579 lines
75 KiB
XML
579 lines
75 KiB
XML
<document id="447B9BE12D4A7931D45F714727F56011" ID-DOI="10.11646/zootaxa.3646.3.1" ID-GBIF-Dataset="cf465fc1-770c-4c46-98a6-4c5e1b40b0b9" ID-ISSN="1175-5326" ID-Zenodo-Dep="222898" ID-ZooBank="6FC50839-9E37-4D30-A3D3-FF28C71B6F2B" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460600059465" checkinUser="plazi" docAuthor="Johnson, Robert A., Overson, Rick P. & Moreau, Corrie S." docDate="2013" docId="03E1432AFFB1F473BCB7E04BFCFCF874" docLanguage="en" docName="zt03646p227.pdf" docOrigin="Zootaxa 3646 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Pogonomyrmex hoelldobleri Johnson & Overson & Moreau 2013, NEW SPECIES" docType="treatment" docVersion="10" lastPageNumber="217" masterDocId="FFD83B52FFBDF463BC20E671FF90FFE0" masterDocTitle="A New Species of Seed-harvester Ant, Pogonomyrmex hoelldobleri (Hymenoptera: Formicidae), from the Mohave and Sonoran Deserts of North America" masterLastPageNumber="227" masterPageNumber="201" pageNumber="213" updateTime="1698327381133" updateUser="plazi">
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<mods:title id="7B368A1F8B9C23BD123DE971CCA6CD9B">A New Species of Seed-harvester Ant, Pogonomyrmex hoelldobleri (Hymenoptera: Formicidae), from the Mohave and Sonoran Deserts of North America</mods:title>
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<mods:namePart id="60CEF808DB1E68B1F413B8175329948F">Johnson, Robert A.</mods:namePart>
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<mods:namePart id="A604E7F6B6EC14352B7C0AACB1F3EB67">Overson, Rick P.</mods:namePart>
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<mods:namePart id="93EFCBBD270CE00DA75BDEA4362FB6F2">Moreau, Corrie S.</mods:namePart>
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<mods:title id="9D9ADF3B5C954D70650E3320988CE051">Zootaxa</mods:title>
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<mods:part id="C673D891EEB9805A788B2EBF0A6600DB">
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<mods:date id="6456897AD3ACA5354F83BA748C9677D6">2013</mods:date>
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<mods:number id="C69D33E9CADB17F366B1FBA13F01D402">2013-05-03</mods:number>
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<mods:number id="FF61EAA8DD6770CCF82FB0628CEEF71C">3646</mods:number>
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<subSubSection id="C352A1B7FFB1F46FBCB7E04BFBA5F9B4" box="[151,1077,1594,1620]" pageId="12" pageNumber="213" type="nomenclature">
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<paragraph id="8BF7F23CFFB1F46FBCB7E04BFBA5F9B4" blockId="12.[151,1077,1594,1653]" box="[151,1077,1594,1620]" pageId="12" pageNumber="213">
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<heading id="D0BF4550FFB1F46FBCB7E04BFBA5F9B4" bold="true" box="[151,1077,1594,1620]" fontSize="11" level="1" pageId="12" pageNumber="213" reason="1">
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<emphasis id="B93C2E2EFFB1F46FBCB7E04BFBA5F9B4" bold="true" box="[151,1077,1594,1620]" pageId="12" pageNumber="213">
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<taxonomicName id="4C4889BFFFB1F46FBCB7E04BFCF0F9B4" authority="Johnson, Overson & Moreau" authorityName="Johnson & Overson & Moreau" authorityYear="2013" box="[151,864,1594,1620]" class="Insecta" family="Formicidae" genus="Pogonomyrmex" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="213" phylum="Arthropoda" rank="species" species="hoelldobleri" status="sp. nov.">
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<emphasis id="B93C2E2EFFB1F46FBCB7E04BFE7DF9B4" bold="true" box="[151,493,1594,1620]" italics="true" pageId="12" pageNumber="213">Pogonomyrmex hoelldobleri</emphasis>
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||
Johnson, Overson & Moreau
|
||
</taxonomicName>
|
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,
|
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<taxonomicNameLabel id="A20F9355FFB1F46FBF4EE04BFBA5F9B4" box="[878,1077,1594,1620]" pageId="12" pageNumber="213" rank="species">NEW SPECIES</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C352A1B7FFB1F46FBCB7E02CFD2EF95B" pageId="12" pageNumber="213" type="description">
|
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<paragraph id="8BF7F23CFFB1F46FBCB7E02CFEDAF995" blockId="12.[151,1077,1594,1653]" box="[151,330,1629,1653]" pageId="12" pageNumber="213">
|
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(
|
||
<figureCitation id="1373EEB9FFB1F46FBCBEE02CFE9BF995" box="[158,267,1629,1653]" captionStart="FIGURE 2" captionStartId="6.[151,250,1724,1747]" captionTargetBox="[171,1402,193,1704]" captionTargetId="figure@6.[169,1404,193,1707]" captionTargetPageId="6" captionText="FIGURE 2. Photographs of diagnostic characters to distinguish between workers of P. magnacanthus Cole and those of P. hoelldobleri Johnson, Overson & Moreau and P. m o h a v e n s i s Johnson. Photograph of P. magnacanthus PARATYPE worker: (A) eye and malar area (MOD = 0.44, OI = 29.5, MR = 0.98), and circumocular whorls indistinct to absent, moderately granulate toward vertex, and (B) circumocular whorls present. Photograph of P. hoelldobleri worker: (C) eye and malar area for HOLOTYPE worker (MOD = 0.36, OI = 23.2, MR = 1.17), and absence of circumocular whorls—rugae converging near vertex, and (D) absence of circumocular whorls—area posterior to eyes with faint rugae. Photograph of P. m o h a v e n s i s worker: (E) eye and malar area for PARATYPE worker (MOD = 0.33, OI = 21.7, MR = 1.36), and absence of circumocular whorlsrugae extending to vertex, with rugae becoming weak to absent on / near vertex, and (F) absence of circumocular whorls—rugae extending to vertex; vertex rugose." httpUri="https://zenodo.org/record/222900/files/figure.png" pageId="12" pageNumber="213">Figures 2</figureCitation>
|
||
,
|
||
<figureCitation id="1373EEB9FFB1F46FBD37E02CFED2F995" box="[279,322,1629,1653]" captionStart-0="FIGURE 6" captionStart-1="FIGURE 7" captionStart-2="FIGURE 8" captionStartId-0="13.[151,250,1958,1981]" captionStartId-1="17.[151,250,1970,1993]" captionStartId-2="18.[151,250,1970,1993]" captionTargetBox-0="[396,1200,187,1942]" captionTargetBox-1="[396,1190,193,1948]" captionTargetBox-2="[383,1204,193,1948]" captionTargetId-0="figure@13.[396,1201,187,1943]" captionTargetId-1="figure@17.[396,1191,193,1949]" captionTargetId-2="figure@18.[383,1204,193,1949]" captionTargetPageId-0="13" captionTargetPageId-1="17" captionTargetPageId-2="18" captionText-0="FIGURE 6. Photograph of Pogonomyrmex hoelldobleri Johnson, Overson & Moreau—HOLOTYPE worker: (A) frontal view of head, (B) lateral view of body, and (C) dorsal view of body." captionText-1="FIGURE 7. Photograph of Pogonomyrmex hoelldobleri Johnson, Overson & Moreau alate queen: (A) frontal view of head, (B) lateral view of body, and (C) dorsal view of body." captionText-2="FIGURE 8. Photograph of Pogonomyrmex hoelldobleri Johnson, Overson & Moreau male: (A) frontal view of head, (B) lateral view of body, and (C) dorsal view of body." httpUri-0="https://zenodo.org/record/222904/files/figure.png" httpUri-1="https://zenodo.org/record/222905/files/figure.png" httpUri-2="https://zenodo.org/record/222906/files/figure.png" pageId="12" pageNumber="213">6–8</figureCitation>
|
||
)
|
||
</paragraph>
|
||
<paragraph id="8BF7F23CFFB1F46FBCB7E0D7FD2EF95B" blockId="12.[151,702,1701,1723]" box="[151,702,1701,1723]" pageId="12" pageNumber="213">
|
||
<taxonomicName id="4C4889BFFFB1F46FBCB7E0D7FDE8F95B" authority="Cole, 1968: 133" authorityName="Cole" authorityPageNumber="133" authorityYear="1968" box="[151,632,1701,1723]" class="Insecta" family="Formicidae" genus="Pogonomyrmex" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="213" phylum="Arthropoda" rank="species" species="magnacanthus">
|
||
<emphasis id="B93C2E2EFFB1F46FBCB7E0D7FE40F95B" box="[151,464,1701,1723]" italics="true" pageId="12" pageNumber="213">Pogonomyrmex magnacanthus</emphasis>
|
||
Cole, 1968: 133
|
||
</taxonomicName>
|
||
[part].
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C352A1B7FFB1F46FBCB7E099FF63F8E0" box="[151,243,1768,1792]" pageId="12" pageNumber="213" type="description">
|
||
<paragraph id="8BF7F23CFFB1F46FBCB7E099FF63F8E0" blockId="12.[151,1436,1768,1972]" box="[151,243,1768,1792]" pageId="12" pageNumber="213">
|
||
<emphasis id="B93C2E2EFFB1F46FBCB7E099FF63F8E0" bold="true" box="[151,243,1768,1792]" pageId="12" pageNumber="213">Worker</emphasis>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C352A1B7FFB1F46DBCB7E17CFA10FEFC" lastPageId="14" lastPageNumber="215" pageId="12" pageNumber="213" type="diagnosis">
|
||
<paragraph id="8BF7F23CFFB1F46FBCB7E17CFED4F854" blockId="12.[151,1436,1768,1972]" pageId="12" pageNumber="213">
|
||
<emphasis id="B93C2E2EFFB1F46FBCB7E17CFDC8F8C4" box="[151,600,1804,1828]" italics="true" pageId="12" pageNumber="213">
|
||
Diagnosis.
|
||
<taxonomicName id="4C4889BFFFB1F46FBD37E17CFDC8F8C4" authorityName="Johnson & Overson & Moreau" authorityYear="2013" box="[279,600,1804,1828]" class="Insecta" family="Formicidae" genus="Pogonomyrmex" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="213" phylum="Arthropoda" rank="species" species="hoelldobleri">Pogonomyrmex hoelldobleri</taxonomicName>
|
||
</emphasis>
|
||
is characterized by: (1) eye not unusually large (MOD usually <0.42, OI usually <27.50, MR usually> 1.05 (
|
||
<figureCitation id="1373EEB9FFB1F46FBE0FE141FD1FF8A8" box="[559,655,1840,1864]" captionStart="FIGURE 3" captionStartId="7.[151,250,1916,1939]" captionTargetBox="[306,1293,185,1892]" captionTargetId="figure@7.[297,1296,139,1895]" captionTargetPageId="7" captionText="FIGURE 3. Bivariate plots for workers: (A) maximum eye diameter versus head width, (B) ocular index versus head width, and (C) malar ratio versus head width (n = 63 for P. magnacanthus Cole, n = 59 for P. hoelldobleri Johnson, Overson & Moreau, plus 16 PARATYPE workers of P. magnacanthus that do not belong to the latter species [see text], n = 25 for P. mohavensis Johnson). Non-type workers were selected to represent the geographic range of each species." httpUri="https://zenodo.org/record/222901/files/figure.png" pageId="12" pageNumber="213">Figure 3</figureCitation>
|
||
), (2) mandible with seven teeth, (3) cephalic rugae converge posterior to eyes, usually near vertex, but not forming circumocular whorls, (4) interrugal spaces on pronotal sides moderately to strongly granulate, dull to weakly shining (see
|
||
<figureCitation id="1373EEB9FFB1F46FBF74E109FC51F870" box="[852,961,1912,1936]" captionStart="FIGURE 2" captionStartId="6.[151,250,1724,1747]" captionTargetBox="[171,1402,193,1704]" captionTargetId="figure@6.[169,1404,193,1707]" captionTargetPageId="6" captionText="FIGURE 2. Photographs of diagnostic characters to distinguish between workers of P. magnacanthus Cole and those of P. hoelldobleri Johnson, Overson & Moreau and P. m o h a v e n s i s Johnson. Photograph of P. magnacanthus PARATYPE worker: (A) eye and malar area (MOD = 0.44, OI = 29.5, MR = 0.98), and circumocular whorls indistinct to absent, moderately granulate toward vertex, and (B) circumocular whorls present. Photograph of P. hoelldobleri worker: (C) eye and malar area for HOLOTYPE worker (MOD = 0.36, OI = 23.2, MR = 1.17), and absence of circumocular whorls—rugae converging near vertex, and (D) absence of circumocular whorls—area posterior to eyes with faint rugae. Photograph of P. m o h a v e n s i s worker: (E) eye and malar area for PARATYPE worker (MOD = 0.33, OI = 21.7, MR = 1.36), and absence of circumocular whorlsrugae extending to vertex, with rugae becoming weak to absent on / near vertex, and (F) absence of circumocular whorls—rugae extending to vertex; vertex rugose." httpUri="https://zenodo.org/record/222900/files/figure.png" pageId="12" pageNumber="213">Figures 2</figureCitation>
|
||
,
|
||
<figureCitation id="1373EEB9FFB1F46FBFEFE109FC4FF870" box="[975,991,1912,1936]" captionStart="FIGURE 6" captionStartId="13.[151,250,1958,1981]" captionTargetBox="[396,1200,187,1942]" captionTargetId="figure@13.[396,1201,187,1943]" captionTargetPageId="13" captionText="FIGURE 6. Photograph of Pogonomyrmex hoelldobleri Johnson, Overson & Moreau—HOLOTYPE worker: (A) frontal view of head, (B) lateral view of body, and (C) dorsal view of body." httpUri="https://zenodo.org/record/222904/files/figure.png" pageId="12" pageNumber="213">6</figureCitation>
|
||
), and (5) gaster concolorous with head and mesosoma.
|
||
</paragraph>
|
||
<caption id="DF37A2B4FFB0F46EBCB7E1D7FC81F83B" httpUri="https://zenodo.org/record/222904/files/figure.png" pageId="13" pageNumber="214" targetBox="[396,1200,187,1942]" targetPageId="13">
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<paragraph id="8BF7F23CFFB0F46EBCB7E1D7FC81F83B" blockId="13.[151,1436,1958,2012]" pageId="13" pageNumber="214">
|
||
<emphasis id="B93C2E2EFFB0F46EBCB7E1D7FE82F85C" bold="true" box="[151,274,1958,1981]" pageId="13" pageNumber="214">FIGURE 6.</emphasis>
|
||
Photograph of
|
||
<taxonomicName id="4C4889BFFFB0F46EBD90E1D9FC1DF85C" authority="Johnson, Overson" authorityName="Johnson, Overson & Moreau" authorityYear="2013" box="[432,909,1959,1981]" class="Insecta" family="Formicidae" genus="Pogonomyrmex" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="214" phylum="Arthropoda" rank="species" species="hoelldobleri">
|
||
<emphasis id="B93C2E2EFFB0F46EBD90E1D9FD40F85D" box="[432,720,1959,1981]" italics="true" pageId="13" pageNumber="214">Pogonomyrmex hoelldobleri</emphasis>
|
||
Johnson, Overson
|
||
</taxonomicName>
|
||
& Moreau—HOLOTYPE worker:
|
||
<emphasis id="B93C2E2EFFB0F46EB8D7E1D7FA8BF85D" bold="true" box="[1271,1307,1958,1981]" pageId="13" pageNumber="214">(A)</emphasis>
|
||
frontal view of head,
|
||
<emphasis id="B93C2E2EFFB0F46EBCCFE1B4FE81F83C" bold="true" box="[239,273,1989,2012]" pageId="13" pageNumber="214">(B)</emphasis>
|
||
lateral view of body, and
|
||
<emphasis id="B93C2E2EFFB0F46EBE38E1B4FDABF83C" bold="true" box="[536,571,1989,2012]" pageId="13" pageNumber="214">(C)</emphasis>
|
||
dorsal view of body.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BF7F23CFFB3F46DBCE7E6E9FA10FEFC" blockId="14.[151,1437,151,1545]" pageId="14" pageNumber="215">
|
||
Measurements (mm)—
|
||
<typeStatus id="54F34C9EFFB3F46DBDE9E6E6FDBCFF4F" box="[457,556,151,175]" pageId="14" pageNumber="215" type="holotype">holotype</typeStatus>
|
||
(
|
||
<emphasis id="B93C2E2EFFB3F46DBE1CE6E8FDDAFF50" box="[572,586,153,176]" italics="true" pageId="14" pageNumber="215">n</emphasis>
|
||
= 75 [15
|
||
<typeStatus id="54F34C9EFFB3F46DBE95E6E9FCB3FF50" box="[693,803,152,176]" pageId="14" pageNumber="215" type="paratype">paratypes</typeStatus>
|
||
, 43 non-types,
|
||
<specimenCount id="9D4E39B5FFB3F46DBFECE6E6FB93FF50" box="[972,1027,151,176]" pageId="14" pageNumber="215" type="generic">
|
||
16
|
||
<taxonomicName id="4C4889BFFFB3F46DBFCEE6E8FB93FF50" authorityName="Cole" authorityYear="1968" box="[1006,1027,153,176]" class="Insecta" family="Formicidae" genus="Pogonomyrmex" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="223" phylum="Arthropoda" rank="species" species="magnacanthus">
|
||
<emphasis id="B93C2E2EFFB3F46DBFCEE6E8FB93FF50" box="[1006,1027,153,176]" italics="true" pageId="14" pageNumber="215">P.</emphasis>
|
||
</taxonomicName>
|
||
</specimenCount>
|
||
magnacanthus
|
||
<typeStatus id="54F34C9EFFB3F46DB892E6E6FAB7FF4F" box="[1202,1319,151,175]" pageId="14" pageNumber="215" type="paratype">Paratypes</typeStatus>
|
||
]). HL 1.57 (1.24–1.76); HW 1.55 (1.17–1.84); MOD 0.36 (0.30–0.42); OMD 0.42 (0.35–0.72); SL 1.20 (0.94–1.34); PNW 0.91 (0.77–1.11); HF 1.67 (1.19–1.89); ML 1.84 (1.35–2.03); PW 0.34 (0.27–0.46): PPW 0.51 (0.38–0.59). Indices: SI 77.42 (66.25–87.18); CI 98.73 (93.98–115.79); OI 23.23 (21.12–29.01); HFI 107.74 (84.97–117.93).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C352A1B7FFB3F46DBCE7E758FF75F9B0" pageId="14" pageNumber="215" type="description">
|
||
<paragraph id="8BF7F23CFFB3F46DBCE7E758FBC5FD64" blockId="14.[151,1437,151,1545]" pageId="14" pageNumber="215">
|
||
<emphasis id="B93C2E2EFFB3F46DBCE7E758FEDBFEA0" box="[199,331,297,320]" italics="true" pageId="14" pageNumber="215">Description</emphasis>
|
||
. Head subquadrate to quadrate (CI = 93.98–115.79), broadest just posterior to eye; posterior margin flat in full-face view. Longitudinal cephalic rugae prominent, in full-face view median rugae usually diverging toward posterior corners near posterior margin of head. In side view, rugae converging slightly near vertex, occasionally becoming faint between posterior margin of eye and vertex. Vertex faintly to strongly rugose, occasionally mostly smooth to weakly granulate, sub-shining to shining. Cephalic interrugal spaces moderately punctate, sub-shining to smooth and shining. Anterior margin of clypeus slightly concave. Mandible with seven teeth; mandibular dorsum coarsely striate. In profile, eyes not unusually large, MOD ranging from 0.22–0.29x HL, OI = 21.12–29.01, MR usually> 1.05; eye situated near middle of head. Antennal scapes moderately long (SI = 66.25–87.18), failing to reach vertex by less than length of basal funicular segment. Basal flange of antennal scape flattened and well developed, margin weakly carinate. Psammophore well developed.
|
||
</paragraph>
|
||
<paragraph id="8BF7F23CFFB3F46DBCE7E4FEFF61FC27" blockId="14.[151,1437,151,1545]" pageId="14" pageNumber="215">Mesosomal profile convex. All mesosomal surfaces with prominent parallel/subparallel rugae. Dorsum of promesonotum with transverse rugae that curve obliquely to posterior on pronotal sides, rugae on pronotal sides often slightly less distinct than on other portions of mesosoma; rugae usually oblique to longitudinal on anterior portion of mesonotum. Mesopleura with subparallel rugae angling posterodorsally. Propodeum lacking spines or teeth; in side view, juncture of propodeum and propodeal declivity evenly convex to weakly angulate; rugae on propodeal dorsum transverse, declivitous face smooth and shining. Propodeal spiracles narrowly ovate. Interrugal spaces on mesosoma moderately granulate-punctate, sub-shining to smooth and shining; interrugal spaces on pronotal sides usually more densely granulate than other portions of mesosoma. Legs moderately to strongly shining.</paragraph>
|
||
<paragraph id="8BF7F23CFFB3F46DBCE7E5A5FEC4FAEC" blockId="14.[151,1437,151,1545]" pageId="14" pageNumber="215">Petiolar peduncle long, ventral margin straight. In side view, posterior face of petiole slightly convex; petiolar node asymmetrical with anterior surface slightly shorter than posterior surface. Apex of node weakly to strongly angulate, posterior surface sometimes curved upward near anterior margin. In dorsal view, petiolar node longer than broad, sides subparallel or diverging slightly toward the smoothly rounded to weakly angulate anterior margin. Sides and dorsum of petiolar node strongly granulate-punctate, dull to sub-shining to smooth and shining, occasionally with several longitudinal to oblique rugae that are restricted to posterior one-third of petiole. Dorsum of postpetiole convex in profile; in dorsal view, widest at or near posterior margin and tapering to anterior margin, maximal width about equal to length, weakly to moderately granulate, dull to sub-shining. Gaster smooth and strongly shining.</paragraph>
|
||
<paragraph id="8BF7F23CFFB3F46DBCE7E369FD67F9E7" blockId="14.[151,1437,151,1545]" pageId="14" pageNumber="215">
|
||
Erect to suberect white pilosity moderately abundant on head, short to medium in length, often with one to few longer hairs, none exceeding MOD. Moderately abundant semidecumbent to decumbent pilosity on scape, abundant semidecumbent to decumbent hairs on funicular segments. Legs with moderately abundant suberect to semidecumbent white setae. Mesosoma, petiole, and postpetiole with a lower density of mostly longer, flexuous hairs mostly concentrated on dorsal surface, longest distinctly shorter than MOD; gastric tergites with moderately abundant, medium length suberect hairs. Entire body concolorous light to dark ferruginous orange, posterior portion of gaster sometimes slightly darker (
|
||
<figureCitation id="1373EEB9FFB3F46DBEA7E39EFD79F9E7" box="[647,745,1519,1543]" captionStart="FIGURE 6" captionStartId="13.[151,250,1958,1981]" captionTargetBox="[396,1200,187,1942]" captionTargetId="figure@13.[396,1201,187,1943]" captionTargetPageId="13" captionText="FIGURE 6. Photograph of Pogonomyrmex hoelldobleri Johnson, Overson & Moreau—HOLOTYPE worker: (A) frontal view of head, (B) lateral view of body, and (C) dorsal view of body." httpUri="https://zenodo.org/record/222904/files/figure.png" pageId="14" pageNumber="215">Figure 6</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BF7F23CFFB3F46DBCB7E046FF75F9B0" blockId="14.[151,1437,1591,2012]" box="[151,229,1591,1616]" pageId="14" pageNumber="215">
|
||
<emphasis id="B93C2E2EFFB3F46DBCB7E046FF75F9B0" bold="true" box="[151,229,1591,1616]" pageId="14" pageNumber="215">Queen</emphasis>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C352A1B7FFB3F46DBCB7E02CFC95F874" pageId="14" pageNumber="215" type="diagnosis">
|
||
<paragraph id="8BF7F23CFFB3F46DBCB7E02CFAB7F8C7" blockId="14.[151,1437,1591,2012]" pageId="14" pageNumber="215">
|
||
<emphasis id="B93C2E2EFFB3F46DBCB7E02CFE80F994" box="[151,272,1629,1652]" italics="true" pageId="14" pageNumber="215">Diagnosis.</emphasis>
|
||
As in worker diagnosis, but with caste-specific structures related to wing-bearing and presence of small ocelli on head. Mandible with seven teeth. Eye not unusually large (MOD <0.45, OI <28.50, MR usually> 1.05). All mesosomal surfaces except for mesoscutum and mesoscutellum with prominent rugae; sculpturing absent on mesoscutum and mesoscutellum except for scattered punctures and occasional faint longitudinal striae (
|
||
<figureCitation id="1373EEB9FFB3F46DB90BE0B6FA1EF93F" box="[1323,1422,1735,1759]" captionStart="FIGURE 7" captionStartId="17.[151,250,1970,1993]" captionTargetBox="[396,1190,193,1948]" captionTargetId="figure@17.[396,1191,193,1949]" captionTargetPageId="17" captionText="FIGURE 7. Photograph of Pogonomyrmex hoelldobleri Johnson, Overson & Moreau alate queen: (A) frontal view of head, (B) lateral view of body, and (C) dorsal view of body." httpUri="https://zenodo.org/record/222905/files/figure.png" pageId="14" pageNumber="215">Figure 7</figureCitation>
|
||
). Posterior face of petiole and dorsum of postpetiole weakly to moderately granulate or with weak tranverse rugae. Base of scape rounded; superior and inferior lobes poorly developed, no wider than width of scape base.
|
||
</paragraph>
|
||
<paragraph id="8BF7F23CFFB3F46DBCE7E144FC95F874" blockId="14.[151,1437,1591,2012]" pageId="14" pageNumber="215">
|
||
Measurements (mm)—(
|
||
<emphasis id="B93C2E2EFFB3F46DBDF7E144FD91F8AC" box="[471,513,1845,1868]" italics="true" pageId="14" pageNumber="215">n =</emphasis>
|
||
8). HL 1.36–1.66; HW 1.44–1.79; MOD 0.40–0.45; OMD 0.40–0.50; SL 0.97– 1.23; PNW 1.24–1.54; HFL 1.38–1.67; ML 2.18–2.62; PW 0.45–0.57; PPW 0.61–0.73. Indices: SI 65.52–72.12; CI 100.00–114.10; OI 22.47–28.47; HFI 83.64–109.03.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C352A1B7FFB3F46CBCE7E1D0FF44FD2C" lastPageId="15" lastPageNumber="216" pageId="14" pageNumber="215" type="description">
|
||
<paragraph id="8BF7F23CFFB3F46CBCE7E1D0FBAAFEDF" blockId="14.[151,1437,1591,2012]" lastBlockId="15.[151,1437,151,644]" lastPageId="15" lastPageNumber="216" pageId="14" pageNumber="215">
|
||
<emphasis id="B93C2E2EFFB3F46DBCE7E1D0FEC3F858" box="[199,339,1953,1976]" italics="true" pageId="14" pageNumber="215">Description.</emphasis>
|
||
As in worker diagnosis, but with caste-specific structures related to wing-bearing, presence of small ocelli on head, and as illustrated in
|
||
<figureCitation id="1373EEB9FFB3F46DBE45E1B5FD55F83C" box="[613,709,1988,2012]" captionStart="FIGURE 7" captionStartId="17.[151,250,1970,1993]" captionTargetBox="[396,1190,193,1948]" captionTargetId="figure@17.[396,1191,193,1949]" captionTargetPageId="17" captionText="FIGURE 7. Photograph of Pogonomyrmex hoelldobleri Johnson, Overson & Moreau alate queen: (A) frontal view of head, (B) lateral view of body, and (C) dorsal view of body." httpUri="https://zenodo.org/record/222905/files/figure.png" pageId="14" pageNumber="215">Figure 7</figureCitation>
|
||
. Small, only slightly larger than conspecific workers. In full-face view, head quadrate to broader than long, posterior margin flat. Dorsum and sides of head conspicuously rugose, in side view rugae forming circumocular whorls posterior to eyes or rugae converging near vertex, interrugal spaces smooth and strongly shining. Mandible with seven teeth, dorsal surface coarsely rugose, strongly shining. Eyes not large (OI = 22.47–28.47), MR usually> 1.05, MOD ranging from 0.25–0.30x HL. Base of scape not flattened; superior and inferior lobes poorly developed, no wider than width of base of scape.
|
||
</paragraph>
|
||
<paragraph id="8BF7F23CFFB2F46CBCE7E73CFC92FD64" blockId="15.[151,1437,151,644]" pageId="15" pageNumber="216">Mesosoma as described above, propodeum unarmed; in side view, juncture of propodeum and propodeal declivity rounded to subangulate, sides and dorsal surface rugose or rugae absent near mid-line, shining, posterior surface smooth and strongly shining. Petiolar peduncle long, ventral margin straight. In side view, petiolar node asymmetrical with anterior surface shorter than posterior surface. Apex of node moderately to strongly angulate, anterior edge of posterior face sometimes curved upward forming a crest. Postpetiole broader than long. Posterior face of petiole and dorsum of postpetiole weakly to moderately granulate or with weak transverse rugae, subshining to shining. Gastric tergites smooth and shining. Most body surfaces with moderately abundant coarse suberect to erect setae. Entire body concolorous light to dark ferruginous orange, except for incomplete to complete darker transverse bands on one or more gastric tergites.</paragraph>
|
||
<paragraph id="8BF7F23CFFB2F46CBCB7E4C5FF44FD2C" blockId="15.[151,1436,692,2015]" box="[151,212,692,716]" pageId="15" pageNumber="216">
|
||
<emphasis id="B93C2E2EFFB2F46CBCB7E4C5FF44FD2C" bold="true" box="[151,212,692,716]" pageId="15" pageNumber="216">Male</emphasis>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C352A1B7FFB2F46CBCB7E4A8FDA9FC27" pageId="15" pageNumber="216" type="diagnosis">
|
||
<paragraph id="8BF7F23CFFB2F46CBCB7E4A8FD2AFCBC" blockId="15.[151,1436,692,2015]" pageId="15" pageNumber="216">
|
||
<emphasis id="B93C2E2EFFB2F46CBCB7E4A8FE80FD10" box="[151,272,729,752]" italics="true" pageId="15" pageNumber="216">Diagnosis.</emphasis>
|
||
Mandible with four teeth on suboblique cutting margin. Mandibular dorsum, clypeus, and antennal scapes lacking sculpture (mandibular dorsum occasionally with faint striae), mostly smooth and shining except for scattered punctures; anterior margin of clypeus weakly concave, lateral lobes indistinct. Eye not unusually large (MOD <0.50, OI <39.4, MR> 0.38) (
|
||
<figureCitation id="1373EEB9FFB2F46CBE6BE535FD3CFCBC" box="[587,684,836,860]" captionStart="FIGURE 8" captionStartId="18.[151,250,1970,1993]" captionTargetBox="[383,1204,193,1948]" captionTargetId="figure@18.[383,1204,193,1949]" captionTargetPageId="18" captionText="FIGURE 8. Photograph of Pogonomyrmex hoelldobleri Johnson, Overson & Moreau male: (A) frontal view of head, (B) lateral view of body, and (C) dorsal view of body." httpUri="https://zenodo.org/record/222906/files/figure.png" pageId="15" pageNumber="216">Figure 8</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BF7F23CFFB2F46CBCE7E519FDA9FC27" blockId="15.[151,1436,692,2015]" pageId="15" pageNumber="216">
|
||
Measurements (mm)—(
|
||
<emphasis id="B93C2E2EFFB2F46CBDF7E518FE75FC60" box="[471,485,873,896]" italics="true" pageId="15" pageNumber="216">n</emphasis>
|
||
= 2). HL 1.15–1.25; HW 1.27–1.30; MOD 0.49–0.50; OMD 0.19–0.20; SL 0.54– 0.56; HFL 1.43–1.56; ML 2.08–2.26; PW 0.54–0.58; PPW 0.65–0.73. Indices: SI 42.52–43.08; CI 104.00–110.43; OI 37.69–39.37; HFI 112.60–120.00.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C352A1B7FFB2F46CBCE7E5A4FA96F9CC" pageId="15" pageNumber="216" type="description">
|
||
<paragraph id="8BF7F23CFFB2F46CBCE7E5A4FC4CFB9C" blockId="15.[151,1436,692,2015]" pageId="15" pageNumber="216">
|
||
<emphasis id="B93C2E2EFFB2F46CBCE7E5A4FEC3FC0C" box="[199,339,981,1004]" italics="true" pageId="15" pageNumber="216">Description.</emphasis>
|
||
Mandible with four teeth on suboblique cutting margin; tip of sub-apical tooth sometimes weakly bifid; basal tooth not offset; mandibular dorsum as described above. Anterior margin of clypeus broadly and shallowly concave, mostly smooth and shining except for scattered punctures, lateral lobes indistinct. Antennal scapes reaching to or near posterior margin of eye, mostly smooth and shining. Cephalic rugae fine and close, slightly wavy to irregular, interrugae weakly punctate, moderately shining.
|
||
</paragraph>
|
||
<paragraph id="8BF7F23CFFB2F46CBCE7E2F9FA96F9CC" blockId="15.[151,1436,692,2015]" pageId="15" pageNumber="216">
|
||
In profile, anterior face of mesonotum forming a mostly straight line with pronotum, slightly less than one-half as long as dorsal surface. In side view, juncture between propodeum and propodeal declivity subangulate, without spines or denticles. Sides of pronotal collar superficially rugoreticulate to punctate-granulate; katepisternum partially to largely covered by very fine wavy to irregular longitudinal striae, sub-shining to shining. Mesonotum shiny with piligerous punctures, notauli weakly impressed. Propodeum mostly smooth and shining to microrugoreticulate, granulate, sub-shining. Ventral margin of petiolar peduncle straight. In side view, petiolar node broadly rounded, anterior surface longer than posterior surface, forming a mostly straight continuous to slightly curved profile with dorsal surface of petiolar peduncle. Dorsal surface of petiole smooth and shining with scattered punctures to microrugoreticulate, sub-shining. Postpetiole broader than long, dorsal surface mostly smooth, sub-shining to shining. Head, mesosoma, petiole, and postpetiole with moderately abundant flexuous white hairs, often similar in length to MOD. Gastric tergites smooth and shining, hairs shorter and less dense than on rest of body. Head and mesosoma brownish-orange, gaster a lighter ferruginous orange (
|
||
<figureCitation id="1373EEB9FFB2F46CB8B6E065FB68F9CC" box="[1174,1272,1556,1580]" captionStart="FIGURE 8" captionStartId="18.[151,250,1970,1993]" captionTargetBox="[383,1204,193,1948]" captionTargetId="figure@18.[383,1204,193,1949]" captionTargetPageId="18" captionText="FIGURE 8. Photograph of Pogonomyrmex hoelldobleri Johnson, Overson & Moreau male: (A) frontal view of head, (B) lateral view of body, and (C) dorsal view of body." httpUri="https://zenodo.org/record/222906/files/figure.png" pageId="15" pageNumber="216">Figure 8</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C352A1B7FFB2F473BCE7E049FC80FB9C" lastPageId="16" lastPageNumber="217" pageId="15" pageNumber="216" type="materials_examined">
|
||
<paragraph id="8BF7F23CFFB2F46CBCE7E049FDF8F8CA" blockId="15.[151,1436,692,2015]" pageId="15" pageNumber="216">
|
||
<emphasis id="B93C2E2EFFB2F46CBCE7E049FEFEF9B0" bold="true" box="[199,366,1591,1616]" italics="true" pageId="15" pageNumber="216">
|
||
<typeStatus id="54F34C9EFFB2F46CBCE7E049FF6EF9AF" box="[199,254,1592,1615]" pageId="15" pageNumber="216">Type</typeStatus>
|
||
material:
|
||
</emphasis>
|
||
<typeStatus id="54F34C9EFFB2F46CBD54E046FE4CF9AF" box="[372,476,1591,1615]" pageId="15" pageNumber="216" type="holotype">Holotype</typeStatus>
|
||
(worker) plus 27
|
||
<typeStatus id="54F34C9EFFB2F46CBE83E049FC82F9B0" box="[675,786,1592,1616]" pageId="15" pageNumber="216" type="paratype">paratypes</typeStatus>
|
||
.
|
||
<emphasis id="B93C2E2EFFB2F46CBF3BE049FBF1F9B0" bold="true" box="[795,1121,1591,1616]" pageId="15" pageNumber="216">
|
||
<collectingCountry id="F35FB2ACFFB2F46CBF3BE049FC6AF9B0" box="[795,1018,1591,1616]" name="United States of America" pageId="15" pageNumber="216">UNITED STATES</collectingCountry>
|
||
:
|
||
<emphasis id="B93C2E2EFFB2F46CB825E046FBF1F9B0" bold="true" box="[1029,1121,1591,1616]" italics="true" pageId="15" pageNumber="216">Arizona</emphasis>
|
||
</emphasis>
|
||
:
|
||
<taxonomicName id="4C4889BFFFB2F46CB84FE048FB4BF9AF" authority="Co." authorityName="Co." box="[1135,1243,1592,1616]" class="Insecta" genus="Yuma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="15" pageNumber="216" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B93C2E2EFFB2F46CB84FE048FB41F9AF" box="[1135,1233,1592,1616]" italics="true" pageId="15" pageNumber="216">Yuma Co</emphasis>
|
||
.
|
||
</taxonomicName>
|
||
:
|
||
<quantity id="4CB05FD9FFB2F46CB8C9E046FAA6F9AF" box="[1257,1334,1591,1615]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="15" pageNumber="216" unit="km" value="0.2">0.2 km</quantity>
|
||
S Tacna,
|
||
<quantity id="4CB05FD9FFB2F46CBCB7E02EFF4EF997" box="[151,222,1631,1655]" metricMagnitude="2" metricUnit="m" metricValue="1.25" pageId="15" pageNumber="216" unit="m" value="125.0">125 m</quantity>
|
||
(
|
||
<geoCoordinate id="EE7C94FBFFB2F46CBCCEE02EFEC6F997" box="[238,342,1628,1655]" direction="south" orientation="latitude" pageId="15" pageNumber="216" precision="92" value="-22.106667">22o 6.4’S</geoCoordinate>
|
||
<geoCoordinate id="EE7C94FBFFB2F46CBD7EE02EFE71F997" box="[350,481,1628,1655]" direction="west" orientation="longitude" pageId="15" pageNumber="216" precision="92" value="-65.613335">65o 36.8’W</geoCoordinate>
|
||
),
|
||
<date id="FFF6D4FCFFB2F46CBDD6E011FD05F997" box="[502,661,1631,1656]" pageId="15" pageNumber="216" value="2010-05-14">May 14, 2010</date>
|
||
, leg. R.A. Johnson #4500. Nests were in Sonoran Desert habitat that was dominated by scattered individuals of
|
||
<taxonomicName id="4C4889BFFFB2F46CBE5FE0F5FCD3F97B" box="[639,835,1667,1691]" class="Magnoliopsida" family="Zygophyllaceae" genus="Larrea" kingdom="Plantae" order="Zygophyllales" pageId="15" pageNumber="216" phylum="Tracheophyta" rank="species" species="tridentata">
|
||
<emphasis id="B93C2E2EFFB2F46CBE5FE0F5FCD3F97B" box="[639,835,1667,1691]" italics="true" pageId="15" pageNumber="216">Larrea tridentata</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C4889BFFFB2F46CBF5DE0F2FBD8F97B" authorityName="Payne" authorityYear="1964" baseAuthorityName="A. Gray" box="[893,1096,1667,1691]" class="Magnoliopsida" family="Asteraceae" genus="Ambrosia" kingdom="Plantae" order="Asterales" pageId="15" pageNumber="216" phylum="Tracheophyta" rank="species" species="dumosa">
|
||
<emphasis id="B93C2E2EFFB2F46CBF5DE0F2FBD8F97B" box="[893,1096,1667,1691]" italics="true" pageId="15" pageNumber="216">Ambrosia dumosa</emphasis>
|
||
</taxonomicName>
|
||
. The
|
||
<typeStatus id="54F34C9EFFB2F46CB8ACE0F3FB7EF97A" box="[1164,1262,1666,1690]" pageId="15" pageNumber="216" type="holotype">holotype</typeStatus>
|
||
is deposited in the MCZ.
|
||
<typeStatus id="54F34C9EFFB2F46CBD2FE0D9FEECF920" box="[271,380,1704,1728]" pageId="15" pageNumber="216" type="paratype">Paratypes</typeStatus>
|
||
(
|
||
<emphasis id="B93C2E2EFFB2F46CBDAEE0D9FE0CF95F" box="[398,412,1704,1727]" italics="true" pageId="15" pageNumber="216">n</emphasis>
|
||
=
|
||
<specimenCount id="9D4E39B5FFB2F46CBD9EE0D6FDAFF95F" box="[446,575,1703,1727]" pageId="15" pageNumber="216" type="worker">27 workers</specimenCount>
|
||
) all from the same locality and date as the
|
||
<typeStatus id="54F34C9EFFB2F46CB814E0D6FB06F95F" box="[1076,1174,1703,1727]" pageId="15" pageNumber="216" type="holotype">holotype</typeStatus>
|
||
and leg. R.A. Johnson #4500 are distributed as follows: MCZ (6w), LACM (3w), UCDC (3w), USNM (6w), WPMC (3w), RAJC (6w). Additional
|
||
<typeStatus id="54F34C9EFFB2F46CBD37E081FEE7F8E8" box="[279,375,1776,1800]" pageId="15" pageNumber="216" type="paratype">paratype</typeStatus>
|
||
series (RAJC), collected on
|
||
<date id="FFF6D4FCFFB2F46CBE9EE081FCF0F8E7" box="[702,864,1775,1799]" pageId="15" pageNumber="216" value="2009-06-24">June 24, 2009</date>
|
||
, include RAJ #4253 (6w) and RAJ#4255 (9w): all series have additional workers in ethanol.
|
||
</paragraph>
|
||
<paragraph id="8BF7F23CFFB2F473BCE7E147FC80FB9C" blockId="15.[151,1436,692,2015]" lastBlockId="16.[151,1437,151,1940]" lastPageId="16" lastPageNumber="217" pageId="15" pageNumber="216">
|
||
<emphasis id="B93C2E2EFFB2F46CBCE7E147FC13F8AF" bold="true" box="[199,899,1846,1871]" pageId="15" pageNumber="216">
|
||
<emphasis id="B93C2E2EFFB2F46CBCE7E147FDB1F8AF" bold="true" box="[199,545,1846,1871]" italics="true" pageId="15" pageNumber="216">Additional material examined</emphasis>
|
||
.
|
||
<collectingCountry id="F35FB2ACFFB2F46CBE10E146FC81F8AF" box="[560,785,1846,1871]" name="United States of America" pageId="15" pageNumber="216">UNITED STATES</collectingCountry>
|
||
:
|
||
<emphasis id="B93C2E2EFFB2F46CBF3EE147FCEAF8AF" bold="true" box="[798,890,1846,1871]" italics="true" pageId="15" pageNumber="216">Arizona</emphasis>
|
||
:
|
||
</emphasis>
|
||
<emphasis id="B93C2E2EFFB2F46CBFABE149FBB9F8AF" box="[907,1065,1848,1871]" italics="true" pageId="15" pageNumber="216">Maricopa Co.</emphasis>
|
||
:
|
||
<quantity id="4CB05FD9FFB2F46CB81BE146FBECF8AF" box="[1083,1148,1847,1871]" metricMagnitude="4" metricUnit="m" metricValue="1.9312128" pageId="15" pageNumber="216" unit="mi" value="12.0">12 mi</quantity>
|
||
E Sentinel,
|
||
<date id="FFF6D4FCFFB2F46CB929E146FA08F8AF" box="[1289,1432,1847,1871]" pageId="15" pageNumber="216" value="1960-07-30">Jul 30, 1960</date>
|
||
, AC Cole AZ-519 (26w
|
||
<typeStatus id="54F34C9EFFB2F46CBD81E12BFDABF892" box="[417,571,1882,1906]" pageId="15" pageNumber="216" type="paratype">PARATYPES</typeStatus>
|
||
of
|
||
<emphasis id="B93C2E2EFFB2F46CBE40E12DFCB6F893" box="[608,806,1883,1907]" italics="true" pageId="15" pageNumber="216">
|
||
<taxonomicName id="4C4889BFFFB2F46CBE40E12DFCB2F893" authorityName="Cole" authorityYear="1968" box="[608,802,1883,1907]" class="Insecta" family="Formicidae" genus="Pogonomyrmex" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hymenoptera" pageId="15" pageNumber="216" phylum="Arthropoda" rank="species" species="magnacanthus">P. magnacanthus</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
LACM).
|
||
<emphasis id="B93C2E2EFFB2F46CBFB5E12AFB8EF892" box="[917,1054,1883,1907]" italics="true" pageId="15" pageNumber="216">Mohave Co.</emphasis>
|
||
: Golden Shores,
|
||
<date id="FFF6D4FCFFB2F46CB8C2E12BFAF9F892" box="[1250,1385,1882,1906]" pageId="15" pageNumber="216" value="1995-09-05">Sep 5, 1995</date>
|
||
, RA Johnson RAJ#691 (8w, RAJC);
|
||
<quantity id="4CB05FD9FFB2F46CBE32E10EFDCEF877" box="[530,606,1919,1943]" metricMagnitude="2" metricUnit="m" metricValue="4.828032" pageId="15" pageNumber="216" unit="mi" value="0.3">0.3 mi</quantity>
|
||
W Golden Shores,
|
||
<date id="FFF6D4FCFFB2F46CBF6EE1F1FC67F877" box="[846,1015,1919,1944]" pageId="15" pageNumber="216" value="2010-05-18">May 18, 2010</date>
|
||
, 620’, RA Johnson RAJ#4490 (6w, RAJC).
|
||
<taxonomicName id="4C4889BFFFB2F46CBCD4E1D5FEF0F85B" authority="Co." authorityName="Co." box="[244,352,1955,1979]" class="Insecta" genus="Yuma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="15" pageNumber="216" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B93C2E2EFFB2F46CBCD4E1D5FEF0F85B" box="[244,352,1955,1979]" italics="true" pageId="15" pageNumber="216">Yuma Co.</emphasis>
|
||
</taxonomicName>
|
||
:
|
||
<date id="FFF6D4FCFFB2F46CBD54E1D3FE04F85A" box="[372,404,1954,1978]" pageId="15" pageNumber="216">I-8</date>
|
||
at Aztec Road, 490’,
|
||
<date id="FFF6D4FCFFB2F46CBEB1E1D2FCBCF85A" box="[657,812,1954,1979]" pageId="15" pageNumber="216" value="2012-04-26">Apr 26, 2012</date>
|
||
, RA Johnson #4919 (6w, RAJC).
|
||
<emphasis id="B93C2E2EFFB2F46CB892E1D3FAA3F85B" bold="true" box="[1202,1331,1954,1979]" pageId="15" pageNumber="216">
|
||
<emphasis id="B93C2E2EFFB2F46CB892E1D3FABAF85B" bold="true" box="[1202,1322,1954,1979]" italics="true" pageId="15" pageNumber="216">California</emphasis>
|
||
:
|
||
</emphasis>
|
||
<taxonomicName id="4C4889BFFFB2F46CB91CE1D2FF2FF83F" authority="Co." authorityName="Co." class="Anthozoa" family="Aurelianidae" genus="Imperial" higherTaxonomySource="GBIF" kingdom="Animalia" order="Actiniaria" pageId="15" pageNumber="216" phylum="Cnidaria" rank="genus">
|
||
<emphasis id="B93C2E2EFFB2F46CB91CE1D2FF2FF83F" italics="true" pageId="15" pageNumber="216">Imperial Co.</emphasis>
|
||
</taxonomicName>
|
||
: Coyote Wells,
|
||
<date id="FFF6D4FCFFB2F46CBD54E1B6FD92F83F" box="[372,514,1991,2015]" pageId="15" pageNumber="216" value="1957-07-29">Jul 29, 1957</date>
|
||
, AC Cole CAL-305 (17w, LACM); El Centro to Jacumba,
|
||
<date id="FFF6D4FCFFB2F46CB8BFE1B6FABDF83F" box="[1183,1325,1991,2015]" pageId="15" pageNumber="216" value="1956-07-10">Jul 10, 1956</date>
|
||
, AC Cole CAL-11 (3w
|
||
<typeStatus id="54F34C9EFFADF473BD1CE6E6FE47FF4F" box="[316,471,151,175]" pageId="16" pageNumber="217" type="paratype">PARATYPES</typeStatus>
|
||
of
|
||
<taxonomicName id="4C4889BFFFADF473BE2CE6E8FD42FF50" authorityName="Cole" authorityYear="1968" box="[524,722,152,176]" class="Insecta" family="Formicidae" genus="Pogonomyrmex" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="217" phylum="Arthropoda" rank="species" species="magnacanthus">
|
||
<emphasis id="B93C2E2EFFADF473BE2CE6E8FD42FF50" box="[524,722,152,176]" italics="true" pageId="16" pageNumber="217">P. magnacanthus</emphasis>
|
||
</taxonomicName>
|
||
, LACM), CAL-12 (3w
|
||
<typeStatus id="54F34C9EFFADF473BFDFE6E6FB0AFF4F" box="[1023,1178,151,175]" pageId="16" pageNumber="217" type="paratype">PARATYPES</typeStatus>
|
||
of
|
||
<taxonomicName id="4C4889BFFFADF473B8EEE6E8FA04FF50" authorityName="Cole" authorityYear="1968" box="[1230,1428,152,176]" class="Insecta" family="Formicidae" genus="Pogonomyrmex" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="217" phylum="Arthropoda" rank="species" species="magnacanthus">
|
||
<emphasis id="B93C2E2EFFADF473B8EEE6E8FA04FF50" box="[1230,1428,152,176]" italics="true" pageId="16" pageNumber="217">P. magnacanthus</emphasis>
|
||
</taxonomicName>
|
||
, LACM);
|
||
<quantity id="4CB05FD9FFADF473BD25E6CDFEDCFF34" box="[261,332,188,212]" metricMagnitude="4" metricUnit="m" metricValue="2.2530816" pageId="16" pageNumber="217" unit="mi" value="14.0">14 mi</quantity>
|
||
W Winterhaven,
|
||
<date id="FFF6D4FCFFADF473BE03E6CDFD2BFF34" box="[547,699,188,212]" pageId="16" pageNumber="217" value="1959-07-28">Jul 28, 1959</date>
|
||
, AC Cole CAL-334A (6w
|
||
<typeStatus id="54F34C9EFFADF473B825E6CDFB0FFF34" box="[1029,1183,188,212]" pageId="16" pageNumber="217" type="paratype">PARATYPES</typeStatus>
|
||
of
|
||
<taxonomicName id="4C4889BFFFADF473B8F0E6CCFA05FF34" authorityName="Cole" authorityYear="1968" box="[1232,1429,188,212]" class="Insecta" family="Formicidae" genus="Pogonomyrmex" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="217" phylum="Arthropoda" rank="species" species="magnacanthus">
|
||
<emphasis id="B93C2E2EFFADF473B8F0E6CCFA05FF34" box="[1232,1429,188,212]" italics="true" pageId="16" pageNumber="217">P. magnacanthus</emphasis>
|
||
</taxonomicName>
|
||
, LACM);
|
||
<quantity id="4CB05FD9FFADF473BD22E6AEFED5FF17" box="[258,325,223,247]" metricMagnitude="4" metricUnit="m" metricValue="2.414016" pageId="16" pageNumber="217" unit="mi" value="15.0">15 mi</quantity>
|
||
E Holtville,
|
||
<date id="FFF6D4FCFFADF473BDFEE6AEFDFFFF17" box="[478,623,223,247]" pageId="16" pageNumber="217" value="1961-07-27">Jul 27, 1961</date>
|
||
, AC Cole CAL-394 (24w
|
||
<typeStatus id="54F34C9EFFADF473BF84E6AEFBAEFF17" box="[932,1086,223,247]" pageId="16" pageNumber="217" type="paratype">PARATYPES</typeStatus>
|
||
of
|
||
<taxonomicName id="4C4889BFFFADF473B849E690FABAFF18" authorityName="Cole" authorityYear="1968" box="[1129,1322,224,248]" class="Insecta" family="Formicidae" genus="Pogonomyrmex" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="217" phylum="Arthropoda" rank="species" species="magnacanthus">
|
||
<emphasis id="B93C2E2EFFADF473B849E690FABAFF18" box="[1129,1322,224,248]" italics="true" pageId="16" pageNumber="217">P. magnacanthus</emphasis>
|
||
</taxonomicName>
|
||
, LACM).
|
||
<taxonomicName id="4C4889BFFFADF473BCB7E774FF67FEFC" authority="Co." authorityName="Co." box="[151,247,261,284]" class="Insecta" family="Bombyliidae" genus="Inyo" kingdom="Animalia" order="Diptera" pageId="16" pageNumber="217" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B93C2E2EFFADF473BCB7E774FF67FEFC" box="[151,247,261,284]" italics="true" pageId="16" pageNumber="217">Inyo Co.</emphasis>
|
||
</taxonomicName>
|
||
: Death Valley National Monument at Ashford Mill, 0’,
|
||
<date id="FFF6D4FCFFADF473BF42E775FC69FEFC" box="[866,1017,260,284]" pageId="16" pageNumber="217" value="1952-04-28">Apr 28, 1952</date>
|
||
, WS Creighton no number (16w, 5aq, LACM), CR-226 (10w, 1aq,
|
||
<quantity id="4CB05FD9FFADF473BDC6E756FD9CFEDF" box="[486,524,295,319]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="16" pageNumber="217" unit="m" value="1.0">1m</quantity>
|
||
, LACM), CR-417 (11w, 1aq, LACM), CR-618 (15w, 1aq,
|
||
<quantity id="4CB05FD9FFADF473B899E756FB4EFEDF" box="[1209,1246,295,319]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="16" pageNumber="217" unit="m" value="1.0">1m</quantity>
|
||
, LACM); Death Valley National Park, Spring 2000, KE Anderson KEA#550 (3w, RAJC);
|
||
<quantity id="4CB05FD9FFADF473BFD0E73DFBC0FE84" box="[1008,1104,332,356]" metricMagnitude="3" metricUnit="m" metricValue="9.05" pageId="16" pageNumber="217" unit="km" value="9.05">9.05 km</quantity>
|
||
W Panamint Springs, 4450',
|
||
<date id="FFF6D4FCFFADF473BCB7E701FEA5FE67" box="[151,309,367,392]" pageId="16" pageNumber="217" value="2006-05-30">May 30, 2006</date>
|
||
, RR Snelling #06-006 (1w, RAJC);
|
||
<emphasis id="B93C2E2EFFADF473BEE9E701FCF1FE67" box="[713,865,368,392]" italics="true" pageId="16" pageNumber="217">Riverside Co.</emphasis>
|
||
:
|
||
<quantity id="4CB05FD9FFADF473BF50E71EFC21FE67" box="[880,945,367,391]" metricMagnitude="4" metricUnit="m" metricValue="3.3796224" pageId="16" pageNumber="217" unit="mi" value="21.0">21 mi</quantity>
|
||
E Indio, 1600’,
|
||
<date id="FFF6D4FCFFADF473B847E701FB60FE67" box="[1127,1264,367,392]" pageId="16" pageNumber="217" value="1952-04-08">Apr 8, 1952</date>
|
||
, WS Creighton CR-324 (19w
|
||
<typeStatus id="54F34C9EFFADF473BD17E7E5FE41FE4C" box="[311,465,404,428]" pageId="16" pageNumber="217" type="paratype">PARATYPES</typeStatus>
|
||
of
|
||
<emphasis id="B93C2E2EFFADF473BDD7E7E4FD2CFE4C" box="[503,700,404,428]" italics="true" pageId="16" pageNumber="217">
|
||
<taxonomicName id="4C4889BFFFADF473BDD7E7E4FD28FE4C" authorityName="Cole" authorityYear="1968" box="[503,696,404,428]" class="Insecta" family="Formicidae" genus="Pogonomyrmex" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="217" phylum="Arthropoda" rank="species" species="magnacanthus">P. magnacanthus</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
LACM).
|
||
<emphasis id="B93C2E2EFFADF473BF0CE7E4FB9CFE4C" box="[812,1036,404,428]" italics="true" pageId="16" pageNumber="217">San Bernardino Co.</emphasis>
|
||
: Needles,
|
||
<date id="FFF6D4FCFFADF473B8A3E7E4FB67FE4C" box="[1155,1271,404,429]" pageId="16" pageNumber="217" value="1905-05">May 1905</date>
|
||
, WM Wheeler (3w, LACM);
|
||
<quantity id="4CB05FD9FFADF473BD19E7C6FE10FE2F" box="[313,384,439,463]" metricMagnitude="3" metricUnit="m" metricValue="5.632704" pageId="16" pageNumber="217" unit="mi" value="3.5">3.5 mi</quantity>
|
||
N Pisgah Crater, 2270’,
|
||
<date id="FFF6D4FCFFADF473BEBAE7C9FCA9FE2F" box="[666,825,439,464]" pageId="16" pageNumber="217" value="2010-05-11">May 11, 2010</date>
|
||
, RA Johnson RAJ#4488 (6w, RAJC), RAJ#4489 (6w, RAJC);
|
||
<quantity id="4CB05FD9FFADF473BCD4E7ADFEACFE14" box="[244,316,476,500]" metricMagnitude="3" metricUnit="m" metricValue="8.851392" pageId="16" pageNumber="217" unit="mi" value="5.5">5.5 mi</quantity>
|
||
NW Pisgah Crater, 1890’,
|
||
<date id="FFF6D4FCFFADF473BE50E7ACFC93FE14" box="[624,771,476,501]" pageId="16" pageNumber="217" value="2010-05-04">May 4, 2010</date>
|
||
, RA Johnson RAJ#4493 (3w, RAJC), RAJ#4494 (5w, 1aq, RAJC);
|
||
<date id="FFF6D4FCFFADF473BCD8E78EFEB6FDF7" box="[248,294,511,535]" pageId="16" pageNumber="217" value="1940-01">I-40</date>
|
||
at Goffs Road, 2040’,
|
||
<date id="FFF6D4FCFFADF473BE1DE78EFD4AFDF7" box="[573,730,511,535]" pageId="16" pageNumber="217" value="2011-09-16">Sep 16, 2011</date>
|
||
, RA Johnson RAJ#4808 (6w, RAJC).
|
||
<emphasis id="B93C2E2EFFADF473B881E471FAD7FDF7" box="[1185,1351,512,536]" italics="true" pageId="16" pageNumber="217">San Diego Co</emphasis>
|
||
.: Anza Borrego State Park, Palm Canyon,
|
||
<date id="FFF6D4FCFFADF473BE3DE455FD24FDDC" box="[541,692,548,572]" pageId="16" pageNumber="217" value="1952-04-17">Apr 17, 1952</date>
|
||
, WS Creighton CR-559 (6w, LACM); Ocotillo Well,
|
||
<date id="FFF6D4FCFFADF473B92AE455FA08FDDC" box="[1290,1432,548,572]" pageId="16" pageNumber="217" value="1960-08-08">Aug 8, 1960</date>
|
||
, AC Cole CAL-366 (4w
|
||
<typeStatus id="54F34C9EFFADF473BD86E436FDD0FDBF" box="[422,576,583,607]" pageId="16" pageNumber="217" type="paratype">PARATYPES</typeStatus>
|
||
of
|
||
<emphasis id="B93C2E2EFFADF473BE46E438FCBCFD80" box="[614,812,584,608]" italics="true" pageId="16" pageNumber="217">
|
||
<taxonomicName id="4C4889BFFFADF473BE46E438FCB7FD80" authorityName="Cole" authorityYear="1968" box="[614,807,584,608]" class="Insecta" family="Formicidae" genus="Pogonomyrmex" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="217" phylum="Arthropoda" rank="species" species="magnacanthus">P. magnacanthus</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
LACM).
|
||
<emphasis id="B93C2E2EFFADF473BFBBE436FC62FD80" bold="true" box="[923,1010,583,608]" italics="true" pageId="16" pageNumber="217">Nevada</emphasis>
|
||
:
|
||
<emphasis id="B93C2E2EFFADF473B821E439FBFFFDBF" box="[1025,1135,584,608]" italics="true" pageId="16" pageNumber="217">Clark Co.</emphasis>
|
||
:
|
||
<quantity id="4CB05FD9FFADF473B85EE439FB21FDBF" box="[1150,1201,583,607]" metricMagnitude="4" metricUnit="m" metricValue="1.1265408" pageId="16" pageNumber="217" unit="mi" value="7.0">7 mi</quantity>
|
||
W Echo Bay, 1800’,
|
||
<date id="FFF6D4FCFFADF473BCB7E41DFEB6FD64" box="[151,294,620,644]" pageId="16" pageNumber="217" value="1976-04-05">Apr 5, 1976</date>
|
||
, G & J Wheeler NEV-725 (3w, LACM);
|
||
<quantity id="4CB05FD9FFADF473BF2BE41DFCD0FD64" box="[779,832,620,644]" metricMagnitude="3" metricUnit="m" metricValue="6.437376" pageId="16" pageNumber="217" unit="mi" value="4.0">4 mi</quantity>
|
||
SW Riverside, 1500’,
|
||
<date id="FFF6D4FCFFADF473B86DE41DFB4CFD64" box="[1101,1244,620,644]" pageId="16" pageNumber="217" value="1970-04-02">Apr 2, 1970</date>
|
||
, G & J Wheeler NEV#707 (3w, LACM);
|
||
<quantity id="4CB05FD9FFADF473BD93E4FEFE7DFD47" box="[435,493,655,679]" metricMagnitude="3" metricUnit="m" metricValue="5.0" pageId="16" pageNumber="217" unit="km" value="5.0">5 km</quantity>
|
||
E Jean,
|
||
<date id="FFF6D4FCFFADF473BE6EE4E1FD78FD47" box="[590,744,655,680]" pageId="16" pageNumber="217" value="2009-04-26">Apr 26, 2009</date>
|
||
, 2780’, RA Johnson RAJ#4222 (3w, RAJC), RAJ#4223 (3w, RAJC); Valley of Fire, 2100’,
|
||
<date id="FFF6D4FCFFADF473BDD9E4C4FD09FD2C" box="[505,665,692,716]" pageId="16" pageNumber="217" value="1970-03-14">Mar 14, 1970</date>
|
||
, G & J Wheeler NEV#640 (3w, LACM), NEV#631 (3w, LACM); Valley of Fire, 2000’, NEV#645 (2w, LACM); W base of Mormon Mesa, 1500’,
|
||
<date id="FFF6D4FCFFADF473B86CE4A9FB4CFD0F" box="[1100,1244,727,752]" pageId="16" pageNumber="217" value="1970-04-03">Apr 3, 1970</date>
|
||
, G & J Wheeler NEV#720 (3w, LACM), NEV#716 (3w, LACM);
|
||
<quantity id="4CB05FD9FFADF473BEE9E48DFD6CFCF4" box="[713,764,764,788]" metricMagnitude="3" metricUnit="m" metricValue="4.828032" pageId="16" pageNumber="217" unit="mi" value="3.0">3 mi</quantity>
|
||
W Cottonwood Cove, 1200’,
|
||
<date id="FFF6D4FCFFADF473B86EE48CFB77FCF4" box="[1102,1255,764,788]" pageId="16" pageNumber="217" value="1970-12-10">Dec 10, 1970</date>
|
||
, G & J Wheeler NEV-1498 (3w, LACM).
|
||
<emphasis id="B93C2E2EFFADF473BD9EE550FD8BFCD7" box="[446,539,800,824]" italics="true" pageId="16" pageNumber="217">Nye Co.</emphasis>
|
||
: Mercury,
|
||
<date id="FFF6D4FCFFADF473BEBBE56EFCBBFCD7" box="[667,811,799,823]" pageId="16" pageNumber="217" value="1961-07-11">Jul 11, 1961</date>
|
||
, #5AA1C (1w, LACM),
|
||
<date id="FFF6D4FCFFADF473B86EE550FB61FCD7" box="[1102,1265,799,825]" pageId="16" pageNumber="217" value="1961-05-24">May 24, 1961</date>
|
||
, #5EA7C (1w, LACM); Pahrump, spring 2000, KE Anderson KEA#532 (3w, RAJC);
|
||
<quantity id="4CB05FD9FFADF473BF90E535FC73FCBC" box="[944,995,836,860]" metricMagnitude="3" metricUnit="m" metricValue="3.218688" pageId="16" pageNumber="217" unit="mi" value="2.0">2 mi</quantity>
|
||
N Pahrump, 2700’,
|
||
<date id="FFF6D4FCFFADF473B8E6E535FADFFCBC" box="[1222,1359,836,860]" pageId="16" pageNumber="217" value="1970-04-04">Apr 4, 1970</date>
|
||
, G & J Wheeler NEV-740 (2w, LACM); Beatty,
|
||
<date id="FFF6D4FCFFADF473BE4AE516FD6AFC9F" box="[618,762,871,895]" pageId="16" pageNumber="217" value="1954-07-17">Jul 17, 1954</date>
|
||
, AC Cole, NEV-374 (6w, LACM), NEV-375 (4w, LACM), NEV-376 (6w, LACM), NEV-377 (22w, LACM), NEV-380 (2w, LACM); Hwy 95 at 4.0 mi NW Lathrop Wells, 2610’,
|
||
<date id="FFF6D4FCFFADF473BCC6E5C1FEECFC27" box="[230,380,943,968]" pageId="16" pageNumber="217" value="2009-04-17">Apr 17, 2009</date>
|
||
, RA Johnson RAJ#4216 (6w, RAJC), RAJ#4217 (9w, RAJC); Rock Valley,
|
||
<quantity id="4CB05FD9FFADF473B8EEE5DEFA90FC27" box="[1230,1280,943,967]" metricMagnitude="4" metricUnit="m" metricValue="1.4484096" pageId="16" pageNumber="217" unit="mi" value="9.0">9 mi</quantity>
|
||
ENE Lathrop Wells,
|
||
<date id="FFF6D4FCFFADF473BCC5E5A5FEEEFC0C" box="[229,382,980,1004]" pageId="16" pageNumber="217" value="1970-04-14">Apr 14, 1970</date>
|
||
, G & J Wheeler, NEV-781 (9w, LACM).
|
||
<emphasis id="B93C2E2EFFADF473BF75E5A2FC5BFC0C" bold="true" box="[853,971,979,1004]" pageId="16" pageNumber="217">
|
||
<collectingCountry id="F35FB2ACFFADF473BF75E5A2FC5BFC0C" box="[853,971,979,1004]" name="Mexico" pageId="16" pageNumber="217">MEXICO</collectingCountry>
|
||
</emphasis>
|
||
:
|
||
<emphasis id="B93C2E2EFFADF473BFFBE5A2FB01FC0C" bold="true" box="[987,1169,979,1004]" italics="true" pageId="16" pageNumber="217">Baja California</emphasis>
|
||
: Valle San Felipe,
|
||
<date id="FFF6D4FCFFADF473B949E5A4FF6DFBEF" pageId="16" pageNumber="217" value="1998-05-10">May 10, 1998</date>
|
||
, RA Johnson RAJ#BC 1376 (9w, RAJC), RAJ#BC 1377 (9w, RAJC).
|
||
<emphasis id="B93C2E2EFFADF473B800E586FBE4FBF0" bold="true" box="[1056,1140,1015,1040]" italics="true" pageId="16" pageNumber="217">Sonora</emphasis>
|
||
: Pinacate Desert, no date, 1982, IR López Moreno #F-11 (2w, LACM), #F-16 (8w, LACM); Puerto Peñasco, 50’,
|
||
<date id="FFF6D4FCFFADF473B846E26DFB61FBD4" box="[1126,1265,1052,1076]" pageId="16" pageNumber="217" value="1950-07-15">Jul 15, 1950</date>
|
||
, WS Creighton #308 (30w, LACM), #304 (3w, LACM);
|
||
<quantity id="4CB05FD9FFADF473BE51E24EFD58FBB7" box="[625,712,1087,1111]" metricMagnitude="4" metricUnit="m" metricValue="1.8507456" pageId="16" pageNumber="217" unit="mi" value="11.5">11.5 mi</quantity>
|
||
E Puerto Peñasco,
|
||
<date id="FFF6D4FCFFADF473BF89E24EFBD7FBB7" box="[937,1095,1087,1111]" pageId="16" pageNumber="217" value="2009-07-17">July 17, 2009</date>
|
||
, RA Johnson RAJ#4274 (9w, RAJC), RAJ#4278 (3w, RAJC), RAJ#4281 (9w, RAJC).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C352A1B7FFADF473BCE7E2F6FC6CFAEC" pageId="16" pageNumber="217" type="etymology">
|
||
<paragraph id="8BF7F23CFFADF473BCE7E2F6FC6CFAEC" blockId="16.[151,1437,151,1940]" pageId="16" pageNumber="217">
|
||
<emphasis id="B93C2E2EFFADF473BCE7E2F6FED7FB40" bold="true" box="[199,327,1159,1184]" pageId="16" pageNumber="217">
|
||
<emphasis id="B93C2E2EFFADF473BCE7E2F6FED0FB40" bold="true" box="[199,320,1159,1184]" italics="true" pageId="16" pageNumber="217">Etymology</emphasis>
|
||
.
|
||
</emphasis>
|
||
The specific epithet honors Prof. Dr. Bert Hölldobler, who was an “ant god” during the tenure of RAJ in graduate school and beyond—and who now is a good friend, colleague, collaborator, and supporter who is dedicated to understanding all aspects of ant biology. His continued child-like enthusiasm for learning about ants and his earnest interest in helping students invigorate all of those around him.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C352A1B7FFADF473BCE7E366FCFCF874" pageId="16" pageNumber="217" type="discussion">
|
||
<paragraph id="8BF7F23CFFADF473BCE7E366FB4EFA04" blockId="16.[151,1437,151,1940]" pageId="16" pageNumber="217">
|
||
<emphasis id="B93C2E2EFFADF473BCE7E366FED5FAD0" bold="true" box="[199,325,1303,1328]" italics="true" pageId="16" pageNumber="217">Discussion</emphasis>
|
||
.
|
||
<taxonomicName id="4C4889BFFFADF473BD73E368FD02FAD0" authorityName="Johnson, Overson & Moreau" authorityYear="2013" box="[339,658,1304,1328]" class="Insecta" family="Formicidae" genus="Pogonomyrmex" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="217" phylum="Arthropoda" rank="species" species="hoelldobleri">
|
||
<emphasis id="B93C2E2EFFADF473BD73E368FD02FAD0" box="[339,658,1304,1328]" italics="true" pageId="16" pageNumber="217">Pogonomyrmex hoelldobleri</emphasis>
|
||
</taxonomicName>
|
||
is most likely to be confused with
|
||
<taxonomicName id="4C4889BFFFADF473B83AE368FB48FAD0" authorityName="Cole" authorityYear="1968" box="[1050,1240,1304,1328]" class="Insecta" family="Formicidae" genus="Pogonomyrmex" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="217" phylum="Arthropoda" rank="species" species="magnacanthus">
|
||
<emphasis id="B93C2E2EFFADF473B83AE368FB48FAD0" box="[1050,1240,1304,1328]" italics="true" pageId="16" pageNumber="217">P. magnacanthus</emphasis>
|
||
</taxonomicName>
|
||
, especially given the large number of series that A.C. Cole erroneously identified as
|
||
<taxonomicName id="4C4889BFFFADF473BF9CE34CFB10FAB4" authorityName="Cole" authorityYear="1968" box="[956,1152,1340,1364]" class="Insecta" family="Formicidae" genus="Pogonomyrmex" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="217" phylum="Arthropoda" rank="species" species="magnacanthus">
|
||
<emphasis id="B93C2E2EFFADF473BF9CE34CFB10FAB4" box="[956,1152,1340,1364]" italics="true" pageId="16" pageNumber="217">P. magnacanthus</emphasis>
|
||
</taxonomicName>
|
||
. The two species occur sympatrically in several locales. The significantly larger eye (MOD and OI) separates
|
||
<taxonomicName id="4C4889BFFFADF473B85CE310FAAEFA98" authorityName="Cole" authorityYear="1968" box="[1148,1342,1376,1400]" class="Insecta" family="Formicidae" genus="Pogonomyrmex" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="217" phylum="Arthropoda" rank="species" species="magnacanthus">
|
||
<emphasis id="B93C2E2EFFADF473B85CE310FAAEFA98" box="[1148,1342,1376,1400]" italics="true" pageId="16" pageNumber="217">P. magnacanthus</emphasis>
|
||
</taxonomicName>
|
||
from
|
||
<taxonomicName id="4C4889BFFFADF473B9A7E310FEB0FA7C" authorityName="Johnson, Overson & Moreau" authorityYear="2013" class="Insecta" family="Formicidae" genus="Pogonomyrmex" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="217" phylum="Arthropoda" rank="species" species="hoelldobleri">
|
||
<emphasis id="B93C2E2EFFADF473B9A7E310FEB0FA7C" italics="true" pageId="16" pageNumber="217">P. hoelldobleri</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="1373EEB9FFADF473BD0FE3F5FE24FA7C" box="[303,436,1412,1436]" captionStart-0="FIGURE 2" captionStart-1="FIGURE 3" captionStartId-0="6.[151,250,1724,1747]" captionStartId-1="7.[151,250,1916,1939]" captionTargetBox-0="[171,1402,193,1704]" captionTargetBox-1="[306,1293,185,1892]" captionTargetId-0="figure@6.[169,1404,193,1707]" captionTargetId-1="figure@7.[297,1296,139,1895]" captionTargetPageId-0="6" captionTargetPageId-1="7" captionText-0="FIGURE 2. Photographs of diagnostic characters to distinguish between workers of P. magnacanthus Cole and those of P. hoelldobleri Johnson, Overson & Moreau and P. m o h a v e n s i s Johnson. Photograph of P. magnacanthus PARATYPE worker: (A) eye and malar area (MOD = 0.44, OI = 29.5, MR = 0.98), and circumocular whorls indistinct to absent, moderately granulate toward vertex, and (B) circumocular whorls present. Photograph of P. hoelldobleri worker: (C) eye and malar area for HOLOTYPE worker (MOD = 0.36, OI = 23.2, MR = 1.17), and absence of circumocular whorls—rugae converging near vertex, and (D) absence of circumocular whorls—area posterior to eyes with faint rugae. Photograph of P. m o h a v e n s i s worker: (E) eye and malar area for PARATYPE worker (MOD = 0.33, OI = 21.7, MR = 1.36), and absence of circumocular whorlsrugae extending to vertex, with rugae becoming weak to absent on / near vertex, and (F) absence of circumocular whorls—rugae extending to vertex; vertex rugose." captionText-1="FIGURE 3. Bivariate plots for workers: (A) maximum eye diameter versus head width, (B) ocular index versus head width, and (C) malar ratio versus head width (n = 63 for P. magnacanthus Cole, n = 59 for P. hoelldobleri Johnson, Overson & Moreau, plus 16 PARATYPE workers of P. magnacanthus that do not belong to the latter species [see text], n = 25 for P. mohavensis Johnson). Non-type workers were selected to represent the geographic range of each species." httpUri-0="https://zenodo.org/record/222900/files/figure.png" httpUri-1="https://zenodo.org/record/222901/files/figure.png" pageId="16" pageNumber="217">Figures 2-3</figureCitation>
|
||
). OI is the best character to separate the two species because it is consistently higher for
|
||
<taxonomicName id="4C4889BFFFADF473BCB7E3D8FEC9FA20" authorityName="Cole" authorityYear="1968" box="[151,345,1448,1472]" class="Insecta" family="Formicidae" genus="Pogonomyrmex" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="217" phylum="Arthropoda" rank="species" species="magnacanthus">
|
||
<emphasis id="B93C2E2EFFADF473BCB7E3D8FEC9FA20" box="[151,345,1448,1472]" italics="true" pageId="16" pageNumber="217">P. magnacanthus</emphasis>
|
||
</taxonomicName>
|
||
(OI = 27.22-33.61) than for
|
||
<emphasis id="B93C2E2EFFADF473BE91E3D8FCC5FA20" box="[689,853,1448,1472]" italics="true" pageId="16" pageNumber="217">P. h o e l l d o b l e r i</emphasis>
|
||
(OI rarely> 27.50)(
|
||
<figureCitation id="1373EEB9FFADF473B86CE3D6FB21FA5F" box="[1100,1201,1447,1471]" captionStart="FIGURE 3" captionStartId="7.[151,250,1916,1939]" captionTargetBox="[306,1293,185,1892]" captionTargetId="figure@7.[297,1296,139,1895]" captionTargetPageId="7" captionText="FIGURE 3. Bivariate plots for workers: (A) maximum eye diameter versus head width, (B) ocular index versus head width, and (C) malar ratio versus head width (n = 63 for P. magnacanthus Cole, n = 59 for P. hoelldobleri Johnson, Overson & Moreau, plus 16 PARATYPE workers of P. magnacanthus that do not belong to the latter species [see text], n = 25 for P. mohavensis Johnson). Non-type workers were selected to represent the geographic range of each species." httpUri="https://zenodo.org/record/222901/files/figure.png" pageId="16" pageNumber="217">Figure 3</figureCitation>
|
||
). The malar ratio is usually <1.0 for
|
||
<taxonomicName id="4C4889BFFFADF473BD77E3BCFD86FA04" authorityName="Cole" authorityYear="1968" box="[343,534,1484,1508]" class="Insecta" family="Formicidae" genus="Pogonomyrmex" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="217" phylum="Arthropoda" rank="species" species="magnacanthus">
|
||
<emphasis id="B93C2E2EFFADF473BD77E3BCFD86FA04" box="[343,534,1484,1508]" italics="true" pageId="16" pageNumber="217">P. magnacanthus</emphasis>
|
||
</taxonomicName>
|
||
, while this ratio is usually> 1.05 for
|
||
<taxonomicName id="4C4889BFFFADF473BF9BE3BCFBCFFA04" authorityName="Johnson, Overson & Moreau" authorityYear="2013" box="[955,1119,1484,1508]" class="Insecta" family="Formicidae" genus="Pogonomyrmex" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="217" phylum="Arthropoda" rank="species" species="hoelldobleri">
|
||
<emphasis id="B93C2E2EFFADF473BF9BE3BCFBCFFA04" box="[955,1119,1484,1508]" italics="true" pageId="16" pageNumber="217">P. hoelldobleri</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="1373EEB9FFADF473B84EE3BDFB40FA04" box="[1134,1232,1484,1508]" captionStart="FIGURE 3" captionStartId="7.[151,250,1916,1939]" captionTargetBox="[306,1293,185,1892]" captionTargetId="figure@7.[297,1296,139,1895]" captionTargetPageId="7" captionText="FIGURE 3. Bivariate plots for workers: (A) maximum eye diameter versus head width, (B) ocular index versus head width, and (C) malar ratio versus head width (n = 63 for P. magnacanthus Cole, n = 59 for P. hoelldobleri Johnson, Overson & Moreau, plus 16 PARATYPE workers of P. magnacanthus that do not belong to the latter species [see text], n = 25 for P. mohavensis Johnson). Non-type workers were selected to represent the geographic range of each species." httpUri="https://zenodo.org/record/222901/files/figure.png" pageId="16" pageNumber="217">Figure 3</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BF7F23CFFADF473BCE7E380FCFCF874" blockId="16.[151,1437,151,1940]" pageId="16" pageNumber="217">
|
||
<taxonomicName id="4C4889BFFFADF473BCE7E380FD99F9E8" authorityName="Johnson, Overson & Moreau" authorityYear="2013" box="[199,521,1520,1544]" class="Insecta" family="Formicidae" genus="Pogonomyrmex" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="217" phylum="Arthropoda" rank="species" species="hoelldobleri">
|
||
<emphasis id="B93C2E2EFFADF473BCE7E380FD99F9E8" box="[199,521,1520,1544]" italics="true" pageId="16" pageNumber="217">Pogonomyrmex hoelldobleri</emphasis>
|
||
</taxonomicName>
|
||
also occurs in sympatry with
|
||
<taxonomicName id="4C4889BFFFADF473BF48E380FB9CF9E8" baseAuthorityName="Buckley" baseAuthorityYear="1867" box="[872,1036,1520,1544]" class="Insecta" family="Formicidae" genus="Pogonomyrmex" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="217" phylum="Arthropoda" rank="species" species="californicus">
|
||
<emphasis id="B93C2E2EFFADF473BF48E380FB9CF9E8" box="[872,1036,1520,1544]" italics="true" pageId="16" pageNumber="217">P. californicus</emphasis>
|
||
</taxonomicName>
|
||
and is likely to occur in sympatry with
|
||
<taxonomicName id="4C4889BFFFADF473BCF0E064FEFFF9CC" box="[208,367,1556,1580]" class="Insecta" family="Formicidae" genus="Pogonomyrmex" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="217" phylum="Arthropoda" rank="species" species="mohavensis">
|
||
<emphasis id="B93C2E2EFFADF473BCF0E064FEFFF9CC" box="[208,367,1556,1580]" italics="true" pageId="16" pageNumber="217">P. mohavensis</emphasis>
|
||
</taxonomicName>
|
||
, but it has a low likelihood of co-occurring with
|
||
<emphasis id="B93C2E2EFFADF473BFB8E064FB8BF9CC" box="[920,1051,1557,1580]" italics="true" pageId="16" pageNumber="217">P. m a r i c o p a</emphasis>
|
||
.
|
||
<taxonomicName id="4C4889BFFFADF473B80EE064FAFEF9CC" authorityName="Johnson, Overson & Moreau" authorityYear="2013" box="[1070,1390,1556,1580]" class="Insecta" family="Formicidae" genus="Pogonomyrmex" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="217" phylum="Arthropoda" rank="species" species="hoelldobleri">
|
||
<emphasis id="B93C2E2EFFADF473B80EE064FAFEF9CC" box="[1070,1390,1556,1580]" italics="true" pageId="16" pageNumber="217">Pogonomyrmex hoelldobleri</emphasis>
|
||
</taxonomicName>
|
||
can be distinguished from
|
||
<taxonomicName id="4C4889BFFFADF473BDB4E048FDA3F9B0" box="[404,563,1592,1616]" class="Insecta" family="Formicidae" genus="Pogonomyrmex" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="217" phylum="Arthropoda" rank="species" species="mohavensis">
|
||
<emphasis id="B93C2E2EFFADF473BDB4E048FDA3F9B0" box="[404,563,1592,1616]" italics="true" pageId="16" pageNumber="217">P. mohavensis</emphasis>
|
||
</taxonomicName>
|
||
based on the following characters: (1)
|
||
<taxonomicName id="4C4889BFFFADF473BFCCE048FB01F9B0" authorityName="Johnson, Overson & Moreau" authorityYear="2013" box="[1004,1169,1592,1616]" class="Insecta" family="Formicidae" genus="Pogonomyrmex" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="217" phylum="Arthropoda" rank="species" species="hoelldobleri">
|
||
<emphasis id="B93C2E2EFFADF473BFCCE048FB01F9B0" box="[1004,1169,1592,1616]" italics="true" pageId="16" pageNumber="217">P. hoelldobleri</emphasis>
|
||
</taxonomicName>
|
||
has seven teeth, (2) the interrugal spaces on the pronotal shoulders are weakly to strongly punctate/granulate, dull to sub-shining, and (3) the cephalic rugae typically converge near the vertex. In
|
||
<taxonomicName id="4C4889BFFFADF473BF15E0F0FC47F978" box="[821,983,1664,1688]" class="Insecta" family="Formicidae" genus="Pogonomyrmex" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="217" phylum="Arthropoda" rank="species" species="mohavensis">
|
||
<emphasis id="B93C2E2EFFADF473BF15E0F0FC47F978" box="[821,983,1664,1688]" italics="true" pageId="16" pageNumber="217">P. mohavensis</emphasis>
|
||
</taxonomicName>
|
||
, the mandible has six teeth (a seventh sometimes occurs as a denticle between the basal and sub-basal teeth), interrugal spaces on the pronotal shoulders are smooth and shining, and the cephalic rugae extend more or less directly to the vertex or converge only slightly near the vertex. Two other
|
||
<taxonomicName id="4C4889BFFFADF473BDE3E09CFDF5F8E4" baseAuthorityName="Buckley" baseAuthorityYear="1867" box="[451,613,1772,1796]" class="Insecta" family="Formicidae" genus="Pogonomyrmex" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="217" phylum="Arthropoda" rank="species" species="californicus">
|
||
<emphasis id="B93C2E2EFFADF473BDE3E09CFDF5F8E4" box="[451,613,1772,1796]" italics="true" pageId="16" pageNumber="217">P. californicus</emphasis>
|
||
</taxonomicName>
|
||
group species (
|
||
<taxonomicName id="4C4889BFFFADF473BF34E09CFC1EF8E4" authorityName="Cole" authorityYear="1968" box="[788,910,1773,1796]" class="Insecta" family="Formicidae" genus="Pogonomyrmex" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="217" phylum="Arthropoda" rank="species" species="anzensis">
|
||
<emphasis id="B93C2E2EFFADF473BF34E09CFC1EF8E4" box="[788,910,1773,1796]" italics="true" pageId="16" pageNumber="217">P. anzensis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C4889BFFFADF473BFE5E09CFBD0F8E4" authorityName="Taber" authorityYear="1998" box="[965,1088,1772,1796]" class="Insecta" family="Formicidae" genus="Pogonomyrmex" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="217" phylum="Arthropoda" rank="species" species="snellingi">
|
||
<emphasis id="B93C2E2EFFADF473BFE5E09CFBD0F8E4" box="[965,1088,1772,1796]" italics="true" pageId="16" pageNumber="217">P. snellingi</emphasis>
|
||
</taxonomicName>
|
||
) also occur in the Sonoran and Mohave Deserts, but it is doubtful that
|
||
<taxonomicName id="4C4889BFFFADF473BE72E160FD66F8C8" authorityName="Johnson, Overson & Moreau" authorityYear="2013" box="[594,758,1808,1832]" class="Insecta" family="Formicidae" genus="Pogonomyrmex" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="217" phylum="Arthropoda" rank="species" species="hoelldobleri">
|
||
<emphasis id="B93C2E2EFFADF473BE72E160FD66F8C8" box="[594,758,1808,1832]" italics="true" pageId="16" pageNumber="217">P. hoelldobleri</emphasis>
|
||
</taxonomicName>
|
||
occurs sympatrically with either species;
|
||
<taxonomicName id="4C4889BFFFADF473B8F0E160FADBF8C8" authorityName="Cole" authorityYear="1968" box="[1232,1355,1809,1832]" class="Insecta" family="Formicidae" genus="Pogonomyrmex" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="217" phylum="Arthropoda" rank="species" species="anzensis">
|
||
<emphasis id="B93C2E2EFFADF473B8F0E160FADBF8C8" box="[1232,1355,1809,1832]" italics="true" pageId="16" pageNumber="217">P. anzensis</emphasis>
|
||
</taxonomicName>
|
||
occurs in unproductive, rocky hillside habitats unlike any sites that are known to be occupied by
|
||
<taxonomicName id="4C4889BFFFADF473B8AEE144FAA3F8AC" authorityName="Johnson, Overson & Moreau" authorityYear="2013" box="[1166,1331,1844,1868]" class="Insecta" family="Formicidae" genus="Pogonomyrmex" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="217" phylum="Arthropoda" rank="species" species="hoelldobleri">
|
||
<emphasis id="B93C2E2EFFADF473B8AEE144FAA3F8AC" box="[1166,1331,1844,1868]" italics="true" pageId="16" pageNumber="217">P. hoelldobleri</emphasis>
|
||
</taxonomicName>
|
||
, while
|
||
<taxonomicName id="4C4889BFFFADF473B9A7E144FF67F890" authorityName="Taber" authorityYear="1998" class="Insecta" family="Formicidae" genus="Pogonomyrmex" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="217" phylum="Arthropoda" rank="species" species="snellingi">
|
||
<emphasis id="B93C2E2EFFADF473B9A7E144FF67F890" italics="true" pageId="16" pageNumber="217">P. snellingi</emphasis>
|
||
</taxonomicName>
|
||
is well removed from the probable geographic distribution of
|
||
<taxonomicName id="4C4889BFFFADF473BF9CE128FBF1F890" authorityName="Johnson, Overson & Moreau" authorityYear="2013" box="[956,1121,1880,1904]" class="Insecta" family="Formicidae" genus="Pogonomyrmex" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="217" phylum="Arthropoda" rank="species" species="hoelldobleri">
|
||
<emphasis id="B93C2E2EFFADF473BF9CE128FBF1F890" box="[956,1121,1880,1904]" italics="true" pageId="16" pageNumber="217">P. hoelldobleri</emphasis>
|
||
</taxonomicName>
|
||
. Regardless, the absence of circumocular whorls separates
|
||
<taxonomicName id="4C4889BFFFADF473BDD2E10CFD07F874" authorityName="Johnson, Overson & Moreau" authorityYear="2013" box="[498,663,1916,1940]" class="Insecta" family="Formicidae" genus="Pogonomyrmex" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="217" phylum="Arthropoda" rank="species" species="hoelldobleri">
|
||
<emphasis id="B93C2E2EFFADF473BDD2E10CFD07F874" box="[498,663,1916,1940]" italics="true" pageId="16" pageNumber="217">P. hoelldobleri</emphasis>
|
||
</taxonomicName>
|
||
from both species.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |