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<document id="2257560BA021D0866F7D13A0FB7C8A6B" ID-CLB-Dataset="45852" ID-DOI="10.5281/zenodo.208130" ID-GBIF-Dataset="990f7a3b-602c-47eb-9717-947c8f1bcb7c" ID-ISSN="1175-5326" ID-Zenodo-Dep="208130" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460416101720" checkinUser="plazi" docAuthor="Breitman, Maria Florencia, Parra, Micaela, Pérez, Cristian Hernán Fulvio &amp; Sites, Jack Walter" docDate="2011" docId="03E78795E724FFBF7DE8FADDFDA6EE0E" docLanguage="en" docName="zt03120p028.pdf" docOrigin="Zootaxa 3120" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Liolaemus morandae Breitman, Parra, Pérez &amp; Sites, 2011, sp. nov." docType="treatment" docVersion="10" lastPageNumber="13" masterDocId="FFDEFFEDE722FFB37D7FFFC6FFF0E976" masterDocTitle="Two new species of lizards from the Liolaemus lineomaculatus section (Squamata: Iguania: Liolaemidae) from southern Patagonia" masterLastPageNumber="28" masterPageNumber="1" pageNumber="7" updateTime="1698328881709" updateUser="ExternalLinkService">
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<mods:title id="E4F5708CD70DF322D3E8C45568B9FBE1">Two new species of lizards from the Liolaemus lineomaculatus section (Squamata: Iguania: Liolaemidae) from southern Patagonia</mods:title>
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<mods:namePart id="246466B8C7766E8A9A6809137971CDF1">Breitman, Maria Florencia</mods:namePart>
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<mods:namePart id="78A75F0D518FAE7933134BB746055446">Parra, Micaela</mods:namePart>
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<mods:namePart id="9AE1C805554E20B001F7D51674BAA8F3">Pérez, Cristian Hernán Fulvio</mods:namePart>
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<mods:role id="D1DB8B4E5BD0E3FB513BA762FAD15C6C">
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<mods:namePart id="86D4B03EC718A56CAF3051269A967FC8">Sites, Jack Walter</mods:namePart>
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<mods:date id="40233B3F1C031D75F11D723BDF634964">2011</mods:date>
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<treatment id="03E78795E724FFBF7DE8FADDFDA6EE0E" ID-DOI="http://doi.org/10.5281/zenodo.5615825" ID-GBIF-Taxon="119425263" ID-Zenodo-Dep="5615825" LSID="urn:lsid:plazi:treatment:03E78795E724FFBF7DE8FADDFDA6EE0E" httpUri="http://treatment.plazi.org/id/03E78795E724FFBF7DE8FADDFDA6EE0E" lastPageId="12" lastPageNumber="13" pageId="6" pageNumber="7">
<subSubSection id="C3546508E724FFB57DE8FADDFE6AECC2" pageId="6" pageNumber="7" type="nomenclature">
<paragraph id="8BF13683E724FFB57DE8FADDFE0CEC42" blockId="6.[151,508,1306,1365]" box="[151,508,1306,1333]" pageId="6" pageNumber="7">
<heading id="D0B981EFE724FFB57DE8FADDFE0CEC42" bold="true" box="[151,508,1306,1333]" fontSize="11" level="1" pageId="6" pageNumber="7" reason="1">
<emphasis id="B93AEA91E724FFB57DE8FADDFE0CEC42" bold="true" box="[151,508,1306,1333]" pageId="6" pageNumber="7">
<taxonomicName id="4C4E4D00E724FFB57DE8FADDFE67EC43" box="[151,407,1307,1333]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="morandae" status="sp. nov.">
<emphasis id="B93AEA91E724FFB57DE8FADDFE67EC43" bold="true" box="[151,407,1307,1333]" italics="true" pageId="6" pageNumber="7">Liolaemus morandae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20957EAE724FFB57CE1FADCFE0CEC42" box="[414,508,1306,1332]" pageId="6" pageNumber="7" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BF13683E724FFB57DE8FAFBFED3EC23" blockId="6.[151,508,1306,1365]" box="[151,291,1341,1365]" pageId="6" pageNumber="7">
(
<figureCitation id="13752A06E724FFB57DE1FAFBFEEBEC23" box="[158,283,1341,1365]" captionStart-0="FIGURE 3" captionStart-1="FIGURE 4" captionStart-2="FIGURE 5" captionStart-3="FIGURE 6" captionStartId-0="7.[151,250,1729,1752]" captionStartId-1="9.[151,250,1941,1964]" captionStartId-2="10.[151,250,1053,1076]" captionStartId-3="12.[151,250,937,960]" captionTargetBox-0="[225,1361,193,1707]" captionTargetBox-1="[231,1356,193,1919]" captionTargetBox-2="[157,1427,198,1017]" captionTargetBox-3="[151,1436,209,931]" captionTargetId-0="figure@7.[225,1362,193,1709]" captionTargetId-1="figure@9.[231,1356,193,1920]" captionTargetId-2="figure@10.[151,1436,193,1032]" captionTargetId-3="figure@12.[151,1436,209,933]" captionTargetPageId-0="7" captionTargetPageId-1="9" captionTargetPageId-2="10" captionTargetPageId-3="12" captionText-0="FIGURE 3. Upper, dorsal and ventral views of male holotype in life of Liolaemus morandae sp. nov. MLP. S 2626; lower, dorsal and ventral views of female (LJAMM-CNP 13020)." captionText-1="FIGURE 4. Type locality of Liolaemus morandae sp. nov., Patagonic phytogeographic province, del Golfo de San Jorge District (45 ° 41 10,6 ” S; 67 ° 53 49,9 ” W, 693 m)." captionText-2="FIGURE 5. Distribution map for species of the lineomaculatus group, with geographical localities sampled in this study superimposed on regional elevation (shading). Black circles: L. morandae sp. nov.; white circle: L. avilae sp. nov.; black squares: L. lineomaculatus; white square: L. kolengh; white triangles: L. hatcheri; black triangle: L. silvanae. Arrows indicate type localities. CH: Chubut province; SC: Santa Cruz province." captionText-3="FIGURE 6. Morphological comparison between Liolaemus morandae sp. nov. (A), L. lineomaculatus (B) and L. avilae sp. nov. (C). Notice differences in color pattern on the tails, size of blotches on the limbs and differences in vertebral and paravertebral lines. Dorsal scales (shown on the right) are less mucronated in L. lineomaculatus. Dorsal scales of hindlimbs are less carinated in L. morandae sp. nov. than in the other species, while they are more mucronated in L. avilae sp. nov." httpUri-0="https://zenodo.org/record/208133/files/figure.png" httpUri-1="https://zenodo.org/record/208134/files/figure.png" httpUri-2="https://zenodo.org/record/208135/files/figure.png" httpUri-3="https://zenodo.org/record/208136/files/figure.png" pageId="6" pageNumber="7">Figure 36</figureCitation>
)
</paragraph>
<paragraph id="8BF13683E724FFB57DE8FA46FE6AECC2" blockId="6.[151,1436,1408,1461]" pageId="6" pageNumber="7">
<bibRef id="52EEA275E724FFB57DE8FA46FE6AECC2" pageId="6" pageNumber="7" type="journal article">
<year id="FFE677D0E724FFB57DE8FA46FF3AECE3" box="[151,202,1408,1429]" pageId="6" pageNumber="7">
<emphasis id="B93AEA91E724FFB57DE8FA46FF3AECE3" bold="true" box="[151,202,1408,1429]" pageId="6" pageNumber="7">2001</emphasis>
</year>
,
<title id="F94ADDB5E724FFB57DA9FA46FAE2ECE0" box="[214,1298,1408,1430]" jopScore="0.48237443" pageId="6" pageNumber="7">
<emphasis id="B93AEA91E724FFB57DA9FA46FE17ECE0" box="[214,487,1408,1430]" italics="true" pageId="6" pageNumber="7">Liolaemus lineomaculatus,</emphasis>
Ibargüengoytía, N., Casalins, L., Schulte II, J.A., Amico, G.A. and Sympson, L
</title>
.,
<journalOrPublisher id="518F3C4FE724FFB5785FFA46FECAECC3" jopScore="1.4735439" pageId="6" pageNumber="7">
<emphasis id="B93AEA91E724FFB5785FFA46FECAECC3" italics="true" pageId="6" pageNumber="7">Herpetological Review</emphasis>
</journalOrPublisher>
,
<part id="FFEA9C1EE724FFB57C38FA59FE93ECC2" box="[327,355,1439,1460]" pageId="6" pageNumber="7" type="volume">32</part>
,
<pagination id="B287C877E724FFB57C12FA59FE66ECC2" box="[365,406,1439,1460]" pageId="6" pageNumber="7" type="pagination">120</pagination>
.
</bibRef>
</paragraph>
</subSubSection>
<subSubSection id="C3546508E724FFB57DE8FA1BFEE9EE36" pageId="6" pageNumber="7" type="materials_examined">
<paragraph id="8BF13683E724FFB57DE8FA1BFBCFEF49" blockId="6.[151,1437,1500,2037]" pageId="6" pageNumber="7">
<emphasis id="B93AEA91E724FFB57DE8FA1BFEFBEC82" bold="true" box="[151,267,1501,1525]" pageId="6" pageNumber="7">
<typeStatus id="54F58821E724FFB57DE8FA1BFEF4EC83" box="[151,260,1501,1525]" pageId="6" pageNumber="7" type="holotype">Holotype</typeStatus>
.
</emphasis>
<collectionCode id="ED5FAE46E724FFB57C6DFA1AFEBEEC82" box="[274,334,1500,1524]" httpUri="http://grbio.org/cool/66si-588n" name="Museo de La Plata" pageId="6" pageNumber="7">MLP</collectionCode>
.S 2626 (
<figureCitation id="13752A06E724FFB57CD3FA1AFDFDEC82" box="[428,525,1500,1524]" captionStart="FIGURE 3" captionStartId="7.[151,250,1729,1752]" captionTargetBox="[225,1361,193,1707]" captionTargetId="figure@7.[225,1362,193,1709]" captionTargetPageId="7" captionText="FIGURE 3. Upper, dorsal and ventral views of male holotype in life of Liolaemus morandae sp. nov. MLP. S 2626; lower, dorsal and ventral views of female (LJAMM-CNP 13020)." httpUri="https://zenodo.org/record/208133/files/figure.png" pageId="6" pageNumber="7">Figure 3</figureCitation>
), an adult male from Provincial Road 37,
<quantity id="4CB69B66E724FFB57E99FA1AFBB2EC82" box="[998,1090,1500,1524]" metricMagnitude="4" metricUnit="m" metricValue="2.28" pageId="6" pageNumber="7" unit="km" value="22.8">22.8 km</quantity>
SW junction National Road 3, Escalante department, Chubut province,
<collectingCountry id="F3597613E724FFB57F13F9C6FD2BEF6E" box="[620,731,1536,1560]" name="Argentina" pageId="6" pageNumber="7">Argentina</collectingCountry>
(
<geoCoordinate id="EE7A5044E724FFB57F91F9C6FC66EF6E" box="[750,918,1536,1560]" direction="south" orientation="latitude" pageId="6" pageNumber="7" precision="1" value="-45.68628">45°4110,6” S</geoCoordinate>
;
<geoCoordinate id="EE7A5044E724FFB57EDAF9C6FBA5EF6E" box="[933,1109,1536,1560]" direction="west" orientation="longitude" pageId="6" pageNumber="7" precision="1" value="-67.897194">67°5349,9” W</geoCoordinate>
,
<quantity id="4CB69B66E724FFB5791CF9C6FB5FEF6E" box="[1123,1199,1536,1560]" metricMagnitude="2" metricUnit="m" metricValue="6.93" pageId="6" pageNumber="7" unit="m" value="693.0">693 m</quantity>
) (
<figureCitation id="13752A06E724FFB579B6F9C6FAC9EF6E" box="[1225,1337,1536,1560]" captionStart="FIGURE 4" captionStartId="9.[151,250,1941,1964]" captionTargetBox="[231,1356,193,1919]" captionTargetId="figure@9.[231,1356,193,1920]" captionTargetPageId="9" captionText="FIGURE 4. Type locality of Liolaemus morandae sp. nov., Patagonic phytogeographic province, del Golfo de San Jorge District (45 ° 41 10,6 ” S; 67 ° 53 49,9 ” W, 693 m)." httpUri="https://zenodo.org/record/208134/files/figure.png" pageId="6" pageNumber="7">Figures 4</figureCitation>
,
<figureCitation id="13752A06E724FFB57837F9C6FAA8EF6E" box="[1352,1368,1536,1560]" captionStart="FIGURE 5" captionStartId="10.[151,250,1053,1076]" captionTargetBox="[157,1427,198,1017]" captionTargetId="figure@10.[151,1436,193,1032]" captionTargetPageId="10" captionText="FIGURE 5. Distribution map for species of the lineomaculatus group, with geographical localities sampled in this study superimposed on regional elevation (shading). Black circles: L. morandae sp. nov.; white circle: L. avilae sp. nov.; black squares: L. lineomaculatus; white square: L. kolengh; white triangles: L. hatcheri; black triangle: L. silvanae. Arrows indicate type localities. CH: Chubut province; SC: Santa Cruz province." httpUri="https://zenodo.org/record/208135/files/figure.png" pageId="6" pageNumber="7">5</figureCitation>
); L.J. Avila, M. Kozykariski, M.F. Breitman and R. Martinez collectors,
<date id="FFF01043E724FFB57EFFF9E1FBCBEF49" box="[896,1083,1572,1599]" pageId="6" pageNumber="7" value="2010-03-12">12th March 2010</date>
.
