894 lines
104 KiB
XML
894 lines
104 KiB
XML
<document id="715E11205A3D50ACDDBBA555E02BC6A4" ID-DOI="10.5281/zenodo.282620" ID-GBIF-Dataset="dc293202-7bfd-4ccd-85b6-fad297545b73" ID-ISSN="1175-5326" ID-Zenodo-Dep="282620" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460544184786" checkinUser="plazi" docAuthor="Rivero-Blanco, Carlos & Schargel, Walter E." docDate="2012" docId="03E77F69FF95E52D19E5F894FBE5FCC5" docLanguage="en" docName="zt03518p078.pdf" docOrigin="Zootaxa 3518" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Gonatodes rozei Rivero-Blanco & Schargel, 2012, sp. nov." docType="treatment" docVersion="6" lastPageNumber="76" masterDocId="FFDE0711FF97E5271972FFD0FFFBFFF2" masterDocTitle="A strikingly polychromatic new species of Gonatodes (Squamata: Sphaerodactylidae) from northern Venezuela" masterLastPageNumber="78" masterPageNumber="66" pageNumber="68" updateTime="1698317520111" updateUser="plazi">
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<mods:mods id="F36FC3798C4D16E51BFB4EF42FB8FA73" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="873F38CBAFF8B5811F9199140C4224E7">
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<mods:title id="816E3F9564EA00539BCB9E8A3FE84798">A strikingly polychromatic new species of Gonatodes (Squamata: Sphaerodactylidae) from northern Venezuela</mods:title>
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</mods:titleInfo>
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<mods:name id="DC4213DD5A914A5423759AC1B6BD7A79" type="personal">
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<mods:role id="8B2F598954275D3AF6339289BC95F183">
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<mods:roleTerm id="B2D5829575F4E947425338D9C81D2224">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="C0CA825E77B80BE0F21D73F703C5AF24">Rivero-Blanco, Carlos</mods:namePart>
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</mods:name>
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<mods:name id="AFD3C7B52B6AFDE040CA43BD8C45180E" type="personal">
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<mods:role id="FE5B5770C2B174C7E0A26DE3CBD90750">
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<mods:roleTerm id="FE4E899660BFC41471014185705C80AC">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="2EA15B84D184568E6037F75A39F01F9D">Schargel, Walter E.</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="4034817EB253A93A1A7DB04A46E70BD2">text</mods:typeOfResource>
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<mods:relatedItem id="40E062A5F3708C38947AD85890F847E8" type="host">
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<mods:titleInfo id="28C0F2405D270FE3D516C006071984F1">
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<mods:title id="AF26E9C3AE612ED01F8D92C10E75B386">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="DDC33F1AFDD5F797C4006CE43C56216C">
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<mods:date id="51DFD310424AB37D6A03385DCA031107">2012</mods:date>
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<mods:detail id="7CC53BDC49894971DDDB23229AE75E1E" type="volume">
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<mods:number id="7DB4402EF0F2E6EC0667F6D6AC80F1B3">3518</mods:number>
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</mods:detail>
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<mods:extent id="60F00F4D25DECAA3AEED55726EE77CFA" unit="page">
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<mods:start id="A8CE83140F2FE7293D51E2BCD3F6319A">66</mods:start>
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<mods:end id="06D8C0DDC4BFF5DF425AAEE1DA5EF3D6">78</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:classification id="145F5EBE249B7CD90F16CA8CF92DD14D">journal article</mods:classification>
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<mods:identifier id="AB15CA8FA92D3C9E1F81C93D17955D56" type="DOI">10.5281/zenodo.282620</mods:identifier>
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<mods:identifier id="378B10A29EBF74D438021ABFABFDF242" type="GBIF-Dataset">dc293202-7bfd-4ccd-85b6-fad297545b73</mods:identifier>
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<mods:identifier id="00AD4832BA087153885E9106349FBCFB" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="E897DBE3DCA8DB2804311BD47B4CE6CF" type="Zenodo-Dep">282620</mods:identifier>
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</mods:mods>
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<treatment id="03E77F69FF95E52D19E5F894FBE5FCC5" ID-DOI="http://doi.org/10.5281/zenodo.6167750" ID-GBIF-Taxon="119528821" ID-Zenodo-Dep="6167750" LSID="urn:lsid:plazi:treatment:03E77F69FF95E52D19E5F894FBE5FCC5" httpUri="http://treatment.plazi.org/id/03E77F69FF95E52D19E5F894FBE5FCC5" lastPageId="10" lastPageNumber="76" pageId="2" pageNumber="68">
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<subSubSection id="C3549DF4FF95E52519E5F894FEB1F88D" pageId="2" pageNumber="68" type="nomenclature">
|
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<paragraph id="8BF1CE7FFF95E52519E5F894FE45F8AC" blockId="2.[151,446,1860,1919]" box="[151,446,1860,1886]" pageId="2" pageNumber="68">
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<heading id="D0B97913FF95E52519E5F894FE45F8AC" bold="true" box="[151,446,1860,1886]" fontSize="11" level="1" pageId="2" pageNumber="68" reason="1">
|
||
<emphasis id="B93A126DFF95E52519E5F894FE45F8AC" bold="true" box="[151,446,1860,1886]" pageId="2" pageNumber="68">
|
||
<taxonomicName id="4C4EB5FCFF95E52519E5F894FEA2F8AC" box="[151,345,1860,1886]" class="Reptilia" family="Sphaerodactylidae" genus="Gonatodes" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="68" phylum="Chordata" rank="species" species="rozei" status="sp. nov.">
|
||
<emphasis id="B93A126DFF95E52519E5F894FEA2F8AC" bold="true" box="[151,345,1860,1886]" italics="true" pageId="2" pageNumber="68">Gonatodes rozei</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A209AF16FF95E5251813F894FE45F8AC" box="[353,446,1860,1886]" pageId="2" pageNumber="68" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BF1CE7FFF95E52519E5F8B7FEB1F88D" blockId="2.[151,446,1860,1919]" box="[151,330,1895,1919]" pageId="2" pageNumber="68">
|
||
(
|
||
<figureCitation id="1375D2FAFF95E52519EDF8B7FF14F88D" box="[159,239,1895,1919]" captionStart="FIGURE 2" captionStartId="3.[151,250,1844,1867]" captionTargetBox="[251,1340,359,1805]" captionTargetId="figure@3.[234,1353,343,1823]" captionTargetPageId="3" captionText="FIGURE 2. Adult male (top, holotype, MHNLS 20780) and adult female (bottom, UTA 60126) of Gonatodes rozei sp. nov. in life." httpUri="https://zenodo.org/record/282622/files/figure.png" pageId="2" pageNumber="68">Figs. 2</figureCitation>
|
||
,
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<figureCitation id="1375D2FAFF95E5251988F8B7FEF1F88D" box="[250,266,1895,1919]" captionStart="FIGURE 3" captionStartId="5.[151,250,1114,1137]" captionTargetBox="[175,1422,589,1102]" captionTargetId="figure@5.[175,1422,589,1102]" captionTargetPageId="5" captionText="FIGURE 3. Scalation pattern in the gular area of Gonatodes rozei sp. nov. (left; illustration based on MHNLS 5532 from Valle de Guanape, Anzoátegui, Venezuela) and the falconensis complex (right; illustration based on specimen of G. taniae, TCWC 59378, from Rancho Grande, Aragua, Venezuela)." httpUri="https://zenodo.org/record/282623/files/figure.png" pageId="2" pageNumber="68">3</figureCitation>
|
||
,
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<figureCitation id="1375D2FAFF95E5251864F8B7FEDDF88D" box="[278,294,1895,1919]" captionStart="FIGURE 4" captionStartId="7.[151,250,1978,2001]" captionTargetBox="[206,1388,184,1932]" captionTargetId="figure@7.[190,1395,159,1970]" captionTargetPageId="7" captionText="FIGURE 4. The four different male color morphs in Gonatodes rozei sp. nov., I-IV, arranged from top to bottom. Illustration by Emil Bröckl." httpUri="https://zenodo.org/record/282624/files/figure.png" pageId="2" pageNumber="68">4</figureCitation>
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,
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<figureCitation id="1375D2FAFF95E5251840F8B7FEBBF88D" box="[306,320,1895,1919]" captionStart="FIGURE 5. A" captionStartId="8.[151,250,1983,2006]" captionTargetBox="[151,1435,1309,1961]" captionTargetId="figure@8.[151,1436,1309,1962]" captionTargetPageId="8" captionText="FIGURE 5. A pair of Gonatodes rozei sp. nov. copulating in captivity." httpUri="https://zenodo.org/record/282625/files/figure.png" pageId="2" pageNumber="68">5</figureCitation>
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||
)
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3549DF4FF95E52319E5F860FCB4FE3D" lastPageId="4" lastPageNumber="70" pageId="2" pageNumber="68" type="materials_examined">
|
||
<paragraph id="8BF1CE7FFF95E52519E5F860FA84F819" blockId="2.[151,1435,1967,2027]" pageId="2" pageNumber="68">
|
||
<emphasis id="B93A126DFF95E52519E5F860FEF0F83A" bold="true" box="[151,267,1968,1992]" pageId="2" pageNumber="68">
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||
<typeStatus id="54F570DDFF95E52519E5F860FEFDF83A" box="[151,262,1968,1992]" pageId="2" pageNumber="68" type="holotype">Holotype</typeStatus>
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.
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</emphasis>
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<collectionCode id="ED5F56BAFF95E5251863F87FFE88F835" box="[273,371,1967,1991]" httpUri="http://grbio.org/cool/4kc6-e4ia" name="Coleccion de Mastozoologia, Museo de Historia Natural de La Salle" pageId="2" pageNumber="68">MHNLS</collectionCode>
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||
20780, an adult male, collected by G. Rivas and W. Schargel,
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<date id="FFF0E8BFFF95E5251D5AF87FFAF1F835" box="[1064,1290,1967,1991]" pageId="2" pageNumber="68" value="2006-09-17">September 17, 2006</date>
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,
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||
<quantity id="4CB6639AFF95E5251C67F87FFAB5F835" box="[1301,1358,1967,1991]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="2" pageNumber="68" unit="km" value="3.0">3 km</quantity>
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SW of Araguita (10 12’58’’ N, 66 29’07’’ W; ca.
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<quantity id="4CB6639AFF95E5251B00F803FD42F819" box="[626,697,2003,2027]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="2" pageNumber="68" unit="m" value="150.0">150 m</quantity>
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asl), near limit of Guatopo National Park, Miranda,
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<collectingCountry id="F3598EEFFF95E5251C76F803FA80F819" box="[1284,1403,2003,2027]" name="Venezuela" pageId="2" pageNumber="68">Venezuela</collectingCountry>
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.
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||
</paragraph>
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||
<paragraph id="8BF1CE7FFF94E52419B5FF48FDC4FECD" blockId="3.[151,1437,151,319]" pageId="3" pageNumber="69">
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||
<emphasis id="B93A126DFF94E52419B5FF48FEC4FF42" bold="true" box="[199,319,152,176]" pageId="3" pageNumber="69">
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||
<typeStatus id="54F570DDFF94E52419B5FF48FEC4FF42" box="[199,319,152,176]" pageId="3" pageNumber="69" type="paratype">Paratypes</typeStatus>
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||
</emphasis>
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(n=39).
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||
<emphasis id="B93A126DFF94E52418C1FF47FD9AFF42" bold="true" box="[435,609,151,176]" pageId="3" pageNumber="69">
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||
<collectingCountry id="F3598EEFFF94E52418C1FF47FD9AFF42" box="[435,609,151,176]" name="Venezuela" pageId="3" pageNumber="69">VENEZUELA</collectingCountry>
|
||
</emphasis>
|
||
:
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||
<emphasis id="B93A126DFF94E5241B06FF47FD05FF42" bold="true" box="[628,766,151,176]" pageId="3" pageNumber="69">MIRANDA</emphasis>
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||
: 32.9 Km N of Altagracia de Orituco, Guatopo National Park:
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<collectionCode id="ED5F56BAFF94E52419ADFF6CFECBFF26" box="[223,304,188,212]" httpUri="http://grbio.org/cool/dj6p-xxc3" name="Texas Cooperative Wildlife Collection" pageId="3" pageNumber="69">TCWC</collectionCode>
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||
59300–59306. Guatopo National Park:
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||
<collectionCode id="ED5F56BAFF94E5241A7EFF6CFC88FF26" box="[780,883,188,212]" httpUri="http://grbio.org/cool/7nxg-etxt" name="Museo de Biologia de la Universidad Central de Venezuela" pageId="3" pageNumber="69">MBUCV</collectionCode>
|
||
5044,
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||
<collectionCode id="ED5F56BAFF94E5241AB9FF6CFBD6FF26" box="[971,1069,188,212]" httpUri="http://grbio.org/cool/4kc6-e4ia" name="Coleccion de Mastozoologia, Museo de Historia Natural de La Salle" pageId="3" pageNumber="69">MHNLS</collectionCode>
|
||
<date id="FFF0E8BFFF94E5241D48FF6CFB41FF26" box="[1082,1210,188,212]" pageId="3" pageNumber="69" value="1666" valueMax="1669">1666–1669</date>
|
||
,
|
||
<date id="FFF0E8BFFF94E5241DB9FF6CFAB7FF26" box="[1227,1356,188,212]" pageId="3" pageNumber="69" value="1674" valueMax="1677">1674–1677</date>
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||
, 2082.
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||
<emphasis id="B93A126DFF94E52419E5FF0FFEDCFF0A" bold="true" box="[151,295,223,248]" pageId="3" pageNumber="69">GUÁRICO:</emphasis>
|
||
<quantity id="4CB6639AFF94E5241846FF0FFE88FF05" box="[308,371,223,247]" metricMagnitude="3" metricUnit="m" metricValue="5.0" pageId="3" pageNumber="69" unit="km" value="5.0">5 km</quantity>
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E of “Puesto La Colonia,” Guatopo National Park:
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<collectionCode id="ED5F56BAFF94E5241A98FF0FFBC0FF05" box="[1002,1083,223,247]" httpUri="http://grbio.org/cool/dj6p-xxc3" name="Texas Cooperative Wildlife Collection" pageId="3" pageNumber="69">TCWC</collectionCode>
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||
59319–59320.
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<quantity id="4CB6639AFF94E5241D87FF0FFAACFF05" box="[1269,1367,223,247]" metricMagnitude="4" metricUnit="m" metricValue="2.52" pageId="3" pageNumber="69" unit="km" value="25.2">25.2 km</quantity>
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N of Altagracia de Orituco, Guatopo National Park:
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<collectionCode id="ED5F56BAFF94E5241BC0FED4FCF8FEEE" box="[690,771,260,284]" httpUri="http://grbio.org/cool/dj6p-xxc3" name="Texas Cooperative Wildlife Collection" pageId="3" pageNumber="69">TCWC</collectionCode>
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59318. Hacienda La Elvira, Guatopo National Park:
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<collectionCode id="ED5F56BAFF94E5241C16FED4FA67FEEE" LSID="urn:lsid:biocol.org:col:34873" box="[1380,1436,260,284]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34873" name="University of Texas at Arlington" pageId="3" pageNumber="69">UTA</collectionCode>
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60126,
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<collectionCode id="ED5F56BAFF94E5241999FEF7FEC7FECD" box="[235,316,295,319]" httpUri="http://grbio.org/cool/dj6p-xxc3" name="Texas Cooperative Wildlife Collection" pageId="3" pageNumber="69">TCWC</collectionCode>
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||
59309–59317, 59321.
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||
</paragraph>
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||
<caption id="DF319EF7FF94E52419E5F8E4FF3BF89A" httpUri="https://zenodo.org/record/282622/files/figure.png" pageId="3" pageNumber="69" targetBox="[251,1340,359,1805]" targetPageId="3">
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||
<paragraph id="8BF1CE7FFF94E52419E5F8E4FF3BF89A" blockId="3.[151,1436,1844,1896]" pageId="3" pageNumber="69">
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||
<emphasis id="B93A126DFF94E52419E5F8E4FEE9F8BB" bold="true" box="[151,274,1844,1867]" pageId="3" pageNumber="69">FIGURE 2.</emphasis>
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||
Adult male (top, holotype, MHNLS 20780) and adult female (bottom, UTA 60126) of
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<taxonomicName id="4C4EB5FCFF94E5241DF7F8E4FAD1F8B8" box="[1157,1322,1844,1866]" class="Reptilia" family="Sphaerodactylidae" genus="Gonatodes" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="69" phylum="Chordata" rank="species" species="rozei" status="sp. nov.">
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||
<emphasis id="B93A126DFF94E5241DF7F8E4FAD1F8B8" box="[1157,1322,1844,1866]" italics="true" pageId="3" pageNumber="69">Gonatodes rozei</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B93A126DFF94E5241C43F8E5FA7AF8B8" bold="true" box="[1329,1409,1845,1866]" pageId="3" pageNumber="69">
|
||
<taxonomicNameLabel id="A209AF16FF94E5241C43F8E5FA7AF8B8" box="[1329,1409,1845,1866]" pageId="3" pageNumber="69" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
in life.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BF1CE7FFF94E52319B5F845FCB4FE3D" blockId="3.[151,1437,1941,2037]" lastBlockId="4.[151,1437,151,2013]" lastPageId="4" lastPageNumber="70" pageId="3" pageNumber="69">
|
||
<emphasis id="B93A126DFF94E52419B5F845FE3BF85C" bold="true" box="[199,448,1941,1966]" pageId="3" pageNumber="69">Referred specimens</emphasis>
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(n=83).
|
||
<emphasis id="B93A126DFF94E5241B4FF845FC28F85C" bold="true" box="[573,979,1941,1966]" pageId="3" pageNumber="69">
|
||
<collectingCountry id="F3598EEFFF94E5241B4FF845FD14F85C" box="[573,751,1941,1966]" name="Venezuela" pageId="3" pageNumber="69">VENEZUELA</collectingCountry>
|
||
: ANZOÁTEGUI:
|
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</emphasis>
|
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Valle de Guanape:
|
||
<collectionCode id="ED5F56BAFF94E5241D94F845FAB3F85F" box="[1254,1352,1941,1965]" httpUri="http://grbio.org/cool/4kc6-e4ia" name="Coleccion de Mastozoologia, Museo de Historia Natural de La Salle" pageId="3" pageNumber="69">MHNLS</collectionCode>
|
||
5532.
|
||
<emphasis id="B93A126DFF94E52419E5F869FEDCF820" bold="true" box="[151,295,1977,2002]" pageId="3" pageNumber="69">GUÁRICO:</emphasis>
|
||
Las Palmas, Altagracia de Orituco:
|
||
<collectionCode id="ED5F56BAFF94E5241B9EF869FCA8F823" box="[748,851,1977,2001]" httpUri="http://grbio.org/cool/7nxg-etxt" name="Museo de Biologia de la Universidad Central de Venezuela" pageId="3" pageNumber="69">MBUCV</collectionCode>
|
||
3253.
|
||
<emphasis id="B93A126DFF94E5241ADDF869FBB8F820" bold="true" box="[943,1091,1977,2002]" pageId="3" pageNumber="69">MIRANDA:</emphasis>
|
||
Araira, Hacienda La Ceiba:
|
||
<collectionCode id="ED5F56BAFF94E52419E5F80DFF02F807" box="[151,249,2013,2037]" httpUri="http://grbio.org/cool/4kc6-e4ia" name="Coleccion de Mastozoologia, Museo de Historia Natural de La Salle" pageId="3" pageNumber="69">MHNLS</collectionCode>
|
||
15177–15178. Capaya:
|
||
<collectionCode id="ED5F56BAFF94E5241B62F80DFD89F807" box="[528,626,2013,2037]" httpUri="http://grbio.org/cool/4kc6-e4ia" name="Coleccion de Mastozoologia, Museo de Historia Natural de La Salle" pageId="3" pageNumber="69">MHNLS</collectionCode>
|
||
2322,
|
||
<date id="FFF0E8BFFF94E5241BB1F80DFCBFF807" box="[707,836,2013,2037]" pageId="3" pageNumber="69" value="2324" valueMax="2325">2324–2325</date>
|
||
, 5143–5145, 5854,
|
||
<collectionCode id="ED5F56BAFF94E5241D55F80DFB81F807" box="[1063,1146,2013,2037]" httpUri="http://grbio.org/cool/sw8c-0xp3" name="Museo de Ciencias Naturales" pageId="3" pageNumber="69">MCNC</collectionCode>
|
||
3719–3726, 3776. Taica:
|
||
<collectionCode id="ED5F56BAFF93E52319E5FF47FF02FF5D" box="[151,249,151,175]" httpUri="http://grbio.org/cool/4kc6-e4ia" name="Coleccion de Mastozoologia, Museo de Historia Natural de La Salle" pageId="4" pageNumber="70">MHNLS</collectionCode>
|
||
2162. Carenero:
|
||
<collectionCode id="ED5F56BAFF93E52318C8FF47FDDAFF5D" box="[442,545,151,175]" httpUri="http://grbio.org/cool/7nxg-etxt" name="Museo de Biologia de la Universidad Central de Venezuela" pageId="4" pageNumber="70">MBUCV</collectionCode>
|
||
5045, Carretera Chuspa – Higuerote,
|
||
<collectionCode id="ED5F56BAFF93E5231AB8FF47FBD7FF5D" box="[970,1068,151,175]" httpUri="http://grbio.org/cool/4kc6-e4ia" name="Coleccion de Mastozoologia, Museo de Historia Natural de La Salle" pageId="4" pageNumber="70">MHNLS</collectionCode>
|
||
3775. Cueva 105, El Encantado:
|
||
<collectionCode id="ED5F56BAFF93E52319E5FF6CFF02FF26" box="[151,249,188,212]" httpUri="http://grbio.org/cool/4kc6-e4ia" name="Coleccion de Mastozoologia, Museo de Historia Natural de La Salle" pageId="4" pageNumber="70">MHNLS</collectionCode>
|
||
1982. Cueva Alfredo Jahn:
|
||
<collectionCode id="ED5F56BAFF93E5231B40FF6CFD62FF26" box="[562,665,188,212]" httpUri="http://grbio.org/cool/7nxg-etxt" name="Museo de Biologia de la Universidad Central de Venezuela" pageId="4" pageNumber="70">MBUCV</collectionCode>
|
||
8415,
|
||
<collectionCode id="ED5F56BAFF93E5231B97FF6CFCBCFF26" box="[741,839,188,212]" httpUri="http://grbio.org/cool/4kc6-e4ia" name="Coleccion de Mastozoologia, Museo de Historia Natural de La Salle" pageId="4" pageNumber="70">MHNLS</collectionCode>
|
||
2323. Cueva Cruxent, Curiepe:
|
||
<collectionCode id="ED5F56BAFF93E5231DDDFF6CFAEAFF26" box="[1199,1297,188,212]" httpUri="http://grbio.org/cool/4kc6-e4ia" name="Coleccion de Mastozoologia, Museo de Historia Natural de La Salle" pageId="4" pageNumber="70">MHNLS</collectionCode>
|
||
2248, 2250. Cueva de la Diaclesa, El Encantado:
|
||
<collectionCode id="ED5F56BAFF93E5231B44FF0FFD63FF05" box="[566,664,223,247]" httpUri="http://grbio.org/cool/4kc6-e4ia" name="Coleccion de Mastozoologia, Museo de Historia Natural de La Salle" pageId="4" pageNumber="70">MHNLS</collectionCode>
|
||
1985. Cueva Graciliano, Curiepe:
|
||
<collectionCode id="ED5F56BAFF93E5231D52FF0FFB79FF05" box="[1056,1154,223,247]" httpUri="http://grbio.org/cool/4kc6-e4ia" name="Coleccion de Mastozoologia, Museo de Historia Natural de La Salle" pageId="4" pageNumber="70">MHNLS</collectionCode>
|
||
2249. Cueva del túnel 4, Los Naranjos:
|
||
<collectionCode id="ED5F56BAFF93E5231837FED4FE5CFEEE" box="[325,423,260,284]" httpUri="http://grbio.org/cool/4kc6-e4ia" name="Coleccion de Mastozoologia, Museo de Historia Natural de La Salle" pageId="4" pageNumber="70">MHNLS</collectionCode>
|
||
<date id="FFF0E8BFFF93E52318C0FED4FDC9FEEE" box="[434,562,260,284]" pageId="4" pageNumber="70" value="1883" valueMax="1884">1883–1884</date>
|
||
, 1981, 2161. El Encantado:
|
||
<collectionCode id="ED5F56BAFF93E5231AF7FED4FC17FEEE" box="[901,1004,260,284]" httpUri="http://grbio.org/cool/7nxg-etxt" name="Museo de Biologia de la Universidad Central de Venezuela" pageId="4" pageNumber="70">MBUCV</collectionCode>
|
||
5043. Los Naranjos:
|
||
<collectionCode id="ED5F56BAFF93E5231D9DFED4FAAAFEEE" box="[1263,1361,260,284]" httpUri="http://grbio.org/cool/4kc6-e4ia" name="Coleccion de Mastozoologia, Museo de Historia Natural de La Salle" pageId="4" pageNumber="70">MHNLS</collectionCode>
|
||
2161. Planta Eléctrica Los Naranjos, Río Guaire:
|
||
<collectionCode id="ED5F56BAFF93E5231BD1FEF8FD0CFECD" box="[675,759,296,319]" pageId="4" pageNumber="70">BMNH</collectionCode>
|
||
1967.256–257,
|
||
<collectionCode id="ED5F56BAFF93E5231AC8FEF7FBF6FECD" box="[954,1037,295,319]" httpUri="http://grbio.org/cool/sw8c-0xp3" name="Museo de Ciencias Naturales" pageId="4" pageNumber="70">MCNC</collectionCode>
|
||
3754–3759. Quebrada Rebollero, Chuspa:
|
||
<collectionCode id="ED5F56BAFF93E5231989FE9CFEB6FE96" box="[251,333,332,356]" httpUri="http://grbio.org/cool/sw8c-0xp3" name="Museo de Ciencias Naturales" pageId="4" pageNumber="70">MCNC</collectionCode>
|
||
3735. Quebrada Urba, Carretera Panaquire – El Sapo:
|
||
<collectionCode id="ED5F56BAFF93E5231AB1FE9CFBDEFE96" box="[963,1061,332,356]" httpUri="http://grbio.org/cool/4kc6-e4ia" name="Coleccion de Mastozoologia, Museo de Historia Natural de La Salle" pageId="4" pageNumber="70">MHNLS</collectionCode>
|
||
16780, 16786.
|
||
<emphasis id="B93A126DFF93E5231DAAFE9BFAABFE96" bold="true" box="[1240,1360,331,356]" pageId="4" pageNumber="70">VA R G A S:</emphasis>
|
||
Cerro Naiguatá, “two hours from toma de agua:”
|
||
<collectionCode id="ED5F56BAFF93E5231BF3FEBFFD18FE75" box="[641,739,367,391]" httpUri="http://grbio.org/cool/4kc6-e4ia" name="Coleccion de Mastozoologia, Museo de Historia Natural de La Salle" pageId="4" pageNumber="70">MHNLS</collectionCode>
|
||
3774. Hacienda El Limón:
|
||
<collectionCode id="ED5F56BAFF93E5231D6DFEBFFB7AFE75" box="[1055,1153,367,391]" httpUri="http://grbio.org/cool/4kc6-e4ia" name="Coleccion de Mastozoologia, Museo de Historia Natural de La Salle" pageId="4" pageNumber="70">MHNLS</collectionCode>
|
||
2060, 4296, 4297, 4362, 4363, 4392, 4532, 4651, 4690, 4813. Pico Naiguatá:
|
||
<collectionCode id="ED5F56BAFF93E5231B8CFE44FCAAFE5E" box="[766,849,404,428]" httpUri="http://grbio.org/cool/sw8c-0xp3" name="Museo de Ciencias Naturales" pageId="4" pageNumber="70">MCNC</collectionCode>
|
||
3733–3734. Piedra del Zamuro, Carayaca:
|
||
<collectionCode id="ED5F56BAFF93E5231C3BFE44FA67FE5E" box="[1353,1436,404,428]" httpUri="http://grbio.org/cool/sw8c-0xp3" name="Museo de Ciencias Naturales" pageId="4" pageNumber="70">MCNC</collectionCode>
|
||
3736–3753. Planta Eléctrica de Naiguatá:
|
||
<collectionCode id="ED5F56BAFF93E5231B1DFE67FD3AFE3D" box="[623,705,439,463]" httpUri="http://grbio.org/cool/sw8c-0xp3" name="Museo de Ciencias Naturales" pageId="4" pageNumber="70">MCNC</collectionCode>
|
||
3727–3732.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3549DF4FF93E52319B5FE0CFD33FDEA" pageId="4" pageNumber="70" type="etymology">
|
||
<paragraph id="8BF1CE7FFF93E52319B5FE0CFD33FDEA" blockId="4.[151,1437,151,2013]" pageId="4" pageNumber="70">
|
||
<emphasis id="B93A126DFF93E52319B5FE0CFEBDFE06" bold="true" box="[199,326,476,500]" pageId="4" pageNumber="70">Etymology</emphasis>
|
||
. We take great pleasure in naming the new species after Dr. Janis Roze, a pioneer of Venezuelan herpetology and a mentor to CRB for many years.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3549DF4FF93E52219B5FDF3FEC8FAC8" lastPageId="5" lastPageNumber="71" pageId="4" pageNumber="70" type="diagnosis">
|
||
<paragraph id="8BF1CE7FFF93E52319B5FDF3FAF0FC35" blockId="4.[151,1437,151,2013]" pageId="4" pageNumber="70">
|
||
<emphasis id="B93A126DFF93E52319B5FDF3FEBAFDCE" bold="true" box="[199,321,547,572]" pageId="4" pageNumber="70">Diagnosis.</emphasis>
|
||
The new species can be distinguished from all congeners by a combination of the following characters: (1) large size (largest specimen examined has a SVL of
|
||
<quantity id="4CB6639AFF93E5231AE0FD97FC00FDAD" box="[914,1019,583,607]" metricMagnitude="-2" metricUnit="m" metricValue="5.91" pageId="4" pageNumber="70" unit="mm" value="59.1">59.1 mm</quantity>
|
||
), (2) 78–100 scales around midbody, (3) single, elongated supraciliary spine absent, (4) three or four lateral scale rows on the digits, (5) subcaudal pattern
|
||
<typeStatus id="54F570DDFF93E523199CFD40FEDDFD5A" box="[238,294,656,680]" pageId="4" pageNumber="70">Type</typeStatus>
|
||
B, (6) gular scales posterior to the postmentals with a sharp transition to small granular scales, (7) males with blue iris (females blue or yellow), and (8) males typically with an immaculate bright yellow to reddish orange hood. The large size of
|
||
<taxonomicName id="4C4EB5FCFF93E5231B78FD08FD99FD02" box="[522,610,728,752]" class="Reptilia" family="Sphaerodactylidae" genus="Gonatodes" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="70" phylum="Chordata" rank="species" species="rozei">
|
||
<emphasis id="B93A126DFF93E5231B78FD08FDE0FD1D" box="[522,539,728,751]" italics="true" pageId="4" pageNumber="70">G</emphasis>
|
||
.
|
||
<emphasis id="B93A126DFF93E5231B5EFD09FD99FD02" box="[556,610,729,752]" italics="true" pageId="4" pageNumber="70">rozei</emphasis>
|
||
</taxonomicName>
|
||
readily distinguishes this species from the small species in the genus, namely
|
||
<taxonomicName id="4C4EB5FCFF93E5231980FD2DFE6FFCE6" box="[242,404,764,788]" class="Reptilia" family="Sphaerodactylidae" genus="Gonatodes" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="70" phylum="Chordata" rank="species" species="albogularis">
|
||
<emphasis id="B93A126DFF93E5231980FD2DFEF8FCE6" box="[242,259,765,788]" italics="true" pageId="4" pageNumber="70">G</emphasis>
|
||
.
|
||
<emphasis id="B93A126DFF93E5231861FD2CFE6FFCE6" box="[275,404,764,788]" italics="true" pageId="4" pageNumber="70">albogularis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C4EB5FCFF93E52318D6FD2DFDC3FCE6" box="[420,568,764,788]" class="Reptilia" family="Sphaerodactylidae" genus="Gonatodes" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="70" phylum="Chordata" rank="species" species="antillensis">
|
||
<emphasis id="B93A126DFF93E52318D6FD2DFE4EFCE6" box="[420,437,765,788]" italics="true" pageId="4" pageNumber="70">G</emphasis>
|
||
.
|
||
<emphasis id="B93A126DFF93E52318B7FD2CFDC3FCE6" box="[453,568,764,788]" italics="true" pageId="4" pageNumber="70">antillensis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C4EB5FCFF93E5231B3BFD2DFCF8FCE6" box="[585,771,764,788]" class="Reptilia" family="Sphaerodactylidae" genus="Gonatodes" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="70" phylum="Chordata" rank="species" species="atricucullaris">
|
||
<emphasis id="B93A126DFF93E5231B3BFD2DFDA1FCE6" box="[585,602,765,788]" italics="true" pageId="4" pageNumber="70">G</emphasis>
|
||
.
|
||
<emphasis id="B93A126DFF93E5231B18FD2CFCF8FCE6" box="[618,771,764,788]" italics="true" pageId="4" pageNumber="70">atricucullaris</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C4EB5FCFF93E5231A60FD2DFC36FCE6" box="[786,973,764,788]" class="Reptilia" family="Sphaerodactylidae" genus="Gonatodes" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="70" phylum="Chordata" rank="species" species="caudiscutatus">
|
||
<emphasis id="B93A126DFF93E5231A60FD2DFCD8FCE6" box="[786,803,765,788]" italics="true" pageId="4" pageNumber="70">G</emphasis>
|
||
.
|
||
<emphasis id="B93A126DFF93E5231A41FD2CFC36FCE6" box="[819,973,764,788]" italics="true" pageId="4" pageNumber="70">caudiscutatus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C4EB5FCFF93E5231AAFFD2DFBA8FCE6" box="[989,1107,764,788]" class="Reptilia" family="Sphaerodactylidae" genus="Gonatodes" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="70" phylum="Chordata" rank="species" species="daudini">
|
||
<emphasis id="B93A126DFF93E5231AAFFD2DFC15FCE6" box="[989,1006,765,788]" italics="true" pageId="4" pageNumber="70">G</emphasis>
|
||
.
|
||
<emphasis id="B93A126DFF93E5231A8CFD2CFBA8FCE6" box="[1022,1107,764,788]" italics="true" pageId="4" pageNumber="70">daudini</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C4EB5FCFF93E5231D16FD2DFB28FCE6" box="[1124,1235,764,788]" class="Reptilia" family="Sphaerodactylidae" genus="Gonatodes" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="70" phylum="Chordata" rank="species" species="eladioi">
|
||
<emphasis id="B93A126DFF93E5231D16FD2DFB8EFCE6" box="[1124,1141,765,788]" italics="true" pageId="4" pageNumber="70">G</emphasis>
|
||
.
|
||
<emphasis id="B93A126DFF93E5231DF7FD2CFB28FCE6" box="[1157,1235,764,788]" italics="true" pageId="4" pageNumber="70">eladioi</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C4EB5FCFF93E5231D91FD2DFA8FFCE6" box="[1251,1396,764,788]" class="Reptilia" family="Sphaerodactylidae" genus="Gonatodes" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="70" phylum="Chordata" rank="species" species="humeralis">
|
||
<emphasis id="B93A126DFF93E5231D91FD2DFB0FFCE6" box="[1251,1268,765,788]" italics="true" pageId="4" pageNumber="70">G</emphasis>
|
||
.
|
||
<emphasis id="B93A126DFF93E5231C76FD2CFA8FFCE6" box="[1284,1396,764,788]" italics="true" pageId="4" pageNumber="70">humeralis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C4EB5FCFF93E5231CF6FD2DFF2CFCCA" class="Reptilia" family="Sphaerodactylidae" genus="Gonatodes" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="70" phylum="Chordata" rank="species" species="ligiae">
|
||
<emphasis id="B93A126DFF93E5231CF6FD2DFA6EFCE6" box="[1412,1429,765,788]" italics="true" pageId="4" pageNumber="70">G</emphasis>
|
||
.
|
||
<emphasis id="B93A126DFF93E52319E5FCF0FF2CFCCA" box="[151,215,800,824]" italics="true" pageId="4" pageNumber="70">ligiae</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C4EB5FCFF93E5231994FCF0FEA9FCCA" box="[230,338,800,824]" class="Reptilia" family="Sphaerodactylidae" genus="Gonatodes" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="70" phylum="Chordata" rank="species" species="petersi">
|
||
<emphasis id="B93A126DFF93E5231994FCF0FF0CFCC5" box="[230,247,800,823]" italics="true" pageId="4" pageNumber="70">G</emphasis>
|
||
.
|
||
<emphasis id="B93A126DFF93E5231874FCF1FEA9FCCA" box="[262,338,801,824]" italics="true" pageId="4" pageNumber="70">petersi</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="4C4EB5FCFF93E52318E1FCF0FDFDFCCA" box="[403,518,800,824]" class="Reptilia" family="Sphaerodactylidae" genus="Gonatodes" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="70" phylum="Chordata" rank="species" species="vittatus">
|
||
<emphasis id="B93A126DFF93E52318E1FCF0FE5FFCC5" box="[403,420,800,823]" italics="true" pageId="4" pageNumber="70">G</emphasis>
|
||
.
|
||
<emphasis id="B93A126DFF93E52318C6FCF1FDFDFCCA" box="[436,518,801,824]" italics="true" pageId="4" pageNumber="70">vittatus</emphasis>
|
||
</taxonomicName>
|
||
, none of which exceed
|
||
<quantity id="4CB6639AFF93E5231A6AFCCFFC93FCCA" box="[792,872,799,824]" metricMagnitude="-2" metricUnit="m" metricValue="4.2" pageId="4" pageNumber="70" unit="mm" value="42.0">42 mm</quantity>
|
||
in SVL.
|
||
<taxonomicName id="4C4EB5FCFF93E5231AA1FCF0FB77FCCA" box="[979,1164,800,824]" class="Reptilia" family="Sphaerodactylidae" genus="Gonatodes" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="70" phylum="Chordata" rank="species" species="rozei">
|
||
<emphasis id="B93A126DFF93E5231AA1FCF0FB77FCCA" box="[979,1164,800,824]" italics="true" pageId="4" pageNumber="70">Gonatodes rozei</emphasis>
|
||
</taxonomicName>
|
||
further differs from the small species, except
|
||
<taxonomicName id="4C4EB5FCFF93E52318E6FC95FDABFCAE" box="[404,592,836,860]" class="Reptilia" family="Sphaerodactylidae" genus="Gonatodes" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="70" phylum="Chordata" rank="species" species="caudiscutatus">
|
||
<emphasis id="B93A126DFF93E52318E6FC95FE5EFCAE" box="[404,421,837,860]" italics="true" pageId="4" pageNumber="70">G</emphasis>
|
||
.
|
||
<emphasis id="B93A126DFF93E52318C4FC94FDABFCAE" box="[438,592,836,860]" italics="true" pageId="4" pageNumber="70">caudiscutatus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C4EB5FCFF93E5231BFFFC95FD14FCAE" box="[653,751,836,860]" class="Reptilia" family="Sphaerodactylidae" genus="Gonatodes" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="70" phylum="Chordata" rank="species" species="ligiae">
|
||
<emphasis id="B93A126DFF93E5231BFFFC95FD65FCAE" box="[653,670,837,860]" italics="true" pageId="4" pageNumber="70">G</emphasis>
|
||
.
|
||
<emphasis id="B93A126DFF93E5231BC2FC94FD14FCAE" box="[688,751,836,860]" italics="true" pageId="4" pageNumber="70">ligiae</emphasis>
|
||
</taxonomicName>
|
||
, in having three (rarely) or four (typically) lateral rows of scales on the distal portion of the digits as opposed to two. Males of the new species generally have a solid yellow or reddish yellow head whereas in
|
||
<taxonomicName id="4C4EB5FCFF93E5231B51FC5DFD26FC56" box="[547,733,908,932]" class="Reptilia" family="Sphaerodactylidae" genus="Gonatodes" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="70" phylum="Chordata" rank="species" species="caudiscutatus">
|
||
<emphasis id="B93A126DFF93E5231B51FC5DFDCFFC56" box="[547,564,909,932]" italics="true" pageId="4" pageNumber="70">G</emphasis>
|
||
.
|
||
<emphasis id="B93A126DFF93E5231B31FC5CFD26FC56" box="[579,733,908,932]" italics="true" pageId="4" pageNumber="70">caudiscutatus</emphasis>
|
||
</taxonomicName>
|
||
the head is always covered with yellow stripes. Specimens of
|
||
<taxonomicName id="4C4EB5FCFF93E52319E5FC60FF10FC3A" box="[151,235,944,968]" class="Reptilia" family="Sphaerodactylidae" genus="Gonatodes" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="70" phylum="Chordata" rank="species" species="rozei">
|
||
<emphasis id="B93A126DFF93E52319E5FC60FF53FC35" box="[151,168,944,967]" italics="true" pageId="4" pageNumber="70">G</emphasis>
|
||
.
|
||
<emphasis id="B93A126DFF93E52319C7FC61FF10FC3A" box="[181,235,945,968]" italics="true" pageId="4" pageNumber="70">rozei</emphasis>
|
||
</taxonomicName>
|
||
have fewer scales around midbody compared to
|
||
<taxonomicName id="4C4EB5FCFF93E5231A63FC60FC94FC3A" box="[785,879,944,968]" class="Reptilia" family="Sphaerodactylidae" genus="Gonatodes" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="70" phylum="Chordata" rank="species" species="ligiae">
|
||
<emphasis id="B93A126DFF93E5231A63FC60FCD9FC35" box="[785,802,944,967]" italics="true" pageId="4" pageNumber="70">G</emphasis>
|
||
.
|
||
<emphasis id="B93A126DFF93E5231A42FC60FC94FC3A" box="[816,879,944,968]" italics="true" pageId="4" pageNumber="70">ligiae</emphasis>
|
||
</taxonomicName>
|
||
(78–100 vs. 109–128, respectively).
|
||
</paragraph>
|
||
<paragraph id="8BF1CE7FFF93E52319B5FC04FAD8F9F5" blockId="4.[151,1437,151,2013]" pageId="4" pageNumber="70">
|
||
Among the large and medium size species (character states in parenthesis),
|
||
<taxonomicName id="4C4EB5FCFF93E5231D65FC05FB96FC1E" box="[1047,1133,981,1004]" class="Reptilia" family="Sphaerodactylidae" genus="Gonatodes" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="70" phylum="Chordata" rank="species" species="rozei">
|
||
<emphasis id="B93A126DFF93E5231D65FC05FBD3FC1E" box="[1047,1064,981,1004]" italics="true" pageId="4" pageNumber="70">G</emphasis>
|
||
.
|
||
<emphasis id="B93A126DFF93E5231D45FC05FB96FC1E" box="[1079,1133,981,1004]" italics="true" pageId="4" pageNumber="70">rozei</emphasis>
|
||
</taxonomicName>
|
||
differs from
|
||
<taxonomicName id="4C4EB5FCFF93E5231C71FC05FA8DFC1E" box="[1283,1398,980,1004]" class="Reptilia" family="Sphaerodactylidae" genus="Gonatodes" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="70" phylum="Chordata" rank="species" species="astralis">
|
||
<emphasis id="B93A126DFF93E5231C71FC05FAEFFC1E" box="[1283,1300,981,1004]" italics="true" pageId="4" pageNumber="70">G</emphasis>
|
||
.
|
||
<emphasis id="B93A126DFF93E5231C50FC04FA8DFC1E" box="[1314,1398,980,1004]" italics="true" pageId="4" pageNumber="70">astralis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C4EB5FCFF93E5231CF6FC05FE98FBE2" class="Reptilia" family="Sphaerodactylidae" genus="Gonatodes" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="70" phylum="Chordata" rank="species" species="alexandermendesi">
|
||
<emphasis id="B93A126DFF93E5231CF6FC05FA6EFC1E" box="[1412,1429,981,1004]" italics="true" pageId="4" pageNumber="70">G</emphasis>
|
||
.
|
||
<emphasis id="B93A126DFF93E52319E5FC28FE98FBE2" box="[151,355,1016,1040]" italics="true" pageId="4" pageNumber="70">alexandermendesi</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C4EB5FCFF93E5231803FC28FE00FBE2" box="[369,507,1016,1040]" class="Reptilia" family="Sphaerodactylidae" genus="Gonatodes" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="70" phylum="Chordata" rank="species" species="hasemani">
|
||
<emphasis id="B93A126DFF93E5231803FC28FE79FBFD" box="[369,386,1016,1039]" italics="true" pageId="4" pageNumber="70">G</emphasis>
|
||
.
|
||
<emphasis id="B93A126DFF93E52318E2FC28FE00FBE2" box="[400,507,1016,1040]" italics="true" pageId="4" pageNumber="70">hasemani</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="4C4EB5FCFF93E5231B48FC28FD1DFBE2" box="[570,742,1016,1040]" class="Reptilia" family="Sphaerodactylidae" genus="Gonatodes" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="70" phylum="Chordata" rank="species" species="superciliaris">
|
||
<emphasis id="B93A126DFF93E5231B48FC28FDB0FBFD" box="[570,587,1016,1039]" italics="true" pageId="4" pageNumber="70">G</emphasis>
|
||
.
|
||
<emphasis id="B93A126DFF93E5231B2BFC28FD1DFBE2" box="[601,742,1016,1040]" italics="true" pageId="4" pageNumber="70">superciliaris</emphasis>
|
||
</taxonomicName>
|
||
in lacking an elongated supraciliary spine (a single elongated spine is present). It differs from
|
||
<taxonomicName id="4C4EB5FCFF93E523188FFBCDFD73FBC6" box="[509,648,1052,1076]" class="Reptilia" family="Sphaerodactylidae" genus="Gonatodes" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="70" phylum="Chordata" rank="species" species="annularis">
|
||
<emphasis id="B93A126DFF93E523188FFBCDFDF5FBC6" box="[509,526,1053,1076]" italics="true" pageId="4" pageNumber="70">G</emphasis>
|
||
.
|
||
<emphasis id="B93A126DFF93E5231B6EFBCCFD73FBC6" box="[540,648,1052,1076]" italics="true" pageId="4" pageNumber="70">annularis</emphasis>
|
||
</taxonomicName>
|
||
in having subcaudal pattern
|
||
<typeStatus id="54F570DDFF93E5231AB5FBCDFC0CFBC7" box="[967,1015,1053,1077]" pageId="4" pageNumber="70">type</typeStatus>
|
||
B (
|
||
<typeStatus id="54F570DDFF93E5231D6DFBCCFBAAFBC6" box="[1055,1105,1052,1076]" pageId="4" pageNumber="70">type</typeStatus>
|
||
A or E) and a body devoid of yellow reticulations in males (body almost always covered with yellow reticulations in males). It differs from
|
||
<taxonomicName id="4C4EB5FCFF93E5231CF6FB90FEF8FB8E" class="Reptilia" family="Sphaerodactylidae" genus="Gonatodes" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="70" phylum="Chordata" rank="species" species="infernalis">
|
||
<emphasis id="B93A126DFF93E5231CF6FB90FA6EFBA5" box="[1412,1429,1088,1111]" italics="true" pageId="4" pageNumber="70">G</emphasis>
|
||
.
|
||
<emphasis id="B93A126DFF93E52319E5FBB4FEF8FB8E" box="[151,259,1124,1148]" italics="true" pageId="4" pageNumber="70">infernalis</emphasis>
|
||
</taxonomicName>
|
||
in having subcaudal pattern
|
||
<typeStatus id="54F570DDFF93E5231B30FBB5FD89FB8F" box="[578,626,1125,1149]" pageId="4" pageNumber="70">type</typeStatus>
|
||
B (
|
||
<typeStatus id="54F570DDFF93E5231BE8FBB4FD37FB8E" box="[666,716,1124,1148]" pageId="4" pageNumber="70">type</typeStatus>
|
||
C) and having females with a brown dorsal coloration with dark and pale markings and a pale vertebral stripe (females with immaculate brown dorsal coloration; males not known for this species).
|
||
<taxonomicName id="4C4EB5FCFF93E523182AFB7CFDEBFB36" box="[344,528,1196,1220]" class="Reptilia" family="Sphaerodactylidae" genus="Gonatodes" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="70" phylum="Chordata" rank="species" species="rozei">
|
||
<emphasis id="B93A126DFF93E523182AFB7CFDEBFB36" box="[344,528,1196,1220]" italics="true" pageId="4" pageNumber="70">Gonatodes rozei</emphasis>
|
||
</taxonomicName>
|
||
differs from
|
||
<taxonomicName id="4C4EB5FCFF93E5231BD4FB7DFCE6FB36" box="[678,797,1196,1220]" class="Reptilia" family="Sphaerodactylidae" genus="Gonatodes" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="70" phylum="Chordata" rank="species" species="ceciliae">
|
||
<emphasis id="B93A126DFF93E5231BD4FB7DFD4CFB36" box="[678,695,1197,1220]" italics="true" pageId="4" pageNumber="70">G</emphasis>
|
||
.
|
||
<emphasis id="B93A126DFF93E5231BB4FB7CFCE6FB36" box="[710,797,1196,1220]" italics="true" pageId="4" pageNumber="70">ceciliae</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C4EB5FCFF93E5231A5EFB7DFC29FB36" box="[812,978,1197,1220]" class="Reptilia" family="Sphaerodactylidae" genus="Gonatodes" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="70" phylum="Chordata" rank="species" species="concinnatus">
|
||
<emphasis id="B93A126DFF93E5231A5EFB7DFCC6FB36" box="[812,829,1197,1220]" italics="true" pageId="4" pageNumber="70">G</emphasis>
|
||
.
|
||
<emphasis id="B93A126DFF93E5231A39FB7DFC29FB36" box="[843,978,1197,1220]" italics="true" pageId="4" pageNumber="70">concinnatus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C4EB5FCFF93E5231A93FB7DFB73FB36" box="[993,1160,1197,1220]" class="Reptilia" family="Sphaerodactylidae" genus="Gonatodes" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="70" phylum="Chordata" rank="species" species="nascimentoi">
|
||
<emphasis id="B93A126DFF93E5231A93FB7DFC09FB36" box="[993,1010,1197,1220]" italics="true" pageId="4" pageNumber="70">G</emphasis>
|
||
.
|
||
<emphasis id="B93A126DFF93E5231D73FB7DFB73FB36" box="[1025,1160,1197,1220]" italics="true" pageId="4" pageNumber="70">nascimentoi</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C4EB5FCFF93E5231DE5FB7DFAE0FB36" box="[1175,1307,1196,1220]" class="Reptilia" family="Sphaerodactylidae" genus="Gonatodes" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="70" phylum="Chordata" rank="species" species="ocellatus">
|
||
<emphasis id="B93A126DFF93E5231DE5FB7DFB53FB36" box="[1175,1192,1197,1220]" italics="true" pageId="4" pageNumber="70">G</emphasis>
|
||
.
|
||
<emphasis id="B93A126DFF93E5231DC4FB7CFAE0FB36" box="[1206,1307,1196,1220]" italics="true" pageId="4" pageNumber="70">ocellatus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C4EB5FCFF93E5231C58FB7DFA6EFB36" box="[1322,1429,1197,1220]" class="Reptilia" family="Sphaerodactylidae" genus="Gonatodes" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="70" phylum="Chordata" rank="species" species="riveroi">
|
||
<emphasis id="B93A126DFF93E5231C58FB7DFAC0FB36" box="[1322,1339,1197,1220]" italics="true" pageId="4" pageNumber="70">G</emphasis>
|
||
.
|
||
<emphasis id="B93A126DFF93E5231C38FB7DFA6EFB36" box="[1354,1429,1197,1220]" italics="true" pageId="4" pageNumber="70">riveroi</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="4C4EB5FCFF93E52319BAFB00FE97FB1A" box="[200,364,1232,1256]" class="Reptilia" family="Sphaerodactylidae" genus="Gonatodes" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="70" phylum="Chordata" rank="species" species="tapajonicus">
|
||
<emphasis id="B93A126DFF93E52319BAFB00FF22FB15" box="[200,217,1232,1255]" italics="true" pageId="4" pageNumber="70">G</emphasis>
|
||
.
|
||
<emphasis id="B93A126DFF93E523199BFB01FE97FB1A" box="[233,364,1233,1256]" italics="true" pageId="4" pageNumber="70">tapajonicus</emphasis>
|
||
</taxonomicName>
|
||
in lacking a suprahumeral/suprascapular pale spot/bar edged by black as opposed to having a distinct (especially in males), pale (white or yellow) spot/bar edged by black located around the scapular area, either anteriorly or posteriorly relative to the front limb insertion. It further differs from
|
||
<taxonomicName id="4C4EB5FCFF93E5231DB6FAC8FA8BFAC2" box="[1220,1392,1304,1328]" class="Reptilia" family="Sphaerodactylidae" genus="Gonatodes" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="70" phylum="Chordata" rank="species" species="concinnatus">
|
||
<emphasis id="B93A126DFF93E5231DB6FAC8FB2EFADD" box="[1220,1237,1304,1327]" italics="true" pageId="4" pageNumber="70">G</emphasis>
|
||
.
|
||
<emphasis id="B93A126DFF93E5231D9BFAC9FA8BFAC2" box="[1257,1392,1305,1328]" italics="true" pageId="4" pageNumber="70">concinnatus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C4EB5FCFF93E5231CF6FAC8FEE5FAA6" class="Reptilia" family="Sphaerodactylidae" genus="Gonatodes" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="70" phylum="Chordata" rank="species" species="nascimentoi">
|
||
<emphasis id="B93A126DFF93E5231CF6FAC8FA6EFADD" box="[1412,1429,1304,1327]" italics="true" pageId="4" pageNumber="70">G</emphasis>
|
||
.
|
||
<emphasis id="B93A126DFF93E52319E5FAEDFEE5FAA6" box="[151,286,1341,1364]" italics="true" pageId="4" pageNumber="70">nascimentoi</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="4C4EB5FCFF93E5231828FAEDFE07FAA6" box="[346,508,1341,1364]" class="Reptilia" family="Sphaerodactylidae" genus="Gonatodes" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="70" phylum="Chordata" rank="species" species="tapajonicus">
|
||
<emphasis id="B93A126DFF93E5231828FAEDFE90FAA6" box="[346,363,1341,1364]" italics="true" pageId="4" pageNumber="70">G</emphasis>
|
||
.
|
||
<emphasis id="B93A126DFF93E523180BFAEDFE07FAA6" box="[377,508,1341,1364]" italics="true" pageId="4" pageNumber="70">tapajonicus</emphasis>
|
||
</taxonomicName>
|
||
in having fewer scales around midbody (<
|
||
<quantity id="4CB6639AFF93E5231AA5FAECFBE5FAA6" box="[983,1054,1340,1364]" metricMagnitude="0" metricUnit="m" metricValue="2.5654" pageId="4" pageNumber="70" unit="in" value="101.0">101 in</quantity>
|
||
<taxonomicName id="4C4EB5FCFF93E5231D56FAEDFB83FAA6" box="[1060,1144,1341,1364]" class="Reptilia" family="Sphaerodactylidae" genus="Gonatodes" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="70" phylum="Chordata" rank="species" species="rozei">
|
||
<emphasis id="B93A126DFF93E5231D56FAEDFBCEFAA6" box="[1060,1077,1341,1364]" italics="true" pageId="4" pageNumber="70">G</emphasis>
|
||
.
|
||
<emphasis id="B93A126DFF93E5231D30FAEDFB83FAA6" box="[1090,1144,1341,1364]" italics="true" pageId="4" pageNumber="70">rozei</emphasis>
|
||
</taxonomicName>
|
||
as opposed to>
|
||
<quantity id="4CB6639AFF93E5231C5FFAECFA88FAA6" box="[1325,1395,1340,1364]" metricMagnitude="0" metricUnit="m" metricValue="2.7432" pageId="4" pageNumber="70" unit="in" value="108.0">108 in</quantity>
|
||
the aforementioned species). Males of these three species, together with
|
||
<taxonomicName id="4C4EB5FCFF93E5231AC6FAB0FBDBFA8A" box="[948,1056,1376,1400]" class="Reptilia" family="Sphaerodactylidae" genus="Gonatodes" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="70" phylum="Chordata" rank="species" species="riveroi">
|
||
<emphasis id="B93A126DFF93E5231AC6FAB0FC3EFA85" box="[948,965,1376,1399]" italics="true" pageId="4" pageNumber="70">G</emphasis>
|
||
.
|
||
<emphasis id="B93A126DFF93E5231AA7FAB1FBDBFA8A" box="[981,1056,1377,1400]" italics="true" pageId="4" pageNumber="70">riveroi</emphasis>
|
||
</taxonomicName>
|
||
, are reddish brown dorsally with small, yellow and white vermiculations densely covering the body (see color variation for
|
||
<taxonomicName id="4C4EB5FCFF93E5231DAFFA55FAC2FA6E" box="[1245,1337,1413,1436]" class="Reptilia" family="Sphaerodactylidae" genus="Gonatodes" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="70" phylum="Chordata" rank="species" species="rozei">
|
||
<emphasis id="B93A126DFF93E5231DAFFA55FB15FA6E" box="[1245,1262,1413,1436]" italics="true" pageId="4" pageNumber="70">G</emphasis>
|
||
.
|
||
<emphasis id="B93A126DFF93E5231C71FA55FAC2FA6E" box="[1283,1337,1413,1436]" italics="true" pageId="4" pageNumber="70">rozei</emphasis>
|
||
</taxonomicName>
|
||
below).
|
||
<taxonomicName id="4C4EB5FCFF93E52319E5FA78FEABFA32" box="[151,336,1448,1472]" class="Reptilia" family="Sphaerodactylidae" genus="Gonatodes" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="70" phylum="Chordata" rank="species" species="rozei">
|
||
<emphasis id="B93A126DFF93E52319E5FA78FEABFA32" box="[151,336,1448,1472]" italics="true" pageId="4" pageNumber="70">Gonatodes rozei</emphasis>
|
||
</taxonomicName>
|
||
lacks red, olive green and yellow reticulations on the body as observed in males of
|
||
<taxonomicName id="4C4EB5FCFF93E5231C6FFA78FA6EFA32" box="[1309,1429,1448,1472]" class="Reptilia" family="Sphaerodactylidae" genus="Gonatodes" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="70" phylum="Chordata" rank="species" species="ceciliae">
|
||
<emphasis id="B93A126DFF93E5231C6FFA78FAD5FA4D" box="[1309,1326,1448,1471]" italics="true" pageId="4" pageNumber="70">G</emphasis>
|
||
.
|
||
<emphasis id="B93A126DFF93E5231C4CFA78FA6EFA32" box="[1342,1429,1448,1472]" italics="true" pageId="4" pageNumber="70">ceciliae</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<taxonomicName id="4C4EB5FCFF93E52319E5FA1CFE84FA16" box="[151,383,1484,1508]" class="Reptilia" family="Sphaerodactylidae" genus="Gonatodes" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="70" phylum="Chordata" rank="species" species="ocellatus">
|
||
<emphasis id="B93A126DFF93E52319E5FA1CFE84FA16" box="[151,383,1484,1508]" italics="true" pageId="4" pageNumber="70">Gonatodes ocellatus</emphasis>
|
||
</taxonomicName>
|
||
is a smaller species that does not exceeds
|
||
<quantity id="4CB6639AFF93E5231A02FA1CFC3AFA16" box="[880,961,1484,1508]" metricMagnitude="-2" metricUnit="m" metricValue="4.9" pageId="4" pageNumber="70" unit="mm" value="49.0">49 mm</quantity>
|
||
in SVL.
|
||
<taxonomicName id="4C4EB5FCFF93E5231D5DFA1CFB13FA16" box="[1071,1256,1484,1508]" class="Reptilia" family="Sphaerodactylidae" genus="Gonatodes" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="70" phylum="Chordata" rank="species" species="rozei">
|
||
<emphasis id="B93A126DFF93E5231D5DFA1CFB13FA16" box="[1071,1256,1484,1508]" italics="true" pageId="4" pageNumber="70">Gonatodes rozei</emphasis>
|
||
</taxonomicName>
|
||
differs from
|
||
<taxonomicName id="4C4EB5FCFF93E5231CF6FA1DFEF5F9FA" class="Reptilia" family="Sphaerodactylidae" genus="Gonatodes" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="70" phylum="Chordata" rank="species" species="lichenosus">
|
||
<emphasis id="B93A126DFF93E5231CF6FA1DFA6EFA16" box="[1412,1429,1485,1508]" italics="true" pageId="4" pageNumber="70">G</emphasis>
|
||
.
|
||
<emphasis id="B93A126DFF93E52319E5FA20FEF5F9FA" box="[151,270,1520,1544]" italics="true" pageId="4" pageNumber="70">lichenosus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C4EB5FCFF93E5231831FA20FE50F9FA" box="[323,427,1520,1544]" class="Reptilia" family="Sphaerodactylidae" genus="Gonatodes" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="70" phylum="Chordata" rank="species" species="seiglei">
|
||
<emphasis id="B93A126DFF93E5231831FA20FEAFF9F5" box="[323,340,1520,1543]" italics="true" pageId="4" pageNumber="70">G</emphasis>
|
||
.
|
||
<emphasis id="B93A126DFF93E5231810FA20FE50F9FA" box="[354,427,1520,1544]" italics="true" pageId="4" pageNumber="70">seiglei</emphasis>
|
||
</taxonomicName>
|
||
in having a subcaudal pattern
|
||
<typeStatus id="54F570DDFF93E5231A73FA20FCCAF9FA" box="[769,817,1520,1544]" pageId="4" pageNumber="70">type</typeStatus>
|
||
B as opposed to
|
||
<typeStatus id="54F570DDFF93E5231A83FA20FBDAF9FA" box="[1009,1057,1520,1544]" pageId="4" pageNumber="70">type</typeStatus>
|
||
G and D, respectively.
|
||
</paragraph>
|
||
<paragraph id="8BF1CE7FFF93E52219B5F9C4FC4AFDE5" blockId="4.[151,1437,151,2013]" lastBlockId="5.[151,1436,151,535]" lastPageId="5" lastPageNumber="71" pageId="4" pageNumber="70">
|
||
<taxonomicName id="4C4EB5FCFF93E52319B5F9C4FE85F9DE" box="[199,382,1556,1580]" class="Reptilia" family="Sphaerodactylidae" genus="Gonatodes" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="70" phylum="Chordata" rank="species" species="rozei">
|
||
<emphasis id="B93A126DFF93E52319B5F9C4FE85F9DE" box="[199,382,1556,1580]" italics="true" pageId="4" pageNumber="70">Gonatodes rozei</emphasis>
|
||
</taxonomicName>
|
||
is found in close geographic proximity (Schargel, unpublished data) with
|
||
<taxonomicName id="4C4EB5FCFF93E5231DBFF9C5FA90F9DE" box="[1229,1387,1556,1580]" class="Reptilia" family="Sphaerodactylidae" genus="Gonatodes" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="70" phylum="Chordata" rank="species" species="falconensis">
|
||
<emphasis id="B93A126DFF93E5231DBFF9C5FB25F9DE" box="[1229,1246,1557,1580]" italics="true" pageId="4" pageNumber="70">G</emphasis>
|
||
.
|
||
<emphasis id="B93A126DFF93E5231D9FF9C4FA90F9DE" box="[1261,1387,1556,1580]" italics="true" pageId="4" pageNumber="70">falconensis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C4EB5FCFF93E52319E5F9E8FEFBF9A2" box="[151,256,1592,1616]" class="Reptilia" family="Sphaerodactylidae" genus="Gonatodes" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="70" phylum="Chordata" rank="species" species="taniae">
|
||
<emphasis id="B93A126DFF93E52319E5F9E8FF53F9BD" box="[151,168,1592,1615]" italics="true" pageId="4" pageNumber="70">G</emphasis>
|
||
.
|
||
<emphasis id="B93A126DFF93E52319C8F9E9FEFBF9A2" box="[186,256,1593,1616]" italics="true" pageId="4" pageNumber="70">taniae</emphasis>
|
||
</taxonomicName>
|
||
, so the comparisons with these two species need to be more exhaustive.
|
||
<taxonomicName id="4C4EB5FCFF93E5231D1FF9E8FA67F9A2" box="[1133,1436,1592,1616]" class="Reptilia" family="Sphaerodactylidae" genus="Gonatodes" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="70" phylum="Chordata" rank="species" species="purpurogularis">
|
||
<emphasis id="B93A126DFF93E5231D1FF9E8FA67F9A2" box="[1133,1436,1592,1616]" italics="true" pageId="4" pageNumber="70">Gonatodes purpurogularis</emphasis>
|
||
</taxonomicName>
|
||
(endemic to the Mérida mountain range) is not found in close geographic proximity to
|
||
<taxonomicName id="4C4EB5FCFF93E5231DFBF98DFB1AF986" box="[1161,1249,1629,1652]" class="Reptilia" family="Sphaerodactylidae" genus="Gonatodes" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="70" phylum="Chordata" rank="species" species="rozei">
|
||
<emphasis id="B93A126DFF93E5231DFBF98DFB61F986" box="[1161,1178,1629,1652]" italics="true" pageId="4" pageNumber="70">G</emphasis>
|
||
.
|
||
<emphasis id="B93A126DFF93E5231DD9F98DFB1AF986" box="[1195,1249,1629,1652]" italics="true" pageId="4" pageNumber="70">rozei</emphasis>
|
||
</taxonomicName>
|
||
but this species forms a well-supported clade with
|
||
<taxonomicName id="4C4EB5FCFF93E5231B6DF950FD47F96A" box="[543,700,1664,1688]" class="Reptilia" family="Sphaerodactylidae" genus="Gonatodes" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="70" phylum="Chordata" rank="species" species="falconensis">
|
||
<emphasis id="B93A126DFF93E5231B6DF950FDCBF965" box="[543,560,1664,1687]" italics="true" pageId="4" pageNumber="70">G</emphasis>
|
||
.
|
||
<emphasis id="B93A126DFF93E5231B4CF950FD47F96A" box="[574,700,1664,1688]" italics="true" pageId="4" pageNumber="70">falconensis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C4EB5FCFF93E5231B86F950FCA2F96A" box="[756,857,1664,1688]" class="Reptilia" family="Sphaerodactylidae" genus="Gonatodes" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="70" phylum="Chordata" rank="species" species="taniae">
|
||
<emphasis id="B93A126DFF93E5231B86F950FCFEF965" box="[756,773,1664,1687]" italics="true" pageId="4" pageNumber="70">G</emphasis>
|
||
.
|
||
<emphasis id="B93A126DFF93E5231A61F951FCA2F96A" box="[787,857,1665,1688]" italics="true" pageId="4" pageNumber="70">taniae</emphasis>
|
||
</taxonomicName>
|
||
(see
|
||
<bibRefCitation id="EFDFB38EFF93E5231AE7F9AFFB81F965" author="Schargel" box="[917,1146,1663,1688]" pageId="4" pageNumber="70" refString="Schargel, W. E., Rivas, G. A., Makowsky, R., Senaris, J. C, Natera, M. A, Barros, T. R., Molina, C. R., & Barrio-Amoros, C. L. (2010) Phylogenetic systematics of the genus Gonatodes (Squamata: Sphaerodactylidae) in the Guayana region, with description of a new species from Venezuela. Systematics and Biodiversity, 8, 321 - 339." type="journal article" year="2010">
|
||
Schargel
|
||
<emphasis id="B93A126DFF93E5231A8CF951FBCBF96A" box="[1022,1072,1664,1688]" italics="true" pageId="4" pageNumber="70">et al</emphasis>
|
||
. 2010
|
||
</bibRefCitation>
|
||
), and all three species are very similar morphologically. These three species will be referred to as the
|
||
<taxonomicName id="4C4EB5FCFF93E5231AAFF974FBA0F94E" box="[989,1115,1700,1724]" class="Reptilia" family="Sphaerodactylidae" genus="Gonatodes" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="70" phylum="Chordata" rank="species" species="falconensis">
|
||
<emphasis id="B93A126DFF93E5231AAFF974FBA0F94E" box="[989,1115,1700,1724]" italics="true" pageId="4" pageNumber="70">falconensis</emphasis>
|
||
</taxonomicName>
|
||
complex in the comparisons in instances for which they all share the same character state. Males of
|
||
<taxonomicName id="4C4EB5FCFF93E5231AAEF918FB80F912" box="[988,1147,1736,1760]" class="Reptilia" family="Sphaerodactylidae" genus="Gonatodes" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="70" phylum="Chordata" rank="species" species="falconensis">
|
||
<emphasis id="B93A126DFF93E5231AAEF918FC16F92D" box="[988,1005,1736,1759]" italics="true" pageId="4" pageNumber="70">G</emphasis>
|
||
.
|
||
<emphasis id="B93A126DFF93E5231A8FF918FB80F912" box="[1021,1147,1736,1760]" italics="true" pageId="4" pageNumber="70">falconensis</emphasis>
|
||
</taxonomicName>
|
||
have dull red or reddish brown markings, typically forming short stripes, on the head, sometimes on the body. Males of
|
||
<taxonomicName id="4C4EB5FCFF93E5231D86F93DFAA6F8F6" box="[1268,1373,1773,1796]" class="Reptilia" family="Sphaerodactylidae" genus="Gonatodes" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="70" phylum="Chordata" rank="species" species="taniae">
|
||
<emphasis id="B93A126DFF93E5231D86F93DFAFEF8F6" box="[1268,1285,1773,1796]" italics="true" pageId="4" pageNumber="70">G</emphasis>
|
||
.
|
||
<emphasis id="B93A126DFF93E5231C65F93DFAA6F8F6" box="[1303,1373,1773,1796]" italics="true" pageId="4" pageNumber="70">taniae</emphasis>
|
||
</taxonomicName>
|
||
have distinct white stripes (which remain conspicuous in museum specimens) on the head, including most noticeably canthal, postocular and postrictal stripes. In
|
||
<taxonomicName id="4C4EB5FCFF93E5231BFAF8E5FCAAF8BE" box="[648,849,1844,1868]" class="Reptilia" family="Sphaerodactylidae" genus="Gonatodes" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="70" phylum="Chordata" rank="species" species="purpurogularis">
|
||
<emphasis id="B93A126DFF93E5231BFAF8E5FD62F8BE" box="[648,665,1845,1868]" italics="true" pageId="4" pageNumber="70">G</emphasis>
|
||
.
|
||
<emphasis id="B93A126DFF93E5231BD5F8E4FCAAF8BE" box="[679,849,1844,1868]" italics="true" pageId="4" pageNumber="70">purpurogularis</emphasis>
|
||
</taxonomicName>
|
||
the white markings on the head are also present but they are short, not forming stripes. In contrast to what is observed in the
|
||
<taxonomicName id="4C4EB5FCFF93E5231A90F888FB9BF882" box="[994,1120,1880,1904]" class="Reptilia" family="Sphaerodactylidae" genus="Gonatodes" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="70" phylum="Chordata" rank="species" species="falconensis">
|
||
<emphasis id="B93A126DFF93E5231A90F888FB9BF882" box="[994,1120,1880,1904]" italics="true" pageId="4" pageNumber="70">falconensis</emphasis>
|
||
</taxonomicName>
|
||
complex, males of
|
||
<taxonomicName id="4C4EB5FCFF93E5231C36F888FA60F882" box="[1348,1435,1880,1904]" class="Reptilia" family="Sphaerodactylidae" genus="Gonatodes" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="70" phylum="Chordata" rank="species" species="rozei">
|
||
<emphasis id="B93A126DFF93E5231C36F888FAAEF89D" box="[1348,1365,1880,1903]" italics="true" pageId="4" pageNumber="70">G</emphasis>
|
||
.
|
||
<emphasis id="B93A126DFF93E5231C17F889FA60F882" box="[1381,1435,1881,1904]" italics="true" pageId="4" pageNumber="70">rozei</emphasis>
|
||
</taxonomicName>
|
||
have most commonly a solid yellow/orange (which fades to cream) head or, less frequently, irregular yellow reticulations on the head. Males of the
|
||
<taxonomicName id="4C4EB5FCFF93E5231B15F870FD1EF84A" box="[615,741,1952,1976]" class="Reptilia" family="Sphaerodactylidae" genus="Gonatodes" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="70" phylum="Chordata" rank="species" species="falconensis">
|
||
<emphasis id="B93A126DFF93E5231B15F870FD1EF84A" box="[615,741,1952,1976]" italics="true" pageId="4" pageNumber="70">falconensis</emphasis>
|
||
</taxonomicName>
|
||
complex have bright yellow coloration (fades in preserved specimens; variable in
|
||
<taxonomicName id="4C4EB5FCFF93E52318D5F815FDB2F82E" box="[423,585,1988,2012]" class="Reptilia" family="Sphaerodactylidae" genus="Gonatodes" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="70" phylum="Chordata" rank="species" species="falconensis">
|
||
<emphasis id="B93A126DFF93E52318D5F815FE43F82E" box="[423,440,1989,2012]" italics="true" pageId="4" pageNumber="70">G</emphasis>
|
||
.
|
||
<emphasis id="B93A126DFF93E52318B9F814FDB2F82E" box="[459,585,1988,2012]" italics="true" pageId="4" pageNumber="70">falconensis</emphasis>
|
||
</taxonomicName>
|
||
) suffused on the sides of the body over a pale gray background. Body coloration is variable (see variation section) in
|
||
<taxonomicName id="4C4EB5FCFF92E5221BC6FF48FCF1FF42" box="[692,778,152,176]" class="Reptilia" family="Sphaerodactylidae" genus="Gonatodes" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="71" phylum="Chordata" rank="species" species="rozei">
|
||
<emphasis id="B93A126DFF92E5221BC6FF48FD3EFF5D" box="[692,709,152,175]" italics="true" pageId="5" pageNumber="71">G</emphasis>
|
||
.
|
||
<emphasis id="B93A126DFF92E5221BA6FF49FCF1FF42" box="[724,778,153,176]" italics="true" pageId="5" pageNumber="71">rozei</emphasis>
|
||
</taxonomicName>
|
||
, but we have not observed specimens with bright yellow coloration on the body. Females of the
|
||
<taxonomicName id="4C4EB5FCFF92E5221B20FF6CFD2BFF26" box="[594,720,188,212]" class="Reptilia" family="Sphaerodactylidae" genus="Gonatodes" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="71" phylum="Chordata" rank="species" species="falconensis">
|
||
<emphasis id="B93A126DFF92E5221B20FF6CFD2BFF26" box="[594,720,188,212]" italics="true" pageId="5" pageNumber="71">falconensis</emphasis>
|
||
</taxonomicName>
|
||
complex have a wide (expanding to the dorsolateral area), pale, middorsal, longitudinal band with straight edges. This band contains either an irregular, dark, vertebral stripe or a series of paired, paravertebral, dark spots. In females of
|
||
<taxonomicName id="4C4EB5FCFF92E5221A56FED5FC80FEEE" box="[804,891,261,284]" class="Reptilia" family="Sphaerodactylidae" genus="Gonatodes" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="71" phylum="Chordata" rank="species" species="rozei">
|
||
<emphasis id="B93A126DFF92E5221A56FED5FCCEFEEE" box="[804,821,261,284]" italics="true" pageId="5" pageNumber="71">G</emphasis>
|
||
.
|
||
<emphasis id="B93A126DFF92E5221A37FED5FC80FEEE" box="[837,891,261,284]" italics="true" pageId="5" pageNumber="71">rozei</emphasis>
|
||
</taxonomicName>
|
||
there is a pale, vertebral stripe which does not expand to the dorsolateral area and, as opposed to the middorsal band of the aforementioned species, the lateral edges are strongly undulating. Females of the
|
||
<taxonomicName id="4C4EB5FCFF92E5221BECFE9CFCE7FE96" box="[670,796,332,356]" class="Reptilia" family="Sphaerodactylidae" genus="Gonatodes" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="71" phylum="Chordata" rank="species" species="falconensis">
|
||
<emphasis id="B93A126DFF92E5221BECFE9CFCE7FE96" box="[670,796,332,356]" italics="true" pageId="5" pageNumber="71">falconensis</emphasis>
|
||
</taxonomicName>
|
||
complex have pale yellow or white transverse dashes on the sides of the body, whereas females of
|
||
<taxonomicName id="4C4EB5FCFF92E5221B0BFEA0FD2BFE7A" box="[633,720,368,392]" class="Reptilia" family="Sphaerodactylidae" genus="Gonatodes" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="71" phylum="Chordata" rank="species" species="rozei">
|
||
<emphasis id="B93A126DFF92E5221B0BFEA0FD71FE75" box="[633,650,368,391]" italics="true" pageId="5" pageNumber="71">G</emphasis>
|
||
.
|
||
<emphasis id="B93A126DFF92E5221BE8FEA1FD2BFE7A" box="[666,720,369,392]" italics="true" pageId="5" pageNumber="71">rozei</emphasis>
|
||
</taxonomicName>
|
||
have white dots on the side of the body. Regarding aspects of morphology other than color pattern, the three species of the
|
||
<taxonomicName id="4C4EB5FCFF92E5221A27FE44FC28FE5E" box="[853,979,404,428]" class="Reptilia" family="Sphaerodactylidae" genus="Gonatodes" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="71" phylum="Chordata" rank="species" species="falconensis">
|
||
<emphasis id="B93A126DFF92E5221A27FE44FC28FE5E" box="[853,979,404,428]" italics="true" pageId="5" pageNumber="71">falconensis</emphasis>
|
||
</taxonomicName>
|
||
complex have a scalation pattern in the gular area in which the scales posterior to the postmentals gradually transition from larger polygonal and juxtaposed scales into smaller imbricate scales (Schargel unpublished); whereas in
|
||
<taxonomicName id="4C4EB5FCFF92E5221D34FE0DFB67FE06" box="[1094,1180,477,500]" class="Reptilia" family="Sphaerodactylidae" genus="Gonatodes" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="71" phylum="Chordata" rank="species" species="rozei">
|
||
<emphasis id="B93A126DFF92E5221D34FE0DFBACFE06" box="[1094,1111,477,500]" italics="true" pageId="5" pageNumber="71">G</emphasis>
|
||
.
|
||
<emphasis id="B93A126DFF92E5221D14FE0DFB67FE06" box="[1126,1180,477,500]" italics="true" pageId="5" pageNumber="71">rozei</emphasis>
|
||
</taxonomicName>
|
||
the scales posterior to the postmentals transition sharply into smaller, granular scales (
|
||
<figureCitation id="1375D2FAFF92E5221A2DFE2FFC58FDE5" box="[863,931,511,535]" captionStart="FIGURE 3" captionStartId="5.[151,250,1114,1137]" captionTargetBox="[175,1422,589,1102]" captionTargetId="figure@5.[175,1422,589,1102]" captionTargetPageId="5" captionText="FIGURE 3. Scalation pattern in the gular area of Gonatodes rozei sp. nov. (left; illustration based on MHNLS 5532 from Valle de Guanape, Anzoátegui, Venezuela) and the falconensis complex (right; illustration based on specimen of G. taniae, TCWC 59378, from Rancho Grande, Aragua, Venezuela)." httpUri="https://zenodo.org/record/282623/files/figure.png" pageId="5" pageNumber="71">Fig. 3</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="DF319EF7FF92E52219E5FB8AFD6FFB5C" httpUri="https://zenodo.org/record/282623/files/figure.png" pageId="5" pageNumber="71" targetBox="[175,1422,589,1102]" targetPageId="5">
|
||
<paragraph id="8BF1CE7FFF92E52219E5FB8AFD6FFB5C" blockId="5.[151,1436,1114,1198]" pageId="5" pageNumber="71">
|
||
<emphasis id="B93A126DFF92E52219E5FB8AFEE9FB82" bold="true" box="[151,274,1114,1137]" pageId="5" pageNumber="71">FIGURE 3.</emphasis>
|
||
Scalation pattern in the gular area of
|
||
<taxonomicName id="4C4EB5FCFF92E5221BFFFB8AFCC9FB82" box="[653,818,1114,1136]" class="Reptilia" family="Sphaerodactylidae" genus="Gonatodes" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="71" phylum="Chordata" rank="species" species="rozei" status="sp. nov.">
|
||
<emphasis id="B93A126DFF92E5221BFFFB8AFCC9FB82" box="[653,818,1114,1136]" italics="true" pageId="5" pageNumber="71">Gonatodes rozei</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B93A126DFF92E5221A4BFB8BFC72FB82" bold="true" box="[825,905,1115,1136]" pageId="5" pageNumber="71">
|
||
<taxonomicNameLabel id="A209AF16FF92E5221A4BFB8BFC72FB82" box="[825,905,1115,1136]" pageId="5" pageNumber="71" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(left; illustration based on MHNLS 5532 from Valle de Guanape, Anzoátegui, Venezuela) and the
|
||
<taxonomicName id="4C4EB5FCFF92E5221B1EFBA9FD25FB7D" box="[620,734,1145,1167]" class="Reptilia" family="Sphaerodactylidae" genus="Gonatodes" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="71" phylum="Chordata" rank="species" species="falconensis">
|
||
<emphasis id="B93A126DFF92E5221B1EFBA9FD25FB7D" box="[620,734,1145,1167]" italics="true" pageId="5" pageNumber="71">falconensis</emphasis>
|
||
</taxonomicName>
|
||
complex (right; illustration based on specimen of
|
||
<taxonomicName id="4C4EB5FCFF92E5221D9AFBAAFABFFB7D" box="[1256,1348,1146,1167]" class="Reptilia" family="Sphaerodactylidae" genus="Gonatodes" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="71" phylum="Chordata" rank="species" species="taniae">
|
||
<emphasis id="B93A126DFF92E5221D9AFBAAFB0CFB7D" box="[1256,1271,1146,1167]" italics="true" pageId="5" pageNumber="71">G</emphasis>
|
||
.
|
||
<emphasis id="B93A126DFF92E5221C77FBAAFABFFB7D" box="[1285,1348,1146,1167]" italics="true" pageId="5" pageNumber="71">taniae</emphasis>
|
||
</taxonomicName>
|
||
, TCWC 59378, from Rancho Grande, Aragua, Venezuela).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BF1CE7FFF92E52219B5FB0AFEC8FAC8" blockId="5.[151,1437,1242,2024]" pageId="5" pageNumber="71">
|
||
Finally,
|
||
<emphasis id="B93A126DFF92E5221851FB0BFE8DFB01" box="[291,374,1243,1267]" italics="true" pageId="5" pageNumber="71">G. ro z e i</emphasis>
|
||
and
|
||
<taxonomicName id="4C4EB5FCFF92E52218C0FB0BFDC7FB01" box="[434,572,1243,1267]" class="Reptilia" family="Sphaerodactylidae" genus="Gonatodes" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="71" phylum="Chordata" rank="species" species="annularis">
|
||
<emphasis id="B93A126DFF92E52218C0FB0BFE38FB00" box="[434,451,1243,1266]" italics="true" pageId="5" pageNumber="71">G</emphasis>
|
||
.
|
||
<emphasis id="B93A126DFF92E52218A3FB0BFDC7FB01" box="[465,572,1243,1267]" italics="true" pageId="5" pageNumber="71">annularis</emphasis>
|
||
</taxonomicName>
|
||
are the only two species of
|
||
<taxonomicName id="4C4EB5FCFF92E5221A09FB0BFC0FFB01" box="[891,1012,1243,1267]" class="Reptilia" family="Sphaerodactylidae" genus="Gonatodes" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="71" phylum="Chordata" rank="genus">
|
||
<emphasis id="B93A126DFF92E5221A09FB0BFC0FFB01" box="[891,1012,1243,1267]" italics="true" pageId="5" pageNumber="71">Gonatodes</emphasis>
|
||
</taxonomicName>
|
||
in which the iris of the eye is blue in life (always the case in males but variable in females). In all other species in the genus the iris is yellow, bronze, copper or red.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3549DF4FF92E52E19B5FA96FD8DF879" lastPageId="9" lastPageNumber="75" pageId="5" pageNumber="71" type="description">
|
||
<paragraph id="8BF1CE7FFF92E52219B5FA96FE85F9A8" blockId="5.[151,1437,1242,2024]" pageId="5" pageNumber="71">
|
||
<emphasis id="B93A126DFF92E52219B5FA96FE12FAAD" bold="true" box="[199,489,1350,1375]" pageId="5" pageNumber="71">
|
||
Description of
|
||
<typeStatus id="54F570DDFF92E522180EFA97FE1EFAAD" box="[380,485,1351,1375]" pageId="5" pageNumber="71" type="holotype">holotype</typeStatus>
|
||
.
|
||
</emphasis>
|
||
An adult male (
|
||
<figureCitation id="1375D2FAFF92E5221BDBFA97FD14FAAD" box="[681,751,1351,1375]" captionStart="FIGURE 2" captionStartId="3.[151,250,1844,1867]" captionTargetBox="[251,1340,359,1805]" captionTargetId="figure@3.[234,1353,343,1823]" captionTargetPageId="3" captionText="FIGURE 2. Adult male (top, holotype, MHNLS 20780) and adult female (bottom, UTA 60126) of Gonatodes rozei sp. nov. in life." httpUri="https://zenodo.org/record/282622/files/figure.png" pageId="5" pageNumber="71">Fig. 2</figureCitation>
|
||
), with snout-vent length of
|
||
<quantity id="4CB6639AFF92E5221D41FA97FB66FAAD" box="[1075,1181,1351,1375]" metricMagnitude="-2" metricUnit="m" metricValue="5.91" pageId="5" pageNumber="71" unit="mm" value="59.1">59.1 mm</quantity>
|
||
. Tail length
|
||
<quantity id="4CB6639AFF92E5221C5CFA97FA63FAAD" box="[1326,1432,1351,1375]" metricMagnitude="-2" metricUnit="m" metricValue="5.7299999999999995" pageId="5" pageNumber="71" unit="mm" value="57.3">57.3 mm</quantity>
|
||
, complete but regenerated. Head about 1.4 times longer than wide (head length:
|
||
<quantity id="4CB6639AFF92E5221D56FABAFB77FA70" box="[1060,1164,1386,1410]" metricMagnitude="-2" metricUnit="m" metricValue="1.3599999999999999" pageId="5" pageNumber="71" unit="mm" value="13.6">13.6 mm</quantity>
|
||
; head width: 10.0 mm). Snout short (eye-nostril distance is
|
||
<quantity id="4CB6639AFF92E5221B46FA5FFD69FA55" box="[564,658,1423,1447]" metricMagnitude="-3" metricUnit="m" metricValue="4.2" pageId="5" pageNumber="71" unit="mm" value="4.2">4.2 mm</quantity>
|
||
), 0.3 times head length, somewhat acutely rounded in dorsal view, sloping toward top of head with an approximate 45º angle in lateral view. Lower jaw slightly projecting past the tip of the snout, visible in dorsal view. Neck
|
||
<quantity id="4CB6639AFF92E5221BF0FA07FD21FA1D" box="[642,730,1495,1519]" metricMagnitude="-3" metricUnit="m" metricValue="7.3" pageId="5" pageNumber="71" unit="mm" value="7.3">7.3 mm</quantity>
|
||
wide (shortest distance), narrower than head and body. Body subcylindrical, wider than high; axilla-groin distance
|
||
<quantity id="4CB6639AFF92E5221B9DFA2AFCADF9E0" box="[751,854,1530,1554]" metricMagnitude="-2" metricUnit="m" metricValue="2.76" pageId="5" pageNumber="71" unit="mm" value="27.6">27.6 mm</quantity>
|
||
. Limbs well developed with moderately long digits, fourth toe length
|
||
<quantity id="4CB6639AFF92E5221825F9CFFE54F9C5" box="[343,431,1567,1591]" metricMagnitude="-3" metricUnit="m" metricValue="7.9" pageId="5" pageNumber="71" unit="mm" value="7.9">7.9 mm</quantity>
|
||
, 0.95 times shank length (
|
||
<quantity id="4CB6639AFF92E5221BA3F9CFFCD0F9C5" box="[721,811,1567,1591]" metricMagnitude="-3" metricUnit="m" metricValue="8.3" pageId="5" pageNumber="71" unit="mm" value="8.3">8.3 mm</quantity>
|
||
, from base of palm to knee). Tail round in cross section, tapering towards tip.
|
||
</paragraph>
|
||
<paragraph id="8BF1CE7FFF92E52219B5F9B8FDD2F935" blockId="5.[151,1437,1242,2024]" pageId="5" pageNumber="71">Tongue relatively wide with bluntly rounded tip, covered by small, scale-like papillae, which become less defined posteriorly; tip of tongue with very short median cleft; anterior portion of tongue gray, fading posteriorly. Teeth very small, subequal, conical.</paragraph>
|
||
<paragraph id="8BF1CE7FFF92E52119B5F902FE4DFE3D" blockId="5.[151,1437,1242,2024]" lastBlockId="6.[151,1437,151,1982]" lastPageId="6" lastPageNumber="72" pageId="5" pageNumber="71">
|
||
Rostral moderately large, visible from above, irregularly shaped, much wider than high, but with a small longitudinal groove that divides the scale almost completely except for a short section towards the lip. Three postrostrals, lateral ones (supranasals) distinctly larger than the single median scale, which is about the same size as the scales on the snout; posterior to the medial postrostral a slightly larger, single scale, separates the lateral postrostrals from contacting each other. Nostril bordered by rostral, four (left) or three (right) postnasals, and lateral postrostral (=supranasal). Three/four postnasals, on average slightly larger than scales on the loreal region. Scales on top of snout and loreal region basally round, granular to subconical, juxtaposed. Loreal scales number about 13/
|
||
<quantity id="4CB6639AFF92E52219E5F81FFF2EF815" box="[151,213,1999,2023]" metricMagnitude="-1" metricUnit="m" metricValue="3.81" pageId="5" pageNumber="71" unit="in" value="15.0">15 in</quantity>
|
||
a line between postnasals and anterior margin of eye socket. Scales decrease slightly in size from the postrostrals toward posterior part of head. Scales on supraorbital slightly smaller and more granular than those on top of head. A moderate supraciliary flap bears three well-developed spines located roughly above the center of the eye. The supraciliary scales anterior to these spines are flat and larger than those on the supraorbital region; posterior to the well-developed spines the supraciliary scales are tiny and granular although a few of these scales project laterally in the form of short and poorly developed spines. Pupil round. Supralabials 6/6 to level of center of eye of which the first is the largest, first few bearing minute tubercles; sixth supralabial followed by about 10/12 tiny, granular scales along the lip to rictus of mouth; these tiny scales about same size as those on temporal region. Scales on temporal region similar to those on posterior top of head. Ear opening (
|
||
<quantity id="4CB6639AFF91E5211D46FE44FB77FE5E" box="[1076,1164,404,428]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" pageId="6" pageNumber="72" unit="mm" value="1.3">1.3 mm</quantity>
|
||
) much smaller than eye (
|
||
<quantity id="4CB6639AFF91E52119EDFE67FF01FE3D" box="[159,250,439,463]" metricMagnitude="-3" metricUnit="m" metricValue="2.7" pageId="6" pageNumber="72" unit="mm" value="2.7">2.7 mm</quantity>
|
||
), obliquely oval.
|
||
</paragraph>
|
||
<paragraph id="8BF1CE7FFF91E52119B5FE0CFC5EFDAD" blockId="6.[151,1437,151,1982]" pageId="6" pageNumber="72">Mental large, with bluntly rounded anterior margin and three roughly straight sides on posterior margin. Postmentals three, distinctly larger than adjacent posterior scales. Scales on chin small, granular, a few larger, roughly elliptical scales adjacent to infralabials, juxtaposed. Infralabials 4/4 to level of center of eye, decreasing in size posteriorly, first two very large and projecting onto ventral plane.</paragraph>
|
||
<paragraph id="8BF1CE7FFF91E52119B5FDBCFBDFFD55" blockId="6.[151,1437,151,1982]" pageId="6" pageNumber="72">Scales on nape and on sides of neck granular to slightly conical. Scales on throat smooth, imbricate, with round posterior margin, with a short transitional area with granular scales on gular area.</paragraph>
|
||
<paragraph id="8BF1CE7FFF91E52119B5FD64FD42FBFD" blockId="6.[151,1437,151,1982]" pageId="6" pageNumber="72">
|
||
Dorsals granular, oval-shaped at base, heterogeneous in size, but majority slightly larger than scales on top of head. Transition between scales on flanks and ventrals abrupt but with a few scales of intermediate morphology. Ventral scales distinctly larger than dorsals, slightly smaller on chest than on belly, smooth, imbricate (each scale overlapping anterior portion of scale lying posteriorly), with round posterior margin, arranged for the most part in oblique rows. There are 47 scales along the midventral line between the level of anterior margin of forelimbs and the small scales anterior to vent. Scales around midbody about 81, of which about 17 are ventrals. Scales on precloacal region similar to ventrals, except proximally to vent, which has minute scales arranged in several rows. Escutcheon distinct, with a vertex projecting anteriorly just past level of groin; posterior margin of escutcheon slightly concave,
|
||
<quantity id="4CB6639AFF91E5211810FC04FE40FC1E" box="[354,443,980,1004]" metricMagnitude="-3" metricUnit="m" metricValue="3.3" pageId="6" pageNumber="72" unit="mm" value="3.3">3.3 mm</quantity>
|
||
from vent. On ventral surfaces of posterior limbs, escutcheon is 3–4 scales wide and extends medially for almost full extent of thighs.
|
||
</paragraph>
|
||
<paragraph id="8BF1CE7FFF91E52119B5FBCCFD56FB15" blockId="6.[151,1437,151,1982]" pageId="6" pageNumber="72">
|
||
Scales dorsally on tail slightly larger than on body, flat, slightly imbricate, with round posterior margin; a short transition area between the granular body scales and the caudal scales just described, extends posteriorly about
|
||
<quantity id="4CB6639AFF91E5211CFCFBEFFF38FB8E" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="6" pageNumber="72" unit="mm" value="4.0">4 mm</quantity>
|
||
from base of tail. Underside of tail covered with flat, wide, short, roughly rectangular plates on regenerated part. Underside of unregenerated part of tail extends about
|
||
<quantity id="4CB6639AFF91E5211A40FB57FC8FFB52" box="[818,884,1159,1184]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" pageId="6" pageNumber="72" unit="mm" value="8.0">8 mm</quantity>
|
||
from vent, of which proximal
|
||
<quantity id="4CB6639AFF91E5211DA2FB57FAE9FB52" box="[1232,1298,1159,1184]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="6" pageNumber="72" unit="mm" value="4.0">4 mm</quantity>
|
||
are covered with scales like those in venter but slightly smaller and distal
|
||
<quantity id="4CB6639AFF91E5211AFFFB7CFC2FFB36" box="[909,980,1196,1220]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="6" pageNumber="72" unit="mm" value="4.0">4 mm</quantity>
|
||
show the beginning of the pattern of differentiated scales (subcaudal pattern
|
||
<typeStatus id="54F570DDFF91E5211B21FB01FD78FB1B" box="[595,643,1233,1257]" pageId="6" pageNumber="72">type</typeStatus>
|
||
B).
|
||
</paragraph>
|
||
<paragraph id="8BF1CE7FFF91E52119B5FB24FB8FF9DE" blockId="6.[151,1437,151,1982]" pageId="6" pageNumber="72">
|
||
Scales on anterior limbs tiny, granular and juxtaposed, except narrowly on anterior margin of forearm, on which scales tend to be bigger, flat and imbricate. Scales on posterior limbs small, granular and juxtaposed on dorsal, posterior and posterior half of ventral surfaces; flat, imbricate, about same size as scales on venter, on anterior and anterior half of ventral surface; on ventral surface of posterior limbs there is a sharp transition between the two
|
||
<typeStatus id="54F570DDFF91E5211987FA55FECBFA6F" box="[245,304,1413,1437]" pageId="6" pageNumber="72">types</typeStatus>
|
||
of scales that coincides with posterior margin of escutcheon. Lamellae under first (
|
||
<emphasis id="B93A126DFF91E5211D9CFA54FB02FA6E" bold="true" box="[1262,1273,1412,1436]" pageId="6" pageNumber="72">I</emphasis>
|
||
) through fifth (
|
||
<emphasis id="B93A126DFF91E52119D2FA78FF4FFA32" bold="true" box="[160,180,1448,1472]" pageId="6" pageNumber="72">V</emphasis>
|
||
) finger (infraproximals in parentheses; right/left):
|
||
<emphasis id="B93A126DFF91E5211B80FA78FD06FA32" bold="true" box="[754,765,1448,1472]" pageId="6" pageNumber="72">I</emphasis>
|
||
: 14/14(3/3),
|
||
<emphasis id="B93A126DFF91E5211AE2FA78FC5DFA32" bold="true" box="[912,934,1448,1472]" pageId="6" pageNumber="72">II</emphasis>
|
||
: 20/19 (6/6),
|
||
<emphasis id="B93A126DFF91E5211D4DFA78FB9BFA32" bold="true" box="[1087,1120,1448,1472]" pageId="6" pageNumber="72">III</emphasis>
|
||
: 23/24(7/7),
|
||
<emphasis id="B93A126DFF91E5211D81FA78FAEBFA32" bold="true" box="[1267,1296,1448,1472]" pageId="6" pageNumber="72">IV</emphasis>
|
||
: 24/24(8/8), and
|
||
<emphasis id="B93A126DFF91E52119B4FA1CFF23FA16" bold="true" box="[198,216,1484,1508]" pageId="6" pageNumber="72">V</emphasis>
|
||
: 19/19(5/5), respectively. Lamellae under first through fifth toe (infraproximals in parentheses):
|
||
<emphasis id="B93A126DFF91E5211C65FA1CFAD9FA16" bold="true" box="[1303,1314,1484,1508]" pageId="6" pageNumber="72">I</emphasis>
|
||
:?/11(?/2),
|
||
<emphasis id="B93A126DFF91E52119E5FA20FF56F9FA" bold="true" box="[151,173,1520,1544]" pageId="6" pageNumber="72">II</emphasis>
|
||
: 18/19(4/6),
|
||
<emphasis id="B93A126DFF91E5211830FA20FE98F9FA" bold="true" box="[322,355,1520,1544]" pageId="6" pageNumber="72">III</emphasis>
|
||
: 24/24(8/7),
|
||
<emphasis id="B93A126DFF91E5211885FA20FDEFF9FA" bold="true" box="[503,532,1520,1544]" pageId="6" pageNumber="72">IV</emphasis>
|
||
: 26/26(10/9), and
|
||
<emphasis id="B93A126DFF91E5211B9AFA20FD01F9FA" bold="true" box="[744,762,1520,1544]" pageId="6" pageNumber="72">V</emphasis>
|
||
: 22/22(6/6), respectively. Fingers and toes with four lateral rows of scales distally. Claws exposed, non-retractile, between two or three basal scales.
|
||
</paragraph>
|
||
<paragraph id="8BF1CE7FFF91E52F19B5F9E7FEA1FF26" blockId="6.[151,1437,151,1982]" lastBlockId="8.[151,1436,151,1255]" lastPageId="8" lastPageNumber="74" pageId="6" pageNumber="72">
|
||
<emphasis id="B93A126DFF91E52119B5F9E7FDD3F9A2" bold="true" box="[199,552,1591,1616]" pageId="6" pageNumber="72">Color in life and preservative</emphasis>
|
||
. In life (
|
||
<figureCitation id="1375D2FAFF91E5211BFEF9E7FD2BF9BD" box="[652,720,1591,1615]" captionStart="FIGURE 2" captionStartId="3.[151,250,1844,1867]" captionTargetBox="[251,1340,359,1805]" captionTargetId="figure@3.[234,1353,343,1823]" captionTargetPageId="3" captionText="FIGURE 2. Adult male (top, holotype, MHNLS 20780) and adult female (bottom, UTA 60126) of Gonatodes rozei sp. nov. in life." httpUri="https://zenodo.org/record/282622/files/figure.png" pageId="6" pageNumber="72">Fig. 2</figureCitation>
|
||
) the dorsal and lateral surfaces of the body and front legs have dark, smoky gray background coloration. This background coloration turns grayish brown dorsally and laterally on the tail and the back legs. The dorsum, flanks, front legs and thighs, are covered with black edged, white ocelli. The black edges are vaguely defined, as they gradually fade into the background coloration. The largest ocelli are about one third (white region only) the size of the eye and are located on the anterior part of the body. The ocelli become smaller and sparser on the posterior part of the body and do not extend past the base of the tail. A bright yellow hood covers the head completely, including ventrally, extending posteriorly to the level of the anterior margin of the front limbs. This yellow hood has some areas suffused with orange, most significantly on a patch on the left temporal region, which seems to be a scar possibly produced by male-male combat. The ventral surface of the body, except the scutcheon area, appears macroscopically as a mottled pale grey pattern on a white background; under magnification the mottled appearance is conferred by the arrangement of melanophores, which are concentrated towards the edges, on the ventral scales. The color in preservative has not changed significantly, except the head, which has faded to cream, and the background coloration of the body, which has faded slightly to a paler gray tone.
|
||
</paragraph>
|
||
<caption id="DF319EF7FF90E52019E5F86AFEC1F81D" httpUri="https://zenodo.org/record/282624/files/figure.png" pageId="7" pageNumber="73" targetBox="[206,1388,184,1932]" targetPageId="7">
|
||
<paragraph id="8BF1CE7FFF90E52019E5F86AFEC1F81D" blockId="7.[151,1436,1978,2031]" pageId="7" pageNumber="73">
|
||
<emphasis id="B93A126DFF90E52019E5F86AFEE8F822" bold="true" box="[151,275,1978,2001]" pageId="7" pageNumber="73">FIGURE 4.</emphasis>
|
||
The four different male color morphs in
|
||
<taxonomicName id="4C4EB5FCFF90E5201BC8F86BFC9AF823" box="[698,865,1979,2001]" class="Reptilia" family="Sphaerodactylidae" genus="Gonatodes" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="73" phylum="Chordata" rank="species" species="rozei" status="sp. nov.">
|
||
<emphasis id="B93A126DFF90E5201BC8F86BFC9AF823" box="[698,865,1979,2001]" italics="true" pageId="7" pageNumber="73">Gonatodes rozei</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B93A126DFF90E5201A1AF86BFC42F822" bold="true" box="[872,953,1979,2000]" pageId="7" pageNumber="73">
|
||
<taxonomicNameLabel id="A209AF16FF90E5201A1AF86BFC42F822" box="[872,953,1979,2000]" pageId="7" pageNumber="73" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, I-IV, arranged from top to bottom. Illustration by Emil Bröckl.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BF1CE7FFF9FE52F19B5FF0FFEAAFDAD" blockId="8.[151,1436,151,1255]" pageId="8" pageNumber="74">
|
||
<emphasis id="B93A126DFF9FE52F19B5FF0FFEC0FF0A" bold="true" box="[199,315,223,248]" pageId="8" pageNumber="74">Variation.</emphasis>
|
||
The largest male is the
|
||
<typeStatus id="54F570DDFF9FE52F1B2FFF0FFD44FF05" box="[605,703,223,247]" pageId="8" pageNumber="74" type="holotype">holotype</typeStatus>
|
||
measuring
|
||
<quantity id="4CB6639AFF9FE52F1A3AFF0FFC54FF05" box="[840,943,223,247]" metricMagnitude="-2" metricUnit="m" metricValue="5.91" pageId="8" pageNumber="74" unit="mm" value="59.1">59.1 mm</quantity>
|
||
in SVL. TCWC 59312 and 59320 are the largest females both measuring
|
||
<quantity id="4CB6639AFF9FE52F1889FED4FDB2FEEE" box="[507,585,260,284]" metricMagnitude="0" metricUnit="m" metricValue="1.40462" pageId="8" pageNumber="74" unit="in" value="55.3">55.3 in</quantity>
|
||
SVL. There are typically five, less frequently four or six supralabials to the level of the center of the eye, followed posteriorly by 1–2 small polygonal scales and subsequently by multiple tiny granular scales to the rictus of the mouth. There are typically four or five, rarely six, infralabials to the level of the center of eye, followed posteriorly by 1–2 small polygonal scales and subsequently by multiple tiny granular scales. Postrostrals 3–4. Postmentals 2–4. Loreal scales 13–16. Scales around the midbody are 78–100 of which about 13–23 are ventrals. There are 35–41 midventral scales along the trunk. Lamellae (infraproximals in parentheses) under the fingers are:
|
||
<emphasis id="B93A126DFF9FE52F1B5EFE0CFDCCFE06" bold="true" box="[556,567,476,500]" pageId="8" pageNumber="74">I</emphasis>
|
||
: 11–16 (2–3),
|
||
<emphasis id="B93A126DFF9FE52F1B96FE0CFD01FE06" bold="true" box="[740,762,476,500]" pageId="8" pageNumber="74">II</emphasis>
|
||
: 18–22 (5–7),
|
||
<emphasis id="B93A126DFF9FE52F1ADBFE0CFC31FE06" bold="true" box="[937,970,476,500]" pageId="8" pageNumber="74">III</emphasis>
|
||
: 22–26 (6–8),
|
||
<emphasis id="B93A126DFF9FE52F1D0AFE0CFB6EFE06" bold="true" box="[1144,1173,476,500]" pageId="8" pageNumber="74">IV</emphasis>
|
||
: 22–27 (7–11), and
|
||
<emphasis id="B93A126DFF9FE52F1CF0FE0CFA6FFE06" bold="true" box="[1410,1428,476,500]" pageId="8" pageNumber="74">V</emphasis>
|
||
: 19–22 (4–7). Lamellae (infraproximals in parentheses) under the toes are:
|
||
<emphasis id="B93A126DFF9FE52F1A8FFDD0FBF3FDEA" bold="true" box="[1021,1032,512,536]" pageId="8" pageNumber="74">I</emphasis>
|
||
: 9–13 (2–3),
|
||
<emphasis id="B93A126DFF9FE52F1DDEFDD0FB39FDEA" bold="true" box="[1196,1218,512,536]" pageId="8" pageNumber="74">II</emphasis>
|
||
: 17–22 (4–7),
|
||
<emphasis id="B93A126DFF9FE52F1C06FDD0FA6EFDEA" bold="true" box="[1396,1429,512,536]" pageId="8" pageNumber="74">III</emphasis>
|
||
: 23–28 (6–10),
|
||
<emphasis id="B93A126DFF9FE52F184EFDF4FEA2FDCE" bold="true" box="[316,345,548,572]" pageId="8" pageNumber="74">IV</emphasis>
|
||
: 25–31 (8–13), and
|
||
<emphasis id="B93A126DFF9FE52F1B49FDF4FDB6FDCE" bold="true" box="[571,589,548,572]" pageId="8" pageNumber="74">V</emphasis>
|
||
: 21–25 (5–9). Fingers and toes typically with four, rarely three, lateral rows of scales distally
|
||
</paragraph>
|
||
<paragraph id="8BF1CE7FFF9FE52F19B5FDBCFAD3FD55" blockId="8.[151,1436,151,1255]" pageId="8" pageNumber="74">
|
||
<collectingCountry id="F3598EEFFF9FE52F19B5FDBCFF1DFD76" box="[199,230,620,644]" name="American Samoa" pageId="8" pageNumber="74">As</collectingCountry>
|
||
mentioned above,
|
||
<taxonomicName id="4C4EB5FCFF9FE52F18CDFDBDFDEEFD76" box="[447,533,621,644]" class="Reptilia" family="Sphaerodactylidae" genus="Gonatodes" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="74" phylum="Chordata" rank="species" species="rozei">
|
||
<emphasis id="B93A126DFF9FE52F18CDFDBDFE2BFD76" box="[447,464,621,644]" italics="true" pageId="8" pageNumber="74">G</emphasis>
|
||
.
|
||
<emphasis id="B93A126DFF9FE52F18ADFDBDFDEEFD76" box="[479,533,621,644]" italics="true" pageId="8" pageNumber="74">rozei</emphasis>
|
||
</taxonomicName>
|
||
is the most polymorphic species of
|
||
<taxonomicName id="4C4EB5FCFF9FE52F1AC0FDBCFBD0FD76" box="[946,1067,620,644]" class="Reptilia" family="Sphaerodactylidae" genus="Gonatodes" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="74" phylum="Chordata" rank="genus">
|
||
<emphasis id="B93A126DFF9FE52F1AC0FDBCFBD0FD76" box="[946,1067,620,644]" italics="true" pageId="8" pageNumber="74">Gonatodes</emphasis>
|
||
</taxonomicName>
|
||
in terms of male coloration. We have been able to group all male specimens into four different color morphs (
|
||
<figureCitation id="1375D2FAFF9FE52F1A8AFD5FFBC0FD55" box="[1016,1083,655,679]" captionStart="FIGURE 4" captionStartId="7.[151,250,1978,2001]" captionTargetBox="[206,1388,184,1932]" captionTargetId="figure@7.[190,1395,159,1970]" captionTargetPageId="7" captionText="FIGURE 4. The four different male color morphs in Gonatodes rozei sp. nov., I-IV, arranged from top to bottom. Illustration by Emil Bröckl." httpUri="https://zenodo.org/record/282624/files/figure.png" pageId="8" pageNumber="74">Fig. 4</figureCitation>
|
||
) as described below:
|
||
</paragraph>
|
||
<paragraph id="8BF1CE7FFF9FE52F19B5FD64FF2FFCC5" blockId="8.[151,1436,151,1255]" pageId="8" pageNumber="74">
|
||
Morph I: This is the color morph of the
|
||
<typeStatus id="54F570DDFF9FE52F1BF7FD64FD10FD3E" box="[645,747,692,716]" pageId="8" pageNumber="74" type="holotype">holotype</typeStatus>
|
||
. The notable variation observed for this color morph includes the color of the cephalic hood which varies from yellow to orange and the number of ocelli on the sides of the body. Some specimens have only a few ocelli in which case they are always concentrated on the anterior portion of the body.
|
||
</paragraph>
|
||
<paragraph id="8BF1CE7FFF9FE52F19B5FC94FE59FC1E" blockId="8.[151,1436,151,1255]" pageId="8" pageNumber="74">Morph II: In this morph individuals have pale grayish brown background coloration on the dorsal and lateral surfaces of the body, tail and limbs. The body is covered by white, vaguely defined spots and ocelli. The latter have diffused dark gray edges and tend to be located laterally on the anterior portion of the body. The head is slightly darker than the body and it is overlaid with irregular and vaguely defined, pale yellow markings. Limbs have suffused pale markings.</paragraph>
|
||
<paragraph id="8BF1CE7FFF9FE52F19B5FC27FA85FB8E" blockId="8.[151,1436,151,1255]" pageId="8" pageNumber="74">Morph III: The dorsal and lateral surfaces of the body and legs are brownish gray to gray, with a few irregular, large, black-edged ocelli located laterally on the anterior portion of the body, most frequently in the scapular region. The head and neck is heavily overlaid with irregular and elongated yellow markings that extend to the gular region. Black suffusions are frequent in the space between the yellow markings. The tail is grayish brown.</paragraph>
|
||
<paragraph id="8BF1CE7FFF9FE52F19B5FB57FE06FB15" blockId="8.[151,1436,151,1255]" pageId="8" pageNumber="74">Morph IV: The dorsal and lateral surfaces of the body are gray and, in contrast to the previous three color morphs, are devoid of any white markings. The head is uniformly dull yellow, including the gular region, which is slightly paler. The tail is brown.</paragraph>
|
||
<caption id="DF319EF7FF9FE52F19E5F86FFC93F827" box="[151,872,1983,2006]" httpUri="https://zenodo.org/record/282625/files/figure.png" pageId="8" pageNumber="74" targetBox="[151,1435,1309,1961]" targetPageId="8">
|
||
<paragraph id="8BF1CE7FFF9FE52F19E5F86FFC93F827" blockId="8.[151,872,1983,2006]" box="[151,872,1983,2006]" pageId="8" pageNumber="74">
|
||
<emphasis id="B93A126DFF9FE52F19E5F86FFEE9F827" bold="true" box="[151,274,1983,2006]" pageId="8" pageNumber="74">FIGURE 5.</emphasis>
|
||
A pair of
|
||
<taxonomicName id="4C4EB5FCFF9FE52F1809F86FFDDBF827" box="[379,544,1983,2005]" class="Reptilia" family="Sphaerodactylidae" genus="Gonatodes" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="74" phylum="Chordata" rank="species" species="rozei" status="sp. nov.">
|
||
<emphasis id="B93A126DFF9FE52F1809F86FFDDBF827" box="[379,544,1983,2005]" italics="true" pageId="8" pageNumber="74">Gonatodes rozei</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B93A126DFF9FE52F1B54F810FD8CF827" bold="true" box="[550,631,1984,2005]" pageId="8" pageNumber="74">
|
||
<taxonomicNameLabel id="A209AF16FF9FE52F1B54F810FD8CF827" box="[550,631,1984,2005]" pageId="8" pageNumber="74" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
copulating in captivity.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BF1CE7FFF9EE52E19B5FF48FB25FE7A" blockId="9.[151,1436,151,392]" pageId="9" pageNumber="75">By far the most common color morph is “I”, which is present in all localities for which several specimens have been obtained. In some localities, like those within Guatopo National Park, almost all male specimens are color morph “I” although color morph “IV” has also been observed at lower frequency in this area. Color morph “IV” has been found in several localities from throughout the distribution of the species and seems to be common in Hacienda El Limón. Color morph “III” has only been recorded in Capaya, Miranda, and Carayaca, Vargas (color morphs “I” and “IV” have also been collected in both localities). Color morph “II” has only been recorded in Carayaca and it is in this locality that all four male color morphs have been observed in sympatry.</paragraph>
|
||
<caption id="DF319EF7FF9EE52E19E5FB0CFDE6FABD" httpUri="https://zenodo.org/record/282626/files/figure.png" pageId="9" pageNumber="75" targetBox="[151,1436,445,1223]" targetPageId="9">
|
||
<paragraph id="8BF1CE7FFF9EE52E19E5FB0CFDE6FABD" blockId="9.[151,1436,1244,1359]" pageId="9" pageNumber="75">
|
||
<emphasis id="B93A126DFF9EE52E19E5FB0CFEEFFB00" bold="true" box="[151,276,1244,1267]" pageId="9" pageNumber="75">FIGURE 6.</emphasis>
|
||
Map of north central Venezuela showing the collecting localities of the specimens examined of
|
||
<taxonomicName id="4C4EB5FCFF9EE52E1D87FB0DFA67FB01" box="[1269,1436,1245,1267]" class="Reptilia" family="Sphaerodactylidae" genus="Gonatodes" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="75" phylum="Chordata" rank="species" species="rozei" status="sp. nov.">
|
||
<emphasis id="B93A126DFF9EE52E1D87FB0DFA67FB01" box="[1269,1436,1245,1267]" italics="true" pageId="9" pageNumber="75">Gonatodes rozei</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B93A126DFF9EE52E19E5FB2CFF11FAE3" bold="true" box="[151,234,1276,1297]" pageId="9" pageNumber="75">
|
||
<taxonomicNameLabel id="A209AF16FF9EE52E19E5FB2CFF11FAE3" box="[151,234,1276,1297]" pageId="9" pageNumber="75" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
. The holotype is indicated by the star and all other localities by dots. The whole type series was collected within Guatopo National Park, which is delimited by a dashed line. The other three national parks are Waraira Repano (top), Macarao (left) and Laguna de Tacarigua (right).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BF1CE7FFF9EE52E19B5FAACFD8DF879" blockId="9.[151,1437,1403,2003]" pageId="9" pageNumber="75">In females and juveniles the ground color of the body, except ventrally, and the head is brown to grayish brown. There is a pale, creamish brown vertebral stripe that extends from the neck or the occipital region onto the tail, where it becomes poorly defined and disappears on the distal portion. The vertebral stripe has undulating edges, which are typically accentuated by a series of paired, small, diffuse black and white spots. On the sides of the body up to three longitudinal rows of white/pale creamish brown spots are present; of which only one row is conspicuous (spots are white rather than pale creamish brown). The largest pale, lateral spots, are about the size of the pupil. Also, the sides of the body have fine, diffuse, irregular black reticulations; on the limbs these reticulations become more conspicuous. The top of the head might be devoid of any conspicuous markings or might have fine and diffuse black markings forming poorly defined canthal, postocular and parietal/occipital stripes. Irregular and diffuse pale marking might also be present on the top of the head. The whole ventral area tends to paler than the dorsum and in large females the belly and underside of the limbs might be suffused with yellow or a ferruginous tinge. The gular area is typically white or cream with longitudinal, fine, diffuse dark brown stripes. The tail commonly has pale (pale pink, pale brown or white) rings which are more conspicuous ventrally and on the posterior portion of the tail. These rings are usually edged anteriorly by black markings. The rings are much less pronounced on large specimens.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3549DF4FF9EE52D19B5F847FC8EFD55" lastPageId="10" lastPageNumber="76" pageId="9" pageNumber="75" type="distribution">
|
||
<paragraph id="8BF1CE7FFF9EE52D19B5F847FBF0FE3D" blockId="9.[151,1437,1403,2003]" lastBlockId="10.[151,1436,151,824]" lastPageId="10" lastPageNumber="76" pageId="9" pageNumber="75">
|
||
<emphasis id="B93A126DFF9EE52E19B5F847FDAEF842" bold="true" box="[199,597,1943,1968]" pageId="9" pageNumber="75">Distribution and natural history.</emphasis>
|
||
The species is found in tropical and premontane humid forests (
|
||
<bibRefCitation id="EFDFB38EFF9EE52E1C33F847FEDCF821" author="Ewel" pageId="9" pageNumber="75" refString="Ewel, J. J. & Madriz, A. (1968). Zonas de vida de Venezuela. Ministerio de Agricultura y Cria, Carcas, Venezuela." type="book" year="1968">Ewel & Madriz 1968</bibRefCitation>
|
||
) on both chains (Serranía del Litoral and Serranía del Interior) of the Cordillera de la Costa Central in north central
|
||
<collectingCountry id="F3598EEFFF9DE52D185CFF47FE5AFF5D" box="[302,417,151,175]" name="Venezuela" pageId="10" pageNumber="76">Venezuela</collectingCountry>
|
||
(
|
||
<figureCitation id="1375D2FAFF9DE52D18C3FF47FE0CFF5D" box="[433,503,151,175]" captionStart="FIGURE 6" captionStartId="9.[151,250,1244,1267]" captionTargetBox="[151,1436,445,1223]" captionTargetId="figure@9.[151,1436,445,1223]" captionTargetPageId="9" captionText="FIGURE 6. Map of north central Venezuela showing the collecting localities of the specimens examined of Gonatodes rozei sp. nov.. The holotype is indicated by the star and all other localities by dots. The whole type series was collected within Guatopo National Park, which is delimited by a dashed line. The other three national parks are Waraira Repano (top), Macarao (left) and Laguna de Tacarigua (right)." httpUri="https://zenodo.org/record/282626/files/figure.png" pageId="10" pageNumber="76">Fig. 6</figureCitation>
|
||
). The species is active during the day and can be commonly observed on trees but close to ground level. It has also been taken from karstic walls outside caves and occasionally individuals have been observed on buildings in areas with large trees near forest habitat. The species has been collected at low elevations (<
|
||
<quantity id="4CB6639AFF9DE52D1855FED4FE99FEEE" box="[295,354,260,284]" metricMagnitude="1" metricUnit="m" metricValue="5.0" pageId="10" pageNumber="76" unit="m" value="50.0">50 m</quantity>
|
||
asl), near the coast, on the western foothills of the Serranía del Litoral (e.g. Carenero and on the road between Chuspa and Higuerote). The locality with highest elevation at which the species has been collected is Taica, Miranda, at about
|
||
<quantity id="4CB6639AFF9DE52D18DCFE9CFDF9FE96" box="[430,514,332,356]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="10" pageNumber="76" unit="m" value="1000.0">1000 m</quantity>
|
||
above sea level. A couple specimens (MCNC 3733-3734) have been collected on “Pico Naiguatá” Mountain, which exceeds
|
||
<quantity id="4CB6639AFF9DE52D1B0AFEBFFD37FE75" box="[632,716,367,391]" metricMagnitude="3" metricUnit="m" metricValue="2.7" pageId="10" pageNumber="76" unit="m" value="2700.0">2700 m</quantity>
|
||
above sea level, but the lack of specific locality data prevents us from determining the precise elevation at which these specimens were collected. However, the other localities with more precise information from this mountain are below
|
||
<quantity id="4CB6639AFF9DE52D1A79FE67FCA9FE3D" box="[779,850,439,463]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="10" pageNumber="76" unit="m" value="800.0">800 m</quantity>
|
||
above sea level.
|
||
</paragraph>
|
||
<paragraph id="8BF1CE7FFF9DE52D19B5FE0DFC8EFD55" blockId="10.[151,1436,151,824]" pageId="10" pageNumber="76">
|
||
We also have a few notes on the reproductive biology of
|
||
<taxonomicName id="4C4EB5FCFF9DE52D1A33FE0DFC6EFE06" box="[833,917,477,500]" class="Reptilia" family="Sphaerodactylidae" genus="Gonatodes" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="76" phylum="Chordata" rank="species" species="rozei">
|
||
<emphasis id="B93A126DFF9DE52D1A33FE0DFCA9FE06" box="[833,850,477,500]" italics="true" pageId="10" pageNumber="76">G</emphasis>
|
||
.
|
||
<emphasis id="B93A126DFF9DE52D1A2DFE0DFC6EFE06" box="[863,917,477,500]" italics="true" pageId="10" pageNumber="76">rozei</emphasis>
|
||
</taxonomicName>
|
||
obtained from individuals kept in captivity by CRB. On
|
||
<date id="FFF0E8BFFF9DE52D187AFDD0FE3CFDE5" box="[264,455,511,536]" pageId="10" pageNumber="76" value="1966-08-18">August 18, 1966</date>
|
||
, at about 08h15, a male and female
|
||
<taxonomicName id="4C4EB5FCFF9DE52D1A16FDD0FC41FDEA" box="[868,954,512,536]" class="Reptilia" family="Sphaerodactylidae" genus="Gonatodes" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="76" phylum="Chordata" rank="species" species="rozei">
|
||
<emphasis id="B93A126DFF9DE52D1A16FDD0FC8EFDE5" box="[868,885,512,535]" italics="true" pageId="10" pageNumber="76">G</emphasis>
|
||
.
|
||
<emphasis id="B93A126DFF9DE52D1AF6FDD1FC41FDEA" box="[900,954,513,536]" italics="true" pageId="10" pageNumber="76">rozei</emphasis>
|
||
</taxonomicName>
|
||
that had been kept in captivity for several months were found copulating (
|
||
<figureCitation id="1375D2FAFF9DE52D1B70FDF4FDB3FDCE" box="[514,584,548,572]" captionStart="FIGURE 5. A" captionStartId="8.[151,250,1983,2006]" captionTargetBox="[151,1435,1309,1961]" captionTargetId="figure@8.[151,1436,1309,1962]" captionTargetPageId="8" captionText="FIGURE 5. A pair of Gonatodes rozei sp. nov. copulating in captivity." httpUri="https://zenodo.org/record/282625/files/figure.png" pageId="10" pageNumber="76">Fig. 5</figureCitation>
|
||
). The male was biting the back of the neck of the female and had both his right front- and hind-legs over the dorsum of the female. The tail of the female was raised over the base of the tail of the male. Regrettably, the pair separated as soon as the first picture was shot. Subsequently, the male retreated to the top part of the cage and proceeded to lick his right hemipenis.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3549DF4FF9DE52D19B5FD64FBE5FCC5" pageId="10" pageNumber="76" type="discussion">
|
||
<paragraph id="8BF1CE7FFF9DE52D19B5FD64FBE5FCC5" blockId="10.[151,1436,151,824]" pageId="10" pageNumber="76">
|
||
<emphasis id="B93A126DFF9DE52D19B5FD64FEC0FD3E" bold="true" box="[199,315,692,716]" pageId="10" pageNumber="76">Remarks.</emphasis>
|
||
<taxonomicName id="4C4EB5FCFF9DE52D1837FD64FDFAFD3E" box="[325,513,692,716]" class="Reptilia" family="Sphaerodactylidae" genus="Gonatodes" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="76" phylum="Chordata" rank="species" species="rozei">
|
||
<emphasis id="B93A126DFF9DE52D1837FD64FDFAFD3E" box="[325,513,692,716]" italics="true" pageId="10" pageNumber="76">Gonatodes rozei</emphasis>
|
||
</taxonomicName>
|
||
has been known among herpetologists for more than 30 years and it has been frequently collected near the city of Caracas. The name
|
||
<taxonomicName id="4C4EB5FCFF9DE52D1A6DFD08FC8DFD02" box="[799,886,728,752]" class="Reptilia" family="Sphaerodactylidae" genus="Gonatodes" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="76" phylum="Chordata" rank="species" species="rozei">
|
||
<emphasis id="B93A126DFF9DE52D1A6DFD08FCCBFD1D" box="[799,816,728,751]" italics="true" pageId="10" pageNumber="76">G</emphasis>
|
||
.
|
||
<emphasis id="B93A126DFF9DE52D1A32FD09FC8DFD02" box="[832,886,729,752]" italics="true" pageId="10" pageNumber="76">rozei</emphasis>
|
||
</taxonomicName>
|
||
was used in the first author’s unpublished PhD dissertation (
|
||
<bibRefCitation id="EFDFB38EFF9DE52D1858FD2CFDEFFCE6" author="Rivero-Blanco" box="[298,532,764,788]" pageId="10" pageNumber="76" refString="Rivero-Blanco, C. (1979) The neotropical lizard genus Gonatodes Fitzinger (Sauria: Sphaerodactylinae). Ph. D. Dissertation. Texas A & M University, College Station, Texas." type="book" year="1979">Rivero-Blanco 1979</bibRefCitation>
|
||
) but, even though herpetologists have used the name informally over internet gecko forums, to our knowledge it has never appeared in the scientific literature.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |