treatments-xml/data/03/E6/ED/03E6ED6DA673BF63FF50B407EC7098EA.xml

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<mods:title id="1B31C85AB67C61CE734A8B144C05DEAA">Revision of the genus Luispenaia Martínez (Coleoptera: Scarabaeidae: Melolonthinae: Tanyproctini), with description of three new species from the Atacama Desert, Chile</mods:title>
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<mods:namePart id="320E2E38D9B9B86A04797F03BAAC9405">Mondaca, José</mods:namePart>
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<mods:namePart id="00287C42941AFB762EA92AA5911BD38C">Pizarro-Araya, Jaime</mods:namePart>
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<heading id="D0B8EB17A673BF65FF50B407E9129B70" bold="true" box="[151,1048,1599,1626]" fontSize="11" level="1" pageId="3" pageNumber="552" reason="1">
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<taxonomicName id="4C4F27F8A673BF65FF50B407EFF79B70" authority="Mondaca, Pizarro-Araya" authorityName="Mondaca, Pizarro-Araya" box="[151,765,1599,1626]" class="Insecta" family="Melolonthidae" genus="Luispenaia" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="552" phylum="Arthropoda" rank="species" species="atacamensis">
<emphasis id="B93B8069A673BF65FF50B407ECB09B73" bold="true" box="[151,442,1600,1626]" italics="true" pageId="3" pageNumber="552">Luispenaia atacamensis</emphasis>
Mondaca, Pizarro-Araya
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, &amp; Alfaro, new species
</emphasis>
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</paragraph>
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<paragraph id="8BF05C7BA673BF65FF59B424EC559B57" blockId="3.[151,1048,1599,1662]" box="[158,351,1635,1662]" pageId="3" pageNumber="552">
(
<figureCitation id="137440FEA673BF65FF61B424EC0E9B57" box="[166,260,1635,1662]" captionStart="FIGURES 10–13" captionStartId="8.[151,264,1969,1994]" captionTargetBox="[175,1412,720,1919]" captionTargetId="figure@8.[151,1436,729,1957]" captionTargetPageId="8" captionText="FIGURES 10–13. Head and pronotum of Luispenaia species. 10, Luispenaia atacamensis; 11, L. paposo; 12, L. paulseni; 13, L. paradoxa. Scale: 0.5 mm." httpUri="https://zenodo.org/record/3246220/files/figure.png" pageId="3" pageNumber="552">Figs. 10</figureCitation>
, 
<figureCitation id="137440FEA673BF65FED7B423EC529B57" box="[272,344,1636,1662]" captionStart-0="FIGURES 14–17" captionStart-1="FIGURES 18–29" captionStartId-0="9.[151,264,1456,1481]" captionStartId-1="10.[151,264,1736,1761]" captionTargetBox-0="[151,1436,170,1444]" captionTargetBox-1="[164,1379,530,1707]" captionTargetId-0="figure@9.[151,1436,170,1444]" captionTargetId-1="figure@10.[151,1436,513,1724]" captionTargetPageId-0="9" captionTargetPageId-1="10" captionText-0="FIGURES 14–17. Antennal club of Luispenaia species. 14, Luispenaia paradoxa Martínez; 15, L. paposo; 16, L. paulseni; 17, L. atacamensis. Scale: 0.5 mm." captionText-1="FIGURES 18–29. Luispenaia species. 18–20, Luispenaia atacamensis; 21–23, L. paposo; 24–26, L. paulseni; 27–29, L. paradoxa. Scale: 1 mm." httpUri-0="https://zenodo.org/record/3246222/files/figure.png" httpUri-1="https://zenodo.org/record/3246224/files/figure.png" pageId="3" pageNumber="552">17–20</figureCitation>
)
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<typeStatus id="54F4E2D9A673BF65FF50B4EBEDD89BEF" box="[151,210,1708,1734]" pageId="3" pageNumber="552">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="F3581CEBA673BF65FEFBB4ECEC719BEC" box="[316,379,1707,1733]" name="Chile" pageId="3" pageNumber="552">Chile</collectingCountry>
, 
<collectingRegion id="498B9299A673BF65FE40B4ECEF6F9BEF" box="[391,613,1707,1734]" country="Chile" name="Atacama" pageId="3" pageNumber="552">Región de Atacama</collectingRegion>
, 
<collectorName id="26BA39ADA673BF65FDB7B4ECEE759BEC" box="[624,895,1707,1733]" pageId="3" pageNumber="552">Quebrada El Algarrobal</collectorName>
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.
</paragraph>
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<emphasis id="B93B8069A673BF65FF00B497EC739BC3" bold="true" box="[199,377,1743,1770]" pageId="3" pageNumber="552">Type material.</emphasis>
<materialsCitation id="3B275626A673BF65FE44B488E83F9A1B" ID-GBIF-Occurrence="2412528346" collectingDate="2015-10-31" collectionCode="MNNC" collectorName="J. Pizarro-F. Alfaro" country="Chile" elevation="694" location="El Algarrobal" longitude="70.520584" municipality="Quebrada" pageId="3" pageNumber="552" specimenCount="1" specimenCount-male="1" stateProvince="REGION ATACAMA" typeStatus="holotype">
<typeStatus id="54F4E2D9A673BF65FE44B488ECE19BC0" box="[387,491,1743,1769]" pageId="3" pageNumber="552" type="holotype">Holotype</typeStatus>
<specimenCount id="9D4997F2A673BF65FE31B488EF269BC0" box="[502,556,1743,1769]" pageId="3" pageNumber="552" type="male">male</specimenCount>
at 
<collectionCode id="ED5EC4BEA673BF65FD91B497EFA09BC3" box="[598,682,1744,1770]" country="Chile" httpUri="http://grbio.org/cool/mf2w-ryyx" name="Museo Nacional de Historia Natural, Santiago" pageId="3" pageNumber="552">MNNC</collectionCode>
labeled: a) “ 
<collectingCountry id="F3581CEBA673BF65FC85B497EE9D9BC3" box="[834,919,1744,1770]" name="Chile" pageId="3" pageNumber="552">CHILE</collectingCountry>
<collectingRegion id="498B9299A673BF65FC65B488E9969BC3" box="[930,1180,1743,1770]" country="Chile" name="Atacama" pageId="3" pageNumber="552">REGIÓN ATACAMA</collectingRegion>
/ Qda. [
<collectingMunicipality id="6B94C601A673BF65FB38B488E8789BC3" box="[1279,1394,1743,1770]" pageId="3" pageNumber="552">Quebrada</collectingMunicipality>
] 
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, 28°16” / 
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- 
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, 
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/ 
<collectingDate id="EFB58353A673BF65FD3FB4B4EE7C9A27" box="[760,886,1779,1806]" pageId="3" pageNumber="552" value="2015-10-31">
<date id="FFF17ABBA673BF65FD3FB4B4EE7C9A27" box="[760,886,1779,1806]" pageId="3" pageNumber="552" value="2015-10-31">31-X-2015</date>
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, t. luz blanca / col. 
<collectorName id="26BA39ADA673BF65FB9DB4B3E83C9A24" box="[1114,1334,1779,1806]" pageId="3" pageNumber="552">J. Pizarro-F. Alfaro</collectorName>
” b) “ 
<taxonomicName id="4C4F27F8A673BF65FAB2B4B3EDFC9A18" authorityName="Martinez" authorityYear="1972" class="Insecta" family="Melolonthidae" genus="Luispenaia" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="552" phylum="Arthropoda" rank="genus">
<emphasis id="B93B8069A673BF65FAB2B4B3EDFC9A18" italics="true" pageId="3" pageNumber="552">Luispenaia</emphasis>
</taxonomicName>
/ 
<taxonomicName id="4C4F27F8A673BF65FEC8B55FEC939A18" authority="Mondaca &amp; Pizarro-Araya &amp; Alfaro, 2019" authorityName="Mondaca &amp; Pizarro-Araya &amp; Alfaro" authorityYear="2019" box="[271,409,1816,1841]" class="Insecta" family="Melolonthidae" genus="Luispenaia" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="552" phylum="Arthropoda" rank="species" species="atacamensis" status="sp. nov.">
<emphasis id="B93B8069A673BF65FEC8B55FEC939A18" box="[271,409,1816,1841]" italics="true" pageId="3" pageNumber="552">atacamensis</emphasis>
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<emphasis id="B93B8069A673BF65FE65B55FECF79A1B" bold="true" box="[418,509,1816,1842]" pageId="3" pageNumber="552">sp. nov.</emphasis>
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/ Mondaca, Pizarro-Araya &amp; Alfaro / HOLOTYPE” (red label, typeset)
</materialsCitation>
. 
<materialsCitation id="3B275626A673BF65FA86B550EC339AB7" ID-GBIF-Occurrence="2412528344" collectionCode="CMNC" country="Chile" pageId="3" pageNumber="552" stateProvince="Region Atacama" typeStatus="paratype">
<specimenCount id="9D4997F2A673BF65FA86B550E8969A18" box="[1345,1436,1815,1841]" pageId="3" pageNumber="552" type="male">64 male</specimenCount>
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at 6 
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, 6 
<collectionCode id="ED5EC4BEA673BF65FE69B57BEF089A7F" box="[430,514,1852,1878]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="3" pageNumber="552" type="Museum">FMNH</collectionCode>
, 6 IADIZA, 14 
<collectionCode id="ED5EC4BEA673BF65FD74B57BEFF79A7F" box="[691,765,1852,1878]" pageId="3" pageNumber="552">JMEC</collectionCode>
, 11 
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, 9 
<collectionCode id="ED5EC4BEA673BF65FC63B57BEEF19A7F" box="[932,1019,1852,1878]" country="Chile" httpUri="http://grbio.org/cool/mf2w-ryyx" name="Museo Nacional de Historia Natural, Santiago" pageId="3" pageNumber="552">MNNC</collectionCode>
, 6 
<collectionCode id="ED5EC4BEA673BF65FBDCB57BE98C9A7F" box="[1051,1158,1852,1878]" pageId="3" pageNumber="552">DNMCH</collectionCode>
, and 6 
<collectionCode id="ED5EC4BEA673BF65FB12B57BE82C9A7F" box="[1237,1318,1852,1878]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34868" lsid="urn:lsid:biocol.org:col:34868" name="University of Nebraska State Museum" pageId="3" pageNumber="552">UNSM</collectionCode>
labeled as 
<typeStatus id="54F4E2D9A673BF65FF50B518EDF39A50" box="[151,249,1887,1913]" pageId="3" pageNumber="552" type="holotype">holotype</typeStatus>
except: b) “ 
<taxonomicName id="4C4F27F8A673BF65FE45B527ECF49A50" authorityName="Martinez" authorityYear="1972" box="[386,510,1888,1913]" class="Insecta" family="Melolonthidae" genus="Luispenaia" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="552" phylum="Arthropoda" rank="genus">
<emphasis id="B93B8069A673BF65FE45B527ECF49A50" box="[386,510,1888,1913]" italics="true" pageId="3" pageNumber="552">Luispenaia</emphasis>
</taxonomicName>
/ 
<taxonomicName id="4C4F27F8A673BF65FDD3B527EF949A50" authority="Mondaca &amp; Pizarro-Araya &amp; Alfaro, 2019" authorityName="Mondaca &amp; Pizarro-Araya &amp; Alfaro" authorityYear="2019" box="[532,670,1888,1913]" class="Insecta" family="Melolonthidae" genus="Luispenaia" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="552" phylum="Arthropoda" rank="species" species="atacamensis" status="sp. nov.">
<emphasis id="B93B8069A673BF65FDD3B527EF949A50" box="[532,670,1888,1913]" italics="true" pageId="3" pageNumber="552">atacamensis</emphasis>
</taxonomicName>
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<emphasis id="B93B8069A673BF65FD62B527EFF49A53" bold="true" box="[677,766,1888,1914]" pageId="3" pageNumber="552">sp. nov.</emphasis>
</taxonomicNameLabel>
/ Mondaca, Pizarro-Araya &amp; Alfaro / 
<typeStatus id="54F4E2D9A673BF65FB6FB527E83D9A53" box="[1192,1335,1888,1914]" pageId="3" pageNumber="552" type="paratype">PARATYPE</typeStatus>
” (yellow label, typeset)
</materialsCitation>
. 
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<specimenCount id="9D4997F2A673BF65FE83B5C3EC949AB4" box="[324,414,1923,1950]" pageId="3" pageNumber="552" type="male">77 male</specimenCount>
<typeStatus id="54F4E2D9A673BF65FE60B5C3EF189AB7" box="[423,530,1924,1950]" pageId="3" pageNumber="552" type="paratype">paratypes</typeStatus>
al 
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labeled: a) “15-025 / 
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, prov. Huasco, / Boquerón Chañar, Llano / Algarrobal, 
<collectingDate id="EFB58353A673BF65FEEAB5EFECBC9AEB" box="[301,438,1959,1986]" pageId="3" pageNumber="552" value="2015-10-30">
<date id="FFF17ABBA673BF65FEEAB5EFECBC9AEB" box="[301,438,1959,1986]" pageId="3" pageNumber="552" value="2015-10-30">30 oct 2015</date>
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, / 
<geoCoordinate id="EE7B3ABCA673BF65FE16B5E0EF6B9AE8" box="[465,609,1959,1985]" degrees="28" direction="south" minutes="16" orientation="latitude" pageId="3" pageNumber="552" precision="1" seconds="0,7" value="-28.266861">28°16’0,7’’S</geoCoordinate>
<geoCoordinate id="EE7B3ABCA673BF65FDAFB5EFEE089AEB" box="[616,770,1960,1986]" degrees="70" direction="west" minutes="31" orientation="longitude" pageId="3" pageNumber="552" precision="1" seconds="1,4" value="-70.51706">70°31’1,4’’W</geoCoordinate>
/ 700 msnm, luz 
<collectionCode id="ED5EC4BEA673BF65FC0FB5EFEEFB9AEB" box="[968,1009,1960,1986]" country="Colombia" name="Departamento de Biologia de la Universidad del Valle" pageId="3" pageNumber="552">UV</collectionCode>
. Leg. J.E. / Barriga-Tuñon, 
<collectorName id="26BA39ADA673BF65FAECB5EFE8929AE8" box="[1323,1432,1959,1986]" pageId="3" pageNumber="552">J. Pizarro</collectorName>
, F. / Alfaro, 
<collectorName id="26BA39ADA673BF65FEE6B58BEC9E9ACF" box="[289,404,1996,2022]" pageId="3" pageNumber="552">R. Botero</collectorName>
, 
<collectorName id="26BA39ADA673BF65FE65B58BEF119ACF" box="[418,539,1996,2022]" pageId="3" pageNumber="552">R. Muñoz</collectorName>
”, b) “ 
<collectionCode id="ED5EC4BEA673BF65FD99B58CEE479ACF" box="[606,845,1995,2022]" pageId="3" pageNumber="552">COLECCIÓN JEBC</collectionCode>
/ Juan Enrique / Barriga-Tuñon / 
<collectingCountry id="F3581CEBA673BF65FB19B58BE83A9ACF" box="[1246,1328,1996,2022]" name="Chile" pageId="3" pageNumber="552">CHILE</collectingCountry>
0212981 / www.coleoptera-neotropical.org”, c) “ 
<taxonomicName id="4C4F27F8A674BF62FD9BB2DDEFD29D9A" authorityName="Martinez" authorityYear="1972" box="[604,728,154,179]" class="Insecta" family="Melolonthidae" genus="Luispenaia" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="553" phylum="Arthropoda" rank="genus">
<emphasis id="B93B8069A674BF62FD9BB2DDEFD29D9A" box="[604,728,154,179]" italics="true" pageId="4" pageNumber="553">Luispenaia</emphasis>
</taxonomicName>
/ 
<taxonomicName id="4C4F27F8A674BF62FD33B2DDEE749D9A" authority="Mondaca &amp; Pizarro-Araya &amp; Alfaro, 2019" authorityName="Mondaca &amp; Pizarro-Araya &amp; Alfaro" authorityYear="2019" box="[756,894,154,179]" class="Insecta" family="Melolonthidae" genus="Luispenaia" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="553" phylum="Arthropoda" rank="species" species="atacamensis" status="sp. nov.">
<emphasis id="B93B8069A674BF62FD33B2DDEE749D9A" box="[756,894,154,179]" italics="true" pageId="4" pageNumber="553">atacamensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2083D12A674BF62FC4EB2DEEEEF9D9A" box="[905,997,153,179]" pageId="4" pageNumber="553" rank="species">
<emphasis id="B93B8069A674BF62FC4EB2DEEEEF9D9A" bold="true" box="[905,997,153,179]" pageId="4" pageNumber="553">sp. nov.</emphasis>
</taxonomicNameLabel>
/ Mondaca, Pizarro-Araya &amp; Alfaro / 
<typeStatus id="54F4E2D9A674BF62FF50B2FAEC2C9DFE" box="[151,294,189,215]" pageId="4" pageNumber="553" type="paratype">PARATYPE</typeStatus>
” (red label, typeset)
</materialsCitation>
. 
<materialsCitation id="3B275626A674BF62FDD6B2FAEF679C36" ID-GBIF-Occurrence="2412528345" collectingDate="2015-10-23" collectionCode="RHSC" country="Chile" pageId="4" pageNumber="553" stateProvince="Region Atacama" typeStatus="paratype">
<specimenCount id="9D4997F2A674BF62FDD6B2FAEF519DFE" box="[529,603,189,215]" pageId="4" pageNumber="553" type="male">1 male</specimenCount>
<typeStatus id="54F4E2D9A674BF62FDA6B2FAEFCB9DFE" box="[609,705,189,215]" pageId="4" pageNumber="553" type="paratype">paratype</typeStatus>
at 
<collectionCode id="ED5EC4BEA674BF62FD25B2FAEE219DFE" box="[738,811,189,215]" pageId="4" pageNumber="553">RHSC</collectionCode>
labeled: a) “ 
<collectingDate id="EFB58353A674BF62FC71B2FAE9489DFE" box="[950,1090,188,215]" pageId="4" pageNumber="553" value="2015-10-23">
<date id="FFF17ABBA674BF62FC71B2FAE9489DFE" box="[950,1090,188,215]" pageId="4" pageNumber="553" value="2015-10-23">23/ X / 2015</date>
</collectingDate>
Entrada / mina Micifuz, S 
<location id="8E900AA0A674BF62FAABB2FAEDED9DD2" LSID="urn:lsid:plazi:treatment:03E6ED6DA673BF63FF50B407EC7098EA:8E900AA0A674BF62FAABB2FAEDED9DD2" country="Chile" latitude="-28.266861" longLatPrecision="1" longitude="70.520584" name="Inca de Oro" pageId="4" pageNumber="553" stateProvince="Atacama">Inca de Oro</location>
, III Reg. / 
<collectingCountry id="F3581CEBA674BF62FEA5B2A6ECBE9DD2" box="[354,436,225,251]" name="Chile" pageId="4" pageNumber="553">CHILE</collectingCountry>
<collectorName id="26BA39ADA674BF62FE7BB2A6EF879DD2" box="[444,653,225,251]" pageId="4" pageNumber="553">Leg. R. Honour S.</collectorName>
”, b) “ 
<taxonomicName id="4C4F27F8A674BF62FD15B2A5EE449DD2" authorityName="Martinez" authorityYear="1972" box="[722,846,226,251]" class="Insecta" family="Melolonthidae" genus="Luispenaia" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="553" phylum="Arthropoda" rank="genus">
<emphasis id="B93B8069A674BF62FD15B2A5EE449DD2" box="[722,846,226,251]" italics="true" pageId="4" pageNumber="553">Luispenaia</emphasis>
</taxonomicName>
/ 
<taxonomicName id="4C4F27F8A674BF62FCA1B2A5EEFA9DD2" authority="Mondaca &amp; Pizarro-Araya &amp; Alfaro, 2019" authorityName="Mondaca &amp; Pizarro-Araya &amp; Alfaro" authorityYear="2019" box="[870,1008,226,251]" class="Insecta" family="Melolonthidae" genus="Luispenaia" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="553" phylum="Arthropoda" rank="species" species="atacamensis" status="sp. nov.">
<emphasis id="B93B8069A674BF62FCA1B2A5EEFA9DD2" box="[870,1008,226,251]" italics="true" pageId="4" pageNumber="553">atacamensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2083D12A674BF62FC3FB2A6E9589DD2" box="[1016,1106,225,251]" pageId="4" pageNumber="553" rank="species">
<emphasis id="B93B8069A674BF62FC3FB2A6E9589DD2" bold="true" box="[1016,1106,225,251]" pageId="4" pageNumber="553">sp. nov.</emphasis>
</taxonomicNameLabel>
/ Mondaca, Pizarro-Araya &amp; Alfaro / 
<typeStatus id="54F4E2D9A674BF62FF31B342EC8F9C36" box="[246,389,261,287]" pageId="4" pageNumber="553" type="paratype">PARATYPE</typeStatus>
” (red label, typeset).
</materialsCitation>
</paragraph>
</subSubSection>
<caption id="DF300CF3A674BF62FF50B4ADE9649A6A" httpUri="https://zenodo.org/record/3246213/files/figure.png" pageId="4" pageNumber="553" startId="4.[151,250,1770,1795]" targetBox="[151,1405,287,1737]" targetPageId="4">
<paragraph id="8BF05C7BA674BF62FF50B4ADE9649A6A" blockId="4.[151,1436,1770,1859]" pageId="4" pageNumber="553">
<emphasis id="B93B8069A674BF62FF50B4ADEC279A2A" bold="true" box="[151,301,1770,1795]" pageId="4" pageNumber="553">FIGURE 1–7.</emphasis>
Generic characters of the genus 
<taxonomicName id="4C4F27F8A674BF62FDBBB4ACEFE79A2B" authorityName="Martinez" authorityYear="1972" box="[636,749,1771,1794]" class="Insecta" family="Melolonthidae" genus="Luispenaia" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="553" phylum="Arthropoda" rank="genus">
<emphasis id="B93B8069A674BF62FDBBB4ACEFE79A2B" box="[636,749,1771,1794]" italics="true" pageId="4" pageNumber="553">Luispenaia</emphasis>
</taxonomicName>
. 
<emphasis id="B93B8069A674BF62FD3DB4ADEE0D9A2A" bold="true" box="[762,775,1770,1795]" pageId="4" pageNumber="553">1</emphasis>
, head and antennal club; 
<emphasis id="B93B8069A674BF62FBCAB4ADE9109A2A" bold="true" box="[1037,1050,1770,1795]" pageId="4" pageNumber="553">2</emphasis>
, eyes and mouth part in ventral view; 
<emphasis id="B93B8069A674BF62FF50B54DEDAE9A0A" bold="true" box="[151,164,1802,1827]" pageId="4" pageNumber="553">3</emphasis>
, eyes and mouth part in lateral view; 
<emphasis id="B93B8069A674BF62FDE7B54DEF279A0A" bold="true" box="[544,557,1802,1827]" pageId="4" pageNumber="553">4</emphasis>
, right wing; 
<emphasis id="B93B8069A674BF62FD69B54CEFB19A0A" bold="true" box="[686,699,1803,1827]" pageId="4" pageNumber="553">5</emphasis>
, pretarsal claw; 
<emphasis id="B93B8069A674BF62FCA6B54DEE649A0A" bold="true" box="[865,878,1802,1827]" pageId="4" pageNumber="553">6</emphasis>
, abdomen in ventral view; 
<emphasis id="B93B8069A674BF62FB45B54CE9859A0A" bold="true" box="[1154,1167,1803,1827]" pageId="4" pageNumber="553">7</emphasis>
, metatibial apex in frontolateral view. Scale: 0.5 mm (Figs. 1–3); 1 mm (Figs. 4, 6); 0.25 mm (Fig. 5); 0.5 mm (Figs. 5, 7).
</paragraph>
</caption>
<subSubSection id="C3550FF0A674BF63FF00B537EE559DFE" lastPageId="5" lastPageNumber="554" pageId="4" pageNumber="553" type="diagnosis">
<paragraph id="8BF05C7BA674BF63FF00B537EE559DFE" blockId="4.[151,1436,1904,2040]" lastBlockId="5.[151,1437,153,1475]" lastPageId="5" lastPageNumber="554" pageId="4" pageNumber="553">
<emphasis id="B93B8069A674BF62FF00B537EC489AA2" bold="true" box="[199,322,1904,1931]" pageId="4" pageNumber="553">Diagnosis.</emphasis>
This species is distinguished from all other species in the genus 
<taxonomicName id="4C4F27F8A674BF62FBCAB535E9839AA2" authorityName="Martinez" authorityYear="1972" box="[1037,1161,1906,1931]" class="Insecta" family="Melolonthidae" genus="Luispenaia" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="553" phylum="Arthropoda" rank="genus">
<emphasis id="B93B8069A674BF62FBCAB535E9839AA2" box="[1037,1161,1906,1931]" italics="true" pageId="4" pageNumber="553">Luispenaia</emphasis>
</taxonomicName>
by the following combination of characters: Body dorsally reddish brown, shiny on legs and elytra; head pronotum and scutellum black or dark brown (
<figureCitation id="137440FEA674BF62FEEFB5FEEC719AFA" box="[296,379,1977,2003]" captionStart="FIGURES 18–29" captionStartId="10.[151,264,1736,1761]" captionTargetBox="[164,1379,530,1707]" captionTargetId="figure@10.[151,1436,513,1724]" captionTargetPageId="10" captionText="FIGURES 18–29. Luispenaia species. 18–20, Luispenaia atacamensis; 21–23, L. paposo; 24–26, L. paulseni; 27–29, L. paradoxa. Scale: 1 mm." httpUri="https://zenodo.org/record/3246224/files/figure.png" pageId="4" pageNumber="553">Fig. 18</figureCitation>
). Clypeus distal and laterally rounded, reflexed upturned; lateral margin between clypeal base and ocular canthus projected externally as a lobule moderately projecting (
<figureCitation id="137440FEA674BF62FC12B59AE9229ADE" box="[981,1064,2013,2039]" captionStart="FIGURES 10–13" captionStartId="8.[151,264,1969,1994]" captionTargetBox="[175,1412,720,1919]" captionTargetId="figure@8.[151,1436,729,1957]" captionTargetPageId="8" captionText="FIGURES 10–13. Head and pronotum of Luispenaia species. 10, Luispenaia atacamensis; 11, L. paposo; 12, L. paulseni; 13, L. paradoxa. Scale: 0.5 mm." httpUri="https://zenodo.org/record/3246220/files/figure.png" pageId="4" pageNumber="553">Fig. 10</figureCitation>
). Antennae with 9 antennomeres; antennal club with 6 antennomeres (
<figureCitation id="137440FEA675BF63FDEBB2DEEF759D9A" box="[556,639,153,179]" captionStart="FIGURES 14–17" captionStartId="9.[151,264,1456,1481]" captionTargetBox="[151,1436,170,1444]" captionTargetId="figure@9.[151,1436,170,1444]" captionTargetPageId="9" captionText="FIGURES 14–17. Antennal club of Luispenaia species. 14, Luispenaia paradoxa Martínez; 15, L. paposo; 16, L. paulseni; 17, L. atacamensis. Scale: 0.5 mm." httpUri="https://zenodo.org/record/3246222/files/figure.png" pageId="5" pageNumber="554">Fig. 17</figureCitation>
). Pronotum approximately subtrapezoidal, broadest in middle, laterally arcuate; surface with longitudinal midline impunctate (
<figureCitation id="137440FEA675BF63FD39B2FAEE5B9DFE" box="[766,849,189,215]" captionStart="FIGURES 10–13" captionStartId="8.[151,264,1969,1994]" captionTargetBox="[175,1412,720,1919]" captionTargetId="figure@8.[151,1436,729,1957]" captionTargetPageId="8" captionText="FIGURES 10–13. Head and pronotum of Luispenaia species. 10, Luispenaia atacamensis; 11, L. paposo; 12, L. paulseni; 13, L. paradoxa. Scale: 0.5 mm." httpUri="https://zenodo.org/record/3246220/files/figure.png" pageId="5" pageNumber="554">Fig. 10</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3550FF0A675BF63FF00B2A7EE809EAA" pageId="5" pageNumber="554" type="description">
<paragraph id="8BF05C7BA675BF63FF00B2A7EF3F9E3E" blockId="5.[151,1437,153,1475]" pageId="5" pageNumber="554">
<emphasis id="B93B8069A675BF63FF00B2A7EF2E9DD2" bold="true" box="[199,548,224,251]" pageId="5" pageNumber="554">
Description of male 
<typeStatus id="54F4E2D9A675BF63FE71B2A6EF159DD2" box="[438,543,225,251]" pageId="5" pageNumber="554" type="holotype">holotype</typeStatus>
.
</emphasis>
Length 
<quantity id="4CB7F19EA675BF63FD46B2A7EFD29DD2" box="[641,728,224,251]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" pageId="5" pageNumber="554" unit="mm" value="5.5">5.5 mm</quantity>
, wide 2.5. 
<emphasis id="B93B8069A675BF63FC96B2A6EE819DD2" box="[849,907,225,251]" italics="true" pageId="5" pageNumber="554">Body</emphasis>
: Slightly convex, dorsally reddish brown shiny on legs and elytra, head, pronotum, and scutellum dark brown (
<figureCitation id="137440FEA675BF63FC93B342EEAC9C36" box="[852,934,261,287]" captionStart="FIGURES 18–29" captionStartId="10.[151,264,1736,1761]" captionTargetBox="[164,1379,530,1707]" captionTargetId="figure@10.[151,1436,513,1724]" captionTargetPageId="10" captionText="FIGURES 18–29. Luispenaia species. 18–20, Luispenaia atacamensis; 21–23, L. paposo; 24–26, L. paulseni; 27–29, L. paradoxa. Scale: 1 mm." httpUri="https://zenodo.org/record/3246224/files/figure.png" pageId="5" pageNumber="554">Fig. 18</figureCitation>
). 
<emphasis id="B93B8069A675BF63FC7DB342EEFD9C36" box="[954,1015,261,287]" italics="true" pageId="5" pageNumber="554">Head</emphasis>
: Clypeus distal and laterally rounded, reflexed upturned, laterally with a prolongation of the margins as external lobules moderately projecting (
<figureCitation id="137440FEA675BF63FAFDB36EE8849C6A" box="[1338,1422,297,323]" captionStart="FIGURES 10–13" captionStartId="8.[151,264,1969,1994]" captionTargetBox="[175,1412,720,1919]" captionTargetId="figure@8.[151,1436,729,1957]" captionTargetPageId="8" captionText="FIGURES 10–13. Head and pronotum of Luispenaia species. 10, Luispenaia atacamensis; 11, L. paposo; 12, L. paulseni; 13, L. paradoxa. Scale: 0.5 mm." httpUri="https://zenodo.org/record/3246220/files/figure.png" pageId="5" pageNumber="554">Fig. 10</figureCitation>
); dorsal surface punctate and setose; punctures large; frontoclypeal suture well defined, frontal width equal to 5.6 times eye width, with surface densely punctate and setose, punctures large; eye canthus short and wide, straight externally. Antennae with 9 antennomeres; antennomere 1 robust, claviform; antennomere 2 submonoliform; antennomere 3 subcylindrical; antennal club with 6 antennomeres glabrous, antennomere 4 lamellate, shorter than antennomeres 5–9; antennomeres 5–9 subequal in length, slightly sinuous in dorsal view (
<figureCitation id="137440FEA675BF63FB9DB39AE9A59CDE" box="[1114,1199,477,503]" captionStart="FIGURES 14–17" captionStartId="9.[151,264,1456,1481]" captionTargetBox="[151,1436,170,1444]" captionTargetId="figure@9.[151,1436,170,1444]" captionTargetPageId="9" captionText="FIGURES 14–17. Antennal club of Luispenaia species. 14, Luispenaia paradoxa Martínez; 15, L. paposo; 16, L. paulseni; 17, L. atacamensis. Scale: 0.5 mm." httpUri="https://zenodo.org/record/3246222/files/figure.png" pageId="5" pageNumber="554">Fig. 17</figureCitation>
). 
<emphasis id="B93B8069A675BF63FB03B399E8399CDE" box="[1220,1331,478,503]" italics="true" pageId="5" pageNumber="554">Pronotum</emphasis>
: Slightly convex, approximately subtrapezoidal, broadest in middle, laterally arcuate; surface moderately punctate and setose; setae recumbent, directed posteriorly; longitudinal midline impunctate, lateral margin with long, moderately dense, laterally directed setae (
<figureCitation id="137440FEA675BF63FE2BB00EEF349F4A" box="[492,574,585,611]" captionStart="FIGURES 10–13" captionStartId="8.[151,264,1969,1994]" captionTargetBox="[175,1412,720,1919]" captionTargetId="figure@8.[151,1436,729,1957]" captionTargetPageId="8" captionText="FIGURES 10–13. Head and pronotum of Luispenaia species. 10, Luispenaia atacamensis; 11, L. paposo; 12, L. paulseni; 13, L. paradoxa. Scale: 0.5 mm." httpUri="https://zenodo.org/record/3246220/files/figure.png" pageId="5" pageNumber="554">Fig. 10</figureCitation>
). Scutellum as wide as long, triangular; sides broadly arcuate. 
<emphasis id="B93B8069A675BF63FB37B00EE83C9F4A" box="[1264,1334,585,611]" italics="true" pageId="5" pageNumber="554">Elytra</emphasis>
: Slightly convex, surface moderately setose; setae recumbent, directed posteriorly, with striae poorly defined; lateral margin with long, moderately dense, laterally directed setae. 
<emphasis id="B93B8069A675BF63FD2DB0D5EE159F82" box="[746,799,658,683]" italics="true" pageId="5" pageNumber="554">Legs</emphasis>
: All them moderately setose and shiny, setae light brown long. Protibiae tridentate, with teeth rounded distally. 
<emphasis id="B93B8069A675BF63FD33B0F2EE549FE6" box="[756,862,693,719]" italics="true" pageId="5" pageNumber="554">Genitalia</emphasis>
: Parameres moderately long, wider at base (frontal view), laterally cuneiform, slightly curved ventrally, distally attenuated and rounded at apex; phallobase 1.13 times longer than parameres (
<figureCitation id="137440FEA675BF63FE58B0BAEF239E3E" box="[415,553,765,791]" captionStart="FIGURES 18–29" captionStartId="10.[151,264,1736,1761]" captionTargetBox="[164,1379,530,1707]" captionTargetId="figure@10.[151,1436,513,1724]" captionTargetPageId="10" captionText="FIGURES 18–29. Luispenaia species. 18–20, Luispenaia atacamensis; 21–23, L. paposo; 24–26, L. paulseni; 27–29, L. paradoxa. Scale: 1 mm." httpUri="https://zenodo.org/record/3246224/files/figure.png" pageId="5" pageNumber="554">Figs. 19–20</figureCitation>
).
</paragraph>
<paragraph id="8BF05C7BA675BF63FF00B166ECA89E12" blockId="5.[151,1437,153,1475]" box="[199,418,801,827]" pageId="5" pageNumber="554">
<emphasis id="B93B8069A675BF63FF00B166EC2F9E12" bold="true" box="[199,293,801,827]" pageId="5" pageNumber="554">Female.</emphasis>
Unknown.
</paragraph>
<paragraph id="8BF05C7BA675BF63FF00B103EE809EAA" blockId="5.[151,1437,153,1475]" pageId="5" pageNumber="554">
<emphasis id="B93B8069A675BF63FF00B103EC359E76" bold="true" box="[199,319,836,863]" pageId="5" pageNumber="554">Variation.</emphasis>
Length 4.8–7.0 mm. Body dorsal color light brown to dark brown, pronotum dark brown to black. Not differing significantly from the 
<typeStatus id="54F4E2D9A675BF63FDEDB12EEF869EAA" box="[554,652,873,899]" pageId="5" pageNumber="554" type="holotype">holotype</typeStatus>
in external characters.
</paragraph>
</subSubSection>
<subSubSection id="C3550FF0A675BF63FF00B1CAE9FA9E8E" box="[199,1264,909,935]" pageId="5" pageNumber="554" type="etymology">
<paragraph id="8BF05C7BA675BF63FF00B1CAE9FA9E8E" blockId="5.[151,1437,153,1475]" box="[199,1264,909,935]" pageId="5" pageNumber="554">
<emphasis id="B93B8069A675BF63FF00B1CAEC449E8E" bold="true" box="[199,334,909,935]" pageId="5" pageNumber="554">Etymology.</emphasis>
Derived from 
<collectingRegion id="498B9299A675BF63FE35B1CAEF529E8E" box="[498,600,909,935]" country="Chile" name="Atacama" pageId="5" pageNumber="554">Atacama</collectingRegion>
, origin region of the new species located in northern 
<collectingCountry id="F3581CEBA675BF63FB6CB1CAE9E19E8E" box="[1195,1259,909,935]" name="Chile" pageId="5" pageNumber="554">Chile</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C3550FF0A675BF63FF00B1F7EEAE9EE2" box="[199,932,944,972]" pageId="5" pageNumber="554" type="distribution">
<paragraph id="8BF05C7BA675BF63FF00B1F7EEAE9EE2" blockId="5.[151,1437,153,1475]" box="[199,932,944,972]" pageId="5" pageNumber="554">
<emphasis id="B93B8069A675BF63FF00B1F7EC6B9EE2" bold="true" box="[199,353,944,971]" pageId="5" pageNumber="554">Distribution.</emphasis>
<collectingCountry id="F3581CEBA675BF63FEA0B1F6ECAC9EE2" box="[359,422,945,971]" name="Chile" pageId="5" pageNumber="554">Chile</collectingCountry>
, 
<collectingRegion id="498B9299A675BF63FE76B1F6EF669EE2" box="[433,620,945,971]" country="Chile" name="Atacama" pageId="5" pageNumber="554">Atacama Region</collectingRegion>
(Huasco province) (
<figureCitation id="137440FEA675BF63FC95B1F6EE9C9EE2" box="[850,918,945,971]" captionStart="FIGURE 8" captionStartId="6.[151,250,1996,2021]" captionTargetBox="[192,1388,211,1949]" captionTargetId="figure@6.[151,1435,170,1984]" captionTargetPageId="6" captionText="FIGURE 8. Distribution of Luispenaia species in the Atacama Desert, northern Chile." httpUri="https://zenodo.org/record/3246215/files/figure.png" pageId="5" pageNumber="554">Fig. 8</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3550FF0A675BF63FF00B193EC7098EA" pageId="5" pageNumber="554" type="materials_examined">
<paragraph id="8BF05C7BA675BF63FF00B193E9529EC6" blockId="5.[151,1437,153,1475]" box="[199,1112,980,1007]" pageId="5" pageNumber="554">
<emphasis id="B93B8069A675BF63FF00B193ECBB9EC6" bold="true" box="[199,433,980,1007]" pageId="5" pageNumber="554">Material examined.</emphasis>
<materialsCitation id="3B275626A675BF63FE70B192E95E9EC6" ID-GBIF-Occurrence="2265605703" box="[439,1108,981,1007]" collectionCode="JEBC, LEULS, RHSC" pageId="5" pageNumber="554" specimenCount="141">
<specimenCount id="9D4997F2A675BF63FE70B192EF579EC6" box="[439,605,981,1007]" pageId="5" pageNumber="554" type="generic">141 specimens</specimenCount>
were examined from 
<collectionCode id="ED5EC4BEA675BF63FC94B192EE9D9EC6" box="[851,919,981,1007]" pageId="5" pageNumber="554">JEBC</collectionCode>
, 
<collectionCode id="ED5EC4BEA675BF63FC65B192EEF69EC6" box="[930,1020,981,1007]" pageId="5" pageNumber="554">LEULS</collectionCode>
, 
<collectionCode id="ED5EC4BEA675BF63FBC0B192E95E9EC6" box="[1031,1108,981,1007]" pageId="5" pageNumber="554">RHSC</collectionCode>
</materialsCitation>
.
</paragraph>
<paragraph id="8BF05C7BA675BF63FF00B1BFEC7098EA" blockId="5.[151,1437,153,1475]" pageId="5" pageNumber="554">
<emphasis id="B93B8069A675BF63FF00B1BFECCF993A" bold="true" box="[199,453,1016,1043]" pageId="5" pageNumber="554">Habitat and ecology.</emphasis>
<taxonomicName id="4C4F27F8A675BF63FE17B1BDEFEB993A" authorityName="Mondaca &amp; Pizarro-Araya &amp; Alfaro" authorityYear="2019" box="[464,737,1018,1043]" class="Insecta" family="Melolonthidae" genus="Luispenaia" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="554" phylum="Arthropoda" rank="species" species="atacamensis">
<emphasis id="B93B8069A675BF63FE17B1BDEFEB993A" box="[464,737,1018,1043]" italics="true" pageId="5" pageNumber="554">Luispenaia atacamensis</emphasis>
</taxonomicName>
has been collected in vegetation floors characterized by an interior Mediterranean desert scrub of 
<taxonomicName id="4C4F27F8A675BF63FD96B65AEE7F991E" authority="Meyen" authorityName="Meyen" box="[593,885,1053,1079]" class="Magnoliopsida" family="Fabaceae" genus="Adesmia" kingdom="Plantae" order="Fabales" pageId="5" pageNumber="554" phylum="Tracheophyta" rank="species" species="argentea">
<emphasis id="B93B8069A675BF63FD96B65AEE14991E" box="[593,798,1053,1079]" italics="true" pageId="5" pageNumber="554">Adesmia argentea</emphasis>
Meyen
</taxonomicName>
and 
<taxonomicName id="4C4F27F8A675BF63FC68B65AEDD69975" authority="Gay (Luebert &amp; Pliscoff 2006)" authorityName="Gay (Luebert &amp; Pliscoff" authorityYear="2006" class="Magnoliopsida" family="Zygophyllaceae" genus="Bulnesia" kingdom="Plantae" order="Zygophyllales" pageId="5" pageNumber="554" phylum="Tracheophyta" rank="species" species="chilensis">
<emphasis id="B93B8069A675BF63FC68B65AE976991E" box="[943,1148,1053,1079]" italics="true" pageId="5" pageNumber="554">Bulnesia chilensis</emphasis>
Gay (
<bibRefCitation id="EFDE218AA675BF63FB02B65AEDDE9975" author="Luebert, F. &amp; Pliscoff, P." pageId="5" pageNumber="554" refId="ref7924" refString="Luebert, F. &amp; Pliscoff, P. (2006) Sinopsis Bioclimatica y Vegetacional de Chile. Editorial Universitaria, Santiago, 316 pp." type="book" year="2006">Luebert &amp; Pliscoff 2006</bibRefCitation>
)
</taxonomicName>
. These areas are represented by a very open thicket with tall bushes and the presence of tree species; among the bushes are 
<taxonomicName id="4C4F27F8A675BF63FE87B622ECC99956" box="[320,451,1125,1151]" class="Magnoliopsida" family="Fabaceae" genus="Adesmia" kingdom="Plantae" order="Fabales" pageId="5" pageNumber="554" phylum="Tracheophyta" rank="species" species="argentea">
<emphasis id="B93B8069A675BF63FE87B622ECC99956" box="[320,451,1125,1151]" italics="true" pageId="5" pageNumber="554">A. argentea</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C4F27F8A675BF63FE15B622EF949956" authorityName="Gay (Luebert &amp; Pliscoff" authorityYear="2006" box="[466,670,1125,1151]" class="Magnoliopsida" family="Zygophyllaceae" genus="Bulnesia" kingdom="Plantae" order="Zygophyllales" pageId="5" pageNumber="554" phylum="Tracheophyta" rank="species" species="chilensis">
<emphasis id="B93B8069A675BF63FE15B622EF949956" box="[466,670,1125,1151]" italics="true" pageId="5" pageNumber="554">Bulnesia chilensis</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C4F27F8A675BF63FD6AB622E92E9956" authority="Clos" authorityName="Clos" authorityYear="1994" box="[685,1060,1125,1151]" class="Magnoliopsida" family="Fabaceae" genus="Balsamocarpon" kingdom="Plantae" order="Fabales" pageId="5" pageNumber="554" phylum="Tracheophyta" rank="species" species="brevifolium">
<emphasis id="B93B8069A675BF63FD6AB622EEEC9956" box="[685,998,1125,1151]" italics="true" pageId="5" pageNumber="554">Balsamocarpon brevifolium</emphasis>
Clos
</taxonomicName>
, 
<taxonomicName id="4C4F27F8A675BF63FBF6B622E8929956" authority="Hook. &amp; Arn." authorityName="Hook. &amp; Arn." box="[1073,1432,1125,1151]" class="Insecta" family="Aphrophoridae" genus="Cordia" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="554" phylum="Arthropoda" rank="species" species="decandra">
<emphasis id="B93B8069A675BF63FBF6B622E9FB9956" box="[1073,1265,1125,1151]" italics="true" pageId="5" pageNumber="554">Cordia decandra</emphasis>
Hook. &amp; Arn.
</taxonomicName>
, 
<taxonomicName id="4C4F27F8A675BF63FF50B6CEEF8D998A" authority="(Miers) I. M. Johnst." authorityName="I. M. Johnst." baseAuthorityName="Miers" box="[151,647,1161,1188]" class="Magnoliopsida" family="Boraginaceae" genus="Heliotropium" kingdom="Plantae" order="Boraginales" pageId="5" pageNumber="554" phylum="Tracheophyta" rank="species" species="sinuatum">
<emphasis id="B93B8069A675BF63FF50B6CEEC91998A" box="[151,411,1161,1187]" italics="true" pageId="5" pageNumber="554">Heliotropium sinuatum</emphasis>
(Miers) I.M. Johnst.
</taxonomicName>
, 
<taxonomicName id="4C4F27F8A675BF63FD55B6CDEE74998A" authority="Gay" authorityName="Gay" box="[658,894,1161,1187]" class="Insecta" family="Trichogrammatidae" genus="Pintoa" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="554" phylum="Arthropoda" rank="species" species="chilensis">
<emphasis id="B93B8069A675BF63FD55B6CDEE4C998A" box="[658,838,1161,1187]" italics="true" pageId="5" pageNumber="554">Pintoa chilensis</emphasis>
Gay
</taxonomicName>
, and 
<taxonomicName id="4C4F27F8A675BF63FC7EB6CDE851998A" authority="Hook. &amp; Arn. Also" authorityName="Hook. &amp; Arn. Also" box="[953,1371,1161,1187]" class="Magnoliopsida" family="Asteraceae" genus="Proustia" kingdom="Plantae" order="Asterales" pageId="5" pageNumber="554" phylum="Tracheophyta" rank="species" species="ilicifolia">
<emphasis id="B93B8069A675BF63FC7EB6CDE975998A" box="[953,1151,1161,1187]" italics="true" pageId="5" pageNumber="554">Proustia ilicifolia</emphasis>
Hook. &amp; Arn. Also
</taxonomicName>
common are low shrubs such as 
<taxonomicName id="4C4F27F8A675BF63FE24B6EAEED199EE" authority="(Hook. &amp; Arn.) Baill." authorityName="Baill." baseAuthorityName="Hook. &amp; Arn." box="[483,987,1197,1223]" class="Magnoliopsida" family="Fabaceae" genus="Caesalpinia" kingdom="Plantae" order="Fabales" pageId="5" pageNumber="554" phylum="Tracheophyta" rank="species" species="angulate">
<emphasis id="B93B8069A675BF63FE24B6EAEFDC99EE" box="[483,726,1197,1223]" italics="true" pageId="5" pageNumber="554">Caesalpinia angulate</emphasis>
(Hook. &amp; Arn.) Baill.
</taxonomicName>
, 
<taxonomicName id="4C4F27F8A675BF63FC2EB6EAE80B99EE" authority="Lam." authorityName="Lam." box="[1001,1281,1197,1223]" class="Magnoliopsida" family="Asteraceae" genus="Encelia" kingdom="Plantae" order="Asterales" pageId="5" pageNumber="554" phylum="Tracheophyta" rank="species" species="canescens">
<emphasis id="B93B8069A675BF63FC2EB6EAE9B099EE" box="[1001,1210,1197,1223]" italics="true" pageId="5" pageNumber="554">Encelia canescens</emphasis>
Lam.
</taxonomicName>
, 
<taxonomicName id="4C4F27F8A675BF63FAC9B6EAEC4499C2" authority="D. Don" authorityName="D. Don" class="Magnoliopsida" family="Lythraceae" genus="Pleurophora" kingdom="Plantae" order="Myrtales" pageId="5" pageNumber="554" phylum="Tracheophyta" rank="species" species="pungens">
<emphasis id="B93B8069A675BF63FAC9B6EAEDFE99C2" italics="true" pageId="5" pageNumber="554">Pleurophora pungens</emphasis>
D. Don
</taxonomicName>
, and the cacti, 
<taxonomicName id="4C4F27F8A675BF63FE37B696E93299C2" authority="(C. F. Forst.) E. F. Anderson" authorityName="E. F. Anderson" baseAuthorityName="C. F. Forst." box="[496,1080,1233,1259]" class="Magnoliopsida" family="Cactaceae" genus="Cumulopuntia" kingdom="Plantae" order="Caryophyllales" pageId="5" pageNumber="554" phylum="Tracheophyta" rank="species" species="sphaerica">
<emphasis id="B93B8069A675BF63FE37B696EE0E99C2" box="[496,772,1233,1259]" italics="true" pageId="5" pageNumber="554">Cumulopuntia sphaerica</emphasis>
(C.F. Först.) E.F. Anderson
</taxonomicName>
, and 
<taxonomicName id="4C4F27F8A675BF63FBB7B696ECEB9826" authority="(Molina) Britton &amp; Rose. The" authorityName="Britton &amp; Rose. The" baseAuthorityName="Molina" class="Magnoliopsida" family="Cactaceae" genus="Trichocereus" higherTaxonomySource="CoL" kingdom="Plantae" order="Caryophyllales" pageId="5" pageNumber="554" phylum="Tracheophyta" rank="species" species="coquimbanus">
<emphasis id="B93B8069A675BF63FBB7B696E89699C2" box="[1136,1436,1233,1259]" italics="true" pageId="5" pageNumber="554">Trichocereus coquimbanus</emphasis>
(Molina) Britton &amp; Rose. The
</taxonomicName>
herbaceous plants are abundant during the rainy season as 
<taxonomicName id="4C4F27F8A675BF63FBAEB6B2E8969826" authority="Clos" authorityName="Clos" authorityYear="1848" box="[1129,1436,1269,1295]" class="Magnoliopsida" family="Rubiaceae" genus="Cruckshanksia" kingdom="Plantae" order="Gentianales" pageId="5" pageNumber="554" phylum="Tracheophyta" rank="species" species="pumila">
<emphasis id="B93B8069A675BF63FBAEB6B2E8699826" box="[1129,1379,1269,1295]" italics="true" pageId="5" pageNumber="554">Cruckshanksia pumila</emphasis>
Clos
</taxonomicName>
and 
<taxonomicName id="4C4F27F8A675BF63FF00B75EEC78981A" box="[199,370,1305,1331]" class="Magnoliopsida" family="Bignoniaceae" genus="Argylia" kingdom="Plantae" order="Lamiales" pageId="5" pageNumber="554" phylum="Tracheophyta" rank="species" species="radiate">
<emphasis id="B93B8069A675BF63FF00B75EEC78981A" box="[199,370,1305,1331]" italics="true" pageId="5" pageNumber="554">Argylia radiate</emphasis>
</taxonomicName>
(L.) D. Don. In the ravines proper such trees include 
<taxonomicName id="4C4F27F8A675BF63FC0DB75DE85B981A" authority="(Molina) Stuntz" authorityName="Stuntz" baseAuthorityName="Molina" box="[970,1361,1305,1332]" class="Magnoliopsida" family="Fabaceae" genus="Prosopis" kingdom="Animalia" order="Fabales" pageId="5" pageNumber="554" phylum="Tracheophyta" rank="species" species="chilensis">
<emphasis id="B93B8069A675BF63FC0DB75DE99F981A" box="[970,1173,1305,1331]" italics="true" pageId="5" pageNumber="554">Prosopis chilensis</emphasis>
(Molina) Stuntz
</taxonomicName>
, 
<taxonomicName id="4C4F27F8A675BF63FA9BB75EEFEB987E" authority="(Gillies ex Hook. &amp; Arn.) Burkart" authorityName="Burkart" authorityYear="1949" baseAuthorityName="Gillies ex Hook. &amp; Arn." class="Magnoliopsida" family="Fabaceae" genus="Geoffroea" kingdom="Plantae" order="Fabales" pageId="5" pageNumber="554" phylum="Tracheophyta" rank="species" species="decorticans">
<emphasis id="B93B8069A675BF63FA9BB75EEC51987E" italics="true" pageId="5" pageNumber="554">Geoffroea decorticans</emphasis>
(Gillies ex Hook. &amp; Arn.) Burkart
</taxonomicName>
, 
<taxonomicName id="4C4F27F8A675BF63FD2AB77AE943987E" authority="(Molina) Molina" authorityName="Molina" baseAuthorityName="Molina" box="[749,1097,1341,1368]" class="Magnoliopsida" family="Fabaceae" genus="Acacia" kingdom="Plantae" order="Fabales" pageId="5" pageNumber="554" phylum="Tracheophyta" rank="species" species="caven">
<emphasis id="B93B8069A675BF63FD2AB77AEE89987E" box="[749,899,1341,1367]" italics="true" pageId="5" pageNumber="554">Acacia caven</emphasis>
(Molina) Molina
</taxonomicName>
, and 
<taxonomicName id="4C4F27F8A675BF63FB43B77AEF289855" authority="(Cav.) Cabrera (Luebert &amp; Pliscoff 2006)" authorityName="Cabrera (Luebert &amp; Pliscoff" authorityYear="2006" class="Magnoliopsida" family="Anacardiaceae" genus="Schinus" kingdom="Plantae" order="Sapindales" pageId="5" pageNumber="554" phylum="Tracheophyta" rank="species" species="polygama">
<emphasis id="B93B8069A675BF63FB43B77AE85A987E" box="[1156,1360,1341,1367]" italics="true" pageId="5" pageNumber="554">Schinus polygama</emphasis>
(Cav.) Cabrera (
<bibRefCitation id="EFDE218AA675BF63FF38B726EF119855" author="Luebert, F. &amp; Pliscoff, P." box="[255,539,1377,1404]" pageId="5" pageNumber="554" refId="ref7924" refString="Luebert, F. &amp; Pliscoff, P. (2006) Sinopsis Bioclimatica y Vegetacional de Chile. Editorial Universitaria, Santiago, 316 pp." type="book" year="2006">Luebert &amp; Pliscoff 2006</bibRefCitation>
)
</taxonomicName>
. Most specimens of 
<taxonomicName id="4C4F27F8A675BF63FCCBB725EEBF9852" authorityName="Mondaca &amp; Pizarro-Araya &amp; Alfaro" authorityYear="2019" box="[780,949,1378,1403]" class="Insecta" family="Melolonthidae" genus="Luispenaia" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="554" phylum="Arthropoda" rank="species" species="atacamensis">
<emphasis id="B93B8069A675BF63FCCBB725EEBF9852" box="[780,949,1378,1403]" italics="true" pageId="5" pageNumber="554">L. atacamensis</emphasis>
</taxonomicName>
were captured during the spring (October) with a light trap installed in an area with the native vegetation described above (
<figureCitation id="137440FEA675BF63FBCBB7C2E97898B6" box="[1036,1138,1413,1439]" captionStart="FIGURES 30" captionStartId="11.[151,264,1675,1700]" captionTargetBox="[151,1435,801,1663]" captionTargetId="figure@11.[151,1435,801,1663]" captionTargetPageId="11" captionText="FIGURES 30. Type locality of Luispenaia species. A, Llanos de Challe National Park (Atacama Region, Chile), habitat of L. paradoxa; B, Boqueron Chañar (Atacama Region, Chile), habitat of L. atacamensis; C, Paposo Norte Natural Monument (Antofagasta Region, Chile), habitat of L. paposo; D, Quebrada Los Yales (Antofagasta Region, Chile), habitat of L. paulseni." httpUri="https://zenodo.org/record/3246226/files/figure.png" pageId="5" pageNumber="554">Fig. 30B</figureCitation>
). One specimen was found buried under stones.
</paragraph>
</subSubSection>
</treatment>
</document>