</paragraph>
<paragraph id="8BF13683E724FFB57DBAF98DFEE9EE36" blockId="6.[151,1437,1500,2037]" pageId="6" pageNumber="7">
<emphasis id="B93AEA91E724FFB57DBAF98DFEB3EF14" bold="true" box="[197,323,1611,1635]" pageId="6" pageNumber="7">
<typeStatus id="54F58821E724FFB57DBAF98DFECDEF15" box="[197,317,1611,1635]" pageId="6" pageNumber="7" type="paratype">Paratypes</typeStatus>
.
</emphasis>
<collectionCode id="ED5FAE46E724FFB57C34F98DFE1DEF15" box="[331,493,1611,1635]" httpUri="http://grbio.org/cool/yjrq-91r7" name="Centro Nacional Patagonico (CENPAT-CONICET)" pageId="6" pageNumber="7">LJAMM-CNP</collectionCode>
13020, adult female and
<collectionCode id="ED5FAE46E724FFB57E71F98DFC40EF15" box="[782,944,1611,1635]" httpUri="http://grbio.org/cool/yjrq-91r7" name="Centro Nacional Patagonico (CENPAT-CONICET)" pageId="6" pageNumber="7">LJAMM-CNP</collectionCode>
13021, juvenile; same locality as
<typeStatus id="54F58821E724FFB5784CF98DFA69EF15" box="[1331,1433,1611,1635]" pageId="6" pageNumber="7" type="holotype">holotype</typeStatus>
. LJAM-CNP 9677-9679, adult females and
<collectionCode id="ED5FAE46E724FFB57FFDF9A9FCD3EFF1" box="[642,803,1647,1671]" httpUri="http://grbio.org/cool/yjrq-91r7" name="Centro Nacional Patagonico (CENPAT-CONICET)" pageId="6" pageNumber="7">LJAMM-CNP</collectionCode>
9680, juvenile; from Holdich station, Escalante department, Chubut province,
<collectingCountry id="F3597613E724FFB57CDAF955FDE4EFDD" box="[421,532,1683,1707]" name="Argentina" pageId="6" pageNumber="7">Argentina</collectingCountry>
(
<geoCoordinate id="EE7A5044E724FFB57F5BF955FD35EFDD" box="[548,709,1683,1707]" direction="south" orientation="latitude" pageId="6" pageNumber="7" precision="1" value="-45.966694">45º5800.1” S</geoCoordinate>
;
<geoCoordinate id="EE7A5044E724FFB57FAEF955FC89EFDD" box="[721,889,1683,1707]" direction="west" orientation="longitude" pageId="6" pageNumber="7" precision="1" value="-68.19967">68º1158.8” W</geoCoordinate>
,
<quantity id="4CB69B66E724FFB57EFBF955FC3EEFDD" box="[900,974,1683,1707]" metricMagnitude="2" metricUnit="m" metricValue="7.609999999999999" pageId="6" pageNumber="7" unit="m" value="761.0">761 m</quantity>
); L.J. Avila, C.H.F. Pérez, M.F. Breitman and N. Feltrin collectors,
<date id="FFF01043E724FFB57CBFF97CFD7EEFA4" box="[448,654,1719,1746]" pageId="6" pageNumber="7" value="2008-02-02">2nd February 2008</date>
.
<collectionCode id="ED5FAE46E724FFB57FE5F97CFCCCEFA4" box="[666,828,1722,1746]" httpUri="http://grbio.org/cool/yjrq-91r7" name="Centro Nacional Patagonico (CENPAT-CONICET)" pageId="6" pageNumber="7">LJAMM-CNP</collectionCode>
10201, adult male and LJAM-CNP 10202, juvenile; from Provincial Road 37,
<quantity id="4CB69B66E724FFB57CC1F918FDFDEF80" box="[446,525,1758,1782]" metricMagnitude="3" metricUnit="m" metricValue="2.5" pageId="6" pageNumber="7" unit="km" value="2.5">2.5 km</quantity>
W junction National Road 3, Escalante department, Chubut province,
<collectingCountry id="F3597613E724FFB57852F918FA6CEF80" box="[1325,1436,1758,1782]" name="Argentina" pageId="6" pageNumber="7">Argentina</collectingCountry>
(
<geoCoordinate id="EE7A5044E724FFB57DE0F8C3FEB1EE6B" box="[159,321,1797,1821]" direction="south" orientation="latitude" pageId="6" pageNumber="7" precision="1" value="-45.62872">45º3743.4” S</geoCoordinate>
;
<geoCoordinate id="EE7A5044E724FFB57C32F8C3FE07EE6B" box="[333,503,1797,1821]" direction="west" orientation="longitude" pageId="6" pageNumber="7" precision="1" value="-67.68433">67º4103.6” W</geoCoordinate>
,
<quantity id="4CB69B66E724FFB57F7DF8C3FDBCEE6B" box="[514,588,1797,1821]" metricMagnitude="2" metricUnit="m" metricValue="6.37" pageId="6" pageNumber="7" unit="m" value="637.0">637 m</quantity>
); L.J. Avila, C.H.F. Pérez, M.F. Breitman and N. Feltrin collectors,
<date id="FFF01043E724FFB5783CF8C3FEE5EE36" pageId="6" pageNumber="7" value="2008-02-02">2 nd February 2008</date>
.
</paragraph>
</subSubSection>
<subSubSection id="C3546508E724FFBB7DBAF88AFB73EE72" lastPageId="8" lastPageNumber="9" pageId="6" pageNumber="7" type="diagnosis">
<paragraph id="8BF13683E724FFB57DBAF88AFEE8EE83" blockId="6.[151,1437,1500,2037]" pageId="6" pageNumber="7">
<emphasis id="B93AEA91E724FFB57DBAF88AFECFEE12" bold="true" box="[197,319,1868,1893]" pageId="6" pageNumber="7">Diagnosis.</emphasis>
<taxonomicName id="4C4E4D00E724FFB57C39F88BFDC5EE13" box="[326,565,1869,1893]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="morandae" status="sp. nov.">
<emphasis id="B93AEA91E724FFB57C39F88BFDC5EE13" box="[326,565,1869,1893]" italics="true" pageId="6" pageNumber="7">Liolaemus morandae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20957EAE724FFB57F42F88BFD67EE13" box="[573,663,1869,1893]" pageId="6" pageNumber="7" rank="species">
<emphasis id="B93AEA91E724FFB57F42F88BFD67EE13" bold="true" box="[573,663,1869,1893]" pageId="6" pageNumber="7">sp. nov.</emphasis>
</taxonomicNameLabel>
is a member of the
<taxonomicName id="4C4E4D00E724FFB57E04F888FBB4EE13" box="[891,1092,1869,1893]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="lineomaculatus">
<emphasis id="B93AEA91E724FFB57E04F888FBB4EE13" box="[891,1092,1869,1893]" italics="true" pageId="6" pageNumber="7">L. lineomaculatus</emphasis>
</taxonomicName>
section, included in the
<taxonomicName id="4C4E4D00E724FFB57824F88BFEFAEEFF" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="lineomaculatus">
<emphasis id="B93AEA91E724FFB57824F88BFEFAEEFF" italics="true" pageId="6" pageNumber="7">lineomaculatus</emphasis>
</taxonomicName>
group, and is characterized by the absence of precloacal pores in both sexes, and presence of dorsal trifid scales (
<bibRefCitation id="EFDF4B72E724FFB57D95F853FE6DEEDB" author="Etheridge" box="[234,413,1941,1965]" pageId="6" pageNumber="7" refString="Etheridge, R. (1995) Redescription of Ctenoblepharys adspersa Tschudi, 1845, and the taxonomy of Liolaeminae (Reptilia: Squamata: Tropiduridae). American Museum Novitates, 3142, 1 - 34." type="journal article" year="1995">Etheridge 1995</bibRefCitation>
). Molecular evidence includes this species in the
<taxonomicName id="4C4E4D00E724FFB57EABF853FB8FEEDB" box="[980,1151,1941,1965]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="lineomaculatus">
<emphasis id="B93AEA91E724FFB57EABF853FB8FEEDB" box="[980,1151,1941,1965]" italics="true" pageId="6" pageNumber="7">lineomaculatus</emphasis>
</taxonomicName>
group, closely related to the clade (
<emphasis id="B93AEA91E724FFB57C74F87CFEB8EEA7" box="[267,328,1978,2001]" italics="true" pageId="6" pageNumber="7">L. sp.</emphasis>
2 +
<taxonomicName id="4C4E4D00E724FFB57C05F87CFDB2EEA7" box="[378,578,1977,2001]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="lineomaculatus">
<emphasis id="B93AEA91E724FFB57C05F87CFDB2EEA7" box="[378,578,1977,2001]" italics="true" pageId="6" pageNumber="7">L. lineomaculatus</emphasis>
</taxonomicName>
)
<figureCitation id="13752A06E724FFB57F2CF87EFD42EEA6" box="[595,690,1976,2000]" captionStart="FIGURE 1" captionStartId="4.[151,250,808,831]" captionTargetBox="[170,1409,190,807]" captionTargetId="figure@4.[151,1436,190,828]" captionTargetPageId="4" captionText="FIGURE 1. Relationships between Liolaemus morandae sp. nov. and L. avilae sp. nov. within the L. lineomaculatus section, and selected species of the subgenus Eulaemus and Liolaemus sensu stricto. On the left, phylogenetic relationships recovered with Bayesian inference using seven nuclear genes (Cmos, ACM 4 tg, PRLR, LDA 8 F, LDA 1 D, LDA 9 C and LDA 9 E, 4180 bp total alignment length), bold branches represent well supported nodes (Pp&gt; 0.95). On the right, the Bayesian tree (modified from Breitman et al. 2011) represents a concatenated analyses including the seven nuclear genes plus two mitochondrial genes (12 S and cyt- b; 5865 bp) and summarizes information from MP and ML methods. Nodes with high support from three methods (MP, jackknife and ML bootstrap&gt; 0.70; Bayesian Pp&gt; 0.95) are identified by bold branches; solid circles show nodes with weak MP support and open circles nodes with weak MP and ML support." httpUri="https://zenodo.org/record/208131/files/figure.png" pageId="6" pageNumber="7">Figure 1</figureCitation>
. All the following differences are summarized in Tables 1 to 4 and in
<figureCitation id="13752A06E724FFB57DCCF81BFEE4EE83" box="[179,276,2013,2037]" captionStart="FIGURE 6" captionStartId="12.[151,250,937,960]" captionTargetBox="[151,1436,209,931]" captionTargetId="figure@12.[151,1436,209,933]" captionTargetPageId="12" captionText="FIGURE 6. Morphological comparison between Liolaemus morandae sp. nov. (A), L. lineomaculatus (B) and L. avilae sp. nov. (C). Notice differences in color pattern on the tails, size of blotches on the limbs and differences in vertebral and paravertebral lines. Dorsal scales (shown on the right) are less mucronated in L. lineomaculatus. Dorsal scales of hindlimbs are less carinated in L. morandae sp. nov. than in the other species, while they are more mucronated in L. avilae sp. nov." httpUri="https://zenodo.org/record/208136/files/figure.png" pageId="6" pageNumber="7">Figure 6</figureCitation>
.
</paragraph>
<caption id="DF31660BE725FFB47DE8F907FD38EF81" httpUri="https://zenodo.org/record/208133/files/figure.png" pageId="7" pageNumber="8" targetBox="[225,1361,193,1707]" targetPageId="7">
<paragraph id="8BF13683E725FFB47DE8F907FD38EF81" blockId="7.[151,1436,1729,1783]" pageId="7" pageNumber="8">
<emphasis id="B93AEA91E725FFB47DE8F907FEE2EFA1" bold="true" box="[151,274,1729,1752]" pageId="7" pageNumber="8">FIGURE 3.</emphasis>
Upper, dorsal and ventral views of male holotype in life of
<taxonomicName id="4C4E4D00E725FFB47E11F904FBB5EFAE" box="[878,1093,1730,1752]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="morandae" status="sp. nov.">
<emphasis id="B93AEA91E725FFB47E11F904FBB5EFAE" box="[878,1093,1730,1752]" italics="true" pageId="7" pageNumber="8">Liolaemus morandae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20957EAE725FFB47934F904FB6BEFA1" box="[1099,1179,1730,1751]" pageId="7" pageNumber="8" rank="species">
<emphasis id="B93AEA91E725FFB47934F904FB6BEFA1" bold="true" box="[1099,1179,1730,1751]" pageId="7" pageNumber="8">sp. nov.</emphasis>
</taxonomicNameLabel>
MLP.S 2626; lower, dorsal and ventral views of female (LJAMM-CNP 13020).
</paragraph>
</caption>
<paragraph id="8BF13683E725FFB47DBAF8D8FA6CEE9C" blockId="7.[151,1436,1822,2027]" pageId="7" pageNumber="8">
Relative to
<taxonomicName id="4C4E4D00E725FFB47C39F8D9FDFFEE40" box="[326,527,1822,1846]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="lineomaculatus">
<emphasis id="B93AEA91E725FFB47C39F8D9FDFFEE40" box="[326,527,1822,1846]" italics="true" pageId="7" pageNumber="8">L. lineomaculatus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C4E4D00E725FFB47F62F8D9FD5DEE40" box="[541,685,1822,1846]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="morandae" status="sp. nov.">
<emphasis id="B93AEA91E725FFB47F62F8D9FD5DEE40" box="[541,685,1822,1846]" italics="true" pageId="7" pageNumber="8">L. morandae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20957EAE725FFB47FCBF8D8FCFEEE40" box="[692,782,1822,1846]" pageId="7" pageNumber="8" rank="species">
<emphasis id="B93AEA91E725FFB47FCBF8D8FCFEEE40" bold="true" box="[692,782,1822,1846]" pageId="7" pageNumber="8">sp. nov.</emphasis>
</taxonomicNameLabel>
has fewer dorsal scales (4757, X = 51.67 vs. 5268, X = 58.41;
<emphasis id="B93AEA91E725FFB47D9BF885FF02EE2C" box="[228,242,1859,1882]" italics="true" pageId="7" pageNumber="8">p</emphasis>
&lt;0.0001), fewer third finger lamellae (1316, X = 15.33 vs. 1419, X = 16,18;
<emphasis id="B93AEA91E725FFB4790BF885FB72EE2C" box="[1140,1154,1859,1882]" italics="true" pageId="7" pageNumber="8">p</emphasis>
&lt;0.0001), shorter fourth toe length (13.416.8, X = 14.9 vs. 12.616.1, X = 14.09;
<emphasis id="B93AEA91E725FFB47E54F8A1FCC9EE08" box="[811,825,1895,1918]" italics="true" pageId="7" pageNumber="8">p</emphasis>
&lt;0.0001) and a larger eye (eye height: 1.72.27, X = 1.99 vs. 1.52.21, X = 1.84, eye length: 2.913.42, X = 3.15 vs. 2.353.3, X = 2.83;
<emphasis id="B93AEA91E725FFB47938F84DFBA5EED4" box="[1095,1109,1931,1954]" italics="true" pageId="7" pageNumber="8">p</emphasis>
&lt;
<quantity id="4CB69B66E725FFB4790DF84CFB2CEED4" box="[1138,1244,1930,1954]" metricMagnitude="-6" metricUnit="m" metricValue="2.54" pageId="7" pageNumber="8" unit="in" value="1.0E-4">0.0001 in</quantity>
both cases).
<taxonomicName id="4C4E4D00E725FFB47812F84DFE90EEB0" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="morandae" status="sp. nov.">
<emphasis id="B93AEA91E725FFB47812F84DFE90EEB0" italics="true" pageId="7" pageNumber="8">Liolaemus morandae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20957EAE725FFB47C16F868FE34EEB0" box="[361,452,1966,1990]" pageId="7" pageNumber="8" rank="species">
<emphasis id="B93AEA91E725FFB47C16F868FE34EEB0" bold="true" box="[361,452,1966,1990]" pageId="7" pageNumber="8">sp. nov.</emphasis>
</taxonomicNameLabel>
has more dorsal blotches than
<emphasis id="B93AEA91E725FFB47E57F869FC09EEB0" box="[808,1017,1966,1990]" italics="true" pageId="7" pageNumber="8">
<taxonomicName id="4C4E4D00E725FFB47E57F869FC05EEB0" box="[808,1013,1966,1990]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="lineomaculatus">L. lineomaculatus</taxonomicName>
,
</emphasis>
and the blotches in
<taxonomicName id="4C4E4D00E725FFB4799EF869FA82EEB0" box="[1249,1394,1966,1990]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="morandae" status="sp. nov.">
<emphasis id="B93AEA91E725FFB4799EF869FA82EEB0" box="[1249,1394,1966,1990]" italics="true" pageId="7" pageNumber="8">L. morandae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20957EAE725FFB47805F868FF38EE9C" pageId="7" pageNumber="8" rank="species">
<emphasis id="B93AEA91E725FFB47805F868FF38EE9C" bold="true" pageId="7" pageNumber="8">sp. nov.</emphasis>
</taxonomicNameLabel>
have (in 50% of the cases) a yellow-orange coloration while
<taxonomicName id="4C4E4D00E725FFB47E05F815FBB3EE9C" box="[890,1091,2002,2026]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="lineomaculatus">
<emphasis id="B93AEA91E725FFB47E05F815FBB3EE9C" box="[890,1091,2002,2026]" italics="true" pageId="7" pageNumber="8">L. lineomaculatus</emphasis>
</taxonomicName>
blotches have dark gray color.
</paragraph>
<paragraph id="8BF13683E72AFFBB7DE8FF5EFB92E9A2" blockId="8.[151,1437,151,2012]" pageId="8" pageNumber="9">
In general, dorsal scales of
<taxonomicName id="4C4E4D00E72AFFBB7CB4FF5FFDAAE9C6" box="[459,602,152,176]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="morandae" status="sp. nov.">
<emphasis id="B93AEA91E72AFFBB7CB4FF5FFDAAE9C6" box="[459,602,152,176]" italics="true" pageId="8" pageNumber="9">L. morandae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20957EAE72AFFBB7F1EFF5EFD4BE9C6" box="[609,699,152,176]" pageId="8" pageNumber="9" rank="species">
<emphasis id="B93AEA91E72AFFBB7F1EFF5EFD4BE9C6" bold="true" box="[609,699,152,176]" pageId="8" pageNumber="9">sp. nov.</emphasis>
</taxonomicNameLabel>
are more mucronated than those of
<emphasis id="B93AEA91E72AFFBB792FFF5FFAD0E9C6" box="[1104,1312,152,176]" italics="true" pageId="8" pageNumber="9">
<taxonomicName id="4C4E4D00E72AFFBB792FFF5FFAECE9C6" box="[1104,1308,152,176]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="lineomaculatus">L. lineomaculatus</taxonomicName>
,
</emphasis>
and dorsal leg scales of
<taxonomicName id="4C4E4D00E72AFFBB7C57FF7BFE47E9A2" box="[296,439,188,212]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="morandae" status="sp. nov.">
<emphasis id="B93AEA91E72AFFBB7C57FF7BFE47E9A2" box="[296,439,188,212]" italics="true" pageId="8" pageNumber="9">L. morandae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20957EAE72AFFBB7CC1FF7AFDE7E9A2" box="[446,535,188,212]" pageId="8" pageNumber="9" rank="species">
<emphasis id="B93AEA91E72AFFBB7CC1FF7AFDE7E9A2" bold="true" box="[446,535,188,212]" pageId="8" pageNumber="9">sp. nov.</emphasis>
</taxonomicNameLabel>
are less carinated with respect to
<taxonomicName id="4C4E4D00E72AFFBB7EEDFF7BFBABE9A2" box="[914,1115,188,212]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="lineomaculatus">
<emphasis id="B93AEA91E72AFFBB7EEDFF7BFBABE9A2" box="[914,1115,188,212]" italics="true" pageId="8" pageNumber="9">L. lineomaculatus</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BF13683E72AFFBB7DBAFF19FC50EB99" blockId="8.[151,1437,151,2012]" pageId="8" pageNumber="9">
Compared to
<emphasis id="B93AEA91E72AFFBB7C1EFF27FD8EE98E" box="[353,638,224,248]" italics="true" pageId="8" pageNumber="9">
<taxonomicName id="4C4E4D00E72AFFBB7C1EFF27FE10E98E" box="[353,480,224,248]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="hatcheri">L. hatcheri</taxonomicName>
,
<taxonomicName id="4C4E4D00E72AFFBB7C92FF27FD8EE98E" box="[493,638,224,248]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="morandae" status="sp. nov.">L. morandae</taxonomicName>
</emphasis>
<taxonomicNameLabel id="A20957EAE72AFFBB7FF8FF26FD12E98E" box="[647,738,224,248]" pageId="8" pageNumber="9" rank="species">
<emphasis id="B93AEA91E72AFFBB7FF8FF26FD12E98E" bold="true" box="[647,738,224,248]" pageId="8" pageNumber="9">sp. nov.</emphasis>
</taxonomicNameLabel>
has higher number of midbody scales (5563, X = 59.83 vs. 4357, X = 48.65
<emphasis id="B93AEA91E72AFFBB7C14FEC3FE89E86A" box="[363,377,261,284]" italics="true" pageId="8" pageNumber="9">p</emphasis>
&lt;0.0001), larger number of scales from occiput to rump (4757, X = 51.67 vs. 4355, X = 48.75;
<emphasis id="B93AEA91E72AFFBB7D9BFEEFFF02E836" box="[228,242,297,320]" italics="true" pageId="8" pageNumber="9">p</emphasis>
&lt;0.0001), more ventral scales (7985, X = 81.33 vs. 6274, X = 67.8;
<emphasis id="B93AEA91E72AFFBB796EFEEFFBEFE836" box="[1041,1055,297,320]" italics="true" pageId="8" pageNumber="9">p</emphasis>
&lt;0.0001), more fourth toe lamellae (1823, X = 21.17 vs. 1822, X = 19.45;
<emphasis id="B93AEA91E72AFFBB7FE8FE8BFD55E812" box="[663,677,333,356]" italics="true" pageId="8" pageNumber="9">p</emphasis>
&lt;0.0001), smaller body size (snout-vent length: 5061, X = 55.67 vs. 5569, X = 61.89, axilla-groin distance: 19.730.8, X = 25.9 vs. 19.535.5, X = 28.95;
<emphasis id="B93AEA91E72AFFBB79BDFEB7FB20E8FE" box="[1218,1232,369,392]" italics="true" pageId="8" pageNumber="9">p</emphasis>
&lt;
<quantity id="4CB69B66E72AFFBB798CFEA9FA90E8F1" box="[1267,1376,367,391]" metricMagnitude="-6" metricUnit="m" metricValue="2.54" pageId="8" pageNumber="9" unit="in" value="1.0E-4">0.0001 in</quantity>
both cases), shorter forelimbs (elbow-wrist length: 4.695.56, X = 5.32 vs. 5.677.33, X = 6.44, third finger length: 6.699.14, X = 7.83 vs. 8.510.57, X = 9.15;
<emphasis id="B93AEA91E72AFFBB7FEBFE7FFD52E8A6" box="[660,674,441,464]" italics="true" pageId="8" pageNumber="9">p</emphasis>
&lt;
<quantity id="4CB69B66E72AFFBB7FBFFE71FCDAE8B9" box="[704,810,439,463]" metricMagnitude="-6" metricUnit="m" metricValue="2.54" pageId="8" pageNumber="9" unit="in" value="1.0E-4">0.0001 in</quantity>
both cases), smaller head (head width: 9.0810.58, X = 9.84 vs. 9.713.05, X = 11.22, head length: 10.9613.06, X = 11.96 vs. 11.4314.65, X = 12.97;
<emphasis id="B93AEA91E72AFFBB79B1FE1BFB2CE882" box="[1230,1244,477,500]" italics="true" pageId="8" pageNumber="9">p</emphasis>
&lt;
<quantity id="4CB69B66E72AFFBB7985FE1AFA93E882" box="[1274,1379,476,500]" metricMagnitude="-6" metricUnit="m" metricValue="2.54" pageId="8" pageNumber="9" unit="in" value="1.0E-4">0.0001 in</quantity>
both cases), and smaller auditory meatus (auditory meatus height: 1.62.44, X = 1.95 vs. 1.852.61, X = 2.15; auditory meatus length: 0.861.81, X = 1.27 vs. 1.382.16, X = 1.77;
<emphasis id="B93AEA91E72AFFBB7E2DFDE3FC90EB4A" box="[850,864,549,572]" italics="true" pageId="8" pageNumber="9">p</emphasis>
&lt;
<quantity id="4CB69B66E72AFFBB7EFEFDE2FC1CEB4A" box="[897,1004,548,572]" metricMagnitude="-6" metricUnit="m" metricValue="2.54" pageId="8" pageNumber="9" unit="in" value="1.0E-4">0.0001 in</quantity>
both cases).
<taxonomicName id="4C4E4D00E72AFFBB79FCFDE2FA82EB4A" box="[1155,1394,548,572]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="morandae" status="sp. nov.">
<emphasis id="B93AEA91E72AFFBB79FCFDE2FA82EB4A" box="[1155,1394,548,572]" italics="true" pageId="8" pageNumber="9">Liolaemus morandae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20957EAE72AFFBB7805FDE2FF38EB16" pageId="8" pageNumber="9" rank="species">
<emphasis id="B93AEA91E72AFFBB7805FDE2FF38EB16" bold="true" pageId="8" pageNumber="9">sp. nov.</emphasis>
</taxonomicNameLabel>
has more dorsal blotches and better defined vertebral and paravertebral lines than
<emphasis id="B93AEA91E72AFFBB7907FD8FFB0DEB16" box="[1144,1277,584,608]" italics="true" pageId="8" pageNumber="9">
<taxonomicName id="4C4E4D00E72AFFBB7907FD8FFB07EB16" box="[1144,1271,584,608]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="hatcheri">L. hatcheri</taxonomicName>
;
</emphasis>
ventral melanism is more pronounced in
<emphasis id="B93AEA91E72AFFBB7CA3FDABFDAEEBF2" box="[476,606,620,644]" italics="true" pageId="8" pageNumber="9">
<taxonomicName id="4C4E4D00E72AFFBB7CA3FDABFDAAEBF2" box="[476,602,620,644]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="hatcheri">L. hatcheri</taxonomicName>
,
</emphasis>
while
<taxonomicName id="4C4E4D00E72AFFBB7FD3FDABFCCDEBF2" box="[684,829,620,644]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="morandae" status="sp. nov.">
<emphasis id="B93AEA91E72AFFBB7FD3FDABFCCDEBF2" box="[684,829,620,644]" italics="true" pageId="8" pageNumber="9">L. morandae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20957EAE72AFFBB7E3AFDAAFC6FEBF2" box="[837,927,620,644]" pageId="8" pageNumber="9" rank="species">
<emphasis id="B93AEA91E72AFFBB7E3AFDAAFC6FEBF2" bold="true" box="[837,927,620,644]" pageId="8" pageNumber="9">sp. nov.</emphasis>
</taxonomicNameLabel>
has fewer melanophores. Dorsal scales of
<taxonomicName id="4C4E4D00E72AFFBB78FAFDABFEF9EBDE" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="morandae" status="sp. nov.">
<emphasis id="B93AEA91E72AFFBB78FAFDABFEF9EBDE" italics="true" pageId="8" pageNumber="9">L. morandae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20957EAE72AFFBB7C6EFD56FE9CEBDE" box="[273,364,656,680]" pageId="8" pageNumber="9" rank="species">
<emphasis id="B93AEA91E72AFFBB7C6EFD56FE9CEBDE" bold="true" box="[273,364,656,680]" pageId="8" pageNumber="9">sp. nov.</emphasis>
</taxonomicNameLabel>
are less “bristly” than those of
<emphasis id="B93AEA91E72AFFBB7FA7FD57FCACEBDE" box="[728,860,656,680]" italics="true" pageId="8" pageNumber="9">
<taxonomicName id="4C4E4D00E72AFFBB7FA7FD57FCA8EBDE" box="[728,856,656,680]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="hatcheri">L. hatcheri</taxonomicName>
.
</emphasis>
Dorsal and ventral limb scales are non-mucronate and carinate in
<taxonomicName id="4C4E4D00E72AFFBB7C36FD73FE2BEBBA" box="[329,475,692,716]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="morandae" status="sp. nov.">
<emphasis id="B93AEA91E72AFFBB7C36FD73FE2BEBBA" box="[329,475,692,716]" italics="true" pageId="8" pageNumber="9">L. morandae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20957EAE72AFFBB7C9BFD72FDB0EBBA" box="[484,576,692,716]" pageId="8" pageNumber="9" rank="species">
<emphasis id="B93AEA91E72AFFBB7C9BFD72FDB0EBBA" bold="true" box="[484,576,692,716]" pageId="8" pageNumber="9">sp. nov.</emphasis>
</taxonomicNameLabel>
while in
<taxonomicName id="4C4E4D00E72AFFBB7FD0FD73FCDCEBBA" box="[687,812,692,716]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="hatcheri">
<emphasis id="B93AEA91E72AFFBB7FD0FD73FCDCEBBA" box="[687,812,692,716]" italics="true" pageId="8" pageNumber="9">L. hatcheri</emphasis>
</taxonomicName>
they are mucronate and non-carinate. Belly scales are round-shaped in
<taxonomicName id="4C4E4D00E72AFFBB7C2CFD1FFE13EB86" box="[339,483,728,752]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="morandae" status="sp. nov.">
<emphasis id="B93AEA91E72AFFBB7C2CFD1FFE13EB86" box="[339,483,728,752]" italics="true" pageId="8" pageNumber="9">L. morandae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20957EAE72AFFBB7C96FD1EFDB3EB86" box="[489,579,728,752]" pageId="8" pageNumber="9" rank="species">
<emphasis id="B93AEA91E72AFFBB7C96FD1EFDB3EB86" bold="true" box="[489,579,728,752]" pageId="8" pageNumber="9">sp. nov.</emphasis>
</taxonomicNameLabel>
and rhomboidal in
<taxonomicName id="4C4E4D00E72AFFBB7E61FD1FFC69EB86" box="[798,921,728,752]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="hatcheri">
<emphasis id="B93AEA91E72AFFBB7E61FD1FFC69EB86" box="[798,921,728,752]" italics="true" pageId="8" pageNumber="9">L. hatcheri</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BF13683E72AFFBB7DBAFD3AFBCBEDB2" blockId="8.[151,1437,151,2012]" pageId="8" pageNumber="9">
<taxonomicName id="4C4E4D00E72AFFBB7DBAFD3AFE43EA62" box="[197,435,764,788]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="morandae" status="sp. nov.">
<emphasis id="B93AEA91E72AFFBB7DBAFD3AFE43EA62" box="[197,435,764,788]" italics="true" pageId="8" pageNumber="9">Liolaemus morandae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20957EAE72AFFBB7CC5FD3AFDE4EA62" box="[442,532,764,788]" pageId="8" pageNumber="9" rank="species">
<emphasis id="B93AEA91E72AFFBB7CC5FD3AFDE4EA62" bold="true" box="[442,532,764,788]" pageId="8" pageNumber="9">sp. nov.</emphasis>
</taxonomicNameLabel>
differs from
<taxonomicName id="4C4E4D00E72AFFBB7FD6FD3BFCD0EA62" box="[681,800,764,788]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="kolengh">
<emphasis id="B93AEA91E72AFFBB7FD6FD3BFCD0EA62" box="[681,800,764,788]" italics="true" pageId="8" pageNumber="9">L. kolengh</emphasis>
</taxonomicName>
in the following traits: larger number of midbody scales (5563, X = 59.83 vs. 4455, X = 50.55;
<emphasis id="B93AEA91E72AFFBB7F0DFCE7FD70EA4E" box="[626,640,801,824]" italics="true" pageId="8" pageNumber="9">p</emphasis>
&lt;0.0001), larger number of scales from occiput to rump (4757, X = 51.67 vs. 4454, X = 49.35;
<emphasis id="B93AEA91E72AFFBB7CA9FC83FE14EA2A" box="[470,484,837,860]" italics="true" pageId="8" pageNumber="9">p</emphasis>
&lt;0.0001), more ventral scales (7985, X = 81.33 vs. 6176, X = 68.58;
<emphasis id="B93AEA91E72AFFBB786CFC83FAD1EA2A" box="[1299,1313,837,860]" italics="true" pageId="8" pageNumber="9">p</emphasis>
&lt;0.0001), larger number of fourth toe lamellae (1823, X = 21.17 vs. 1622, X = 18.74;
<emphasis id="B93AEA91E72AFFBB7943FCAFFBBAEAF6" box="[1084,1098,873,896]" italics="true" pageId="8" pageNumber="9">p</emphasis>
&lt;0.0001), shorter forelimbs (elbow-wrist length: 4.695.56, X = 5.32 vs. 4.67.03, X = 6.16, third finger length: 6.699.14, X = 7.83 vs. 8.31 11.12, X = 9.65;
<emphasis id="B93AEA91E72AFFBB7C2CFC77FE91EABE" box="[339,353,945,968]" italics="true" pageId="8" pageNumber="9">p</emphasis>
&lt;
<quantity id="4CB69B66E72AFFBB7C01FC69FE18EAB1" box="[382,488,943,967]" metricMagnitude="-6" metricUnit="m" metricValue="2.54" pageId="8" pageNumber="9" unit="in" value="1.0E-4">0.0001 in</quantity>
both cases), smaller auditory meatus (auditory meatus height: 1.62.44, X = 1.95 vs. 1.632.95, X = 2.22, auditory meatus length: 0.861.81, X = 1.27 vs. 1.222.24, X = 1.67;
<emphasis id="B93AEA91E72AFFBB79BDFC13FB20EA9A" box="[1218,1232,981,1004]" italics="true" pageId="8" pageNumber="9">p</emphasis>
&lt;
<quantity id="4CB69B66E72AFFBB798BFC12FA91EA9A" box="[1268,1377,980,1004]" metricMagnitude="-6" metricUnit="m" metricValue="2.54" pageId="8" pageNumber="9" unit="in" value="1.0E-4">0.0001 in</quantity>
both cases) and longer tail (6179, X = 70.33 vs. 4770, X = 60.52;
<emphasis id="B93AEA91E72AFFBB7E2AFC3FFC93ED66" box="[853,867,1017,1040]" italics="true" pageId="8" pageNumber="9">p</emphasis>
= 0.0019).
<taxonomicName id="4C4E4D00E72AFFBB7E9BFC3EFB21ED66" box="[996,1233,1016,1040]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="morandae" status="sp. nov.">
<emphasis id="B93AEA91E72AFFBB7E9BFC3EFB21ED66" box="[996,1233,1016,1040]" italics="true" pageId="8" pageNumber="9">Liolaemus morandae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20957EAE72AFFBB79A8FC3EFAC0ED66" box="[1239,1328,1016,1040]" pageId="8" pageNumber="9" rank="species">
<emphasis id="B93AEA91E72AFFBB79A8FC3EFAC0ED66" bold="true" box="[1239,1328,1016,1040]" pageId="8" pageNumber="9">sp. nov.</emphasis>
</taxonomicNameLabel>
has more dorsal blotches and better defined vertebral and paravertebral lines than
<emphasis id="B93AEA91E72AFFBB7EC3FBDBFBCBED42" box="[956,1083,1052,1076]" italics="true" pageId="8" pageNumber="9">
<taxonomicName id="4C4E4D00E72AFFBB7EC3FBDBFBC4ED42" box="[956,1076,1052,1076]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="kolengh">L. kolengh</taxonomicName>
;
</emphasis>
ventral melanism is stronger in
<taxonomicName id="4C4E4D00E72AFFBB7DE8FB87FEFFED2E" box="[151,271,1088,1112]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="kolengh">
<emphasis id="B93AEA91E72AFFBB7DE8FB87FEFFED2E" box="[151,271,1088,1112]" italics="true" pageId="8" pageNumber="9">L. kolengh</emphasis>
</taxonomicName>
than in
<taxonomicName id="4C4E4D00E72AFFBB7C0FFB87FDF2ED2E" box="[368,514,1088,1112]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="morandae" status="sp. nov.">
<emphasis id="B93AEA91E72AFFBB7C0FFB87FDF2ED2E" box="[368,514,1088,1112]" italics="true" pageId="8" pageNumber="9">L. morandae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20957EAE72AFFBB7F75FB86FD96ED2E" box="[522,614,1088,1112]" pageId="8" pageNumber="9" rank="species">
<emphasis id="B93AEA91E72AFFBB7F75FB86FD96ED2E" bold="true" box="[522,614,1088,1112]" pageId="8" pageNumber="9">sp. nov.</emphasis>
</taxonomicNameLabel>
; the general background coloration of
<taxonomicName id="4C4E4D00E72AFFBB795AFB87FB6EED2E" box="[1061,1182,1088,1112]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="kolengh">
<emphasis id="B93AEA91E72AFFBB795AFB87FB6EED2E" box="[1061,1182,1088,1112]" italics="true" pageId="8" pageNumber="9">L. kolengh</emphasis>
</taxonomicName>
is dark gray or black, though in almost 50% of our samples the dorsal pattern cannot be recognized. In general, the dorsal scales of
<taxonomicName id="4C4E4D00E72AFFBB78FAFBA3FEF9EDD6" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="morandae" status="sp. nov.">
<emphasis id="B93AEA91E72AFFBB78FAFBA3FEF9EDD6" italics="true" pageId="8" pageNumber="9">L. morandae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20957EAE72AFFBB7C6EFB4EFE9CEDD6" box="[273,364,1160,1184]" pageId="8" pageNumber="9" rank="species">
<emphasis id="B93AEA91E72AFFBB7C6EFB4EFE9CEDD6" bold="true" box="[273,364,1160,1184]" pageId="8" pageNumber="9">sp. nov.</emphasis>
</taxonomicNameLabel>
are less “bristly” than those of
<emphasis id="B93AEA91E72AFFBB7FAAFB4FFCA4EDD6" box="[725,852,1160,1184]" italics="true" pageId="8" pageNumber="9">
<taxonomicName id="4C4E4D00E72AFFBB7FAAFB4FFCA0EDD6" box="[725,848,1160,1184]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="kolengh">L. kolengh</taxonomicName>
.
</emphasis>
Dorsal and ventral limbs scales are non-mucronate and non-bristly in
<taxonomicName id="4C4E4D00E72AFFBB7C1AFB6BFE04EDB2" box="[357,500,1196,1220]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="morandae" status="sp. nov.">
<emphasis id="B93AEA91E72AFFBB7C1AFB6BFE04EDB2" box="[357,500,1196,1220]" italics="true" pageId="8" pageNumber="9">L. morandae</emphasis>
</taxonomicName>
<emphasis id="B93AEA91E72AFFBB7C84FB6AFDA5EDB2" bold="true" box="[507,597,1196,1220]" pageId="8" pageNumber="9">
<taxonomicNameLabel id="A20957EAE72AFFBB7C84FB6AFDA5EDB2" box="[507,597,1196,1220]" pageId="8" pageNumber="9" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, but mucronate and “bristly” in
<taxonomicName id="4C4E4D00E72AFFBB7EC1FB6BFBC4EDB2" box="[958,1076,1196,1220]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="kolengh">
<emphasis id="B93AEA91E72AFFBB7EC1FB6BFBC4EDB2" box="[958,1076,1196,1220]" italics="true" pageId="8" pageNumber="9">L. kolengh</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BF13683E72AFFBB7DBAFB16FB73EE72" blockId="8.[151,1437,151,2012]" pageId="8" pageNumber="9">
<taxonomicName id="4C4E4D00E72AFFBB7DBAFB16FE44ED9E" box="[197,436,1232,1256]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="morandae" status="sp. nov.">
<emphasis id="B93AEA91E72AFFBB7DBAFB16FE44ED9E" box="[197,436,1232,1256]" italics="true" pageId="8" pageNumber="9">Liolaemus morandae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20957EAE72AFFBB7CC1FB16FDEAED9E" box="[446,538,1232,1256]" pageId="8" pageNumber="9" rank="species">
<emphasis id="B93AEA91E72AFFBB7CC1FB16FDEAED9E" bold="true" box="[446,538,1232,1256]" pageId="8" pageNumber="9">sp. nov.</emphasis>
</taxonomicNameLabel>
differs from
<taxonomicName id="4C4E4D00E72AFFBB7FC9FB17FCC4ED9E" box="[694,820,1232,1256]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="silvanae">
<emphasis id="B93AEA91E72AFFBB7FC9FB17FCC4ED9E" box="[694,820,1232,1256]" italics="true" pageId="8" pageNumber="9">L. silvanae</emphasis>
</taxonomicName>
in several characters, including: a smaller number of scales from occiput to rump (4757, X = 51.67 vs. 5264, X = 56;
<emphasis id="B93AEA91E72AFFBB7EE1FB33FC5CEC7A" box="[926,940,1269,1292]" italics="true" pageId="8" pageNumber="9">p</emphasis>
&lt;0.0001), higher number of ventral scales (7985, X = 81.33 vs. 7085, X = 74,59;
<emphasis id="B93AEA91E72AFFBB7F1BFADFFD82EC46" box="[612,626,1305,1328]" italics="true" pageId="8" pageNumber="9">p</emphasis>
&lt;0.0001), fewer third finger lamellae (1316, X = 15.33 vs. 1418, X = 16.24;
<emphasis id="B93AEA91E72AFFBB7D98FAFBFF05EC22" box="[231,245,1341,1364]" italics="true" pageId="8" pageNumber="9">p</emphasis>
&lt;0.0001), smaller body size (snout-vent length: 5061, X = 55.67 vs. 6578, X = 73, axilla-groin distance: 19.730.8, X = 25.9 vs. 25.139.7, X = 32.38; p&lt;
<quantity id="4CB69B66E72AFFBB7E49FAA6FC53EC0E" box="[822,931,1376,1400]" metricMagnitude="-6" metricUnit="m" metricValue="2.54" pageId="8" pageNumber="9" unit="in" value="1.0E-4">0.0001 in</quantity>
both cases), shorter forelimbs (elbow-wrist length: 4.695.56, X = 5.32 vs. 6.838.82, X = 7.71, third finger length: 6.699.14, X = 7.83 vs. 10.6513.76 X = 12.23;
<emphasis id="B93AEA91E72AFFBB7D98FA6FFF05ECB6" box="[231,245,1449,1472]" italics="true" pageId="8" pageNumber="9">p</emphasis>
&lt;
<quantity id="4CB69B66E72AFFBB7C68FA6EFE73ECB6" box="[279,387,1448,1472]" metricMagnitude="-6" metricUnit="m" metricValue="2.54" pageId="8" pageNumber="9" unit="in" value="1.0E-4">0.0001 in</quantity>
both cases), smaller head (head height: 6.528.21, X = 7.23 vs. 6.849.06, X = 8.18, head width: 9.0810.58, X = 9.84 vs. 11.3114.71, X = 13.14, head length: 10.9613.06, X = 11.96 vs. 12.4715.9, X = 14.42;
<emphasis id="B93AEA91E72AFFBB7D98FA37FF05EF7E" box="[231,245,1521,1544]" italics="true" pageId="8" pageNumber="9">p</emphasis>
&lt;
<quantity id="4CB69B66E72AFFBB7C67FA36FE74EF7E" box="[280,388,1520,1544]" metricMagnitude="-6" metricUnit="m" metricValue="2.54" pageId="8" pageNumber="9" unit="in" value="1.0E-4">0.0001 in</quantity>
all cases), smaller eye (eye height: 1.72.27, X = 1.99 vs. 2.092.9, X = 2.46, eye length: 2.913.42, X = 3.15 vs. 3.594.18, X = 3.86;
<emphasis id="B93AEA91E72AFFBB7FD1F9D3FD4CEF5A" box="[686,700,1557,1580]" italics="true" pageId="8" pageNumber="9">p</emphasis>
&lt;
<quantity id="4CB69B66E72AFFBB7F9FF9D2FCBCEF5A" box="[736,844,1556,1580]" metricMagnitude="-6" metricUnit="m" metricValue="2.54" pageId="8" pageNumber="9" unit="in" value="1.0E-4">0.0001 in</quantity>
both cases), and smaller auditory meatus (auditory meatus height: 1.62.44, X = 1.95 vs. 1.952.95, X = 2.54; auditory meatus length: 0.861.81, X = 1.27 vs. 1.49 2.57, X = 2.04;
<emphasis id="B93AEA91E72AFFBB7C31F99BFEACEF02" box="[334,348,1629,1652]" italics="true" pageId="8" pageNumber="9">p</emphasis>
&lt;
<quantity id="4CB69B66E72AFFBB7C02F99AFE19EF02" box="[381,489,1628,1652]" metricMagnitude="-6" metricUnit="m" metricValue="2.54" pageId="8" pageNumber="9" unit="in" value="1.0E-4">0.0001 in</quantity>
both cases). The general background coloration of
<taxonomicName id="4C4E4D00E72AFFBB7949F99BFB42EF02" box="[1078,1202,1628,1652]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="silvanae">
<emphasis id="B93AEA91E72AFFBB7949F99BFB42EF02" box="[1078,1202,1628,1652]" italics="true" pageId="8" pageNumber="9">L. silvanae</emphasis>
</taxonomicName>
is black; there is no recognizable dorsal pattern, and the venter is also strongly melanistic, whereas the venter is weakly melanistic in
<taxonomicName id="4C4E4D00E72AFFBB78FAF947FEF9EFCA" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="morandae" status="sp. nov.">
<emphasis id="B93AEA91E72AFFBB78FAF947FEF9EFCA" italics="true" pageId="8" pageNumber="9">L. morandae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20957EAE72AFFBB7C6FF962FE9AEFCA" box="[272,362,1700,1724]" pageId="8" pageNumber="9" rank="species">
<emphasis id="B93AEA91E72AFFBB7C6FF962FE94EFCA" bold="true" box="[272,356,1700,1724]" pageId="8" pageNumber="9">sp. nov</emphasis>
.
</taxonomicNameLabel>
<taxonomicName id="4C4E4D00E72AFFBB7C0DF962FD90EFCA" box="[370,608,1700,1724]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="morandae" status="sp. nov.">
<emphasis id="B93AEA91E72AFFBB7C0DF962FD90EFCA" box="[370,608,1700,1724]" italics="true" pageId="8" pageNumber="9">Liolaemus morandae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20957EAE72AFFBB7F17F962FD33EFCA" box="[616,707,1700,1724]" pageId="8" pageNumber="9" rank="species">
<emphasis id="B93AEA91E72AFFBB7F17F962FD33EFCA" bold="true" box="[616,707,1700,1724]" pageId="8" pageNumber="9">sp. nov.</emphasis>
</taxonomicNameLabel>
possesses well-defined vertebral and paravertebral lines that are not present in
<emphasis id="B93AEA91E72AFFBB7C46F90FFE4CEF96" box="[313,444,1736,1760]" italics="true" pageId="8" pageNumber="9">
<taxonomicName id="4C4E4D00E72AFFBB7C46F90FFE48EF96" box="[313,440,1736,1760]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="silvanae">L. silvanae</taxonomicName>
.
</emphasis>
In general,
<taxonomicName id="4C4E4D00E72AFFBB7F3BF90FFD4FEF96" box="[580,703,1736,1760]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="silvanae">
<emphasis id="B93AEA91E72AFFBB7F3BF90FFD4FEF96" box="[580,703,1736,1760]" italics="true" pageId="8" pageNumber="9">L. silvanae</emphasis>
</taxonomicName>
possesses a “bristly” appearance that is not present in
<taxonomicName id="4C4E4D00E72AFFBB7854F90FFF30EE72" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="morandae" status="sp. nov.">
<emphasis id="B93AEA91E72AFFBB7854F90FFF30EE72" italics="true" pageId="8" pageNumber="9">L. morandae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20957EAE72AFFBB7DB9F92AFEEFEE72" box="[198,287,1772,1796]" pageId="8" pageNumber="9" rank="species">
<emphasis id="B93AEA91E72AFFBB7DB9F92AFEE9EE72" bold="true" box="[198,281,1772,1796]" pageId="8" pageNumber="9">sp. nov</emphasis>
.
</taxonomicNameLabel>
Limb scales are less mucronated in
<taxonomicName id="4C4E4D00E72AFFBB7FC9F92BFCB5EE72" box="[694,837,1772,1796]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="morandae" status="sp. nov.">
<emphasis id="B93AEA91E72AFFBB7FC9F92BFCB5EE72" box="[694,837,1772,1796]" italics="true" pageId="8" pageNumber="9">L. morandae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20957EAE72AFFBB7E33F92AFC56EE72" box="[844,934,1772,1796]" pageId="8" pageNumber="9" rank="species">
<emphasis id="B93AEA91E72AFFBB7E33F92AFC56EE72" bold="true" box="[844,934,1772,1796]" pageId="8" pageNumber="9">sp. nov.</emphasis>
</taxonomicNameLabel>
than in
<taxonomicName id="4C4E4D00E72AFFBB797EF92BFB8BEE72" box="[1025,1147,1772,1796]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="silvanae">
<emphasis id="B93AEA91E72AFFBB797EF92BFB8BEE72" box="[1025,1147,1772,1796]" italics="true" pageId="8" pageNumber="9">L. silvanae</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C3546508E72AFFBF7DBAF8C9FAACEC62" lastPageId="12" lastPageNumber="13" pageId="8" pageNumber="9" type="description">
<paragraph id="8BF13683E72AFFBB7DBAF8C9FBA4EEAA" blockId="8.[151,1437,151,2012]" pageId="8" pageNumber="9">
<emphasis id="B93AEA91E72AFFBB7DBAF8C9FE12EE5E" bold="true" box="[197,482,1807,1832]" pageId="8" pageNumber="9">
Description of
<typeStatus id="54F58821E72AFFBB7C0BF8D6FE2DEE5E" box="[372,477,1808,1832]" pageId="8" pageNumber="9" type="holotype">holotype</typeStatus>
.
</emphasis>
Adult male. Snout-vent length 56.0 mm. Tail length (complete, not regenerated) 79.0 mm. Axilla-groin distance
<quantity id="4CB69B66E72AFFBB7CBCF8F2FDDAEE3A" box="[451,554,1844,1868]" metricMagnitude="-2" metricUnit="m" metricValue="2.28" pageId="8" pageNumber="9" unit="mm" value="22.8">22.8 mm</quantity>
. Auditory meatus-eye distance
<quantity id="4CB69B66E72AFFBB7EF7F8F2FC11EE3A" box="[904,993,1844,1868]" metricMagnitude="-3" metricUnit="m" metricValue="4.8" pageId="8" pageNumber="9" unit="mm" value="4.8">4.8 mm</quantity>
. External auditory meatus conspicuous, higher (
<quantity id="4CB69B66E72AFFBB7D8FF891FEBAEE19" box="[240,330,1879,1903]" metricMagnitude="-3" metricUnit="m" metricValue="2.1" pageId="8" pageNumber="9" unit="mm" value="2.1">2.1 mm</quantity>
) than wide (
<quantity id="4CB69B66E72AFFBB7CA4F891FDC8EE19" box="[475,568,1879,1903]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="8" pageNumber="9" unit="mm" value="1.2">1.2 mm</quantity>
). Head length 13.0 mm (from anterior border of tympanum to tip of snout),
<quantity id="4CB69B66E72AFFBB7DE8F8BAFF0CEEE2" box="[151,252,1916,1940]" metricMagnitude="-2" metricUnit="m" metricValue="1.05" pageId="8" pageNumber="9" unit="mm" value="10.5">10.5 mm</quantity>
wide (at anterior border of tympanum),
<quantity id="4CB69B66E72AFFBB7FB9F8BAFCEDEEE2" box="[710,797,1916,1940]" metricMagnitude="-3" metricUnit="m" metricValue="7.8" pageId="8" pageNumber="9" unit="mm" value="7.8">7.8 mm</quantity>
high (at anterior border of tympanum). Snout length 4.0 mm (orbit-tip of snout distance). Interorbital distance
<quantity id="4CB69B66E72AFFBB7F85F866FCA4EECE" box="[762,852,1952,1976]" metricMagnitude="-3" metricUnit="m" metricValue="4.2" pageId="8" pageNumber="9" unit="mm" value="4.2">4.2 mm</quantity>
. Eye-nostril distance
<quantity id="4CB69B66E72AFFBB7937F866FB52EECE" box="[1096,1186,1952,1976]" metricMagnitude="-3" metricUnit="m" metricValue="3.1" pageId="8" pageNumber="9" unit="mm" value="3.1">3.1 mm</quantity>
. Forelimb length
<quantity id="4CB69B66E72AFFBB7814F866FF36EEAA" metricMagnitude="-2" metricUnit="m" metricValue="1.67" pageId="8" pageNumber="9" unit="mm" value="16.7">16.7 mm</quantity>
. Tibial length
<quantity id="4CB69B66E72AFFBB7C19F802FE3DEEAA" box="[358,461,1988,2012]" metricMagnitude="-2" metricUnit="m" metricValue="1.03" pageId="8" pageNumber="9" unit="mm" value="10.3">10.3 mm</quantity>
. Foot length
<quantity id="4CB69B66E72AFFBB7F20F802FD33EEAA" box="[607,707,1988,2012]" metricMagnitude="-2" metricUnit="m" metricValue="1.68" pageId="8" pageNumber="9" unit="mm" value="16.8">16.8 mm</quantity>
(ankle to tip of claw on fourth toe).
</paragraph>
<caption id="DF31660BE72BFFBA7DE8F853FD90EEBC" httpUri="https://zenodo.org/record/208134/files/figure.png" pageId="9" pageNumber="10" targetBox="[231,1356,193,1919]" targetPageId="9">
<paragraph id="8BF13683E72BFFBA7DE8F853FD90EEBC" blockId="9.[151,1436,1941,1994]" pageId="9" pageNumber="10">
<emphasis id="B93AEA91E72BFFBA7DE8F853FEE3EEDC" bold="true" box="[151,275,1941,1964]" pageId="9" pageNumber="10">FIGURE 4.</emphasis>
Type locality of
<taxonomicName id="4C4E4D00E72BFFBA7CBDF853FD68EEDD" box="[450,664,1941,1963]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="morandae" status="sp. nov.">
<emphasis id="B93AEA91E72BFFBA7CBDF853FD68EEDD" box="[450,664,1941,1963]" italics="true" pageId="9" pageNumber="10">Liolaemus morandae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20957EAE72BFFBA7FDFF850FD01EEDD" box="[672,753,1942,1963]" pageId="9" pageNumber="10" rank="species">
<emphasis id="B93AEA91E72BFFBA7FDFF850FD01EEDD" bold="true" box="[672,753,1942,1963]" pageId="9" pageNumber="10">sp. nov.</emphasis>
</taxonomicNameLabel>
, Patagonic phytogeographic province, del Golfo de San Jorge District (45°4110,6” S; 67°5349,9” W, 693 m).
</paragraph>
</caption>
<caption id="DF31660BE728FFB97DE8FBDBFD40EDF9" httpUri="https://zenodo.org/record/208135/files/figure.png" pageId="10" pageNumber="11" targetBox="[157,1427,198,1017]" targetPageId="10">
<paragraph id="8BF13683E728FFB97DE8FBDBFD40EDF9" blockId="10.[151,1436,1053,1167]" pageId="10" pageNumber="11">
<emphasis id="B93AEA91E728FFB97DE8FBDBFEE2ED45" bold="true" box="[151,274,1053,1076]" pageId="10" pageNumber="11">FIGURE 5.</emphasis>
Distribution map for species of the
<taxonomicName id="4C4E4D00E728FFB97F03FBD8FCE7ED42" box="[636,791,1054,1076]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="lineomaculatus">
<emphasis id="B93AEA91E728FFB97F03FBD8FCE7ED42" box="[636,791,1054,1076]" italics="true" pageId="10" pageNumber="11">lineomaculatus</emphasis>
</taxonomicName>
group, with geographical localities sampled in this study superimposed on regional elevation (shading). Black circles:
<taxonomicName id="4C4E4D00E728FFB97FB5FBFBFCBBED24" box="[714,843,1084,1106]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="morandae" status="sp. nov.">
<emphasis id="B93AEA91E728FFB97FB5FBFBFCBBED24" box="[714,843,1084,1106]" italics="true" pageId="10" pageNumber="11">L. morandae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20957EAE728FFB97E2EFBFBFC51ED24" box="[849,929,1085,1106]" pageId="10" pageNumber="11" rank="species">
<emphasis id="B93AEA91E728FFB97E2EFBFBFC51ED24" bold="true" box="[849,929,1085,1106]" pageId="10" pageNumber="11">sp. nov.</emphasis>
</taxonomicNameLabel>
; white circle:
<taxonomicName id="4C4E4D00E728FFB9794BFBFBFB7CED24" box="[1076,1164,1084,1106]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="avilae" status="sp. nov.">
<emphasis id="B93AEA91E728FFB9794BFBFBFB7CED24" box="[1076,1164,1084,1106]" italics="true" pageId="10" pageNumber="11">L. avilae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20957EAE728FFB979EDFBFBFB12ED24" box="[1170,1250,1085,1106]" pageId="10" pageNumber="11" rank="species">
<emphasis id="B93AEA91E728FFB979EDFBFBFB12ED24" bold="true" box="[1170,1250,1085,1106]" pageId="10" pageNumber="11">sp. nov.</emphasis>
</taxonomicNameLabel>
; black squares:
<emphasis id="B93AEA91E728FFB978F8FBFBFECAED07" italics="true" pageId="10" pageNumber="11">
<taxonomicName id="4C4E4D00E728FFB978F8FBFBFEC4ED07" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="lineomaculatus">L. lineomaculatus</taxonomicName>
;
</emphasis>
white square:
<emphasis id="B93AEA91E728FFB97CB0FB9AFDB2ED07" box="[463,578,1115,1137]" italics="true" pageId="10" pageNumber="11">
<taxonomicName id="4C4E4D00E728FFB97CB0FB9AFDCCED07" box="[463,572,1115,1137]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="kolengh">L. kolengh</taxonomicName>
;
</emphasis>
white triangles:
<emphasis id="B93AEA91E728FFB97F92FB9AFC94ED07" box="[749,868,1115,1137]" italics="true" pageId="10" pageNumber="11">
<taxonomicName id="4C4E4D00E728FFB97F92FB9AFCAEED07" box="[749,862,1115,1137]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="hatcheri">L. hatcheri</taxonomicName>
;
</emphasis>
black triangle:
<emphasis id="B93AEA91E728FFB9797BFB9AFB8AED07" box="[1028,1146,1115,1137]" italics="true" pageId="10" pageNumber="11">
<taxonomicName id="4C4E4D00E728FFB9797BFB9AFB86ED07" box="[1028,1142,1115,1137]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="silvanae">L. silvanae</taxonomicName>
.
</emphasis>
Arrows indicate type localities. CH: Chubut province; SC: Santa Cruz province.
</paragraph>
</caption>
<paragraph id="8BF13683E728FFB87DBAFB71FD2FE849" blockId="10.[151,1437,1207,2023]" lastBlockId="11.[151,1437,151,1977]" lastPageId="11" lastPageNumber="12" pageId="10" pageNumber="11">
Dorsal head scales bulged, smooth, 15 between occiput, at the level of anterior border of tympanum, to rostral, pitted with numerous scale organs in the anterior region, and reducing to a single organ, or lack, in the posterior half of the head. Rostral scale wider (
<quantity id="4CB69B66E728FFB97F40FB39FD6AEC61" box="[575,666,1279,1303]" metricMagnitude="-3" metricUnit="m" metricValue="2.4" pageId="10" pageNumber="11" unit="mm" value="2.4">2.4 mm</quantity>
) than high (
<quantity id="4CB69B66E728FFB97E5DFB39FC8FEC61" box="[802,895,1279,1303]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" pageId="10" pageNumber="11" unit="mm" value="0.9">0.9 mm</quantity>
). Two postrostrals, together with anterior lorilabial, separate nasal scales from rostral, surrounded by six scales. Nasal scales longer than wide, irregularly hexagonal; nostril one-half length of nasal, posterior in position. Scales surrounding nasals 7 on the left side and 8 on the right side. Four internasals. Frontonasals six, irregular in size and position. Prefrontals 6, a small rhomboidal scale in the center (
<quantity id="4CB69B66E728FFB97C4FFA49FE7BECD1" box="[304,395,1423,1447]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" pageId="10" pageNumber="11" unit="mm" value="0.9">0.9 mm</quantity>
). Three dorso-lateral larger scales, one for the fragmentation of the right scale, and a pair lateral medium-sized scales (
<quantity id="4CB69B66E728FFB97CBCFA75FDEFECBD" box="[451,543,1459,1483]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" pageId="10" pageNumber="11" unit="mm" value="1.3">1.3 mm</quantity>
), approximately quadrangular. Two frontal scales. Frontoparietals in two rows, one anterior and one posterior scale, unfragmented. Interparietal pentagonal, surrounded by six scales; four smaller and irregular in front and sides, two larger in back. Parietal eye evident. Parietals slightly bulged, irregularly shaped, equal in size to interparietal (1.5 vs.
<quantity id="4CB69B66E728FFB97FF7F9D9FD15EF41" box="[648,741,1567,1591]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="10" pageNumber="11" unit="mm" value="1.5">1.5 mm</quantity>
). Circumorbitals:1310. Transversally expanded supraoculars 43. Smaller lateral supraoculars: 1617. One canthal higher than wide, separated from nasal by one postnasal. Loreal scales bulged, two on the left side (by fusion of the posterior loreal scale with preocular scale) and three on the rigth side. Lorilabials longer than wide (86), approximately equal to labials. Superciliaries 77, flattened and elongated, anterior five broadly overlapping dorsally. Orbit with 1517 upper and 1213 lower ciliaries on each side. Orbit diameter 4.5 x
<quantity id="4CB69B66E728FFB97CB8F915FDD2EF9D" box="[455,546,1747,1771]" metricMagnitude="-3" metricUnit="m" metricValue="1.9" pageId="10" pageNumber="11" unit="mm" value="1.9">1.9 mm</quantity>
. Preocular small, unfragmented, longer than wide. Subocular scale elongated, approximately nine times longer than wide (3.7 x
<quantity id="4CB69B66E728FFB97FBEF931FCEAEE79" box="[705,794,1783,1807]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" pageId="10" pageNumber="11" unit="mm" value="0.6">0.6 mm</quantity>
). A well marked longitudinal ridge along upper margin of preocular and subocular scales. Postocular small, slightly bulged, quarter superimposed to subocular, with a marked longitudinal ridge. Palpebral scales small granular and bulged. Supralabials 910, convex. Temporals smooth, convex, juxtaposed with one scale organ in the tip. Anterior auriculars smaller than adjacent posterior temporals, slightly projecting outward (32). Posterior auriculars small and granular. External auditory meatus conspicuous, higher (
<quantity id="4CB69B66E728FFB97C1CF86DFE4EEEB5" box="[355,446,1963,1987]" metricMagnitude="-3" metricUnit="m" metricValue="2.1" pageId="10" pageNumber="11" unit="mm" value="2.1">2.1 mm</quantity>
) than wide (
<quantity id="4CB69B66E728FFB97F2EF86DFD5FEEB5" box="[593,687,1963,1987]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="10" pageNumber="11" unit="mm" value="1.2">1.2 mm</quantity>
). Lateral scales of neck granular with inflated skin. Mental scale wider (
<quantity id="4CB69B66E728FFB97D9AF809FECEEE91" box="[229,318,1999,2023]" metricMagnitude="-3" metricUnit="m" metricValue="2.9" pageId="10" pageNumber="11" unit="mm" value="2.9">2.9 mm</quantity>
) than high (
<quantity id="4CB69B66E728FFB97CBCF809FDEEEE91" box="[451,542,1999,2023]" metricMagnitude="-3" metricUnit="m" metricValue="1.4" pageId="10" pageNumber="11" unit="mm" value="1.4">1.4 mm</quantity>
), in contact with four scales. Mental posterior followed by two postmentals, and two rows of three chinshields on each side. Six infralabials on each side, first on each side quadrangular two times wider than supralabials, all others elongated, slightly smaller than supralabials. Gular scales smooth, flat, imbricate, with rounded posterior margins, with melanophores. Scales of throat between chinshields slightly juxtaposed, becoming slightly imbricate toward auditory meatus. Thirty gulars between tympanum openings. Infralabials separated from chinshields by one to two rows of scales.
</paragraph>
<paragraph id="8BF13683E729FFB87DBAFE8AFEA4E8F1" blockId="11.[151,1437,151,1977]" pageId="11" pageNumber="12">Antehumeral, longitudinal and postauricular neck folds well developed; gular, rictal, dorsolateral and oblique not conspicuous.</paragraph>
<paragraph id="8BF13683E729FFB87DBAFE52FC2BEBF2" blockId="11.[151,1437,151,1977]" pageId="11" pageNumber="12">Scales of dorsal neck region rhomboidal, imbricate and strongly keeled. Fifty dorsal scales between occiput and anterior surface of thighs. Dorsal body scales rhomboidal, imbricate, strongly keeled, mucronate, very few trifid scales. Dorsal scales grade laterally into slightly smaller, slightly keeled scales at midbody. Scales immediately anterior and posterior to forelimb and hindlimb insertion small, smooth, granular, and non-overlapping. Body lateral scales grading smaller to larger at midbody. Ventral body scales rhomboidal, smooth, flat, imbricate, larger than dorsal scales. Sixty-one midbody scales; scales between mental and precloacal pores 85. Scales of cloacal region about equal in size to ventral body scales; without precloacal pores.</paragraph>
<paragraph id="8BF13683E729FFB87DBAFD49FDAFEA2A" blockId="11.[151,1437,151,1977]" pageId="11" pageNumber="12">Anterior suprabrachials rhomboidal, imbricate, smooth, slightly larger in size to dorsal body scales. Postabrachials smaller, smooth, becoming granular near axilla. Supra-antebrachials similar to suprabrachial. Infra-antebrachials rhomboidals, imbricate, smooth, toward the hand slightly mucronate. Supracarpals imbricated, rhomboidal, smooth. Infracarpals strongly imbricate, rhomboidal, slightly keeled, 3-mucronate. Subdigital lamellae with 25 keels, each terminating in a short mucron, 25 mucronate, numbering: I: 8, II: 12, III: 16, IV: 17, V: 11. Claws robust, curved and sharp, opaque brown.</paragraph>
<paragraph id="8BF13683E729FFB87DBAFCA1FDB9ED0A" blockId="11.[151,1437,151,1977]" pageId="11" pageNumber="12">Suprafemorals as large as dorsal body scales, rhomboidal, imbricated, smooth near the body, toward to the knee slightly keeled. Postfemorals small, granular shape. Supratibials rhomboidal, imbricated, keeled, some mucronate, smaller and smooth toward to the foot. Infrafemoral scales small, granular and smooth. Supratarsals rhomboidal, imbricated and smooth. Infratarsals small, rhomboidal, imbricate, smooth, mucronate, slightly keeled near the digit. Subdigital scales 13 keeled, 15 mucronate, numbering: I: 9, II: 14, III: 20, IV: 21, V: 14. Claws robust, curved and sharp, opaque brown. Tail complete, non-regenerated. Dorsal and lateral caudal scales, rhomboidal in the first half of the tail, becoming quadrangular toward the tip, strongly keeled. Ventrals subtriangular and smooth, toward posterior half moderate keeled.</paragraph>
<paragraph id="8BF13683E729FFB87DBAFB41FE92ECB6" blockId="11.[151,1437,151,1977]" pageId="11" pageNumber="12">
<emphasis id="B93AEA91E729FFB87DBAFB41FE1CEDD6" bold="true" box="[197,492,1159,1184]" pageId="11" pageNumber="12">
Color of
<typeStatus id="54F58821E729FFB87C4EFB4EFE67EDD6" box="[305,407,1160,1184]" pageId="11" pageNumber="12" type="holotype">holotype</typeStatus>
in life.
</emphasis>
Dark grey dorsal background (
<figureCitation id="13752A06E729FFB87E2FFB4EFC41EDD6" box="[848,945,1160,1184]" captionStart="FIGURE 3" captionStartId="7.[151,250,1729,1752]" captionTargetBox="[225,1361,193,1707]" captionTargetId="figure@7.[225,1362,193,1709]" captionTargetPageId="7" captionText="FIGURE 3. Upper, dorsal and ventral views of male holotype in life of Liolaemus morandae sp. nov. MLP. S 2626; lower, dorsal and ventral views of female (LJAMM-CNP 13020)." httpUri="https://zenodo.org/record/208133/files/figure.png" pageId="11" pageNumber="12">Figure 3</figureCitation>
) that becomes lighter on the lateral region, between axilla and groin. Dorsal pattern presents twelve paravertebral, quadrangular, black blotches, extending from the nuchal to the postcloacal region, those series fuse into a dark line that is present to the tip of the tail. Dorsolateral region, in between the occipital and the tip of the tail, formed by black and white-bordered blotches. Lateral region, between axilla and groin, is characterized by a black and white reticulate pattern. In between the series of blotches, one white vertebral line and two light orange paravertebral longitudinal lines appear, one and one and a half scale wide, respectively. Vertebral line extends from the nuchal region, while the paravertebral lines extend from the temporal region; all of them continue to the tip of the tail. Dorsal region of the limbs are dark gray with a black reticulation.
</paragraph>
<paragraph id="8BF13683E729FFB87DBAFA0AFAF0EFEE" blockId="11.[151,1437,151,1977]" pageId="11" pageNumber="12">Dark gray head background. Lateral white line along the longitudinal folds is present from the top of the auditory meatus to the antehumeral fold. White subocular scale with black longitudinal ridge. Dorsal surface with black blotches, one longitudinal blotch is present in the anterior part of the head, crossing through internasals, postnasals and prefrontals; two transverse blotches are crossing through postfrontals and first circumorbitals, and extend back to the last circumorbitals, forming a cross with the longitudinal blotch. The interparietal scale is surrounded by three small dark-brown blotches and the occipitals show a white longitudinal band of the same color.</paragraph>
<paragraph id="8BF13683E729FFB87DBAF962FDEAEE5E" blockId="11.[151,1437,151,1977]" pageId="11" pageNumber="12">White ventral color with a black reticulate pattern over the entire body, most strongly marked in the pectoral region and throat, and extending to the adjacent malar and maxilla region. Ventral scales with melanophores. Ventral area of chest, belly, cloacal region and limbs tinged with a light yellowish color. Cloacal and postcloacal region with few black blotches. Gray tail.</paragraph>
<paragraph id="8BF13683E729FFB87DBAF8F5FAF7EECE" blockId="11.[151,1437,151,1977]" pageId="11" pageNumber="12">
<emphasis id="B93AEA91E729FFB87DBAF8F5FDA9EE3D" bold="true" box="[197,601,1843,1868]" pageId="11" pageNumber="12">
Color of
<typeStatus id="54F58821E729FFB87C4EF8F2FE68EE3A" box="[305,408,1844,1868]" pageId="11" pageNumber="12" type="holotype">holotype</typeStatus>
in preservative.
</emphasis>
After one year in preservative, the dorsal coloration of the head, dorsum, body flanks and tail becomes darker while maintaining the contrast, but the two yellowish paravertebral lines turned gray. Ventral scales of throat, neck, chest, belly and forelimbs retain the same coloration as in life, and the distinctive light ventral yellowish tinge of chest, belly, cloacal region and limbs turns gray (
<figureCitation id="13752A06E729FFB879E8F866FB09EECE" box="[1175,1273,1952,1976]" captionStart="FIGURE 6" captionStartId="12.[151,250,937,960]" captionTargetBox="[151,1436,209,931]" captionTargetId="figure@12.[151,1436,209,933]" captionTargetPageId="12" captionText="FIGURE 6. Morphological comparison between Liolaemus morandae sp. nov. (A), L. lineomaculatus (B) and L. avilae sp. nov. (C). Notice differences in color pattern on the tails, size of blotches on the limbs and differences in vertebral and paravertebral lines. Dorsal scales (shown on the right) are less mucronated in L. lineomaculatus. Dorsal scales of hindlimbs are less carinated in L. morandae sp. nov. than in the other species, while they are more mucronated in L. avilae sp. nov." httpUri="https://zenodo.org/record/208136/files/figure.png" pageId="11" pageNumber="12">Figure 6</figureCitation>
).
</paragraph>
<caption id="DF31660BE72EFFBF7DE8FC6FFAFBED6C" httpUri="https://zenodo.org/record/208136/files/figure.png" pageId="12" pageNumber="13" targetBox="[151,1436,209,931]" targetPageId="12">
<paragraph id="8BF13683E72EFFBF7DE8FC6FFAFBED6C" blockId="12.[151,1436,937,1051]" pageId="12" pageNumber="13">
<emphasis id="B93AEA91E72EFFBF7DE8FC6FFEE4EAC8" bold="true" box="[151,276,937,960]" pageId="12" pageNumber="13">FIGURE 6.</emphasis>
Morphological comparison between
<taxonomicName id="4C4E4D00E72EFFBF7FE9FC6FFC9FEAC9" box="[662,879,937,959]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="morandae" status="sp. nov.">
<emphasis id="B93AEA91E72EFFBF7FE9FC6FFC9FEAC9" box="[662,879,937,959]" italics="true" pageId="12" pageNumber="13">Liolaemus morandae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20957EAE72EFFBF7E08FC6CFC3AEAC9" box="[887,970,938,959]" pageId="12" pageNumber="13" rank="species">
<emphasis id="B93AEA91E72EFFBF7E08FC6CFC3AEAC9" bold="true" box="[887,970,938,959]" pageId="12" pageNumber="13">sp. nov.</emphasis>
</taxonomicNameLabel>
(A),
<taxonomicName id="4C4E4D00E72EFFBF797BFC6CFB4CEAC9" box="[1028,1212,937,959]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="lineomaculatus">
<emphasis id="B93AEA91E72EFFBF797BFC6CFB4CEAC9" box="[1028,1212,937,959]" italics="true" pageId="12" pageNumber="13">L. lineomaculatus</emphasis>
</taxonomicName>
(B) and
<taxonomicName id="4C4E4D00E72EFFBF7864FC6CFA85EAC9" box="[1307,1397,937,959]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="avilae" status="sp. nov.">
<emphasis id="B93AEA91E72EFFBF7864FC6CFA85EAC9" box="[1307,1397,937,959]" italics="true" pageId="12" pageNumber="13">L. avilae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20957EAE72EFFBF7802FC6CFF33EAAB" pageId="12" pageNumber="13" rank="species">
<emphasis id="B93AEA91E72EFFBF7802FC6CFF33EAAB" bold="true" pageId="12" pageNumber="13">sp. nov.</emphasis>
</taxonomicNameLabel>
(C). Notice differences in color pattern on the tails, size of blotches on the limbs and differences in vertebral and paravertebral lines. Dorsal scales (shown on the right) are less mucronated in
<emphasis id="B93AEA91E72EFFBF7E0FFC21FBDEEA8A" box="[880,1070,998,1020]" italics="true" pageId="12" pageNumber="13">
<taxonomicName id="4C4E4D00E72EFFBF7E0FFC21FBDAEA8A" box="[880,1066,998,1020]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="lineomaculatus">L. lineomaculatus</taxonomicName>
.
</emphasis>
Dorsal scales of hindlimbs are less carinated in
<taxonomicName id="4C4E4D00E72EFFBF7C6BFBC0FE65ED6D" box="[276,405,1029,1051]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="morandae" status="sp. nov.">
<emphasis id="B93AEA91E72EFFBF7C6BFBC0FE65ED6D" box="[276,405,1029,1051]" italics="true" pageId="12" pageNumber="13">L. morandae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20957EAE72EFFBF7CE3FBC3FE1CED6C" box="[412,492,1029,1050]" pageId="12" pageNumber="13" rank="species">
<emphasis id="B93AEA91E72EFFBF7CE3FBC3FE1CED6C" bold="true" box="[412,492,1029,1050]" pageId="12" pageNumber="13">sp. nov.</emphasis>
</taxonomicNameLabel>
than in the other species, while they are more mucronated in
<taxonomicName id="4C4E4D00E72EFFBF7923FBC0FB45ED6D" box="[1116,1205,1029,1051]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="avilae" status="sp. nov.">
<emphasis id="B93AEA91E72EFFBF7923FBC0FB45ED6D" box="[1116,1205,1029,1051]" italics="true" pageId="12" pageNumber="13">L. avilae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20957EAE72EFFBF79C4FBC3FAFBED6C" box="[1211,1291,1029,1050]" pageId="12" pageNumber="13" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</caption>
<paragraph id="8BF13683E72EFFBF7DBAFB81FAACEC62" blockId="12.[151,1436,1095,1912]" pageId="12" pageNumber="13">
<emphasis id="B93AEA91E72EFFBF7DBAFB81FEC9ED16" bold="true" box="[197,313,1095,1120]" pageId="12" pageNumber="13">Variation.</emphasis>
Morphological and meristic character variation between males and females of
<taxonomicName id="4C4E4D00E72EFFBF79D0FB8EFA6CED16" box="[1199,1436,1096,1120]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="morandae" status="sp. nov.">
<emphasis id="B93AEA91E72EFFBF79D0FB8EFA6CED16" box="[1199,1436,1096,1120]" italics="true" pageId="12" pageNumber="13">Liolaemus morandae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20957EAE72EFFBF7DE8FBAAFF02EDF2" box="[151,242,1132,1156]" pageId="12" pageNumber="13" rank="species">
<emphasis id="B93AEA91E72EFFBF7DE8FBAAFF02EDF2" bold="true" box="[151,242,1132,1156]" pageId="12" pageNumber="13">sp. nov.</emphasis>
</taxonomicNameLabel>
, are shown in
<tableCitation id="C6CC0338E72EFFBF7CE1FBAAFE05EDF2" box="[414,501,1132,1156]" captionStart="TABLE 6" captionStartId="14.[151,235,632,653]" captionTargetBox="[151,1436,711,1852]" captionTargetPageId="14" captionText="TABLE 6. Variation between females and males of L. morandae sp. nov. and L. avilae sp. nov. Summarized as mean ± SD (min max) values for all meristic and morphometric (in millimeters) variables." httpUri="http://table.plazi.org/id/DF31660BE72CFFBD7DE8FDBEFC35EBDA" pageId="12" pageNumber="13" tableUuid="DF31660BE72CFFBD7DE8FDBEFC35EBDA">Table 6</tableCitation>
. Females in life present basically the same dorsal and lateral pattern as present in males, but females generally have a lighter brown background coloration; vertebral and paravertebral lines are strongly defined, and in both sexes paravertebral lines vary from white to yellow. Ventrally, one male shows a light orange color on the belly, while in other males and all females the ventral region ranged from white to gray due to different degrees of melanism. All distinctive ventral light orange coloration changed to gray in preservative.
</paragraph>
</subSubSection>
<subSubSection id="C3546508E72EFFBF7DBAFAE6FEC1EC09" pageId="12" pageNumber="13" type="etymology">
<paragraph id="8BF13683E72EFFBF7DBAFAE6FEC1EC09" blockId="12.[151,1436,1095,1912]" pageId="12" pageNumber="13">
<emphasis id="B93AEA91E72EFFBF7DBAFAE6FEBCEC4E" bold="true" box="[197,332,1312,1336]" pageId="12" pageNumber="13">Etymology.</emphasis>
The specific epithet of this species “
<taxonomicName id="4C4E4D00E72EFFBF7F92FAE6FCAEEC4E" box="[749,862,1312,1336]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="morandae">
<emphasis id="B93AEA91E72EFFBF7F92FAE6FCAEEC4E" box="[749,862,1312,1336]" italics="true" pageId="12" pageNumber="13">morandae</emphasis>
</taxonomicName>
” refers to our colleague and the first authors PhD advisor Dr. Mariana Morando, to honor her after more than ten years of research and teaching on the Patagonian herpetofauna.
</paragraph>
</subSubSection>
<subSubSection id="C3546508E72EFFBF7DBAFA4DFDA6EE0E" pageId="12" pageNumber="13" type="distribution">
<paragraph id="8BF13683E72EFFBF7DBAFA4DFE7CEF0A" blockId="12.[151,1436,1095,1912]" pageId="12" pageNumber="13">
<emphasis id="B93AEA91E72EFFBF7DBAFA4DFEAEECD5" bold="true" box="[197,350,1419,1444]" pageId="12" pageNumber="13">Distribution.</emphasis>
<taxonomicName id="4C4E4D00E72EFFBF7C18FA4AFDA6ECD2" box="[359,598,1420,1444]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="morandae" status="sp. nov.">
<emphasis id="B93AEA91E72EFFBF7C18FA4AFDA6ECD2" box="[359,598,1420,1444]" italics="true" pageId="12" pageNumber="13">Liolaemus morandae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20957EAE72EFFBF7F21FA4AFD4AECD2" box="[606,698,1420,1444]" pageId="12" pageNumber="13" rank="species">
<emphasis id="B93AEA91E72EFFBF7F21FA4AFD4AECD2" bold="true" box="[606,698,1420,1444]" pageId="12" pageNumber="13">sp. nov.</emphasis>
</taxonomicNameLabel>
is known only from the
<typeStatus id="54F58821E72EFFBF7EA7FA4BFBF8ECD3" box="[984,1032,1421,1445]" pageId="12" pageNumber="13">type</typeStatus>
locality and a few localities on the Pampa del Castillo Plateau; and from three localities from southern Chubut and northern Santa Cruz provinces. The
<typeStatus id="54F58821E72EFFBF7DE8FA12FF09EC9A" box="[151,249,1492,1516]" pageId="12" pageNumber="13" type="holotype">holotype</typeStatus>
and some
<typeStatus id="54F58821E72EFFBF7C01FA13FE19EC9B" box="[382,489,1493,1517]" pageId="12" pageNumber="13" type="paratype">paratypes</typeStatus>
are from Provincial Road 37,
<quantity id="4CB69B66E72EFFBF7E28FA12FC47EC9A" box="[855,951,1492,1516]" metricMagnitude="4" metricUnit="m" metricValue="2.28" pageId="12" pageNumber="13" unit="km" value="22.8">22.8 km</quantity>
SW junction National Road 3, Escalante department, Chubut province,
<collectingCountry id="F3597613E72EFFBF7C8FFA3EFDAFEF66" box="[496,607,1528,1552]" name="Argentina" pageId="12" pageNumber="13">Argentina</collectingCountry>
(
<figureCitation id="13752A06E72EFFBF7F10FA3EFD2DEF66" box="[623,733,1528,1552]" captionStart="FIGURE 4" captionStartId="9.[151,250,1941,1964]" captionTargetBox="[231,1356,193,1919]" captionTargetId="figure@9.[231,1356,193,1920]" captionTargetPageId="9" captionText="FIGURE 4. Type locality of Liolaemus morandae sp. nov., Patagonic phytogeographic province, del Golfo de San Jorge District (45 ° 41 10,6 ” S; 67 ° 53 49,9 ” W, 693 m)." httpUri="https://zenodo.org/record/208134/files/figure.png" pageId="12" pageNumber="13">Figures 4</figureCitation>
,
<figureCitation id="13752A06E72EFFBF7F95FA3EFD0AEF66" box="[746,762,1528,1552]" captionStart="FIGURE 5" captionStartId="10.[151,250,1053,1076]" captionTargetBox="[157,1427,198,1017]" captionTargetId="figure@10.[151,1436,193,1032]" captionTargetPageId="10" captionText="FIGURE 5. Distribution map for species of the lineomaculatus group, with geographical localities sampled in this study superimposed on regional elevation (shading). Black circles: L. morandae sp. nov.; white circle: L. avilae sp. nov.; black squares: L. lineomaculatus; white square: L. kolengh; white triangles: L. hatcheri; black triangle: L. silvanae. Arrows indicate type localities. CH: Chubut province; SC: Santa Cruz province." httpUri="https://zenodo.org/record/208135/files/figure.png" pageId="12" pageNumber="13">5</figureCitation>
). The other
<typeStatus id="54F58821E72EFFBF7EF9FA3FFC01EF67" box="[902,1009,1529,1553]" pageId="12" pageNumber="13" type="paratype">paratypes</typeStatus>
are from Provincial Road 37,
<quantity id="4CB69B66E72EFFBF7832FA3EFA6CEF66" box="[1357,1436,1528,1552]" metricMagnitude="3" metricUnit="m" metricValue="2.5" pageId="12" pageNumber="13" unit="km" value="2.5">2.5 km</quantity>
W junction National Road
<quantity id="4CB69B66E72EFFBF7CB8F9DAFE03EF42" box="[455,499,1564,1588]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="12" pageNumber="13" unit="in" value="3.0">3 in</quantity>
Holdich Station, Escalante department, Chubut province,
<collectingCountry id="F3597613E72EFFBF79FBF9DAFB04EF42" box="[1156,1268,1564,1588]" name="Argentina" pageId="12" pageNumber="13">Argentina</collectingCountry>
(
<figureCitation id="13752A06E72EFFBF787CF9DAFA81EF42" box="[1283,1393,1564,1588]" captionStart="FIGURE 4" captionStartId="9.[151,250,1941,1964]" captionTargetBox="[231,1356,193,1919]" captionTargetId="figure@9.[231,1356,193,1920]" captionTargetPageId="9" captionText="FIGURE 4. Type locality of Liolaemus morandae sp. nov., Patagonic phytogeographic province, del Golfo de San Jorge District (45 ° 41 10,6 ” S; 67 ° 53 49,9 ” W, 693 m)." httpUri="https://zenodo.org/record/208134/files/figure.png" pageId="12" pageNumber="13">Figures 4</figureCitation>
,
<figureCitation id="13752A06E72EFFBF7802F9DAFA7DEF42" box="[1405,1421,1564,1588]" captionStart="FIGURE 5" captionStartId="10.[151,250,1053,1076]" captionTargetBox="[157,1427,198,1017]" captionTargetId="figure@10.[151,1436,193,1032]" captionTargetPageId="10" captionText="FIGURE 5. Distribution map for species of the lineomaculatus group, with geographical localities sampled in this study superimposed on regional elevation (shading). Black circles: L. morandae sp. nov.; white circle: L. avilae sp. nov.; black squares: L. lineomaculatus; white square: L. kolengh; white triangles: L. hatcheri; black triangle: L. silvanae. Arrows indicate type localities. CH: Chubut province; SC: Santa Cruz province." httpUri="https://zenodo.org/record/208135/files/figure.png" pageId="12" pageNumber="13">5</figureCitation>
). The distance from the
<typeStatus id="54F58821E72EFFBF7CEBF987FE34EF2F" box="[404,452,1601,1625]" pageId="12" pageNumber="13">type</typeStatus>
locality to the first locality is more than
<quantity id="4CB69B66E72EFFBF7EEFF986FC1CEF2E" box="[912,1004,1600,1624]" metricMagnitude="4" metricUnit="m" metricValue="1.78" pageId="12" pageNumber="13" unit="km" value="17.8">17.8 km</quantity>
airline, and more than
<quantity id="4CB69B66E72EFFBF798EF986FABDEF2E" box="[1265,1357,1600,1624]" metricMagnitude="4" metricUnit="m" metricValue="3.88" pageId="12" pageNumber="13" unit="km" value="38.8">38.8 km</quantity>
airline to the second locality.
</paragraph>
<paragraph id="8BF13683E72EFFBF7DBAF94EFDA6EE0E" blockId="12.[151,1436,1095,1912]" pageId="12" pageNumber="13">
<emphasis id="B93AEA91E72EFFBF7DBAF94EFE73EFD6" bold="true" box="[197,387,1671,1696]" pageId="12" pageNumber="13">Natural history.</emphasis>
<taxonomicName id="4C4E4D00E72EFFBF7CF4F94EFD89EFD6" box="[395,633,1672,1696]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="morandae" status="sp. nov.">
<emphasis id="B93AEA91E72EFFBF7CF4F94EFD89EFD6" box="[395,633,1672,1696]" italics="true" pageId="12" pageNumber="13">Liolaemus morandae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20957EAE72EFFBF7FFFF94EFD2AEFD6" box="[640,730,1672,1696]" pageId="12" pageNumber="13" rank="species">
<emphasis id="B93AEA91E72EFFBF7FFFF94EFD2AEFD6" bold="true" box="[640,730,1672,1696]" pageId="12" pageNumber="13">sp. nov.</emphasis>
</taxonomicNameLabel>
occurs in Patagónica province, Golfo de San Jorge District, in environments characterized as grasslands and steppe highlands (
<bibRefCitation id="EFDF4B72E72EFFBF7EFEF96AFC0EEFB2" author="Roig" box="[897,1022,1708,1732]" pageId="12" pageNumber="13" refString="Roig, A. F. (1998) La vegetacion de la Patagonia. In: Correa M. N. (Ed.), Flora Patagonica. Coleccion cientifica del INTA, Tomo VIII, parte I, Buenos Aires, Argentina, pp. 48 - 174." type="book chapter" year="1998">Roig 1998</bibRefCitation>
), with vegetation dominated by the shrubs
<taxonomicName id="4C4E4D00E72EFFBF7D98F916FEBDEF9E" box="[231,333,1744,1768]" class="Liliopsida" family="Poaceae" genus="Stipa" kingdom="Plantae" order="Poales" pageId="12" pageNumber="13" phylum="Tracheophyta" rank="species" species="spp">
<emphasis id="B93AEA91E72EFFBF7D98F916FEBDEF9E" box="[231,333,1744,1768]" italics="true" pageId="12" pageNumber="13">Stipa spp</emphasis>
</taxonomicName>
.,
<taxonomicName id="4C4E4D00E72EFFBF7C1CF916FDB6EF9E" box="[355,582,1744,1768]" class="Magnoliopsida" family="Asteraceae" genus="Senecio" kingdom="Plantae" order="Asterales" pageId="12" pageNumber="13" phylum="Tracheophyta" rank="species" species="filaginoides">
<emphasis id="B93AEA91E72EFFBF7C1CF916FDB6EF9E" box="[355,582,1744,1768]" italics="true" pageId="12" pageNumber="13">Senecio filaginoides</emphasis>
</taxonomicName>
,
<emphasis id="B93AEA91E72EFFBF7F2BF916FC13EF9E" box="[596,995,1744,1768]" italics="true" pageId="12" pageNumber="13">
<taxonomicName id="4C4E4D00E72EFFBF7F2BF916FC5DEF9E" authority="Nassauvia" authorityName="Nassauvia" box="[596,941,1744,1768]" class="Magnoliopsida" family="Apiaceae" genus="Mulinum" kingdom="Plantae" order="Apiales" pageId="12" pageNumber="13" phylum="Tracheophyta" rank="species" species="spinosum">Mulinum spinosum, Nassauvia</taxonomicName>
<taxonomicName id="4C4E4D00E72EFFBF7ECBF917FC2EEF9E" box="[948,990,1745,1768]" class="Liliopsida" family="Poaceae" genus="Stipa" kingdom="Plantae" order="Poales" pageId="12" pageNumber="13" phylum="Tracheophyta" rank="species" species="spp">spp</taxonomicName>
.
</emphasis>
, among others. This species was found in open substrates, sharing the habitat with
<emphasis id="B93AEA91E72EFFBF7FEFF932FB2CEE7A" box="[656,1244,1780,1804]" italics="true" pageId="12" pageNumber="13">
<taxonomicName id="4C4E4D00E72EFFBF7FEFF932FCA1EE7A" box="[656,849,1780,1804]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="kingii">Liolaemus kingii</taxonomicName>
,
<taxonomicName id="4C4E4D00E72EFFBF7E20F933FB89EE7A" authority="Diplolaemus" authorityName="Diplolaemus" box="[863,1145,1780,1804]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="bibronii">L. bibronii, Diplolaemus</taxonomicName>
<taxonomicName id="4C4E4D00E72EFFBF79FCF932FB2CEE7A" box="[1155,1244,1780,1804]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="bibronii">bibronii</taxonomicName>
</emphasis>
and
<emphasis id="B93AEA91E72EFFBF7867F933FA6CEE7A" box="[1304,1436,1780,1804]" italics="true" pageId="12" pageNumber="13">D. darwinii</emphasis>
(
<figureCitation id="13752A06E72EFFBF7DE1F8DEFEF1EE46" box="[158,257,1816,1840]" captionStart="FIGURE 4" captionStartId="9.[151,250,1941,1964]" captionTargetBox="[231,1356,193,1919]" captionTargetId="figure@9.[231,1356,193,1920]" captionTargetPageId="9" captionText="FIGURE 4. Type locality of Liolaemus morandae sp. nov., Patagonic phytogeographic province, del Golfo de San Jorge District (45 ° 41 10,6 ” S; 67 ° 53 49,9 ” W, 693 m)." httpUri="https://zenodo.org/record/208134/files/figure.png" pageId="12" pageNumber="13">Figure 4</figureCitation>
). Based on the natural history of its close relative
<emphasis id="B93AEA91E72EFFBF7E53F8DFFC0BEE46" box="[812,1019,1816,1840]" italics="true" pageId="12" pageNumber="13">
<taxonomicName id="4C4E4D00E72EFFBF7E53F8DFFC07EE46" box="[812,1015,1816,1840]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="lineomaculatus">L. lineomaculatus</taxonomicName>
,
</emphasis>
we hypothesize that
<taxonomicName id="4C4E4D00E72EFFBF799AF8DFFA85EE46" box="[1253,1397,1816,1840]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="morandae" status="sp. nov.">
<emphasis id="B93AEA91E72EFFBF799AF8DFFA85EE46" box="[1253,1397,1816,1840]" italics="true" pageId="12" pageNumber="13">L. morandae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20957EAE72EFFBF7805F8DEFF38EE22" pageId="12" pageNumber="13" rank="species">
<emphasis id="B93AEA91E72EFFBF7805F8DEFF38EE22" bold="true" pageId="12" pageNumber="13">sp. nov.</emphasis>
</taxonomicNameLabel>
is herbivorous and likely to also be viviparous, perhaps giving birth to between three and six neonates per clutch (
<bibRefCitation id="EFDF4B72E72EFFBF7D95F8A6FEA4EE0E" author="Cei" box="[234,340,1888,1912]" pageId="12" pageNumber="13" refString="Cei, J. M. (1986) Reptiles del centro, centro-oeste y sur de la Argentina. Herpetofauna de las zonas aridas y semiaridas. Museo Regionale di Scienze Naturali Torino, 4, 1 - 527." type="journal article" year="1986">Cei 1986</bibRefCitation>
;
<bibRefCitation id="EFDF4B72E72EFFBF7C1FF8A6FDBAEE0E" author="Espinoza" box="[352,586,1888,1912]" pageId="12" pageNumber="13" refString="Espinoza, R. E., Wiens, J. J. &amp; Tracy, C. R. (2004) Recurrent evolution of herbivory in small, cold-climate lizards: breaking the ecophysiological rules of reptilian herbivory. Proceedings of the National Academy of Sciences, 101, 16819 - 16824." type="journal article" year="2004">
Espinoza
<emphasis id="B93AEA91E72EFFBF7CB1F8A7FDF6EE0E" box="[462,518,1888,1912]" italics="true" pageId="12" pageNumber="13">et al.</emphasis>
2004
</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>