treatments-xml/data/03/DF/BE/03DFBE05E025FFC9FF64D145FC41FB48.xml

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<mods:title id="64D8276BE1BA9997C2D479496E017B88">Review of the genus Beckhoplia Dombrow, 2005 with the description of fifteen new species from South Africa and observations on its biogeography (Coleoptera: Scarabaeidae: Melolonthinae: Hopliini)</mods:title>
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<mods:namePart id="75621EDAA9B4EA201201570286AAE5EA">Dombrow, Holger E.</mods:namePart>
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<mods:namePart id="52A2E9C401EB6D706D9FFC727538F083">Colville, Jonathan F.</mods:namePart>
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<mods:date id="08D4F357F6415D1D042DF4873F806E94">2020</mods:date>
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<subSubSection id="C36C5C98E025FFCCFF64D145FD0EFAF5" box="[151,691,1374,1401]" pageId="57" pageNumber="58" type="nomenclature">
<paragraph id="8BC90F13E025FFCCFF64D145FD0EFAF5" blockId="57.[151,854,1374,1472]" box="[151,691,1374,1401]" pageId="57" pageNumber="58">
<heading id="D081B87FE025FFCCFF64D145FD0EFAF5" bold="true" box="[151,691,1374,1401]" fontSize="11" level="2" pageId="57" pageNumber="58" reason="4">
<emphasis id="B902D301E025FFCCFF64D145FD0EFAF5" bold="true" box="[151,691,1374,1401]" pageId="57" pageNumber="58">
<taxonomicName id="4C767490E025FFCCFF64D145FDABFAF4" authority="Dombrow" authorityName="Dombrow" authorityYear="2020" box="[151,534,1374,1401]" class="Insecta" family="Melolonthidae" genus="Beckhoplia" kingdom="Animalia" order="Coleoptera" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="suturalis" status="sp. nov.">
<emphasis id="B902D301E025FFCCFF64D145FE2CFAF4" bold="true" box="[151,401,1374,1401]" italics="true" pageId="57" pageNumber="58">Beckhoplia suturalis</emphasis>
Dombrow
</taxonomicName>
, 
<taxonomicNameLabel id="A2316E7AE025FFCCFDD0D144FD0EFAF5" box="[547,691,1374,1401]" pageId="57" pageNumber="58" rank="species">new species</taxonomicNameLabel>
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<paragraph id="8BC90F13E025FFCCFF64D198FE32FA10" blockId="57.[151,854,1374,1472]" box="[151,399,1411,1437]" pageId="57" pageNumber="58">
(
<figureCitation id="134D1396E025FFCCFF6CD198FF40FA10" box="[159,253,1411,1437]" captionStart="FIGURES 29–30" captionStartId="16.[151,264,1974,1998]" captionTargetBox="[467,1125,1593,1907]" captionTargetId="figure-177@16.[458,1129,1590,1914]" captionTargetPageId="16" captionText="FIGURES 29–30. General drawings of the shape of the elytra of Beckhoplia Dombrow, 2005 species, ♂♂. 29, ovate; 30, oblong Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4401934" httpUri="https://zenodo.org/record/4401934/files/figure.png" pageId="57" pageNumber="58">Figs. 29</figureCitation>
, 
<figureCitation id="134D1396E025FFCCFEFAD198FEEFFA10" box="[265,338,1411,1437]" captionStart="FIGURES 62–97" captionStartId="28.[151,264,1743,1767]" captionTargetBox="[161,1426,187,1710]" captionTargetId="figure-25@28.[151,1436,181,1719]" captionTargetPageId="28" captionText="FIGURES 62–97. Parameres and phallobase of Beckhoplia Dombrow, 2005 species, in dorsal view (left side), in lateral view (right side), ♂♂. 62–63, B. nigra Dombrow, new species; 64–65, B. pallidibrunnea Dombrow, new species; 66–67, B. fusca Dombrow, new species; 68–69, B. setosa Dombrow, new species; 70–71, B. castanea Dombrow, new species; 72–73, B. colvillei Dombrow, 2005; 74–75, B. pulchra Dombrow, new species; 76–77, B. nigrofasciata Dombrow, new species; 78–79, B. caliginosa Dombrow, new species; 80–81, B. pumila Dombrow, new species; 82–83, B. gifbergensis Dombrow, new species; 84–85, B. julianae Dombrow, 2005; 86–87, B. elkeae Dombrow, new species; 88–89, B. suturalis Dombrow, new species; 90–91, B. nigrosetosa Dombrow, new species; 92–93, B. occidentalis Dombrow, 2005; 94–95, B. bicolor Dombrow, new species; 96–97, B. dolichiocnemis Dombrow, new species; 98–99, Anisochelus inornatus Burmeister, 1844. Scale bar 1mm." figureDoi="http://doi.org/10.5281/zenodo.4401954" httpUri="https://zenodo.org/record/4401954/files/figure.png" pageId="57" pageNumber="58">88–89</figureCitation>
, 
<figureCitation id="134D1396E025FFCCFEAED198FE3AFA10" box="[349,391,1411,1437]" captionStart="FIGURES 100–117" captionStartId="42.[151,264,1665,1689]" captionTargetBox="[162,1424,188,1625]" captionTargetId="figure-25@42.[151,1436,181,1641]" captionTargetPageId="42" captionText="FIGURES 100–117. Detailed drawings of the parameres for the key and the differential diagnosis of Beckhoplia Dombrow, 2005 species, in dorsal view, ♂♂. Arrows indicate characters of importance. 100, B. nigra Dombrow, new species; 101, B. pallidibrunnea Dombrow, new species; 102, B. fusca Dombrow, new species; 103, B. setosa Dombrow, new species; 104, B. castanea Dombrow, new species; 105, B. colvillei Dombrow, 2005; 106, B. pulchra Dombrow, new species; 107, B. nigrofasciata Dombrow, new species; 108, B. caliginosa Dombrow, new species; 109, B. pumila Dombrow, new species; 110, B. gifbergensis Dombrow, new species; 111, B. julianae Dombrow, 2005; 112, B. elkeae Dombrow, new species; 113, B. suturalis Dombrow, new species; 114, B. nigrosetosa Dombrow, new species; 115, B. occidentalis Dombrow, 2005; 116, B. bicolor Dombrow, new species; 117, B. dolichiocnemis Dombrow sp.n. Scale bar 1mm." figureDoi="http://doi.org/10.5281/zenodo.4401956" httpUri="https://zenodo.org/record/4401956/files/figure.png" pageId="57" pageNumber="58">113</figureCitation>
)
</paragraph>
<paragraph id="8BC90F13E025FFCCFF64D1B3FCEBFA4D" blockId="57.[151,854,1374,1472]" box="[151,854,1448,1472]" pageId="57" pageNumber="58">
http://zoobank.org/ 
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</paragraph>
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<subSubSection id="C36C5C98E025FFCCFF64D1F6FD23F989" box="[151,670,1517,1541]" pageId="57" pageNumber="58" type="reference_group">
<paragraph id="8BC90F13E025FFCCFF64D1F6FD23F989" blockId="57.[151,670,1517,1541]" box="[151,670,1517,1541]" pageId="57" pageNumber="58">
<treatmentCitationGroup id="AB66283DE025FFCCFF64D1F6FD23F989" box="[151,670,1517,1541]" pageId="57" pageNumber="58">
<treatmentCitation id="0AD72902E025FFCCFF64D1F6FD82F988" author="Dombrow, H." box="[151,575,1517,1541]" page="128" pageId="57" pageNumber="58" year="2005">
<taxonomicName id="4C767490E025FFCCFF64D1F6FD82F988" authority="Dombrow, 2005: 128" authorityName="Dombrow" authorityPageNumber="128" authorityYear="2005" box="[151,575,1517,1541]" class="Insecta" family="Melolonthidae" genus="Beckhoplia" kingdom="Animalia" order="Coleoptera" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="colvillei">
<emphasis id="B902D301E025FFCCFF64D1F6FEDCF988" box="[151,353,1517,1541]" italics="true" pageId="57" pageNumber="58">Beckhoplia colvillei</emphasis>
<bibRefCitation id="EFE772E2E025FFCCFE94D1F6FD82F988" author="Dombrow, H." box="[359,575,1517,1541]" pageId="57" pageNumber="58" pagination="125 - 130" refId="ref61191" refString="Dombrow, H. (2005) Beckhoplia a new genus of Hopliini from the Northern Cape of the Republic of South Africa. (Coleoptera: Scarabaeidae: Melolonthinae: Hopliini). Coleoptera, 9, 125 - 130." type="journal article" year="2005">Dombrow, 2005: 128</bibRefCitation>
</taxonomicName>
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ex parte.
</treatmentCitationGroup>
</paragraph>
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<subSubSection id="C36C5C98E025FFCFFF64D228FB04FF3C" lastPageId="58" lastPageNumber="59" pageId="57" pageNumber="58" type="materials_examined">
<paragraph id="8BC90F13E025FFCCFF64D228FD32F9C0" blockId="57.[151,655,1586,1613]" box="[151,655,1586,1613]" pageId="57" pageNumber="58">
<materialsCitation id="3B1E054EE025FFCCFF64D228FD36F9C0" ID-GBIF-Occurrence="3022772313" box="[151,651,1586,1613]" collectionCode="NCP" country="South Africa" location="RSA" pageId="57" pageNumber="58" specimenCount="1" typeStatus="holotype">
<emphasis id="B902D301E025FFCCFF64D228FE8BF9C0" bold="true" box="[151,310,1587,1613]" pageId="57" pageNumber="58">
<typeStatus id="54CDB1B1E025FFCCFF64D228FF6FF9C0" box="[151,210,1587,1613]" pageId="57" pageNumber="58">Type</typeStatus>
locality.
</emphasis>
[
<collectingCountry id="F3614F83E025FFCCFEB0D228FEC2F9C0" box="[323,383,1587,1613]" name="South Africa" pageId="57" pageNumber="58">RSA</collectingCountry>
, 
<collectionCode id="ED6797D6E025FFCCFE79D229FE7AF9C0" box="[394,455,1586,1613]" country="United Kingdom" name="National Collection of Passiflora" pageId="57" pageNumber="58" type="Herbarium">NCP</collectionCode>
]: Nieuwoudtville
</materialsCitation>
.
</paragraph>
<paragraph id="8BC90F13E025FFCCFF64D260FC5AF854" blockId="57.[151,1437,1658,2010]" pageId="57" pageNumber="58">
<emphasis id="B902D301E025FFCCFF64D260FEFAF918" bold="true" box="[151,327,1659,1685]" pageId="57" pageNumber="58">Type material.</emphasis>
[49] 
<emphasis id="B902D301E025FFCCFE7BD260FE48F918" bold="true" box="[392,501,1659,1685]" pageId="57" pageNumber="58">
<typeStatus id="54CDB1B1E025FFCCFE7BD260FE48F918" box="[392,501,1659,1685]" pageId="57" pageNumber="58" type="holotype">Holotype</typeStatus>
</emphasis>
<specimenCount id="9D70C49AE025FFCCFE0DD267FDABF91B" box="[510,534,1660,1686]" pageId="57" pageNumber="58" type="male">♂</specimenCount>
: white label pr « 
<collectingCountry id="F3614F83E025FFCCFD2BD260FC14F918" box="[728,937,1659,1685]" name="South Africa" pageId="57" pageNumber="58">SOUTH AFRICA</collectingCountry>
, Northern | 
<collectingRegion id="49B2C1F1E025FFCCFBC0D260FBD2F918" box="[1075,1135,1659,1685]" country="South Africa" name="Western Cape" pageId="57" pageNumber="58">Cape</collectingRegion>
, Hantam | Nieuwoudtville at | 31.22.33S–19.07.43E» // pr «Time 12:30, 15. Sept. 2001 | alt. 
<quantity id="4C8EA2F6E025FFCCFC74D284FC6FF937" box="[903,978,1695,1722]" metricMagnitude="2" metricUnit="m" metricValue="7.33" pageId="57" pageNumber="58" unit="m" value="733.0">733 m</quantity>
, 
<taxonomicName id="4C767490E025FFCCFC2CD284FBC7F934" box="[991,1146,1695,1721]" class="Liliopsida" family="Iridaceae" kingdom="Plantae" order="Asparagales" pageId="57" pageNumber="58" phylum="Tracheophyta" rank="family">IRIDACEAE</taxonomicName>
: | 
<taxonomicName id="4C767490E025FFCCFB67D2BBFAE8F934" authorityName="Sweet" authorityYear="1826" baseAuthorityName="Andrews" box="[1172,1365,1696,1721]" class="Liliopsida" family="Iridaceae" genus="Homeria" higherTaxonomySource="CoL" kingdom="Plantae" order="Asparagales" pageId="57" pageNumber="58" phylum="Tracheophyta" rank="species" species="miniata">
<emphasis id="B902D301E025FFCCFB67D2BBFAE8F934" box="[1172,1365,1696,1721]" italics="true" pageId="57" pageNumber="58">Homeria miniata</emphasis>
</taxonomicName>
| leg.: Holger Dombrow #327» || yellow label hw «18» pr «.485 | coll. Dombrow» || red label pr « 
<typeStatus id="54CDB1B1E025FFCCFB6BD2DFFB41F953" box="[1176,1276,1732,1758]" pageId="57" pageNumber="58" type="paratype">Paratype</typeStatus>
| 
<taxonomicName id="4C767490E025FFCCFAE2D2D8FA32F950" authorityName="Dombrow" authorityYear="2005" box="[1297,1423,1731,1757]" class="Insecta" family="Melolonthidae" genus="Beckhoplia" kingdom="Animalia" order="Coleoptera" pageId="57" pageNumber="58" phylum="Arthropoda" rank="genus">
<emphasis id="B902D301E025FFCCFAE2D2D8FA32F950" box="[1297,1423,1731,1757]" italics="true" pageId="57" pageNumber="58">Beckhoplia</emphasis>
</taxonomicName>
| 
<taxonomicName id="4C767490E025FFCCFF64D2FCFF4CF88C" authority="Dombrow &amp; Colville, 2020" authorityName="Dombrow" authorityYear="2020" box="[151,241,1767,1793]" class="Insecta" family="Melolonthidae" genus="Beckhoplia" kingdom="Animalia" order="Coleoptera" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="colvillei" status="sp. nov.">
<emphasis id="B902D301E025FFCCFF64D2FCFF4CF88C" box="[151,241,1767,1793]" italics="true" pageId="57" pageNumber="58">colvillei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2316E7AE025FFCCFF0AD2F3FE93F88F" box="[249,302,1768,1794]" pageId="57" pageNumber="58" rank="species">sp.n.</taxonomicNameLabel>
| det.: H. Dombrow 2003» || blue label pr «Bioregion: | Shale Renosterveld | Vegetation Type: FRs2 – | Nieuwoudtville Shale Renosterveld» || yellow label pr «SAM–COL | A071220» || red label pr « 
<typeStatus id="54CDB1B1E025FFCCFB2CD310FAF4F8A8" box="[1247,1353,1803,1829]" pageId="57" pageNumber="58" type="holotype">Holotype</typeStatus>
| 
<taxonomicName id="4C767490E025FFCCFAAED310FF62F8C4" authorityName="Dombrow" authorityYear="2005" class="Insecta" family="Melolonthidae" genus="Beckhoplia" kingdom="Animalia" order="Coleoptera" pageId="57" pageNumber="58" phylum="Arthropoda" rank="genus">
<emphasis id="B902D301E025FFCCFAAED310FF62F8C4" italics="true" pageId="57" pageNumber="58">Beckhoplia</emphasis>
</taxonomicName>
| 
<taxonomicName id="4C767490E025FFCCFF06D334FEEAF8C4" authority="Dombrow &amp; Colville, 2020" authorityName="Dombrow" authorityYear="2020" box="[245,343,1839,1865]" class="Insecta" family="Melolonthidae" genus="Beckhoplia" kingdom="Animalia" order="Coleoptera" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="suturalis" status="sp. nov.">
<emphasis id="B902D301E025FFCCFF06D334FEEAF8C4" box="[245,343,1839,1865]" italics="true" pageId="57" pageNumber="58">suturalis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2316E7AE025FFCCFEACD32BFE29F8C7" box="[351,404,1840,1866]" pageId="57" pageNumber="58" rank="species">sp.n.</taxonomicNameLabel>
| det.: H. Dombrow 2019» (
<collectionCode id="ED6797D6E025FFCCFD26D334FC96F8C4" box="[725,811,1839,1865]" country="South Africa" httpUri="http://grbio.org/cool/94p5-pepz" name="Iziko Museums of Cape Town" pageId="57" pageNumber="58" type="Museum">SAMC</collectionCode>
). – 
<emphasis id="B902D301E025FFCCFCA6D334FC70F8C4" bold="true" box="[853,973,1839,1865]" pageId="57" pageNumber="58">
<typeStatus id="54CDB1B1E025FFCCFCA6D334FC70F8C4" box="[853,973,1839,1865]" pageId="57" pageNumber="58" type="paratype">Paratypes</typeStatus>
</emphasis>
<specimenCount id="9D70C49AE025FFCCFC26D334FB98F8C7" box="[981,1061,1839,1866]" pageId="57" pageNumber="58" type="male">29 ♂♂</specimenCount>
, 
<specimenCount id="9D70C49AE025FFCCFBC1D334FB3FF8C7" box="[1074,1154,1839,1866]" pageId="57" pageNumber="58" type="female">13 ♀♀</specimenCount>
: same labels as 
<typeStatus id="54CDB1B1E025FFCCFAC9D334FA21F8C4" box="[1338,1436,1839,1865]" pageId="57" pageNumber="58" type="holotype">holotype</typeStatus>
but || yellow label hw «18» pr «.486,.488–.514,.517–.529,.534–.553 | coll. Dombrow» || red label pr « 
<typeStatus id="54CDB1B1E025FFCCFAD9D34FFA33F8E3" box="[1322,1422,1876,1902]" pageId="57" pageNumber="58" type="paratype">Paratype</typeStatus>
| 
<taxonomicName id="4C767490E025FFCCFF64D36CFEA8F81C" authorityName="Dombrow" authorityYear="2005" box="[151,277,1911,1937]" class="Insecta" family="Melolonthidae" genus="Beckhoplia" kingdom="Animalia" order="Coleoptera" pageId="57" pageNumber="58" phylum="Arthropoda" rank="genus">
<emphasis id="B902D301E025FFCCFF64D36CFEA8F81C" box="[151,277,1911,1937]" italics="true" pageId="57" pageNumber="58">Beckhoplia</emphasis>
</taxonomicName>
| 
<taxonomicName id="4C767490E025FFCCFEDBD36CFE37F81C" authority="Dombrow &amp; Colville, 2020" authorityName="Dombrow" authorityYear="2020" box="[296,394,1911,1937]" class="Insecta" family="Melolonthidae" genus="Beckhoplia" kingdom="Animalia" order="Coleoptera" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="suturalis" status="sp. nov.">
<emphasis id="B902D301E025FFCCFEDBD36CFE37F81C" box="[296,394,1911,1937]" italics="true" pageId="57" pageNumber="58">suturalis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2316E7AE025FFCCFE62D363FE7BF81F" box="[401,454,1912,1938]" pageId="57" pageNumber="58" rank="species">sp.n.</taxonomicNameLabel>
| det.: H. Dombrow 2019» (
<collectionCode id="ED6797D6E025FFCCFCF2D36CFCEAF81C" box="[769,855,1911,1937]" pageId="57" pageNumber="58">ESDW</collectionCode>
); 
<specimenCount id="9D70C49AE025FFCCFC9AD36CFC17F81F" box="[873,938,1911,1938]" pageId="57" pageNumber="58" type="male">2 ♂♂</specimenCount>
, 
<specimenCount id="9D70C49AE025FFCCFC45D36CFC4AF81F" box="[950,1015,1911,1938]" pageId="57" pageNumber="58" type="female">4 ♀♀</specimenCount>
: same labels as 
<typeStatus id="54CDB1B1E025FFCCFB59D36CFAB1F81C" box="[1194,1292,1911,1937]" pageId="57" pageNumber="58" type="holotype">holotype</typeStatus>
but || yellow label hw «18» pr «.515–.516,.530–.533 | coll. Dombrow» || yellow label pr «SAM–COL | A071220» || red label pr « 
<typeStatus id="54CDB1B1E025FFCCFF50D3DBFEBAF857" box="[163,263,1984,2010]" pageId="57" pageNumber="58" type="paratype">Paratype</typeStatus>
| 
<taxonomicName id="4C767490E025FFCCFEE8D3A4FE24F854" authorityName="Dombrow" authorityYear="2005" box="[283,409,1983,2009]" class="Insecta" family="Melolonthidae" genus="Beckhoplia" kingdom="Animalia" order="Coleoptera" pageId="57" pageNumber="58" phylum="Arthropoda" rank="genus">
<emphasis id="B902D301E025FFCCFEE8D3A4FE24F854" box="[283,409,1983,2009]" italics="true" pageId="57" pageNumber="58">Beckhoplia</emphasis>
</taxonomicName>
| 
<taxonomicName id="4C767490E025FFCCFE5FD3A4FDB3F854" authority="Dombrow &amp; Colville, 2020" authorityName="Dombrow" authorityYear="2020" box="[428,526,1983,2009]" class="Insecta" family="Melolonthidae" genus="Beckhoplia" kingdom="Animalia" order="Coleoptera" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="suturalis" status="sp. nov.">
<emphasis id="B902D301E025FFCCFE5FD3A4FDB3F854" box="[428,526,1983,2009]" italics="true" pageId="57" pageNumber="58">suturalis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2316E7AE025FFCCFDE6D3DBFDF7F857" box="[533,586,1984,2010]" pageId="57" pageNumber="58" rank="species">sp.n.</taxonomicNameLabel>
| det.: H. Dombrow 2019» (
<collectionCode id="ED6797D6E025FFCCFC75D3A4FC61F854" box="[902,988,1983,2009]" country="South Africa" httpUri="http://grbio.org/cool/94p5-pepz" name="Iziko Museums of Cape Town" pageId="57" pageNumber="58" type="Museum">SAMC</collectionCode>
).
</paragraph>
<paragraph id="8BC90F13E026FFCFFF64D48DFB04FF3C" blockId="58.[151,1209,150,177]" box="[151,1209,150,177]" pageId="58" pageNumber="59">
<emphasis id="B902D301E026FFCFFF64D48DFE1FFF3C" bold="true" box="[151,418,150,177]" pageId="58" pageNumber="59">
Condition of 
<typeStatus id="54CDB1B1E026FFCFFEC7D48CFE20FF3C" box="[308,413,151,177]" pageId="58" pageNumber="59" type="holotype">holotype</typeStatus>
.
</emphasis>
Excellent. Glued on a card. Dissected, parameres glued behind beetle.
</paragraph>
</subSubSection>
<subSubSection id="C36C5C98E026FFCFFF64D4C5FC28FE78" pageId="58" pageNumber="59" type="diagnosis">
<paragraph id="8BC90F13E026FFCFFF64D4C5FC28FE78" blockId="58.[151,1437,222,501]" pageId="58" pageNumber="59">
<emphasis id="B902D301E026FFCFFF64D4C5FE22FF75" bold="true" box="[151,415,222,248]" pageId="58" pageNumber="59">Differential diagnosis.</emphasis>
Differs from 
<taxonomicName id="4C767490E026FFCFFDC8D4C4FC8DFF74" authority=": Elytra" authorityName="Elytra" box="[571,816,223,249]" class="Insecta" family="Melolonthidae" genus="Beckhoplia" kingdom="Animalia" order="Coleoptera" pageId="58" pageNumber="59" phylum="Arthropoda" rank="species" species="nigrosetosa">
<emphasis id="B902D301E026FFCFFDC8D4C4FD60FF75" box="[571,733,223,248]" italics="true" pageId="58" pageNumber="59">B. nigrosetosa</emphasis>
: Elytra
</taxonomicName>
of 
<taxonomicName id="4C767490E026FFCFFCA4D4C4FC64FF74" authorityName="Dombrow" authorityYear="2020" box="[855,985,223,249]" class="Insecta" family="Melolonthidae" genus="Beckhoplia" kingdom="Animalia" order="Coleoptera" pageId="58" pageNumber="59" phylum="Arthropoda" rank="species" species="suturalis">
<emphasis id="B902D301E026FFCFFCA4D4C4FC64FF74" box="[855,985,223,249]" italics="true" pageId="58" pageNumber="59">B. suturalis</emphasis>
</taxonomicName>
widest in anterior third. Elytra of 
<taxonomicName id="4C767490E026FFCFFAAED4C4FEBEFE91" class="Insecta" family="Melolonthidae" genus="Beckhoplia" kingdom="Animalia" order="Coleoptera" pageId="58" pageNumber="59" phylum="Arthropoda" rank="species" species="nigrosetosa">
<emphasis id="B902D301E026FFCFFAAED4C4FEBEFE91" italics="true" pageId="58" pageNumber="59">B. nigrosetosa</emphasis>
</taxonomicName>
widest medially. Lateral margin of elytra of 
<taxonomicName id="4C767490E026FFCFFD07D518FCC8FE90" authorityName="Dombrow" authorityYear="2020" box="[756,885,259,285]" class="Insecta" family="Melolonthidae" genus="Beckhoplia" kingdom="Animalia" order="Coleoptera" pageId="58" pageNumber="59" phylum="Arthropoda" rank="species" species="suturalis">
<emphasis id="B902D301E026FFCFFD07D518FCC8FE90" box="[756,885,259,285]" italics="true" pageId="58" pageNumber="59">B. suturalis</emphasis>
</taxonomicName>
broadly blackened. Lateral margin of elytra of 
<taxonomicName id="4C767490E026FFCFFA77D518FEA4FECD" class="Insecta" family="Melolonthidae" genus="Beckhoplia" kingdom="Animalia" order="Coleoptera" pageId="58" pageNumber="59" phylum="Arthropoda" rank="species" species="nigrosetosa">
<emphasis id="B902D301E026FFCFFA77D518FEA4FECD" italics="true" pageId="58" pageNumber="59">B. nigrosetosa</emphasis>
</taxonomicName>
narrowly darkened. Interno-posterior margin of parameres of 
<taxonomicName id="4C767490E026FFCFFC3DD53CFBF2FECC" authorityName="Dombrow" authorityYear="2020" box="[974,1103,295,321]" class="Insecta" family="Melolonthidae" genus="Beckhoplia" kingdom="Animalia" order="Coleoptera" pageId="58" pageNumber="59" phylum="Arthropoda" rank="species" species="suturalis">
<emphasis id="B902D301E026FFCFFC3DD53CFBF2FECC" box="[974,1103,295,321]" italics="true" pageId="58" pageNumber="59">B. suturalis</emphasis>
</taxonomicName>
rounded in dorsal view (
<figureCitation id="134D1396E026FFCFFA97D53CFF08FEE8" captionStart="FIGURES 62–97" captionStartId="28.[151,264,1743,1767]" captionTargetBox="[161,1426,187,1710]" captionTargetId="figure-25@28.[151,1436,181,1719]" captionTargetPageId="28" captionText="FIGURES 62–97. Parameres and phallobase of Beckhoplia Dombrow, 2005 species, in dorsal view (left side), in lateral view (right side), ♂♂. 62–63, B. nigra Dombrow, new species; 64–65, B. pallidibrunnea Dombrow, new species; 66–67, B. fusca Dombrow, new species; 68–69, B. setosa Dombrow, new species; 70–71, B. castanea Dombrow, new species; 72–73, B. colvillei Dombrow, 2005; 74–75, B. pulchra Dombrow, new species; 76–77, B. nigrofasciata Dombrow, new species; 78–79, B. caliginosa Dombrow, new species; 80–81, B. pumila Dombrow, new species; 82–83, B. gifbergensis Dombrow, new species; 84–85, B. julianae Dombrow, 2005; 86–87, B. elkeae Dombrow, new species; 88–89, B. suturalis Dombrow, new species; 90–91, B. nigrosetosa Dombrow, new species; 92–93, B. occidentalis Dombrow, 2005; 94–95, B. bicolor Dombrow, new species; 96–97, B. dolichiocnemis Dombrow, new species; 98–99, Anisochelus inornatus Burmeister, 1844. Scale bar 1mm." figureDoi="http://doi.org/10.5281/zenodo.4401954" httpUri="https://zenodo.org/record/4401954/files/figure.png" pageId="58" pageNumber="59">Figs. 88</figureCitation>
, 
<figureCitation id="134D1396E026FFCFFF32D550FF53FEE8" box="[193,238,331,357]" captionStart="FIGURES 100–117" captionStartId="42.[151,264,1665,1689]" captionTargetBox="[162,1424,188,1625]" captionTargetId="figure-25@42.[151,1436,181,1641]" captionTargetPageId="42" captionText="FIGURES 100–117. Detailed drawings of the parameres for the key and the differential diagnosis of Beckhoplia Dombrow, 2005 species, in dorsal view, ♂♂. Arrows indicate characters of importance. 100, B. nigra Dombrow, new species; 101, B. pallidibrunnea Dombrow, new species; 102, B. fusca Dombrow, new species; 103, B. setosa Dombrow, new species; 104, B. castanea Dombrow, new species; 105, B. colvillei Dombrow, 2005; 106, B. pulchra Dombrow, new species; 107, B. nigrofasciata Dombrow, new species; 108, B. caliginosa Dombrow, new species; 109, B. pumila Dombrow, new species; 110, B. gifbergensis Dombrow, new species; 111, B. julianae Dombrow, 2005; 112, B. elkeae Dombrow, new species; 113, B. suturalis Dombrow, new species; 114, B. nigrosetosa Dombrow, new species; 115, B. occidentalis Dombrow, 2005; 116, B. bicolor Dombrow, new species; 117, B. dolichiocnemis Dombrow sp.n. Scale bar 1mm." figureDoi="http://doi.org/10.5281/zenodo.4401956" httpUri="https://zenodo.org/record/4401956/files/figure.png" pageId="58" pageNumber="59">113</figureCitation>
). Interno-posterior margin of parameres of 
<taxonomicName id="4C767490E026FFCFFD20D550FCC9FEE9" box="[723,884,331,356]" class="Insecta" family="Melolonthidae" genus="Beckhoplia" kingdom="Animalia" order="Coleoptera" pageId="58" pageNumber="59" phylum="Arthropoda" rank="species" species="nigrosetosa">
<emphasis id="B902D301E026FFCFFD20D550FCC9FEE9" box="[723,884,331,356]" italics="true" pageId="58" pageNumber="59">B. nigrosetosa</emphasis>
</taxonomicName>
parabola in dorsal view (
<figureCitation id="134D1396E026FFCFFB62D550FB52FEE8" box="[1169,1263,331,357]" captionStart="FIGURES 62–97" captionStartId="28.[151,264,1743,1767]" captionTargetBox="[161,1426,187,1710]" captionTargetId="figure-25@28.[151,1436,181,1719]" captionTargetPageId="28" captionText="FIGURES 62–97. Parameres and phallobase of Beckhoplia Dombrow, 2005 species, in dorsal view (left side), in lateral view (right side), ♂♂. 62–63, B. nigra Dombrow, new species; 64–65, B. pallidibrunnea Dombrow, new species; 66–67, B. fusca Dombrow, new species; 68–69, B. setosa Dombrow, new species; 70–71, B. castanea Dombrow, new species; 72–73, B. colvillei Dombrow, 2005; 74–75, B. pulchra Dombrow, new species; 76–77, B. nigrofasciata Dombrow, new species; 78–79, B. caliginosa Dombrow, new species; 80–81, B. pumila Dombrow, new species; 82–83, B. gifbergensis Dombrow, new species; 84–85, B. julianae Dombrow, 2005; 86–87, B. elkeae Dombrow, new species; 88–89, B. suturalis Dombrow, new species; 90–91, B. nigrosetosa Dombrow, new species; 92–93, B. occidentalis Dombrow, 2005; 94–95, B. bicolor Dombrow, new species; 96–97, B. dolichiocnemis Dombrow, new species; 98–99, Anisochelus inornatus Burmeister, 1844. Scale bar 1mm." figureDoi="http://doi.org/10.5281/zenodo.4401954" httpUri="https://zenodo.org/record/4401954/files/figure.png" pageId="58" pageNumber="59">Figs. 90</figureCitation>
, 
<figureCitation id="134D1396E026FFCFFB09D550FA9AFEE8" box="[1274,1319,331,357]" captionStart="FIGURES 100–117" captionStartId="42.[151,264,1665,1689]" captionTargetBox="[162,1424,188,1625]" captionTargetId="figure-25@42.[151,1436,181,1641]" captionTargetPageId="42" captionText="FIGURES 100–117. Detailed drawings of the parameres for the key and the differential diagnosis of Beckhoplia Dombrow, 2005 species, in dorsal view, ♂♂. Arrows indicate characters of importance. 100, B. nigra Dombrow, new species; 101, B. pallidibrunnea Dombrow, new species; 102, B. fusca Dombrow, new species; 103, B. setosa Dombrow, new species; 104, B. castanea Dombrow, new species; 105, B. colvillei Dombrow, 2005; 106, B. pulchra Dombrow, new species; 107, B. nigrofasciata Dombrow, new species; 108, B. caliginosa Dombrow, new species; 109, B. pumila Dombrow, new species; 110, B. gifbergensis Dombrow, new species; 111, B. julianae Dombrow, 2005; 112, B. elkeae Dombrow, new species; 113, B. suturalis Dombrow, new species; 114, B. nigrosetosa Dombrow, new species; 115, B. occidentalis Dombrow, 2005; 116, B. bicolor Dombrow, new species; 117, B. dolichiocnemis Dombrow sp.n. Scale bar 1mm." figureDoi="http://doi.org/10.5281/zenodo.4401956" httpUri="https://zenodo.org/record/4401956/files/figure.png" pageId="58" pageNumber="59">114</figureCitation>
). Externoposterior margin of parameres of 
<taxonomicName id="4C767490E026FFCFFDE4D574FD27FE04" authorityName="Dombrow" authorityYear="2020" box="[535,666,367,393]" class="Insecta" family="Melolonthidae" genus="Beckhoplia" kingdom="Animalia" order="Coleoptera" pageId="58" pageNumber="59" phylum="Arthropoda" rank="species" species="suturalis">
<emphasis id="B902D301E026FFCFFDE4D574FD27FE04" box="[535,666,367,393]" italics="true" pageId="58" pageNumber="59">B. suturalis</emphasis>
</taxonomicName>
convex in dorsal view (
<figureCitation id="134D1396E026FFCFFC43D574FBB2FE04" box="[944,1039,367,393]" captionStart="FIGURES 62–97" captionStartId="28.[151,264,1743,1767]" captionTargetBox="[161,1426,187,1710]" captionTargetId="figure-25@28.[151,1436,181,1719]" captionTargetPageId="28" captionText="FIGURES 62–97. Parameres and phallobase of Beckhoplia Dombrow, 2005 species, in dorsal view (left side), in lateral view (right side), ♂♂. 62–63, B. nigra Dombrow, new species; 64–65, B. pallidibrunnea Dombrow, new species; 66–67, B. fusca Dombrow, new species; 68–69, B. setosa Dombrow, new species; 70–71, B. castanea Dombrow, new species; 72–73, B. colvillei Dombrow, 2005; 74–75, B. pulchra Dombrow, new species; 76–77, B. nigrofasciata Dombrow, new species; 78–79, B. caliginosa Dombrow, new species; 80–81, B. pumila Dombrow, new species; 82–83, B. gifbergensis Dombrow, new species; 84–85, B. julianae Dombrow, 2005; 86–87, B. elkeae Dombrow, new species; 88–89, B. suturalis Dombrow, new species; 90–91, B. nigrosetosa Dombrow, new species; 92–93, B. occidentalis Dombrow, 2005; 94–95, B. bicolor Dombrow, new species; 96–97, B. dolichiocnemis Dombrow, new species; 98–99, Anisochelus inornatus Burmeister, 1844. Scale bar 1mm." figureDoi="http://doi.org/10.5281/zenodo.4401954" httpUri="https://zenodo.org/record/4401954/files/figure.png" pageId="58" pageNumber="59">Figs. 88</figureCitation>
, 
<figureCitation id="134D1396E026FFCFFBEED574FBF7FE04" box="[1053,1098,367,393]" captionStart="FIGURES 100–117" captionStartId="42.[151,264,1665,1689]" captionTargetBox="[162,1424,188,1625]" captionTargetId="figure-25@42.[151,1436,181,1641]" captionTargetPageId="42" captionText="FIGURES 100–117. Detailed drawings of the parameres for the key and the differential diagnosis of Beckhoplia Dombrow, 2005 species, in dorsal view, ♂♂. Arrows indicate characters of importance. 100, B. nigra Dombrow, new species; 101, B. pallidibrunnea Dombrow, new species; 102, B. fusca Dombrow, new species; 103, B. setosa Dombrow, new species; 104, B. castanea Dombrow, new species; 105, B. colvillei Dombrow, 2005; 106, B. pulchra Dombrow, new species; 107, B. nigrofasciata Dombrow, new species; 108, B. caliginosa Dombrow, new species; 109, B. pumila Dombrow, new species; 110, B. gifbergensis Dombrow, new species; 111, B. julianae Dombrow, 2005; 112, B. elkeae Dombrow, new species; 113, B. suturalis Dombrow, new species; 114, B. nigrosetosa Dombrow, new species; 115, B. occidentalis Dombrow, 2005; 116, B. bicolor Dombrow, new species; 117, B. dolichiocnemis Dombrow sp.n. Scale bar 1mm." figureDoi="http://doi.org/10.5281/zenodo.4401956" httpUri="https://zenodo.org/record/4401956/files/figure.png" pageId="58" pageNumber="59">113</figureCitation>
). Externo-posterior margin of parameres of 
<taxonomicName id="4C767490E026FFCFFEC3D588FE6CFE21" box="[304,465,403,428]" class="Insecta" family="Melolonthidae" genus="Beckhoplia" kingdom="Animalia" order="Coleoptera" pageId="58" pageNumber="59" phylum="Arthropoda" rank="species" species="nigrosetosa">
<emphasis id="B902D301E026FFCFFEC3D588FE6CFE21" box="[304,465,403,428]" italics="true" pageId="58" pageNumber="59">B. nigrosetosa</emphasis>
</taxonomicName>
linear with a convex median extension in dorsal view (
<figureCitation id="134D1396E026FFCFFBCCD588FB20FE20" box="[1087,1181,403,429]" captionStart="FIGURES 62–97" captionStartId="28.[151,264,1743,1767]" captionTargetBox="[161,1426,187,1710]" captionTargetId="figure-25@28.[151,1436,181,1719]" captionTargetPageId="28" captionText="FIGURES 62–97. Parameres and phallobase of Beckhoplia Dombrow, 2005 species, in dorsal view (left side), in lateral view (right side), ♂♂. 62–63, B. nigra Dombrow, new species; 64–65, B. pallidibrunnea Dombrow, new species; 66–67, B. fusca Dombrow, new species; 68–69, B. setosa Dombrow, new species; 70–71, B. castanea Dombrow, new species; 72–73, B. colvillei Dombrow, 2005; 74–75, B. pulchra Dombrow, new species; 76–77, B. nigrofasciata Dombrow, new species; 78–79, B. caliginosa Dombrow, new species; 80–81, B. pumila Dombrow, new species; 82–83, B. gifbergensis Dombrow, new species; 84–85, B. julianae Dombrow, 2005; 86–87, B. elkeae Dombrow, new species; 88–89, B. suturalis Dombrow, new species; 90–91, B. nigrosetosa Dombrow, new species; 92–93, B. occidentalis Dombrow, 2005; 94–95, B. bicolor Dombrow, new species; 96–97, B. dolichiocnemis Dombrow, new species; 98–99, Anisochelus inornatus Burmeister, 1844. Scale bar 1mm." figureDoi="http://doi.org/10.5281/zenodo.4401954" httpUri="https://zenodo.org/record/4401954/files/figure.png" pageId="58" pageNumber="59">Figs. 90</figureCitation>
, 
<figureCitation id="134D1396E026FFCFFB5BD588FB68FE20" box="[1192,1237,403,429]" captionStart="FIGURES 100–117" captionStartId="42.[151,264,1665,1689]" captionTargetBox="[162,1424,188,1625]" captionTargetId="figure-25@42.[151,1436,181,1641]" captionTargetPageId="42" captionText="FIGURES 100–117. Detailed drawings of the parameres for the key and the differential diagnosis of Beckhoplia Dombrow, 2005 species, in dorsal view, ♂♂. Arrows indicate characters of importance. 100, B. nigra Dombrow, new species; 101, B. pallidibrunnea Dombrow, new species; 102, B. fusca Dombrow, new species; 103, B. setosa Dombrow, new species; 104, B. castanea Dombrow, new species; 105, B. colvillei Dombrow, 2005; 106, B. pulchra Dombrow, new species; 107, B. nigrofasciata Dombrow, new species; 108, B. caliginosa Dombrow, new species; 109, B. pumila Dombrow, new species; 110, B. gifbergensis Dombrow, new species; 111, B. julianae Dombrow, 2005; 112, B. elkeae Dombrow, new species; 113, B. suturalis Dombrow, new species; 114, B. nigrosetosa Dombrow, new species; 115, B. occidentalis Dombrow, 2005; 116, B. bicolor Dombrow, new species; 117, B. dolichiocnemis Dombrow sp.n. Scale bar 1mm." figureDoi="http://doi.org/10.5281/zenodo.4401956" httpUri="https://zenodo.org/record/4401956/files/figure.png" pageId="58" pageNumber="59">114</figureCitation>
). Anterior margin of phallobase of 
<taxonomicName id="4C767490E026FFCFFEA7D5ACFE6BFE5C" authorityName="Dombrow" authorityYear="2020" box="[340,470,439,465]" class="Insecta" family="Melolonthidae" genus="Beckhoplia" kingdom="Animalia" order="Coleoptera" pageId="58" pageNumber="59" phylum="Arthropoda" rank="species" species="suturalis">
<emphasis id="B902D301E026FFCFFEA7D5ACFE6BFE5C" box="[340,470,439,465]" italics="true" pageId="58" pageNumber="59">B. suturalis</emphasis>
</taxonomicName>
curly bracket-shaped in dorsal view (
<figureCitation id="134D1396E026FFCFFC73D5ACFC63FE5C" box="[896,990,439,465]" captionStart="FIGURES 62–97" captionStartId="28.[151,264,1743,1767]" captionTargetBox="[161,1426,187,1710]" captionTargetId="figure-25@28.[151,1436,181,1719]" captionTargetPageId="28" captionText="FIGURES 62–97. Parameres and phallobase of Beckhoplia Dombrow, 2005 species, in dorsal view (left side), in lateral view (right side), ♂♂. 62–63, B. nigra Dombrow, new species; 64–65, B. pallidibrunnea Dombrow, new species; 66–67, B. fusca Dombrow, new species; 68–69, B. setosa Dombrow, new species; 70–71, B. castanea Dombrow, new species; 72–73, B. colvillei Dombrow, 2005; 74–75, B. pulchra Dombrow, new species; 76–77, B. nigrofasciata Dombrow, new species; 78–79, B. caliginosa Dombrow, new species; 80–81, B. pumila Dombrow, new species; 82–83, B. gifbergensis Dombrow, new species; 84–85, B. julianae Dombrow, 2005; 86–87, B. elkeae Dombrow, new species; 88–89, B. suturalis Dombrow, new species; 90–91, B. nigrosetosa Dombrow, new species; 92–93, B. occidentalis Dombrow, 2005; 94–95, B. bicolor Dombrow, new species; 96–97, B. dolichiocnemis Dombrow, new species; 98–99, Anisochelus inornatus Burmeister, 1844. Scale bar 1mm." figureDoi="http://doi.org/10.5281/zenodo.4401954" httpUri="https://zenodo.org/record/4401954/files/figure.png" pageId="58" pageNumber="59">Figs. 88</figureCitation>
, 
<figureCitation id="134D1396E026FFCFFC19D5ACFBAAFE5C" box="[1002,1047,439,465]" captionStart="FIGURES 100–117" captionStartId="42.[151,264,1665,1689]" captionTargetBox="[162,1424,188,1625]" captionTargetId="figure-25@42.[151,1436,181,1641]" captionTargetPageId="42" captionText="FIGURES 100–117. Detailed drawings of the parameres for the key and the differential diagnosis of Beckhoplia Dombrow, 2005 species, in dorsal view, ♂♂. Arrows indicate characters of importance. 100, B. nigra Dombrow, new species; 101, B. pallidibrunnea Dombrow, new species; 102, B. fusca Dombrow, new species; 103, B. setosa Dombrow, new species; 104, B. castanea Dombrow, new species; 105, B. colvillei Dombrow, 2005; 106, B. pulchra Dombrow, new species; 107, B. nigrofasciata Dombrow, new species; 108, B. caliginosa Dombrow, new species; 109, B. pumila Dombrow, new species; 110, B. gifbergensis Dombrow, new species; 111, B. julianae Dombrow, 2005; 112, B. elkeae Dombrow, new species; 113, B. suturalis Dombrow, new species; 114, B. nigrosetosa Dombrow, new species; 115, B. occidentalis Dombrow, 2005; 116, B. bicolor Dombrow, new species; 117, B. dolichiocnemis Dombrow sp.n. Scale bar 1mm." figureDoi="http://doi.org/10.5281/zenodo.4401956" httpUri="https://zenodo.org/record/4401956/files/figure.png" pageId="58" pageNumber="59">113</figureCitation>
). Anterior margin of phallobase of 
<taxonomicName id="4C767490E026FFCFFF64D5C0FE85FE79" box="[151,312,475,500]" class="Insecta" family="Melolonthidae" genus="Beckhoplia" kingdom="Animalia" order="Coleoptera" pageId="58" pageNumber="59" phylum="Arthropoda" rank="species" species="nigrosetosa">
<emphasis id="B902D301E026FFCFFF64D5C0FE85FE79" box="[151,312,475,500]" italics="true" pageId="58" pageNumber="59">B. nigrosetosa</emphasis>
</taxonomicName>
broad parabola-shaped in dorsal view (
<figureCitation id="134D1396E026FFCFFD01D5C0FCEDFE78" box="[754,848,475,501]" captionStart="FIGURES 62–97" captionStartId="28.[151,264,1743,1767]" captionTargetBox="[161,1426,187,1710]" captionTargetId="figure-25@28.[151,1436,181,1719]" captionTargetPageId="28" captionText="FIGURES 62–97. Parameres and phallobase of Beckhoplia Dombrow, 2005 species, in dorsal view (left side), in lateral view (right side), ♂♂. 62–63, B. nigra Dombrow, new species; 64–65, B. pallidibrunnea Dombrow, new species; 66–67, B. fusca Dombrow, new species; 68–69, B. setosa Dombrow, new species; 70–71, B. castanea Dombrow, new species; 72–73, B. colvillei Dombrow, 2005; 74–75, B. pulchra Dombrow, new species; 76–77, B. nigrofasciata Dombrow, new species; 78–79, B. caliginosa Dombrow, new species; 80–81, B. pumila Dombrow, new species; 82–83, B. gifbergensis Dombrow, new species; 84–85, B. julianae Dombrow, 2005; 86–87, B. elkeae Dombrow, new species; 88–89, B. suturalis Dombrow, new species; 90–91, B. nigrosetosa Dombrow, new species; 92–93, B. occidentalis Dombrow, 2005; 94–95, B. bicolor Dombrow, new species; 96–97, B. dolichiocnemis Dombrow, new species; 98–99, Anisochelus inornatus Burmeister, 1844. Scale bar 1mm." figureDoi="http://doi.org/10.5281/zenodo.4401954" httpUri="https://zenodo.org/record/4401954/files/figure.png" pageId="58" pageNumber="59">Figs. 90</figureCitation>
, 
<figureCitation id="134D1396E026FFCFFCA8D5C0FC35FE78" box="[859,904,475,501]" captionStart="FIGURES 100–117" captionStartId="42.[151,264,1665,1689]" captionTargetBox="[162,1424,188,1625]" captionTargetId="figure-25@42.[151,1436,181,1641]" captionTargetPageId="42" captionText="FIGURES 100–117. Detailed drawings of the parameres for the key and the differential diagnosis of Beckhoplia Dombrow, 2005 species, in dorsal view, ♂♂. Arrows indicate characters of importance. 100, B. nigra Dombrow, new species; 101, B. pallidibrunnea Dombrow, new species; 102, B. fusca Dombrow, new species; 103, B. setosa Dombrow, new species; 104, B. castanea Dombrow, new species; 105, B. colvillei Dombrow, 2005; 106, B. pulchra Dombrow, new species; 107, B. nigrofasciata Dombrow, new species; 108, B. caliginosa Dombrow, new species; 109, B. pumila Dombrow, new species; 110, B. gifbergensis Dombrow, new species; 111, B. julianae Dombrow, 2005; 112, B. elkeae Dombrow, new species; 113, B. suturalis Dombrow, new species; 114, B. nigrosetosa Dombrow, new species; 115, B. occidentalis Dombrow, 2005; 116, B. bicolor Dombrow, new species; 117, B. dolichiocnemis Dombrow sp.n. Scale bar 1mm." figureDoi="http://doi.org/10.5281/zenodo.4401956" httpUri="https://zenodo.org/record/4401956/files/figure.png" pageId="58" pageNumber="59">114</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C36C5C98E026FFC9FF64D639FB42FC28" lastPageId="60" lastPageNumber="61" pageId="58" pageNumber="59" type="description">
<paragraph id="8BC90F13E026FFCFFF64D639FE9CFDB1" blockId="58.[151,1437,546,2013]" box="[151,289,546,572]" pageId="58" pageNumber="59">
<emphasis id="B902D301E026FFCFFF64D639FE9CFDB1" bold="true" box="[151,289,546,572]" pageId="58" pageNumber="59">Description</emphasis>
</paragraph>
<paragraph id="8BC90F13E026FFCFFF34D65CFC83FDEC" blockId="58.[151,1437,546,2013]" box="[199,830,583,609]" pageId="58" pageNumber="59">
<emphasis id="B902D301E026FFCFFF34D65CFE3DFDEC" bold="true" box="[199,384,583,609]" pageId="58" pageNumber="59">Measurements:</emphasis>
length 
<quantity id="4C8EA2F6E026FFCFFE27D65CFDE3FDEC" box="[468,606,583,609]" metricMagnitude="-3" metricUnit="m" metricValue="8.7" metricValueMax="9.6" metricValueMin="7.8" pageId="58" pageNumber="59" unit="mm" value="8.7" valueMax="9.6" valueMin="7.8">7.8–9.6 mm</quantity>
; width 4.0– 
<quantity id="4C8EA2F6E026FFCFFD12D65CFC87FDEC" box="[737,826,583,609]" metricMagnitude="-3" metricUnit="m" metricValue="4.7" pageId="58" pageNumber="59" unit="mm" value="4.7">4.7 mm</quantity>
.
</paragraph>
<paragraph id="8BC90F13E026FFCFFF34D670FEADFD24" blockId="58.[151,1437,546,2013]" pageId="58" pageNumber="59">
<emphasis id="B902D301E026FFCFFF34D670FEE6FD08" bold="true" box="[199,347,619,645]" pageId="58" pageNumber="59">Integument:</emphasis>
Black; funiculum brown annulated; elytra transparently brown, lateral and posterior margin blackened.
</paragraph>
<paragraph id="8BC90F13E026FFCFFF34D6A8FDBDF96C" blockId="58.[151,1437,546,2013]" pageId="58" pageNumber="59">
<emphasis id="B902D301E026FFCFFF34D6A8FEFAFD40" bold="true" box="[199,327,691,717]" pageId="58" pageNumber="59">Tegument:</emphasis>
<emphasis id="B902D301E026FFCFFEA3D6A8FE17FD40" box="[336,426,691,717]" italics="true" pageId="58" pageNumber="59">Clypeus</emphasis>
bare; loose setose with short, erect, black setae on the inner side of the reflexed anterior and lateral margins. 
<emphasis id="B902D301E026FFCFFE89D6CCFE0AFD7C" box="[378,439,727,753]" italics="true" pageId="58" pageNumber="59">Head</emphasis>
loose setose with long, erect, black setae. 
<emphasis id="B902D301E026FFCFFC7ED6CCFBA8FD7C" box="[909,1045,727,753]" italics="true" pageId="58" pageNumber="59">Eye canthus</emphasis>
with a dense setose ciliary of long, black setae. 
<emphasis id="B902D301E026FFCFFED6D6E0FEC9FC98" box="[293,372,763,789]" italics="true" pageId="58" pageNumber="59">Scapus</emphasis>
loose setose with long, black setae, whitish ventrally, sparse and short on 
<emphasis id="B902D301E026FFCFFB3FD6E0FF1DFCB5" italics="true" pageId="58" pageNumber="59">pedicellus; funicle I</emphasis>
bare; 
<emphasis id="B902D301E026FFCFFF19D704FEC7FCB5" box="[234,378,799,825]" italics="true" pageId="58" pageNumber="59">funicle II–IV</emphasis>
with few extremely short, semi-erect, pale-brown setae; 
<emphasis id="B902D301E026FFCFFBF5D704FBF3FCB4" box="[1030,1102,799,825]" italics="true" pageId="58" pageNumber="59">clavus</emphasis>
sparse setose with extremely short, reclinate pale-brown setae, very short, semi-erect, black setae interspersed. 
<emphasis id="B902D301E026FFCFFBDBD758FB2AFCD1" box="[1064,1175,835,860]" italics="true" pageId="58" pageNumber="59">Pronotum</emphasis>
loose setose with long, erect, black setae, shorter towards posterior margin; with a loose setose ciliary of long, black setae. 
<emphasis id="B902D301E026FFCFFB1FD77CFAE7FC0C" box="[1260,1370,871,897]" italics="true" pageId="58" pageNumber="59">Scutellum</emphasis>
loose setose with short and medium-long, semi-erect, brown setae. 
<emphasis id="B902D301E026FFCFFCB0D790FC34FC28" box="[835,905,907,933]" italics="true" pageId="58" pageNumber="59">Elytra</emphasis>
without pattern. Sparse setose with short, reclinate and semi-erect, black setae; few short, reclinate, white setae interspersed below apical torus; with a loose juxtasutural row of long, erect, black setae from anterior margin crossing scutellar torus towards apical torus. Suture with a loose row of medium-long and long, robust, erect, black setae. 
<emphasis id="B902D301E026FFCFFC98D7ECFC01FB9C" box="[875,956,1015,1041]" italics="true" pageId="58" pageNumber="59">Thorax</emphasis>
and 
<emphasis id="B902D301E026FFCFFC00D7ECFBE8FB9C" box="[1011,1109,1015,1041]" italics="true" pageId="58" pageNumber="59">pleurites</emphasis>
loose pilose with long, black hairs. 
<emphasis id="B902D301E026FFCFFF28D000FEDDFBB9" box="[219,352,1051,1076]" italics="true" pageId="58" pageNumber="59">Ventrite I–V</emphasis>
loose setose with short, reclinate, black setae; medium-long and long, semi-erect, black setae and short reclinate white setae interspersed. 
<emphasis id="B902D301E026FFCFFDACD024FD4DFBD4" box="[607,752,1087,1113]" italics="true" pageId="58" pageNumber="59">Propygidium</emphasis>
and 
<emphasis id="B902D301E026FFCFFCD9D024FC2FFBD4" box="[810,914,1087,1113]" italics="true" pageId="58" pageNumber="59">pygidium</emphasis>
loose setose with long, reclinate, black setae; short reclinate white setae interspersed on 
<emphasis id="B902D301E026FFCFFD72D078FCB2FBF0" box="[641,783,1123,1149]" italics="true" pageId="58" pageNumber="59">propygidium</emphasis>
; 
<emphasis id="B902D301E026FFCFFCD3D078FC73FBF0" box="[800,974,1123,1149]" italics="true" pageId="58" pageNumber="59">tip of pygidium</emphasis>
with a sparse setose ciliary of mediumlong and long, black setae. 
<emphasis id="B902D301E026FFCFFE38D09CFD27FB2C" box="[459,666,1159,1185]" italics="true" pageId="58" pageNumber="59">Prothoracic femur</emphasis>
loose pilose with long black hairs; bare internally. 
<emphasis id="B902D301E026FFCFFB28D09CFA21FB2C" box="[1243,1436,1159,1185]" italics="true" pageId="58" pageNumber="59">Prothoracic tibia</emphasis>
loose setose with short, reclinate, greyish setae; medium-long, semi-erect, black setae interspersed; erect and long ventrally; with a sparse setose longitudinal row of medium-long, erect, black setae dorsally; a loose setose ciliary of short, black setae on internal ridge; a setaceous ciliary of short, brown setae on anterior margin, more robust towards internal ridge; a medium-long downwards pointing setose ciliary of pale-brown setae along anterior margin of apical tooth. 
<emphasis id="B902D301E026FFCFFEFAD120FE50FAD8" box="[265,493,1339,1365]" italics="true" pageId="58" pageNumber="59">Mesothoracic femur</emphasis>
loose pilose with long black hairs; bare internally. 
<emphasis id="B902D301E026FFCFFBC5D120FAB6FAD8" box="[1078,1291,1339,1365]" italics="true" pageId="58" pageNumber="59">Mesothoracic tibia</emphasis>
loose setose with short, reclinate, whitish setae, sparser internally; medium-long, semi-erect, robust, black setae interspersed, long and finer internally. Posterior margin with a bristled ciliary of short, black bristles; missing internally. 
<emphasis id="B902D301E026FFCFFAAAD198FE81FA4C" italics="true" pageId="58" pageNumber="59">Metathoracic femur</emphasis>
loose setose with long, semi-erect, black setae; distinctly long and whitish ventrally; bare internally. 
<emphasis id="B902D301E026FFCFFF64D1D0FED7FA68" box="[151,362,1483,1509]" italics="true" pageId="58" pageNumber="59">Metathoracic tibia</emphasis>
sparse setose with short, reclinate, brown setae; medium-long, semi-erect, robust, black setae interspersed, long, denser and finer internally. Posterior margin with a loose bristled outer marginal ciliary of short, brown bristles, lengthening dorsally and ventrally, more robust ventrally. 
<emphasis id="B902D301E026FFCFFC25D208FAB2F9A1" box="[982,1295,1555,1581]" italics="true" pageId="58" pageNumber="59">Prothoracic tarsomere I–IV</emphasis>
bare. 
<emphasis id="B902D301E026FFCFFAA5D208FF49F9DC" italics="true" pageId="58" pageNumber="59">Mesothoracic</emphasis>
and 
<emphasis id="B902D301E026FFCFFEDCD22CFDCAF9DD" box="[303,631,1591,1617]" italics="true" pageId="58" pageNumber="59">metathoracic tarsomere I–IV</emphasis>
sparse setose with short, reclinate, brown setae. 
<emphasis id="B902D301E026FFCFFB54D22CFAEAF9DD" box="[1191,1367,1591,1616]" italics="true" pageId="58" pageNumber="59">Tarsomere I–IV</emphasis>
of all thoracic legs with a setaceous, apical, marginal ciliary of short, black setae lengthening towards 
<emphasis id="B902D301E026FFCFFB33D240FF64F915" italics="true" pageId="58" pageNumber="59">metathoracic tarsomeres</emphasis>
, fine dorsally and more robust ventrally. 
<emphasis id="B902D301E026FFCFFD6CD264FC8AF915" box="[671,823,1663,1688]" italics="true" pageId="58" pageNumber="59">Pretarsomere</emphasis>
of all thoracic legs sparse setose with short, semi-erect, brown setae, lengthening towards 
<emphasis id="B902D301E026FFCFFDEFD2B8FCF0F931" box="[540,845,1699,1725]" italics="true" pageId="58" pageNumber="59">metathoracic pretarsomere</emphasis>
; few, long, semi-erect, black setae at apical margin, shorter and red-brown ventrally.
</paragraph>
<paragraph id="8BC90F13E026FFCFFF34D2F0FC06F8C0" blockId="58.[151,1437,546,2013]" pageId="58" pageNumber="59">
<emphasis id="B902D301E026FFCFFF34D2F0FEADF888" bold="true" box="[199,272,1771,1797]" pageId="58" pageNumber="59">Head:</emphasis>
Suture between head and clypeus linear, raised. Head dense punctate with small, shallow, roundish, not merging punctures. Funicle I–II cylindrical, slightly thickened medially; funicle III asymmetric frustum-shaped; clavus elliptic, slightly longer than pedicellus and funiculum combined.
</paragraph>
<paragraph id="8BC90F13E026FFCFFF34D34CFEF4F850" blockId="58.[151,1437,546,2013]" pageId="58" pageNumber="59">
<emphasis id="B902D301E026FFCFFF34D34CFEF4F8FC" bold="true" box="[199,329,1879,1905]" pageId="58" pageNumber="59">Pronotum:</emphasis>
Widest medially; loose punctate with small, shallow, roundish punctures, dense posteriorly and within medial furrow. Anterior corner orthogonal. Lateral margin linearly attenuate anteriorly and subparallel posteriorly. With an indistinct, shortened, shallow medial furrow from posterior margin terminating on disk. Posterior corner rounded.
</paragraph>
<paragraph id="8BC90F13E027FFCEFF34D48DFCE7FF75" blockId="59.[151,1437,150,2013]" pageId="59" pageNumber="60">
<emphasis id="B902D301E027FFCEFF34D48DFEF5FF3D" bold="true" box="[199,328,150,176]" pageId="59" pageNumber="60">Scutellum:</emphasis>
Parabola-shaped; as broad as long; dense punctate with small to large, shallow, roundish punctures, micropunctate within; with a shortened, median, longitudinal, narrow, smooth stripe from anterior margin terminating in apical third. Lateral margin slightly convex. Tip pointed.
</paragraph>
<paragraph id="8BC90F13E027FFCEFF34D519FF6AFE5C" blockId="59.[151,1437,150,2013]" pageId="59" pageNumber="60">
<emphasis id="B902D301E027FFCEFF34D519FEA0FE91" bold="true" box="[199,285,258,284]" pageId="59" pageNumber="60">Elytra:</emphasis>
Ovate; longer than broad; widest in anterior third; protruding propygidium. Lateral margin slightly convex attenuate apically. Sculpture indistinct; almost complanate laterally; humerus indistinct; with a broad roundish torus beside scutellum, expanding towards disk; lateral trench indistinct; complanate along scutellum and suture. Loose punctate with extremely small to very small, shallow, roundish punctures, micropunctures in-between; a loose juxta-sutural row of small, deep, roundish punctures from anterior margin, crossing scutellar torus to apical torus.
</paragraph>
<paragraph id="8BC90F13E027FFCEFF34D5C1FC53FDB1" blockId="59.[151,1437,150,2013]" pageId="59" pageNumber="60">
<emphasis id="B902D301E027FFCEFF34D5C1FEF8FE79" bold="true" box="[199,325,474,500]" pageId="59" pageNumber="60">Abdomen:</emphasis>
Ventrite I distinctly shortened medially; ventrite V not constricted ventrally. Propygidium with a slightly projected stoma at posterior corner. Pygidium distinctly pulvinate longitudinally and laterally; upper margin slightly convex, linear medially; lateral margin slightly convex; tip rounded.
</paragraph>
<paragraph id="8BC90F13E027FFCEFF34D65DFD59FD7D" blockId="59.[151,1437,150,2013]" pageId="59" pageNumber="60">
<emphasis id="B902D301E027FFCEFF34D65DFE37FDED" bold="true" box="[199,394,582,608]" pageId="59" pageNumber="60">Prothoracic leg:</emphasis>
Femur and tibia slender. Tibia smooth ventrally. Median tooth elongate. Basal tooth very small. Interspace between teeth almost equal. Shape of interspace between teeth concave. Tarsomere I–IV combined extremely short; equal in length; slightly less than 
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length of tibia. Pretarsomere distinctly thickened; smooth ventrally; slightly less than 
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length of tarsomere I–IV combined. Internal claw as long as pretarsomere. External claw slightly more than 
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length of internal claw.
</paragraph>
<paragraph id="8BC90F13E027FFCEFF34D6E1FCF2FC0D" blockId="59.[151,1437,150,2013]" pageId="59" pageNumber="60">
<emphasis id="B902D301E027FFCEFF34D6E1FE21FC99" bold="true" box="[199,412,762,788]" pageId="59" pageNumber="60">Mesothoracic leg:</emphasis>
Tibia slightly shorter than femur. Tarsomere I–IV combined extremely short; slightly less than 
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length of tibia. Pretarsomere with a shortened smooth ridge ventrally followed by a roundish cutting apically; slightly shorter than tarsomere I–IV combined. External claw slightly more than 
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length of pretarsomere. Internal claw slightly more than 
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/ 
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in
</quantity>
length of external claw.
</paragraph>
<paragraph id="8BC90F13E027FFCEFF34D791FC5AFB9D" blockId="59.[151,1437,150,2013]" pageId="59" pageNumber="60">
<emphasis id="B902D301E027FFCEFF34D791FE21FC29" bold="true" box="[199,412,906,932]" pageId="59" pageNumber="60">Metathoracic leg:</emphasis>
Tibia slightly less than 
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/ 
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more in length of femur. Tarsomere I–IV combined distinctly short; slightly less than 
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/ 
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in
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length of tibia. Pretarsomere slightly thickened; with an undulate ridge ventrally; slightly less than 
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/ 
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in
</quantity>
length of tarsomere I–IV combined. External claw slightly less than 
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more in length of pretarsomere. Internal claw slightly more than 
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/ 
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<subScript id="17F20D56E027FFCEFD47D013FD01FB9B" attach="left" box="[692,700,1032,1046]" fontSize="6" pageId="59" pageNumber="60">5</subScript>
in
</quantity>
length of external claw.
</paragraph>
<paragraph id="8BC90F13E027FFCEFF34D001FD90FBB9" blockId="59.[151,1437,150,2013]" box="[199,557,1050,1076]" pageId="59" pageNumber="60">
<emphasis id="B902D301E027FFCEFF34D001FEECFBB9" bold="true" box="[199,337,1050,1076]" pageId="59" pageNumber="60">Parameres:</emphasis>
(
<figureCitation id="134D1396E027FFCEFE93D001FE54FBB9" box="[352,489,1050,1076]" captionStart="FIGURES 62–97" captionStartId="28.[151,264,1743,1767]" captionTargetBox="[161,1426,187,1710]" captionTargetId="figure-25@28.[151,1436,181,1719]" captionTargetPageId="28" captionText="FIGURES 62–97. Parameres and phallobase of Beckhoplia Dombrow, 2005 species, in dorsal view (left side), in lateral view (right side), ♂♂. 62–63, B. nigra Dombrow, new species; 64–65, B. pallidibrunnea Dombrow, new species; 66–67, B. fusca Dombrow, new species; 68–69, B. setosa Dombrow, new species; 70–71, B. castanea Dombrow, new species; 72–73, B. colvillei Dombrow, 2005; 74–75, B. pulchra Dombrow, new species; 76–77, B. nigrofasciata Dombrow, new species; 78–79, B. caliginosa Dombrow, new species; 80–81, B. pumila Dombrow, new species; 82–83, B. gifbergensis Dombrow, new species; 84–85, B. julianae Dombrow, 2005; 86–87, B. elkeae Dombrow, new species; 88–89, B. suturalis Dombrow, new species; 90–91, B. nigrosetosa Dombrow, new species; 92–93, B. occidentalis Dombrow, 2005; 94–95, B. bicolor Dombrow, new species; 96–97, B. dolichiocnemis Dombrow, new species; 98–99, Anisochelus inornatus Burmeister, 1844. Scale bar 1mm." figureDoi="http://doi.org/10.5281/zenodo.4401954" httpUri="https://zenodo.org/record/4401954/files/figure.png" pageId="59" pageNumber="60">Figs. 88–89</figureCitation>
, 
<figureCitation id="134D1396E027FFCEFE07D001FD9CFBB9" box="[500,545,1050,1076]" captionStart="FIGURES 100–117" captionStartId="42.[151,264,1665,1689]" captionTargetBox="[162,1424,188,1625]" captionTargetId="figure-25@42.[151,1436,181,1641]" captionTargetPageId="42" captionText="FIGURES 100–117. Detailed drawings of the parameres for the key and the differential diagnosis of Beckhoplia Dombrow, 2005 species, in dorsal view, ♂♂. Arrows indicate characters of importance. 100, B. nigra Dombrow, new species; 101, B. pallidibrunnea Dombrow, new species; 102, B. fusca Dombrow, new species; 103, B. setosa Dombrow, new species; 104, B. castanea Dombrow, new species; 105, B. colvillei Dombrow, 2005; 106, B. pulchra Dombrow, new species; 107, B. nigrofasciata Dombrow, new species; 108, B. caliginosa Dombrow, new species; 109, B. pumila Dombrow, new species; 110, B. gifbergensis Dombrow, new species; 111, B. julianae Dombrow, 2005; 112, B. elkeae Dombrow, new species; 113, B. suturalis Dombrow, new species; 114, B. nigrosetosa Dombrow, new species; 115, B. occidentalis Dombrow, 2005; 116, B. bicolor Dombrow, new species; 117, B. dolichiocnemis Dombrow sp.n. Scale bar 1mm." figureDoi="http://doi.org/10.5281/zenodo.4401956" httpUri="https://zenodo.org/record/4401956/files/figure.png" pageId="59" pageNumber="60">113</figureCitation>
).
</paragraph>
<paragraph id="8BC90F13E027FFCEFF34D05BFCCBFBD4" blockId="59.[151,1437,150,2013]" box="[199,886,1086,1115]" pageId="59" pageNumber="60">
<emphasis id="B902D301E027FFCEFF34D05BFE05FBD5" bold="true" box="[199,440,1086,1115]" pageId="59" pageNumber="60">
<specimenCount id="9D70C49AE027FFCEFF34D05BFF4FFBD6" box="[199,242,1088,1115]" pageId="59" pageNumber="60" type="female">♀♀</specimenCount>
. Measurements:
</emphasis>
length 
<quantity id="4C8EA2F6E027FFCEFDFFD025FD2BFBD4" box="[524,662,1086,1113]" metricMagnitude="-3" metricUnit="m" metricValue="9.05" metricValueMax="9.5" metricValueMin="8.6" pageId="59" pageNumber="60" unit="mm" value="9.05" valueMax="9.5" valueMin="8.6">8.6–9.5 mm</quantity>
; width 
<quantity id="4C8EA2F6E027FFCEFD1BD025FCCFFBD4" box="[744,882,1086,1113]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" metricValueMax="5.2" metricValueMin="4.8" pageId="59" pageNumber="60" unit="mm" value="5.0" valueMax="5.2" valueMin="4.8">4.8–5.2 mm</quantity>
.
</paragraph>
<paragraph id="8BC90F13E027FFCEFF34D079FE3DFB2D" blockId="59.[151,1437,150,2013]" pageId="59" pageNumber="60">
<emphasis id="B902D301E027FFCEFF34D079FEE6FBF1" bold="true" box="[199,347,1122,1148]" pageId="59" pageNumber="60">Integument:</emphasis>
Lateral margins of elytra darkened; thoracic legs dark red-brown; metathoracic tibia and thoracic tarsi pale red-brown.
</paragraph>
<paragraph id="8BC90F13E027FFCEFF34D0B1FDBBF835" blockId="59.[151,1437,150,2013]" pageId="59" pageNumber="60">
<emphasis id="B902D301E027FFCEFF34D0B1FEFAFB49" bold="true" box="[199,327,1194,1220]" pageId="59" pageNumber="60">Tegument:</emphasis>
<emphasis id="B902D301E027FFCEFEA2D0B1FE16FB49" box="[337,427,1194,1220]" italics="true" pageId="59" pageNumber="60">Clypeus</emphasis>
loose setose with short, erect, pale-brown setae; medium-long and brown on head. 
<emphasis id="B902D301E027FFCEFA81D0B0FF53FB65" italics="true" pageId="59" pageNumber="60">Eye canthus</emphasis>
with a loose setose ciliary of medium-long, brown setae. 
<emphasis id="B902D301E027FFCEFC7AD0D4FC65FB64" box="[905,984,1231,1257]" italics="true" pageId="59" pageNumber="60">Scapus</emphasis>
loose setose with medium-long, brown setae, paler ventrally, shorter and pale-brown on 
<emphasis id="B902D301E027FFCEFD46D0E9FC2FFA81" box="[693,914,1266,1292]" italics="true" pageId="59" pageNumber="60">pedicellus; funicle I</emphasis>
bare, 
<emphasis id="B902D301E027FFCEFC25D0E9FBD9FA81" box="[982,1124,1266,1292]" italics="true" pageId="59" pageNumber="60">funicle II–IV</emphasis>
with extremely short, semierect, pale-brown setae, reclinate on 
<emphasis id="B902D301E027FFCEFDB0D10DFD36FABD" box="[579,651,1302,1328]" italics="true" pageId="59" pageNumber="60">clavus</emphasis>
. 
<emphasis id="B902D301E027FFCEFD6FD10CFCB6FABD" box="[668,779,1303,1328]" italics="true" pageId="59" pageNumber="60">Pronotum</emphasis>
without pattern. Loose setose with short and long, erect brown setae; with a loose setose ciliary of long, brown setae. 
<emphasis id="B902D301E027FFCEFCB5D121FC09FAD9" box="[838,948,1338,1364]" italics="true" pageId="59" pageNumber="60">Scutellum</emphasis>
loose setose with short, semi-erect, whitish setae. 
<emphasis id="B902D301E027FFCEFF2ED145FE9EFAF5" box="[221,291,1374,1400]" italics="true" pageId="59" pageNumber="60">Elytra</emphasis>
without pattern. Loose setose with short, reclinate, yellowish-brown setae; long, semi-erect, brown setae interspersed, denser towards lateral margin. Suture with a loose row of short and long, robust, semi-erect, black setae. 
<emphasis id="B902D301E027FFCEFF2DD1BDFE92FA4D" box="[222,303,1446,1472]" italics="true" pageId="59" pageNumber="60">Thorax</emphasis>
and 
<emphasis id="B902D301E027FFCEFE9BD1BDFE77FA4D" box="[360,458,1446,1472]" italics="true" pageId="59" pageNumber="60">pleurites</emphasis>
loose pilose with long, yellowish hairs. 
<emphasis id="B902D301E027FFCEFC66D1BCFBA1FA4D" box="[917,1052,1447,1472]" italics="true" pageId="59" pageNumber="60">Ventrite I–V</emphasis>
loose setose with short, reclinate, yellowish setae; long, semi-erect, yellowish and brown setae interspersed. 
<emphasis id="B902D301E027FFCEFC2ED1D1FBD3FA69" box="[989,1134,1482,1508]" italics="true" pageId="59" pageNumber="60">Propygidium</emphasis>
and 
<emphasis id="B902D301E027FFCEFB55D1D1FAB3FA69" box="[1190,1294,1482,1508]" italics="true" pageId="59" pageNumber="60">pygidium</emphasis>
loose setose with short and medium-long, reclinate, yellowish setae; medium-long, semi-erect, brown setae interspersed; tip of 
<emphasis id="B902D301E027FFCEFF64D209FF42F9A1" box="[151,255,1554,1580]" italics="true" pageId="59" pageNumber="60">pygidium</emphasis>
with a sparse setose ciliary long, brown setae. 
<emphasis id="B902D301E027FFCEFCF6D209FC6FF9A1" box="[773,978,1554,1580]" italics="true" pageId="59" pageNumber="60">Prothoracic femur</emphasis>
loose pilose with long, brown hairs. 
<emphasis id="B902D301E027FFCEFA99D208FE93F9DD" italics="true" pageId="59" pageNumber="60">Prothoracic tibia</emphasis>
loose setose with short, reclinate, brown setae; erect and long ventrally; a loose setose ciliary of short and medium-long, black and yellowish setae on internal ridge; with a setaceous ciliary of loose short pale-brown setae on anterior margin; a medium-long downwards pointing setose ciliary of pale-brown setae along anterior margin of apical tooth. 
<emphasis id="B902D301E027FFCEFE6AD2B9FDC6F931" box="[409,635,1698,1724]" italics="true" pageId="59" pageNumber="60">Mesothoracic femur</emphasis>
loose pilose with long, yellowish-brown hairs; distinctly long ventrally; long semi-erect brown setae interspersed externally. 
<emphasis id="B902D301E027FFCEFD15D2DDFC07F96D" box="[742,954,1734,1760]" italics="true" pageId="59" pageNumber="60">Mesothoracic tibia</emphasis>
sparse setose with short, reclinate, whitish setae; medium-long, semi-erect, robust, brown setae interspersed, long and finer internally; ciliary of posterior margin as in male but red-brown. 
<emphasis id="B902D301E027FFCEFE1AD315FD75F8A5" box="[489,712,1806,1832]" italics="true" pageId="59" pageNumber="60">Metathoracic femur</emphasis>
loose setose with long, semi-erect, yellowish-brown setae; paler ventrally. 
<emphasis id="B902D301E027FFCEFEF5D329FE6AF8C1" box="[262,471,1842,1868]" italics="true" pageId="59" pageNumber="60">Metathoracic tibia</emphasis>
sparse setose with short, reclinate, whitish setae; medium-long and robust, semi-erect, red-brown setae interspersed, loose, long and finer internally. Posterior margin with a bristled outer marginal ciliary of short, loose, transparently red-brown bristles, lengthening dorsally and ventrally and more robust ventrally. Setae of 
<emphasis id="B902D301E027FFCEFF45D385FEF0F835" box="[182,333,1950,1976]" italics="true" pageId="59" pageNumber="60">thoracic tarsi</emphasis>
pale red-brown.
</paragraph>
<paragraph id="8BC90F13E027FFC9FF34D3D9FBE9FF74" blockId="59.[151,1437,150,2013]" lastBlockId="60.[151,1437,151,939]" lastPageId="60" lastPageNumber="61" pageId="59" pageNumber="60">
<emphasis id="B902D301E027FFCEFF34D3D9FEADF851" bold="true" box="[199,272,1986,2012]" pageId="59" pageNumber="60">Head:</emphasis>
Clypeus very slightly depressed laterally; dense punctate with large and very large, deep, roundish punctures. Anterior margin of clypeus slightly vertically reflexed. Lateral margin slightly reflexed. Suture between head and clypeus undulate, crenated. Head dense punctate with small and large, deep, roundish punctures. Eye on head level. Funicle II–II cylindrical; clavus as long as pedicellus and funiculum combined.
</paragraph>
<paragraph id="8BC90F13E020FFC9FF34D518FD16FEE8" blockId="60.[151,1437,151,939]" pageId="60" pageNumber="61">
<emphasis id="B902D301E020FFC9FF34D518FEF4FE90" bold="true" box="[199,329,259,285]" pageId="60" pageNumber="61">Pronotum:</emphasis>
Widest in anterior third; moderate pulvinate; loose punctate with very small, shallow, roundish punctures, dense in posterior half of medial furrow. Anterior corner obtuse. Lateral margin convex. With an indistinct, complete, shallow, narrow medial furrow.
</paragraph>
<paragraph id="8BC90F13E020FFC9FF34D574FD96FE04" blockId="60.[151,1437,151,939]" box="[199,555,367,393]" pageId="60" pageNumber="61">
<emphasis id="B902D301E020FFC9FF34D574FEA0FE04" bold="true" box="[199,285,367,393]" pageId="60" pageNumber="61">Elytra:</emphasis>
Covering propygidium.
</paragraph>
<paragraph id="8BC90F13E020FFC9FF34D589FF60FE78" blockId="60.[151,1437,151,939]" pageId="60" pageNumber="61">
<emphasis id="B902D301E020FFC9FF34D589FEF8FE21" bold="true" box="[199,325,402,428]" pageId="60" pageNumber="61">Abdomen:</emphasis>
Distinctly pulvinate; ventrite II–V broadened medially. Pygidium slightly pulvinate longitudinally and laterally; slightly declined posteriorly; lateral margin subparallel in upper fifth, linearly attenuate apically; tip linear.
</paragraph>
<paragraph id="8BC90F13E020FFC9FF34D5E5FAD4FD08" blockId="60.[151,1437,151,939]" pageId="60" pageNumber="61">
<emphasis id="B902D301E020FFC9FF34D5E5FE3BFD94" bold="true" box="[199,390,510,537]" pageId="60" pageNumber="61">Prothoracic leg:</emphasis>
Femur robust. Median tooth of tibia broad triangular-shaped. Tarsomere I–IV combined very short; slightly more than 
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/ 
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<subScript id="17F20D56E020FFC9FE33D62EFE6DFDCE" attach="left" box="[448,464,565,579]" fontSize="6" pageId="60" pageNumber="61">10</subScript>
in
</quantity>
length of tibia. Pretarsomere slightly thickened; with a shortened smooth ridge ventrally, roundish cutting missing; slightly less than 
<superScript id="7C03A25BE020FFC9FD30D65CFD76FDD8" attach="right" box="[707,715,583,597]" fontSize="6" pageId="60" pageNumber="61">7</superScript>
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<subScript id="17F20D56E020FFC9FD20D642FD5EFDEA" attach="left" box="[723,739,601,615]" fontSize="6" pageId="60" pageNumber="61">10</subScript>
length of tarsomere I–IV combined. Internal claw slender; not broadened medially; slightly more than 
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/ 
<subScript id="17F20D56E020FFC9FD95D666FDCBFD06" attach="left" box="[614,630,637,651]" fontSize="6" pageId="60" pageNumber="61">10</subScript>
length of pretarsomere. External claw 
<superScript id="7C03A25BE020FFC9FBD8D671FB8EFDF5" attach="right" box="[1067,1075,618,632]" fontSize="6" pageId="60" pageNumber="61">4</superScript>
/ 
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<subScript id="17F20D56E020FFC9FBC8D666FBFEFD06" attach="left" box="[1083,1091,637,651]" fontSize="6" pageId="60" pageNumber="61">5</subScript>
in
</quantity>
length of internal claw.
</paragraph>
<paragraph id="8BC90F13E020FFC9FF34D695FDABFC98" blockId="60.[151,1437,151,939]" pageId="60" pageNumber="61">
<emphasis id="B902D301E020FFC9FF34D695FE21FD24" bold="true" box="[199,412,654,681]" pageId="60" pageNumber="61">Mesothoracic leg:</emphasis>
Tibia 
<superScript id="7C03A25BE020FFC9FE16D695FE50FD11" attach="right" box="[485,493,654,668]" fontSize="6" pageId="60" pageNumber="61">9</superScript>
/ 
<quantity id="4C8EA2F6E020FFC9FE06D6BAFD9EFD24" box="[501,547,655,687]" metricMagnitude="-1" metricUnit="m" metricValue="2.54" pageId="60" pageNumber="61" unit="in" value="10.0">
<subScript id="17F20D56E020FFC9FE06D6BAFDB8FD22" attach="left" box="[501,517,673,687]" fontSize="6" pageId="60" pageNumber="61">10</subScript>
in
</quantity>
length of femur. Tarsomere I–IV combined distinctly short; slightly more than 
<superScript id="7C03A25BE020FFC9FF64D6A9FF22FD4D" attach="none" box="[151,159,690,704]" fontSize="6" pageId="60" pageNumber="61">1</superScript>
/ 
<quantity id="4C8EA2F6E020FFC9FF54D6DEFF70FD40" box="[167,205,691,723]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="60" pageNumber="61" unit="in" value="2.0">
<subScript id="17F20D56E020FFC9FF54D6DEFF12FD5E" attach="left" box="[167,175,709,723]" fontSize="6" pageId="60" pageNumber="61">2</subScript>
in
</quantity>
length of tibia. Pretarsomere slightly thickened; roundish cutting missing; slightly more than 
<superScript id="7C03A25BE020FFC9FB02D6A8FB44FD4C" attach="right" box="[1265,1273,691,705]" fontSize="6" pageId="60" pageNumber="61">7</superScript>
/ 
<quantity id="4C8EA2F6E020FFC9FAF2D6DEFA92FD40" box="[1281,1327,691,723]" metricMagnitude="-1" metricUnit="m" metricValue="2.54" pageId="60" pageNumber="61" unit="in" value="10.0">
<subScript id="17F20D56E020FFC9FAF2D6DEFAACFD5E" attach="left" box="[1281,1297,709,723]" fontSize="6" pageId="60" pageNumber="61">10</subScript>
in
</quantity>
length of than tarsomere I–IV combined. External claw slender; not broadened medially; as long as pretarsomere. Internal claw 
<superScript id="7C03A25BE020FFC9FF20D6E1FF66FC85" attach="right" box="[211,219,762,776]" fontSize="6" pageId="60" pageNumber="61">4</superScript>
/ 
<quantity id="4C8EA2F6E020FFC9FF10D716FEB5FC98" box="[227,264,763,795]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="60" pageNumber="61" unit="in" value="5.0">
<subScript id="17F20D56E020FFC9FF10D716FF56FC96" attach="left" box="[227,235,781,795]" fontSize="6" pageId="60" pageNumber="61">5</subScript>
in
</quantity>
length of external claw.
</paragraph>
<paragraph id="8BC90F13E020FFC9FF34D705FB42FC28" blockId="60.[151,1437,151,939]" pageId="60" pageNumber="61">
<emphasis id="B902D301E020FFC9FF34D705FE21FCB4" bold="true" box="[199,412,798,825]" pageId="60" pageNumber="61">Metathoracic leg:</emphasis>
Femur slightly thickened. Tibia slightly extended dorso-apically; slightly less than 
<superScript id="7C03A25BE020FFC9FAAED705FAD8FCA1" attach="right" box="[1373,1381,798,812]" fontSize="6" pageId="60" pageNumber="61">9</superScript>
/ 
<quantity id="4C8EA2F6E020FFC9FA9ED72AFA21FCB4" box="[1389,1436,799,831]" metricMagnitude="-1" metricUnit="m" metricValue="2.54" pageId="60" pageNumber="61" unit="in" value="10.0">
<subScript id="17F20D56E020FFC9FA9ED72AFAC0FCB2" attach="left" box="[1389,1405,817,831]" fontSize="6" pageId="60" pageNumber="61">10</subScript>
in
</quantity>
length of femur. Tarsomere I–IV combined slightly more than 
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length of tibia. Pretarsomere slender; with a smooth ridge ventrally; slightly more than 
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length of tarsomere I–IV combined. External claw slightly more than 
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more in length of pretarsomere. Internal claw slightly less than 
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in
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length of external claw.
</paragraph>
</subSubSection>
<subSubSection id="C36C5C98E020FFC9FF64D7C9FCEBFBB8" pageId="60" pageNumber="61" type="distribution">
<paragraph id="8BC90F13E020FFC9FF64D7C9FBEDFC60" blockId="60.[151,1104,978,1005]" box="[151,1104,978,1005]" pageId="60" pageNumber="61">
<emphasis id="B902D301E020FFC9FF64D7C9FE8CFC61" bold="true" box="[151,305,978,1004]" pageId="60" pageNumber="61">Distribution.</emphasis>
<collectingRegion id="49B2C1F1E020FFC9FEC4D7C8FDF5FC60" box="[311,584,979,1005]" country="South Africa" name="Northern Cape" pageId="60" pageNumber="61">Northern Cape Province</collectingRegion>
(
<figureCitation id="134D1396E020FFC9FDA4D7C8FD05FC60" box="[599,696,979,1005]" captionStart="FIGURE 140" captionStartId="72.[151,250,1944,1968]" captionTargetBox="[195,1393,188,1910]" captionTargetId="figure-25@72.[185,1402,181,1920]" captionTargetPageId="72" captionText="FIGURE 140. Distributions of 18 Beckhoplia Dombrow, 2005 species, showing the concentration of species around the Northern Cape town of Nieuwoudtville." figureDoi="http://doi.org/10.5281/zenodo.4401964" httpUri="https://zenodo.org/record/4401964/files/figure.png" pageId="60" pageNumber="61">Fig. 140</figureCitation>
), only known from the 
<typeStatus id="54CDB1B1E020FFC9FC4CD7C8FC52FC60" box="[959,1007,979,1005]" pageId="60" pageNumber="61">type</typeStatus>
locality.
</paragraph>
<paragraph id="8BC90F13E020FFC9FF64D001FCEBFBB8" blockId="60.[151,854,1050,1077]" box="[151,854,1050,1077]" pageId="60" pageNumber="61">
<emphasis id="B902D301E020FFC9FF64D001FEE3FBB8" bold="true" box="[151,350,1050,1077]" pageId="60" pageNumber="61">
Vegetation 
<typeStatus id="54CDB1B1E020FFC9FEEFD000FEE4FBB8" box="[284,345,1051,1077]" pageId="60" pageNumber="61">Type</typeStatus>
.
</emphasis>
Nieuwoudtville Shale Renosterveld (FRs 2).
</paragraph>
</subSubSection>
<subSubSection id="C36C5C98E020FFC9FF64D078FD5DFBF3" box="[151,736,1123,1150]" pageId="60" pageNumber="61" type="biology_ecology">
<paragraph id="8BC90F13E020FFC9FF64D078FD5DFBF3" blockId="60.[151,736,1123,1150]" box="[151,736,1123,1150]" pageId="60" pageNumber="61">
<emphasis id="B902D301E020FFC9FF64D078FE95FBF0" bold="true" box="[151,296,1123,1149]" pageId="60" pageNumber="61">Host Plants.</emphasis>
<taxonomicName id="4C767490E020FFC9FEDCD078FE27FBF0" box="[303,410,1123,1149]" class="Liliopsida" family="Iridaceae" kingdom="Plantae" order="Asparagales" pageId="60" pageNumber="61" phylum="Tracheophyta" rank="family">Iridaceae</taxonomicName>
: 
<taxonomicName id="4C767490E020FFC9FE57D078FD61FBF3" authority="(Andrews)" baseAuthorityName="Andrews" box="[420,732,1123,1150]" class="Liliopsida" family="Iridaceae" genus="Moraea" kingdom="Plantae" order="Asparagales" pageId="60" pageNumber="61" phylum="Tracheophyta" rank="species" species="miniata">
<emphasis id="B902D301E020FFC9FE57D078FDE7FBF1" box="[420,602,1123,1148]" italics="true" pageId="60" pageNumber="61">Moraea miniata</emphasis>
(Andrews)
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C36C5C98E020FFC9FF64D0B0FC41FB48" box="[151,1020,1194,1221]" pageId="60" pageNumber="61" type="etymology">
<paragraph id="8BC90F13E020FFC9FF64D0B0FC41FB48" blockId="60.[151,1020,1194,1221]" box="[151,1020,1194,1221]" pageId="60" pageNumber="61">
<emphasis id="B902D301E020FFC9FF64D0B0FEA3FB48" bold="true" box="[151,286,1195,1221]" pageId="60" pageNumber="61">Etymology.</emphasis>
The name derives from the elevated suture [Latin: sutur = seam].
</paragraph>
</subSubSection>
</treatment>
</document>