448 lines
58 KiB
XML
448 lines
58 KiB
XML
<document id="080C46C23BD9ABB6D7C9BC299470464C" ID-DOI="10.5281/zenodo.278361" ID-GBIF-Dataset="6f260d4c-919b-406a-a811-3733fa76ea14" ID-ISSN="1175-5326" ID-Zenodo-Dep="278361" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460333061231" checkinUser="plazi" docAuthor="Huang, Hao, He, Yang & Shi, Zheng" docDate="2011" docId="03E55A29DA1BFFD2FF66F9EB9C704F38" docLanguage="en" docName="zt02987p064.pdf" docOrigin="Zootaxa 2987" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Lucanus cheni Huang, new species" docType="treatment" docVersion="8" lastPageNumber="59" masterDocId="FFDC2251DA1BFFD1FFF1FFC09E4F4C2C" masterDocTitle="Description of two new species of Lucanus Scopoli (Coleoptera: Scarabaeoidea: Lucanidae) from southeastern Tibet, China" masterLastPageNumber="64" masterPageNumber="56" pageNumber="56" updateTime="1698259095457" updateUser="plazi">
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<mods:mods id="82A1277FE0D1E8DCCE6ECBAEEF06F72A" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="DAFA45DB30551957490D704DE7D406AC">
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<mods:title id="ACF16CDC404CD582C14307CF7956721C">Description of two new species of Lucanus Scopoli (Coleoptera: Scarabaeoidea: Lucanidae) from southeastern Tibet, China</mods:title>
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</mods:titleInfo>
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<mods:name id="6A3A93CD2A12B6E1CDACA31728A17BCF" type="personal">
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<mods:role id="417F91B32C1024B11E93EC14E1C2F2B2">
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<mods:roleTerm id="3A9E1E0F4B480161D52902FC9C6F1F20">Author</mods:roleTerm>
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<mods:namePart id="A4327209FF62FF3FB91785131454DF7E">Huang, Hao</mods:namePart>
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</mods:name>
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<mods:name id="97BC6D475F3A0D1ACD4343623F878D6B" type="personal">
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<mods:roleTerm id="5E828CACAC8E4696D0EACD9FE366F1AB">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="3A17817CF920778360A7862A52C8D170">He, Yang</mods:namePart>
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</mods:name>
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<mods:name id="D21CCF2F4C17D11132BFE7F824E7FF78" type="personal">
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<mods:roleTerm id="B82F6B620915A353F02EDBA38F2BAE1F">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="541E8126F1D0E0E824B924F5C02D3B28">Shi, Zheng</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="3966272B0FEAC3E85A1CE2D18032255D">text</mods:typeOfResource>
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<mods:titleInfo id="43578C31F03F45B98021E7B8A4238CBC">
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<mods:title id="4CBFD034E7DF6FCE44937DCAB25145D8">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="68BA5E19B7A0E6162B5F5DA31C52AE97">
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<mods:date id="F05BC14E1055DC618928194B494AE100">2011</mods:date>
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<mods:detail id="7CF34B7A298B6D1175EBBD06A6EF9FBC" type="volume">
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<mods:number id="F3C54961BCCD11FC3DF544EC56D3106E">2987</mods:number>
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<mods:start id="7D2DF3DE11C9454C051CF881B1D38CF5">56</mods:start>
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<mods:classification id="C3A9D6489B110F44BE0FFCF94DB7785B">journal article</mods:classification>
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<mods:identifier id="722284CE90A2EB587414DBE6DDCAE271" type="DOI">10.5281/zenodo.278361</mods:identifier>
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<mods:identifier id="06C5D3B97E3675C57359D5DFCDFC331B" type="GBIF-Dataset">6f260d4c-919b-406a-a811-3733fa76ea14</mods:identifier>
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<mods:identifier id="D35ACE5746103CC5CB1D905FDA72B598" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="CBEB13B59909A32297804E5F50EB6B4F" type="Zenodo-Dep">278361</mods:identifier>
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<treatment id="03E55A29DA1BFFD2FF66F9EB9C704F38" ID-DOI="http://doi.org/10.5281/zenodo.6194825" ID-GBIF-Taxon="119404770" ID-Zenodo-Dep="6194825" LSID="urn:lsid:plazi:treatment:03E55A29DA1BFFD2FF66F9EB9C704F38" httpUri="http://treatment.plazi.org/id/03E55A29DA1BFFD2FF66F9EB9C704F38" lastPageId="3" lastPageNumber="59" pageId="0" pageNumber="56">
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<subSubSection id="C356B8B4DA1BFFD1FF66F9EB9C714A69" box="[151,574,1579,1605]" pageId="0" pageNumber="56" type="nomenclature">
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<paragraph id="8BF3EB3FDA1BFFD1FF66F9EB9C714A69" blockId="0.[151,574,1579,1605]" box="[151,574,1579,1605]" pageId="0" pageNumber="56">
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<heading id="D0BB5C53DA1BFFD1FF66F9EB9C714A69" bold="true" box="[151,574,1579,1605]" fontSize="11" level="1" pageId="0" pageNumber="56" reason="1">
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<emphasis id="B938372DDA1BFFD1FF66F9EB9C714A69" bold="true" box="[151,574,1579,1605]" pageId="0" pageNumber="56">
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||
<taxonomicName id="4C4C90BCDA1BFFD1FF66F9EB9FE84A69" ID-CoL="72P4Z" authority="Huang" authorityName="Huang" box="[151,423,1579,1605]" class="Insecta" family="Lucanidae" genus="Lucanus" kingdom="Animalia" order="Coleoptera" pageId="0" pageNumber="56" phylum="Arthropoda" rank="species" species="cheni" status="sp. nov.">
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<emphasis id="B938372DDA1BFFD1FF66F9EB9F044A69" bold="true" box="[151,331,1579,1605]" italics="true" pageId="0" pageNumber="56">Lucanus cheni</emphasis>
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Huang
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</taxonomicName>
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<taxonomicNameLabel id="A20B8A56DA1BFFD1FE5FF9EB9C714A69" box="[430,574,1579,1605]" pageId="0" pageNumber="56" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C356B8B4DA1BFFD1FF66F9B39E9C4BAA" pageId="0" pageNumber="56" type="reference_group">
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<paragraph id="8BF3EB3FDA1BFFD1FF66F9B39E9C4BAA" blockId="0.[151,1436,1650,2034]" pageId="0" pageNumber="56">
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<taxonomicName id="4C4C90BCDA1BFFD1FF66F9B39C594AA6" ID-CoL="7X2LP" authority="Arrow, 1950: 46" authorityName="Arrow" authorityPageNumber="46" authorityYear="1950" box="[151,534,1650,1674]" class="Insecta" family="Lucanidae" genus="Lucanus" kingdom="Animalia" order="Coleoptera" pageId="0" pageNumber="56" phylum="Arthropoda" rank="species" species="furcifer">
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<emphasis id="B938372DDA1BFFD1FF66F9B39F1E4AA6" box="[151,337,1650,1674]" italics="true" pageId="0" pageNumber="56">Lucanus furcifer</emphasis>
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Arrow, 1950: 46
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</taxonomicName>
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, partim [specimens from Sikkim (Lachen Lachung)], plate 3,
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<figureCitation id="1377F7BADA1BFFD1FB24F9B29B554AA6" box="[1237,1306,1650,1674]" captionStart="FIGURES 4 – 5" captionStartId="1.[151,264,1641,1664]" captionText="FIGURES 4 – 5. Lucanus cheni. 4 — male genitalia (holotype); 5 — female genitalia (paratype, same specimen as in Fig. 2). Scale bars for Figs. 4 and 5: 1 mm. Abbreviations: lt, last abdominal tergite; lv, last abdominal ventrite; v 9, ventral plate of 9 th abdominal segment; d 9, dorsal and lateral plates of 9 th abdominal segment; ad, aedeagus in dorsal view; al, aedeagus in lateral view; dp, dorsal pouch of penis; pl, penis in lateral view; pv, penis in ventral view; f, flagellum; af, apex of flagellum; sfv, sclerotized female genitalia in ventral view; sfd, sclerotized female genitalia in dorsal view; sg, spermathecal gland; s, spermatheca; sd, spermathecal duct; ag, accessory gland; mo, median oviduct." pageId="0" pageNumber="56">Fig. 4</figureCitation>
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; male from Sikkim;
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<bibRefCitation id="EFDD96CEDA1BFFD1FF07F9569FA54A82" author="Didier" box="[246,490,1686,1710]" pageId="0" pageNumber="56" refString="Didier, R. & Seguy, E. (1953) Note sur le Lucanus singularis Planet et sur les especes voisines. Revue francaise d'entomologie, 20, 28 - 32." type="journal article" year="1953">Didier & Séguy 1953</bibRefCitation>
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: 30,
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<figureCitation id="1377F7BADA1BFFD1FDEEF9569C2F4A82" box="[543,608,1686,1710]" captionStart="FIGURES 1 – 2" captionStartId="1.[151,264,1580,1603]" captionTargetBox="[160,1436,373,1558]" captionTargetId="figure@1.[151,1436,373,1559]" captionTargetPageId="1" captionText="FIGURES 1 – 2. Lucanus cheni habitus in dorsal view. 1 — male holotype; 2 — female paratype." httpUri="https://zenodo.org/record/278362/files/figure.png" pageId="0" pageNumber="56">Fig. 2</figureCitation>
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for male head,
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<figureCitation id="1377F7BADA1BFFD1FCFEF9569D1F4A82" box="[783,848,1686,1710]" captionStart="FIGURES 6 – 9" captionStartId="5.[151,264,1634,1657]" captionTargetBox="[255,1340,202,1610]" captionTargetId="figure@5.[250,1352,193,1625]" captionTargetPageId="5" captionText="FIGURES 6 – 9. Lucanus langi habitus in dorsal view. 6 — male holotype; 7, 8 — male paratypes; 9 — female paratype. FIGURES 10 – 13. Male head in dorsal or anterodorsal view to show clypeolabrum and details of mandibles. 10 — Lucanus cheni holotype; 11 — Lucanus langi paratype; 12 — Lucanus langi paratype (same specimen as in Fig. 8); 13 — Lucanus smithii (Kurseong, India, photo by A. Galant). Abbreviations: s, basal part of mandible swollen and serrate on inner margin; b, clypeolabrum bifurcate at tip; nb, clypeolabrum non-bifurcate; ns, basal part of mandible non-serrate and not swollen. FIGURES 14 – 15. Lucanus langi, part of habitus in ventral view. 14 — male paratype (same specimen as in Fig. 7); 15 — female paratype (same specimen as in Fig. 9)." httpUri="https://zenodo.org/record/278363/files/figure.png" pageId="0" pageNumber="56">Fig. 6</figureCitation>
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for male “androcotype”;
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<bibRefCitation id="EFDD96CEDA1BFFD1FB9FF9569B404A82" author="Bacchus" box="[1134,1295,1686,1710]" pageId="0" pageNumber="56" refString="Bacchus, M. E. (1978) A catalogue of the type-specimens of the Scarabaeinae (Scarabaeidae) and the smaller lamellicorn families (Coleoptera) described by G. J. Arrow. Bulletin of the British Museum (Natural History) B. Entomology, 37 (3), 97 - 115." type="journal article" year="1978">Bacchus 1978</bibRefCitation>
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: 109, part of
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<typeStatus id="54F7559DDA1BFFD1FF66F97A9F734AFE" box="[151,316,1722,1746]" pageId="0" pageNumber="56" type="paralectotype">paralectotypes</typeStatus>
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,
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<specimenCount id="9D4A20B6DA1BFFD1FEB6F97A9FDD4AFE" box="[327,402,1722,1746]" pageId="0" pageNumber="56" type="male">1 male</specimenCount>
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||
from “Sikkim, Lachen Lochung, viii (MNHN, Paris)”;
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<bibRefCitation id="EFDD96CEDA1BFFD1FBF2F97B9ADE4AFE" author="Huang" box="[1027,1169,1722,1747]" pageId="0" pageNumber="56" refString="Huang, H. (2006) New descriptions and notes on Chinese stag-beetles, with discovery of the second species of Noseolucanus from SE Tibet. Coleoptera, 10, 11 - 34." type="journal article" year="2006">Huang 2006</bibRefCitation>
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||
: 14, 18, Hanmi, Motuo, southeastern Tibet;
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||
<bibRefCitation id="EFDD96CEDA1BFFD1FE83F91F9C114ADA" author="Huang" box="[370,606,1758,1783]" pageId="0" pageNumber="56" refString="Huang, H. & Chen, C. - C. (2010) Stag Beetles of China 1. Formosan Biology Company, Taipei, 288 pp." type="book" year="2010">Huang & Chen 2010</bibRefCitation>
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||
: 156, figures for habitus, maxilla, labium and clypeolabrum of both sexes, male and female genitalia. (Taxonomic change according to the
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||
<typeStatus id="54F7559DDA1BFFD1FC94F8C29D834B36" box="[869,972,1794,1818]" pageId="0" pageNumber="56" type="lectotype">lectotype</typeStatus>
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||
designation of
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||
<bibRefCitation id="EFDD96CEDA1BFFD1FB88F8C29B634B36" author="Bacchus" box="[1145,1324,1794,1818]" pageId="0" pageNumber="56" refString="Bacchus, M. E. (1978) A catalogue of the type-specimens of the Scarabaeinae (Scarabaeidae) and the smaller lamellicorn families (Coleoptera) described by G. J. Arrow. Bulletin of the British Museum (Natural History) B. Entomology, 37 (3), 97 - 115." type="journal article" year="1978">Bacchus [1978]</bibRefCitation>
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||
).
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<taxonomicName id="4C4C90BCDA1BFFD1FF66F8E79C2F4B12" ID-CoL="3WB4B" authority="Lacroix, 1971: 566" authorityName="Lacroix" authorityPageNumber="566" authorityYear="1971" box="[151,608,1830,1854]" class="Insecta" family="Lucanidae" genus="Lucanus" kingdom="Animalia" order="Coleoptera" pageId="0" pageNumber="56" phylum="Arthropoda" rank="species" species="singularis">
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<emphasis id="B938372DDA1BFFD1FF66F8E79F3F4B12" box="[151,368,1830,1854]" italics="true" pageId="0" pageNumber="56">Lucanus singularis</emphasis>
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:
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<bibRefCitation id="EFDD96CEDA1BFFD1FE70F8E69C6A4B12" author="Lacroix" box="[385,549,1830,1854]" pageId="0" pageNumber="56" refString="Lacroix, J. P. (1971) Revision des Lucanus du groupe de lunifer. Annales de la Societe Entomologique de France (New Series), 7 (3), 553 - 570." type="journal article" year="1971">Lacroix, 1971</bibRefCitation>
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: 566
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</taxonomicName>
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, partim [“
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<typeStatus id="54F7559DDA1BFFD1FD23F8E69D744B12" box="[722,827,1830,1854]" pageId="0" pageNumber="56" type="paratype">paratype</typeStatus>
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||
:
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<specimenCount id="9D4A20B6DA1BFFD1FCB9F8E69DD64B12" box="[840,921,1830,1854]" pageId="0" pageNumber="56" type="male">1 male</specimenCount>
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, forme mineure, region thibetaine, Lacheng- Lachung”],
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<figureCitation id="1377F7BADA1BFFD1FEECF88A9F104B4E" box="[285,351,1866,1890]" captionStart="FIGURES 6 – 9" captionStartId="5.[151,264,1634,1657]" captionTargetBox="[255,1340,202,1610]" captionTargetId="figure@5.[250,1352,193,1625]" captionTargetPageId="5" captionText="FIGURES 6 – 9. Lucanus langi habitus in dorsal view. 6 — male holotype; 7, 8 — male paratypes; 9 — female paratype. FIGURES 10 – 13. Male head in dorsal or anterodorsal view to show clypeolabrum and details of mandibles. 10 — Lucanus cheni holotype; 11 — Lucanus langi paratype; 12 — Lucanus langi paratype (same specimen as in Fig. 8); 13 — Lucanus smithii (Kurseong, India, photo by A. Galant). Abbreviations: s, basal part of mandible swollen and serrate on inner margin; b, clypeolabrum bifurcate at tip; nb, clypeolabrum non-bifurcate; ns, basal part of mandible non-serrate and not swollen. FIGURES 14 – 15. Lucanus langi, part of habitus in ventral view. 14 — male paratype (same specimen as in Fig. 7); 15 — female paratype (same specimen as in Fig. 9)." httpUri="https://zenodo.org/record/278363/files/figure.png" pageId="0" pageNumber="56">Fig. 8</figureCitation>
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for male;
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<bibRefCitation id="EFDD96CEDA1BFFD1FE25F88A9C3E4B4E" author="Schenk" box="[468,625,1866,1890]" pageId="0" pageNumber="56" refString="Schenk, K. - D. (2009) Contribution to the knowledge of the stag beetles of Asia and description of several new taxa (2). Beetles World, 4, 2 - 16." type="journal article" year="2009">Schenk, 2009</bibRefCitation>
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||
: 7,
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<figureCitation id="1377F7BADA1BFFD1FD69F88A9C9D4B4E" box="[664,722,1866,1890]" captionStart="FIGURES 1 – 2" captionStartId="1.[151,264,1580,1603]" captionTargetBox="[160,1436,373,1558]" captionTargetId="figure@1.[151,1436,373,1559]" captionTargetPageId="1" captionText="FIGURES 1 – 2. Lucanus cheni habitus in dorsal view. 1 — male holotype; 2 — female paratype." httpUri="https://zenodo.org/record/278362/files/figure.png" pageId="0" pageNumber="56">Fig.1</figureCitation>
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[male from Sikkim],
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||
<figureCitation id="1377F7BADA1BFFD1FC36F88A9A464B4E" box="[967,1033,1866,1890]" captionStart="FIGURES 1 – 2" captionStartId="1.[151,264,1580,1603]" captionTargetBox="[160,1436,373,1558]" captionTargetId="figure@1.[151,1436,373,1559]" captionTargetPageId="1" captionText="FIGURES 1 – 2. Lucanus cheni habitus in dorsal view. 1 — male holotype; 2 — female paratype." httpUri="https://zenodo.org/record/278362/files/figure.png" pageId="0" pageNumber="56">Fig. 2</figureCitation>
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||
[male from
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<collectingCountry id="F35BABAFDA1BFFD1FB67F88A9AA54B4E" box="[1174,1258,1866,1890]" name="Bhutan" pageId="0" pageNumber="56">Bhutan</collectingCountry>
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] (misidentification).
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||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C356B8B4DA1BFFD3FF36F8529EAB4E3B" lastPageId="2" lastPageNumber="58" pageId="0" pageNumber="56" type="materials_examined">
|
||
<paragraph id="8BF3EB3FDA1BFFD0FF36F8529C2D4D13" blockId="0.[151,1436,1650,2034]" lastBlockId="1.[151,1436,151,320]" lastPageId="1" lastPageNumber="57" pageId="0" pageNumber="56">
|
||
<emphasis id="B938372DDA1BFFD1FF36F8529D944B86" bold="true" box="[199,987,1937,1962]" pageId="0" pageNumber="56">
|
||
<typeStatus id="54F7559DDA1BFFD1FF36F8529F4F4B86" box="[199,256,1938,1962]" pageId="0" pageNumber="56">Type</typeStatus>
|
||
material.
|
||
<typeStatus id="54F7559DDA1BFFD1FE76F8529FBB4B86" box="[391,500,1938,1962]" pageId="0" pageNumber="56" type="holotype">Holotype</typeStatus>
|
||
(
|
||
<figureCitation id="1377F7BADA1BFFD1FDF9F8519C124B86" box="[520,605,1937,1962]" captionStart="FIGURES 1 – 2" captionStartId="1.[151,264,1580,1603]" captionTargetBox="[160,1436,373,1558]" captionTargetId="figure@1.[151,1436,373,1559]" captionTargetPageId="1" captionText="FIGURES 1 – 2. Lucanus cheni habitus in dorsal view. 1 — male holotype; 2 — female paratype." httpUri="https://zenodo.org/record/278362/files/figure.png" pageId="0" pageNumber="56">Figs. 1</figureCitation>
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||
, 10):
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||
<collectingCountry id="F35BABAFDA1BFFD1FD5DF8519D424B86" box="[684,781,1937,1962]" name="China" pageId="0" pageNumber="56">CHINA</collectingCountry>
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||
: Xizang (Tibet):
|
||
</emphasis>
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||
3, Linzhi Prefecture, Motuo County, Hanmi,
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||
<quantity id="4CB446DADA1BFFD1FF00F8769F0D4BE2" box="[241,322,1974,1998]" metricMagnitude="3" metricUnit="m" metricValue="2.15" pageId="0" pageNumber="56" unit="m" value="2150.0">2150m</quantity>
|
||
,
|
||
<date id="FFF2CDFFDA1BFFD1FEBDF8769FAE4BE2" box="[332,481,1974,1998]" pageId="0" pageNumber="56" value="2005-08-17">17.VIII.2005</date>
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||
, H. Huang leg.; deposited in the entomological collection of Shanghai Entomological Museum, Chinese Academy of Science, Shanghai,
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||
<collectingCountry id="F35BABAFDA1BFFD1FD10F81A9D674BDE" box="[737,808,2010,2034]" name="China" pageId="0" pageNumber="56">China</collectingCountry>
|
||
.
|
||
<emphasis id="B938372DDA1BFFD1FCC5F81A9AC44BDE" bold="true" box="[820,1163,2009,2034]" pageId="0" pageNumber="56">
|
||
<typeStatus id="54F7559DDA1BFFD1FCC5F81A9DFF4BDE" box="[820,944,2010,2034]" pageId="0" pageNumber="56" type="paratype">Paratypes</typeStatus>
|
||
:
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||
<collectingCountry id="F35BABAFDA1BFFD1FC4FF8199A504BDE" box="[958,1055,2009,2034]" name="China" pageId="0" pageNumber="56">CHINA</collectingCountry>
|
||
: Xizang:
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||
</emphasis>
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23 3, 20 ƤƤ, Linzhi Prefecture, Motuo County, Hanmi,
|
||
<quantity id="4CB446DADA1AFFD0FDD0FF579C204C83" box="[545,623,151,175]" metricMagnitude="3" metricUnit="m" metricValue="2.15" pageId="1" pageNumber="57" unit="m" value="2150.0">2150m</quantity>
|
||
,
|
||
<date id="FFF2CDFFDA1AFFD0FD8DFF579D0E4C83" box="[636,833,151,175]" pageId="1" pageNumber="57" value="2005-08-11" valueMax="2005-08-27" valueMin="2005-08-11">11–27.VIII. 2005</date>
|
||
, H. Huang, L. Tang & D.-K. Zhou leg.; 2
|
||
<typeStatus id="54F7559DDA1AFFD0FAE2FF589B314C9C" box="[1299,1406,152,176]" pageId="1" pageNumber="57" type="paratype">paratypes</typeStatus>
|
||
(1 3, 1 Ƥ) in Natural History Museum, London; 12
|
||
<typeStatus id="54F7559DDA1AFFD0FD25FF7D9D704CF9" box="[724,831,189,213]" pageId="1" pageNumber="57" type="paratype">paratypes</typeStatus>
|
||
(6 3, 6 ƤƤ) in Hao Huang’s collection (Qingdao,
|
||
<collectingCountry id="F35BABAFDA1AFFD0FF66FF1F9E904CDB" box="[151,223,223,247]" name="China" pageId="1" pageNumber="57">China</collectingCountry>
|
||
); 5
|
||
<typeStatus id="54F7559DDA1AFFD0FEF6FF209F3D4CD4" box="[263,370,224,248]" pageId="1" pageNumber="57" type="paratype">paratypes</typeStatus>
|
||
(2 3, 3 ƤƤ) in Chang-Chin Chen’s collection (
|
||
<collectingCountry id="F35BABAFDA1AFFD0FC6FFF1F9DBC4CDB" box="[926,1011,223,247]" name="China" pageId="1" pageNumber="57">Tianjin</collectingCountry>
|
||
,
|
||
<collectingCountry id="F35BABAFDA1AFFD0FC0FFF1F9A094CDB" box="[1022,1094,223,247]" name="China" pageId="1" pageNumber="57">China</collectingCountry>
|
||
); 8
|
||
<typeStatus id="54F7559DDA1AFFD0FB9FFF209A964CD4" box="[1134,1241,224,248]" pageId="1" pageNumber="57" type="paratype">paratypes</typeStatus>
|
||
(4 3, 4 ƤƤ) in the entomological collection of Shanghai Normal University, Shanghai,
|
||
<collectingCountry id="F35BABAFDA1AFFD0FC35FEC49A444D30" box="[964,1035,260,284]" name="China" pageId="1" pageNumber="57">China</collectingCountry>
|
||
; 16
|
||
<typeStatus id="54F7559DDA1AFFD0FBCBFEC59AE94D31" box="[1082,1190,261,285]" pageId="1" pageNumber="57" type="paratype">paratypes</typeStatus>
|
||
(9 3, 7 ƤƤ) in Da- Kang Zhou’s collection (Beijing,
|
||
<collectingCountry id="F35BABAFDA1AFFD0FDFFFEE79C194D13" box="[526,598,295,319]" name="China" pageId="1" pageNumber="57">China</collectingCountry>
|
||
).
|
||
</paragraph>
|
||
<caption id="DF33BBB7DA1AFFD0FF66F9EC9A184A6E" box="[151,1111,1580,1603]" httpUri="https://zenodo.org/record/278362/files/figure.png" pageId="1" pageNumber="57" targetBox="[160,1436,373,1558]" targetPageId="1">
|
||
<paragraph id="8BF3EB3FDA1AFFD0FF66F9EC9A184A6E" blockId="1.[151,1437,1580,1817]" box="[151,1111,1580,1603]" pageId="1" pageNumber="57">
|
||
<emphasis id="B938372DDA1AFFD0FF66F9EC9F754A6D" bold="true" box="[151,314,1580,1603]" pageId="1" pageNumber="57">FIGURES 1–2.</emphasis>
|
||
<taxonomicName id="4C4C90BCDA1AFFD0FEB1F9ED9F9C4A6E" box="[320,467,1580,1602]" class="Insecta" family="Lucanidae" genus="Lucanus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="57" phylum="Arthropoda" rank="species" species="cheni">
|
||
<emphasis id="B938372DDA1AFFD0FEB1F9ED9F9C4A6E" box="[320,467,1580,1602]" italics="true" pageId="1" pageNumber="57">Lucanus cheni</emphasis>
|
||
</taxonomicName>
|
||
habitus in dorsal view. 1—male holotype; 2—female paratype.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF33BBB7DA1AFFD0FF66F98A9D9E4A4D" box="[151,977,1610,1633]" pageId="1" pageNumber="57">
|
||
<paragraph id="8BF3EB3FDA1AFFD0FF66F98A9D9E4A4D" blockId="1.[151,1437,1580,1817]" box="[151,977,1610,1633]" pageId="1" pageNumber="57">
|
||
<emphasis id="B938372DDA1AFFD0FF66F98A9F5D4A4C" bold="true" box="[151,274,1610,1633]" pageId="1" pageNumber="57">FIGURE 3.</emphasis>
|
||
<taxonomicName id="4C4C90BCDA1AFFD0FEE8F98C9C7F4A4D" box="[281,560,1611,1633]" class="Insecta" family="Lucanidae" genus="Lucanus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="57" phylum="Arthropoda" rank="subSpecies" species="thibetanus" subSpecies="furcifer">
|
||
<emphasis id="B938372DDA1AFFD0FEE8F98C9C7F4A4D" box="[281,560,1611,1633]" italics="true" pageId="1" pageNumber="57">Lucanus thibetanus furcifer</emphasis>
|
||
</taxonomicName>
|
||
habitus of male lectotype in dorsal view.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF33BBB7DA1AFFD0FF66F9A99D1D4B35" pageId="1" pageNumber="57">
|
||
<paragraph id="8BF3EB3FDA1AFFD0FF66F9A99D1D4B35" blockId="1.[151,1437,1580,1817]" pageId="1" pageNumber="57">
|
||
<emphasis id="B938372DDA1AFFD0FF66F9A99F724A53" bold="true" box="[151,317,1641,1664]" pageId="1" pageNumber="57">FIGURES 4–5.</emphasis>
|
||
<taxonomicName id="4C4C90BCDA1AFFD0FEB6F9AB9F924AAC" box="[327,477,1642,1664]" class="Insecta" family="Lucanidae" genus="Lucanus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="57" phylum="Arthropoda" rank="species" species="cheni">
|
||
<emphasis id="B938372DDA1AFFD0FEB6F9AB9F924AAC" box="[327,477,1642,1664]" italics="true" pageId="1" pageNumber="57">Lucanus cheni</emphasis>
|
||
</taxonomicName>
|
||
. 4—male genitalia (holotype); 5—female genitalia (paratype, same specimen as in Fig. 2). Scale bars for Figs. 4 and 5: 1 mm. Abbreviations: lt, last abdominal tergite; lv, last abdominal ventrite; v9, ventral plate of 9th abdominal segment; d9, dorsal and lateral plates of 9th abdominal segment; ad, aedeagus in dorsal view; al, aedeagus in lateral view; dp, dorsal pouch of penis; pl, penis in lateral view; pv, penis in ventral view; f, flagellum; af, apex of flagellum; sfv, sclerotized female genitalia in ventral view; sfd, sclerotized female genitalia in dorsal view; sg, spermathecal gland; s, spermatheca; sd, spermathecal duct; ag, accessory gland; mo, median oviduct.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BF3EB3FDA1AFFD0FF36F8869A154BC2" blockId="1.[151,1436,1861,2030]" pageId="1" pageNumber="57">
|
||
<emphasis id="B938372DDA1AFFD0FF36F8869F884B72" bold="true" box="[199,455,1861,1886]" pageId="1" pageNumber="57">
|
||
<typeStatus id="54F7559DDA1AFFD0FF36F8869F7B4B72" box="[199,308,1862,1886]" pageId="1" pageNumber="57" type="holotype">Holotype</typeStatus>
|
||
description.
|
||
</emphasis>
|
||
Body length measured from apex of mandible to terminal tip of elytra:
|
||
<quantity id="4CB446DADA1AFFD0FB1BF8869B734B72" box="[1258,1340,1862,1886]" metricMagnitude="-2" metricUnit="m" metricValue="6.1" pageId="1" pageNumber="57" unit="mm" value="61.0">61 mm</quantity>
|
||
.
|
||
<emphasis id="B938372DDA1AFFD0FF34F8A99F934BAE" bold="true" box="[197,476,1897,1922]" pageId="1" pageNumber="57">Color and pubescence:</emphasis>
|
||
Head, pronotum and elytra blackish brown, densely clad with yellow pubescence. Ventral surface of head, thorax and abdominal ventrites blackish brown, densely clad with yellow pubescence; pubescence on metasternum markedly longer than on other areas. All femora and tarsi on both dorsal and ventral surfaces blackish brown. All tibiae on both dorsal and ventral surfaces reddish brown.
|
||
</paragraph>
|
||
<paragraph id="8BF3EB3FDA19FFD3FF34FF589BD34D80" blockId="2.[151,1437,151,2013]" pageId="2" pageNumber="58">
|
||
<emphasis id="B938372DDA19FFD3FF34FF589F414C9C" bold="true" box="[197,270,152,176]" pageId="2" pageNumber="58">Head:</emphasis>
|
||
Frontal ridge clearly defined, more elevated in the middle. Lateral and posterior ridges evenly convex and rounded. Clypeolabrum (
|
||
<emphasis id="B938372DDA19FFD3FE2EFF7D9C524CF8" box="[479,541,189,212]" italics="true" pageId="2" pageNumber="58">sensu</emphasis>
|
||
<bibRefCitation id="EFDD96CEDA19FFD3FDD5FF7C9CF14CF8" author="Gravely" box="[548,702,188,212]" pageId="2" pageNumber="58" refString="Gravely, F. H. (1915) A catalogue of the Lucanidae in the collection of the Indian Museum. Records of the Indian Museum, 11, 407 - 431." type="journal article" year="1915">Gravely 1915</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFDD96CEDA19FFD3FD3BFF7D9DFC4CF8" author="Huang" box="[714,947,188,213]" pageId="2" pageNumber="58" refString="Huang, H. & Chen, C. - C. (2010) Stag Beetles of China 1. Formosan Biology Company, Taipei, 288 pp." type="book" year="2010">Huang & Chen 2010</bibRefCitation>
|
||
) fused with frons, not defined by transverse suture, about 4 times as long as wide, with dorsal branch near apex; dorsal branch bifurcate at tip. Mandible about 3 times as long as head, strongly incurved at basal 1/3, with widely opened apical fork formed by apex and subapical ventral tooth; basal part swollen at the inner margin and serrate; the major inner tooth preceded by 4–6 smaller teeth in continuation to the base of the subapical tooth, and followed by 4–5 smaller teeth that are not continued to the base of the mandible. Antennal club with 4 antennomeres; antennomere 7 slender and sharply pointed at tip; antennomeres 8–10 lamellate. Maxilla with a long setose galea. Ligula bilobed, branches pointed at apex (
|
||
<figureCitation id="1377F7BADA19FFD3FACAFE549BC04D80" box="[1339,1423,404,428]" captionStart="FIGURES 19 – 21" captionStartId="5.[151,264,1972,1995]" captionText="FIGURES 19 – 21. Maxilla and labium of male. 19 — maxilla of Lucanus langi (same specimen as in Fig. 8); 20 — labium of Lucanus langi (same specimen as in Fig. 8); 21 — labium of Lucanus cheni (holotype). Scale bars 1 mm for all." pageId="2" pageNumber="58">Fig. 21</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BF3EB3FDA19FFD3FF34FE789EAB4E3B" blockId="2.[151,1437,151,2013]" pageId="2" pageNumber="58">
|
||
<emphasis id="B938372DDA19FFD3FF34FE789F494DFC" bold="true" box="[197,262,440,464]" pageId="2" pageNumber="58">Legs:</emphasis>
|
||
Protibia with 4–6 distinct teeth along lateral margin; apex bifurcate, acute. Mesotibia with 4–6 lateral spines in addition to terminal spurs and spines. Metatibia with three lateral spines in addition to terminal spurs and spines.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C356B8B4DA19FFD3FF34FDE49EF34A7C" pageId="2" pageNumber="58" type="description">
|
||
<paragraph id="8BF3EB3FDA19FFD3FF34FDE49F7F4F38" blockId="2.[151,1437,151,2013]" pageId="2" pageNumber="58">
|
||
<emphasis id="B938372DDA19FFD3FF34FDE49F9A4E10" bold="true" box="[197,469,547,572]" pageId="2" pageNumber="58">
|
||
Male genitalia (
|
||
<figureCitation id="1377F7BADA19FFD3FE8FFDE39F894E10" box="[382,454,547,572]" captionStart="FIGURES 4 – 5" captionStartId="1.[151,264,1641,1664]" captionText="FIGURES 4 – 5. Lucanus cheni. 4 — male genitalia (holotype); 5 — female genitalia (paratype, same specimen as in Fig. 2). Scale bars for Figs. 4 and 5: 1 mm. Abbreviations: lt, last abdominal tergite; lv, last abdominal ventrite; v 9, ventral plate of 9 th abdominal segment; d 9, dorsal and lateral plates of 9 th abdominal segment; ad, aedeagus in dorsal view; al, aedeagus in lateral view; dp, dorsal pouch of penis; pl, penis in lateral view; pv, penis in ventral view; f, flagellum; af, apex of flagellum; sfv, sclerotized female genitalia in ventral view; sfd, sclerotized female genitalia in dorsal view; sg, spermathecal gland; s, spermatheca; sd, spermathecal duct; ag, accessory gland; mo, median oviduct." pageId="2" pageNumber="58">Fig. 4</figureCitation>
|
||
):
|
||
</emphasis>
|
||
Last abdominal tergite with ill-defined lateral angles. Last abdominal ventrite without apparent membranous area. Ventral plate of 9th abdominal segment with basal part almost even in width, and with thin longitudinal membranous stripe along the midline of the terminal expansion. Aedeagus in dorsal view about 2.30 times as long as wide. Basal piece in dorsal view rather oblong, nearly twice as long as parameres, with pair of sclerotized dorsal plates; ventral plate at caudal end of the basal piece membranous. Paramere with slender basal process. Penis about as long as parameres, with flagellum (permanently everted internal sac) about 5 times as long as parameres.
|
||
</paragraph>
|
||
<paragraph id="8BF3EB3FDA19FFD3FF36FCE09B644F14" blockId="2.[151,1437,151,2013]" box="[199,1323,799,824]" pageId="2" pageNumber="58">
|
||
<emphasis id="B938372DDA19FFD3FF36FCE09FC74F14" bold="true" box="[199,392,800,824]" pageId="2" pageNumber="58">
|
||
Male
|
||
<typeStatus id="54F7559DDA19FFD3FEFAFCE09FCC4F14" box="[267,387,800,824]" pageId="2" pageNumber="58" type="paratype">paratypes</typeStatus>
|
||
.
|
||
</emphasis>
|
||
Body length measured from apex of mandible to terminal tip of elytra:
|
||
<quantity id="4CB446DADA19FFD3FB5AFCDF9B684F14" box="[1195,1319,799,824]" metricMagnitude="-2" metricUnit="m" metricValue="6.15" metricValueMax="8.1" metricValueMin="4.2" pageId="2" pageNumber="58" unit="mm" value="61.5" valueMax="81.0" valueMin="42.0">42–81 mm</quantity>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BF3EB3FDA19FFD3FF36FC839C2D4FC0" blockId="2.[151,1437,151,2013]" pageId="2" pageNumber="58">
|
||
<emphasis id="B938372DDA19FFD3FF36FC839F744F70" bold="true" box="[199,315,835,860]" pageId="2" pageNumber="58">Variation.</emphasis>
|
||
Apical fork of the mandible widely opened in large-sized males, narrowly opened in medium-sized males, absent in small-sized males. Clypeolabrum longer in large-sized males, shorter in medium-sized males, with upper branch not bifurcate at apex in small-sized males. Number of inner teeth on the mandibles varying with specimen size. Frontal ridge of head in small-sized males not elevated medially. Lateral ridges of head in small-sized males not convex laterally or posteriorly.
|
||
</paragraph>
|
||
<paragraph id="8BF3EB3FDA19FFD3FF36FC389BD2483C" blockId="2.[151,1437,151,2013]" box="[199,1437,1015,1040]" pageId="2" pageNumber="58">
|
||
<emphasis id="B938372DDA19FFD3FF36FC389FD5483C" bold="true" box="[199,410,1016,1040]" pageId="2" pageNumber="58">
|
||
Female
|
||
<typeStatus id="54F7559DDA19FFD3FED5FC389FD5483C" box="[292,410,1016,1040]" pageId="2" pageNumber="58" type="paratype">paratypes</typeStatus>
|
||
</emphasis>
|
||
(
|
||
<figureCitation id="1377F7BADA19FFD3FE5BFC379FA04823" box="[426,495,1015,1039]" captionStart="FIGURES 1 – 2" captionStartId="1.[151,264,1580,1603]" captionTargetBox="[160,1436,373,1558]" captionTargetId="figure@1.[151,1436,373,1559]" captionTargetPageId="1" captionText="FIGURES 1 – 2. Lucanus cheni habitus in dorsal view. 1 — male holotype; 2 — female paratype." httpUri="https://zenodo.org/record/278362/files/figure.png" pageId="2" pageNumber="58">Fig. 2</figureCitation>
|
||
). Body length measured from apex of mandible to terminal tip of elytra:
|
||
<quantity id="4CB446DADA19FFD3FAEDFC379BD7483C" box="[1308,1432,1015,1040]" metricMagnitude="-2" metricUnit="m" metricValue="3.45" metricValueMax="4.1" metricValueMin="2.8" pageId="2" pageNumber="58" unit="mm" value="34.5" valueMax="41.0" valueMin="28.0">28–41 mm</quantity>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BF3EB3FDA19FFD3FF34FBDB9D49488C" blockId="2.[151,1437,151,2013]" pageId="2" pageNumber="58">
|
||
<emphasis id="B938372DDA19FFD3FF34FBDB9F9B4818" bold="true" box="[197,468,1051,1076]" pageId="2" pageNumber="58">Color and pubescence:</emphasis>
|
||
All structures except the tibiae black or blackish brown on both dorsal and ventral surfaces. Tibiae varying from entirely black to entirely reddish brown among individuals. Dorsal surface of body entirely glabrous, only sparsely clad with short yellow pubescence along lateral elytral margins in fresh specimens. Ventral surface of body with dense, yellow pubescence.
|
||
</paragraph>
|
||
<paragraph id="8BF3EB3FDA19FFD3FF34FB6C9F4F4978" blockId="2.[151,1437,151,2013]" pageId="2" pageNumber="58">
|
||
<emphasis id="B938372DDA19FFD3FF34FB6C9F4148E8" bold="true" box="[197,270,1196,1220]" pageId="2" pageNumber="58">Head:</emphasis>
|
||
Canthus with both anterior and posterior angles clearly defined; anterior angle outside of eye and about as far as the posterior angle; lateral margin straight. Frontal and lateral ridges obsolete, not defined. Clypeolabrum in dorsal view transverse, flat at tip, not protruding forwards medially. Both left and right mandibles with clear dorsal tooth, 2 widely-separated inner teeth and large gap behind apex; lateral margin between 2 inner teeth concave and long.
|
||
</paragraph>
|
||
<paragraph id="8BF3EB3FDA19FFD3FF34FAA09AC149B0" blockId="2.[151,1437,151,2013]" pageId="2" pageNumber="58">
|
||
<emphasis id="B938372DDA19FFD3FF34FAA09F084954" bold="true" box="[197,327,1376,1400]" pageId="2" pageNumber="58">Pronotum:</emphasis>
|
||
About twice as wide as long, widest behind middle; anterior angle produced; lateral angle clearly defined, pointed; posterior angle obtuse; lateral margin before lateral angle evenly convex.
|
||
</paragraph>
|
||
<paragraph id="8BF3EB3FDA19FFD3FF34FA689EF34A7C" blockId="2.[151,1437,151,2013]" pageId="2" pageNumber="58">
|
||
<emphasis id="B938372DDA19FFD3FF34FA689FBC49EC" bold="true" box="[197,499,1447,1472]" pageId="2" pageNumber="58">
|
||
Female genitalia (
|
||
<figureCitation id="1377F7BADA19FFD3FE6AFA679FAC49EC" box="[411,483,1447,1472]" captionStart="FIGURES 4 – 5" captionStartId="1.[151,264,1641,1664]" captionText="FIGURES 4 – 5. Lucanus cheni. 4 — male genitalia (holotype); 5 — female genitalia (paratype, same specimen as in Fig. 2). Scale bars for Figs. 4 and 5: 1 mm. Abbreviations: lt, last abdominal tergite; lv, last abdominal ventrite; v 9, ventral plate of 9 th abdominal segment; d 9, dorsal and lateral plates of 9 th abdominal segment; ad, aedeagus in dorsal view; al, aedeagus in lateral view; dp, dorsal pouch of penis; pl, penis in lateral view; pv, penis in ventral view; f, flagellum; af, apex of flagellum; sfv, sclerotized female genitalia in ventral view; sfd, sclerotized female genitalia in dorsal view; sg, spermathecal gland; s, spermatheca; sd, spermathecal duct; ag, accessory gland; mo, median oviduct." pageId="2" pageNumber="58">Fig. 5</figureCitation>
|
||
):
|
||
</emphasis>
|
||
Last abdominal tergite with poorly-defined lateral angles. Last abdominal ventrite with large membranous area medially. Hemisternite with outer apex protruding beyond inner apex; the inner lateral margin of sclerotized part short. Spermatheca sclerotized and J-shaped. Spermathecal duct about 2–3 times as long as spermatheca. Spermathecal gland slightly longer than spermatheca. Central conjunction of 9th tergites broad at tip.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C356B8B4DA19FFD3FF36F99C9CA54A58" box="[199,746,1627,1652]" pageId="2" pageNumber="58" type="materials_examined">
|
||
<paragraph id="8BF3EB3FDA19FFD3FF36F99C9CA54A58" blockId="2.[151,1437,151,2013]" box="[199,746,1627,1652]" pageId="2" pageNumber="58">
|
||
<emphasis id="B938372DDA19FFD3FF36F99C9F2A4A58" bold="true" box="[199,357,1627,1652]" pageId="2" pageNumber="58">
|
||
<typeStatus id="54F7559DDA19FFD3FF36F99C9F4D4A58" box="[199,258,1628,1652]" pageId="2" pageNumber="58">Type</typeStatus>
|
||
locality.
|
||
</emphasis>
|
||
<collectingCountry id="F35BABAFDA19FFD3FE9DF99C9FFD4A58" box="[364,434,1628,1652]" name="China" pageId="2" pageNumber="58">China</collectingCountry>
|
||
, southeastern Tibet, Motuo.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C356B8B4DA19FFD3FF36F9409F374A90" pageId="2" pageNumber="58" type="etymology">
|
||
<paragraph id="8BF3EB3FDA19FFD3FF36F9409F374A90" blockId="2.[151,1437,151,2013]" pageId="2" pageNumber="58">
|
||
<emphasis id="B938372DDA19FFD3FF36F9409F014AB4" bold="true" box="[199,334,1664,1688]" pageId="2" pageNumber="58">Etymology.</emphasis>
|
||
This species is dedicated to Mr. Chang-Chin Chen (
|
||
<collectingCountry id="F35BABAFDA19FFD3FC63F9409DA84AB4" box="[914,999,1664,1688]" name="China" pageId="2" pageNumber="58">Tianjin</collectingCountry>
|
||
,
|
||
<collectingCountry id="F35BABAFDA19FFD3FC00F9409A774AB4" box="[1009,1080,1664,1688]" name="China" pageId="2" pageNumber="58">China</collectingCountry>
|
||
), a collaborator and good friend of the senior author.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C356B8B4DA19FFD2FF36F9089C704F38" lastPageId="3" lastPageNumber="59" pageId="2" pageNumber="58" type="discussion">
|
||
<paragraph id="8BF3EB3FDA19FFD3FF36F9089BD94BF0" blockId="2.[151,1437,151,2013]" pageId="2" pageNumber="58">
|
||
<emphasis id="B938372DDA19FFD3FF36F9089F744ACC" bold="true" box="[199,315,1736,1760]" pageId="2" pageNumber="58">Remarks.</emphasis>
|
||
This new species is similar to
|
||
<taxonomicName id="4C4C90BCDA19FFD3FD6FF9099D354ACC" box="[670,890,1736,1760]" class="Insecta" family="Lucanidae" genus="Lucanus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="58" phylum="Arthropoda" rank="species" species="thibetanus">
|
||
<emphasis id="B938372DDA19FFD3FD6FF9099D354ACC" box="[670,890,1736,1760]" italics="true" pageId="2" pageNumber="58">Lucanus thibetanus</emphasis>
|
||
</taxonomicName>
|
||
, but it can be distinguished using the following characters: male mandible with greater number of inner teeth behind the major inner tooth; lateral and posterior ridges of the male head more convex laterally and less convex posteriorly; female clypeolabrum shorter and narrower, with the lateral corners of the clypeus indistinct; anterior angle of the female head usually indistinct; apical fork of the female protibia markedly shorter; size of the aedeagus smaller, with the apical part of the penis concave at the outer lateral margin (the male genitalia of all known subspecies of
|
||
<taxonomicName id="4C4C90BCDA19FFD3FC34F8BD9AD04BB8" box="[965,1183,1916,1940]" class="Insecta" family="Lucanidae" genus="Lucanus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="58" phylum="Arthropoda" rank="species" species="thibetanus">
|
||
<emphasis id="B938372DDA19FFD3FC34F8BD9AD04BB8" box="[965,1183,1916,1940]" italics="true" pageId="2" pageNumber="58">Lucanus thibetanus</emphasis>
|
||
</taxonomicName>
|
||
have been examined); spermathecal duct of female genitalia shorter (all subspecies of
|
||
<taxonomicName id="4C4C90BCDA19FFD3FC77F8619A2A4B94" box="[902,1125,1952,1976]" class="Insecta" family="Lucanidae" genus="Lucanus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="58" phylum="Arthropoda" rank="species" species="thibetanus">
|
||
<emphasis id="B938372DDA19FFD3FC77F8619A2A4B94" box="[902,1125,1952,1976]" italics="true" pageId="2" pageNumber="58">Lucanus thibetanus</emphasis>
|
||
</taxonomicName>
|
||
except
|
||
<taxonomicName id="4C4C90BCDA19FFD3FB34F8619F6F4BF0" authority="Nagai, 2000" authorityName="Nagai" authorityYear="2000" class="Insecta" family="Lucanidae" genus="Lucanus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="58" phylum="Arthropoda" rank="subSpecies" species="thibetanus" subSpecies="isaki">
|
||
<emphasis id="B938372DDA19FFD3FB34F8619A9B4B94" box="[1221,1236,1953,1976]" italics="true" pageId="2" pageNumber="58">L</emphasis>
|
||
.
|
||
<emphasis id="B938372DDA19FFD3FB16F8609BD34B94" box="[1255,1436,1952,1976]" italics="true" pageId="2" pageNumber="58">thibetanus isaki</emphasis>
|
||
Nagai, 2000
|
||
</taxonomicName>
|
||
from northern
|
||
<collectingCountry id="F35BABAFDA19FFD3FE3BF8059C784BF1" box="[458,567,1989,2013]" name="Myanmar" pageId="2" pageNumber="58">Myanmar</collectingCountry>
|
||
have been examined); hemisternite of the female genitalia markedly smaller.
|
||
</paragraph>
|
||
<paragraph id="8BF3EB3FDA18FFD2FF66FF579C8A4D48" blockId="3.[151,1436,151,788]" pageId="3" pageNumber="59">
|
||
This new species can be easily distinguished from the unique female
|
||
<typeStatus id="54F7559DDA18FFD2FC59FF579A454C83" box="[936,1034,151,175]" pageId="3" pageNumber="59" type="holotype">holotype</typeStatus>
|
||
of
|
||
<taxonomicName id="4C4C90BCDA18FFD2FBC1FF599BD34C83" authority="Planet, 1900" authorityName="Planet" authorityYear="1900" box="[1072,1436,151,176]" class="Insecta" family="Lucanidae" genus="Lucanus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="59" phylum="Arthropoda" rank="species" species="singularis">
|
||
<emphasis id="B938372DDA18FFD2FBC1FF599B484C9C" box="[1072,1287,152,176]" italics="true" pageId="3" pageNumber="59">Lucanus singularis</emphasis>
|
||
Planet, 1900
|
||
</taxonomicName>
|
||
by the following characters: clypeolabrum not protruding medially, canthus with the anterior angle clearly defined and outside of the eye. A series of specimens identified as
|
||
<taxonomicName id="4C4C90BCDA18FFD2FCC6FF219A404CD4" box="[823,1039,224,248]" class="Insecta" family="Lucanidae" genus="Lucanus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="59" phylum="Arthropoda" rank="species" species="singularis">
|
||
<emphasis id="B938372DDA18FFD2FCC6FF219A404CD4" box="[823,1039,224,248]" italics="true" pageId="3" pageNumber="59">Lucanus singularis</emphasis>
|
||
</taxonomicName>
|
||
were recently discovered from the Chayu area, southeastern Tibet, which match the most characters of the
|
||
<typeStatus id="54F7559DDA18FFD2FC35FEC49A694D30" box="[964,1062,260,284]" pageId="3" pageNumber="59" type="holotype">holotype</typeStatus>
|
||
of
|
||
<taxonomicName id="4C4C90BCDA18FFD2FBBCFEC59B6B4D30" box="[1101,1316,260,284]" class="Insecta" family="Lucanidae" genus="Lucanus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="59" phylum="Arthropoda" rank="species" species="singularis">
|
||
<emphasis id="B938372DDA18FFD2FBBCFEC59B6B4D30" box="[1101,1316,260,284]" italics="true" pageId="3" pageNumber="59">Lucanus singularis</emphasis>
|
||
</taxonomicName>
|
||
. The male of
|
||
<taxonomicName id="4C4C90BCDA18FFD2FF47FEE99FC34D6C" box="[182,396,296,320]" class="Insecta" family="Lucanidae" genus="Lucanus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="59" phylum="Arthropoda" rank="species" species="singularis">
|
||
<emphasis id="B938372DDA18FFD2FF47FEE99FC34D6C" box="[182,396,296,320]" italics="true" pageId="3" pageNumber="59">Lucanus singularis</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<emphasis id="B938372DDA18FFD2FE6FFEE99F934D6C" box="[414,476,297,320]" italics="true" pageId="3" pageNumber="59">sensu</emphasis>
|
||
<bibRefCitation id="EFDD96CEDA18FFD2FE15FEE89C9E4D13" author="Huang" box="[484,721,295,320]" pageId="3" pageNumber="59" refString="Huang, H. & Chen, C. - C. (2010) Stag Beetles of China 1. Formosan Biology Company, Taipei, 288 pp." type="book" year="2010">Huang & Chen 2010</bibRefCitation>
|
||
) has many differences compared to the male of
|
||
<taxonomicName id="4C4C90BCDA18FFD2FB03FEE99BDB4D6C" box="[1266,1428,296,320]" class="Insecta" family="Lucanidae" genus="Lucanus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="59" phylum="Arthropoda" rank="species" species="cheni">
|
||
<emphasis id="B938372DDA18FFD2FB03FEE99BDB4D6C" box="[1266,1428,296,320]" italics="true" pageId="3" pageNumber="59">Lucanus cheni</emphasis>
|
||
</taxonomicName>
|
||
, such as a much shorter and less dentate mandible.
|
||
</paragraph>
|
||
<paragraph id="8BF3EB3FDA18FFD2FF34FEAF9C704F38" blockId="3.[151,1436,151,788]" pageId="3" pageNumber="59">
|
||
This new species had been identified as
|
||
<taxonomicName id="4C4C90BCDA18FFD2FD65FEB19D014DA4" box="[660,846,368,392]" class="Insecta" family="Lucanidae" genus="Lucanus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="59" phylum="Arthropoda" rank="species" species="furcifer">
|
||
<emphasis id="B938372DDA18FFD2FD65FEB19D014DA4" box="[660,846,368,392]" italics="true" pageId="3" pageNumber="59">Lucanus furcifer</emphasis>
|
||
</taxonomicName>
|
||
until the senior author noticed that
|
||
<bibRefCitation id="EFDD96CEDA18FFD2FB1BFEAF9BD34DAB" author="Bacchus" box="[1258,1436,367,391]" pageId="3" pageNumber="59" refString="Bacchus, M. E. (1978) A catalogue of the type-specimens of the Scarabaeinae (Scarabaeidae) and the smaller lamellicorn families (Coleoptera) described by G. J. Arrow. Bulletin of the British Museum (Natural History) B. Entomology, 37 (3), 97 - 115." type="journal article" year="1978">Bacchus (1978)</bibRefCitation>
|
||
published a
|
||
<typeStatus id="54F7559DDA18FFD2FED2FE549FC54D80" box="[291,394,404,428]" pageId="3" pageNumber="59" type="lectotype">lectotype</typeStatus>
|
||
designation of
|
||
<taxonomicName id="4C4C90BCDA18FFD2FDCCFE559CB74D80" box="[573,760,404,428]" class="Insecta" family="Lucanidae" genus="Lucanus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="59" phylum="Arthropoda" rank="species" species="furcifer">
|
||
<emphasis id="B938372DDA18FFD2FDCCFE559CB74D80" box="[573,760,404,428]" italics="true" pageId="3" pageNumber="59">Lucanus furcifer</emphasis>
|
||
</taxonomicName>
|
||
and selected a male specimen of
|
||
<taxonomicName id="4C4C90BCDA18FFD2FB8EFE559B124D80" box="[1151,1373,404,428]" class="Insecta" family="Lucanidae" genus="Lucanus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="59" phylum="Arthropoda" rank="species" species="thibetanus">
|
||
<emphasis id="B938372DDA18FFD2FB8EFE559B124D80" box="[1151,1373,404,428]" italics="true" pageId="3" pageNumber="59">Lucanus thibetanus</emphasis>
|
||
</taxonomicName>
|
||
from Yunnan as the
|
||
<typeStatus id="54F7559DDA18FFD2FEB0FE779FE74DE3" box="[321,424,439,463]" pageId="3" pageNumber="59" type="lectotype">lectotype</typeStatus>
|
||
of
|
||
<emphasis id="B938372DDA18FFD2FE23FE799CDE4DFC" box="[466,657,440,464]" italics="true" pageId="3" pageNumber="59">
|
||
<taxonomicName id="4C4C90BCDA18FFD2FE23FE799CC14DFC" box="[466,654,440,464]" class="Insecta" family="Lucanidae" genus="Lucanus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="59" phylum="Arthropoda" rank="species" species="furcifer">Lucanus furcifer</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
As fully discussed in the recent publication (
|
||
<bibRefCitation id="EFDD96CEDA18FFD2FB6AFE779BDF4DE3" author="Huang" box="[1179,1424,439,463]" pageId="3" pageNumber="59" refString="Huang, H. & Chen, C. - C. (2010) Stag Beetles of China 1. Formosan Biology Company, Taipei, 288 pp." type="book" year="2010">Huang & Chen 2010</bibRefCitation>
|
||
), Arrow (1950) made the mistake of mixing two species into one taxon: a Sikkim species represented by the figures of the males in Arrow (1950: plate 3,
|
||
<figureCitation id="1377F7BADA18FFD2FDB4FE3F9CC64E3B" box="[581,649,511,535]" captionStart="FIGURES 4 – 5" captionStartId="1.[151,264,1641,1664]" captionText="FIGURES 4 – 5. Lucanus cheni. 4 — male genitalia (holotype); 5 — female genitalia (paratype, same specimen as in Fig. 2). Scale bars for Figs. 4 and 5: 1 mm. Abbreviations: lt, last abdominal tergite; lv, last abdominal ventrite; v 9, ventral plate of 9 th abdominal segment; d 9, dorsal and lateral plates of 9 th abdominal segment; ad, aedeagus in dorsal view; al, aedeagus in lateral view; dp, dorsal pouch of penis; pl, penis in lateral view; pv, penis in ventral view; f, flagellum; af, apex of flagellum; sfv, sclerotized female genitalia in ventral view; sfd, sclerotized female genitalia in dorsal view; sg, spermathecal gland; s, spermatheca; sd, spermathecal duct; ag, accessory gland; mo, median oviduct." pageId="3" pageNumber="59">Fig. 4</figureCitation>
|
||
) and
|
||
<bibRefCitation id="EFDD96CEDA18FFD2FD3BFE3F9A034E3B" author="Didier" box="[714,1100,511,535]" pageId="3" pageNumber="59" refString="Didier, R. & Seguy, E. (1953) Note sur le Lucanus singularis Planet et sur les especes voisines. Revue francaise d'entomologie, 20, 28 - 32." type="journal article" year="1953">
|
||
Didier & Séguy (1953:
|
||
<figureCitation id="1377F7BADA18FFD2FC27FE3F9A6A4E3B" box="[982,1061,511,535]" captionStart="FIGURES 1 – 2" captionStartId="1.[151,264,1580,1603]" captionTargetBox="[160,1436,373,1558]" captionTargetId="figure@1.[151,1436,373,1559]" captionTargetPageId="1" captionText="FIGURES 1 – 2. Lucanus cheni habitus in dorsal view. 1 — male holotype; 2 — female paratype." httpUri="https://zenodo.org/record/278362/files/figure.png" pageId="3" pageNumber="59">Figs. 2</figureCitation>
|
||
,
|
||
<figureCitation id="1377F7BADA18FFD2FBC2FE3F9A0C4E3B" box="[1075,1091,511,535]" captionStart="FIGURES 6 – 9" captionStartId="5.[151,264,1634,1657]" captionTargetBox="[255,1340,202,1610]" captionTargetId="figure@5.[250,1352,193,1625]" captionTargetPageId="5" captionText="FIGURES 6 – 9. Lucanus langi habitus in dorsal view. 6 — male holotype; 7, 8 — male paratypes; 9 — female paratype. FIGURES 10 – 13. Male head in dorsal or anterodorsal view to show clypeolabrum and details of mandibles. 10 — Lucanus cheni holotype; 11 — Lucanus langi paratype; 12 — Lucanus langi paratype (same specimen as in Fig. 8); 13 — Lucanus smithii (Kurseong, India, photo by A. Galant). Abbreviations: s, basal part of mandible swollen and serrate on inner margin; b, clypeolabrum bifurcate at tip; nb, clypeolabrum non-bifurcate; ns, basal part of mandible non-serrate and not swollen. FIGURES 14 – 15. Lucanus langi, part of habitus in ventral view. 14 — male paratype (same specimen as in Fig. 7); 15 — female paratype (same specimen as in Fig. 9)." httpUri="https://zenodo.org/record/278363/files/figure.png" pageId="3" pageNumber="59">6</figureCitation>
|
||
)
|
||
</bibRefCitation>
|
||
, and a subspecies of
|
||
<taxonomicName id="4C4C90BCDA18FFD2FACCFDC19F434E10" class="Insecta" family="Lucanidae" genus="Lucanus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="59" phylum="Arthropoda" rank="species" species="thibetanus">
|
||
<emphasis id="B938372DDA18FFD2FACCFDC19F434E10" italics="true" pageId="3" pageNumber="59">Lucanus thibetanus</emphasis>
|
||
</taxonomicName>
|
||
from Yunnan represented by the figures of the males and females in
|
||
<bibRefCitation id="EFDD96CEDA18FFD2FBFEFDE49B514E10" author="Planet" box="[1039,1310,548,572]" pageId="3" pageNumber="59" refString="Planet, L. (1903) Description d'un Lucanide nouveau. Le Naturaliste, 25, 12 - 13." type="journal article" year="1903">
|
||
Planet (1903:
|
||
<figureCitation id="1377F7BADA18FFD2FB5DFDE49B594E10" box="[1196,1302,548,572]" captionStart="FIGURES 1 – 2" captionStartId="1.[151,264,1580,1603]" captionTargetBox="[160,1436,373,1558]" captionTargetId="figure@1.[151,1436,373,1559]" captionTargetPageId="1" captionText="FIGURES 1 – 2. Lucanus cheni habitus in dorsal view. 1 — male holotype; 2 — female paratype." httpUri="https://zenodo.org/record/278362/files/figure.png" pageId="3" pageNumber="59">Figs. 1, 2</figureCitation>
|
||
)
|
||
</bibRefCitation>
|
||
and
|
||
<bibRefCitation id="EFDD96CEDA18FFD2FAA5FDE49FA14E73" author="Didier" pageId="3" pageNumber="59" refString="Didier, R. & Seguy, E. (1953) Note sur le Lucanus singularis Planet et sur les especes voisines. Revue francaise d'entomologie, 20, 28 - 32." type="journal article" year="1953">
|
||
Didier & Séguy (1953:
|
||
<figureCitation id="1377F7BADA18FFD2FEA6FD879FE74E73" box="[343,424,583,607]" captionStart="FIGURE 3" captionStartId="1.[151,250,1610,1633]" captionText="FIGURE 3. Lucanus thibetanus furcifer habitus of male lectotype in dorsal view." pageId="3" pageNumber="59">Figs. 3</figureCitation>
|
||
,
|
||
<figureCitation id="1377F7BADA18FFD2FE47FD879F894E73" box="[438,454,583,607]" captionStart="FIGURES 4 – 5" captionStartId="1.[151,264,1641,1664]" captionText="FIGURES 4 – 5. Lucanus cheni. 4 — male genitalia (holotype); 5 — female genitalia (paratype, same specimen as in Fig. 2). Scale bars for Figs. 4 and 5: 1 mm. Abbreviations: lt, last abdominal tergite; lv, last abdominal ventrite; v 9, ventral plate of 9 th abdominal segment; d 9, dorsal and lateral plates of 9 th abdominal segment; ad, aedeagus in dorsal view; al, aedeagus in lateral view; dp, dorsal pouch of penis; pl, penis in lateral view; pv, penis in ventral view; f, flagellum; af, apex of flagellum; sfv, sclerotized female genitalia in ventral view; sfd, sclerotized female genitalia in dorsal view; sg, spermathecal gland; s, spermatheca; sd, spermathecal duct; ag, accessory gland; mo, median oviduct." pageId="3" pageNumber="59">5</figureCitation>
|
||
,
|
||
<figureCitation id="1377F7BADA18FFD2FE25FD879FAB4E73" box="[468,484,583,607]" captionStart="FIGURES 6 – 9" captionStartId="5.[151,264,1634,1657]" captionTargetBox="[255,1340,202,1610]" captionTargetId="figure@5.[250,1352,193,1625]" captionTargetPageId="5" captionText="FIGURES 6 – 9. Lucanus langi habitus in dorsal view. 6 — male holotype; 7, 8 — male paratypes; 9 — female paratype. FIGURES 10 – 13. Male head in dorsal or anterodorsal view to show clypeolabrum and details of mandibles. 10 — Lucanus cheni holotype; 11 — Lucanus langi paratype; 12 — Lucanus langi paratype (same specimen as in Fig. 8); 13 — Lucanus smithii (Kurseong, India, photo by A. Galant). Abbreviations: s, basal part of mandible swollen and serrate on inner margin; b, clypeolabrum bifurcate at tip; nb, clypeolabrum non-bifurcate; ns, basal part of mandible non-serrate and not swollen. FIGURES 14 – 15. Lucanus langi, part of habitus in ventral view. 14 — male paratype (same specimen as in Fig. 7); 15 — female paratype (same specimen as in Fig. 9)." httpUri="https://zenodo.org/record/278363/files/figure.png" pageId="3" pageNumber="59">7</figureCitation>
|
||
)
|
||
</bibRefCitation>
|
||
. Since
|
||
<bibRefCitation id="EFDD96CEDA18FFD2FDB2FD879CBA4E73" author="Bacchus" box="[579,757,583,607]" pageId="3" pageNumber="59" refString="Bacchus, M. E. (1978) A catalogue of the type-specimens of the Scarabaeinae (Scarabaeidae) and the smaller lamellicorn families (Coleoptera) described by G. J. Arrow. Bulletin of the British Museum (Natural History) B. Entomology, 37 (3), 97 - 115." type="journal article" year="1978">Bacchus (1978)</bibRefCitation>
|
||
selected a male specimen from Yunnan as the
|
||
<typeStatus id="54F7559DDA18FFD2FAE4FD879B344E73" box="[1301,1403,583,607]" pageId="3" pageNumber="59" type="lectotype">lectotype</typeStatus>
|
||
of
|
||
<taxonomicName id="4C4C90BCDA18FFD2FF66FDAD9F1E4EA8" box="[151,337,620,644]" class="Insecta" family="Lucanidae" genus="Lucanus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="59" phylum="Arthropoda" rank="species" species="furcifer">
|
||
<emphasis id="B938372DDA18FFD2FF66FDAD9F1E4EA8" box="[151,337,620,644]" italics="true" pageId="3" pageNumber="59">Lucanus furcifer</emphasis>
|
||
</taxonomicName>
|
||
, the Sikkim species is an unnamed species, and
|
||
<taxonomicName id="4C4C90BCDA18FFD2FC77FDAD9A0E4EA8" box="[902,1089,620,644]" class="Insecta" family="Lucanidae" genus="Lucanus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="59" phylum="Arthropoda" rank="species" species="furcifer">
|
||
<emphasis id="B938372DDA18FFD2FC77FDAD9A0E4EA8" box="[902,1089,620,644]" italics="true" pageId="3" pageNumber="59">Lucanus furcifer</emphasis>
|
||
</taxonomicName>
|
||
now refers to a subspecies of
|
||
<taxonomicName id="4C4C90BCDA18FFD2FF66FD519F3E4E84" box="[151,369,656,680]" class="Insecta" family="Lucanidae" genus="Lucanus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="59" phylum="Arthropoda" rank="species" species="thibetanus">
|
||
<emphasis id="B938372DDA18FFD2FF66FD519F3E4E84" box="[151,369,656,680]" italics="true" pageId="3" pageNumber="59">Lucanus thibetanus</emphasis>
|
||
</taxonomicName>
|
||
. The
|
||
<typeStatus id="54F7559DDA18FFD2FE41FD4F9C584E8B" box="[432,535,655,679]" pageId="3" pageNumber="59" type="lectotype">lectotype</typeStatus>
|
||
of
|
||
<taxonomicName id="4C4C90BCDA18FFD2FDCAFD519CBC4E84" box="[571,755,656,680]" class="Insecta" family="Lucanidae" genus="Lucanus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="59" phylum="Arthropoda" rank="species" species="furcifer">
|
||
<emphasis id="B938372DDA18FFD2FDCAFD519CBC4E84" box="[571,755,656,680]" italics="true" pageId="3" pageNumber="59">Lucanus furcifer</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="1377F7BADA18FFD2FCF3FD4F9D0B4E8B" box="[770,836,655,679]" captionStart="FIGURE 3" captionStartId="1.[151,250,1610,1633]" captionText="FIGURE 3. Lucanus thibetanus furcifer habitus of male lectotype in dorsal view." pageId="3" pageNumber="59">Fig. 3</figureCitation>
|
||
) matches all the diagnostic characters of
|
||
<taxonomicName id="4C4C90BCDA18FFD2FAE1FD519C884EE0" authority="Didier & Seguy, 1953" authorityName="Didier & Seguy" authorityYear="1953" class="Insecta" family="Lucanidae" genus="Lucanus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="59" phylum="Arthropoda" rank="subSpecies" species="thibetanus" subSpecies="pseudosingularis">
|
||
<emphasis id="B938372DDA18FFD2FAE1FD519FF74EE0" italics="true" pageId="3" pageNumber="59">Lucanus thibetanus pseudosingularis</emphasis>
|
||
<bibRefCitation id="EFDD96CEDA18FFD2FE32FD749C884EE0" author="Didier" box="[451,711,692,716]" pageId="3" pageNumber="59" refString="Didier, R. & Seguy, E. (1953) Note sur le Lucanus singularis Planet et sur les especes voisines. Revue francaise d'entomologie, 20, 28 - 32." type="journal article" year="1953">Didier & Séguy, 1953</bibRefCitation>
|
||
</taxonomicName>
|
||
, therefore it is a senior synonym of the latter. For the species delimitation of
|
||
<taxonomicName id="4C4C90BCDA18FFD2FEB5FD199C504EDC" box="[324,543,728,752]" class="Insecta" family="Lucanidae" genus="Lucanus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="59" phylum="Arthropoda" rank="species" species="thibetanus">
|
||
<emphasis id="B938372DDA18FFD2FEB5FD199C504EDC" box="[324,543,728,752]" italics="true" pageId="3" pageNumber="59">Lucanus thibetanus</emphasis>
|
||
</taxonomicName>
|
||
, see
|
||
<bibRefCitation id="EFDD96CEDA18FFD2FDA9FD189D194EC3" author="Huang" box="[600,854,727,752]" pageId="3" pageNumber="59" refString="Huang, H. & Chen, C. - C. (2010) Stag Beetles of China 1. Formosan Biology Company, Taipei, 288 pp." type="book" year="2010">Huang & Chen (2010)</bibRefCitation>
|
||
. The taxonomic changes of
|
||
<taxonomicName id="4C4C90BCDA18FFD2FB60FD199E844F38" class="Insecta" family="Lucanidae" genus="Lucanus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="59" phylum="Arthropoda" rank="subSpecies" species="thibetanus" subSpecies="furcifer">
|
||
<emphasis id="B938372DDA18FFD2FB60FD199E844F38" italics="true" pageId="3" pageNumber="59">Lucanus thibetanus furcifer</emphasis>
|
||
</taxonomicName>
|
||
in the literature are stated below.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |