treatments-xml/data/03/E7/87/03E78795E72DFFA77DE8FF51FD57ED9E.xml

<document id="76648D23777C31C053D28654249ADFA7" ID-CLB-Dataset="45852" ID-DOI="10.5281/zenodo.208130" ID-GBIF-Dataset="990f7a3b-602c-47eb-9717-947c8f1bcb7c" ID-ISSN="1175-5326" ID-Zenodo-Dep="208130" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460416101720" checkinUser="plazi" docAuthor="Breitman, Maria Florencia, Parra, Micaela, Pérez, Cristian Hernán Fulvio &amp; Sites, Jack Walter" docDate="2011" docId="03E78795E72DFFA77DE8FF51FD57ED9E" docLanguage="en" docName="zt03120p028.pdf" docOrigin="Zootaxa 3120" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Liolaemus avilae Breitman, Parra, Pérez &amp; Sites, 2011, sp. nov." docType="treatment" docVersion="10" lastPageNumber="21" masterDocId="FFDEFFEDE722FFB37D7FFFC6FFF0E976" masterDocTitle="Two new species of lizards from the Liolaemus lineomaculatus section (Squamata: Iguania: Liolaemidae) from southern Patagonia" masterLastPageNumber="28" masterPageNumber="1" pageNumber="16" updateTime="1698328881709" updateUser="ExternalLinkService">
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<mods:title id="892668D53B8B6A2458C73F7C115A3872">Two new species of lizards from the Liolaemus lineomaculatus section (Squamata: Iguania: Liolaemidae) from southern Patagonia</mods:title>
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<mods:namePart id="11054FF36EF4BA4688101ACC0BB7D640">Breitman, Maria Florencia</mods:namePart>
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<mods:namePart id="30DEB738DAEC66C40DA74A9D755ECB5B">Parra, Micaela</mods:namePart>
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<mods:namePart id="D0648C3852DB3B26E9EC8E6F7A19F6D7">Pérez, Cristian Hernán Fulvio</mods:namePart>
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<mods:namePart id="D4DB4821EA46AF3EF86FD4D28FCE7942">Sites, Jack Walter</mods:namePart>
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<mods:date id="533D0EF5EDAF473F0335EA4CF2C99257">2011</mods:date>
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<treatment id="03E78795E72DFFA77DE8FF51FD57ED9E" ID-DOI="http://doi.org/10.5281/zenodo.5615827" ID-GBIF-Taxon="119425262" ID-Zenodo-Dep="5615827" LSID="urn:lsid:plazi:treatment:03E78795E72DFFA77DE8FF51FD57ED9E" httpUri="http://treatment.plazi.org/id/03E78795E72DFFA77DE8FF51FD57ED9E" lastPageId="20" lastPageNumber="21" pageId="15" pageNumber="16">
<subSubSection id="C3546508E72DFFBC7DE8FF51FC0CE802" pageId="15" pageNumber="16" type="nomenclature">
<paragraph id="8BF13683E72DFFBC7DE8FF51FE39E9C7" blockId="15.[151,457,151,210]" box="[151,457,151,177]" pageId="15" pageNumber="16">
<heading id="D0B981EFE72DFFBC7DE8FF51FE39E9C7" bold="true" box="[151,457,151,177]" fontSize="11" level="1" pageId="15" pageNumber="16" reason="1">
<emphasis id="B93AEA91E72DFFBC7DE8FF51FE39E9C7" bold="true" box="[151,457,151,177]" pageId="15" pageNumber="16">
<taxonomicName id="4C4E4D00E72DFFBC7DE8FF51FE95E9C7" box="[151,357,151,177]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="avilae" status="sp. nov.">
<emphasis id="B93AEA91E72DFFBC7DE8FF51FE95E9C7" bold="true" box="[151,357,151,177]" italics="true" pageId="15" pageNumber="16">Liolaemus avilae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20957EAE72DFFBC7C13FF51FE39E9C7" box="[364,457,151,177]" pageId="15" pageNumber="16" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BF13683E72DFFBC7DE8FF7CFEDFE9A4" blockId="15.[151,457,151,210]" box="[151,303,186,210]" pageId="15" pageNumber="16">
(
<figureCitation id="13752A06E72DFFBC7DE1FF7CFEFBE9A4" box="[158,267,186,210]" captionStart="FIGURE 6" captionStartId="12.[151,250,937,960]" captionTargetBox="[151,1436,209,931]" captionTargetId="figure@12.[151,1436,209,933]" captionTargetPageId="12" captionText="FIGURE 6. Morphological comparison between Liolaemus morandae sp. nov. (A), L. lineomaculatus (B) and L. avilae sp. nov. (C). Notice differences in color pattern on the tails, size of blotches on the limbs and differences in vertebral and paravertebral lines. Dorsal scales (shown on the right) are less mucronated in L. lineomaculatus. Dorsal scales of hindlimbs are less carinated in L. morandae sp. nov. than in the other species, while they are more mucronated in L. avilae sp. nov." httpUri="https://zenodo.org/record/208136/files/figure.png" pageId="15" pageNumber="16">Figures 6</figureCitation>
, 
<figureCitation id="13752A06E72DFFBC7C68FF7CFED6E9A4" box="[279,294,186,210]" captionStart="FIGURE 7" captionStartId="18.[151,250,1929,1952]" captionTargetBox="[198,1398,305,1899]" captionTargetId="figure@18.[194,1398,301,1900]" captionTargetPageId="18" captionText="FIGURE 7. Upper, dorsal and ventral views of holotype in life of L. avilae sp. nov. MLP. S 2627; below, dorsal and ventral views of female (LJAMM-CNP 9399)." httpUri="https://zenodo.org/record/208137/files/figure.png" pageId="15" pageNumber="16">7</figureCitation>
)
</paragraph>
<paragraph id="8BF13683E72DFFBC7DE8FEC5FC8AE841" blockId="15.[151,1182,258,373]" pageId="15" pageNumber="16">
<bibRef id="52EEA275E72DFFBC7DE8FEC5FC8AE841" pageId="15" pageNumber="16" type="journal article">
<title id="F94ADDB5E72DFFBC7DE8FEC5FE13E86E" box="[151,483,258,280]" jopScore="0.3613745" pageId="15" pageNumber="16" type="_" typeScore="1">
<emphasis id="B93AEA91E72DFFBC7DE8FEC5FF3AE86E" bold="true" box="[151,202,259,280]" pageId="15" pageNumber="16">1971</emphasis>
, 
<emphasis id="B93AEA91E72DFFBC7DA9FEC4FE13E86E" box="[214,483,258,280]" italics="true" pageId="15" pageNumber="16">Liolaemus lineomaculatus</emphasis>
</title>
, 
<author id="53F3DE66E72DFFBC7C92FEC5FD5CE86E" box="[493,684,259,280]" fns="I" npo="LF" pageId="15" pageNumber="16">Donoso-Barros, R.</author>
and 
<author id="53F3DE66E72DFFBC7FA2FEC4FCCCE86E" box="[733,828,258,280]" fns="I" npo="LF" pageId="15" pageNumber="16">Cei, J.M.</author>
, 
<emphasis id="B93AEA91E72DFFBC7E39FEC4FBC2E86E" box="[838,1074,258,280]" italics="true" pageId="15" pageNumber="16">Journal of Herpetology</emphasis>
, 5, 89–95. 
<year id="FFE677D0E72DFFBC7DE8FEE7FF3AE840" box="[151,202,289,310]" pageId="15" pageNumber="16">
<emphasis id="B93AEA91E72DFFBC7DE8FEE7FF3AE840" bold="true" box="[151,202,289,310]" pageId="15" pageNumber="16">1975</emphasis>
</year>
, 
<journalOrPublisher id="518F3C4FE72DFFBC7DA9FEE7FD19E841" box="[214,745,289,311]" jopScore="5.0592427" pageId="15" pageNumber="16">
<emphasis id="B93AEA91E72DFFBC7DA9FEE7FE17E841" box="[214,487,289,311]" italics="true" pageId="15" pageNumber="16">Liolaemus lineomaculatus,</emphasis>
Cei, J.M., 
<emphasis id="B93AEA91E72DFFBC7F28FEE7FD19E841" box="[599,745,289,311]" italics="true" pageId="15" pageNumber="16">Herpetologica</emphasis>
</journalOrPublisher>
, 
<part id="FFEA9C1EE72DFFBC7F8AFEE4FCE0E841" box="[757,784,290,311]" pageId="15" pageNumber="16" type="volume">31</part>
, 
<pagination id="B287C877E72DFFBC7E63FEE7FCB2E841" box="[796,834,289,311]" pageId="15" pageNumber="16" type="pagination">109</pagination>
–116.
</bibRef>
</paragraph>
<paragraph id="8BF13683E72DFFBC7DE8FE86FC0CE802" blockId="15.[151,1182,258,373]" pageId="15" pageNumber="16">
<bibRef id="52EEA275E72DFFBC7DE8FE86FC0CE802" pageId="15" pageNumber="16" type="journal article">
<title id="F94ADDB5E72DFFBC7DE8FE86FE13E820" box="[151,483,320,342]" jopScore="0.24050441" pageId="15" pageNumber="16" type="_" typeScore="1">
<emphasis id="B93AEA91E72DFFBC7DE8FE86FF20E823" bold="true" box="[151,208,320,341]" pageId="15" pageNumber="16">1982,</emphasis>
<emphasis id="B93AEA91E72DFFBC7DA9FE86FE13E820" box="[214,483,320,342]" italics="true" pageId="15" pageNumber="16">Liolaemus lineomaculatus</emphasis>
</title>
, 
<author id="53F3DE66E72DFFBC7C92FE86FDBAE820" box="[493,586,320,342]" fns="I" npo="LF" pageId="15" pageNumber="16">Cei, J.M.</author>
and 
<author id="53F3DE66E72DFFBC7F04FE86FCF0E823" box="[635,768,320,342]" fns="I" npo="LF" pageId="15" pageNumber="16">Scolaro, J.A.</author>
, 
<emphasis id="B93AEA91E72DFFBC7E74FE86FC07E820" box="[779,1015,320,342]" italics="true" pageId="15" pageNumber="16">Journal of Herpetology</emphasis>
, 16, 354–363. 
<year id="FFE677D0E72DFFBC7DE8FE99FF3AE802" box="[151,202,351,372]" pageId="15" pageNumber="16">
<emphasis id="B93AEA91E72DFFBC7DE8FE99FF3AE802" bold="true" box="[151,202,351,372]" pageId="15" pageNumber="16">1992</emphasis>
</year>
, 
<journalOrPublisher id="518F3C4FE72DFFBC7DA9FE99FC99E803" box="[214,873,350,373]" jopScore="3.367062" pageId="15" pageNumber="16">
<emphasis id="B93AEA91E72DFFBC7DA9FE99FE17E803" box="[214,487,351,373]" italics="true" pageId="15" pageNumber="16">Liolaemus lineomaculatus,</emphasis>
Scolaro, J.A., 
<emphasis id="B93AEA91E72DFFBC7F03FE99FC99E803" box="[636,873,351,373]" italics="true" pageId="15" pageNumber="16">Acta zoologica lilloana</emphasis>
</journalOrPublisher>
, 
<part id="FFEA9C1EE72DFFBC7E0AFE99FC60E802" box="[885,912,351,372]" pageId="15" pageNumber="16" type="volume">41</part>
, 
<pagination id="B287C877E72DFFBC7EE4FE99FC32E802" box="[923,962,351,372]" pageId="15" pageNumber="16" type="pagination">287</pagination>
–293.
</bibRef>
</paragraph>
</subSubSection>
<subSubSection id="C3546508E72DFFBC7DE8FE67FEFFEBA9" pageId="15" pageNumber="16" type="materials_examined">
<paragraph id="8BF13683E72DFFBC7DE8FE67FBC5EB72" blockId="15.[151,1437,417,2031]" pageId="15" pageNumber="16">
<emphasis id="B93AEA91E72DFFBC7DE8FE67FEFBE8CF" bold="true" box="[151,267,417,441]" pageId="15" pageNumber="16">
<typeStatus id="54F58821E72DFFBC7DE8FE67FEF4E8CF" box="[151,260,417,441]" pageId="15" pageNumber="16" type="holotype">Holotype</typeStatus>
.
</emphasis>
<collectionCode id="ED5FAE46E72DFFBC7C6DFE67FEBEE8CF" box="[274,334,417,441]" httpUri="http://grbio.org/cool/66si-588n" name="Museo de La Plata" pageId="15" pageNumber="16">MLP</collectionCode>
.S 2627 (
<figureCitation id="13752A06E72DFFBC7CD1FE67FDE1E8CF" box="[430,529,417,441]" captionStart="FIGURE 7" captionStartId="18.[151,250,1929,1952]" captionTargetBox="[198,1398,305,1899]" captionTargetId="figure@18.[194,1398,301,1900]" captionTargetPageId="18" captionText="FIGURE 7. Upper, dorsal and ventral views of holotype in life of L. avilae sp. nov. MLP. S 2627; below, dorsal and ventral views of female (LJAMM-CNP 9399)." httpUri="https://zenodo.org/record/208137/files/figure.png" pageId="15" pageNumber="16">Figure 7</figureCitation>
), an adult male from Lago Buenos Aires plateau, 18.7 SW Puesto Lebrun, Lago Buenos Aires department, Santa Cruz province, 
<collectingCountry id="F3597613E72DFFBC7FC9FE03FCD5E8AB" box="[694,805,453,477]" name="Argentina" pageId="15" pageNumber="16">Argentina</collectingCountry>
(
<geoCoordinate id="EE7A5044E72DFFBC7E4AFE03FC26E8AB" box="[821,982,453,477]" direction="south" orientation="latitude" pageId="15" pageNumber="16" precision="1" value="-47.09139">47º05’29.0” S</geoCoordinate>
, 
<geoCoordinate id="EE7A5044E72DFFBC7E9CFE03FB7DE8AB" box="[995,1165,453,477]" direction="west" orientation="longitude" pageId="15" pageNumber="16" precision="1" value="-71.020256">71º01’12.9” W</geoCoordinate>
, 
<quantity id="4CB69B66E72DFFBC79E7FE03FB1DE8AB" box="[1176,1261,453,477]" metricMagnitude="3" metricUnit="m" metricValue="1.154" pageId="15" pageNumber="16" unit="m" value="1154.0">1154 m</quantity>
) (
<figureCitation id="13752A06E72DFFBC787BFE03FA81E8AB" box="[1284,1393,453,477]" captionStart="FIGURE 5" captionStartId="10.[151,250,1053,1076]" captionTargetBox="[157,1427,198,1017]" captionTargetId="figure@10.[151,1436,193,1032]" captionTargetPageId="10" captionText="FIGURE 5. Distribution map for species of the lineomaculatus group, with geographical localities sampled in this study superimposed on regional elevation (shading). Black circles: L. morandae sp. nov.; white circle: L. avilae sp. nov.; black squares: L. lineomaculatus; white square: L. kolengh; white triangles: L. hatcheri; black triangle: L. silvanae. Arrows indicate type localities. CH: Chubut province; SC: Santa Cruz province." httpUri="https://zenodo.org/record/208135/files/figure.png" pageId="15" pageNumber="16">Figures 5</figureCitation>
, 
<figureCitation id="13752A06E72DFFBC7801FE03FA7DE8AB" box="[1406,1421,453,477]" captionStart="FIGURE 8" captionStartId="19.[151,250,1965,1988]" captionTargetBox="[212,1380,193,1945]" captionTargetId="figure@19.[212,1380,193,1945]" captionTargetPageId="19" captionText="FIGURE 8. Type locality of Liolaemus avilae sp. nov. Altoandina phytogeographic province, Altoandino Austral District (47 º 05 ’ 29.0 ” S, 71 º 01 ’ 12.9 ” W, 1154 m)." httpUri="https://zenodo.org/record/208138/files/figure.png" pageId="15" pageNumber="16">8</figureCitation>
), L.J. Avila, C.H.F. Pérez, M.F. Breitman and N. Feltrin collectors, 
<date id="FFF01043E72DFFBC7E09FE2AFBC2EB72" box="[886,1074,489,517]" pageId="15" pageNumber="16" value="2008-01-09">9th January 2008</date>
.
</paragraph>
<paragraph id="8BF13683E72DFFBC7DBAFDD6FEFFEBA9" blockId="15.[151,1437,417,2031]" pageId="15" pageNumber="16">
<typeStatus id="54F58821E72DFFBC7DBAFDD6FECDEB5E" box="[197,317,528,552]" pageId="15" pageNumber="16" type="paratype">
<emphasis id="B93AEA91E72DFFBC7DBAFDD6FECDEB5E" bold="true" box="[197,317,528,552]" pageId="15" pageNumber="16">Paratypes</emphasis>
</typeStatus>
. 
<collectionCode id="ED5FAE46E72DFFBC7C33FDD6FE1EEB5E" box="[332,494,528,552]" httpUri="http://grbio.org/cool/yjrq-91r7" name="Centro Nacional Patagonico (CENPAT-CONICET)" pageId="15" pageNumber="16">LJAMM-CNP</collectionCode>
9250, 9253, 9274, adult males, 
<collectionCode id="ED5FAE46E72DFFBC7E24FDD6FC0DEB5E" box="[859,1021,528,552]" httpUri="http://grbio.org/cool/yjrq-91r7" name="Centro Nacional Patagonico (CENPAT-CONICET)" pageId="15" pageNumber="16">LJAMM-CNP</collectionCode>
9276-9277, 9399, adult females and 
<collectionCode id="ED5FAE46E72DFFBC7DE8FDF2FEC9EB3A" box="[151,313,564,588]" httpUri="http://grbio.org/cool/yjrq-91r7" name="Centro Nacional Patagonico (CENPAT-CONICET)" pageId="15" pageNumber="16">LJAMM-CNP</collectionCode>
9251, 9252, juveniles; from same locality as 
<typeStatus id="54F58821E72DFFBC7E42FDF2FC52EB3A" box="[829,930,564,588]" pageId="15" pageNumber="16" type="holotype">holotype</typeStatus>
, L.J. Avila, C.H.F. Pérez, M.F. Breitman and N. Feltrin collectors, 
<date id="FFF01043E72DFFBC7CF9FD9EFDC4EB06" box="[390,564,600,625]" pageId="15" pageNumber="16" value="2008-01-09">9 January 2008</date>
. 
<collectionCode id="ED5FAE46E72DFFBC7F41FD9EFD10EB06" box="[574,736,600,624]" httpUri="http://grbio.org/cool/yjrq-91r7" name="Centro Nacional Patagonico (CENPAT-CONICET)" pageId="15" pageNumber="16">LJAMM-CNP</collectionCode>
9243, an adult male from Puesto Lebrun, 
<quantity id="4CB69B66E72DFFBC79C6FD9EFAE5EB06" box="[1209,1301,600,624]" metricMagnitude="4" metricUnit="m" metricValue="2.73" pageId="15" pageNumber="16" unit="km" value="27.3">27.3 km</quantity>
W Estancia La Vizcaina, Lago Buenos Aires Plateau, Lago Buenos Aires department, Santa Cruz province, 
<collectingCountry id="F3597613E72DFFBC7852FDBAFA6CEBE2" box="[1325,1436,636,660]" name="Argentina" pageId="15" pageNumber="16">Argentina</collectingCountry>
(
<geoCoordinate id="EE7A5044E72DFFBC7DE0FD65FECFEBCD" box="[159,319,675,699]" direction="south" orientation="latitude" pageId="15" pageNumber="16" precision="1" value="-46.96439">46º57’51.8” S</geoCoordinate>
, 
<geoCoordinate id="EE7A5044E72DFFBC7C33FD65FE05EBCD" box="[332,501,675,699]" direction="west" orientation="longitude" pageId="15" pageNumber="16" precision="1" value="-71.10755">71º06’27.2” W</geoCoordinate>
, 
<quantity id="4CB69B66E72DFFBC7F7FFD65FDA8EBCD" box="[512,600,675,699]" metricMagnitude="3" metricUnit="m" metricValue="1.353" pageId="15" pageNumber="16" unit="m" value="1353.0">1353 m</quantity>
), L.J. Avila, C.H.F. Pérez, M.F. Breitman and N. Feltrin collectors, 
<date id="FFF01043E72DFFBC7833FD65FEFCEBA9" pageId="15" pageNumber="16" value="2008-01-08">8th January 2008</date>
.
</paragraph>
</subSubSection>
<subSubSection id="C3546508E72DFFA37DBAFD2CFB21ECB6" lastPageId="16" lastPageNumber="17" pageId="15" pageNumber="16" type="diagnosis">
<paragraph id="8BF13683E72DFFBC7DBAFD2CFDCBEA19" blockId="15.[151,1437,417,2031]" pageId="15" pageNumber="16">
<emphasis id="B93AEA91E72DFFBC7DBAFD2CFECFEA74" bold="true" box="[197,319,746,771]" pageId="15" pageNumber="16">Diagnosis.</emphasis>
<taxonomicName id="4C4E4D00E72DFFBC7C37FD2DFDFBEA75" box="[328,523,747,771]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="avilae" status="sp. nov.">
<emphasis id="B93AEA91E72DFFBC7C37FD2DFDFBEA75" box="[328,523,747,771]" italics="true" pageId="15" pageNumber="16">Liolaemus avilae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20957EAE72DFFBC7F6BFD2DFD9FEA75" box="[532,623,747,771]" pageId="15" pageNumber="16" rank="species">
<emphasis id="B93AEA91E72DFFBC7F6BFD2DFD9FEA75" bold="true" box="[532,623,747,771]" pageId="15" pageNumber="16">sp. nov.</emphasis>
</taxonomicNameLabel>
, a member of the 
<taxonomicName id="4C4E4D00E72DFFBC7E39FD2AFBE1EA75" box="[838,1041,747,771]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="lineomaculatus">
<emphasis id="B93AEA91E72DFFBC7E39FD2AFBE1EA75" box="[838,1041,747,771]" italics="true" pageId="15" pageNumber="16">L. lineomaculatus</emphasis>
</taxonomicName>
section and specifically the 
<taxonomicName id="4C4E4D00E72DFFBC7824FD2DFEFAEA51" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="lineomaculatus">
<emphasis id="B93AEA91E72DFFBC7824FD2DFEFAEA51" italics="true" pageId="15" pageNumber="16">lineomaculatus</emphasis>
</taxonomicName>
group, has dorsal trifid scales but lacks of precloacal pores in either sex (Etheridge 1995); molecular evidence places this species in the 
<taxonomicName id="4C4E4D00E72DFFBC7F7AFCF5FD40EA3D" box="[517,688,819,843]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="lineomaculatus">
<emphasis id="B93AEA91E72DFFBC7F7AFCF5FD40EA3D" box="[517,688,819,843]" italics="true" pageId="15" pageNumber="16">lineomaculatus</emphasis>
</taxonomicName>
group, as the sister species to 
<taxonomicName id="4C4E4D00E72DFFBC796BFCF2FB2EEA3D" box="[1044,1246,819,843]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="lineomaculatus">
<emphasis id="B93AEA91E72DFFBC796BFCF2FB2EEA3D" box="[1044,1246,819,843]" italics="true" pageId="15" pageNumber="16">L. lineomaculatus</emphasis>
</taxonomicName>
(Breitman 
<emphasis id="B93AEA91E72DFFBC781DFCF2FA65EA3D" box="[1378,1429,819,843]" italics="true" pageId="15" pageNumber="16">et al</emphasis>
. 2011; see Tables 1 to 4 and 
<figureCitation id="13752A06E72DFFBC7CB2FC91FDDEEA19" box="[461,558,855,879]" captionStart="FIGURE 6" captionStartId="12.[151,250,937,960]" captionTargetBox="[151,1436,209,931]" captionTargetId="figure@12.[151,1436,209,933]" captionTargetPageId="12" captionText="FIGURE 6. Morphological comparison between Liolaemus morandae sp. nov. (A), L. lineomaculatus (B) and L. avilae sp. nov. (C). Notice differences in color pattern on the tails, size of blotches on the limbs and differences in vertebral and paravertebral lines. Dorsal scales (shown on the right) are less mucronated in L. lineomaculatus. Dorsal scales of hindlimbs are less carinated in L. morandae sp. nov. than in the other species, while they are more mucronated in L. avilae sp. nov." httpUri="https://zenodo.org/record/208136/files/figure.png" pageId="15" pageNumber="16">Figure 6</figureCitation>
).
</paragraph>
<paragraph id="8BF13683E72DFFBC7DBAFCBDFE54EDC5" blockId="15.[151,1437,417,2031]" pageId="15" pageNumber="16">
Relative to 
<taxonomicName id="4C4E4D00E72DFFBC7C38FCBAFE28EAE5" box="[327,472,891,915]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="morandae" status="sp. nov.">
<emphasis id="B93AEA91E72DFFBC7C38FCBAFE28EAE5" box="[327,472,891,915]" italics="true" pageId="15" pageNumber="16">L. morandae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20957EAE72DFFBC7CA0FCBDFDC9EAE5" box="[479,569,891,915]" pageId="15" pageNumber="16" rank="species">
<emphasis id="B93AEA91E72DFFBC7CA0FCBDFDC9EAE5" bold="true" box="[479,569,891,915]" pageId="15" pageNumber="16">sp. nov.</emphasis>
</taxonomicNameLabel>
, 
<taxonomicName id="4C4E4D00E72DFFBC7F37FCBAFD5BEAE5" box="[584,683,891,915]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="avilae" status="sp. nov.">
<emphasis id="B93AEA91E72DFFBC7F37FCBAFD5BEAE5" box="[584,683,891,915]" italics="true" pageId="15" pageNumber="16">L. avilae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20957EAE72DFFBC7FCCFCBDFCFDEAE5" box="[691,781,891,915]" pageId="15" pageNumber="16" rank="species">
<emphasis id="B93AEA91E72DFFBC7FCCFCBDFCFDEAE5" bold="true" box="[691,781,891,915]" pageId="15" pageNumber="16">sp. nov.</emphasis>
</taxonomicNameLabel>
has more scales in contact with the interparietal scale (7– 10, X = 8,13 vs. 6–7, X = 6.33; 
<emphasis id="B93AEA91E72DFFBC7F67FC66FDD6EAC1" box="[536,550,928,951]" italics="true" pageId="15" pageNumber="16">p</emphasis>
= 0.0027), fewer supralabial scales (5–6, X = 5.25 vs. 5–10, X = 7.17; 
<emphasis id="B93AEA91E72DFFBC780BFC66FA72EAC1" box="[1396,1410,928,951]" italics="true" pageId="15" pageNumber="16">p</emphasis>
= 0.0008), fewer midbody scales (53–59, X = 55.5 vs. 55–63, X = 59.83; 
<emphasis id="B93AEA91E72DFFBC7ECEFC02FC4FEAAD" box="[945,959,964,987]" italics="true" pageId="15" pageNumber="16">p</emphasis>
&lt;0.0001), more third finger lamellae (15– 18, X = 16.25 vs. 13–16, X = 15.33; 
<emphasis id="B93AEA91E72DFFBC7F4FFC2EFDCEEA89" box="[560,574,1000,1023]" italics="true" pageId="15" pageNumber="16">p</emphasis>
&lt;0.0001), a longer elbow-wrist distance (4.89–6.81, X = 5.76 vs. 4.69–5.56, X = 5.32; 
<emphasis id="B93AEA91E72DFFBC7C6CFBCAFED1ED55" box="[275,289,1036,1059]" italics="true" pageId="15" pageNumber="16">p</emphasis>
&lt;0.0001), and a shorter rostral-nasal distance (1.69–2.25, X = 1.95 vs. 2.02–2.42, X = 2.15; 
<emphasis id="B93AEA91E72DFFBC780BFBCAFA72ED55" box="[1396,1410,1036,1059]" italics="true" pageId="15" pageNumber="16">p</emphasis>
&lt;0.0001). 
<taxonomicName id="4C4E4D00E72DFFBC7D81FBE9FE30ED31" box="[254,448,1071,1095]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="avilae" status="sp. nov.">
<emphasis id="B93AEA91E72DFFBC7D81FBE9FE30ED31" box="[254,448,1071,1095]" italics="true" pageId="15" pageNumber="16">Liolaemus avilae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20957EAE72DFFBC7CB4FBE9FDD7ED31" box="[459,551,1071,1095]" pageId="15" pageNumber="16" rank="species">
<emphasis id="B93AEA91E72DFFBC7CB4FBE9FDD7ED31" bold="true" box="[459,551,1071,1095]" pageId="15" pageNumber="16">sp. nov.</emphasis>
</taxonomicNameLabel>
has fewer dorsal blotches than 
<taxonomicName id="4C4E4D00E72DFFBC7EE7FBF6FBDAED31" box="[920,1066,1071,1095]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="morandae" status="sp. nov.">
<emphasis id="B93AEA91E72DFFBC7EE7FBF6FBDAED31" box="[920,1066,1071,1095]" italics="true" pageId="15" pageNumber="16">L. morandae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20957EAE72DFFBC794BFBE9FB60ED31" box="[1076,1168,1071,1095]" pageId="15" pageNumber="16" rank="species">
<emphasis id="B93AEA91E72DFFBC794BFBE9FB60ED31" bold="true" box="[1076,1168,1071,1095]" pageId="15" pageNumber="16">sp. nov.</emphasis>
</taxonomicNameLabel>
, and the blotches in 
<taxonomicName id="4C4E4D00E72DFFBC78FAFBF6FEF9ED1D" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="morandae" status="sp. nov.">
<emphasis id="B93AEA91E72DFFBC78FAFBF6FEF9ED1D" italics="true" pageId="15" pageNumber="16">L. morandae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20957EAE72DFFBC7C70FB95FE98ED1D" box="[271,360,1107,1131]" pageId="15" pageNumber="16" rank="species">
<emphasis id="B93AEA91E72DFFBC7C70FB95FE98ED1D" bold="true" box="[271,360,1107,1131]" pageId="15" pageNumber="16">sp. nov.</emphasis>
</taxonomicNameLabel>
have (50% of cases) brownish coloration, while 
<taxonomicName id="4C4E4D00E72DFFBC7EF8FB92FC18ED1D" box="[903,1000,1107,1131]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="avilae" status="sp. nov.">
<emphasis id="B93AEA91E72DFFBC7EF8FB92FC18ED1D" box="[903,1000,1107,1131]" italics="true" pageId="15" pageNumber="16">L. avilae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20957EAE72DFFBC7E91FB95FBB7ED1D" box="[1006,1095,1107,1131]" pageId="15" pageNumber="16" rank="species">
<emphasis id="B93AEA91E72DFFBC7E91FB95FBB7ED1D" bold="true" box="[1006,1095,1107,1131]" pageId="15" pageNumber="16">sp. nov.</emphasis>
</taxonomicNameLabel>
blotches have dark gray color. Ventral hand and foot surfaces of 
<taxonomicName id="4C4E4D00E72DFFBC7F6BFBBEFD54EDF9" box="[532,676,1143,1167]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="morandae" status="sp. nov.">
<emphasis id="B93AEA91E72DFFBC7F6BFBBEFD54EDF9" box="[532,676,1143,1167]" italics="true" pageId="15" pageNumber="16">L. morandae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20957EAE72DFFBC7FD3FBB1FCF6EDF9" box="[684,774,1143,1167]" pageId="15" pageNumber="16" rank="species">
<emphasis id="B93AEA91E72DFFBC7FD3FBB1FCF6EDF9" bold="true" box="[684,774,1143,1167]" pageId="15" pageNumber="16">sp. nov.</emphasis>
</taxonomicNameLabel>
present mucronated and “bristly” scales that are less obvious in 
<taxonomicName id="4C4E4D00E72DFFBC7D9EFB5AFEB4EDC5" box="[225,324,1179,1203]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="avilae" status="sp. nov.">
<emphasis id="B93AEA91E72DFFBC7D9EFB5AFEB4EDC5" box="[225,324,1179,1203]" italics="true" pageId="15" pageNumber="16">L. avilae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20957EAE72DFFBC7C35FB5DFE54EDC5" box="[330,420,1179,1203]" pageId="15" pageNumber="16" rank="species">
<emphasis id="B93AEA91E72DFFBC7C35FB5DFE54EDC5" bold="true" box="[330,420,1179,1203]" pageId="15" pageNumber="16">sp. nov.</emphasis>
</taxonomicNameLabel>
</paragraph>
<paragraph id="8BF13683E72DFFBC7DBAFB79FBA4EF6D" blockId="15.[151,1437,417,2031]" pageId="15" pageNumber="16">
<taxonomicName id="4C4E4D00E72DFFBC7DBAFB79FE74EDA1" box="[197,388,1215,1239]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="avilae" status="sp. nov.">
<emphasis id="B93AEA91E72DFFBC7DBAFB79FE74EDA1" box="[197,388,1215,1239]" italics="true" pageId="15" pageNumber="16">Liolaemus avilae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20957EAE72DFFBC7CF4FB79FE14EDA1" box="[395,484,1215,1239]" pageId="15" pageNumber="16" rank="species">
<emphasis id="B93AEA91E72DFFBC7CF4FB79FE14EDA1" bold="true" box="[395,484,1215,1239]" pageId="15" pageNumber="16">sp. nov.</emphasis>
</taxonomicNameLabel>
differs from 
<taxonomicName id="4C4E4D00E72DFFBC7F09FB06FCCFEDA1" box="[630,831,1215,1239]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="lineomaculatus">
<emphasis id="B93AEA91E72DFFBC7F09FB06FCCFEDA1" box="[630,831,1215,1239]" italics="true" pageId="15" pageNumber="16">L. lineomaculatus</emphasis>
</taxonomicName>
in the following traits: more scales in contact with the interparietal (7–10, X = 8,13 vs. 6–10, X = 7.18; 
<emphasis id="B93AEA91E72DFFBC7FBFFB22FD3EED8D" box="[704,718,1252,1275]" italics="true" pageId="15" pageNumber="16">p</emphasis>
= 0.0027), fewer supralabial scales (5–6, X = 5.25 vs. 5–8, X = 6.71; 
<emphasis id="B93AEA91E72DFFBC7DA5FACEFF18EC69" box="[218,232,1288,1311]" italics="true" pageId="15" pageNumber="16">p</emphasis>
= 0.0008), fewer midbody scales (53–59, X = 55.5 vs. 54–65, X = 60; 
<emphasis id="B93AEA91E72DFFBC794BFACEFBB2EC69" box="[1076,1090,1288,1311]" italics="true" pageId="15" pageNumber="16">p</emphasis>
&lt;0.0001), fewer scales from occiput to rump (51–58, X = 53.75 vs. 52–68, X = 58.41; 
<emphasis id="B93AEA91E72DFFBC7E5EFAEAFCDFEC35" box="[801,815,1324,1347]" italics="true" pageId="15" pageNumber="16">p</emphasis>
&lt;0.0001), fewer ventral scales (72–91, X = 83 vs. 80– 95, X = 85.29; 
<emphasis id="B93AEA91E72DFFBC7C3EFA96FEBFEC11" box="[321,335,1360,1383]" italics="true" pageId="15" pageNumber="16">p</emphasis>
&lt;0.0001), larger eyes (eye height: 2–2.62, X = 2.2 vs. 1.5–2.21, X = 1.84, eye length: 2.63–3.32, X = 3.07 vs. 2.35–3.3, X = 2.83; 
<emphasis id="B93AEA91E72DFFBC7F76FAB2FDE7ECFD" box="[521,535,1396,1419]" italics="true" pageId="15" pageNumber="16">p</emphasis>
&lt;
<quantity id="4CB69B66E72DFFBC7F4AFAB5FD6FECFD" box="[565,671,1395,1419]" metricMagnitude="-6" metricUnit="m" metricValue="2.54" pageId="15" pageNumber="16" unit="in" value="1.0E-4">0.0001 in</quantity>
both cases), and longer auditory meatus lengths (1.11–2.1, X = 1.51 vs. 0.87–1.39, X = 1.15; 
<emphasis id="B93AEA91E72DFFBC7CCEFA5EFE4FECD9" box="[433,447,1432,1455]" italics="true" pageId="15" pageNumber="16">p</emphasis>
&lt;0.0001). 
<taxonomicName id="4C4E4D00E72DFFBC7F3CFA51FCF3ECD9" box="[579,771,1431,1455]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="avilae" status="sp. nov.">
<emphasis id="B93AEA91E72DFFBC7F3CFA51FCF3ECD9" box="[579,771,1431,1455]" italics="true" pageId="15" pageNumber="16">Liolaemus avilae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20957EAE72DFFBC7E73FA51FC96ECD9" box="[780,870,1431,1455]" pageId="15" pageNumber="16" rank="species">
<emphasis id="B93AEA91E72DFFBC7E73FA51FC96ECD9" bold="true" box="[780,870,1431,1455]" pageId="15" pageNumber="16">sp. nov.</emphasis>
</taxonomicNameLabel>
has fewer dorsal blotches than 
<emphasis id="B93AEA91E72DFFBC79B3FA5EFA6CECD9" box="[1228,1436,1431,1455]" italics="true" pageId="15" pageNumber="16">
<taxonomicName id="4C4E4D00E72DFFBC79B3FA5EFA68ECD9" box="[1228,1432,1431,1455]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="lineomaculatus">L. lineomaculatus</taxonomicName>
.
</emphasis>
The vertebral line is less well defined in 
<taxonomicName id="4C4E4D00E72DFFBC7F21FA7AFD31ECA5" box="[606,705,1467,1491]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="avilae" status="sp. nov.">
<emphasis id="B93AEA91E72DFFBC7F21FA7AFD31ECA5" box="[606,705,1467,1491]" italics="true" pageId="15" pageNumber="16">L. avilae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20957EAE72DFFBC7FB7FA7DFCD1ECA5" box="[712,801,1467,1491]" pageId="15" pageNumber="16" rank="species">
<emphasis id="B93AEA91E72DFFBC7FB7FA7DFCD1ECA5" bold="true" box="[712,801,1467,1491]" pageId="15" pageNumber="16">sp. nov.</emphasis>
</taxonomicNameLabel>
than in 
<taxonomicName id="4C4E4D00E72DFFBC7E03FA7AFBB5ECA5" box="[892,1093,1467,1491]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="lineomaculatus">
<emphasis id="B93AEA91E72DFFBC7E03FA7AFBB5ECA5" box="[892,1093,1467,1491]" italics="true" pageId="15" pageNumber="16">L. lineomaculatus</emphasis>
</taxonomicName>
. In general, dorsal scales of 
<taxonomicName id="4C4E4D00E72DFFBC78FAFA7AFF2BEC81" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="avilae" status="sp. nov.">
<emphasis id="B93AEA91E72DFFBC78FAFA7AFF2BEC81" italics="true" pageId="15" pageNumber="16">L. avilae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20957EAE72DFFBC7D9BFA19FECEEC81" box="[228,318,1503,1527]" pageId="15" pageNumber="16" rank="species">
<emphasis id="B93AEA91E72DFFBC7D9BFA19FECEEC81" bold="true" box="[228,318,1503,1527]" pageId="15" pageNumber="16">sp. nov.</emphasis>
</taxonomicNameLabel>
are more mucronated than in 
<emphasis id="B93AEA91E72DFFBC7FE9FA26FBC1EC81" box="[662,1073,1503,1527]" italics="true" pageId="15" pageNumber="16">
<taxonomicName id="4C4E4D00E72DFFBC7FE9FA26FC94EC81" box="[662,868,1503,1527]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="lineomaculatus">L. lineomaculatus</taxonomicName>
. 
<taxonomicName id="4C4E4D00E72DFFBC7E0FFA19FBC1EC81" box="[880,1073,1503,1527]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="avilae" status="sp. nov.">Liolaemus avilae</taxonomicName>
</emphasis>
<taxonomicNameLabel id="A20957EAE72DFFBC7946FA19FB64EC81" box="[1081,1172,1503,1527]" pageId="15" pageNumber="16" rank="species">
<emphasis id="B93AEA91E72DFFBC7946FA19FB64EC81" bold="true" box="[1081,1172,1503,1527]" pageId="15" pageNumber="16">sp. nov.</emphasis>
</taxonomicNameLabel>
possesses scales in the dorsal surface of the forelimbs that are less carinated than those of 
<taxonomicName id="4C4E4D00E72DFFBC7EFBF9C2FBBDEF6D" box="[900,1101,1539,1563]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="lineomaculatus">
<emphasis id="B93AEA91E72DFFBC7EFBF9C2FBBDEF6D" box="[900,1101,1539,1563]" italics="true" pageId="15" pageNumber="16">L. lineomaculatus</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BF13683E72DFFA37DBAF9E1FEE9E9A2" blockId="15.[151,1437,417,2031]" lastBlockId="16.[151,1437,151,2012]" lastPageId="16" lastPageNumber="17" pageId="15" pageNumber="16">
Compared with 
<emphasis id="B93AEA91E72DFFBC7C06F9EEFD94EF49" box="[377,612,1575,1599]" italics="true" pageId="15" pageNumber="16">
<taxonomicName id="4C4E4D00E72DFFBC7C06F9EEFE07EF49" box="[377,503,1575,1599]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="hatcheri">L. hatcheri</taxonomicName>
, 
<taxonomicName id="4C4E4D00E72DFFBC7F7DF9EEFD94EF49" box="[514,612,1575,1599]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="avilae" status="sp. nov.">L. avilae</taxonomicName>
</emphasis>
<taxonomicNameLabel id="A20957EAE72DFFBC7F13F9E1FD36EF49" box="[620,710,1575,1599]" pageId="15" pageNumber="16" rank="species">
<emphasis id="B93AEA91E72DFFBC7F13F9E1FD36EF49" bold="true" box="[620,710,1575,1599]" pageId="15" pageNumber="16">sp. nov.</emphasis>
</taxonomicNameLabel>
has more scales in contact with the interparietal (7–10, X = 8,13 vs. 5–10, X = 6.85; 
<emphasis id="B93AEA91E72DFFBC7C09F98AFE74EF15" box="[374,388,1612,1635]" italics="true" pageId="15" pageNumber="16">p</emphasis>
= 0.0027), fewer supralabial scales (5–6, X = 5.25 vs. 6–8, X = 6.85; 
<emphasis id="B93AEA91E72DFFBC79E9F98AFB54EF15" box="[1174,1188,1612,1635]" italics="true" pageId="15" pageNumber="16">p</emphasis>
= 0.0008), more midbody scales (53–59, X = 55.5 vs. 43–57, X = 48.65; 
<emphasis id="B93AEA91E72DFFBC7F8CF9B6FCF1EFF1" box="[755,769,1648,1671]" italics="true" pageId="15" pageNumber="16">p</emphasis>
&lt;0.0001), more scales from occiput to rump (51–58, X = 53.75 vs. 43–55, X = 48.75; 
<emphasis id="B93AEA91E72DFFBC7C9CF952FE01EFDD" box="[483,497,1684,1707]" italics="true" pageId="15" pageNumber="16">p</emphasis>
&lt;0.0001), more ventral scales (72–91, X = 83 vs. 62–74, X = 67.8; 
<emphasis id="B93AEA91E72DFFBC7870F952FAEDEFDD" box="[1295,1309,1684,1707]" italics="true" pageId="15" pageNumber="16">p</emphasis>
&lt;0.0001), more third finger lamellae (15–18, X = 16.25 vs. 13–18, X = 14.85; 
<emphasis id="B93AEA91E72DFFBC7EE7F97EFC56EFB9" box="[920,934,1720,1743]" italics="true" pageId="15" pageNumber="16">p</emphasis>
&lt;0.0001), more fourth toe lamellae (20–23, X = 21.29 vs. 18–22, X = 19.45; 
<emphasis id="B93AEA91E72DFFBC7F75F91AFDE8EF85" box="[522,536,1756,1779]" italics="true" pageId="15" pageNumber="16">p</emphasis>
&lt;0.0001), smaller body size (snout-vent length: 48–59, X = 55.38 vs. 55–69, X = 61.89, axilla-groin distance: 19.4–29.5, X = 25.5 vs. 19.5–35.5, X = 28.95; 
<emphasis id="B93AEA91E72DFFBC7966F8C6FBD7EE61" box="[1049,1063,1792,1815]" italics="true" pageId="15" pageNumber="16">p</emphasis>
&lt;
<quantity id="4CB69B66E72DFFBC7936F939FB45EE61" box="[1097,1205,1791,1815]" metricMagnitude="-6" metricUnit="m" metricValue="2.54" pageId="15" pageNumber="16" unit="in" value="1.0E-4">0.0001 in</quantity>
both cases), shorter forelimbs (elbow-wrist length: 4.89–6.81, X = 5.76 vs. 5.67–7.33, X = 6.44, third finger length: 7.27–9.52, X = 8.02 vs. 8.5–10.57, X = 9.15; 
<emphasis id="B93AEA91E72DFFBC7C8DF88EFDF0EE29" box="[498,512,1864,1887]" italics="true" pageId="15" pageNumber="16">p</emphasis>
&lt;
<quantity id="4CB69B66E72DFFBC7F5DF881FD7EEE29" box="[546,654,1863,1887]" metricMagnitude="-6" metricUnit="m" metricValue="2.54" pageId="15" pageNumber="16" unit="in" value="1.0E-4">0.0001 in</quantity>
both cases), and smaller head width (9.02–10.19, X = 9.59 vs. 9.7– 13.05, X = 11.22; 
<emphasis id="B93AEA91E72DFFBC7C1EF8AAFE9FEEF5" box="[353,367,1900,1923]" italics="true" pageId="15" pageNumber="16">p</emphasis>
&lt;0.0001). 
<taxonomicName id="4C4E4D00E72DFFBC7C8FF8ADFD40EEF5" box="[496,688,1899,1923]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="avilae" status="sp. nov.">
<emphasis id="B93AEA91E72DFFBC7C8FF8ADFD40EEF5" box="[496,688,1899,1923]" italics="true" pageId="15" pageNumber="16">Liolaemus avilae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20957EAE72DFFBC7FC9F8ADFCE0EEF5" box="[694,784,1899,1923]" pageId="15" pageNumber="16" rank="species">
<emphasis id="B93AEA91E72DFFBC7FC9F8ADFCE0EEF5" bold="true" box="[694,784,1899,1923]" pageId="15" pageNumber="16">sp. nov.</emphasis>
</taxonomicNameLabel>
possesses fewer dorsal blotches and a more defined vertebral line relative to 
<taxonomicName id="4C4E4D00E72DFFBC7C01F856FE0CEED1" box="[382,508,1935,1959]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="hatcheri">
<emphasis id="B93AEA91E72DFFBC7C01F856FE0CEED1" box="[382,508,1935,1959]" italics="true" pageId="15" pageNumber="16">L. hatcheri</emphasis>
</taxonomicName>
. Ventral melanism is more pronounced in 
<taxonomicName id="4C4E4D00E72DFFBC7E91F856FB9CEED1" box="[1006,1132,1935,1959]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="hatcheri">
<emphasis id="B93AEA91E72DFFBC7E91F856FB9CEED1" box="[1006,1132,1935,1959]" italics="true" pageId="15" pageNumber="16">L. hatcheri</emphasis>
</taxonomicName>
than in 
<taxonomicName id="4C4E4D00E72DFFBC79AFF856FAC5EED1" box="[1232,1333,1935,1959]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="avilae" status="sp. nov.">
<emphasis id="B93AEA91E72DFFBC79AFF856FAC5EED1" box="[1232,1333,1935,1959]" italics="true" pageId="15" pageNumber="16">L. avilae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20957EAE72DFFBC7840F849FA6CEED1" box="[1343,1436,1935,1959]" pageId="15" pageNumber="16" rank="species">
<emphasis id="B93AEA91E72DFFBC7840F849FA6CEED1" bold="true" box="[1343,1436,1935,1959]" pageId="15" pageNumber="16">sp. nov.</emphasis>
</taxonomicNameLabel>
(fewer melanophores). Dorsal scales of 
<taxonomicName id="4C4E4D00E72DFFBC7F20F872FD32EEBD" box="[607,706,1971,1995]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="avilae" status="sp. nov.">
<emphasis id="B93AEA91E72DFFBC7F20F872FD32EEBD" box="[607,706,1971,1995]" italics="true" pageId="15" pageNumber="16">L. avilae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20957EAE72DFFBC7FB3F875FCD7EEBD" box="[716,807,1971,1995]" pageId="15" pageNumber="16" rank="species">
<emphasis id="B93AEA91E72DFFBC7FB3F875FCD7EEBD" bold="true" box="[716,807,1971,1995]" pageId="15" pageNumber="16">sp. nov.</emphasis>
</taxonomicNameLabel>
are less “bristly” than those of 
<emphasis id="B93AEA91E72DFFBC79EAF872FAE9EEBD" box="[1173,1305,1971,1995]" italics="true" pageId="15" pageNumber="16">
<taxonomicName id="4C4E4D00E72DFFBC79EAF872FAE5EEBD" box="[1173,1301,1971,1995]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="hatcheri">L. hatcheri</taxonomicName>
.
</emphasis>
In the new species, limb surface scales (fore-hind and dorsal-ventral) are non-mucronate and carinated (in 
<taxonomicName id="4C4E4D00E72DFFBC798FF81EFA9FEE99" box="[1264,1391,2007,2031]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="hatcheri">
<emphasis id="B93AEA91E72DFFBC798FF81EFA9FEE99" box="[1264,1391,2007,2031]" italics="true" pageId="15" pageNumber="16">L. hatcheri</emphasis>
</taxonomicName>
the majority are mucronated and non-carinate). Ventral scales are round-shaped in 
<taxonomicName id="4C4E4D00E732FFA37978FF5FFB98E9C6" box="[1031,1128,152,176]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="avilae" status="sp. nov.">
<emphasis id="B93AEA91E732FFA37978FF5FFB98E9C6" box="[1031,1128,152,176]" italics="true" pageId="16" pageNumber="17">L. avilae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20957EAE732FFA37910FF5EFB39E9C6" box="[1135,1225,152,176]" pageId="16" pageNumber="17" rank="species">
<emphasis id="B93AEA91E732FFA37910FF5EFB39E9C6" bold="true" box="[1135,1225,152,176]" pageId="16" pageNumber="17">sp. nov.</emphasis>
</taxonomicNameLabel>
and rhomboidal in 
<taxonomicName id="4C4E4D00E732FFA37DE8FF7BFEE1E9A2" box="[151,273,188,212]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="hatcheri">
<emphasis id="B93AEA91E732FFA37DE8FF7BFEE1E9A2" box="[151,273,188,212]" italics="true" pageId="16" pageNumber="17">L. hatcheri</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BF13683E732FFA37DBAFF19FEF0EA62" blockId="16.[151,1437,151,2012]" pageId="16" pageNumber="17">
Compared with 
<emphasis id="B93AEA91E732FFA37C06FF27FDAFE98E" box="[377,607,224,248]" italics="true" pageId="16" pageNumber="17">
<taxonomicName id="4C4E4D00E732FFA37C06FF27FE02E98E" box="[377,498,224,248]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="kolengh">L. kolengh</taxonomicName>
, 
<taxonomicName id="4C4E4D00E732FFA37C82FF27FDAFE98E" box="[509,607,224,248]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="avilae" status="sp. nov.">L. avilae</taxonomicName>
</emphasis>
<taxonomicNameLabel id="A20957EAE732FFA37F18FF26FD30E98E" box="[615,704,224,248]" pageId="16" pageNumber="17" rank="species">
<emphasis id="B93AEA91E732FFA37F18FF26FD30E98E" bold="true" box="[615,704,224,248]" pageId="16" pageNumber="17">sp. nov.</emphasis>
</taxonomicNameLabel>
has fewer scales in contact with the interparietal (7–10, X = 8,13 vs. 4–8, X = 6.26; 
<emphasis id="B93AEA91E732FFA37C14FEC3FE89E86A" box="[363,377,261,284]" italics="true" pageId="16" pageNumber="17">p</emphasis>
= 0.0027), fewer supralabial scales (5–6, X = 5.25 vs. 5–8, X = 6.65; 
<emphasis id="B93AEA91E732FFA379EBFEC3FB52E86A" box="[1172,1186,261,284]" italics="true" pageId="16" pageNumber="17">p</emphasis>
= 0.0008), more midbody scales (53–59, X = 55.5 vs. 44–55, X = 50.55; 
<emphasis id="B93AEA91E732FFA37F8CFEEFFCF1E836" box="[755,769,297,320]" italics="true" pageId="16" pageNumber="17">p</emphasis>
&lt;0.0001), more scales from occiput to rump (51–58, X = 53.75 vs. 44–54, X = 49.35; 
<emphasis id="B93AEA91E732FFA37CA0FE8BFE1DE812" box="[479,493,333,356]" italics="true" pageId="16" pageNumber="17">p</emphasis>
&lt;0.0001), more ventral scales (72–91, X = 83 vs. 61–76, X = 68.58; 
<emphasis id="B93AEA91E732FFA3786FFE8BFAEEE812" box="[1296,1310,333,356]" italics="true" pageId="16" pageNumber="17">p</emphasis>
&lt;0.0001), more third finger lamellae (15–18, X = 16.25 vs. 13–16, X = 14.6; 
<emphasis id="B93AEA91E732FFA37EFEFEB7FC7FE8FE" box="[897,911,369,392]" italics="true" pageId="16" pageNumber="17">p</emphasis>
&lt;0.0001), more fourth toe lamellae (20–23, X = 21.29 vs. 16–22, X = 18.74; 
<emphasis id="B93AEA91E732FFA37F7DFE53FDE0E8DA" box="[514,528,405,428]" italics="true" pageId="16" pageNumber="17">p</emphasis>
&lt;0.0001), longer tails (64–77, X = 68.88 vs. 47–70, X = 60.52, 
<emphasis id="B93AEA91E732FFA37872FE53FAEBE8DA" box="[1293,1307,405,428]" italics="true" pageId="16" pageNumber="17">p</emphasis>
= 0.0019), shorter forelimbs (elbow-wrist length: 4.89–6.81, X = 5.76 vs. 4.6–7.03, X = 6.16, third finger length: 7.27–9.52, X = 8.02 vs. 8.31–11.12, X = 9.65; 
<emphasis id="B93AEA91E732FFA37F68FE1BFDD5E882" box="[535,549,477,500]" italics="true" pageId="16" pageNumber="17">p</emphasis>
&lt;
<quantity id="4CB69B66E732FFA37F38FE1AFD43E882" box="[583,691,476,500]" metricMagnitude="-6" metricUnit="m" metricValue="2.54" pageId="16" pageNumber="17" unit="in" value="1.0E-4">0.0001 in</quantity>
both cases), smaller head width (9.02–10.19, X = 9.59 vs. 8.95– 11.71, X = 10.73; 
<emphasis id="B93AEA91E732FFA37C1DFDC7FE80EB6E" box="[354,368,513,536]" italics="true" pageId="16" pageNumber="17">p</emphasis>
&lt;0.0001), and smaller rostral scale (rostral height: 0.55–1.09, X = 0.85 vs. 0.87–1.2, X = 1.03, rostral length 2.31–2.69, X = 2.51 vs. 2.46–3.12, X = 2.77; 
<emphasis id="B93AEA91E732FFA37E3BFDE3FCA2EB4A" box="[836,850,549,572]" italics="true" pageId="16" pageNumber="17">p</emphasis>
&lt;
<quantity id="4CB69B66E732FFA37E0CFDE2FC2FEB4A" box="[883,991,548,572]" metricMagnitude="-6" metricUnit="m" metricValue="2.54" pageId="16" pageNumber="17" unit="in" value="1.0E-4">0.0001 in</quantity>
both cases). 
<taxonomicName id="4C4E4D00E732FFA37909FDE2FAC8EB4A" box="[1142,1336,548,572]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="avilae" status="sp. nov.">
<emphasis id="B93AEA91E732FFA37909FDE2FAC8EB4A" box="[1142,1336,548,572]" italics="true" pageId="16" pageNumber="17">Liolaemus avilae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20957EAE732FFA3783EFDE2FA6CEB4A" box="[1345,1436,548,572]" pageId="16" pageNumber="17" rank="species">
<emphasis id="B93AEA91E732FFA3783EFDE2FA6CEB4A" bold="true" box="[1345,1436,548,572]" pageId="16" pageNumber="17">sp. nov.</emphasis>
</taxonomicNameLabel>
exhibits a light background coloration, while the general background coloration of 
<taxonomicName id="4C4E4D00E732FFA3794BFD8FFB5AEB16" box="[1076,1194,584,608]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="kolengh">
<emphasis id="B93AEA91E732FFA3794BFD8FFB5AEB16" box="[1076,1194,584,608]" italics="true" pageId="16" pageNumber="17">L. kolengh</emphasis>
</taxonomicName>
is dark gray or black; however, the dorsal pattern could not be recognized in around 50% of our samples. 
<taxonomicName id="4C4E4D00E732FFA3791CFDAAFACAEBF2" box="[1123,1338,620,644]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="kolengh">
<emphasis id="B93AEA91E732FFA3791CFDAAFACAEBF2" box="[1123,1338,620,644]" italics="true" pageId="16" pageNumber="17">Liolaemus kolengh</emphasis>
</taxonomicName>
exhibits more dorsal blotches than 
<taxonomicName id="4C4E4D00E732FFA37CC1FD57FDEFEBDE" box="[446,543,656,680]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="avilae" status="sp. nov.">
<emphasis id="B93AEA91E732FFA37CC1FD57FDEFEBDE" box="[446,543,656,680]" italics="true" pageId="16" pageNumber="17">L. avilae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20957EAE732FFA37F58FD56FD70EBD1" box="[551,640,656,680]" pageId="16" pageNumber="17" rank="species">
<emphasis id="B93AEA91E732FFA37F58FD56FD89EBDE" bold="true" box="[551,633,656,680]" pageId="16" pageNumber="17">sp. nov</emphasis>
.
</taxonomicNameLabel>
Ventral melanism is stronger in 
<taxonomicName id="4C4E4D00E732FFA37E94FD57FB91EBDE" box="[1003,1121,656,680]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="kolengh">
<emphasis id="B93AEA91E732FFA37E94FD57FB91EBDE" box="[1003,1121,656,680]" italics="true" pageId="16" pageNumber="17">L. kolengh</emphasis>
</taxonomicName>
than in 
<taxonomicName id="4C4E4D00E732FFA379C3FD57FAEDEBDE" box="[1212,1309,656,680]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="avilae" status="sp. nov.">
<emphasis id="B93AEA91E732FFA379C3FD57FAEDEBDE" box="[1212,1309,656,680]" italics="true" pageId="16" pageNumber="17">L. avilae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20957EAE732FFA3785BFD56FA8EEBD1" box="[1316,1406,656,680]" pageId="16" pageNumber="17" rank="species">
<emphasis id="B93AEA91E732FFA3785BFD56FA87EBDE" bold="true" box="[1316,1399,656,680]" pageId="16" pageNumber="17">sp. nov</emphasis>
.
</taxonomicNameLabel>
In general, the dorsal scales of 
<taxonomicName id="4C4E4D00E732FFA37CA4FD73FDCEEBBA" box="[475,574,692,716]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="avilae" status="sp. nov.">
<emphasis id="B93AEA91E732FFA37CA4FD73FDCEEBBA" box="[475,574,692,716]" italics="true" pageId="16" pageNumber="17">L. avilae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20957EAE732FFA37F38FD72FD52EBBA" box="[583,674,692,716]" pageId="16" pageNumber="17" rank="species">
<emphasis id="B93AEA91E732FFA37F38FD72FD52EBBA" bold="true" box="[583,674,692,716]" pageId="16" pageNumber="17">sp. nov.</emphasis>
</taxonomicNameLabel>
are less “bristly” than scales of 
<emphasis id="B93AEA91E732FFA3796CFD73FB62EBBA" box="[1043,1170,692,716]" italics="true" pageId="16" pageNumber="17">
<taxonomicName id="4C4E4D00E732FFA3796CFD73FB7EEBBA" box="[1043,1166,692,716]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="kolengh">L. kolengh</taxonomicName>
.
</emphasis>
Limb scales (fore-hind and dorsal-ventral) are non-mucronate and non-“bristly” in 
<taxonomicName id="4C4E4D00E732FFA37E2EFD1FFC47EB86" box="[849,951,728,752]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="avilae" status="sp. nov.">
<emphasis id="B93AEA91E732FFA37E2EFD1FFC47EB86" box="[849,951,728,752]" italics="true" pageId="16" pageNumber="17">L. avilae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20957EAE732FFA37EBCFD1EFBD1EB86" box="[963,1057,728,752]" pageId="16" pageNumber="17" rank="species">
<emphasis id="B93AEA91E732FFA37EBCFD1EFBD1EB86" bold="true" box="[963,1057,728,752]" pageId="16" pageNumber="17">sp. nov.</emphasis>
</taxonomicNameLabel>
(mucronated and “bristly” in 
<taxonomicName id="4C4E4D00E732FFA378FAFD1FFF00EA62" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="kolengh">
<emphasis id="B93AEA91E732FFA378FAFD1FFF00EA62" italics="true" pageId="16" pageNumber="17">L. kolengh</emphasis>
</taxonomicName>
).
</paragraph>
<paragraph id="8BF13683E732FFA37DBAFCE6FB21ECB6" blockId="16.[151,1437,151,2012]" pageId="16" pageNumber="17">
<taxonomicName id="4C4E4D00E732FFA37DBAFCE6FE75EA4E" box="[197,389,800,824]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="avilae" status="sp. nov.">
<emphasis id="B93AEA91E732FFA37DBAFCE6FE75EA4E" box="[197,389,800,824]" italics="true" pageId="16" pageNumber="17">Liolaemus avilae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20957EAE732FFA37CF2FCE6FE17EA4E" box="[397,487,800,824]" pageId="16" pageNumber="17" rank="species">
<emphasis id="B93AEA91E732FFA37CF2FCE6FE17EA4E" bold="true" box="[397,487,800,824]" pageId="16" pageNumber="17">sp. nov.</emphasis>
</taxonomicNameLabel>
differs from 
<taxonomicName id="4C4E4D00E732FFA37F02FCE7FD09EA4E" box="[637,761,800,824]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="silvanae">
<emphasis id="B93AEA91E732FFA37F02FCE7FD09EA4E" box="[637,761,800,824]" italics="true" pageId="16" pageNumber="17">L. silvanae</emphasis>
</taxonomicName>
in several characters, including: a smaller number of scales in contact with the interparietal (7–10, X = 8,13 vs. 5–8, X = 6.94; 
<emphasis id="B93AEA91E732FFA37EF5FC83FC68EA2A" box="[906,920,837,860]" italics="true" pageId="16" pageNumber="17">p</emphasis>
= 0.0027), fewer supralabial scales (5–6, X = 5.25 vs. 5–7, X = 6.18; 
<emphasis id="B93AEA91E732FFA37CD6FCAFFE47EAF6" box="[425,439,873,896]" italics="true" pageId="16" pageNumber="17">p</emphasis>
= 0.0008), fewer midbody scales (53–59, X = 55.5 vs. 55–65, X = 59.65; 
<emphasis id="B93AEA91E732FFA37870FCAFFAEDEAF6" box="[1295,1309,873,896]" italics="true" pageId="16" pageNumber="17">p</emphasis>
&lt;0.0001), fewer scales from occiput to rump (51–58, X = 53.75 vs. 52–64, X = 56; 
<emphasis id="B93AEA91E732FFA37EACFC4BFC11EAD2" box="[979,993,909,932]" italics="true" pageId="16" pageNumber="17">p</emphasis>
&lt;0.0001), more ventral scales (72–91, X = 83 vs. 70–80, X = 74.59; 
<emphasis id="B93AEA91E732FFA37C94FC77FE09EABE" box="[491,505,945,968]" italics="true" pageId="16" pageNumber="17">p</emphasis>
&lt;0.0001), smaller body size (snout-vent length: 48–59, X = 55.38 vs. 65–78, X = 73, axilla-groin distance: 19.4–29.5, X = 25.3 vs. 25.1–39.7, X = 32.38; 
<emphasis id="B93AEA91E732FFA37EADFC13FC10EA9A" box="[978,992,981,1004]" italics="true" pageId="16" pageNumber="17">p</emphasis>
&lt;
<quantity id="4CB69B66E732FFA3797FFC12FB9BEA9A" box="[1024,1131,980,1004]" metricMagnitude="-6" metricUnit="m" metricValue="2.54" pageId="16" pageNumber="17" unit="in" value="1.0E-4">0.0001 in</quantity>
both cases), shorter fourth toe length (13.82–15.98, X = 14.91 vs. 16.5–20.1, X = 18.46), shorter forelimbs (elbow-wrist length: 4.89–6.81, X = 5.76 vs. 6.83–8.82, X = 7.71, third finger length: 7.27–9.52, X = 8.02 vs. 10.65–13.76, X = 12.23; 
<emphasis id="B93AEA91E732FFA3787AFBDBFAE3ED42" box="[1285,1299,1053,1076]" italics="true" pageId="16" pageNumber="17">p</emphasis>
&lt;
<quantity id="4CB69B66E732FFA3784EFBDAFA6CED42" box="[1329,1436,1052,1076]" metricMagnitude="-6" metricUnit="m" metricValue="2.54" pageId="16" pageNumber="17" unit="in" value="1.0E-4">0.0001 in</quantity>
both cases), smaller head (head height: 6.43–7.67, X = 7.07 vs. 6.84–9.06, X = 8.18, head width: 9.02–10.19, X = 9.59 vs. 11.31–14.71, X = 13.14, head length: 10.94–12.9, X = 11.78 vs. 12.47–15.9, X = 14.42; 
<emphasis id="B93AEA91E732FFA3799FFBA3FB1EED0A" box="[1248,1262,1125,1148]" italics="true" pageId="16" pageNumber="17">p</emphasis>
&lt;
<quantity id="4CB69B66E732FFA37871FBA2FA88ED0A" box="[1294,1400,1124,1148]" metricMagnitude="-6" metricUnit="m" metricValue="2.54" pageId="16" pageNumber="17" unit="in" value="1.0E-4">0.0001 in</quantity>
all cases), smaller eyes (eye height: 2–2.62, X = 2.2 vs. 2.09–2.9, X = 2.46, eye length: 2.63–3.32, X = 3.07 vs. 3.59– 4.18, X = 3.86; 
<emphasis id="B93AEA91E732FFA37C39FB6BFEA4EDB2" box="[326,340,1197,1220]" italics="true" pageId="16" pageNumber="17">p</emphasis>
&lt;
<quantity id="4CB69B66E732FFA37C0EFB6AFE2BEDB2" box="[369,475,1196,1220]" metricMagnitude="-6" metricUnit="m" metricValue="2.54" pageId="16" pageNumber="17" unit="in" value="1.0E-4">0.0001 in</quantity>
both cases), smaller rostral scale (rostral height: 0.55–1.09, X = 0.85 vs. 1.01–1.35, X = 1.19, rostral length: 2.31–2.69, X = 2.51 vs. 2.61–3.47, X = 3.13; 
<emphasis id="B93AEA91E732FFA37EF1FB17FC6CED9E" box="[910,924,1233,1256]" italics="true" pageId="16" pageNumber="17">p</emphasis>
&lt;
<quantity id="4CB69B66E732FFA37EC5FB09FBD4ED91" box="[954,1060,1231,1255]" metricMagnitude="-6" metricUnit="m" metricValue="2.54" pageId="16" pageNumber="17" unit="in" value="1.0E-4">0.0001 in</quantity>
both cases), and smaller auditory meatus (auditory meatus height: 1.95–2.56, X = 1.95 vs. 1.95–2.95, X = 2.54, auditory meatus length: 1.11–2.01, X = 1.51 vs. 1.49–2.57, X = 2.04; 
<emphasis id="B93AEA91E732FFA37C80FADFFDFDEC46" box="[511,525,1305,1328]" italics="true" pageId="16" pageNumber="17">p</emphasis>
&lt;
<quantity id="4CB69B66E732FFA37F54FAD1FD65EC59" box="[555,661,1303,1327]" metricMagnitude="-6" metricUnit="m" metricValue="2.54" pageId="16" pageNumber="17" unit="in" value="1.0E-4">0.0001 in</quantity>
both cases). Background coloration in 
<taxonomicName id="4C4E4D00E732FFA37931FADFFB39EC46" box="[1102,1225,1304,1328]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="silvanae">
<emphasis id="B93AEA91E732FFA37931FADFFB39EC46" box="[1102,1225,1304,1328]" italics="true" pageId="16" pageNumber="17">L. silvanae</emphasis>
</taxonomicName>
is black, there is a no recognizable dorsal pattern, and ventral area is strongly melanistic, whereas the ventral area is weakly melanistic in 
<taxonomicName id="4C4E4D00E732FFA37DAAFAA7FEC7EC0E" box="[213,311,1376,1400]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="avilae" status="sp. nov.">
<emphasis id="B93AEA91E732FFA37DAAFAA7FEC7EC0E" box="[213,311,1376,1400]" italics="true" pageId="16" pageNumber="17">L. avilae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20957EAE732FFA37C42FAA6FE66EC0E" box="[317,406,1376,1400]" pageId="16" pageNumber="17" rank="species">
<emphasis id="B93AEA91E732FFA37C42FAA6FE66EC0E" bold="true" box="[317,406,1376,1400]" pageId="16" pageNumber="17">sp. nov.</emphasis>
</taxonomicNameLabel>
<taxonomicName id="4C4E4D00E732FFA37CE3FAA6FDACEC0E" box="[412,604,1376,1400]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="avilae" status="sp. nov.">
<emphasis id="B93AEA91E732FFA37CE3FAA6FDACEC0E" box="[412,604,1376,1400]" italics="true" pageId="16" pageNumber="17">Liolaemus avilae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20957EAE732FFA37F1DFAA6FD4BEC0E" box="[610,699,1376,1400]" pageId="16" pageNumber="17" rank="species">
<emphasis id="B93AEA91E732FFA37F1DFAA6FD4BEC0E" bold="true" box="[610,699,1376,1400]" pageId="16" pageNumber="17">sp. nov.</emphasis>
</taxonomicNameLabel>
has a defined vertebral and paravertebral lines that are not present in 
<emphasis id="B93AEA91E732FFA37DCBFA43FEC7ECEA" box="[180,311,1412,1436]" italics="true" pageId="16" pageNumber="17">
<taxonomicName id="4C4E4D00E732FFA37DCBFA43FEC3ECEA" box="[180,307,1412,1436]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="silvanae">L. silvanae</taxonomicName>
.
</emphasis>
In general, 
<taxonomicName id="4C4E4D00E732FFA37CC2FA43FDC8ECEA" box="[445,568,1412,1436]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="silvanae">
<emphasis id="B93AEA91E732FFA37CC2FA43FDC8ECEA" box="[445,568,1412,1436]" italics="true" pageId="16" pageNumber="17">L. silvanae</emphasis>
</taxonomicName>
possesses a “bristly” appearance that is absent in 
<taxonomicName id="4C4E4D00E732FFA37914FA43FB3DECEA" box="[1131,1229,1412,1436]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="avilae" status="sp. nov.">
<emphasis id="B93AEA91E732FFA37914FA43FB3DECEA" box="[1131,1229,1412,1436]" italics="true" pageId="16" pageNumber="17">L. avilae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20957EAE732FFA379ABFA42FADEECEA" box="[1236,1326,1412,1436]" pageId="16" pageNumber="17" rank="species">
<emphasis id="B93AEA91E732FFA379ABFA42FADEECEA" bold="true" box="[1236,1326,1412,1436]" pageId="16" pageNumber="17">sp. nov.</emphasis>
</taxonomicNameLabel>
Scales of limbs (fore-hind and dorsal-ventral) are less mucronated in 
<taxonomicName id="4C4E4D00E732FFA37E4DFA6FFC64ECB6" box="[818,916,1448,1472]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="avilae" status="sp. nov.">
<emphasis id="B93AEA91E732FFA37E4DFA6FFC64ECB6" box="[818,916,1448,1472]" italics="true" pageId="16" pageNumber="17">L. avilae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20957EAE732FFA37EE4FA6EFC05ECB6" box="[923,1013,1448,1472]" pageId="16" pageNumber="17" rank="species">
<emphasis id="B93AEA91E732FFA37EE4FA6EFC05ECB6" bold="true" box="[923,1013,1448,1472]" pageId="16" pageNumber="17">sp. nov.</emphasis>
</taxonomicNameLabel>
than in 
<taxonomicName id="4C4E4D00E732FFA37930FA6FFB3AECB6" box="[1103,1226,1448,1472]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="silvanae">
<emphasis id="B93AEA91E732FFA37930FA6FFB3AECB6" box="[1103,1226,1448,1472]" italics="true" pageId="16" pageNumber="17">L. silvanae</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C3546508E732FFA77DBAFA0DFCCDEBBA" lastPageId="20" lastPageNumber="21" pageId="16" pageNumber="17" type="description">
<paragraph id="8BF13683E732FFA37DBAFA0DFEE5EFEE" blockId="16.[151,1437,151,2012]" pageId="16" pageNumber="17">
<emphasis id="B93AEA91E732FFA37DBAFA0DFE12EC92" bold="true" box="[197,482,1483,1508]" pageId="16" pageNumber="17">
Description of 
<typeStatus id="54F58821E732FFA37C0BFA0AFE2DEC92" box="[372,477,1484,1508]" pageId="16" pageNumber="17" type="holotype">holotype</typeStatus>
.
</emphasis>
Adult male. Snout-vent length 59.0 mm. Tail length (complete, not regenerated) 77.0 mm. Axilla-groin distance 
<quantity id="4CB69B66E732FFA37CBCFA36FDD9EF7E" box="[451,553,1520,1544]" metricMagnitude="-2" metricUnit="m" metricValue="2.69" pageId="16" pageNumber="17" unit="mm" value="26.9">26.9 mm</quantity>
. Head length 
<quantity id="4CB69B66E732FFA37FBEFA36FCD4EF7E" box="[705,804,1520,1544]" metricMagnitude="-2" metricUnit="m" metricValue="1.23" pageId="16" pageNumber="17" unit="mm" value="12.3">12.3 mm</quantity>
(from anterior border of tympanum to tip of snout), 
<quantity id="4CB69B66E732FFA37814FA36FF33EF5A" metricMagnitude="-2" metricUnit="m" metricValue="1.01" pageId="16" pageNumber="17" unit="mm" value="10.1">10.1 mm</quantity>
wide (at anterior border of tympanum), 
<quantity id="4CB69B66E732FFA37FF0F9D2FD16EF5A" box="[655,742,1556,1580]" metricMagnitude="-3" metricUnit="m" metricValue="7.6" pageId="16" pageNumber="17" unit="mm" value="7.6">7.6 mm</quantity>
high (at anterior border of tympanum). Snout length 
<quantity id="4CB69B66E732FFA3783AF9D2FA6CEF5A" box="[1349,1436,1556,1580]" metricMagnitude="-3" metricUnit="m" metricValue="4.2" pageId="16" pageNumber="17" unit="mm" value="4.2">4.2 mm</quantity>
(orbit-tip of snout distance). Interorbital distance 
<quantity id="4CB69B66E732FFA37FB7F9FEFCD2EF26" box="[712,802,1592,1616]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" pageId="16" pageNumber="17" unit="mm" value="4.5">4.5 mm</quantity>
. Eye-nostril distance 
<quantity id="4CB69B66E732FFA37968F9FEFB81EF26" box="[1047,1137,1592,1616]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="16" pageNumber="17" unit="mm" value="3.5">3.5 mm</quantity>
. Auditory meatus-eye distance 
<quantity id="4CB69B66E732FFA37DA4F99AFEC7EF02" box="[219,311,1628,1652]" metricMagnitude="-3" metricUnit="m" metricValue="4.9" pageId="16" pageNumber="17" unit="mm" value="4.9">4.9 mm</quantity>
. Forelimb length 
<quantity id="4CB69B66E732FFA37F7AF99AFD9EEF02" box="[517,622,1628,1652]" metricMagnitude="-2" metricUnit="m" metricValue="1.63" pageId="16" pageNumber="17" unit="mm" value="16.3">16.3 mm</quantity>
. Tibial length 11.0 mm. Foot length 
<quantity id="4CB69B66E732FFA37965F99AFB70EF02" box="[1050,1152,1628,1652]" metricMagnitude="-2" metricUnit="m" metricValue="1.59" pageId="16" pageNumber="17" unit="mm" value="15.9">15.9 mm</quantity>
(ankle to tip of claw on fourth toe).
</paragraph>
<paragraph id="8BF13683E732FFA27DBAF962FBE9EA41" blockId="16.[151,1437,151,2012]" lastBlockId="17.[151,1437,151,2012]" lastPageId="17" lastPageNumber="18" pageId="16" pageNumber="17">
Dorsal head scales bulged, smooth, 15 between occiput at level of anterior border of tympanum to rostral, pitted with numerous scale organs in the anterior region, and reducing to a single organ, or absence at the posterior half of the head. Rostral scale wider (
<quantity id="4CB69B66E732FFA37F40F92AFD6AEE72" box="[575,666,1772,1796]" metricMagnitude="-3" metricUnit="m" metricValue="2.8" pageId="16" pageNumber="17" unit="mm" value="2.8">2.8 mm</quantity>
) than high (
<quantity id="4CB69B66E732FFA37E5DF92AFC8FEE72" box="[802,895,1772,1796]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="16" pageNumber="17" unit="mm" value="0.8">0.8 mm</quantity>
). Two postrostrals, together with anterior lorilabial, separate nasal scales from rostral, surrounded by six scales. Nasal scales longer than wide, irregularly hexagonal, nostril one-half length of nasal, in a posterior position. Scales surrounding nasals 7 on the left side and 6 on the right side. Four internasals. Frontonasals four, irregular in size and position. Prefrontals 5, a small quadrangular scale in the center (
<quantity id="4CB69B66E732FFA37C01F8BAFE2CEEE2" box="[382,476,1916,1940]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" pageId="16" pageNumber="17" unit="mm" value="1.1">1.1 mm</quantity>
), two dorso-lateral larger scales roughly hexagonal (
<quantity id="4CB69B66E732FFA37940F8BAFB6EEEE2" box="[1087,1182,1916,1940]" metricMagnitude="-3" metricUnit="m" metricValue="1.7" pageId="16" pageNumber="17" unit="mm" value="1.7">1.7 mm</quantity>
), and a pair of lateral medium-sized scales (
<quantity id="4CB69B66E732FFA37CEAF866FE03EECE" box="[405,499,1952,1976]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="16" pageNumber="17" unit="mm" value="1.2">1.2 mm</quantity>
), approximately triangular. Three frontal scales. Frontoparietals in two rows, one anterior and two posterior scales. Interparietal pentagonal (
<quantity id="4CB69B66E732FFA37E4EF802FC7DEEAA" box="[817,909,1988,2012]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" pageId="16" pageNumber="17" unit="mm" value="1.3">1.3 mm</quantity>
), surrounded by seven scales; five in front and sides, irregular in shape and size, and two larger posterior scales. Parietal eye evident. Parietals slightly bulged, irregularly shaped, similar in size to interparietal. Circumorbitals: 12–10. Transversally expanded supraoculars 5– 4. Smaller lateral supraoculars: 16–19. One canthal higher than wide, separate from nasal by one postnasals. Loreal scales bulged, four on each side. Lorilabials longer than wide (7–6), approximately equal to labials. Superciliaries 7–8, flattened and elongated, anterior four broadly overlapping dorsally. Orbit with 12–15 upper and 13–11 lower ciliaries on each side. Orbit diameter 4.3 x 2.0 mm. Preocular small, unfragmented, square. Subocular scale elongated, approximately nine times longer than wide (3.8 x 
<quantity id="4CB69B66E733FFA27E5FFEA9FC8CE8F1" box="[800,892,367,391]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" pageId="17" pageNumber="18" unit="mm" value="0.6">0.6 mm</quantity>
). A well marked longitudinal ridge along upper margin of preocular and subocular scales. Postocular small, slightly bulged, ~ 25% superimposed onto subocular, with a marked longitudinal ridge. Palpebral scales small granular and bulged. Supralabials 6–5, convex. Temporals smooth, convex, juxtaposed with one scale organ in the tip. Anterior auriculars smaller than adjacent posterior temporals, projecting slightly outward (3–2). Posterior auricular scales small and granular. External auditory meatus conspicuous, higher (
<quantity id="4CB69B66E733FFA27CF8FDE2FE2FEB4A" box="[391,479,548,572]" metricMagnitude="-3" metricUnit="m" metricValue="1.7" pageId="17" pageNumber="18" unit="mm" value="1.7">1.7 mm</quantity>
) than wide (
<quantity id="4CB69B66E733FFA27F13FDE2FD37EB4A" box="[620,711,548,572]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="17" pageNumber="18" unit="mm" value="1.2">1.2 mm</quantity>
). Lateral scales of neck granular with inflated skin. Mental scale wider (3.0 mm) than high (1.0 mm), in contact with four scales. Mental followed posteriorly by two postmentals, and two rows of three chinshields on each side. Infralabials 5-4 on each side, first on each side quadrangular two times wider than supralabials, all others elongated, slightly smaller than supralabials. Gular scales smooth, flat, imbricate, with rounded posterior margins, with melanophores. Scales of throat between chinshields slightly juxtaposed, becoming slightly imbricate toward auditory meatus. Thirty-seven gulars between tympanum openings. Infralabials separated from chinshields by one to two rows of scales. Antehumeral and longitudinal neck folds well developed; postauricular, rictal, dorsolateral, oblique and gular not conspicuous.
</paragraph>
<paragraph id="8BF13683E733FFA27DBAFC82FCF3ED42" blockId="17.[151,1437,151,2012]" pageId="17" pageNumber="18">Scales of dorsal neck region rhomboidal, imbricate, strongly keeled. Very few trifids scales. Fifty-five dorsal scales between occiput and anterior surface of thighs. Dorsal body scales rhomboidal, imbricate, strongly keeled. Dorsal scales grade laterally into slightly smaller and keeled scales at midbody. Scales immediately anterior and posterior to forelimb and hindlimb insertion small, smooth, granular, and non-overlapping. Body lateral scales grading smaller to larger at midbody. Ventral body scales rhomboidals, smooth, flat, imbricate, larger than dorsal scales. Fifty-eight midbody scales; scales between mental and precloacal pores 90. Scales of cloacal region similar in size to ventral body scales; without precloacal pores.</paragraph>
<paragraph id="8BF13683E733FFA27DBAFB86FF18EF7E" blockId="17.[151,1437,151,2012]" pageId="17" pageNumber="18">Anterior suprabrachials rhomboidal, imbricate, smooth, slightly larger in size to dorsal body scales. Postabrachials smaller, smooth, becoming granular near axilla. Supra-antebrachials similar to suprabrachial. Infra-antebrachials rhomboidal, imbricate, smooth. Supracarpals imbricated, rhomboidal, smooth. Infracarpals imbricate, rhomboidal, very slightly keeled. Subdigital lamellae with 2–3 keels, each terminating in a short mucron, 2–3 mucronate, numbering: I: 8, II: 14, III: 17, IV: 19, V: 12. Claws robust, curved and sharp, opaque brown. Suprafemorals larger as dorsal body scales, rhomboidal, imbricated, smooth upper thigh, toward to the lateral thigh keeled. Postfemorals small, granular shape. Supratibials rhomboidal, imbricated, moderately keeled, smaller and smooth toward to the foot. Infrafemoral scale small, granular and smooth. Supratarsals rhomboidal, imbricated and smooth. Infratarsals small, rhomboidal, imbricate, smooth, some keeled and mucronate near the digit. Subdigital scales have 3 keels, 3–4 mucronate, numbering: I: 9, II: 14, III: 19, IV: 22, V: 16. Claws robust, curved and sharp, opaque brown. Tail complete, non-regenerated. Dorsal and lateral caudal scales, rhombic in the first half of the tail, quadrangulars toward the tip, strongly keeled. Ventral subtriangular and smooth, toward posterior half moderatly keeled.</paragraph>
<paragraph id="8BF13683E733FFA27DBAF9D5FD2CEE3A" blockId="17.[151,1437,151,2012]" pageId="17" pageNumber="18">
<emphasis id="B93AEA91E733FFA27DBAF9D5FE1EEF5A" bold="true" box="[197,494,1555,1580]" pageId="17" pageNumber="18">
Color of 
<typeStatus id="54F58821E733FFA27C4DF9D2FE69EF5A" box="[306,409,1556,1580]" pageId="17" pageNumber="18" type="holotype">holotype</typeStatus>
in life.
</emphasis>
Dark gray dorsal background (
<figureCitation id="13752A06E733FFA27E2CF9D2FC46EF5A" box="[851,950,1556,1580]" captionStart="FIGURE 7" captionStartId="18.[151,250,1929,1952]" captionTargetBox="[198,1398,305,1899]" captionTargetId="figure@18.[194,1398,301,1900]" captionTargetPageId="18" captionText="FIGURE 7. Upper, dorsal and ventral views of holotype in life of L. avilae sp. nov. MLP. S 2627; below, dorsal and ventral views of female (LJAMM-CNP 9399)." httpUri="https://zenodo.org/record/208137/files/figure.png" pageId="17" pageNumber="18">Figure 7</figureCitation>
) that becomes lighter on the lateral region between axilla and groin. Dorsal pattern presents nine paravertebral, quadrangulars, dark brown blotches, extending from the nuchal to the pelvic region, these series or blotches are combined at the beginning of the pelvic region into a pattern of transversal bands, which is present to the tip of the tail. Dorsolateral region between the occipital and the pelvic region is characterized by eight irregular, black and white-bordered blotches that are combined with the transversal bands of the tail. Lateral region, between axilla and groin, is characterized by a black and white reticulate pattern. In between the series of blotches a white vertebral and two yellowish paravertebral lines, onescale wide, are present. Vertebral line extends from the nuchal region, while the paravertebral lines extend from the temporal region; all are present to the pelvic region.
</paragraph>
<paragraph id="8BF13683E733FFA17DBAF89EFB3AE981" blockId="17.[151,1437,151,2012]" lastBlockId="18.[151,1436,151,248]" lastPageId="18" lastPageNumber="19" pageId="17" pageNumber="18">Dark gray head background. Two lateral white lines along the longitudinal fold are present from the top of the auditory meatus to the antehumeral fold. White subocular scale with a black longitudinal ridge. One longitudinal band is present in the anterior part of the head, crossing though internasals, postnasals, prefrontals, frontals, postfrontals and interparietal scales, ending in the interparietal scale that is surrounded by small brown spots; however, a lighter central region is observable in the frontal, postfrontal and interparietal scales. Anterior circumorbitals present a transverse dark brown band. Nasals scales dark brown. Small dark brown blotches on postocular, temporal and occipital region. Dorsal region of the limbs are dark gray with a dark brown reticulation.</paragraph>
<caption id="DF31660BE730FFA17DE8F84FFDD2EEC9" httpUri="https://zenodo.org/record/208137/files/figure.png" pageId="18" pageNumber="19" targetBox="[198,1398,305,1899]" targetPageId="18">
<paragraph id="8BF13683E730FFA17DE8F84FFDD2EEC9" blockId="18.[151,1436,1929,1983]" pageId="18" pageNumber="19">
<emphasis id="B93AEA91E730FFA17DE8F84FFEE4EEE9" bold="true" box="[151,276,1929,1952]" pageId="18" pageNumber="19">FIGURE 7.</emphasis>
Upper, dorsal and ventral views of holotype in life of 
<taxonomicName id="4C4E4D00E730FFA17E33F84DFC57EED6" box="[844,935,1930,1952]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="avilae" status="sp. nov.">
<emphasis id="B93AEA91E730FFA17E33F84DFC57EED6" box="[844,935,1930,1952]" italics="true" pageId="18" pageNumber="19">L. avilae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20957EAE730FFA17ED0F84CFBF1EEE9" box="[943,1025,1930,1951]" pageId="18" pageNumber="19" rank="species">
<emphasis id="B93AEA91E730FFA17ED0F84CFBF1EEE9" bold="true" box="[943,1025,1930,1951]" pageId="18" pageNumber="19">sp. nov.</emphasis>
</taxonomicNameLabel>
MLP.S 2627; below, dorsal and ventral views of female (LJAMM-CNP 9399).
</paragraph>
</caption>
<caption id="DF31660BE731FFA07DE8F86BFDC8EE94" httpUri="https://zenodo.org/record/208138/files/figure.png" pageId="19" pageNumber="20" targetBox="[212,1380,193,1945]" targetPageId="19">
<paragraph id="8BF13683E731FFA07DE8F86BFDC8EE94" blockId="19.[151,1436,1965,2018]" pageId="19" pageNumber="20">
<emphasis id="B93AEA91E731FFA07DE8F86BFEE6EEB4" bold="true" box="[151,278,1965,1988]" pageId="19" pageNumber="20">FIGURE 8.</emphasis>
Type locality of 
<taxonomicName id="4C4E4D00E731FFA07CAFF86BFD72EEB5" box="[464,642,1965,1987]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="19" pageNumber="20" phylum="Chordata" rank="species" species="avilae" status="sp. nov.">
<emphasis id="B93AEA91E731FFA07CAFF86BFD72EEB5" box="[464,642,1965,1987]" italics="true" pageId="19" pageNumber="20">Liolaemus avilae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20957EAE731FFA07FF4F868FD2FEEB5" box="[651,735,1966,1987]" pageId="19" pageNumber="20" rank="species">
<emphasis id="B93AEA91E731FFA07FF4F868FD2FEEB5" bold="true" box="[651,735,1966,1987]" pageId="19" pageNumber="20">sp. nov.</emphasis>
</taxonomicNameLabel>
Altoandina phytogeographic province, Altoandino Austral District (47º05’29.0” S, 71º01’12.9” W, 1154 m).
</paragraph>
</caption>
<paragraph id="8BF13683E736FFA77DBAFF51FB97E981" blockId="20.[151,1437,151,1256]" pageId="20" pageNumber="21">Gray ventral color with a dark reticulate pattern over the entire body, it is strongly marked in throat, chest, tail and body laterals, and extend to the adjacent malar and maxilla region. Ventral scales with melanophores. Ventral area of chest, belly, cloacal region and limbs tinged with a light orange color. Gray tail.</paragraph>
<paragraph id="8BF13683E736FFA77DBAFEC5FB36E8F1" blockId="20.[151,1437,151,1256]" pageId="20" pageNumber="21">
<emphasis id="B93AEA91E736FFA77DBAFEC5FDA3E86A" bold="true" box="[197,595,259,284]" pageId="20" pageNumber="21">
Color of 
<typeStatus id="54F58821E736FFA77C52FEC2FE64E86A" box="[301,404,260,284]" pageId="20" pageNumber="21" type="holotype">holotype</typeStatus>
in preservative.
</emphasis>
After three years in preservative, the dorsal coloration of the head, dorsum, body flanks and tail become darker while maintaining the contrast, but the two yellowish paravertebral lines turned gray. Ventral scales of throat, neck, chest, belly and forelimbs retain the same coloration as in life, and the distinctive light ventral orange color of the chest, belly, cloacal region and limbs turns gray (
<figureCitation id="13752A06E736FFA77929FEA9FB49E8F1" box="[1110,1209,367,391]" captionStart="FIGURE 6" captionStartId="12.[151,250,937,960]" captionTargetBox="[151,1436,209,931]" captionTargetId="figure@12.[151,1436,209,933]" captionTargetPageId="12" captionText="FIGURE 6. Morphological comparison between Liolaemus morandae sp. nov. (A), L. lineomaculatus (B) and L. avilae sp. nov. (C). Notice differences in color pattern on the tails, size of blotches on the limbs and differences in vertebral and paravertebral lines. Dorsal scales (shown on the right) are less mucronated in L. lineomaculatus. Dorsal scales of hindlimbs are less carinated in L. morandae sp. nov. than in the other species, while they are more mucronated in L. avilae sp. nov." httpUri="https://zenodo.org/record/208136/files/figure.png" pageId="20" pageNumber="21">Figure 6</figureCitation>
).
</paragraph>
<paragraph id="8BF13683E736FFA77DBAFE55FCCDEBBA" blockId="20.[151,1437,151,1256]" pageId="20" pageNumber="21">
<emphasis id="B93AEA91E736FFA77DBAFE55FECCE8DD" bold="true" box="[197,316,403,428]" pageId="20" pageNumber="21">Variation.</emphasis>
Morphological and meristic character variation between male and female 
<taxonomicName id="4C4E4D00E736FFA779D5FE52FA9EE8DA" box="[1194,1390,404,428]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="20" pageNumber="21" phylum="Chordata" rank="species" species="avilae" status="sp. nov.">
<emphasis id="B93AEA91E736FFA779D5FE52FA9EE8DA" box="[1194,1390,404,428]" italics="true" pageId="20" pageNumber="21">Liolaemus avilae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20957EAE736FFA77805FE52FF38E8A6" pageId="20" pageNumber="21" rank="species">
<emphasis id="B93AEA91E736FFA77805FE52FF38E8A6" bold="true" pageId="20" pageNumber="21">sp. nov.</emphasis>
</taxonomicNameLabel>
, is shown in 
<tableCitation id="C6CC0338E736FFA77C23FE71FE43E8B9" box="[348,435,439,463]" captionStart="TABLE 6" captionStartId="14.[151,235,632,653]" captionTargetBox="[151,1436,711,1852]" captionTargetPageId="14" captionText="TABLE 6. Variation between females and males of L. morandae sp. nov. and L. avilae sp. nov. Summarized as mean ± SD (min – max) values for all meristic and morphometric (in millimeters) variables." httpUri="http://table.plazi.org/id/DF31660BE72CFFBD7DE8FDBEFC35EBDA" pageId="20" pageNumber="21" tableUuid="DF31660BE72CFFBD7DE8FDBEFC35EBDA">Table 6</tableCitation>
. Females in life present basically the same dorsal and lateral patterns as present in males, but females generally have a lighter brown background coloration; vertebral and paravertebral lines are variable, in some individuals they are strongly defined while in others they are less conspicuous; lines vary from white to yellow in females and from light orange to pink in males. Ventrally dark reticulate pattern in the entire body is present and is strongly marked in the throat, tail and lateral body scales. Ventral, ventro-lateral, limb, cloacal and postcloacal regions vary from males to females, in the former a ventral light orange coloration is present, while in one female (out of three) a light pink ventral coloration is present. All distinctive ventral and dorsal coloration changed from yellow, orange or pink to gray or white in preservative.
</paragraph>
</subSubSection>
<subSubSection id="C3546508E736FFA77DBAFD1EFEC1EA41" pageId="20" pageNumber="21" type="etymology">
<paragraph id="8BF13683E736FFA77DBAFD1EFEC1EA41" blockId="20.[151,1437,151,1256]" pageId="20" pageNumber="21">
<emphasis id="B93AEA91E736FFA77DBAFD1EFEB4EB86" bold="true" box="[197,324,728,752]" pageId="20" pageNumber="21">Etymology</emphasis>
. The specific epithet of this species “ 
<taxonomicName id="4C4E4D00E736FFA77F9AFD1EFCD9EB86" box="[741,809,728,752]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="20" pageNumber="21" phylum="Chordata" rank="species" species="avilae">
<emphasis id="B93AEA91E736FFA77F9AFD1EFCD9EB86" box="[741,809,728,752]" italics="true" pageId="20" pageNumber="21">avilae</emphasis>
</taxonomicName>
” refers to our colleague and the first author’s PhD advisor Dr. Luciano Javier Avila, to honor him after more than fifteen years of research and teaching on the Patagonian herpetofauna.
</paragraph>
</subSubSection>
<subSubSection id="C3546508E736FFA77DBAFC85FD57ED9E" pageId="20" pageNumber="21" type="distribution">
<paragraph id="8BF13683E736FFA77DBAFC85FE02EAD2" blockId="20.[151,1437,151,1256]" pageId="20" pageNumber="21">
<emphasis id="B93AEA91E736FFA77DBAFC85FEAEEA2D" bold="true" box="[197,350,835,860]" pageId="20" pageNumber="21">Distribution.</emphasis>
<taxonomicName id="4C4E4D00E736FFA77C17FC82FDDBEA2A" box="[360,555,836,860]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="20" pageNumber="21" phylum="Chordata" rank="species" species="avilae" status="sp. nov.">
<emphasis id="B93AEA91E736FFA77C17FC82FDDBEA2A" box="[360,555,836,860]" italics="true" pageId="20" pageNumber="21">Liolaemus avilae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20957EAE736FFA77F4BFC82FD61EA2A" box="[564,657,836,860]" pageId="20" pageNumber="21" rank="species">
<emphasis id="B93AEA91E736FFA77F4BFC82FD61EA2A" bold="true" box="[564,657,836,860]" pageId="20" pageNumber="21">sp. nov.</emphasis>
</taxonomicNameLabel>
is known only from the 
<typeStatus id="54F58821E736FFA77ECAFC83FC15EA2B" box="[949,997,837,861]" pageId="20" pageNumber="21">type</typeStatus>
locality and surroundings, from Lago Buenos Aires Plateau in Puesto Lebrun and 18.7 SW from there, Santa Cruz province, Lago Buenos Aires department, 
<collectingCountry id="F3597613E736FFA77DA2FC4AFEBCEAD2" box="[221,332,908,932]" name="Argentina" pageId="20" pageNumber="21">Argentina</collectingCountry>
(
<figureCitation id="13752A06E736FFA77C24FC4AFE37EAD2" box="[347,455,908,932]" captionStart="FIGURE 5" captionStartId="10.[151,250,1053,1076]" captionTargetBox="[157,1427,198,1017]" captionTargetId="figure@10.[151,1436,193,1032]" captionTargetPageId="10" captionText="FIGURE 5. Distribution map for species of the lineomaculatus group, with geographical localities sampled in this study superimposed on regional elevation (shading). Black circles: L. morandae sp. nov.; white circle: L. avilae sp. nov.; black squares: L. lineomaculatus; white square: L. kolengh; white triangles: L. hatcheri; black triangle: L. silvanae. Arrows indicate type localities. CH: Chubut province; SC: Santa Cruz province." httpUri="https://zenodo.org/record/208135/files/figure.png" pageId="20" pageNumber="21">Figures 5</figureCitation>
, 
<figureCitation id="13752A06E736FFA77CABFC4AFE14EAD2" box="[468,484,908,932]" captionStart="FIGURE 8" captionStartId="19.[151,250,1965,1988]" captionTargetBox="[212,1380,193,1945]" captionTargetId="figure@19.[212,1380,193,1945]" captionTargetPageId="19" captionText="FIGURE 8. Type locality of Liolaemus avilae sp. nov. Altoandina phytogeographic province, Altoandino Austral District (47 º 05 ’ 29.0 ” S, 71 º 01 ’ 12.9 ” W, 1154 m)." httpUri="https://zenodo.org/record/208138/files/figure.png" pageId="20" pageNumber="21">8</figureCitation>
).
</paragraph>
<paragraph id="8BF13683E736FFA77DBAFC76FD57ED9E" blockId="20.[151,1437,151,1256]" pageId="20" pageNumber="21">
<emphasis id="B93AEA91E736FFA77DBAFC76FE74EABE" bold="true" box="[197,388,943,968]" pageId="20" pageNumber="21">Natural history.</emphasis>
Out of three females, only one presents a ventral pinkish coloration and a different one was pregnant; thus we infer that the pinkish coloration in females is not related to pregnancy. The species occurs in the Altoandina Phytogeographic province, Altoandino Austral district, in environments characterized as sparse grasslands and chamaephytes such as 
<emphasis id="B93AEA91E736FFA77F70FBDBFC22ED42" box="[527,978,1052,1076]" italics="true" pageId="20" pageNumber="21">
<taxonomicName id="4C4E4D00E736FFA77F70FBDBFC94ED42" authority="Nassauvia" authorityName="Nassauvia" box="[527,868,1052,1076]" class="Magnoliopsida" family="Ericaceae" genus="Empetrum" kingdom="Plantae" order="Ericales" pageId="20" pageNumber="21" phylum="Tracheophyta" rank="species" species="rubrum">Empetrum rubrum, Nassauvia</taxonomicName>
pygmaea
</emphasis>
and 
<emphasis id="B93AEA91E736FFA77973FBDAFB0AED42" box="[1036,1274,1052,1076]" italics="true" pageId="20" pageNumber="21">
<taxonomicName id="4C4E4D00E736FFA77973FBDAFB06ED42" box="[1036,1270,1052,1076]" class="Reptilia" family="Liolaemidae" genus="Azorrella" kingdom="Animalia" order="Squamata" pageId="20" pageNumber="21" phylum="Chordata" rank="species" species="ameghinoi">Azorrella ameghinoi</taxonomicName>
,
</emphasis>
among others (Roig 1998); some 
<taxonomicName id="4C4E4D00E736FFA77C0DFB87FE3BED2E" box="[370,459,1089,1112]" class="Liliopsida" family="Poaceae" genus="Festuca" kingdom="Plantae" order="Poales" pageId="20" pageNumber="21" phylum="Tracheophyta" rank="genus">
<emphasis id="B93AEA91E736FFA77C0DFB87FE3BED2E" box="[370,459,1089,1112]" italics="true" pageId="20" pageNumber="21">Festuca</emphasis>
</taxonomicName>
<taxonomicName id="4C4E4D00E736FFA77CACFB87FE0DED2E" box="[467,509,1089,1112]" class="Liliopsida" family="Poaceae" genus="Stipa" kingdom="Plantae" order="Poales" pageId="20" pageNumber="21" phylum="Tracheophyta" rank="species" species="spp">spp</taxonomicName>
. and 
<taxonomicName id="4C4E4D00E736FFA77F46FB86FD60ED21" box="[569,656,1088,1111]" class="Magnoliopsida" family="Asteraceae" genus="Senecio" kingdom="Plantae" order="Asterales" pageId="20" pageNumber="21" phylum="Tracheophyta" rank="genus">
<emphasis id="B93AEA91E736FFA77F46FB86FD60ED21" box="[569,656,1088,1111]" italics="true" pageId="20" pageNumber="21">Senecio</emphasis>
</taxonomicName>
<taxonomicName id="4C4E4D00E736FFA77FE7FB87FD33ED2E" box="[664,707,1089,1112]" class="Liliopsida" family="Poaceae" genus="Stipa" kingdom="Plantae" order="Poales" pageId="20" pageNumber="21" phylum="Tracheophyta" rank="species" species="spp">spp</taxonomicName>
. were also observed. This new species was found in open volcanic-sand substrates between patches of basaltic rocks, sharing the habitat with 
<taxonomicName id="4C4E4D00E736FFA77E80FBA2FB02ED0A" box="[1023,1266,1124,1148]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="20" pageNumber="21" phylum="Chordata" rank="species" species="archeforus">
<emphasis id="B93AEA91E736FFA77E80FBA2FB02ED0A" box="[1023,1266,1124,1148]" italics="true" pageId="20" pageNumber="21">Liolaemus archeforus</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C4E4D00E736FFA77858FBA2FF04EDD6" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="20" pageNumber="21" phylum="Chordata" rank="species" species="silvanae">
<emphasis id="B93AEA91E736FFA77858FBA2FF04EDD6" italics="true" pageId="20" pageNumber="21">Liolaemus silvanae</emphasis>
</taxonomicName>
(
<figureCitation id="13752A06E736FFA77C7CFB4EFE95EDD6" box="[259,357,1160,1184]" captionStart="FIGURE 8" captionStartId="19.[151,250,1965,1988]" captionTargetBox="[212,1380,193,1945]" captionTargetId="figure@19.[212,1380,193,1945]" captionTargetPageId="19" captionText="FIGURE 8. Type locality of Liolaemus avilae sp. nov. Altoandina phytogeographic province, Altoandino Austral District (47 º 05 ’ 29.0 ” S, 71 º 01 ’ 12.9 ” W, 1154 m)." httpUri="https://zenodo.org/record/208138/files/figure.png" pageId="20" pageNumber="21">Figure 8</figureCitation>
). Based on the natural history of its sister species 
<emphasis id="B93AEA91E736FFA77EE9FB4FFB96EDD6" box="[918,1126,1160,1184]" italics="true" pageId="20" pageNumber="21">
<taxonomicName id="4C4E4D00E736FFA77EE9FB4FFB92EDD6" box="[918,1122,1160,1184]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="20" pageNumber="21" phylum="Chordata" rank="species" species="lineomaculatus">L. lineomaculatus</taxonomicName>
,
</emphasis>
we hypothesize that 
<taxonomicName id="4C4E4D00E736FFA7782CFB4FFF49EDB2" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="20" pageNumber="21" phylum="Chordata" rank="species" species="avilae" status="sp. nov.">
<emphasis id="B93AEA91E736FFA7782CFB4FFF49EDB2" italics="true" pageId="20" pageNumber="21">L. avilae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20957EAE736FFA77DBFFB6AFEEAEDB2" box="[192,282,1196,1220]" pageId="20" pageNumber="21" rank="species">
<emphasis id="B93AEA91E736FFA77DBFFB6AFEEAEDB2" bold="true" box="[192,282,1196,1220]" pageId="20" pageNumber="21">sp. nov.</emphasis>
</taxonomicNameLabel>
is herbivorous and likely to also be viviparous, perhaps giving birth to between three and six individuals per clutch (Cei 1986; Espinoza 
<emphasis id="B93AEA91E736FFA77F60FB17FDA7ED9E" box="[543,599,1232,1256]" italics="true" pageId="20" pageNumber="21">et al.</emphasis>
2004).
</paragraph>
</subSubSection>
</treatment>
</document>