionStart="Fig" captionStartId="3.[151,182,1558,1577]" captionTargetBox="[255,1334,191,1536]" captionTargetId="figure@3.[254,1334,189,1538]" captionTargetPageId="3" captionText="Fig. 2. Helotrephes nainamalaii sp. nov. (holotype, brachypterous male).A – habitus, dorsal view; B – head, frontal view; C – pronotal and genal plate, ventral view (arrows indicate incision of pronotal plate and sub-basal tubercle of genal plate); D – ventromedian carinae, lateral view (arrows indicate strongly protruded prosternal carina and blunt posterior corner of metasternal carina); E – genital capsule, lateral view; F – apex of right paramere, dorsolateral view; G – aedeagus, lateral view; H–, right paramere, lateral view; I – left paramere, lateral view." figureDoi="http://doi.org/10.5281/zenodo.3699244" httpUri="https://zenodo.org/record/3699244/files/figure.png" pageId="3" pageNumber="246">Fig. 2G</figureCitation>
|
||
). The first two characters should probably easily facilitate the recognition of the hitherto unknown female. Although the hook-shap- ed apex of the aedeagus resembles that of
|
||
<taxonomicName id="4C4E4D473856FFBC13718B7811D7C991" authorityName="Zettel & Polhemus" authorityYear="1998" box="[607,733,1913,1937]" class="Insecta" family="Helotrephidae" genus="Helotrephes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="246" phylum="Arthropoda" rank="species" species="australis">
|
||
<emphasis id="B93AEAD63856FFBC13718B7811D7C991" box="[607,733,1913,1937]" italics="true" pageId="3" pageNumber="246">H. australis</emphasis>
|
||
</taxonomicName>
|
||
, a common and widespread species in Southeast Asia,
|
||
<taxonomicName id="4C4E4D473856FFBC13F08B98121DC9D1" authorityName="Jehamalar & Chandra & Zettel" authorityYear="2018" class="Insecta" family="Helotrephidae" genus="Helotrephes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="246" phylum="Arthropoda" rank="species" species="nainamalaii">
|
||
<emphasis id="B93AEAD63856FFBC13F08B98121DC9D1" italics="true" pageId="3" pageNumber="246">H. nainamalaii</emphasis>
|
||
</taxonomicName>
|
||
differs from
|
||
<taxonomicName id="4C4E4D473856FFBC10B08BB81110C9D1" authorityName="Zettel & Polhemus" authorityYear="1998" box="[414,538,1977,2001]" class="Insecta" family="Helotrephidae" genus="Helotrephes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="246" phylum="Arthropoda" rank="species" species="australis">
|
||
<emphasis id="B93AEAD63856FFBC10B08BB81110C9D1" box="[414,538,1977,2001]" italics="true" pageId="3" pageNumber="246">H. australis</emphasis>
|
||
</taxonomicName>
|
||
by the apical hook of the aedeagus being bent backward and then curved forward (
|
||
<figureCitation id="13752A413856FFBC11B18BFB13F8C611" box="[159,242,2041,2064]" captionStart="Fig" captionStartId="3.[151,182,1558,1577]" captionTargetBox="[255,1334,191,1536]" captionTargetId="figure@3.[254,1334,189,1538]" captionTargetPageId="3" captionText="Fig. 2. Helotrephes nainamalaii sp. nov. (holotype, brachypterous male).A – habitus, dorsal view; B – head, frontal view; C – pronotal and genal plate, ventral view (arrows indicate incision of pronotal plate and sub-basal tubercle of genal plate); D – ventromedian carinae, lateral view (arrows indicate strongly protruded prosternal carina and blunt posterior corner of metasternal carina); E – genital capsule, lateral view; F – apex of right paramere, dorsolateral view; G – aedeagus, lateral view; H–, right paramere, lateral view; I – left paramere, lateral view." figureDoi="http://doi.org/10.5281/zenodo.3699244" httpUri="https://zenodo.org/record/3699244/files/figure.png" pageId="3" pageNumber="246">Fig. 2G</figureCitation>
|
||
), while in
|
||
<taxonomicName id="4C4E4D473856FFBC10708BF812D3C611" authorityName="Zettel & Polhemus" authorityYear="1998" box="[350,473,2041,2065]" class="Insecta" family="Helotrephidae" genus="Helotrephes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="246" phylum="Arthropoda" rank="species" species="australis">
|
||
<emphasis id="B93AEAD63856FFBC10708BF812D3C611" box="[350,473,2041,2065]" italics="true" pageId="3" pageNumber="246">H. australis</emphasis>
|
||
</taxonomicName>
|
||
it is simply curved forward (
|
||
<figureCitation id="13752A413856FFBC11B1841B13FFC631" box="[159,245,2073,2096]" captionStart="Fig" captionStartId="1.[151,182,1930,1949]" captionTargetBox="[299,1290,638,1908]" captionTargetId="figure@1.[298,1290,635,1911]" captionTargetPageId="1" captionText="Fig. 1. Key characters of species belonging to the Helotrephes australis species-group.A–E – aedeagi in right side view:A – H. recurvatus Zettel, 2000; B – H. australis Zettel & Polhemus, 1998; C – H. nieserianus Zettel & Polhemus, 1998; D – H. incisus Zettel & Polhemus, 1998; E – H. vietnamensis Zettel, 2005. F–I – pronotal and genal plate: F – H. recurvatus; G – H. globulus Zettel, 2004; H – H. australis; I – H. incisus. J – right paramere of H. australis. K – left paramere of H. australis. L–O – medioventral carinae: L – H. globulus; M – H. australis; N – H. recurvatus; O – H. incisus. Abbreviations: pr – prosternum; ms – mesosternum, mt – metasternum, 2–6 – carinae of abdominal sterna 2–6. Figures not on scale. Redrawn (with slight modifications) from ZETTEL & POLHEMUS (1998): B–D, H–K, M, O; ZETTEL (2000a): A, F, N; ZETTEL (2004a): G, L; and ZETTEL (2005): E." figureDoi="http://doi.org/10.5281/zenodo.3699242" httpUri="https://zenodo.org/record/3699242/files/figure.png" pageId="3" pageNumber="246">Fig. 1B</figureCitation>
|
||
). Moreover in
|
||
<taxonomicName id="4C4E4D473856FFBC108E84181128C631" authorityName="Zettel & Polhemus" authorityYear="1998" box="[416,546,2073,2097]" class="Insecta" family="Helotrephidae" genus="Helotrephes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="246" phylum="Arthropoda" rank="species" species="australis">
|
||
<emphasis id="B93AEAD63856FFBC108E84181128C631" box="[416,546,2073,2097]" italics="true" pageId="3" pageNumber="246">H. australis</emphasis>
|
||
</taxonomicName>
|
||
the right paramere is very slender, and the left paramere has a curved apex (
|
||
<figureCitation id="13752A413856FFBC126D8AFB10BFC911" box="[835,949,1785,1808]" captionStart="Fig" captionStartId="1.[151,182,1930,1949]" captionTargetBox="[299,1290,638,1908]" captionTargetId="figure@1.[298,1290,635,1911]" captionTargetPageId="1" captionText="Fig. 1. Key characters of species belonging to the Helotrephes australis species-group.A–E – aedeagi in right side view:A – H. recurvatus Zettel, 2000; B – H. australis Zettel & Polhemus, 1998; C – H. nieserianus Zettel & Polhemus, 1998; D – H. incisus Zettel & Polhemus, 1998; E – H. vietnamensis Zettel, 2005. F–I – pronotal and genal plate: F – H. recurvatus; G – H. globulus Zettel, 2004; H – H. australis; I – H. incisus. J – right paramere of H. australis. K – left paramere of H. australis. L–O – medioventral carinae: L – H. globulus; M – H. australis; N – H. recurvatus; O – H. incisus. Abbreviations: pr – prosternum; ms – mesosternum, mt – metasternum, 2–6 – carinae of abdominal sterna 2–6. Figures not on scale. Redrawn (with slight modifications) from ZETTEL & POLHEMUS (1998): B–D, H–K, M, O; ZETTEL (2000a): A, F, N; ZETTEL (2004a): G, L; and ZETTEL (2005): E." figureDoi="http://doi.org/10.5281/zenodo.3699242" httpUri="https://zenodo.org/record/3699242/files/figure.png" pageId="3" pageNumber="246">Figs 1J–K</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF31664C3856FFBC11B98A14174AC878" ID-DOI="http://doi.org/10.5281/zenodo.3699244" ID-Zenodo-Dep="3699244" httpUri="https://zenodo.org/record/3699244/files/figure.png" pageId="3" pageNumber="246" startId="3.[151,182,1558,1577]" targetBox="[255,1334,191,1536]" targetPageId="3">
|
||
<paragraph id="8BF136C43856FFBC11B98A14174AC878" blockId="3.[151,1437,1558,1657]" pageId="3" pageNumber="246">
|
||
Fig. 2.
|
||
<taxonomicName id="4C4E4D473856FFBC11FD8A1412ADC828" authority="Jehamalar & Chandra & Zettel, 2018" authorityName="Jehamalar & Chandra & Zettel" authorityYear="2018" box="[211,423,1558,1577]" class="Insecta" family="Helotrephidae" genus="Helotrephes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="246" phylum="Arthropoda" rank="species" species="nainamalaii" status="sp. nov.">
|
||
<emphasis id="B93AEAD63856FFBC11FD8A1412ADC828" box="[211,423,1558,1577]" italics="true" pageId="3" pageNumber="246">Helotrephes nainamalaii</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A20957AD3856FFBC10828A1412E5C828" box="[428,495,1558,1577]" pageId="3" pageNumber="246" rank="species">sp. nov.</taxonomicNameLabel>
|
||
(holotype, brachypterous male).A – habitus, dorsal view; B – head, frontal view; C – pronotal and genal plate, ventral view (arrows indicate incision of pronotal plate and sub-basal tubercle of genal plate); D – ventromedian carinae, lateral view (arrows indicate strongly protruded prosternal carina and blunt posterior corner of metasternal carina); E – genital capsule, lateral view; F – apex of right paramere, dorsolateral view; G – aedeagus, lateral view; H–, right paramere, lateral view; I – left paramere, lateral view.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C354654F3856FFBC12128B1B10C4C991" pageId="3" pageNumber="246" type="etymology">
|
||
<paragraph id="8BF136C43856FFBC12128B1B10C4C991" blockId="3.[828,1437,1753,2096]" pageId="3" pageNumber="246">
|
||
<emphasis id="B93AEAD63856FFBC12128B1B10B6C931" bold="true" box="[828,956,1817,1840]" pageId="3" pageNumber="246">Etymology.</emphasis>
|
||
<taxonomicName id="4C4E4D473856FFBC12EA8B1B17C4C931" authorityName="Jehamalar & Chandra & Zettel" authorityYear="2018" box="[964,1230,1817,1840]" class="Insecta" family="Helotrephidae" genus="Helotrephes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="246" phylum="Arthropoda" rank="species" species="nainamalaii">
|
||
<emphasis id="B93AEAD63856FFBC12EA8B1B17C4C931" box="[964,1230,1817,1840]" italics="true" pageId="3" pageNumber="246">Helotrephes nainamalaii</emphasis>
|
||
</taxonomicName>
|
||
is named after Mr. R. Nainamalai, Divisional Forest Officer, Jaintia Hills,
|
||
<collectingRegion id="498AF8263856FFBC12128B5B10BFC971" box="[828,949,1881,1904]" country="India" name="Meghalaya" pageId="3" pageNumber="246">Meghalaya</collectingRegion>
|
||
, for arranging the field trip in the East Khasi Hills District.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C354654F3856FFBB12128B9B17D0CF74" lastPageId="4" lastPageNumber="247" pageId="3" pageNumber="246" type="distribution">
|
||
<paragraph id="8BF136C43856FFBB12128B9B11F2CA79" blockId="3.[828,1437,1753,2096]" lastBlockId="4.[151,760,961,1144]" lastPageId="4" lastPageNumber="247" pageId="3" pageNumber="246">
|
||
<emphasis id="B93AEAD63856FFBC12128B9B1769C9B1" bold="true" box="[828,1123,1945,1968]" pageId="3" pageNumber="246">
|
||
<typeStatus id="54F588663856FFBC12128B9B107EC9B1" box="[828,884,1945,1968]" pageId="3" pageNumber="246">Type</typeStatus>
|
||
locality and habitat
|
||
</emphasis>
|
||
(
|
||
<figureCitation id="13752A413856FFBC155B8B9B17F9C9B1" box="[1141,1267,1945,1968]" captionStart="Figs 3" captionStartId="4.[151,188,817,836]" captionTargetBox="[151,760,189,807]" captionTargetId="figure@4.[151,760,187,807]" captionTargetPageId="4" captionText="Figs 3A–C. Type locality of Helotrephes nainamalaii sp. nov. A – map of India; B – map of Meghalaya; triangle denotes the distribution of the new species; C – photograph of the type locality, Phudjynniaw Stream in Janiaw Village." figureDoi="http://doi.org/10.5281/zenodo.3699246" httpUri="https://zenodo.org/record/3699246/files/figure.png" pageId="3" pageNumber="246">Figs 3A–C</figureCitation>
|
||
): Meghalaya is located between
|
||
<geoCoordinate id="EE7A50033856FFBC12C68BBB1735C9D1" box="[1000,1087,1977,2000]" degrees="25" direction="north" minutes="01" orientation="latitude" pageId="3" pageNumber="246" precision="925" value="25.016666">25°01′N</geoCoordinate>
|
||
and
|
||
<geoCoordinate id="EE7A50033856FFBC15408BBB17CFC9D1" box="[1134,1221,1977,2000]" degrees="26" direction="north" minutes="05" orientation="latitude" pageId="3" pageNumber="246" precision="925" value="26.083334">26°05′N</geoCoordinate>
|
||
latitude and
|
||
<geoCoordinate id="EE7A50033856FFBC14668BBB1696C9D1" box="[1352,1436,1977,2000]" degrees="85" direction="east" minutes="49" orientation="longitude" pageId="3" pageNumber="246" precision="925" value="85.816666">85°49′E</geoCoordinate>
|
||
and
|
||
<geoCoordinate id="EE7A50033856FFBC12458BDB10CAC9F1" box="[875,960,2009,2032]" degrees="92" direction="east" minutes="52" orientation="longitude" pageId="3" pageNumber="246" precision="925" value="92.86667">92°52′E</geoCoordinate>
|
||
longitude. The East Khasi Hills district experiences an average annual rainfall of about
|
||
<quantity id="4CB69B213856FFBC140C8BFB1696C611" box="[1314,1436,2041,2064]" metricMagnitude="1" metricUnit="m" metricValue="1.2" pageId="3" pageNumber="246" unit="mm" value="12000.0">12,000 mm</quantity>
|
||
and is considered as one of the wettest regions on earth. About 39% of the district’s total area is forested (
|
||
<quantity id="4CB69B213851FFBB13B18FC313B3CDF9" metricMagnitude="6" metricUnit="m" metricValue="1.06752" pageId="4" pageNumber="247" unit="km" value="1067.52">1067.52 km</quantity>
|
||
<superScript id="7C3B9B8C3851FFBB11978FDD13CBCDEC" attach="left" box="[185,193,991,1005]" fontSize="6" pageId="4" pageNumber="247">2</superScript>
|
||
out of
|
||
<quantity id="4CB69B213851FFBB103E8FE31265CDF9" box="[272,367,993,1016]" metricMagnitude="6" metricUnit="m" metricValue="2.748" pageId="4" pageNumber="247" unit="km" value="2748.0">2748 km</quantity>
|
||
<superScript id="7C3B9B8C3851FFBB10418FDD127DCDEC" attach="left" box="[367,375,991,1005]" fontSize="6" pageId="4" pageNumber="247">2</superScript>
|
||
). The district is dissected by plenty of streams; the streams and rivers in the north flow toward the Brahmaputra River and those in the south toward the Surma River of
|
||
<collectingCountry id="F35976543851FFBB1011884312B5CA59" box="[319,447,1089,1112]" name="Bangladesh" pageId="4" pageNumber="247">Bangladesh</collectingCountry>
|
||
. Phudjynniaw Stream located in East Khasi Hills District, is the tributary of Umngi River
|
||
<subSubSection id="C354654F3851FFBB13DA886311F2CA79" box="[756,760,1121,1144]" pageId="4" pageNumber="247">,</subSubSection>
|
||
</paragraph>
|
||
<paragraph id="8BF136C43851FFBB12128CBC1758CF14" blockId="4.[828,1437,190,373]" pageId="4" pageNumber="247">
|
||
which flows into
|
||
<collectingCountry id="F35976543851FFBB12C08CBC1760CED4" box="[1006,1130,190,213]" name="Bangladesh" pageId="4" pageNumber="247">Bangladesh</collectingCountry>
|
||
and meets with Surma River. It is a limestone-bedded slow flowing shallow stream with an open canopy (
|
||
<figureCitation id="13752A413851FFBB12DC8CFC174CCF14" box="[1010,1094,254,277]" captionStart="Figs 3" captionStartId="4.[151,188,817,836]" captionTargetBox="[151,760,189,807]" captionTargetId="figure@4.[151,760,187,807]" captionTargetPageId="4" captionText="Figs 3A–C. Type locality of Helotrephes nainamalaii sp. nov. A – map of India; B – map of Meghalaya; triangle denotes the distribution of the new species; C – photograph of the type locality, Phudjynniaw Stream in Janiaw Village." figureDoi="http://doi.org/10.5281/zenodo.3699246" httpUri="https://zenodo.org/record/3699246/files/figure.png" pageId="4" pageNumber="247">Fig. 3C</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BF136C43851FFBB12128D1C17D0CF74" blockId="4.[828,1437,190,373]" pageId="4" pageNumber="247">
|
||
<emphasis id="B93AEAD63851FFBB12128D1C10C0CF34" bold="true" box="[828,970,286,309]" pageId="4" pageNumber="247">Distribution.</emphasis>
|
||
Known only from the
|
||
<typeStatus id="54F588663851FFBB159C8D1C17D5CF34" box="[1202,1247,286,309]" pageId="4" pageNumber="247">type</typeStatus>
|
||
locality, Phudjynniaw Stream in Janiaw Village, Mawsynram Block, East Khasi Hills District,
|
||
<collectingRegion id="498AF8263851FFBB15368D5C179BCF74" box="[1048,1169,350,373]" country="India" name="Meghalaya" pageId="4" pageNumber="247">Meghalaya</collectingRegion>
|
||
,
|
||
<collectingCountry id="F35976543851FFBB15B28D5C17DCCF74" box="[1180,1238,350,373]" name="India" pageId="4" pageNumber="247">India</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |