treatments-xml/data/03/E6/ED/03E6ED6DA679BF6AFF50B474EEEE9C41.xml

<document id="EDA64E41D755EA09B4B9BC2B9BADCB64" ID-CLB-Dataset="26477" ID-DOI="10.11646/zootaxa.4615.3.8" ID-GBIF-Dataset="9238318c-4409-45cc-96cf-1aa17c4006d0" ID-ISSN="1175-5326" ID-Zenodo-Dep="3246211" ID-ZooBank="2AA8C6E7-11D5-4443-8E0E-9FB19856B0CB" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1560509668797" checkinUser="plazi" docAuthor="Mondaca, José, Pizarro-Araya, Jaime &amp; Alfaro, Fermín M." docDate="2019" docId="03E6ED6DA679BF6AFF50B474EEEE9C41" docLanguage="en" docName="zootaxa.4615.3.8.pdf" docOrigin="Zootaxa 4615 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Luispenaia paulseni Mondaca &amp; Pizarro-Araya &amp; Alfaro 2019, new species" docType="treatment" docUuid="5FB6F8DC-8572-454C-902C-1A21A6FCB402" docUuidSource="ZooBank" docVersion="8" lastPageNumber="560" masterDocId="FFDF9515A670BF66FFC7B247ED0A9D29" masterDocTitle="Revision of the genus Luispenaia Martínez (Coleoptera: Scarabaeidae: Melolonthinae: Tanyproctini), with description of three new species from the Atacama Desert, Chile" masterLastPageNumber="562" masterPageNumber="549" pageNumber="558" updateTime="1698734911197" updateUser="ExternalLinkService">
<mods:mods id="C4324D0754454206579FE14708A28521" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="A970990D74A948A1393D9B54CA66B842">
<mods:title id="92BE75A40C3F095D1235C996DCC4BF50">Revision of the genus Luispenaia Martínez (Coleoptera: Scarabaeidae: Melolonthinae: Tanyproctini), with description of three new species from the Atacama Desert, Chile</mods:title>
</mods:titleInfo>
<mods:name id="38564557E7D316CF3AF7B6601400100C" type="personal">
<mods:role id="03402492B0426A290E0CF58519EA44DD">
<mods:roleTerm id="31B1E6B57F6B7B1425FA67779E679B63">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="82025394AC62658E5F89FB061A4726F8">Mondaca, José</mods:namePart>
</mods:name>
<mods:name id="CFB19D6F014F37B418814607C47C3E33" type="personal">
<mods:role id="510829C9A2B50DB45A20136E8312BAC3">
<mods:roleTerm id="8F63F1CC6D2276557DD19389B2807B30">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="C732F108737AB6C061FBF17497313654">Pizarro-Araya, Jaime</mods:namePart>
</mods:name>
<mods:name id="182953C0955BE89373DAAEF42E52189F" type="personal">
<mods:role id="D148581EDA54994EAACE4E4DDC4AD37B">
<mods:roleTerm id="2FA11FF75BE886A710847FC9287BBF5C">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="1A0302F20D964F7398E43EF44F13714F">Alfaro, Fermín M.</mods:namePart>
</mods:name>
<mods:typeOfResource id="BF064B4F9F647503DBB509D2F31AAD26">text</mods:typeOfResource>
<mods:relatedItem id="7E04A98D0CA3D546B958A28FA40AED7B" type="host">
<mods:titleInfo id="2F0715B225E7DA00130A9A713B96D334">
<mods:title id="B0C247028A6BF5E8868E3AFD5171AE4E">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="9F08654E1263AC868C3F40A92430FA03">
<mods:date id="376B1940A0EC4E116DD340865FA91698">2019</mods:date>
<mods:detail id="BCE4AB3BB198C4BB4B5780B71A5FC374" type="pubDate">
<mods:number id="1EBC72C4BAD54D1FCB3C81AB3CDBDC3E">2019-06-14</mods:number>
</mods:detail>
<mods:detail id="4216727DAB9EB7BB216A745F4709A426" type="volume">
<mods:number id="10C69F306038DDB2D0B3602710F87A9C">4615</mods:number>
</mods:detail>
<mods:detail id="5B61FA6FE6FC4749E92DC8D0B58B2D17" type="issue">
<mods:number id="AFAE5BC7D34E9EC08D373C014F0A39A8">3</mods:number>
</mods:detail>
<mods:extent id="7FE57D8FAA5E6E9136354B378E933E88" unit="page">
<mods:start id="5A5FA4CAAA4FD63489551DD280DC82D5">549</mods:start>
<mods:end id="7CAB05AEC9D354E48BDEF4F003D3FD27">562</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="C02BD8BCD3658611ADD3B12989F67CC2">journal article</mods:classification>
<mods:identifier id="B9A4EE87ECABAF7A70F9A859C987A27C" type="CLB-Dataset">26477</mods:identifier>
<mods:identifier id="E86B728BAFAA4D16698AE00FDEE926A6" type="DOI">10.11646/zootaxa.4615.3.8</mods:identifier>
<mods:identifier id="FD1C77B938DB5B40F51B0F503ACB8194" type="GBIF-Dataset">9238318c-4409-45cc-96cf-1aa17c4006d0</mods:identifier>
<mods:identifier id="9305DB99323EFBE74B7EA0BA6DF46C83" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="32A40721F105379B7CB14C4E85CBD900" type="Zenodo-Dep">3246211</mods:identifier>
<mods:identifier id="16F516AA307307BE861FC65435FBED1C" type="ZooBank">2AA8C6E7-11D5-4443-8E0E-9FB19856B0CB</mods:identifier>
</mods:mods>
<treatment id="03E6ED6DA679BF6AFF50B474EEEE9C41" ID-GBIF-Taxon="158223212" LSID="urn:lsid:zoobank.org:act:5FB6F8DC-8572-454C-902C-1A21A6FCB402" httpUri="http://treatment.plazi.org/id/03E6ED6DA679BF6AFF50B474EEEE9C41" lastPageId="12" lastPageNumber="560" pageId="9" pageNumber="558">
<subSubSection id="C3550FF0A679BF6FFF50B474EED19B64" box="[151,987,1587,1614]" pageId="9" pageNumber="558" type="nomenclature">
<paragraph id="8BF05C7BA679BF6FFF50B474EED19B64" blockId="9.[151,987,1587,1650]" box="[151,987,1587,1614]" pageId="9" pageNumber="558">
<heading id="D0B8EB17A679BF6FFF50B474EED19B64" bold="true" box="[151,987,1587,1614]" fontSize="11" level="1" pageId="9" pageNumber="558" reason="1">
<emphasis id="B93B8069A679BF6FFF50B474EED19B64" bold="true" box="[151,987,1587,1614]" pageId="9" pageNumber="558">
<taxonomicName id="4C4F27F8A679BF6FFF50B474EE4E9B64" authority="Mondaca, Pizarro-Araya &amp; Alfaro" authorityName="Mondaca &amp; Pizarro-Araya &amp; Alfaro" authorityYear="2019" box="[151,836,1587,1614]" class="Insecta" family="Melolonthidae" genus="Luispenaia" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="558" phylum="Arthropoda" rank="species" species="paulseni" status="sp. nov.">
<emphasis id="B93B8069A679BF6FFF50B474EC879B67" bold="true" box="[151,397,1587,1614]" italics="true" pageId="9" pageNumber="558">Luispenaia paulseni</emphasis>
Mondaca, Pizarro-Araya &amp; Alfaro
</taxonomicName>
<taxonomicNameLabel id="A2083D12A679BF6FFC8CB474EED19B64" box="[843,987,1587,1613]" pageId="9" pageNumber="558" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3550FF0A679BF6FFF59B410EC839B5B" box="[158,393,1623,1650]" pageId="9" pageNumber="558" type="description">
<paragraph id="8BF05C7BA679BF6FFF59B410EC839B5B" blockId="9.[151,987,1587,1650]" box="[158,393,1623,1650]" pageId="9" pageNumber="558">
(
<figureCitation id="137440FEA679BF6FFF61B410EC0E9B58" box="[166,260,1623,1649]" captionStart="FIGURES 10–13" captionStartId="8.[151,264,1969,1994]" captionTargetBox="[175,1412,720,1919]" captionTargetId="figure@8.[151,1436,729,1957]" captionTargetPageId="8" captionText="FIGURES 10–13. Head and pronotum of Luispenaia species. 10, Luispenaia atacamensis; 11, L. paposo; 12, L. paulseni; 13, L. paradoxa. Scale: 0.5 mm." httpUri="https://zenodo.org/record/3246220/files/figure.png" pageId="9" pageNumber="558">Figs. 12</figureCitation>
, 
<figureCitation id="137440FEA679BF6FFED7B410EC249B58" box="[272,302,1623,1649]" captionStart="FIGURES 14–17" captionStartId="9.[151,264,1456,1481]" captionTargetBox="[151,1436,170,1444]" captionTargetId="figure@9.[151,1436,170,1444]" captionTargetPageId="9" captionText="FIGURES 14–17. Antennal club of Luispenaia species. 14, Luispenaia paradoxa Martínez; 15, L. paposo; 16, L. paulseni; 17, L. atacamensis. Scale: 0.5 mm." httpUri="https://zenodo.org/record/3246222/files/figure.png" pageId="9" pageNumber="558">16</figureCitation>
, 
<figureCitation id="137440FEA679BF6FFEFDB410EC8B9B5B" box="[314,385,1623,1650]" captionStart="FIGURES 18–29" captionStartId="10.[151,264,1736,1761]" captionTargetBox="[164,1379,530,1707]" captionTargetId="figure@10.[151,1436,513,1724]" captionTargetPageId="10" captionText="FIGURES 18–29. Luispenaia species. 18–20, Luispenaia atacamensis; 21–23, L. paposo; 24–26, L. paulseni; 27–29, L. paradoxa. Scale: 1 mm." httpUri="https://zenodo.org/record/3246224/files/figure.png" pageId="9" pageNumber="558">24–26</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3550FF0A679BF6CFF50B4D8E9B09C36" lastPageId="10" lastPageNumber="559" pageId="9" pageNumber="558" type="materials_examined">
<paragraph id="8BF05C7BA679BF6FFF50B4D8EED19B90" blockId="9.[151,1437,1694,2046]" box="[151,987,1694,1722]" pageId="9" pageNumber="558">
<materialsCitation id="3B275626A679BF6FFF50B4D8EED29B90" ID-GBIF-Occurrence="2265605696" box="[151,984,1694,1722]" country="Chile" location="Quebrada Los Yales" municipality="Paposo" pageId="9" pageNumber="558" specimenCount="1" stateProvince="Antofagasta" typeStatus="holotype">
<emphasis id="B93B8069A679BF6FFF50B4D8EC3C9B90" bold="true" box="[151,310,1694,1721]" pageId="9" pageNumber="558">
<typeStatus id="54F4E2D9A679BF6FFF50B4D8EDD89B90" box="[151,210,1695,1721]" pageId="9" pageNumber="558">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="F3581CEBA679BF6FFEFBB4D8EC719B90" box="[316,379,1695,1721]" name="Chile" pageId="9" pageNumber="558">Chile</collectingCountry>
, 
<collectingRegion id="498B9299A679BF6FFE40B4D8EF839B90" box="[391,649,1695,1721]" country="Chile" name="Antofagasta" pageId="9" pageNumber="558">Región de Antofagasta</collectingRegion>
, 
<collectingMunicipality id="6B94C601A679BF6FFD53B4E7EFE29B93" box="[660,744,1696,1722]" pageId="9" pageNumber="558">Paposo</collectingMunicipality>
, 
<location id="8E900AA0A679BF6FFD34B4D8EED29B90" LSID="urn:lsid:plazi:treatment:03E6ED6DA679BF6AFF50B474EEEE9C41:8E900AA0A679BF6FFD34B4D8EED29B90" box="[755,984,1695,1722]" country="Chile" municipality="Paposo" name="Quebrada Los Yales" pageId="9" pageNumber="558" stateProvince="Antofagasta">Quebrada Los Yales</location>
</materialsCitation>
.
</paragraph>
<paragraph id="8BF05C7BA679BF6CFF00B484E9B09C36" blockId="9.[151,1437,1694,2046]" lastBlockId="10.[151,1437,152,503]" lastPageId="10" lastPageNumber="559" pageId="9" pageNumber="558">
<emphasis id="B93B8069A679BF6FFF00B484EC7C9BF4" bold="true" box="[199,374,1730,1757]" pageId="9" pageNumber="558">Type material:</emphasis>
a) “ 
<collectingCountry id="F3581CEBA679BF6FFE65B484ECFD9BF4" box="[418,503,1731,1757]" name="Chile" pageId="9" pageNumber="558">CHILE</collectingCountry>
<collectingRegion id="498B9299A679BF6FFE3BB484EFCF9BF4" box="[508,709,1731,1757]" country="Chile" name="Antofagasta" pageId="9" pageNumber="558">ANTOFAGASTA</collectingRegion>
/ Qda. Los Yales, Paposo / 
<geoCoordinate id="EE7B3ABCA679BF6FFC33B485E9989BF4" box="[1012,1170,1730,1757]" degrees="25" direction="south" minutes="00" orientation="latitude" pageId="9" pageNumber="558" precision="1" seconds="27.7" value="-25.007694">25°00’27.7’’S</geoCoordinate>
/ 
<geoCoordinate id="EE7B3ABCA679BF6FFB61B484E8469BF4" box="[1190,1356,1731,1757]" degrees="70" direction="east" minutes="26" orientation="longitude" pageId="9" pageNumber="558" precision="1" seconds="48,3" value="70.446754">70°26’48,3’’O</geoCoordinate>
; 
<quantity id="4CB7F19EA679BF6FFA91B484E8969BF7" box="[1366,1436,1731,1758]" metricMagnitude="2" metricUnit="m" metricValue="6.17" pageId="9" pageNumber="558" unit="m" value="617.0">617 m</quantity>
/ 
<date id="FFF17ABBA679BF6FFF61B4A1EC2E9A28" box="[166,292,1766,1793]" pageId="9" pageNumber="558" value="2015-10-27">27-X-2015</date>
, t. luz blanca / col. J. Pizarro-F. Alfaro”, b) “ 
<taxonomicName id="4C4F27F8A679BF6FFCDAB4AFEE939A28" authorityName="Martinez" authorityYear="1972" box="[797,921,1768,1793]" class="Insecta" family="Melolonthidae" genus="Luispenaia" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="558" phylum="Arthropoda" rank="genus">
<emphasis id="B93B8069A679BF6FFCDAB4AFEE939A28" box="[797,921,1768,1793]" italics="true" pageId="9" pageNumber="558">Luispenaia</emphasis>
</taxonomicName>
/ 
<taxonomicName id="4C4F27F8A679BF6FFC77B4A0E9059A28" authority="Mondaca &amp; Pizarro-Araya &amp; Alfaro, 2019" authorityName="Mondaca &amp; Pizarro-Araya &amp; Alfaro" authorityYear="2019" box="[944,1039,1767,1793]" class="Insecta" family="Melolonthidae" genus="Luispenaia" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="558" phylum="Arthropoda" rank="species" species="paulseni" status="sp. nov.">
<emphasis id="B93B8069A679BF6FFC77B4A0E9059A28" box="[944,1039,1767,1793]" italics="true" pageId="9" pageNumber="558">paulseni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2083D12A679BF6FFBD1B4A0E97A9A28" box="[1046,1136,1767,1793]" pageId="9" pageNumber="558" rank="species">
<emphasis id="B93B8069A679BF6FFBD1B4A0E97A9A28" bold="true" box="[1046,1136,1767,1793]" pageId="9" pageNumber="558">sp. nov.</emphasis>
</taxonomicNameLabel>
/ Mondaca, Pizarro-Araya &amp; Alfaro / 
<typeStatus id="54F4E2D9A679BF6FFEC8B54CECAC9A0C" box="[271,422,1803,1829]" pageId="9" pageNumber="558" type="holotype">HOLOTYPE</typeStatus>
” (red label, typeset). One male 
<typeStatus id="54F4E2D9A679BF6FFCC3B54BEE6E9A0F" box="[772,868,1804,1830]" pageId="9" pageNumber="558" type="paratype">paratype</typeStatus>
at 
<collectionCode id="ED5EC4BEA679BF6FFC42B54CEEC79A0C" box="[901,973,1803,1829]" pageId="9" pageNumber="558">JMEC</collectionCode>
and one male 
<typeStatus id="54F4E2D9A679BF6FFBAAB54BE9C79A0F" box="[1133,1229,1804,1830]" pageId="9" pageNumber="558" type="paratype">paratype</typeStatus>
at 
<collectionCode id="ED5EC4BEA679BF6FFB29B54CE84F9A0C" box="[1262,1349,1803,1829]" pageId="9" pageNumber="558">LEULS</collectionCode>
labeled as 
<typeStatus id="54F4E2D9A679BF6FFF71B568EC129A60" box="[182,280,1839,1865]" pageId="9" pageNumber="558" type="holotype">holotype</typeStatus>
except: b) “ 
<taxonomicName id="4C4F27F8A679BF6FFE63B577EF2A9A60" authorityName="Martinez" authorityYear="1972" box="[420,544,1840,1865]" class="Insecta" family="Melolonthidae" genus="Luispenaia" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="558" phylum="Arthropoda" rank="genus">
<emphasis id="B93B8069A679BF6FFE63B577EF2A9A60" box="[420,544,1840,1865]" italics="true" pageId="9" pageNumber="558">Luispenaia</emphasis>
</taxonomicName>
/ 
<taxonomicName id="4C4F27F8A679BF6FFDF0B568EF9C9A60" authority="Mondaca &amp; Pizarro-Araya &amp; Alfaro, 2019" authorityName="Mondaca &amp; Pizarro-Araya &amp; Alfaro" authorityYear="2019" box="[567,662,1839,1865]" class="Insecta" family="Melolonthidae" genus="Luispenaia" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="558" phylum="Arthropoda" rank="species" species="paulseni" status="sp. nov.">
<emphasis id="B93B8069A679BF6FFDF0B568EF9C9A60" box="[567,662,1839,1865]" italics="true" pageId="9" pageNumber="558">paulseni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2083D12A679BF6FFD59B568EFF29A60" box="[670,760,1839,1865]" pageId="9" pageNumber="558" rank="species">
<emphasis id="B93B8069A679BF6FFD59B568EFF29A60" bold="true" box="[670,760,1839,1865]" pageId="9" pageNumber="558">sp. nov.</emphasis>
</taxonomicNameLabel>
/ Mondaca, Pizarro-Araya &amp; Alfaro / 
<typeStatus id="54F4E2D9A679BF6FFB6FB568E83D9A60" box="[1192,1335,1839,1865]" pageId="9" pageNumber="558" type="paratype">PARATYPE</typeStatus>
” (yellow label, typeset)”. One male 
<typeStatus id="54F4E2D9A679BF6FFE01B513EF2C9A47" box="[454,550,1876,1902]" pageId="9" pageNumber="558" type="paratype">paratype</typeStatus>
at 
<collectionCode id="ED5EC4BEA679BF6FFD8CB514EF869A44" box="[587,652,1875,1901]" pageId="9" pageNumber="558">JEBC</collectionCode>
labeled: a) “15-018 / 
<collectingCountry id="F3581CEBA679BF6FFC40B514EED69A44" box="[903,988,1875,1901]" name="Chile" pageId="9" pageNumber="558">CHILE</collectingCountry>
, prov. 
<collectingRegion id="498B9299A679BF6FFBEFB514E9B89A44" box="[1064,1202,1875,1901]" country="Chile" name="Antofagasta" pageId="9" pageNumber="558">Antofagasta</collectingRegion>
, / Paposo, quebrada, 
<date id="FFF17ABBA679BF6FFF50B530EC099ABB" box="[151,259,1911,1938]" pageId="9" pageNumber="558" value="1927-10-30">27/30 oct</date>
/ 2015, 770 msnm, luz 
<collectionCode id="ED5EC4BEA679BF6FFDDAB53FEF4F9ABB" box="[541,581,1912,1938]" country="Colombia" name="Departamento de Biologia de la Universidad del Valle" pageId="9" pageNumber="558">UV</collectionCode>
/ 25º00’16.7’’2S 
<geoCoordinate id="EE7B3ABCA679BF6FFCD4B530EECD9AB8" box="[787,967,1911,1937]" degrees="70" direction="west" minutes="26" orientation="longitude" pageId="9" pageNumber="558" precision="1" seconds="36,38" value="-70.44344">70º26’36,38’’W</geoCoordinate>
/ leg. J.E. Barriga-Tuñon J. / Pizarro, F. Alfaro, R. Botero, R. Muñoz”, b) “ 
<taxonomicName id="4C4F27F8A679BF6FFDD1B5DBEF989A9C" authorityName="Martinez" authorityYear="1972" box="[534,658,1948,1973]" class="Insecta" family="Melolonthidae" genus="Luispenaia" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="558" phylum="Arthropoda" rank="genus">
<emphasis id="B93B8069A679BF6FFDD1B5DBEF989A9C" box="[534,658,1948,1973]" italics="true" pageId="9" pageNumber="558">Luispenaia</emphasis>
</taxonomicName>
/ 
<taxonomicName id="4C4F27F8A679BF6FFD61B5DCEE0F9A9C" authority="Mondaca &amp; Pizarro-Araya &amp; Alfaro, 2019" authorityName="Mondaca &amp; Pizarro-Araya &amp; Alfaro" authorityYear="2019" box="[678,773,1947,1973]" class="Insecta" family="Melolonthidae" genus="Luispenaia" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="558" phylum="Arthropoda" rank="species" species="paulseni" status="sp. nov.">
<emphasis id="B93B8069A679BF6FFD61B5DCEE0F9A9C" box="[678,773,1947,1973]" italics="true" pageId="9" pageNumber="558">paulseni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2083D12A679BF6FFCCDB5DCEE699A9C" box="[778,867,1947,1973]" pageId="9" pageNumber="558" rank="species">
<emphasis id="B93B8069A679BF6FFCCDB5DCEE699A9C" bold="true" box="[778,867,1947,1973]" pageId="9" pageNumber="558">sp. nov.</emphasis>
</taxonomicNameLabel>
/ Mondaca, Pizarro-Araya &amp; Alfaro / 
<typeStatus id="54F4E2D9A679BF6FFAC2B5DCE89E9A9C" box="[1285,1428,1947,1973]" pageId="9" pageNumber="558" type="paratype">PARATYPE</typeStatus>
” (yellow label, typeset)”. One male 
<typeStatus id="54F4E2D9A679BF6FFDDFB587EF729AF3" box="[536,632,1984,2010]" pageId="9" pageNumber="558" type="paratype">paratype</typeStatus>
at 
<collectionCode id="ED5EC4BEA679BF6FFD5FB5F8EFEA9AF0" box="[664,736,1983,2009]" pageId="9" pageNumber="558">JMEC</collectionCode>
labeled: a) “ 
<collectingCountry id="F3581CEBA679BF6FFCAEB5F8EEB49AF0" box="[873,958,1983,2009]" name="Chile" pageId="9" pageNumber="558">CHILE</collectingCountry>
<collectingRegion id="498B9299A679BF6FFC05B5F8E9819AF0" box="[962,1163,1983,2009]" country="Chile" name="Antofagasta" pageId="9" pageNumber="558">ANTOFAGASTA</collectingRegion>
/ Paposo / Oct/ 
<date id="FFF17ABBA679BF6FFAF4B5F8E8969AF0" box="[1331,1436,1983,2009]" pageId="9" pageNumber="558" value="2002-12">Dic-2002</date>
/ col. M. Ferrú”, b) “ 
<taxonomicName id="4C4F27F8A679BF6FFE46B5A3ECF79AD4" authorityName="Martinez" authorityYear="1972" box="[385,509,2020,2045]" class="Insecta" family="Melolonthidae" genus="Luispenaia" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="558" phylum="Arthropoda" rank="genus">
<emphasis id="B93B8069A679BF6FFE46B5A3ECF79AD4" box="[385,509,2020,2045]" italics="true" pageId="9" pageNumber="558">Luispenaia</emphasis>
</taxonomicName>
/ 
<taxonomicName id="4C4F27F8A679BF6FFDD3B5A4EF799AD4" authority="Mondaca &amp; Pizarro-Araya &amp; Alfaro, 2019" authorityName="Mondaca &amp; Pizarro-Araya &amp; Alfaro" authorityYear="2019" box="[532,627,2019,2045]" class="Insecta" family="Melolonthidae" genus="Luispenaia" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="558" phylum="Arthropoda" rank="species" species="paulseni" status="sp. nov.">
<emphasis id="B93B8069A679BF6FFDD3B5A4EF799AD4" box="[532,627,2019,2045]" italics="true" pageId="9" pageNumber="558">paulseni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2083D12A679BF6FFDBDB5A4EFDE9AD4" box="[634,724,2019,2045]" pageId="9" pageNumber="558" rank="species">
<emphasis id="B93B8069A679BF6FFDBDB5A4EFDE9AD4" bold="true" box="[634,724,2019,2045]" pageId="9" pageNumber="558">sp. nov.</emphasis>
</taxonomicNameLabel>
/ Mondaca, Pizarro-Araya &amp; Alfaro / 
<typeStatus id="54F4E2D9A679BF6FFB44B5A4E8189AD4" box="[1155,1298,2019,2045]" pageId="9" pageNumber="558" type="paratype">PARATYPE</typeStatus>
” (yellow la- bel, typeset)”. One male 
<typeStatus id="54F4E2D9A67ABF6CFE69B2DEEF049D9A" box="[430,526,153,179]" pageId="10" pageNumber="559" type="paratype">paratype</typeStatus>
at 
<collectionCode id="ED5EC4BEA67ABF6CFDF6B2DEEF739D9A" box="[561,633,153,179]" pageId="10" pageNumber="559">JMEC</collectionCode>
labeled: a) “ 
<collectingCountry id="F3581CEBA67ABF6CFCC0B2DEEE569D9A" box="[775,860,153,179]" name="Chile" pageId="10" pageNumber="559">CHILE</collectingCountry>
<collectingRegion id="498B9299A67ABF6CFCA5B2DEEEE39D9A" box="[866,1001,153,179]" country="Chile" name="Atacama" pageId="10" pageNumber="559">ATACAMA</collectingRegion>
/ 
<quantity id="4CB7F19EA67ABF6CFC39B2DEE9329D9A" box="[1022,1080,153,179]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="10" pageNumber="559" unit="km" value="1.0">1 km</quantity>
N. de Bahia Salada / 
<date id="FFF17ABBA67ABF6CFAEBB2DFE8969D9A" box="[1324,1436,152,179]" pageId="10" pageNumber="559" value="1991-01-25">25-I-1991</date>
/ leg. M. Guerrero”, b) “ 
<taxonomicName id="4C4F27F8A67ABF6CFE6EB2F9EF2F9DFE" authorityName="Martinez" authorityYear="1972" box="[425,549,190,215]" class="Insecta" family="Melolonthidae" genus="Luispenaia" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="559" phylum="Arthropoda" rank="genus">
<emphasis id="B93B8069A67ABF6CFE6EB2F9EF2F9DFE" box="[425,549,190,215]" italics="true" pageId="10" pageNumber="559">Luispenaia</emphasis>
</taxonomicName>
/ 
<taxonomicName id="4C4F27F8A67ABF6CFDFCB2FAEF909DFE" authority="Mondaca &amp; Pizarro-Araya &amp; Alfaro, 2019" authorityName="Mondaca &amp; Pizarro-Araya &amp; Alfaro" authorityYear="2019" box="[571,666,189,215]" class="Insecta" family="Melolonthidae" genus="Luispenaia" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="559" phylum="Arthropoda" rank="species" species="paulseni" status="sp. nov.">
<emphasis id="B93B8069A67ABF6CFDFCB2FAEF909DFE" box="[571,666,189,215]" italics="true" pageId="10" pageNumber="559">paulseni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2083D12A67ABF6CFD66B2FAEFF19DFE" box="[673,763,189,215]" pageId="10" pageNumber="559" rank="species">
<emphasis id="B93B8069A67ABF6CFD66B2FAEFF19DFE" bold="true" box="[673,763,189,215]" pageId="10" pageNumber="559">sp. nov.</emphasis>
</taxonomicNameLabel>
/ Mondaca, Pizarro-Araya &amp; Alfaro”. One male 
<typeStatus id="54F4E2D9A67ABF6CFAE7B2FAE88A9DFE" box="[1312,1408,189,215]" pageId="10" pageNumber="559" type="paratype">paratype</typeStatus>
at 
<collectionCode id="ED5EC4BEA67ABF6CFF50B2A6EDE49DD2" box="[151,238,225,251]" pageId="10" pageNumber="559">LEULS</collectionCode>
labeled: a) “ 
<collectingCountry id="F3581CEBA67ABF6CFE46B2A6ECDC9DD2" box="[385,470,225,251]" name="Chile" pageId="10" pageNumber="559">CHILE</collectingCountry>
<collectingRegion id="498B9299A67ABF6CFE18B2A6EF7D9DD2" box="[479,631,225,251]" country="Chile" name="Coquimbo" pageId="10" pageNumber="559">COQUIMBO</collectingRegion>
/ Ovalle, Lagunillas / 
<date id="FFF17ABBA67ABF6CFCBCB2A6EEFC9DD2" box="[891,1014,225,251]" pageId="10" pageNumber="559" value="1989-10-27">27-X-1989</date>
/ col. E. Vasquez”, b). “ 
<taxonomicName id="4C4F27F8A67ABF6CFAC8B2A5E8819DD2" authorityName="Martinez" authorityYear="1972" box="[1295,1419,226,251]" class="Insecta" family="Melolonthidae" genus="Luispenaia" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="559" phylum="Arthropoda" rank="genus">
<emphasis id="B93B8069A67ABF6CFAC8B2A5E8819DD2" box="[1295,1419,226,251]" italics="true" pageId="10" pageNumber="559">Luispenaia</emphasis>
</taxonomicName>
/ 
<taxonomicName id="4C4F27F8A67ABF6CFF50B342EDFC9C36" authority="Mondaca &amp; Pizarro-Araya &amp; Alfaro, 2019" authorityName="Mondaca &amp; Pizarro-Araya &amp; Alfaro" authorityYear="2019" box="[151,246,261,287]" class="Insecta" family="Melolonthidae" genus="Luispenaia" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="559" phylum="Arthropoda" rank="species" species="paulseni" status="sp. nov.">
<emphasis id="B93B8069A67ABF6CFF50B342EDFC9C36" box="[151,246,261,287]" italics="true" pageId="10" pageNumber="559">paulseni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2083D12A67ABF6CFF3AB342EC5D9C36" box="[253,343,261,287]" pageId="10" pageNumber="559" rank="species">
<emphasis id="B93B8069A67ABF6CFF3AB342EC5D9C36" bold="true" box="[253,343,261,287]" pageId="10" pageNumber="559">sp. nov.</emphasis>
</taxonomicNameLabel>
/ Mondaca, Pizarro-Araya and Alfaro / 
<typeStatus id="54F4E2D9A67ABF6CFCD2B342EEAE9C36" box="[789,932,261,287]" pageId="10" pageNumber="559" type="paratype">PARATYPE</typeStatus>
” (yellow label, typeset)”
</paragraph>
</subSubSection>
<subSubSection id="C3550FF0A67ABF6CFF00B36FEFC69CDE" pageId="10" pageNumber="559" type="diagnosis">
<paragraph id="8BF05C7BA67ABF6CFF00B36FEFC69CDE" blockId="10.[151,1437,152,503]" pageId="10" pageNumber="559">
<emphasis id="B93B8069A67ABF6CFF00B36FEC489C6A" bold="true" box="[199,322,296,323]" pageId="10" pageNumber="559">Diagnosis.</emphasis>
This species is distinguished from all other species in the genus 
<taxonomicName id="4C4F27F8A67ABF6CFBDAB36DE9939C6A" authorityName="Martinez" authorityYear="1972" box="[1053,1177,298,323]" class="Insecta" family="Melolonthidae" genus="Luispenaia" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="559" phylum="Arthropoda" rank="genus">
<emphasis id="B93B8069A67ABF6CFBDAB36DE9939C6A" box="[1053,1177,298,323]" italics="true" pageId="10" pageNumber="559">Luispenaia</emphasis>
</taxonomicName>
by the following combination of characters: Body flattened, dorsally dark brown or black on head, pronotum and scutellum, legs, elytra light brown, shiny (
<figureCitation id="137440FEA67ABF6CFEB9B336ECDC9CA2" box="[382,470,369,395]" captionStart="FIGURES 18–29" captionStartId="10.[151,264,1736,1761]" captionTargetBox="[164,1379,530,1707]" captionTargetId="figure@10.[151,1436,513,1724]" captionTargetPageId="10" captionText="FIGURES 18–29. Luispenaia species. 18–20, Luispenaia atacamensis; 21–23, L. paposo; 24–26, L. paulseni; 27–29, L. paradoxa. Scale: 1 mm." httpUri="https://zenodo.org/record/3246224/files/figure.png" pageId="10" pageNumber="559">Fig. 24</figureCitation>
). Clypeus distally straight and upturned, with lateral margins rounded and reflexed upturned; lateral margin between clypeal base and ocular canthus very projected as external rounded lobule (
<figureCitation id="137440FEA67ABF6CFAA8B3D2EDBC9CFA" captionStart="FIGURES 10–13" captionStartId="8.[151,264,1969,1994]" captionTargetBox="[175,1412,720,1919]" captionTargetId="figure@8.[151,1436,729,1957]" captionTargetPageId="8" captionText="FIGURES 10–13. Head and pronotum of Luispenaia species. 10, Luispenaia atacamensis; 11, L. paposo; 12, L. paulseni; 13, L. paradoxa. Scale: 0.5 mm." httpUri="https://zenodo.org/record/3246220/files/figure.png" pageId="10" pageNumber="559">Fig. 12</figureCitation>
). Antennae with 8 antennomeres; club with 3 antennomeres (
<figureCitation id="137440FEA67ABF6CFCB7B3FEEECE9CFD" box="[880,964,441,468]" captionStart="FIGURES 14–17" captionStartId="9.[151,264,1456,1481]" captionTargetBox="[151,1436,170,1444]" captionTargetId="figure@9.[151,1436,170,1444]" captionTargetPageId="9" captionText="FIGURES 14–17. Antennal club of Luispenaia species. 14, Luispenaia paradoxa Martínez; 15, L. paposo; 16, L. paulseni; 17, L. atacamensis. Scale: 0.5 mm." httpUri="https://zenodo.org/record/3246222/files/figure.png" pageId="10" pageNumber="559">Fig. 16</figureCitation>
). Pronotum approximately subhexagonal, laterally angled, with scarce dorsal setae (
<figureCitation id="137440FEA67ABF6CFDACB39AEFB49CDE" box="[619,702,477,503]" captionStart="FIGURES 10–13" captionStartId="8.[151,264,1969,1994]" captionTargetBox="[175,1412,720,1919]" captionTargetId="figure@8.[151,1436,729,1957]" captionTargetPageId="8" captionText="FIGURES 10–13. Head and pronotum of Luispenaia species. 10, Luispenaia atacamensis; 11, L. paposo; 12, L. paulseni; 13, L. paradoxa. Scale: 0.5 mm." httpUri="https://zenodo.org/record/3246220/files/figure.png" pageId="10" pageNumber="559">Fig. 12</figureCitation>
).
</paragraph>
</subSubSection>
<caption id="DF300CF3A67ABF6CFF50B48FEC519A29" httpUri="https://zenodo.org/record/3246224/files/figure.png" pageId="10" pageNumber="559" startId="10.[151,264,1736,1761]" targetBox="[164,1379,530,1707]" targetPageId="10">
<paragraph id="8BF05C7BA67ABF6CFF50B48FEC519A29" blockId="10.[151,1436,1736,1793]" pageId="10" pageNumber="559">
<emphasis id="B93B8069A67ABF6CFF50B48FEC5E9BC8" bold="true" box="[151,340,1736,1761]" pageId="10" pageNumber="559">FIGURES 18–29.</emphasis>
<taxonomicName id="4C4F27F8A67ABF6CFE9DB48EECC19BC9" authorityName="Martinez" authorityYear="1972" box="[346,459,1737,1760]" class="Insecta" family="Melolonthidae" genus="Luispenaia" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="559" phylum="Arthropoda" rank="genus">
<emphasis id="B93B8069A67ABF6CFE9DB48EECC19BC9" box="[346,459,1737,1760]" italics="true" pageId="10" pageNumber="559">Luispenaia</emphasis>
</taxonomicName>
species. 
<emphasis id="B93B8069A67ABF6CFDE0B48FEF6D9BC8" bold="true" box="[551,615,1736,1761]" pageId="10" pageNumber="559">18–20</emphasis>
, 
<taxonomicName id="4C4F27F8A67ABF6CFDB4B48EEE6D9BC9" authorityName="Mondaca &amp; Pizarro-Araya &amp; Alfaro" authorityYear="2019" box="[627,871,1737,1760]" class="Insecta" family="Melolonthidae" genus="Luispenaia" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="559" phylum="Arthropoda" rank="species" species="atacamensis">
<emphasis id="B93B8069A67ABF6CFDB4B48EEE6D9BC9" box="[627,871,1737,1760]" italics="true" pageId="10" pageNumber="559">Luispenaia atacamensis</emphasis>
</taxonomicName>
; 
<emphasis id="B93B8069A67ABF6CFCB2B48FEEBE9BC8" bold="true" box="[885,948,1736,1761]" pageId="10" pageNumber="559">21–23</emphasis>
, 
<taxonomicName id="4C4F27F8A67ABF6CFC06B48EE92F9BC9" authorityName="Mondaca &amp; Pizarro-Araya &amp; Alfaro" authorityYear="2019" box="[961,1061,1737,1760]" class="Insecta" family="Melolonthidae" genus="Luispenaia" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="559" phylum="Arthropoda" rank="species" species="paposo">
<emphasis id="B93B8069A67ABF6CFC06B48EE92F9BC9" box="[961,1061,1737,1760]" italics="true" pageId="10" pageNumber="559">L. paposo</emphasis>
</taxonomicName>
; 
<emphasis id="B93B8069A67ABF6CFBF4B48FE9789BC8" bold="true" box="[1075,1138,1736,1761]" pageId="10" pageNumber="559">24–26</emphasis>
, 
<taxonomicName id="4C4F27F8A67ABF6CFBB8B48EE9FA9BC8" authorityName="Mondaca &amp; Pizarro-Araya &amp; Alfaro" authorityYear="2019" box="[1151,1264,1737,1761]" class="Insecta" family="Melolonthidae" genus="Luispenaia" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="559" phylum="Arthropoda" rank="species" species="paulseni">
<emphasis id="B93B8069A67ABF6CFBB8B48EE9FA9BC8" box="[1151,1264,1737,1761]" italics="true" pageId="10" pageNumber="559">L. paulseni</emphasis>
</taxonomicName>
; 
<emphasis id="B93B8069A67ABF6CFB3AB48FE8369BC8" bold="true" box="[1277,1340,1736,1761]" pageId="10" pageNumber="559">27–29</emphasis>
, 
<taxonomicName id="4C4F27F8A67ABF6CFA8EB48EEDC29A28" authorityName="Martinez, and Ptiophys" authorityYear="1972" class="Insecta" family="Melolonthidae" genus="Luispenaia" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="559" phylum="Arthropoda" rank="species" species="paradoxa">
<emphasis id="B93B8069A67ABF6CFA8EB48EEDC29A28" italics="true" pageId="10" pageNumber="559">L. paradoxa</emphasis>
</taxonomicName>
. Scale: 1 mm.
</paragraph>
</caption>
<subSubSection id="C3550FF0A67ABF6DFF00B56DEF549F16" lastPageId="11" lastPageNumber="560" pageId="10" pageNumber="559" type="description">
<paragraph id="8BF05C7BA67ABF6DFF00B56DEF169CFA" blockId="10.[151,1437,1834,2042]" lastBlockId="11.[151,1437,153,791]" lastPageId="11" lastPageNumber="560" pageId="10" pageNumber="559">
<emphasis id="B93B8069A67ABF6CFF00B56DEF229A6C" bold="true" box="[199,552,1834,1861]" pageId="10" pageNumber="559">
Description of male 
<typeStatus id="54F4E2D9A67ABF6CFE7DB56CEF299A6C" box="[442,547,1835,1861]" pageId="10" pageNumber="559" type="holotype">holotype</typeStatus>
.
</emphasis>
Length 
<quantity id="4CB7F19EA67ABF6CFD40B56DEFEA9A6F" box="[647,736,1834,1862]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" pageId="10" pageNumber="559" unit="mm" value="6.5">6.5 mm</quantity>
, wide 
<quantity id="4CB7F19EA67ABF6CFCECB56CEE8F9A6F" box="[811,901,1835,1862]" metricMagnitude="-3" metricUnit="m" metricValue="2.7" pageId="10" pageNumber="559" unit="mm" value="2.7">2.7 mm</quantity>
. 
<emphasis id="B93B8069A67ABF6CFC56B56CEEC19A6C" box="[913,971,1835,1861]" italics="true" pageId="10" pageNumber="559">Body</emphasis>
: Flattened, dorsally dark brown on head, pronotum and scutellum, legs and elytra light brown, shiny (
<figureCitation id="137440FEA67ABF6CFCFEB508EE849A40" box="[825,910,1871,1897]" captionStart="FIGURES 18–29" captionStartId="10.[151,264,1736,1761]" captionTargetBox="[164,1379,530,1707]" captionTargetId="figure@10.[151,1436,513,1724]" captionTargetPageId="10" captionText="FIGURES 18–29. Luispenaia species. 18–20, Luispenaia atacamensis; 21–23, L. paposo; 24–26, L. paulseni; 27–29, L. paradoxa. Scale: 1 mm." httpUri="https://zenodo.org/record/3246224/files/figure.png" pageId="10" pageNumber="559">Fig. 24</figureCitation>
). 
<emphasis id="B93B8069A67ABF6CFC66B508EED49A40" box="[929,990,1871,1897]" italics="true" pageId="10" pageNumber="559">Head</emphasis>
: Clypeus distally straight and upturned, with lateral margins rounded and reflexed upturned, laterally with a prolongation of the margins as external rounded lobules; dorsal surface punctate and setose; punctures large; clypeofrontal suture well defined, frontal width equal to 4.5 times the width of the eye, with surface densely punctate, punctures large; eye canthus short and narrow, slightly rounded externally (
<figureCitation id="137440FEA67ABF6CFE46B598ECD29AD0" box="[385,472,2015,2041]" captionStart="FIGURES 10–13" captionStartId="8.[151,264,1969,1994]" captionTargetBox="[175,1412,720,1919]" captionTargetId="figure@8.[151,1436,729,1957]" captionTargetPageId="8" captionText="FIGURES 10–13. Head and pronotum of Luispenaia species. 10, Luispenaia atacamensis; 11, L. paposo; 12, L. paulseni; 13, L. paradoxa. Scale: 0.5 mm." httpUri="https://zenodo.org/record/3246220/files/figure.png" pageId="10" pageNumber="559">Fig. 12</figureCitation>
). Antennae with 8 antennomeres; antennomere 1 robust, claviform; antennomere 2 submonoliform; antennomere 3 cylindrical; antennomere 4 subcylindrical, wide and flattened at apex; antennomere 5 discoidal; antennal club slightly longer than antennomeres 2–4 combined, with 3 antennomeres glabrous and micropunctate, subequal in length, straight in dorsal view (
<figureCitation id="137440FEA67BBF6DFCC5B2A6EE5F9DD5" box="[770,853,225,252]" captionStart="FIGURES 14–17" captionStartId="9.[151,264,1456,1481]" captionTargetBox="[151,1436,170,1444]" captionTargetId="figure@9.[151,1436,170,1444]" captionTargetPageId="9" captionText="FIGURES 14–17. Antennal club of Luispenaia species. 14, Luispenaia paradoxa Martínez; 15, L. paposo; 16, L. paulseni; 17, L. atacamensis. Scale: 0.5 mm." httpUri="https://zenodo.org/record/3246222/files/figure.png" pageId="11" pageNumber="560">Fig. 16</figureCitation>
). 
<emphasis id="B93B8069A67BBF6DFCAFB2A5EEDD9DD2" box="[872,983,226,251]" italics="true" pageId="11" pageNumber="560">Pronotum</emphasis>
: Approximately subpentagonal, laterally angled; surface scarcely punctate and setose, lateral margin with long and moderately dense setae (
<figureCitation id="137440FEA67BBF6DFB20B342E8309C36" box="[1255,1338,261,287]" captionStart="FIGURES 10–13" captionStartId="8.[151,264,1969,1994]" captionTargetBox="[175,1412,720,1919]" captionTargetId="figure@8.[151,1436,729,1957]" captionTargetPageId="8" captionText="FIGURES 10–13. Head and pronotum of Luispenaia species. 10, Luispenaia atacamensis; 11, L. paposo; 12, L. paulseni; 13, L. paradoxa. Scale: 0.5 mm." httpUri="https://zenodo.org/record/3246220/files/figure.png" pageId="11" pageNumber="560">Fig. 12</figureCitation>
). 
<emphasis id="B93B8069A67BBF6DFA89B342E89E9C36" box="[1358,1428,261,287]" italics="true" pageId="11" pageNumber="560">Elytra</emphasis>
: Flattened, surface scarcely setose, with 8 striae well defined; setae erect, slightly recumbent posteriorly; lateral margin with long, moderately dense, laterally directed setae. 
<emphasis id="B93B8069A67BBF6DFCF9B309EE799C4E" box="[830,883,334,359]" italics="true" pageId="11" pageNumber="560">Legs</emphasis>
: Moderately setose and shiny; setae yellow, long. Protibiae tridentate, with teeth rounded distally. 
<emphasis id="B93B8069A67BBF6DFD6AB336EE1D9CA2" box="[685,791,369,395]" italics="true" pageId="11" pageNumber="560">Genitalia</emphasis>
: Parameres long and slender, wider at base (frontal view), laterally slender, slightly curved ventrally, widened in the apical third and rounded at apex; phallobase 1.0 times as longer as parameres (
<figureCitation id="137440FEA67BBF6DFE41B3FEEF1A9CFA" box="[390,528,440,467]" captionStart="FIGURES 18–29" captionStartId="10.[151,264,1736,1761]" captionTargetBox="[164,1379,530,1707]" captionTargetId="figure@10.[151,1436,513,1724]" captionTargetPageId="10" captionText="FIGURES 18–29. Luispenaia species. 18–20, Luispenaia atacamensis; 21–23, L. paposo; 24–26, L. paulseni; 27–29, L. paradoxa. Scale: 1 mm." httpUri="https://zenodo.org/record/3246224/files/figure.png" pageId="11" pageNumber="560">Figs. 25–26</figureCitation>
).
</paragraph>
<paragraph id="8BF05C7BA67BBF6DFF00B39AECA89CDE" blockId="11.[151,1437,153,791]" box="[199,418,477,503]" pageId="11" pageNumber="560">
<emphasis id="B93B8069A67BBF6DFF00B39AEC2F9CDE" bold="true" box="[199,293,477,503]" pageId="11" pageNumber="560">Female.</emphasis>
Unknown.
</paragraph>
<paragraph id="8BF05C7BA67BBF6DFF00B047EF549F16" blockId="11.[151,1437,153,791]" pageId="11" pageNumber="560">
<emphasis id="B93B8069A67BBF6DFF00B047EC359F32" bold="true" box="[199,319,512,539]" pageId="11" pageNumber="560">Variation.</emphasis>
Length 
<quantity id="4CB7F19EA67BBF6DFE5DB047EF289F32" box="[410,546,512,539]" metricMagnitude="-3" metricUnit="m" metricValue="6.15" metricValueMax="6.5" metricValueMin="5.8" pageId="11" pageNumber="560" unit="mm" value="6.15" valueMax="6.5" valueMin="5.8">5.8–6.5 mm</quantity>
. Dorsal color of pronotum light brown to dark brown. Not differing significantly from the 
<typeStatus id="54F4E2D9A67BBF6DFF39B062EC6A9F16" box="[254,352,549,575]" pageId="11" pageNumber="560" type="holotype">holotype</typeStatus>
in external characters.
</paragraph>
</subSubSection>
<subSubSection id="C3550FF0A67BBF6DFF00B00EEFD99F82" pageId="11" pageNumber="560" type="etymology">
<paragraph id="8BF05C7BA67BBF6DFF00B00EEFD99F82" blockId="11.[151,1437,153,791]" pageId="11" pageNumber="560">
<emphasis id="B93B8069A67BBF6DFF00B00EEC449F4A" bold="true" box="[199,334,585,611]" pageId="11" pageNumber="560">Etymology.</emphasis>
Named in honor of our colleague and friend M.J. Paulsen from the University of 
<collectingRegion id="498B9299A67BBF6DFB35B00EE8969F4A" box="[1266,1436,585,611]" country="United States of America" name="Nebraska" pageId="11" pageNumber="560">Nebraska State</collectingRegion>
Museum, Lincoln, 
<collectingRegion id="498B9299A67BBF6DFEAAB02AECD09FAE" box="[365,474,621,647]" country="United States of America" name="Nebraska" pageId="11" pageNumber="560">Nebraska</collectingRegion>
, 
<collectingCountry id="F3581CEBA67BBF6DFE22B02AEE0E9FAE" box="[485,772,621,647]" name="United States of America" pageId="11" pageNumber="560">United States of America</collectingCountry>
. He has contributed greatly to the taxonomy of the Lucani- dae and 
<taxonomicName id="4C4F27F8A67BBF6DFF33B0D6EC669F82" box="[244,364,657,683]" class="Insecta" family="Glaresidae" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="560" phylum="Arthropoda" rank="family">Glaresidae</taxonomicName>
from 
<collectingCountry id="F3581CEBA67BBF6DFE76B0D6ECE49F82" box="[433,494,657,683]" name="Chile" pageId="11" pageNumber="560">Chile</collectingCountry>
and South America.
</paragraph>
</subSubSection>
<subSubSection id="C3550FF0A67BBF6DFF00B0F3EC6C9FDA" pageId="11" pageNumber="560" type="distribution">
<paragraph id="8BF05C7BA67BBF6DFF00B0F3EC6C9FDA" blockId="11.[151,1437,153,791]" pageId="11" pageNumber="560">
<emphasis id="B93B8069A67BBF6DFF00B0F3EC6B9FE6" bold="true" box="[199,353,692,719]" pageId="11" pageNumber="560">Distribution.</emphasis>
<collectingCountry id="F3581CEBA67BBF6DFEAFB0F2ECAD9FE6" box="[360,423,693,719]" name="Chile" pageId="11" pageNumber="560">Chile</collectingCountry>
, 
<collectingRegion id="498B9299A67BBF6DFE75B0F2EF339FE6" box="[434,569,693,719]" country="Chile" name="Antofagasta" pageId="11" pageNumber="560">Antofagasta</collectingRegion>
(
<collectingRegion id="498B9299A67BBF6DFD8FB0F2EE4B9FF9" box="[584,833,693,720]" country="Chile" name="Antofagasta" pageId="11" pageNumber="560">Antofagasta Province</collectingRegion>
), 
<collectingRegion id="498B9299A67BBF6DFC95B0F2EEBC9FE6" box="[850,950,693,719]" country="Chile" name="Atacama" pageId="11" pageNumber="560">Atacama</collectingRegion>
(Copiapo Province), and 
<collectingRegion id="498B9299A67BBF6DFB1FB0F2E8449FE6" box="[1240,1358,693,719]" country="Chile" name="Coquimbo" pageId="11" pageNumber="560">Coquimbo</collectingRegion>
(Elqui Province) (
<figureCitation id="137440FEA67BBF6DFED4B09EEC529FDA" box="[275,344,729,755]" captionStart="FIGURE 8" captionStartId="6.[151,250,1996,2021]" captionTargetBox="[192,1388,211,1949]" captionTargetId="figure@6.[151,1435,170,1984]" captionTargetPageId="6" captionText="FIGURE 8. Distribution of Luispenaia species in the Atacama Desert, northern Chile." httpUri="https://zenodo.org/record/3246215/files/figure.png" pageId="11" pageNumber="560">Fig. 8</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3550FF0A67BBF6AFF00B0BBEEEE9C41" lastPageId="12" lastPageNumber="561" pageId="11" pageNumber="560" type="materials_examined">
<paragraph id="8BF05C7BA67BBF6DFF00B0BBE9309E3E" blockId="11.[151,1437,153,791]" box="[199,1082,764,791]" pageId="11" pageNumber="560">
<emphasis id="B93B8069A67BBF6DFF00B0BBECBB9E3E" bold="true" box="[199,433,764,791]" pageId="11" pageNumber="560">Material examined.</emphasis>
<materialsCitation id="3B275626A67BBF6DFE70B0BAE93D9E3E" ID-GBIF-Occurrence="2265605699" box="[439,1079,765,791]" collectionCode="JEBC, JMEC, LEULS" pageId="11" pageNumber="560" specimenCount="6">
<specimenCount id="9D4997F2A67BBF6DFE70B0BAEF4B9E3E" box="[439,577,765,791]" pageId="11" pageNumber="560" type="generic">6 specimens</specimenCount>
were examined from 
<collectionCode id="ED5EC4BEA67BBF6DFCF0B0BAEE719E3E" box="[823,891,765,791]" pageId="11" pageNumber="560">JEBC</collectionCode>
, 
<collectionCode id="ED5EC4BEA67BBF6DFC41B0BAEEDA9E3E" box="[902,976,765,791]" pageId="11" pageNumber="560">JMEC</collectionCode>
, 
<collectionCode id="ED5EC4BEA67BBF6DFC1BB0BAE93D9E3E" box="[988,1079,765,791]" pageId="11" pageNumber="560">LEULS</collectionCode>
</materialsCitation>
.
</paragraph>
<caption id="DF300CF3A67BBF6DFF50B4CCE8669BCA" httpUri="https://zenodo.org/record/3246226/files/figure.png" pageId="11" pageNumber="560" startId="11.[151,264,1675,1700]" targetBox="[151,1435,801,1663]" targetPageId="11">
<paragraph id="8BF05C7BA67BBF6DFF50B4CCE8669BCA" blockId="11.[151,1437,1675,1764]" pageId="11" pageNumber="560">
<emphasis id="B93B8069A67BBF6DFF50B4CCEC259B8D" bold="true" box="[151,303,1675,1700]" pageId="11" pageNumber="560">FIGURES 30.</emphasis>
Type locality of 
<taxonomicName id="4C4F27F8A67BBF6DFE19B4CBEF459B8A" authorityName="Martinez" authorityYear="1972" box="[478,591,1676,1699]" class="Insecta" family="Melolonthidae" genus="Luispenaia" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="560" phylum="Arthropoda" rank="genus">
<emphasis id="B93B8069A67BBF6DFE19B4CBEF459B8A" box="[478,591,1676,1699]" italics="true" pageId="11" pageNumber="560">Luispenaia</emphasis>
</taxonomicName>
species. 
<emphasis id="B93B8069A67BBF6DFD6BB4CCEFB49B8D" bold="true" box="[684,702,1675,1700]" pageId="11" pageNumber="560">A</emphasis>
, Llanos de Challe National Park (Atacama Region, Chile), habitat of 
<taxonomicName id="4C4F27F8A67BBF6DFA4FB4CBEDF29BED" authorityName="Martinez, and Ptiophys" authorityYear="1972" class="Insecta" family="Melolonthidae" genus="Luispenaia" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="560" phylum="Arthropoda" rank="species" species="paradoxa">
<emphasis id="B93B8069A67BBF6DFA4FB4CBEDF29BED" italics="true" pageId="11" pageNumber="560">L. paradoxa</emphasis>
</taxonomicName>
; 
<emphasis id="B93B8069A67BBF6DFEC1B4EBEC1D9BED" bold="true" box="[262,279,1708,1732]" pageId="11" pageNumber="560">B</emphasis>
, Boqueron Chañar (Atacama Region, Chile), habitat of 
<taxonomicName id="4C4F27F8A67BBF6DFC97B4EBEEE29BEA" authorityName="Mondaca &amp; Pizarro-Araya &amp; Alfaro" authorityYear="2019" box="[848,1000,1708,1731]" class="Insecta" family="Melolonthidae" genus="Luispenaia" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="560" phylum="Arthropoda" rank="species" species="atacamensis">
<emphasis id="B93B8069A67BBF6DFC97B4EBEEE29BEA" box="[848,1000,1708,1731]" italics="true" pageId="11" pageNumber="560">L. atacamensis</emphasis>
</taxonomicName>
; 
<emphasis id="B93B8069A67BBF6DFC32B4ECE90D9BED" bold="true" box="[1013,1031,1707,1732]" pageId="11" pageNumber="560">C</emphasis>
, Paposo Norte Natural Monument (Antofagasta Region, Chile), habitat of 
<taxonomicName id="4C4F27F8A67BBF6DFDC6B48BEF6F9BCA" authorityName="Mondaca &amp; Pizarro-Araya &amp; Alfaro" authorityYear="2019" box="[513,613,1740,1763]" class="Insecta" family="Melolonthidae" genus="Luispenaia" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="560" phylum="Arthropoda" rank="species" species="paposo">
<emphasis id="B93B8069A67BBF6DFDC6B48BEF6F9BCA" box="[513,613,1740,1763]" italics="true" pageId="11" pageNumber="560">L. paposo</emphasis>
</taxonomicName>
; 
<emphasis id="B93B8069A67BBF6DFDB4B48BEF8F9BCD" bold="true" box="[627,645,1740,1764]" pageId="11" pageNumber="560">D</emphasis>
, Quebrada Los Yales (Antofagasta Region, Chile), habitat of 
<taxonomicName id="4C4F27F8A67BBF6DFB32B48BE86C9BCD" authorityName="Mondaca &amp; Pizarro-Araya &amp; Alfaro" authorityYear="2019" box="[1269,1382,1740,1764]" class="Insecta" family="Melolonthidae" genus="Luispenaia" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="560" phylum="Arthropoda" rank="species" species="paulseni">
<emphasis id="B93B8069A67BBF6DFB32B48BE86C9BCD" box="[1269,1382,1740,1764]" italics="true" pageId="11" pageNumber="560">L. paulseni</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<paragraph id="8BF05C7BA67BBF6AFF00B54AEEEE9C41" blockId="11.[151,1436,1805,2013]" lastBlockId="12.[151,1437,153,360]" lastPageId="12" lastPageNumber="561" pageId="11" pageNumber="560">
<emphasis id="B93B8069A67BBF6DFF00B54AECCE9A01" bold="true" box="[199,452,1805,1832]" pageId="11" pageNumber="560">Habitat and ecology.</emphasis>
<taxonomicName id="4C4F27F8A67BBF6DFE09B548EFBE9A01" authorityName="Mondaca &amp; Pizarro-Araya &amp; Alfaro" authorityYear="2019" box="[462,692,1806,1832]" class="Insecta" family="Melolonthidae" genus="Luispenaia" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="560" phylum="Arthropoda" rank="species" species="paulseni">
<emphasis id="B93B8069A67BBF6DFE09B548EFBE9A01" box="[462,692,1806,1832]" italics="true" pageId="11" pageNumber="560">Luispenaia paulseni</emphasis>
</taxonomicName>
has been collected during the spring (October–November) with mixed light trap installed in an area with mixed native vegetation, while a one specimen from Paposo were captured with a pitfall trap (
<figureCitation id="137440FEA67BBF6DFEA8B511ECCF9A59" box="[367,453,1878,1904]" captionStart="FIGURES 30" captionStartId="11.[151,264,1675,1700]" captionTargetBox="[151,1435,801,1663]" captionTargetId="figure@11.[151,1435,801,1663]" captionTargetPageId="11" captionText="FIGURES 30. Type locality of Luispenaia species. A, Llanos de Challe National Park (Atacama Region, Chile), habitat of L. paradoxa; B, Boqueron Chañar (Atacama Region, Chile), habitat of L. atacamensis; C, Paposo Norte Natural Monument (Antofagasta Region, Chile), habitat of L. paposo; D, Quebrada Los Yales (Antofagasta Region, Chile), habitat of L. paulseni." httpUri="https://zenodo.org/record/3246226/files/figure.png" pageId="11" pageNumber="560">Fig. 30</figureCitation>
). The 
<typeStatus id="54F4E2D9A67BBF6DFDC9B510EF349A58" box="[526,574,1879,1905]" pageId="11" pageNumber="560">type</typeStatus>
locality corresponds to a habitat belonging to the vegetational floor coastal Mediterranean desert scrub of 
<taxonomicName id="4C4F27F8A67BBF6DFE36B53DEE179ABC" authority="(Phil.)" authorityName="Phil." baseAuthorityName="Phil." box="[497,797,1914,1941]" class="Magnoliopsida" family="Euphorbiaceae" genus="Euphorbia" kingdom="Plantae" order="Malpighiales" pageId="11" pageNumber="560" phylum="Tracheophyta" rank="species" species="lactiflua">
<emphasis id="B93B8069A67BBF6DFE36B53DEFC49ABD" box="[497,718,1914,1940]" italics="true" pageId="11" pageNumber="560">Euphorbia lactiflua</emphasis>
(Phil.)
</taxonomicName>
and 
<taxonomicName id="4C4F27F8A67BBF6DFC91B53DE8929ABC" authority="(Schumann) Britton &amp; Rose" authorityName="Britton &amp; Rose" baseAuthorityName="Schumann" box="[854,1432,1914,1941]" class="Magnoliopsida" family="Cactaceae" genus="Eulychnia" kingdom="Plantae" order="Caryophyllales" pageId="11" pageNumber="560" phylum="Tracheophyta" rank="species" species="iquiquensis">
<emphasis id="B93B8069A67BBF6DFC91B53DE95A9ABD" box="[854,1104,1914,1940]" italics="true" pageId="11" pageNumber="560">Eulychnia iquiquensis</emphasis>
(Schumann) Britton &amp; Rose
</taxonomicName>
, which is characterized by being a dense shrubland with columnar succulents. This habitat is dominated by a diverse set of tall bushes such as 
<taxonomicName id="4C4F27F8A67BBF6DFE00B584EF4D9AF5" box="[455,583,1986,2012]" class="Magnoliopsida" family="Euphorbiaceae" genus="Euphorbia" kingdom="Plantae" order="Malpighiales" pageId="11" pageNumber="560" phylum="Tracheophyta" rank="species" species="lactiflua">
<emphasis id="B93B8069A67BBF6DFE00B584EF4D9AF5" box="[455,583,1986,2012]" italics="true" pageId="11" pageNumber="560">E. lactiflua</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C4F27F8A67BBF6DFD9FB584EE3D9AF5" authority="Phil." authorityName="Phil." box="[600,823,1986,2012]" class="Magnoliopsida" family="Asteraceae" genus="Proustia" kingdom="Plantae" order="Asterales" pageId="11" pageNumber="560" phylum="Tracheophyta" rank="species" species="tipia">
<emphasis id="B93B8069A67BBF6DFD9FB584EFFF9AF5" box="[600,757,1987,2012]" italics="true" pageId="11" pageNumber="560">Proustia tipia</emphasis>
Phil.
</taxonomicName>
, 
<taxonomicName id="4C4F27F8A67BBF6DFC81B584E9259AF5" authority="Phil." authorityName="Phil." box="[838,1071,1986,2012]" class="Magnoliopsida" family="Solanaceae" genus="Lycium" kingdom="Plantae" order="Solanales" pageId="11" pageNumber="560" phylum="Tracheophyta" rank="species" species="deserti">
<emphasis id="B93B8069A67BBF6DFC81B584EEE49AF5" box="[838,1006,1986,2012]" italics="true" pageId="11" pageNumber="560">Lycium deserti</emphasis>
Phil.
</taxonomicName>
, 
<taxonomicName id="4C4F27F8A67BBF6AFBF9B585EC289D9A" authority="(Phil.) I. M. Johnst." authorityName="I. M. Johnst." baseAuthorityName="Phil." class="Magnoliopsida" family="Boraginaceae" genus="Heliotropium" kingdom="Plantae" lastPageId="12" lastPageNumber="561" order="Boraginales" pageId="11" pageNumber="560" phylum="Tracheophyta" rank="species" species="taltalense">
<emphasis id="B93B8069A67BBF6DFBF9B585E8419AF5" box="[1086,1355,1986,2012]" italics="true" pageId="11" pageNumber="560">Heliotropium taltalense</emphasis>
(Phil.) I.M. Johnst.
</taxonomicName>
, 
<taxonomicName id="4C4F27F8A67CBF6AFEE8B2DDECE29D9A" box="[303,488,153,179]" class="Magnoliopsida" family="Boraginaceae" genus="Heliotropium" kingdom="Plantae" order="Boraginales" pageId="12" pageNumber="561" phylum="Tracheophyta" rank="species" species="philippianum">
<emphasis id="B93B8069A67CBF6AFEE8B2DDECE29D9A" box="[303,488,153,179]" italics="true" pageId="12" pageNumber="561">H. philippianum</emphasis>
</taxonomicName>
I.M., Johnst., 
<taxonomicName id="4C4F27F8A67CBF6AFD48B2DEEEB69D9A" authority="Barneoud" authorityName="Barneoud" box="[655,956,153,179]" class="Magnoliopsida" family="Oxalidaceae" genus="Oxalis" kingdom="Plantae" order="Oxalidales" pageId="12" pageNumber="561" phylum="Tracheophyta" rank="species" species="gigantea">
<emphasis id="B93B8069A67CBF6AFD48B2DEEE499D9A" box="[655,835,153,179]" italics="true" pageId="12" pageNumber="561">Oxalis gigantea</emphasis>
Barnéoud
</taxonomicName>
, among which grows the columnar cactus 
<taxonomicName id="4C4F27F8A67CBF6AFF50B2FAEC9A9DFE" box="[151,400,189,215]" class="Magnoliopsida" family="Cactaceae" genus="Eulychnia" kingdom="Plantae" order="Caryophyllales" pageId="12" pageNumber="561" phylum="Tracheophyta" rank="species" species="iquiquensis">
<emphasis id="B93B8069A67CBF6AFF50B2FAEC9A9DFE" box="[151,400,189,215]" italics="true" pageId="12" pageNumber="561">Eulychnia iquiquensis</emphasis>
</taxonomicName>
and the arrosetadas 
<taxonomicName id="4C4F27F8A67CBF6AFDB0B2F9EE7C9DFE" authority="Molina" authorityName="Molina" authorityYear="1782" box="[631,886,189,215]" class="Liliopsida" family="Bromeliaceae" genus="Puya" kingdom="Plantae" order="Poales" pageId="12" pageNumber="561" phylum="Tracheophyta" rank="species" species="chilensis">
<emphasis id="B93B8069A67CBF6AFDB0B2F9EE109DFE" box="[631,794,189,215]" italics="true" pageId="12" pageNumber="561">Puya chilensis</emphasis>
Molina
</taxonomicName>
, 
<taxonomicName id="4C4F27F8A67CBF6AFC46B2FAEC539DD5" authority="(Phil.) Mez (Luebert &amp; Pliscoff 2006)" authorityName="Mez (Luebert &amp; Pliscoff" authorityYear="2006" class="Liliopsida" family="Bromeliaceae" genus="Deuterocohnia" kingdom="Plantae" order="Poales" pageId="12" pageNumber="561" phylum="Tracheophyta" rank="species" species="chrysantha">
<emphasis id="B93B8069A67CBF6AFC46B2FAE9A69DFE" box="[897,1196,189,215]" italics="true" pageId="12" pageNumber="561">Deuterocohnia chrysantha</emphasis>
(Phil.) Mez (
<bibRefCitation id="EFDE218AA67CBF6AFA84B2FAEC5B9DD5" author="Luebert, F. &amp; Pliscoff, P." pageId="12" pageNumber="561" refId="ref7924" refString="Luebert, F. &amp; Pliscoff, P. (2006) Sinopsis Bioclimatica y Vegetacional de Chile. Editorial Universitaria, Santiago, 316 pp." type="book" year="2006">Luebert &amp; Pliscoff 2006</bibRefCitation>
)
</taxonomicName>
. The herbaceous plants are abundant in the rainy years, such as 
<taxonomicName id="4C4F27F8A67CBF6AFBF4B2A6E8669DD2" authority="Phil." authorityName="Phil." box="[1075,1388,225,251]" class="Magnoliopsida" family="Oxalidaceae" genus="Oxalis" kingdom="Plantae" order="Oxalidales" pageId="12" pageNumber="561" phylum="Tracheophyta" rank="species" species="bulbocastanum">
<emphasis id="B93B8069A67CBF6AFBF4B2A6E8259DD2" box="[1075,1327,225,251]" italics="true" pageId="12" pageNumber="561">Oxalis bulbocastanum</emphasis>
Phil.
</taxonomicName>
and 
<taxonomicName id="4C4F27F8A67CBF6AFF50B342ECC59C36" authority="Phil." authorityName="Phil." box="[151,463,261,287]" class="Liliopsida" family="Alstroemeriaceae" genus="Alstroemeria" kingdom="Plantae" order="Liliales" pageId="12" pageNumber="561" phylum="Tracheophyta" rank="species" species="violacea">
<emphasis id="B93B8069A67CBF6AFF50B342EC9A9C36" box="[151,400,261,287]" italics="true" pageId="12" pageNumber="561">Alstroemeria violacea</emphasis>
Phil.
</taxonomicName>
, in addition to the vascular epiphytes 
<taxonomicName id="4C4F27F8A67CBF6AFCB9B342E98A9C36" authority="Phil." authorityName="Phil." box="[894,1152,261,287]" class="Liliopsida" family="Bromeliaceae" genus="Tillandsia" kingdom="Plantae" order="Poales" pageId="12" pageNumber="561" phylum="Tracheophyta" rank="species" species="geissei">
<emphasis id="B93B8069A67CBF6AFCB9B342E9499C36" box="[894,1091,261,287]" italics="true" pageId="12" pageNumber="561">Tillandsia geissei</emphasis>
Phil.
</taxonomicName>
and 
<taxonomicName id="4C4F27F8A67CBF6AFB7FB342EDF19C6A" authority="Sweet. It" authorityName="Sweet. It" authorityYear="1828" class="Magnoliopsida" family="Tropaeolaceae" genus="Tropaeolum" kingdom="Plantae" order="Capparales" pageId="12" pageNumber="561" phylum="Tracheophyta" rank="species" species="tricolor">
<emphasis id="B93B8069A67CBF6AFB7FB342E8979C36" box="[1208,1437,261,287]" italics="true" pageId="12" pageNumber="561">Tropaeolum tricolor</emphasis>
Sweet. It
</taxonomicName>
should be noted that it is a part of the coastal desert that is very rigorous and that is compensated in some sectors by the occurrence of dense camanchacas (
<bibRefCitation id="EFDE218AA67CBF6AFD06B30AEED29C41" author="Luebert, F. &amp; Pliscoff, P." box="[705,984,333,360]" pageId="12" pageNumber="561" refId="ref7924" refString="Luebert, F. &amp; Pliscoff, P. (2006) Sinopsis Bioclimatica y Vegetacional de Chile. Editorial Universitaria, Santiago, 316 pp." type="book" year="2006">Luebert &amp; Pliscoff 2006</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>