752 lines
94 KiB
XML
752 lines
94 KiB
XML
<document id="41FC7E1C43A1D2C5D8DE4BC21581174D" ID-CLB-Dataset="8721" ID-DOI="10.11646/zootaxa.4823.1.1" ID-GBIF-Dataset="a43d0118-9df5-4970-a316-0dae5c9caebf" ID-ISSN="1175-5326" ID-Zenodo-Dep="4401916" ID-ZooBank="3DF06F26-FD35-452E-9056-1D05B49946EA" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1609340622775" checkinUser="plazi" docAuthor="Dombrow, Holger E. & Colville, Jonathan F." docDate="2020" docId="03DFBE05E009FFECFF64D65DFBB9F818" docLanguage="en" docName="zootaxa.4823.1.1.pdf" docOrigin="Zootaxa 4823" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Beckhoplia fusca Dombrow 2020, new species" docType="treatment" docUuid="9E5F22D0-AFDD-4010-B842-BD1B617E7533" docUuidSource="ZooBank" docVersion="9" lastPageNumber="26" masterDocId="FFE6C67DE01CFFF5FFF3D41BFFBDFF8D" masterDocTitle="Review of the genus Beckhoplia Dombrow, 2005 with the description of fifteen new species from South Africa and observations on its biogeography (Coleoptera: Scarabaeidae: Melolonthinae: Hopliini)" masterLastPageNumber="79" masterPageNumber="1" pageNumber="22" updateTime="1698849823920" updateUser="ExternalLinkService">
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<mods:titleInfo id="946D054943EC699E441E54F05504DB4B">
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<mods:title id="FD6B615FCB3FFF3D7C1093B7C123BCED">Review of the genus Beckhoplia Dombrow, 2005 with the description of fifteen new species from South Africa and observations on its biogeography (Coleoptera: Scarabaeidae: Melolonthinae: Hopliini)</mods:title>
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</mods:titleInfo>
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<mods:name id="88120F917953BF7C067FB3099A40E325" type="personal">
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<mods:role id="77056B92578F8259E1A8D3B6B1CAA988">
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<mods:roleTerm id="8349B9F837CFAC34BC8DE04EA6CBBB6D">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="C3C5844B7677CD13F36A443FB4E3790A">Dombrow, Holger E.</mods:namePart>
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<mods:nameIdentifier id="CA9ED3DA89150DB28F18E50992AD63D7" type="ORCID">0000-0003-2176-3077</mods:nameIdentifier>
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<mods:name id="27784E880897546C1D2312EBA642CB71" type="personal">
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<mods:roleTerm id="D6D980E9B58430FC55776466EFC30EB3">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="B5DD3C8CCC0601F804FE3D5F8A2F65AC">Colville, Jonathan F.</mods:namePart>
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<mods:nameIdentifier id="94E36274BE7908CB9A2ADD81607F9FD9" type="ORCID">0000-0003-2176-3077</mods:nameIdentifier>
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<mods:nameIdentifier id="D32C153FE5150CF3179DDEDA7D1E00E8" type="email">jonathan.colville@gmail.com</mods:nameIdentifier>
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<mods:typeOfResource id="8DDB44FF27E16970515BE03074139C92">text</mods:typeOfResource>
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<mods:title id="AD3BB362118ABDBB84B1C040B0EED77C">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="AA2F20767EACB815ABAF5CE534DE580C">
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<mods:date id="33B055EBAEE938B75DE0E1452AF90F3D">2020</mods:date>
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<mods:number id="537C3CD9DC067B81990136D2BA368A52">2020-08-10</mods:number>
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<mods:number id="771DAAEE54636746015B68EF9D2CF77F">4823</mods:number>
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<mods:start id="9FE8A005DEA3D1E696DF99E774B99BE5">1</mods:start>
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<mods:classification id="36050A8A614F5732017580EBA8BCB355">journal article</mods:classification>
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<mods:identifier id="0B9D7E11FFFB2DE6B6C0B81F864BCCAC" type="DOI">10.11646/zootaxa.4823.1.1</mods:identifier>
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<mods:identifier id="C83E000EFDB3796C5755DF02A48A2B2A" type="ISSN">1175-5326</mods:identifier>
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<treatment id="03DFBE05E009FFECFF64D65DFBB9F818" ID-DOI="http://doi.org/10.5281/zenodo.4455277" ID-GBIF-Taxon="177013530" ID-Zenodo-Dep="4455277" LSID="urn:lsid:zoobank.org:act:9E5F22D0-AFDD-4010-B842-BD1B617E7533" httpUri="http://treatment.plazi.org/id/03DFBE05E009FFECFF64D65DFBB9F818" lastPageId="25" lastPageNumber="26" pageId="21" pageNumber="22">
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<subSubSection id="C36C5C98E009FFE0FF64D65DFD28FDED" box="[151,661,582,609]" pageId="21" pageNumber="22" type="nomenclature">
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<paragraph id="8BC90F13E009FFE0FF64D65DFD28FDED" blockId="21.[151,883,582,680]" box="[151,661,582,609]" pageId="21" pageNumber="22">
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<heading id="D081B87FE009FFE0FF64D65DFD28FDED" bold="true" box="[151,661,582,609]" fontSize="11" level="2" pageId="21" pageNumber="22" reason="4">
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<emphasis id="B902D301E009FFE0FF64D65DFD28FDED" bold="true" box="[151,661,582,609]" pageId="21" pageNumber="22">
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<taxonomicName id="4C767490E009FFE0FF64D65DFE49FDEC" authority="Dombrow" authorityName="Dombrow" authorityYear="2020" box="[151,500,582,609]" class="Insecta" family="Melolonthidae" genus="Beckhoplia" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="fusca" status="sp. nov.">
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<emphasis id="B902D301E009FFE0FF64D65DFED3FDED" bold="true" box="[151,366,582,609]" italics="true" pageId="21" pageNumber="22">Beckhoplia fusca</emphasis>
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Dombrow
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</taxonomicName>
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,
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<taxonomicNameLabel id="A2316E7AE009FFE0FDF1D65CFD28FDED" box="[514,661,582,609]" pageId="21" pageNumber="22" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C36C5C98E009FFE0FF64D670FCCEFD25" pageId="21" pageNumber="22" type="description">
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<paragraph id="8BC90F13E009FFE0FF64D670FE0EFD08" blockId="21.[151,883,582,680]" box="[151,435,619,645]" pageId="21" pageNumber="22">
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(
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<figureCitation id="134D1396E009FFE0FF53D670FF4CFD08" box="[160,241,619,645]" captionStart="FIGURES 8–10" captionStartId="10.[151,264,1454,1478]" captionTargetBox="[155,1430,1277,1425]" captionTargetId="figure-142@10.[151,1436,1273,1430]" captionTargetPageId="10" captionText="FIGURES 8–10. General drawings of the position of the eye of Beckhoplia Dombrow, 2005 species, in lateral view. 8, eye below head level, ♀♀; 9, eye on head level, ♀♀; 10, eye above head level, ♂♂. Scale bar 0.5 mm" figureDoi="http://doi.org/10.5281/zenodo.4401924" httpUri="https://zenodo.org/record/4401924/files/figure.png" pageId="21" pageNumber="22">Figs. 9</figureCitation>
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,
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<figureCitation id="134D1396E009FFE0FF0DD670FEA0FD08" box="[254,285,619,645]" captionStart="FIGURES 15–18" captionStartId="16.[151,264,509,533]" captionTargetBox="[199,1387,184,446]" captionTargetId="figure-25@16.[195,1392,181,449]" captionTargetPageId="16" captionText="FIGURES 15–18. General shape of the antenna and clavus of Beckhoplia Dombrow, 2005 species, ♂♂, in lateral view, other- wise indicated. 15, nine segmented antenna: a, scapus; b, pedicellus; c, funicle I–IV = funiculum; d, three segmented lamellar clavus. Scale bar 1 mm. 16, clavus short elliptic, ♀♀; 17, clavus elliptic; 18, clavus long elliptic. Scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4401928" httpUri="https://zenodo.org/record/4401928/files/figure.png" pageId="21" pageNumber="22">16</figureCitation>
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,
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<figureCitation id="134D1396E009FFE0FEDAD670FECEFD08" box="[297,371,619,645]" captionStart="FIGURES 62–97" captionStartId="28.[151,264,1743,1767]" captionTargetBox="[161,1426,187,1710]" captionTargetId="figure-25@28.[151,1436,181,1719]" captionTargetPageId="28" captionText="FIGURES 62–97. Parameres and phallobase of Beckhoplia Dombrow, 2005 species, in dorsal view (left side), in lateral view (right side), ♂♂. 62–63, B. nigra Dombrow, new species; 64–65, B. pallidibrunnea Dombrow, new species; 66–67, B. fusca Dombrow, new species; 68–69, B. setosa Dombrow, new species; 70–71, B. castanea Dombrow, new species; 72–73, B. colvillei Dombrow, 2005; 74–75, B. pulchra Dombrow, new species; 76–77, B. nigrofasciata Dombrow, new species; 78–79, B. caliginosa Dombrow, new species; 80–81, B. pumila Dombrow, new species; 82–83, B. gifbergensis Dombrow, new species; 84–85, B. julianae Dombrow, 2005; 86–87, B. elkeae Dombrow, new species; 88–89, B. suturalis Dombrow, new species; 90–91, B. nigrosetosa Dombrow, new species; 92–93, B. occidentalis Dombrow, 2005; 94–95, B. bicolor Dombrow, new species; 96–97, B. dolichiocnemis Dombrow, new species; 98–99, Anisochelus inornatus Burmeister, 1844. Scale bar 1mm." figureDoi="http://doi.org/10.5281/zenodo.4401954" httpUri="https://zenodo.org/record/4401954/files/figure.png" pageId="21" pageNumber="22">66–67</figureCitation>
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,
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<figureCitation id="134D1396E009FFE0FE8CD670FE16FD08" box="[383,427,619,645]" captionStart="FIGURES 100–117" captionStartId="42.[151,264,1665,1689]" captionTargetBox="[162,1424,188,1625]" captionTargetId="figure-25@42.[151,1436,181,1641]" captionTargetPageId="42" captionText="FIGURES 100–117. Detailed drawings of the parameres for the key and the differential diagnosis of Beckhoplia Dombrow, 2005 species, in dorsal view, ♂♂. Arrows indicate characters of importance. 100, B. nigra Dombrow, new species; 101, B. pallidibrunnea Dombrow, new species; 102, B. fusca Dombrow, new species; 103, B. setosa Dombrow, new species; 104, B. castanea Dombrow, new species; 105, B. colvillei Dombrow, 2005; 106, B. pulchra Dombrow, new species; 107, B. nigrofasciata Dombrow, new species; 108, B. caliginosa Dombrow, new species; 109, B. pumila Dombrow, new species; 110, B. gifbergensis Dombrow, new species; 111, B. julianae Dombrow, 2005; 112, B. elkeae Dombrow, new species; 113, B. suturalis Dombrow, new species; 114, B. nigrosetosa Dombrow, new species; 115, B. occidentalis Dombrow, 2005; 116, B. bicolor Dombrow, new species; 117, B. dolichiocnemis Dombrow sp.n. Scale bar 1mm." figureDoi="http://doi.org/10.5281/zenodo.4401956" httpUri="https://zenodo.org/record/4401956/files/figure.png" pageId="21" pageNumber="22">102</figureCitation>
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)
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</paragraph>
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<paragraph id="8BC90F13E009FFE0FF64D68BFCCEFD25" blockId="21.[151,883,582,680]" box="[151,883,656,680]" pageId="21" pageNumber="22">
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http://zoobank.org/
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<uuid id="FFD035C6E009FFE0FE98D68BFCCEFD25" box="[363,883,656,680]" pageId="21" pageNumber="22">9E5F22D0-AFDD-4010-B842-BD1B617E7533</uuid>
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</paragraph>
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||
</subSubSection>
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<subSubSection id="C36C5C98E009FFE0FF64D6C3FF5EFBD7" pageId="21" pageNumber="22" type="materials_examined">
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<paragraph id="8BC90F13E009FFE0FF64D6C3FCF9FD7F" blockId="21.[151,836,728,755]" box="[151,836,728,755]" pageId="21" pageNumber="22">
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<materialsCitation id="3B1E054EE009FFE0FF64D6C3FC82FD7F" ID-GBIF-Occurrence="3022772308" box="[151,831,728,755]" collectionCode="NCP" country="South Africa" location="RSA" pageId="21" pageNumber="22" specimenCount="1" typeStatus="holotype">
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<emphasis id="B902D301E009FFE0FF64D6C3FE84FD7F" bold="true" box="[151,313,728,754]" pageId="21" pageNumber="22">
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||
<typeStatus id="54CDB1B1E009FFE0FF64D6C3FF6EFD7F" box="[151,211,728,754]" pageId="21" pageNumber="22">Type</typeStatus>
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locality.
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</emphasis>
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[
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<collectingCountry id="F3614F83E009FFE0FEB4D6C3FE39FD7F" box="[327,388,728,754]" name="South Africa" pageId="21" pageNumber="22">RSA</collectingCountry>
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,
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||
<collectionCode id="ED6797D6E009FFE0FE63D6C3FE72FD7E" box="[400,463,728,755]" country="United Kingdom" name="National Collection of Passiflora" pageId="21" pageNumber="22" type="Herbarium">NCP</collectionCode>
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]:
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<quantity id="4C8EA2F6E009FFE0FE13D6C3FDA7FD7F" box="[480,538,728,754]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="21" pageNumber="22" unit="km" value="1.0">1 km</quantity>
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nördl. Farm Wilemsriver
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</materialsCitation>
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.
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</paragraph>
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<paragraph id="8BC90F13E009FFE0FF64D73BFB44FC63" blockId="21.[151,1437,800,1007]" pageId="21" pageNumber="22">
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<emphasis id="B902D301E009FFE0FF64D73BFEF7FCB7" bold="true" box="[151,330,800,826]" pageId="21" pageNumber="22">Type material.</emphasis>
|
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[
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<materialsCitation id="3B1E054EE009FFE0FEA8D73BFECAFC47" ID-GBIF-Occurrence="3022772306" collectingDate="1989-08-30" collectionCode="R, ZSM" collectorName="Dombrow" country="South Africa" determinerName="H. Dombrow" elevation="750" latitude="-31.337223" location="Farm Wilemsriver" longLatPrecision="20" longitude="19.086945" pageId="21" pageNumber="22" specimenCount="1" specimenCount-male="1" stateProvince="Western Cape" typeStatus="holotype">
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||
9]
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<emphasis id="B902D301E009FFE0FE8FD73BFE56FCB7" bold="true" box="[380,491,800,826]" pageId="21" pageNumber="22">
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<typeStatus id="54CDB1B1E009FFE0FE8FD73BFE56FCB7" box="[380,491,800,826]" pageId="21" pageNumber="22" type="holotype">Holotype</typeStatus>
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||
</emphasis>
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||
<specimenCount id="9D70C49AE009FFE0FE07D73AFDB0FCB6" box="[500,525,801,827]" pageId="21" pageNumber="22" type="male">♂</specimenCount>
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: yellow label hw «54», pr «.920» | coll.
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<collectorName id="26836AC5E009FFE0FC17D73BFBE5FCB7" box="[996,1112,800,826]" pageId="21" pageNumber="22">Dombrow</collectorName>
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|| white label pr «AFRIKA:
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||
<collectingCountry id="F3614F83E009FFE0FF64D75FFEB7FCD3" box="[151,266,836,862]" name="South Africa" pageId="21" pageNumber="22">Südafrika</collectingCountry>
|
||
,
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||
<collectingRegion id="49B2C1F1E009FFE0FEE5D75FFE20FCD3" box="[278,413,836,862]" country="South Africa" name="Western Cape" pageId="21" pageNumber="22">Kapprovinz</collectingRegion>
|
||
|
|
||
<quantity id="4C8EA2F6E009FFE0FE46D75FFE52FCD3" box="[437,495,836,862]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="21" pageNumber="22" unit="km" value="1.0">1 km</quantity>
|
||
nördl.
|
||
<location id="8EA959C8E009FFE0FDB6D75EFCAAFCD3" LSID="urn:lsid:plazi:treatment:03DFBE05E009FFECFF64D65DFBB9F818:8EA959C8E009FFE0FDB6D75EFCAAFCD3" box="[581,791,836,863]" country="South Africa" latitude="-31.337223" longLatPrecision="20" longitude="19.086945" name="Farm Wilemsriver" pageId="21" pageNumber="22" stateProvince="Western Cape">Farm Wilemsriver</location>
|
||
|
|
||
<geoCoordinate id="EE4269D4E009FFE0FCDDD75FFC05FCD3" box="[814,952,836,862]" degrees="31" direction="south" minutes="20" orientation="latitude" pageId="21" pageNumber="22" precision="15" seconds="14" value="-31.337223">31°20’14”S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE4269D4E009FFE0FC36D75FFBECFCD2" box="[965,1105,836,863]" degrees="19" direction="east" minutes="05" orientation="longitude" pageId="21" pageNumber="22" precision="15" seconds="13" value="19.086945">19°05’13”E</geoCoordinate>
|
||
,
|
||
<quantity id="4C8EA2F6E009FFE0FBADD75FFB1AFCD2" box="[1118,1191,836,863]" metricMagnitude="2" metricUnit="m" metricValue="7.5" pageId="21" pageNumber="22" unit="m" value="750.0">
|
||
<elevation id="005BE820E009FFE0FBADD75FFB1AFCD2" box="[1118,1191,836,863]" metricMagnitude="2" metricUnit="m" metricValue="7.5" pageId="21" pageNumber="22" unit="m" value="750.0">750 m</elevation>
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||
</quantity>
|
||
|
|
||
<date id="FFC829D3E009FFE0FB4CD75FFAFFFCD3" box="[1215,1346,836,862]" pageId="21" pageNumber="22" value="1989-08-30">
|
||
<collectingDate id="EF8CD03BE009FFE0FB4CD75FFAFFFCD3" box="[1215,1346,836,862]" pageId="21" pageNumber="22" value="1989-08-30">30.08.1989</collectingDate>
|
||
</date>
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||
, leg.
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||
<collectionCode id="ED6797D6E009FFE0FA71D75EFA25FCD2" box="[1410,1432,837,863]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="21" pageNumber="22">R</collectionCode>
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||
. Fritz» || green label pr «Host plant: |
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||
<taxonomicName id="4C767490E009FFE0FDC4D772FD5AFC0F" authorityName="L.Bolus" authorityYear="1920" box="[567,743,872,898]" class="Liliopsida" family="Iridaceae" genus="Homeria" higherTaxonomySource="CoL" kingdom="Plantae" order="Asparagales" pageId="21" pageNumber="22" phylum="Tracheophyta" rank="species" species="bifida">
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||
<emphasis id="B902D301E009FFE0FDC4D772FD5AFC0F" box="[567,743,872,898]" italics="true" pageId="21" pageNumber="22">Homeria bifida</emphasis>
|
||
</taxonomicName>
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||
» || blue label pr «Bioregion: | Sandstone Fynbos | Vegetation Type: FFs1 – | Bokkeveld Sandstone Fynbos» || red label pr «
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||
<typeStatus id="54CDB1B1E009FFE0FCA4D797FC79FC2B" box="[855,964,908,934]" pageId="21" pageNumber="22" type="holotype">Holotype</typeStatus>
|
||
|
|
||
<taxonomicName id="4C767490E009FFE0FC2AD797FBE4FC2B" authorityName="Dombrow" authorityYear="2005" box="[985,1113,908,934]" class="Insecta" family="Melolonthidae" genus="Beckhoplia" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B902D301E009FFE0FC2AD797FBE4FC2B" box="[985,1113,908,934]" italics="true" pageId="21" pageNumber="22">Beckhoplia</emphasis>
|
||
</taxonomicName>
|
||
|
|
||
<taxonomicName id="4C767490E009FFE0FB9DD797FB17FC2B" authority="Dombrow & Colville, 2020" authorityName="Dombrow" authorityYear="2020" box="[1134,1194,908,934]" class="Insecta" family="Melolonthidae" genus="Beckhoplia" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="fusca" status="sp. nov.">
|
||
<emphasis id="B902D301E009FFE0FB9DD797FB17FC2B" box="[1134,1194,908,934]" italics="true" pageId="21" pageNumber="22">fusca</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2316E7AE009FFE0FB41D796FB55FC2A" box="[1202,1256,909,935]" pageId="21" pageNumber="22" rank="species">sp.n.</taxonomicNameLabel>
|
||
| det.:
|
||
<determinerName id="7E15E3D7E009FFE0FAC4D796FF6FFC47" pageId="21" pageNumber="22">H. Dombrow</determinerName>
|
||
2019» (
|
||
<collectionCode id="ED6797D6E009FFE0FEDDD7ABFED2FC47" box="[302,367,944,970]" country="Germany" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="21" pageNumber="22" type="Museum">ZSM</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
. –
|
||
<materialsCitation id="3B1E054EE009FFE0FE65D7ABFB48FC63" ID-GBIF-Occurrence="3022772314" collectionCode="ESDW" collectorName="Dombrow" country="South Africa" determinerName="H. Dombrow" pageId="21" pageNumber="22" specimenCount="8" specimenCount-female="8" stateProvince="Western Cape" typeStatus="paratype">
|
||
<emphasis id="B902D301E009FFE0FE65D7ABFDADFC47" bold="true" box="[406,528,944,970]" pageId="21" pageNumber="22">
|
||
<typeStatus id="54CDB1B1E009FFE0FE65D7ABFDADFC47" box="[406,528,944,970]" pageId="21" pageNumber="22" type="paratype">Paratypes</typeStatus>
|
||
</emphasis>
|
||
<specimenCount id="9D70C49AE009FFE0FDE4D7ABFDE4FC46" box="[535,601,944,971]" pageId="21" pageNumber="22" type="female">8 ♀♀</specimenCount>
|
||
: same labels as
|
||
<typeStatus id="54CDB1B1E009FFE0FCFDD7ABFCCFFC47" box="[782,882,944,970]" pageId="21" pageNumber="22" type="holotype">holotype</typeStatus>
|
||
but || yellow label hw «54», pr «.982–.989 | coll.
|
||
<collectorName id="26836AC5E009FFE0FF64D7CFFEB0FC63" box="[151,269,980,1006]" pageId="21" pageNumber="22">Dombrow</collectorName>
|
||
» || red label pr «
|
||
<typeStatus id="54CDB1B1E009FFE0FE3AD7CEFD8DFC62" box="[457,560,981,1007]" pageId="21" pageNumber="22" type="paratype">Paratype</typeStatus>
|
||
|
|
||
<taxonomicName id="4C767490E009FFE0FDB7D7CFFD79FC63" authorityName="Dombrow" authorityYear="2005" box="[580,708,980,1006]" class="Insecta" family="Melolonthidae" genus="Beckhoplia" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B902D301E009FFE0FDB7D7CFFD79FC63" box="[580,708,980,1006]" italics="true" pageId="21" pageNumber="22">Beckhoplia</emphasis>
|
||
</taxonomicName>
|
||
|
|
||
<taxonomicName id="4C767490E009FFE0FD29D7CFFCABFC63" authority="Dombrow & Colville, 2020" authorityName="Dombrow" authorityYear="2020" box="[730,790,980,1006]" class="Insecta" family="Melolonthidae" genus="Beckhoplia" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="fusca" status="sp. nov.">
|
||
<emphasis id="B902D301E009FFE0FD29D7CFFCABFC63" box="[730,790,980,1006]" italics="true" pageId="21" pageNumber="22">fusca</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2316E7AE009FFE0FCEED7CEFCE9FC62" box="[797,852,981,1007]" pageId="21" pageNumber="22" rank="species">sp.n.</taxonomicNameLabel>
|
||
| det.:
|
||
<determinerName id="7E15E3D7E009FFE0FC51D7CEFB84FC63" box="[930,1081,980,1007]" pageId="21" pageNumber="22">H. Dombrow</determinerName>
|
||
2019» (
|
||
<collectionCode id="ED6797D6E009FFE0FB65D7CFFB50FC63" box="[1174,1261,980,1006]" pageId="21" pageNumber="22">ESDW</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BC90F13E009FFE0FF64D007FF5EFBD7" blockId="21.[151,1437,1052,1114]" pageId="21" pageNumber="22">
|
||
<emphasis id="B902D301E009FFE0FF64D007FE0DFBBB" bold="true" box="[151,432,1052,1078]" pageId="21" pageNumber="22">
|
||
Condition of
|
||
<typeStatus id="54CDB1B1E009FFE0FECCD007FE17FBBB" box="[319,426,1052,1078]" pageId="21" pageNumber="22" type="holotype">holotype</typeStatus>
|
||
.
|
||
</emphasis>
|
||
Excellent. Glued on card. Dissected, parameres and spiculum gastrale glued behind beetle.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36C5C98E009FFE0FF64D093FA81FADB" pageId="21" pageNumber="22" type="diagnosis">
|
||
<paragraph id="8BC90F13E009FFE0FF64D093FA81FADB" blockId="21.[151,1437,1160,1367]" pageId="21" pageNumber="22">
|
||
<emphasis id="B902D301E009FFE0FF64D093FE18FB2F" bold="true" box="[151,421,1160,1186]" pageId="21" pageNumber="22">Differential diagnosis.</emphasis>
|
||
Differs from
|
||
<taxonomicName id="4C767490E009FFE0FDBAD092FC24FB2F" authority=": Scutellum" authorityName="Scutellum" box="[585,921,1160,1187]" class="Insecta" family="Melolonthidae" genus="Beckhoplia" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="pallidibrunnea">
|
||
<emphasis id="B902D301E009FFE0FDBAD092FCA9FB2F" box="[585,788,1160,1186]" italics="true" pageId="21" pageNumber="22">B. pallidibrunnea</emphasis>
|
||
: Scutellum
|
||
</taxonomicName>
|
||
of
|
||
<taxonomicName id="4C767490E009FFE0FC30D092FB9CFB2F" authorityName="Dombrow" authorityYear="2020" box="[963,1057,1160,1186]" class="Insecta" family="Melolonthidae" genus="Beckhoplia" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="fusca">
|
||
<emphasis id="B902D301E009FFE0FC30D092FB9CFB2F" box="[963,1057,1160,1186]" italics="true" pageId="21" pageNumber="22">B. fusca</emphasis>
|
||
</taxonomicName>
|
||
longer than broad. Scutellum of
|
||
<taxonomicName id="4C767490E009FFE0FF64D0B6FED9FB4B" box="[151,356,1196,1222]" class="Insecta" family="Melolonthidae" genus="Beckhoplia" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="pallidibrunnea">
|
||
<emphasis id="B902D301E009FFE0FF64D0B6FED9FB4B" box="[151,356,1196,1222]" italics="true" pageId="21" pageNumber="22">B. pallidibrunnea</emphasis>
|
||
</taxonomicName>
|
||
as broad as long. Pretarsomere of metathoracic tarsus of
|
||
<taxonomicName id="4C767490E009FFE0FBE5D0B6FBC8FB4B" authorityName="Dombrow" authorityYear="2020" box="[1046,1141,1196,1222]" class="Insecta" family="Melolonthidae" genus="Beckhoplia" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="fusca">
|
||
<emphasis id="B902D301E009FFE0FBE5D0B6FBC8FB4B" box="[1046,1141,1196,1222]" italics="true" pageId="21" pageNumber="22">B. fusca</emphasis>
|
||
</taxonomicName>
|
||
slender. Pretarsomere of metathoracic tarsus of
|
||
<taxonomicName id="4C767490E009FFE0FE51D0CAFDD3FB67" box="[418,622,1232,1258]" class="Insecta" family="Melolonthidae" genus="Beckhoplia" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="pallidibrunnea">
|
||
<emphasis id="B902D301E009FFE0FE51D0CAFDD3FB67" box="[418,622,1232,1258]" italics="true" pageId="21" pageNumber="22">B. pallidibrunnea</emphasis>
|
||
</taxonomicName>
|
||
slightly thickened. Interno-posterior margin of parameres of
|
||
<taxonomicName id="4C767490E009FFE0FACDD0CAFA21FB67" authorityName="Dombrow" authorityYear="2020" box="[1342,1436,1232,1258]" class="Insecta" family="Melolonthidae" genus="Beckhoplia" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="fusca">
|
||
<emphasis id="B902D301E009FFE0FACDD0CAFA21FB67" box="[1342,1436,1232,1258]" italics="true" pageId="21" pageNumber="22">B. fusca</emphasis>
|
||
</taxonomicName>
|
||
narrowed curly bracket-shaped (
|
||
<figureCitation id="134D1396E009FFE0FDFDD0EFFDD2FA83" box="[526,623,1268,1294]" captionStart="FIGURES 62–97" captionStartId="28.[151,264,1743,1767]" captionTargetBox="[161,1426,187,1710]" captionTargetId="figure-25@28.[151,1436,181,1719]" captionTargetPageId="28" captionText="FIGURES 62–97. Parameres and phallobase of Beckhoplia Dombrow, 2005 species, in dorsal view (left side), in lateral view (right side), ♂♂. 62–63, B. nigra Dombrow, new species; 64–65, B. pallidibrunnea Dombrow, new species; 66–67, B. fusca Dombrow, new species; 68–69, B. setosa Dombrow, new species; 70–71, B. castanea Dombrow, new species; 72–73, B. colvillei Dombrow, 2005; 74–75, B. pulchra Dombrow, new species; 76–77, B. nigrofasciata Dombrow, new species; 78–79, B. caliginosa Dombrow, new species; 80–81, B. pumila Dombrow, new species; 82–83, B. gifbergensis Dombrow, new species; 84–85, B. julianae Dombrow, 2005; 86–87, B. elkeae Dombrow, new species; 88–89, B. suturalis Dombrow, new species; 90–91, B. nigrosetosa Dombrow, new species; 92–93, B. occidentalis Dombrow, 2005; 94–95, B. bicolor Dombrow, new species; 96–97, B. dolichiocnemis Dombrow, new species; 98–99, Anisochelus inornatus Burmeister, 1844. Scale bar 1mm." figureDoi="http://doi.org/10.5281/zenodo.4401954" httpUri="https://zenodo.org/record/4401954/files/figure.png" pageId="21" pageNumber="22">Figs. 66</figureCitation>
|
||
,
|
||
<figureCitation id="134D1396E009FFE0FD8FD0EFFD16FA83" box="[636,683,1268,1294]" captionStart="FIGURES 100–117" captionStartId="42.[151,264,1665,1689]" captionTargetBox="[162,1424,188,1625]" captionTargetId="figure-25@42.[151,1436,181,1641]" captionTargetPageId="42" captionText="FIGURES 100–117. Detailed drawings of the parameres for the key and the differential diagnosis of Beckhoplia Dombrow, 2005 species, in dorsal view, ♂♂. Arrows indicate characters of importance. 100, B. nigra Dombrow, new species; 101, B. pallidibrunnea Dombrow, new species; 102, B. fusca Dombrow, new species; 103, B. setosa Dombrow, new species; 104, B. castanea Dombrow, new species; 105, B. colvillei Dombrow, 2005; 106, B. pulchra Dombrow, new species; 107, B. nigrofasciata Dombrow, new species; 108, B. caliginosa Dombrow, new species; 109, B. pumila Dombrow, new species; 110, B. gifbergensis Dombrow, new species; 111, B. julianae Dombrow, 2005; 112, B. elkeae Dombrow, new species; 113, B. suturalis Dombrow, new species; 114, B. nigrosetosa Dombrow, new species; 115, B. occidentalis Dombrow, 2005; 116, B. bicolor Dombrow, new species; 117, B. dolichiocnemis Dombrow sp.n. Scale bar 1mm." figureDoi="http://doi.org/10.5281/zenodo.4401956" httpUri="https://zenodo.org/record/4401956/files/figure.png" pageId="21" pageNumber="22">102</figureCitation>
|
||
). Interno-posterior margin of parameres of
|
||
<taxonomicName id="4C767490E009FFE0FB57D0EEFAD2FA83" box="[1188,1391,1268,1294]" class="Insecta" family="Melolonthidae" genus="Beckhoplia" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="pallidibrunnea">
|
||
<emphasis id="B902D301E009FFE0FB57D0EEFAD2FA83" box="[1188,1391,1268,1294]" italics="true" pageId="21" pageNumber="22">B. pallidibrunnea</emphasis>
|
||
</taxonomicName>
|
||
parabola-shaped (
|
||
<figureCitation id="134D1396E009FFE0FEB4D103FE1BFABF" box="[327,422,1304,1330]" captionStart="FIGURES 62–97" captionStartId="28.[151,264,1743,1767]" captionTargetBox="[161,1426,187,1710]" captionTargetId="figure-25@28.[151,1436,181,1719]" captionTargetPageId="28" captionText="FIGURES 62–97. Parameres and phallobase of Beckhoplia Dombrow, 2005 species, in dorsal view (left side), in lateral view (right side), ♂♂. 62–63, B. nigra Dombrow, new species; 64–65, B. pallidibrunnea Dombrow, new species; 66–67, B. fusca Dombrow, new species; 68–69, B. setosa Dombrow, new species; 70–71, B. castanea Dombrow, new species; 72–73, B. colvillei Dombrow, 2005; 74–75, B. pulchra Dombrow, new species; 76–77, B. nigrofasciata Dombrow, new species; 78–79, B. caliginosa Dombrow, new species; 80–81, B. pumila Dombrow, new species; 82–83, B. gifbergensis Dombrow, new species; 84–85, B. julianae Dombrow, 2005; 86–87, B. elkeae Dombrow, new species; 88–89, B. suturalis Dombrow, new species; 90–91, B. nigrosetosa Dombrow, new species; 92–93, B. occidentalis Dombrow, 2005; 94–95, B. bicolor Dombrow, new species; 96–97, B. dolichiocnemis Dombrow, new species; 98–99, Anisochelus inornatus Burmeister, 1844. Scale bar 1mm." figureDoi="http://doi.org/10.5281/zenodo.4401954" httpUri="https://zenodo.org/record/4401954/files/figure.png" pageId="21" pageNumber="22">Figs. 64</figureCitation>
|
||
,
|
||
<figureCitation id="134D1396E009FFE0FE41D103FE5DFABF" box="[434,480,1304,1330]" captionStart="FIGURES 100–117" captionStartId="42.[151,264,1665,1689]" captionTargetBox="[162,1424,188,1625]" captionTargetId="figure-25@42.[151,1436,181,1641]" captionTargetPageId="42" captionText="FIGURES 100–117. Detailed drawings of the parameres for the key and the differential diagnosis of Beckhoplia Dombrow, 2005 species, in dorsal view, ♂♂. Arrows indicate characters of importance. 100, B. nigra Dombrow, new species; 101, B. pallidibrunnea Dombrow, new species; 102, B. fusca Dombrow, new species; 103, B. setosa Dombrow, new species; 104, B. castanea Dombrow, new species; 105, B. colvillei Dombrow, 2005; 106, B. pulchra Dombrow, new species; 107, B. nigrofasciata Dombrow, new species; 108, B. caliginosa Dombrow, new species; 109, B. pumila Dombrow, new species; 110, B. gifbergensis Dombrow, new species; 111, B. julianae Dombrow, 2005; 112, B. elkeae Dombrow, new species; 113, B. suturalis Dombrow, new species; 114, B. nigrosetosa Dombrow, new species; 115, B. occidentalis Dombrow, 2005; 116, B. bicolor Dombrow, new species; 117, B. dolichiocnemis Dombrow sp.n. Scale bar 1mm." figureDoi="http://doi.org/10.5281/zenodo.4401956" httpUri="https://zenodo.org/record/4401956/files/figure.png" pageId="21" pageNumber="22">101</figureCitation>
|
||
). Externo-posterior margin of parameres of
|
||
<taxonomicName id="4C767490E009FFE0FC26D102FB8CFABF" authorityName="Dombrow" authorityYear="2020" box="[981,1073,1304,1330]" class="Insecta" family="Melolonthidae" genus="Beckhoplia" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="fusca">
|
||
<emphasis id="B902D301E009FFE0FC26D102FB8CFABF" box="[981,1073,1304,1330]" italics="true" pageId="21" pageNumber="22">B. fusca</emphasis>
|
||
</taxonomicName>
|
||
convex, incised medially (
|
||
<figureCitation id="134D1396E009FFE0FA90D103FF0BFADB" captionStart="FIGURES 62–97" captionStartId="28.[151,264,1743,1767]" captionTargetBox="[161,1426,187,1710]" captionTargetId="figure-25@28.[151,1436,181,1719]" captionTargetPageId="28" captionText="FIGURES 62–97. Parameres and phallobase of Beckhoplia Dombrow, 2005 species, in dorsal view (left side), in lateral view (right side), ♂♂. 62–63, B. nigra Dombrow, new species; 64–65, B. pallidibrunnea Dombrow, new species; 66–67, B. fusca Dombrow, new species; 68–69, B. setosa Dombrow, new species; 70–71, B. castanea Dombrow, new species; 72–73, B. colvillei Dombrow, 2005; 74–75, B. pulchra Dombrow, new species; 76–77, B. nigrofasciata Dombrow, new species; 78–79, B. caliginosa Dombrow, new species; 80–81, B. pumila Dombrow, new species; 82–83, B. gifbergensis Dombrow, new species; 84–85, B. julianae Dombrow, 2005; 86–87, B. elkeae Dombrow, new species; 88–89, B. suturalis Dombrow, new species; 90–91, B. nigrosetosa Dombrow, new species; 92–93, B. occidentalis Dombrow, 2005; 94–95, B. bicolor Dombrow, new species; 96–97, B. dolichiocnemis Dombrow, new species; 98–99, Anisochelus inornatus Burmeister, 1844. Scale bar 1mm." figureDoi="http://doi.org/10.5281/zenodo.4401954" httpUri="https://zenodo.org/record/4401954/files/figure.png" pageId="21" pageNumber="22">Figs. 66</figureCitation>
|
||
,
|
||
<figureCitation id="134D1396E009FFE0FF31D127FF4CFADB" box="[194,241,1340,1366]" captionStart="FIGURES 100–117" captionStartId="42.[151,264,1665,1689]" captionTargetBox="[162,1424,188,1625]" captionTargetId="figure-25@42.[151,1436,181,1641]" captionTargetPageId="42" captionText="FIGURES 100–117. Detailed drawings of the parameres for the key and the differential diagnosis of Beckhoplia Dombrow, 2005 species, in dorsal view, ♂♂. Arrows indicate characters of importance. 100, B. nigra Dombrow, new species; 101, B. pallidibrunnea Dombrow, new species; 102, B. fusca Dombrow, new species; 103, B. setosa Dombrow, new species; 104, B. castanea Dombrow, new species; 105, B. colvillei Dombrow, 2005; 106, B. pulchra Dombrow, new species; 107, B. nigrofasciata Dombrow, new species; 108, B. caliginosa Dombrow, new species; 109, B. pumila Dombrow, new species; 110, B. gifbergensis Dombrow, new species; 111, B. julianae Dombrow, 2005; 112, B. elkeae Dombrow, new species; 113, B. suturalis Dombrow, new species; 114, B. nigrosetosa Dombrow, new species; 115, B. occidentalis Dombrow, 2005; 116, B. bicolor Dombrow, new species; 117, B. dolichiocnemis Dombrow sp.n. Scale bar 1mm." figureDoi="http://doi.org/10.5281/zenodo.4401956" httpUri="https://zenodo.org/record/4401956/files/figure.png" pageId="21" pageNumber="22">102</figureCitation>
|
||
). Externo-posterior margin of parameres of
|
||
<taxonomicName id="4C767490E009FFE0FD19D126FC09FADB" box="[746,948,1340,1366]" class="Insecta" family="Melolonthidae" genus="Beckhoplia" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="pallidibrunnea">
|
||
<emphasis id="B902D301E009FFE0FD19D126FC09FADB" box="[746,948,1340,1366]" italics="true" pageId="21" pageNumber="22">B. pallidibrunnea</emphasis>
|
||
</taxonomicName>
|
||
concave medially (
|
||
<figureCitation id="134D1396E009FFE0FB67D127FB49FADB" box="[1172,1268,1340,1366]" captionStart="FIGURES 62–97" captionStartId="28.[151,264,1743,1767]" captionTargetBox="[161,1426,187,1710]" captionTargetId="figure-25@28.[151,1436,181,1719]" captionTargetPageId="28" captionText="FIGURES 62–97. Parameres and phallobase of Beckhoplia Dombrow, 2005 species, in dorsal view (left side), in lateral view (right side), ♂♂. 62–63, B. nigra Dombrow, new species; 64–65, B. pallidibrunnea Dombrow, new species; 66–67, B. fusca Dombrow, new species; 68–69, B. setosa Dombrow, new species; 70–71, B. castanea Dombrow, new species; 72–73, B. colvillei Dombrow, 2005; 74–75, B. pulchra Dombrow, new species; 76–77, B. nigrofasciata Dombrow, new species; 78–79, B. caliginosa Dombrow, new species; 80–81, B. pumila Dombrow, new species; 82–83, B. gifbergensis Dombrow, new species; 84–85, B. julianae Dombrow, 2005; 86–87, B. elkeae Dombrow, new species; 88–89, B. suturalis Dombrow, new species; 90–91, B. nigrosetosa Dombrow, new species; 92–93, B. occidentalis Dombrow, 2005; 94–95, B. bicolor Dombrow, new species; 96–97, B. dolichiocnemis Dombrow, new species; 98–99, Anisochelus inornatus Burmeister, 1844. Scale bar 1mm." figureDoi="http://doi.org/10.5281/zenodo.4401954" httpUri="https://zenodo.org/record/4401954/files/figure.png" pageId="21" pageNumber="22">Figs. 64</figureCitation>
|
||
,
|
||
<figureCitation id="134D1396E009FFE0FAF3D127FA93FADB" box="[1280,1326,1340,1366]" captionStart="FIGURES 100–117" captionStartId="42.[151,264,1665,1689]" captionTargetBox="[162,1424,188,1625]" captionTargetId="figure-25@42.[151,1436,181,1641]" captionTargetPageId="42" captionText="FIGURES 100–117. Detailed drawings of the parameres for the key and the differential diagnosis of Beckhoplia Dombrow, 2005 species, in dorsal view, ♂♂. Arrows indicate characters of importance. 100, B. nigra Dombrow, new species; 101, B. pallidibrunnea Dombrow, new species; 102, B. fusca Dombrow, new species; 103, B. setosa Dombrow, new species; 104, B. castanea Dombrow, new species; 105, B. colvillei Dombrow, 2005; 106, B. pulchra Dombrow, new species; 107, B. nigrofasciata Dombrow, new species; 108, B. caliginosa Dombrow, new species; 109, B. pumila Dombrow, new species; 110, B. gifbergensis Dombrow, new species; 111, B. julianae Dombrow, 2005; 112, B. elkeae Dombrow, new species; 113, B. suturalis Dombrow, new species; 114, B. nigrosetosa Dombrow, new species; 115, B. occidentalis Dombrow, 2005; 116, B. bicolor Dombrow, new species; 117, B. dolichiocnemis Dombrow sp.n. Scale bar 1mm." figureDoi="http://doi.org/10.5281/zenodo.4401956" httpUri="https://zenodo.org/record/4401956/files/figure.png" pageId="21" pageNumber="22">101</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36C5C98E009FFECFF64D19FFBDEF9F8" lastPageId="25" lastPageNumber="26" pageId="21" pageNumber="22" type="description">
|
||
<paragraph id="8BC90F13E009FFE0FF64D19FFE99FA13" blockId="21.[151,1437,1412,2015]" box="[151,292,1412,1438]" pageId="21" pageNumber="22">
|
||
<emphasis id="B902D301E009FFE0FF64D19FFE99FA13" bold="true" box="[151,292,1412,1438]" pageId="21" pageNumber="22">Description</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BC90F13E009FFE0FF34D1B2FD5AFA4E" blockId="21.[151,1437,1412,2015]" box="[199,743,1448,1475]" pageId="21" pageNumber="22">
|
||
<emphasis id="B902D301E009FFE0FF34D1B2FE3EFA4E" bold="true" box="[199,387,1449,1475]" pageId="21" pageNumber="22">Measurements:</emphasis>
|
||
length
|
||
<quantity id="4C8EA2F6E009FFE0FE29D1B3FD89FA4E" box="[474,564,1448,1475]" metricMagnitude="-3" metricUnit="m" metricValue="8.2" pageId="21" pageNumber="22" unit="mm" value="8.2">8.2 mm</quantity>
|
||
, width
|
||
<quantity id="4C8EA2F6E009FFE0FD7BD1B3FD5EFA4E" box="[648,739,1448,1475]" metricMagnitude="-3" metricUnit="m" metricValue="4.4" pageId="21" pageNumber="22" unit="mm" value="4.4">4.4 mm</quantity>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BC90F13E009FFE0FF34D1D7FD53FA6B" blockId="21.[151,1437,1412,2015]" box="[199,750,1484,1510]" pageId="21" pageNumber="22">
|
||
<emphasis id="B902D301E009FFE0FF34D1D7FEE3FA6B" bold="true" box="[199,350,1484,1510]" pageId="21" pageNumber="22">Integument:</emphasis>
|
||
Black; elytra transparently brown.
|
||
</paragraph>
|
||
<paragraph id="8BC90F13E009FFE3FF34D1EBFB01FD62" blockId="21.[151,1437,1412,2015]" lastBlockId="22.[151,1437,151,2037]" lastPageId="22" lastPageNumber="23" pageId="21" pageNumber="22">
|
||
<emphasis id="B902D301E009FFE0FF34D1EBFEF7F987" bold="true" box="[199,330,1520,1546]" pageId="21" pageNumber="22">Tegument:</emphasis>
|
||
<emphasis id="B902D301E009FFE0FEA7D1EBFE0DF987" box="[340,432,1520,1546]" italics="true" pageId="21" pageNumber="22">Clypeus</emphasis>
|
||
bare; sparse setose with extremely short, erect, brown setae on the inner side of the vertically reflexed anterior and lateral margin.
|
||
<emphasis id="B902D301E009FFE0FD43D20FFD50F9A3" box="[688,749,1556,1582]" italics="true" pageId="21" pageNumber="22">Head</emphasis>
|
||
loose setose with long, erect, black setae.
|
||
<emphasis id="B902D301E009FFE0FB24D20EFADFF9A3" box="[1239,1378,1556,1582]" italics="true" pageId="21" pageNumber="22">Eye canthus</emphasis>
|
||
with a dense setose ciliary of long, black setae.
|
||
<emphasis id="B902D301E009FFE0FD71D222FD6EF9DE" box="[642,723,1593,1619]" italics="true" pageId="21" pageNumber="22">Scapus</emphasis>
|
||
loose setose with medium-long, black setae, dark-brown ventrally, sparse and short on
|
||
<emphasis id="B902D301E009FFE0FE39D247FD83F9FB" box="[458,574,1628,1654]" italics="true" pageId="21" pageNumber="22">pedicellus</emphasis>
|
||
,
|
||
<emphasis id="B902D301E009FFE0FDBDD247FD7DF9FB" box="[590,704,1628,1654]" italics="true" pageId="21" pageNumber="22">funiculum</emphasis>
|
||
and
|
||
<emphasis id="B902D301E009FFE0FD08D247FCF9F9FB" box="[763,836,1628,1654]" italics="true" pageId="21" pageNumber="22">clavus</emphasis>
|
||
sparse setose with very short and short, semi-erect, brown setae.
|
||
<emphasis id="B902D301E009FFE0FEC1D29AFE1EF917" box="[306,419,1665,1690]" italics="true" pageId="21" pageNumber="22">Pronotum</emphasis>
|
||
loose setose with medium-long and long, erect, black setae; with a loose setose ciliary of long to distinctly long, black setae on lateral margin.
|
||
<emphasis id="B902D301E009FFE0FCD9D2BFFC26F933" box="[810,923,1700,1726]" italics="true" pageId="21" pageNumber="22">Scutellum</emphasis>
|
||
loose setose with medium-long, semi-erect, black setae, shortening apically; with a longitudinal, median, bare strip.
|
||
<emphasis id="B902D301E009FFE0FC34D2D3FBB3F96F" box="[967,1038,1736,1762]" italics="true" pageId="21" pageNumber="22">Elytra</emphasis>
|
||
without pattern; sparse setose with medium-long, semi-erect, black setae; lengthening towards humerus; with a sparse juxta-sutural row of robust, medium-long, semi-erect, black setae, from anterior margin, crossing scutellar torus towards apical torus; narrowly-lanceolate, semi-erect, offwhite scales interspersed along lateral margin, denser around apical torus. Suture with a loose row of short to long, robust, semi-erect, black setae, lengthening apically.
|
||
<emphasis id="B902D301E009FFE0FB5DD343FABDF8FF" box="[1198,1280,1880,1906]" italics="true" pageId="21" pageNumber="22">Thorax</emphasis>
|
||
and
|
||
<emphasis id="B902D301E009FFE0FACBD343FA21F8FF" box="[1336,1436,1880,1906]" italics="true" pageId="21" pageNumber="22">pleurites</emphasis>
|
||
loose pilose with long, black hairs.
|
||
<emphasis id="B902D301E009FFE0FDD9D366FD0EF81B" box="[554,691,1917,1942]" italics="true" pageId="21" pageNumber="22">Ventrite I–V</emphasis>
|
||
sparse setose with short, reclinate, brown setae; long, semi-erect, black setae interspersed.
|
||
<emphasis id="B902D301E009FFE0FE4ED3BBFDECF837" box="[445,593,1952,1978]" italics="true" pageId="21" pageNumber="22">Propygidium</emphasis>
|
||
loose setose with long, semi-erect, black setae; posterior margin with a loose row of reclinate, narrowly-lanceolate, whitish scales.
|
||
<emphasis id="B902D301E009FFE0FCB1D3DFFC12F853" box="[834,943,1988,2014]" italics="true" pageId="21" pageNumber="22">Pygidium</emphasis>
|
||
sparse setose with long, erect, black setae; tip with a loose ciliary of medium-long, black setae.
|
||
<emphasis id="B902D301E00AFFE3FD00D48CFC78FF3C" box="[755,965,151,177]" italics="true" pageId="22" pageNumber="23">Prothoracic femur</emphasis>
|
||
loose setose with long, semi-erect, black setae; bare internally.
|
||
<emphasis id="B902D301E00AFFE3FE6CD4A0FDD5FF58" box="[415,616,187,213]" italics="true" pageId="22" pageNumber="23">Prothoracic tibia</emphasis>
|
||
sparse setose with short, reclinate and semi-erect, pale-brown setae, sparser ventrally; with a sparse setose longitudinal row of short and medium-long, erect, black setae dorsally; a loose setose ciliary of short, black setae on internal ridge; a setaceous ciliary of short, dark red-brown setae on anterior margin, more robust towards internal ridge; a medium-long, downwards pointing setose ciliary of brown setae along anterior margin of apical tooth.
|
||
<emphasis id="B902D301E00AFFE3FD78D550FCCCFEE8" box="[651,881,331,357]" italics="true" pageId="22" pageNumber="23">Mesothoracic femur</emphasis>
|
||
sparse setose with long, semi-erect, black setae, denser and brown ventrally; bare internally.
|
||
<emphasis id="B902D301E00AFFE3FD65D574FCD2FE04" box="[662,879,367,393]" italics="true" pageId="22" pageNumber="23">Mesothoracic tibia</emphasis>
|
||
loose setose with short, semi-erect, offwhite setae; medium-long, semi-erect, robust, black setae interspersed, sparser and finer internally. Posterior margin with a bristled ciliary of short to medium-long, black bristles; missing internally.
|
||
<emphasis id="B902D301E00AFFE3FC00D5ACFB68FE5C" box="[1011,1237,439,465]" italics="true" pageId="22" pageNumber="23">Metathoracic femur</emphasis>
|
||
loose setose with medium-long to distinctly long, semi-erect, black setae; offwhite ventrally; bare internally.
|
||
<emphasis id="B902D301E00AFFE3FB37D5C0FA21FE78" box="[1220,1436,475,501]" italics="true" pageId="22" pageNumber="23">Metathoracic tibia</emphasis>
|
||
sparse setose with medium-long to long, semi-erect, robust black setae; finer internally; very short and short, reclinate, brown setae interspersed. Posterior margin with a loose bristled outer marginal ciliary of short, black bristles, lengthening dorsally and ventrally.
|
||
<emphasis id="B902D301E00AFFE3FD6FD65CFCEDFDED" box="[668,848,583,608]" italics="true" pageId="22" pageNumber="23">Tarsomere I–IV</emphasis>
|
||
of all thoracic legs sparse setose with very-short, reclinate, brown, setae lengthening towards
|
||
<emphasis id="B902D301E00AFFE3FD66D670FC01FD09" box="[661,956,619,645]" italics="true" pageId="22" pageNumber="23">metathoracic tarsomeres;</emphasis>
|
||
with a setaceous, apical, marginal ciliary of medium-long, black setae, lengthening and more robust towards
|
||
<emphasis id="B902D301E00AFFE3FC58D694FB74FD25" box="[939,1225,655,681]" italics="true" pageId="22" pageNumber="23">metathoracic tarsomeres</emphasis>
|
||
, fine dorsally and more robust ventrally.
|
||
<emphasis id="B902D301E00AFFE3FE64D6A8FD8FFD41" box="[407,562,691,716]" italics="true" pageId="22" pageNumber="23">Pretarsomere</emphasis>
|
||
of all thoracic legs sparse setose with very short, reclinate, pale-brown setae; few, medium-long, semi-erect, black setae at apical margin, shorter and red-brown ventrally.
|
||
</paragraph>
|
||
<paragraph id="8BC90F13E00AFFE3FF34D6E3FA94FCDA" blockId="22.[151,1437,151,2037]" pageId="22" pageNumber="23">
|
||
<emphasis id="B902D301E00AFFE3FF34D6E3FEACFC9F" bold="true" box="[199,273,760,786]" pageId="22" pageNumber="23">Head:</emphasis>
|
||
Suture between head and clypeus undulated, crenated. Head dense punctate with small, shallow, roundish, not merging punctures, micropunctate within. Funicle I-II cylindrical, slightly thickened medially; funicle III asymmetric frustum-shaped; clavus elliptic, as long as pedicellus and funiculum combined.
|
||
</paragraph>
|
||
<paragraph id="8BC90F13E00AFFE3FF34D77BFB38FC32" blockId="22.[151,1437,151,2037]" pageId="22" pageNumber="23">
|
||
<emphasis id="B902D301E00AFFE3FF34D77BFEF1FCF7" bold="true" box="[199,332,864,890]" pageId="22" pageNumber="23">Pronotum:</emphasis>
|
||
Widest medially; loose punctate with small, shallow, roundish punctures, dense within medial furrow and posteriorly. Anterior corner orthogonal. Lateral margin linearly attenuate anteriorly, subparallel posteriorly. With an indistinct, complete, shallow, medial furrow. Posterior corner rounded.
|
||
</paragraph>
|
||
<paragraph id="8BC90F13E00AFFE3FF34D7D3FF44FBA4" blockId="22.[151,1437,151,2037]" pageId="22" pageNumber="23">
|
||
<emphasis id="B902D301E00AFFE3FF34D7D3FEF7FC6F" bold="true" box="[199,330,968,994]" pageId="22" pageNumber="23">Scutellum:</emphasis>
|
||
Parabola-shaped; as broad as long; dense punctate with small, shallow, roundish punctures, micropunctate within; with a complete, median, longitudinal, narrow, smooth stripe. Lateral margin slightly convex. Tip rounded.
|
||
</paragraph>
|
||
<paragraph id="8BC90F13E00AFFE3FF34D028FAD0FA8C" blockId="22.[151,1437,151,2037]" pageId="22" pageNumber="23">
|
||
<emphasis id="B902D301E00AFFE3FF34D028FEA3FBC0" bold="true" box="[199,286,1075,1101]" pageId="22" pageNumber="23">Elytra:</emphasis>
|
||
Ovate; as broad as long; widest medially; protruding propygidium. Lateral margin slightly convex attenuate apically. Sculpture indistinct; slightly pulvinate laterally; humerus indistinct; with a roundish torus beside scutellum; lateral trench indistinct; broadly, slightly, longitudinally depressed along suture. Loose punctate with extremely small, shallow, roundish punctures, micropunctures in-between; a sparse juxta-sutural row of large, roundish tubercles from anterior margin, crossing scutellar torus towards apical torus; a sparse supra-humeral row of small, roundish tubercles from anterior margin between humerus and scutellar torus towards apical torus.
|
||
</paragraph>
|
||
<paragraph id="8BC90F13E00AFFE3FF34D111FAEAFAE0" blockId="22.[151,1437,151,2037]" pageId="22" pageNumber="23">
|
||
<emphasis id="B902D301E00AFFE3FF34D111FEFAFAA9" bold="true" box="[199,327,1290,1316]" pageId="22" pageNumber="23">Abdomen:</emphasis>
|
||
Ventrite I extremely shortened medially; ventrite V broadest, not constricted ventrally. Propygidium with a slightly projected stoma at posterior corner. Pygidium distinctly pulvinate longitudinally and laterally; upper margin slightly convex, linear medially; lateral margin slightly convex attenuate apically; tip linear.
|
||
</paragraph>
|
||
<paragraph id="8BC90F13E00AFFE3FF34D16DFCF1F9E4" blockId="22.[151,1437,151,2037]" pageId="22" pageNumber="23">
|
||
<emphasis id="B902D301E00AFFE3FF34D16DFE30FA1C" bold="true" box="[199,397,1398,1425]" pageId="22" pageNumber="23">Prothoracic leg:</emphasis>
|
||
Femur and tibia robust. Tibia smooth ventrally. Median tooth elongate. Basal tooth very small. Interspace between anterior and median tooth double in width than between median and basal tooth. Shape of interspace between anterior and median tooth u-shaped and between median and basal tooth concave. Tarsomere I–IV combined extremely short; lengthening from tarsomere I–IV; slightly more than
|
||
<superScript id="7C03A25BE00AFFE3FB46D1F9FB00FA7D" attach="right" box="[1205,1213,1506,1520]" fontSize="6" pageId="22" pageNumber="23">2</superScript>
|
||
/
|
||
<quantity id="4C8EA2F6E00AFFE3FB36D1EEFB56FA70" box="[1221,1259,1507,1539]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="22" pageNumber="23" unit="in" value="5.0">
|
||
<subScript id="17F20D56E00AFFE3FB36D1EEFB70F98E" attach="left" box="[1221,1229,1525,1539]" fontSize="6" pageId="22" pageNumber="23">5</subScript>
|
||
in
|
||
</quantity>
|
||
length of tibia. Pretarsomere distinctly thickened; with a small, triangular-shaped, median tooth ventrally; slightly less than
|
||
<superScript id="7C03A25BE00AFFE3FA77D21DFA31F999" attach="right" box="[1412,1420,1542,1556]" fontSize="6" pageId="22" pageNumber="23">1</superScript>
|
||
/
|
||
<subScript id="17F20D56E00AFFE3FA67D202FA21F9AA" attach="none" box="[1428,1436,1561,1575]" fontSize="6" pageId="22" pageNumber="23">5</subScript>
|
||
more in length of tarsomere I–IV combined. Internal claw slightly less than
|
||
<superScript id="7C03A25BE00AFFE3FBFAD231FBACF9B5" attach="right" box="[1033,1041,1578,1592]" fontSize="6" pageId="22" pageNumber="23">1</superScript>
|
||
/
|
||
<subScript id="17F20D56E00AFFE3FBEAD226FB97F9C6" attach="left" box="[1049,1066,1597,1611]" fontSize="6" pageId="22" pageNumber="23">10</subScript>
|
||
more in length of pretarsomere. External claw slightly less than
|
||
<superScript id="7C03A25BE00AFFE3FDF6D255FDB0F9D1" attach="right" box="[517,525,1614,1628]" fontSize="6" pageId="22" pageNumber="23">4</superScript>
|
||
/
|
||
<quantity id="4C8EA2F6E00AFFE3FDE5D27AFD86F9E4" box="[534,571,1615,1647]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="22" pageNumber="23" unit="in" value="5.0">
|
||
<subScript id="17F20D56E00AFFE3FDE5D27AFDA3F9E2" attach="left" box="[534,542,1633,1647]" fontSize="6" pageId="22" pageNumber="23">5</subScript>
|
||
in
|
||
</quantity>
|
||
length of internal claw.
|
||
</paragraph>
|
||
<paragraph id="8BC90F13E00AFFE3FF34D269FB3CF974" blockId="22.[151,1437,151,2037]" pageId="22" pageNumber="23">
|
||
<emphasis id="B902D301E00AFFE3FF34D269FE1CF900" bold="true" box="[199,417,1650,1677]" pageId="22" pageNumber="23">Mesothoracic leg:</emphasis>
|
||
Tibia
|
||
<superScript id="7C03A25BE00AFFE3FE18D269FE4EF90D" attach="right" box="[491,499,1650,1664]" fontSize="6" pageId="22" pageNumber="23">9</superScript>
|
||
/
|
||
<quantity id="4C8EA2F6E00AFFE3FE08D29EFD97F900" box="[507,554,1651,1683]" metricMagnitude="-1" metricUnit="m" metricValue="2.54" pageId="22" pageNumber="23" unit="in" value="10.0">
|
||
<subScript id="17F20D56E00AFFE3FE08D29EFDB6F91E" attach="left" box="[507,523,1669,1683]" fontSize="6" pageId="22" pageNumber="23">10</subScript>
|
||
in
|
||
</quantity>
|
||
length femur. Tarsomere I–IV combined extremely short; slightly more than
|
||
<superScript id="7C03A25BE00AFFE3FF64D28DFF22F929" attach="none" box="[151,159,1686,1700]" fontSize="6" pageId="22" pageNumber="23">3</superScript>
|
||
/
|
||
<quantity id="4C8EA2F6E00AFFE3FF5BD2B2FF6BF93C" box="[168,214,1687,1719]" metricMagnitude="-1" metricUnit="m" metricValue="2.54" pageId="22" pageNumber="23" unit="in" value="10.0">
|
||
<subScript id="17F20D56E00AFFE3FF5BD2B2FF05F93A" attach="left" box="[168,184,1705,1719]" fontSize="6" pageId="22" pageNumber="23">10</subScript>
|
||
in
|
||
</quantity>
|
||
length of tibia. Pretarsomere with a shortened smooth ridge ventrally, followed by a roundish cutting apically; slightly more than
|
||
<superScript id="7C03A25BE00AFFE3FE41D2A1FE07F945" attach="right" box="[434,442,1722,1736]" fontSize="6" pageId="22" pageNumber="23">1</superScript>
|
||
/
|
||
<subScript id="17F20D56E00AFFE3FE31D2D6FE6FF956" attach="left" box="[450,466,1741,1755]" fontSize="6" pageId="22" pageNumber="23">10</subScript>
|
||
more in length of tarsomere I–IV combined. External claw slightly less than
|
||
<superScript id="7C03A25BE00AFFE3FAC8D2A1FAFEF945" attach="right" box="[1339,1347,1722,1736]" fontSize="6" pageId="22" pageNumber="23">1</superScript>
|
||
/
|
||
<subScript id="17F20D56E00AFFE3FAB8D2D6FAE6F956" attach="left" box="[1355,1371,1741,1755]" fontSize="6" pageId="22" pageNumber="23">10</subScript>
|
||
more in length of pretarsomere. Internal claw slightly less than
|
||
<superScript id="7C03A25BE00AFFE3FCDED2C5FC88F961" attach="right" box="[813,821,1758,1772]" fontSize="6" pageId="22" pageNumber="23">9</superScript>
|
||
/
|
||
<quantity id="4C8EA2F6E00AFFE3FCCDD2EAFCD1F974" box="[830,876,1759,1791]" metricMagnitude="-1" metricUnit="m" metricValue="2.54" pageId="22" pageNumber="23" unit="in" value="10.0">
|
||
<subScript id="17F20D56E00AFFE3FCCDD2EAFCF3F972" attach="left" box="[830,846,1777,1791]" fontSize="6" pageId="22" pageNumber="23">10</subScript>
|
||
in
|
||
</quantity>
|
||
length of external claw.
|
||
</paragraph>
|
||
<paragraph id="8BC90F13E00AFFE3FF34D319FCE5F804" blockId="22.[151,1437,151,2037]" pageId="22" pageNumber="23">
|
||
<emphasis id="B902D301E00AFFE3FF34D319FE22F890" bold="true" box="[199,415,1794,1821]" pageId="22" pageNumber="23">Metathoracic leg:</emphasis>
|
||
Tibia slightly less than
|
||
<superScript id="7C03A25BE00AFFE3FD43D319FD05F89D" attach="right" box="[688,696,1794,1808]" fontSize="6" pageId="22" pageNumber="23">9</superScript>
|
||
/
|
||
<quantity id="4C8EA2F6E00AFFE3FD32D30EFD52F890" box="[705,751,1795,1827]" metricMagnitude="-1" metricUnit="m" metricValue="2.54" pageId="22" pageNumber="23" unit="in" value="10.0">
|
||
<subScript id="17F20D56E00AFFE3FD32D30EFD6CF8AE" attach="left" box="[705,721,1813,1827]" fontSize="6" pageId="22" pageNumber="23">10</subScript>
|
||
in
|
||
</quantity>
|
||
length of femur. Tarsomere I–IV combined distinctly short; slightly more than
|
||
<superScript id="7C03A25BE00AFFE3FE86D33CFEC0F8B8" attach="right" box="[373,381,1831,1845]" fontSize="6" pageId="22" pageNumber="23">7</superScript>
|
||
/
|
||
<quantity id="4C8EA2F6E00AFFE3FE75D322FE0BF8CC" box="[390,438,1831,1863]" metricMagnitude="-1" metricUnit="m" metricValue="2.54" pageId="22" pageNumber="23" unit="in" value="10.0">
|
||
<subScript id="17F20D56E00AFFE3FE75D322FE2BF8CA" attach="left" box="[390,406,1849,1863]" fontSize="6" pageId="22" pageNumber="23">10</subScript>
|
||
in
|
||
</quantity>
|
||
length of tibia. Pretarsomere slender; with an undulate ridge ventrally; slightly more than
|
||
<superScript id="7C03A25BE00AFFE3FF3CD351FF6AF8D5" attach="right" box="[207,215,1866,1880]" fontSize="6" pageId="22" pageNumber="23">2</superScript>
|
||
/
|
||
<quantity id="4C8EA2F6E00AFFE3FF2CD346FEB9F8E8" box="[223,260,1867,1899]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="22" pageNumber="23" unit="in" value="5.0">
|
||
<subScript id="17F20D56E00AFFE3FF2CD346FF5AF8E6" attach="left" box="[223,231,1885,1899]" fontSize="6" pageId="22" pageNumber="23">5</subScript>
|
||
in
|
||
</quantity>
|
||
length of tarsomere I–IV combined. External claw slightly less than
|
||
<superScript id="7C03A25BE00AFFE3FBFCD351FBAAF8D5" attach="right" box="[1039,1047,1866,1880]" fontSize="6" pageId="22" pageNumber="23">3</superScript>
|
||
/
|
||
<subScript id="17F20D56E00AFFE3FBD3D346FB8DF8E6" attach="left" box="[1056,1072,1885,1899]" fontSize="6" pageId="22" pageNumber="23">10</subScript>
|
||
more in length of pretarsomere. Internal claw slightly more than
|
||
<superScript id="7C03A25BE00AFFE3FDFED375FDA8F8F1" attach="right" box="[525,533,1902,1916]" fontSize="6" pageId="22" pageNumber="23">4</superScript>
|
||
/
|
||
<quantity id="4C8EA2F6E00AFFE3FDEED39AFDFEF804" box="[541,579,1903,1935]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="22" pageNumber="23" unit="in" value="5.0">
|
||
<subScript id="17F20D56E00AFFE3FDEED39AFD98F802" attach="left" box="[541,549,1921,1935]" fontSize="6" pageId="22" pageNumber="23">5</subScript>
|
||
in
|
||
</quantity>
|
||
length of external claw.
|
||
</paragraph>
|
||
<paragraph id="8BC90F13E00AFFE3FF34D388FD8BF820" blockId="22.[151,1437,151,2037]" box="[199,566,1939,1965]" pageId="22" pageNumber="23">
|
||
<emphasis id="B902D301E00AFFE3FF34D388FEE9F820" bold="true" box="[199,340,1939,1965]" pageId="22" pageNumber="23">Parameres:</emphasis>
|
||
(
|
||
<figureCitation id="134D1396E00AFFE3FE97D388FE52F820" box="[356,495,1939,1965]" captionStart="FIGURES 62–97" captionStartId="28.[151,264,1743,1767]" captionTargetBox="[161,1426,187,1710]" captionTargetId="figure-25@28.[151,1436,181,1719]" captionTargetPageId="28" captionText="FIGURES 62–97. Parameres and phallobase of Beckhoplia Dombrow, 2005 species, in dorsal view (left side), in lateral view (right side), ♂♂. 62–63, B. nigra Dombrow, new species; 64–65, B. pallidibrunnea Dombrow, new species; 66–67, B. fusca Dombrow, new species; 68–69, B. setosa Dombrow, new species; 70–71, B. castanea Dombrow, new species; 72–73, B. colvillei Dombrow, 2005; 74–75, B. pulchra Dombrow, new species; 76–77, B. nigrofasciata Dombrow, new species; 78–79, B. caliginosa Dombrow, new species; 80–81, B. pumila Dombrow, new species; 82–83, B. gifbergensis Dombrow, new species; 84–85, B. julianae Dombrow, 2005; 86–87, B. elkeae Dombrow, new species; 88–89, B. suturalis Dombrow, new species; 90–91, B. nigrosetosa Dombrow, new species; 92–93, B. occidentalis Dombrow, 2005; 94–95, B. bicolor Dombrow, new species; 96–97, B. dolichiocnemis Dombrow, new species; 98–99, Anisochelus inornatus Burmeister, 1844. Scale bar 1mm." figureDoi="http://doi.org/10.5281/zenodo.4401954" httpUri="https://zenodo.org/record/4401954/files/figure.png" pageId="22" pageNumber="23">Figs. 66–67</figureCitation>
|
||
,
|
||
<figureCitation id="134D1396E00AFFE3FE08D388FD97F820" box="[507,554,1939,1965]" captionStart="FIGURES 100–117" captionStartId="42.[151,264,1665,1689]" captionTargetBox="[162,1424,188,1625]" captionTargetId="figure-25@42.[151,1436,181,1641]" captionTargetPageId="42" captionText="FIGURES 100–117. Detailed drawings of the parameres for the key and the differential diagnosis of Beckhoplia Dombrow, 2005 species, in dorsal view, ♂♂. Arrows indicate characters of importance. 100, B. nigra Dombrow, new species; 101, B. pallidibrunnea Dombrow, new species; 102, B. fusca Dombrow, new species; 103, B. setosa Dombrow, new species; 104, B. castanea Dombrow, new species; 105, B. colvillei Dombrow, 2005; 106, B. pulchra Dombrow, new species; 107, B. nigrofasciata Dombrow, new species; 108, B. caliginosa Dombrow, new species; 109, B. pumila Dombrow, new species; 110, B. gifbergensis Dombrow, new species; 111, B. julianae Dombrow, 2005; 112, B. elkeae Dombrow, new species; 113, B. suturalis Dombrow, new species; 114, B. nigrosetosa Dombrow, new species; 115, B. occidentalis Dombrow, 2005; 116, B. bicolor Dombrow, new species; 117, B. dolichiocnemis Dombrow sp.n. Scale bar 1mm." figureDoi="http://doi.org/10.5281/zenodo.4401956" httpUri="https://zenodo.org/record/4401956/files/figure.png" pageId="22" pageNumber="23">102</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BC90F13E00AFFE3FF34D3A3FC38F85C" blockId="22.[151,1437,151,2037]" box="[199,901,1974,2003]" pageId="22" pageNumber="23">
|
||
<emphasis id="B902D301E00AFFE3FF34D3A3FE01F85C" bold="true" box="[199,444,1975,2003]" pageId="22" pageNumber="23">
|
||
<specimenCount id="9D70C49AE00AFFE3FF34D3A3FF4FF85E" box="[199,242,1976,2003]" pageId="22" pageNumber="23" type="female">♀♀</specimenCount>
|
||
. Measurements:
|
||
</emphasis>
|
||
length
|
||
<quantity id="4C8EA2F6E00AFFE3FDE0D3ACFD1DF85C" box="[531,672,1975,2001]" metricMagnitude="-3" metricUnit="m" metricValue="8.4" metricValueMax="8.9" metricValueMin="7.9" pageId="22" pageNumber="23" unit="mm" value="8.4" valueMax="8.9" valueMin="7.9">7.9–8.9 mm</quantity>
|
||
; width
|
||
<quantity id="4C8EA2F6E00AFFE3FD07D3ADFC3CF85C" box="[756,897,1974,2001]" metricMagnitude="-3" metricUnit="m" metricValue="4.8" metricValueMax="5.2" metricValueMin="4.4" pageId="22" pageNumber="23" unit="mm" value="4.8" valueMax="5.2" valueMin="4.4">4.4–5.2 mm</quantity>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BC90F13E00AFFE3FF34D3C0FD25F878" blockId="22.[151,1437,151,2037]" box="[199,664,2011,2037]" pageId="22" pageNumber="23">
|
||
<emphasis id="B902D301E00AFFE3FF34D3C0FEE3F878" bold="true" box="[199,350,2011,2037]" pageId="22" pageNumber="23">Integument:</emphasis>
|
||
Thoracic tarsi dark-brown.
|
||
</paragraph>
|
||
<caption id="DF095F9BE00BFFE2FF64D589FB3CFD88" ID-DOI="http://doi.org/10.5281/zenodo.4455243" ID-Zenodo-Dep="4455243" httpUri="https://zenodo.org/record/4455243/files/figure.png" pageId="23" pageNumber="24" startId="23.[151,275,402,428]" targetBox="[304,1284,185,355]" targetPageId="23">
|
||
<paragraph id="8BC90F13E00BFFE2FF64D589FB3CFD88" blockId="23.[151,1436,402,517]" pageId="23" pageNumber="24">
|
||
<emphasis id="B902D301E00BFFE2FF64D589FEDAFE20" bold="true" box="[151,359,402,429]" pageId="23" pageNumber="24">FIGURES 36–38.</emphasis>
|
||
General drawings of the pygidium of
|
||
<taxonomicName id="4C767490E00BFFE2FCFED588FBF8FE23" authority="Dombrow, 2005" authorityName="Dombrow" authorityYear="2005" box="[781,1093,403,430]" class="Insecta" family="Melolonthidae" genus="Beckhoplia" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B902D301E00BFFE2FCFED588FC36FE20" box="[781,907,403,429]" italics="true" pageId="23" pageNumber="24">Beckhoplia</emphasis>
|
||
<bibRefCitation id="EFE772E2E00BFFE2FC62D588FBF8FE23" author="Dombrow, H." box="[913,1093,403,430]" pageId="23" pageNumber="24" pagination="125 - 130" refId="ref61191" refString="Dombrow, H. (2005) Beckhoplia a new genus of Hopliini from the Northern Cape of the Republic of South Africa. (Coleoptera: Scarabaeidae: Melolonthinae: Hopliini). Coleoptera, 9, 125 - 130." type="journal article" year="2005">Dombrow, 2005</bibRefCitation>
|
||
</taxonomicName>
|
||
species, in posterior view
|
||
<specimenCount id="9D70C49AE00BFFE2FA98D588FA25FE20" box="[1387,1432,403,429]" pageId="23" pageNumber="24" type="male">♂♂</specimenCount>
|
||
. 36, upper margin moderately convex, linear medially, lateral margin subparallel in upper fifth, linearly attenuate apically; 37, upper margin slightly convex, lateral margin linearly attenuate apically; 38, upper margin moderately convex, linear medially, lateral margin slightly convex attenuate apically. Scale bar
|
||
<quantity id="4C8EA2F6E00BFFE2FBB3D5F0FB3CFD88" box="[1088,1153,491,517]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="23" pageNumber="24" unit="mm" value="1.0">1 mm</quantity>
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF095F9BE00BFFE2FF64D6F6FD96FCAA" ID-DOI="http://doi.org/10.5281/zenodo.4401942" ID-Zenodo-Dep="4401942" httpUri="https://zenodo.org/record/4401942/files/figure.png" pageId="23" pageNumber="24" startId="23.[151,264,749,773]" targetBox="[432,1159,536,689]" targetPageId="23">
|
||
<paragraph id="8BC90F13E00BFFE2FF64D6F6FD96FCAA" blockId="23.[151,1436,749,808]" pageId="23" pageNumber="24">
|
||
<emphasis id="B902D301E00BFFE2FF64D6F6FEEEFC88" bold="true" box="[151,339,749,773]" pageId="23" pageNumber="24">FIGURES 39–40.</emphasis>
|
||
General drawings of the tip of
|
||
<taxonomicName id="4C767490E00BFFE2FD79D6F6FD56FC88" box="[650,747,749,773]" family="Trichomycteridae" genus="Pygidium" higherTaxonomySource="CoL" kingdom="Animalia" order="Siluriformes" pageId="23" pageNumber="70" phylum="Chordata" rank="genus">pygidium</taxonomicName>
|
||
of
|
||
<taxonomicName id="4C767490E00BFFE2FCFFD6F6FB9AFC88" authority="Dombrow, 2005" authorityName="Dombrow" authorityYear="2005" box="[780,1063,749,773]" class="Insecta" family="Melolonthidae" genus="Beckhoplia" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B902D301E00BFFE2FCFFD6F6FCC3FC88" box="[780,894,749,773]" italics="true" pageId="23" pageNumber="24">Beckhoplia</emphasis>
|
||
<bibRefCitation id="EFE772E2E00BFFE2FC70D6F6FB9AFC88" author="Dombrow, H." box="[899,1063,749,773]" pageId="23" pageNumber="24" pagination="125 - 130" refId="ref61191" refString="Dombrow, H. (2005) Beckhoplia a new genus of Hopliini from the Northern Cape of the Republic of South Africa. (Coleoptera: Scarabaeidae: Melolonthinae: Hopliini). Coleoptera, 9, 125 - 130." type="journal article" year="2005">Dombrow, 2005</bibRefCitation>
|
||
</taxonomicName>
|
||
species in posterior view ♂♂. 39, tip linear; 40, tip rounded. Scale bar 1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF095F9BE00BFFE2FF64D116FD77FA58" ID-DOI="http://doi.org/10.5281/zenodo.4401944" ID-Zenodo-Dep="4401944" httpUri="https://zenodo.org/record/4401944/files/figure.png" pageId="23" pageNumber="24" startId="23.[151,264,1293,1317]" targetBox="[208,1379,833,1240]" targetPageId="23">
|
||
<paragraph id="8BC90F13E00BFFE2FF64D116FD77FA58" blockId="23.[151,1437,1293,1493]" pageId="23" pageNumber="24">
|
||
<emphasis id="B902D301E00BFFE2FF64D116FEE8FAA8" bold="true" box="[151,341,1293,1317]" pageId="23" pageNumber="24">FIGURES 41–46.</emphasis>
|
||
General drawing of the prothoracic tibia of
|
||
<taxonomicName id="4C767490E00BFFE2FCEDD116FB80FAA8" authority="Dombrow, 2005" authorityName="Dombrow" authorityYear="2005" box="[798,1085,1293,1317]" class="Insecta" family="Melolonthidae" genus="Beckhoplia" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B902D301E00BFFE2FCEDD116FC2DFAA8" box="[798,912,1293,1317]" italics="true" pageId="23" pageNumber="24">Beckhoplia</emphasis>
|
||
<bibRefCitation id="EFE772E2E00BFFE2FC64D116FB80FAA8" author="Dombrow, H." box="[919,1085,1293,1317]" pageId="23" pageNumber="24" pagination="125 - 130" refId="ref61191" refString="Dombrow, H. (2005) Beckhoplia a new genus of Hopliini from the Northern Cape of the Republic of South Africa. (Coleoptera: Scarabaeidae: Melolonthinae: Hopliini). Coleoptera, 9, 125 - 130." type="journal article" year="2005">Dombrow, 2005</bibRefCitation>
|
||
</taxonomicName>
|
||
species, in dorsal view, otherwise indicated, ♂♂. 41, slender, basal tooth very small, interspace between teeth almost equal, shape of interspace concave, with a dense longitudinal row of deep, elongate roundish punctures; 42, basal tooth rudimentary, interspace between teeth almost equal, interspace between anterior and median tooth concave and between median and basal tooth linear; 43, interspace between anterior and median tooth u-shaped and between median and basal tooth concave; 44, with a medio-ventral ridge; 45, with a medio-ventral ridge, cross-section: a, internal ridge; b, external ridge; c, medio-ventral ridge; 46, smooth ventrally, cross-section: a, internal ridge; b, external ridge. Scale bar 1mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF095F9BE00BFFE2FF64D399FD29F851" ID-DOI="http://doi.org/10.5281/zenodo.4401946" ID-Zenodo-Dep="4401946" httpUri="https://zenodo.org/record/4401946/files/figure.png" pageId="23" pageNumber="24" startId="23.[151,264,1922,1946]" targetBox="[404,1181,1515,1867]" targetPageId="23">
|
||
<paragraph id="8BC90F13E00BFFE2FF64D399FD29F851" blockId="23.[151,1437,1922,2012]" pageId="23" pageNumber="24">
|
||
<emphasis id="B902D301E00BFFE2FF64D399FEE8F816" bold="true" box="[151,341,1922,1947]" pageId="23" pageNumber="24">FIGURES 47–49.</emphasis>
|
||
General drawing of the prothoracic tarsomeres of
|
||
<taxonomicName id="4C767490E00BFFE2FCAAD398FBCAF816" authority="Dombrow, 2005" authorityName="Dombrow" authorityYear="2005" box="[857,1143,1923,1947]" class="Insecta" family="Melolonthidae" genus="Beckhoplia" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B902D301E00BFFE2FCAAD398FC76F816" box="[857,971,1923,1947]" italics="true" pageId="23" pageNumber="24">Beckhoplia</emphasis>
|
||
<bibRefCitation id="EFE772E2E00BFFE2FC21D398FBCAF816" author="Dombrow, H." box="[978,1143,1923,1947]" pageId="23" pageNumber="24" pagination="125 - 130" refId="ref61191" refString="Dombrow, H. (2005) Beckhoplia a new genus of Hopliini from the Northern Cape of the Republic of South Africa. (Coleoptera: Scarabaeidae: Melolonthinae: Hopliini). Coleoptera, 9, 125 - 130." type="journal article" year="2005">Dombrow, 2005</bibRefCitation>
|
||
</taxonomicName>
|
||
species, in dorsal view, ♂♂. Scale bar 1 mm, otherwise indicated. 47, tarsomere I–IV equal in length; 48, lengthening from tarsomere I–IV; 49, tarsomere II–IV with a ventral extension, scale bar 0.5 mm..
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF095F9BE004FFEDFF64D6BDFB5CFCA7" ID-DOI="http://doi.org/10.5281/zenodo.4401948" ID-Zenodo-Dep="4401948" httpUri="https://zenodo.org/record/4401948/files/figure.png" pageId="24" pageNumber="25" startId="24.[151,264,678,702]" targetBox="[212,1375,186,616]" targetPageId="24">
|
||
<paragraph id="8BC90F13E004FFEDFF64D6BDFB5CFCA7" blockId="24.[151,1436,678,810]" pageId="24" pageNumber="25">
|
||
<emphasis id="B902D301E004FFEDFF64D6BDFEE8FD33" bold="true" box="[151,341,678,702]" pageId="24" pageNumber="25">FIGURES 50–54.</emphasis>
|
||
General drawing of the types of prothoracic pretarsomere of
|
||
<taxonomicName id="4C767490E004FFEDFC34D6BDFB5BFD33" authority="Dombrow, 2005" authorityName="Dombrow" authorityYear="2005" box="[967,1254,678,702]" class="Insecta" family="Melolonthidae" genus="Beckhoplia" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B902D301E004FFEDFC34D6BDFB84FD33" box="[967,1081,678,702]" italics="true" pageId="24" pageNumber="25">Beckhoplia</emphasis>
|
||
<bibRefCitation id="EFE772E2E004FFEDFBB3D6BDFB5BFD33" author="Dombrow, H." box="[1088,1254,678,702]" pageId="24" pageNumber="25" pagination="125 - 130" refId="ref61191" refString="Dombrow, H. (2005) Beckhoplia a new genus of Hopliini from the Northern Cape of the Republic of South Africa. (Coleoptera: Scarabaeidae: Melolonthinae: Hopliini). Coleoptera, 9, 125 - 130." type="journal article" year="2005">Dombrow, 2005</bibRefCitation>
|
||
</taxonomicName>
|
||
species, in lateral view, with roundish cutting ventrally, ♂♂. 50, distinctly thickened, smooth ventrally; 51, slightly thickened, with a shortened smooth ridge ventrally; 52, distinctly thickened, with a median tooth ventrally; 53, slightly thickened, with two median teeth consecutive ventrally; 54, distinctly thickened, with a shortened granulate ridge ventrally. Scale bar 0.5 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF095F9BE004FFEDFF64D170FDE1FA62" ID-DOI="http://doi.org/10.5281/zenodo.4401950" ID-Zenodo-Dep="4401950" httpUri="https://zenodo.org/record/4401950/files/figure.png" pageId="24" pageNumber="25" startId="24.[151,264,1387,1411]" targetBox="[155,1432,874,1323]" targetPageId="24">
|
||
<paragraph id="8BC90F13E004FFEDFF64D170FDE1FA62" blockId="24.[151,1436,1387,1519]" pageId="24" pageNumber="25">
|
||
<emphasis id="B902D301E004FFEDFF64D170FEE8FA0E" bold="true" box="[151,341,1387,1411]" pageId="24" pageNumber="25">FIGURES 55–58.</emphasis>
|
||
General drawing of the types of mesothoracic pretarsomere of
|
||
<taxonomicName id="4C767490E004FFEDFC11D170FABCFA0E" authority="Dombrow, 2005" authorityName="Dombrow" authorityYear="2005" box="[994,1281,1387,1411]" class="Insecta" family="Melolonthidae" genus="Beckhoplia" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B902D301E004FFEDFC11D170FBE9FA0E" box="[994,1108,1387,1411]" italics="true" pageId="24" pageNumber="25">Beckhoplia</emphasis>
|
||
<bibRefCitation id="EFE772E2E004FFEDFBA8D170FABCFA0E" author="Dombrow, H." box="[1115,1281,1387,1411]" pageId="24" pageNumber="25" pagination="125 - 130" refId="ref61191" refString="Dombrow, H. (2005) Beckhoplia a new genus of Hopliini from the Northern Cape of the Republic of South Africa. (Coleoptera: Scarabaeidae: Melolonthinae: Hopliini). Coleoptera, 9, 125 - 130." type="journal article" year="2005">Dombrow, 2005</bibRefCitation>
|
||
</taxonomicName>
|
||
species, in lateral view showing the distinctly thickened, roundish cutting missing ventrally, ♂♂, otherwise indicated. 55, with a shortened smooth ridge, with roundish cutting; 56, with a shortened granulate ridge; 57, slender, with a full undulate ridge, ♀♀; 58, with a shortened smooth ridge. Scale bar 0.5 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF095F9BE004FFEDFF64D35BFE06F812" ID-DOI="http://doi.org/10.5281/zenodo.4401952" ID-Zenodo-Dep="4401952" httpUri="https://zenodo.org/record/4401952/files/figure.png" pageId="24" pageNumber="25" startId="24.[151,264,1856,1880]" targetBox="[155,1430,1585,1791]" targetPageId="24">
|
||
<paragraph id="8BC90F13E004FFEDFF64D35BFE06F812" blockId="24.[151,1436,1856,1952]" pageId="24" pageNumber="25">
|
||
<emphasis id="B902D301E004FFEDFF64D35BFEE9F8D5" bold="true" box="[151,340,1856,1880]" pageId="24" pageNumber="25">FIGURES 59–61.</emphasis>
|
||
General drawing of the types of metathoracic pretarsomere of
|
||
<taxonomicName id="4C767490E004FFEDFC3FD35BFB55F8D5" authority="Dombrow, 2005" authorityName="Dombrow" authorityYear="2005" box="[972,1256,1856,1880]" class="Insecta" family="Melolonthidae" genus="Beckhoplia" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B902D301E004FFEDFC3FD35BFB83F8D5" box="[972,1086,1856,1880]" italics="true" pageId="24" pageNumber="25">Beckhoplia</emphasis>
|
||
<bibRefCitation id="EFE772E2E004FFEDFBB7D35BFB55F8D5" author="Dombrow, H." box="[1092,1256,1856,1880]" pageId="24" pageNumber="25" pagination="125 - 130" refId="ref61191" refString="Dombrow, H. (2005) Beckhoplia a new genus of Hopliini from the Northern Cape of the Republic of South Africa. (Coleoptera: Scarabaeidae: Melolonthinae: Hopliini). Coleoptera, 9, 125 - 130." type="journal article" year="2005">Dombrow, 2005</bibRefCitation>
|
||
</taxonomicName>
|
||
species, in lateral view, ♂♂. 59, slender, with a smooth ridge; 60, slightly thickened, with a granulate ridge; 61, slightly thickened, with an undulate ridge. Scale bar 0.5 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BC90F13E005FFECFF34D48CFC0FFD7C" blockId="25.[151,1437,151,1659]" pageId="25" pageNumber="26">
|
||
<emphasis id="B902D301E005FFECFF34D48CFEF7FF3C" bold="true" box="[199,330,151,177]" pageId="25" pageNumber="26">Tegument:</emphasis>
|
||
<emphasis id="B902D301E005FFECFEA6D48CFE0CFF3C" box="[341,433,151,177]" italics="true" pageId="25" pageNumber="26">Clypeus</emphasis>
|
||
loose setose with short, erect, pale-brown setae, medium-long and brown on
|
||
<emphasis id="B902D301E005FFECFAADD48CFA28FF3C" box="[1374,1429,151,177]" italics="true" pageId="25" pageNumber="26">head</emphasis>
|
||
.
|
||
<emphasis id="B902D301E005FFECFF64D4A0FE99FF58" box="[151,292,187,213]" italics="true" pageId="25" pageNumber="26">Eye canthus</emphasis>
|
||
with a dense setose ciliary of medium-long, brown setae. Setae on
|
||
<taxonomicName id="4C767490E005FFECFBCCD4A0FB30FF59" authorityName="Norman" authorityYear="1875" box="[1087,1165,187,212]" genus="Scapus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="25" pageNumber="70" phylum="Cnidaria" rank="genus">
|
||
<emphasis id="B902D301E005FFECFBCCD4A0FB30FF59" box="[1087,1165,187,212]" italics="true" pageId="25" pageNumber="26">scapus</emphasis>
|
||
</taxonomicName>
|
||
brown and pale-brown ventrally; sparse and short on
|
||
<emphasis id="B902D301E005FFECFE0BD4C4FDD1FF74" box="[504,620,223,249]" italics="true" pageId="25" pageNumber="26">pedicellus</emphasis>
|
||
. Setae on
|
||
<emphasis id="B902D301E005FFECFD1AD4C4FCEAFF75" box="[745,855,223,248]" italics="true" pageId="25" pageNumber="26">pronotum</emphasis>
|
||
including ciliary brown. Setae on
|
||
<taxonomicName id="4C767490E005FFECFB1FD4C4FAE7FF74" box="[1260,1370,223,249]" class="Trilobita" family="Styginidae" genus="Scutellum" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Corynexochida" pageId="25" pageNumber="70" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B902D301E005FFECFB1FD4C4FAE7FF74" box="[1260,1370,223,249]" italics="true" pageId="25" pageNumber="26">scutellum</emphasis>
|
||
</taxonomicName>
|
||
short and brown, with a shortened bare stripe in anterior half.
|
||
<emphasis id="B902D301E005FFECFCC2D518FCC5FE90" box="[817,888,259,285]" italics="true" pageId="25" pageNumber="26">Elytra</emphasis>
|
||
without pattern; loose setose with short, reclinate, pale-brown setae; long, semi-erect, brown setae interspersed, lengthening towards humerus; with a sparse juxta-sutural row of robust, medium-long, semi-erect, brown setae, from anterior margin, crossing scutellar torus towards apical torus. Suture with a loose row of short to long, robust, semi-erect, brown setae, lengthening apically. Hairs on
|
||
<emphasis id="B902D301E005FFECFEB6D588FE2DFE20" box="[325,400,403,429]" italics="true" pageId="25" pageNumber="26">thorax</emphasis>
|
||
and
|
||
<emphasis id="B902D301E005FFECFE3BD588FD91FE20" box="[456,556,403,429]" italics="true" pageId="25" pageNumber="26">pleurites</emphasis>
|
||
brown.
|
||
<emphasis id="B902D301E005FFECFD7FD588FCABFE21" box="[652,790,403,428]" italics="true" pageId="25" pageNumber="26">Ventrite I–V</emphasis>
|
||
loose setose with short, reclinate, yellowish setae; long, semi-erect, brown setae interspersed.
|
||
<taxonomicName id="4C767490E005FFECFDA3D5ACFD59FE5C" authorityName="Bocchino" authorityYear="1964" box="[592,740,439,465]" class="Actinopterygii" genus="Propygidium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="25" pageNumber="26" phylum="Chordata" rank="genus">
|
||
<emphasis id="B902D301E005FFECFDA3D5ACFD59FE5C" box="[592,740,439,465]" italics="true" pageId="25" pageNumber="26">Propygidium</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C767490E005FFECFCD3D5ACFC37FE5C" box="[800,906,439,465]" family="Trichomycteridae" genus="Pygidium" higherTaxonomySource="CoL" kingdom="Animalia" order="Siluriformes" pageId="25" pageNumber="70" phylum="Chordata" rank="genus">
|
||
<emphasis id="B902D301E005FFECFCD3D5ACFC37FE5C" box="[800,906,439,465]" italics="true" pageId="25" pageNumber="26">pygidium</emphasis>
|
||
</taxonomicName>
|
||
loose setose with short, reclinate, yellowishbrown setae, medium-long to long, semi-erect and erect, pale-brown setae interspersed; tip of
|
||
<taxonomicName id="4C767490E005FFECFB12D5C0FAF6FE78" box="[1249,1355,475,501]" family="Trichomycteridae" genus="Pygidium" higherTaxonomySource="CoL" kingdom="Animalia" order="Siluriformes" pageId="25" pageNumber="70" phylum="Chordata" rank="genus">
|
||
<emphasis id="B902D301E005FFECFB12D5C0FAF6FE78" box="[1249,1355,475,501]" italics="true" pageId="25" pageNumber="26">pygidium</emphasis>
|
||
</taxonomicName>
|
||
with a loose ciliary of medium-long, brown setae. Setae on
|
||
<emphasis id="B902D301E005FFECFD1AD5E4FC05FD94" box="[745,952,511,537]" italics="true" pageId="25" pageNumber="26">prothoracic femur</emphasis>
|
||
brown. Setae of longitudinal dorsal row of
|
||
<emphasis id="B902D301E005FFECFF64D638FEE4FDB0" box="[151,345,547,573]" italics="true" pageId="25" pageNumber="26">prothoracic tibia</emphasis>
|
||
brown. Setae on
|
||
<emphasis id="B902D301E005FFECFDD2D638FCB8FDB0" box="[545,773,547,573]" italics="true" pageId="25" pageNumber="26">mesothoracic femur</emphasis>
|
||
brown, pale-brown ventrally. Interspersed setae on
|
||
<emphasis id="B902D301E005FFECFAABD638FE8CFDEC" italics="true" pageId="25" pageNumber="26">mesothoracic tibia</emphasis>
|
||
brown. Bristles of ciliary red-brown.
|
||
<emphasis id="B902D301E005FFECFD12D65CFC7EFDEC" box="[737,963,583,609]" italics="true" pageId="25" pageNumber="26">Metathoracic femur</emphasis>
|
||
sparse setose with short to medium-long, reclinate, pale-brown setae; long to distinctly long, semi-erect, brown setae interspersed; denser dorsally. Setae of
|
||
<emphasis id="B902D301E005FFECFF64D694FED7FD24" box="[151,362,655,681]" italics="true" pageId="25" pageNumber="26">metathoracic tibia</emphasis>
|
||
brown, interspersed ones offwhite. Bristles of ciliary very short, transparently red-brown and distinctly lengthening dorsally and ventrally. Ciliary of
|
||
<emphasis id="B902D301E005FFECFCE9D6A8FC75FD41" box="[794,968,691,716]" italics="true" pageId="25" pageNumber="26">tarsomere I–IV</emphasis>
|
||
of thoracic legs red-brown. Interspersed setae on
|
||
<emphasis id="B902D301E005FFECFF0FD6CCFE22FD7D" box="[252,415,727,752]" italics="true" pageId="25" pageNumber="26">pretarsomeres</emphasis>
|
||
of thoracic legs transparently pale red-brown.
|
||
</paragraph>
|
||
<paragraph id="8BC90F13E005FFECFF34D6E0FD73FC0C" blockId="25.[151,1437,151,1659]" pageId="25" pageNumber="26">
|
||
<emphasis id="B902D301E005FFECFF34D6E0FEACFC98" bold="true" box="[199,273,763,789]" pageId="25" pageNumber="26">Head:</emphasis>
|
||
Clypeus slightly longitudinally depressed laterally; dense punctate with small, deep, roundish punctures, micropunctate within; anterior margin distinctly reflexed. Head dense punctate with small to large, deep, roundish, partly merging punctures. Eye on head level. Funicle II asymmetric frustum-shaped; funicle III short asymmetric frustum-shaped; clavus short elliptic.
|
||
</paragraph>
|
||
<paragraph id="8BC90F13E005FFECFF34D790FF4CFC44" blockId="25.[151,1437,151,1659]" pageId="25" pageNumber="26">
|
||
<emphasis id="B902D301E005FFECFF34D790FEF1FC28" bold="true" box="[199,332,907,933]" pageId="25" pageNumber="26">Pronotum:</emphasis>
|
||
Widest in anterior third; moderate pulvinate laterally. Anterior corner obtuse. Lateral margin convex.
|
||
</paragraph>
|
||
<paragraph id="8BC90F13E005FFECFF34D7C9FC43FB9C" blockId="25.[151,1437,151,1659]" pageId="25" pageNumber="26">
|
||
<taxonomicName id="4C767490E005FFECFF34D7C9FE27FC60" authority=": Dense" authorityName="Dense" box="[199,410,978,1005]" class="Trilobita" family="Styginidae" genus="Scutellum" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Corynexochida" pageId="25" pageNumber="26" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B902D301E005FFECFF34D7C9FEF6FC61" bold="true" box="[199,331,978,1004]" pageId="25" pageNumber="26">Scutellum:</emphasis>
|
||
Dense
|
||
</taxonomicName>
|
||
punctate with very small, deep, roundish punctures; with a shortened, median, longitudinal, narrow, smooth stripe from anterior margin terminating in anterior half.
|
||
</paragraph>
|
||
<paragraph id="8BC90F13E005FFECFF34D000FB09FBD4" blockId="25.[151,1437,151,1659]" pageId="25" pageNumber="26">
|
||
<emphasis id="B902D301E005FFECFF34D000FEA3FBB8" bold="true" box="[199,286,1051,1077]" pageId="25" pageNumber="26">Elytra:</emphasis>
|
||
Longer than broad; widest in anterior quarter. Sculpture indistinct; scutellar torus missing; complanate along suture. Tubercles of juxta-sutural row of very small; supra-humeral row missing.
|
||
</paragraph>
|
||
<paragraph id="8BC90F13E005FFECFF34D079FAFEFB48" blockId="25.[151,1437,151,1659]" pageId="25" pageNumber="26">
|
||
<emphasis id="B902D301E005FFECFF34D079FEFAFBF1" bold="true" box="[199,327,1122,1148]" pageId="25" pageNumber="26">Abdomen:</emphasis>
|
||
Distinctly pulvinate; ventrite I distinctly shortened medially; ventrite II–V broadened medially.
|
||
<taxonomicName id="4C767490E005FFECFF64D09CFE91FB2C" authorityName="Bocchino" authorityYear="1964" box="[151,300,1159,1185]" class="Actinopterygii" genus="Propygidium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="25" pageNumber="70" phylum="Chordata" rank="genus">Propygidium</taxonomicName>
|
||
with an indistinct stoma at posterior corner.
|
||
<taxonomicName id="4C767490E005FFECFCC9D09CFC14FB2C" box="[826,937,1159,1185]" family="Trichomycteridae" genus="Pygidium" higherTaxonomySource="CoL" kingdom="Animalia" order="Siluriformes" pageId="25" pageNumber="70" phylum="Chordata" rank="genus">Pygidium</taxonomicName>
|
||
slightly pulvinate longitudinally and laterally; slightly declined posteriorly; lateral margin subparallel in upper quarter, linearly attenuate apically.
|
||
</paragraph>
|
||
<paragraph id="8BC90F13E005FFECFF34D0D5FD1BFAF4" blockId="25.[151,1437,151,1659]" pageId="25" pageNumber="26">
|
||
<emphasis id="B902D301E005FFECFF34D0D5FE31FB64" bold="true" box="[199,396,1230,1257]" pageId="25" pageNumber="26">Prothoracic leg:</emphasis>
|
||
Median tooth broad triangular-shaped. Shape of interspace between teeth concave. Tarsomere I–IV combined distinctly short; equal in length; slightly more than
|
||
<superScript id="7C03A25BE005FFECFC2AD0E9FC5CFA8D" attach="right" box="[985,993,1266,1280]" fontSize="6" pageId="25" pageNumber="26">3</superScript>
|
||
/
|
||
<quantity id="4C8EA2F6E005FFECFC1AD11EFBB2FA80" box="[1001,1039,1267,1299]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="25" pageNumber="26" unit="in" value="5.0">
|
||
<subScript id="17F20D56E005FFECFC1AD11EFC4CFA9E" attach="left" box="[1001,1009,1285,1299]" fontSize="6" pageId="25" pageNumber="26">5</subScript>
|
||
in
|
||
</quantity>
|
||
length of tibia. Pretarsomere slender; with smooth ventrally, roundish cutting missing; slightly less than
|
||
<superScript id="7C03A25BE005FFECFC33D10CFC75FAA8" attach="right" box="[960,968,1303,1317]" fontSize="6" pageId="25" pageNumber="26">7</superScript>
|
||
/
|
||
<quantity id="4C8EA2F6E005FFECFC23D132FC40FABC" box="[976,1021,1303,1335]" metricMagnitude="-1" metricUnit="m" metricValue="2.54" pageId="25" pageNumber="26" unit="in" value="10.0">
|
||
<subScript id="17F20D56E005FFECFC23D132FC5DFABA" attach="left" box="[976,992,1321,1335]" fontSize="6" pageId="25" pageNumber="26">10</subScript>
|
||
in
|
||
</quantity>
|
||
length of tarsomere I–IV combined. Internal claw slender; not broadened medially; slightly more than
|
||
<superScript id="7C03A25BE005FFECFC6FD121FC19FAC5" attach="right" box="[924,932,1338,1352]" fontSize="6" pageId="25" pageNumber="26">9</superScript>
|
||
/
|
||
<quantity id="4C8EA2F6E005FFECFC5FD156FC61FAD8" box="[940,988,1339,1371]" metricMagnitude="-1" metricUnit="m" metricValue="2.54" pageId="25" pageNumber="26" unit="in" value="10.0">
|
||
<subScript id="17F20D56E005FFECFC5FD156FC01FAD6" attach="left" box="[940,956,1357,1371]" fontSize="6" pageId="25" pageNumber="26">10</subScript>
|
||
in
|
||
</quantity>
|
||
length of pretarsomere. External claw slightly less than
|
||
<superScript id="7C03A25BE005FFECFE93D145FED5FAE1" attach="right" box="[352,360,1374,1388]" fontSize="6" pageId="25" pageNumber="26">3</superScript>
|
||
/
|
||
<quantity id="4C8EA2F6E005FFECFE83D16AFE2BFAF4" box="[368,406,1375,1407]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="25" pageNumber="26" unit="in" value="5.0">
|
||
<subScript id="17F20D56E005FFECFE83D16AFEC5FAF2" attach="left" box="[368,376,1393,1407]" fontSize="6" pageId="25" pageNumber="26">5</subScript>
|
||
in
|
||
</quantity>
|
||
length of internal claw.
|
||
</paragraph>
|
||
<paragraph id="8BC90F13E005FFECFF34D199FB54F984" blockId="25.[151,1437,151,1659]" pageId="25" pageNumber="26">
|
||
<emphasis id="B902D301E005FFECFF34D199FE1DFA10" bold="true" box="[199,416,1410,1437]" pageId="25" pageNumber="26">Mesothoracic leg:</emphasis>
|
||
Tibia slightly less than
|
||
<superScript id="7C03A25BE005FFECFD5CD199FD0AFA1D" attach="right" box="[687,695,1410,1424]" fontSize="6" pageId="25" pageNumber="26">9</superScript>
|
||
/
|
||
<quantity id="4C8EA2F6E005FFECFD4CD18EFD51FA10" box="[703,748,1411,1443]" metricMagnitude="-1" metricUnit="m" metricValue="2.54" pageId="25" pageNumber="26" unit="in" value="10.0">
|
||
<subScript id="17F20D56E005FFECFD4CD18EFD72FA2E" attach="left" box="[703,719,1429,1443]" fontSize="6" pageId="25" pageNumber="26">10</subScript>
|
||
in
|
||
</quantity>
|
||
length femur. Tarsomere I–IV combined very short; slightly more than
|
||
<superScript id="7C03A25BE005FFECFEE6D1BDFEA0FA39" attach="right" box="[277,285,1446,1460]" fontSize="6" pageId="25" pageNumber="26">2</superScript>
|
||
/
|
||
<quantity id="4C8EA2F6E005FFECFED6D1A2FEF0FA4C" box="[293,333,1447,1479]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="25" pageNumber="26" unit="in" value="5.0">
|
||
<subScript id="17F20D56E005FFECFED6D1A2FE90FA4A" attach="left" box="[293,301,1465,1479]" fontSize="6" pageId="25" pageNumber="26">5</subScript>
|
||
in
|
||
</quantity>
|
||
length of tibia. Pretarsomere slender; with a smooth ridge ventrally, roundish cutting missing; slightly less than
|
||
<superScript id="7C03A25BE005FFECFE9FD1D1FEC9FA55" attach="right" box="[364,372,1482,1496]" fontSize="6" pageId="25" pageNumber="26">4</superScript>
|
||
/
|
||
<quantity id="4C8EA2F6E005FFECFE8ED1C6FE1BFA68" box="[381,422,1483,1515]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="25" pageNumber="26" unit="in" value="5.0">
|
||
<subScript id="17F20D56E005FFECFE8ED1C6FE38FA66" attach="left" box="[381,389,1501,1515]" fontSize="6" pageId="25" pageNumber="26">5</subScript>
|
||
in
|
||
</quantity>
|
||
length of tarsomere I–IV combined. External claw slender; not broadened medially; slightly shorter than pretarsomere. Internal claw slightly more than
|
||
<superScript id="7C03A25BE005FFECFC6DD1F5FC1BFA71" attach="right" box="[926,934,1518,1532]" fontSize="6" pageId="25" pageNumber="26">4</superScript>
|
||
/
|
||
<quantity id="4C8EA2F6E005FFECFC5CD21AFC69F984" box="[943,980,1519,1551]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="25" pageNumber="26" unit="in" value="5.0">
|
||
<subScript id="17F20D56E005FFECFC5CD21AFC0AF982" attach="left" box="[943,951,1537,1551]" fontSize="6" pageId="25" pageNumber="26">5</subScript>
|
||
in
|
||
</quantity>
|
||
length of external claw.
|
||
</paragraph>
|
||
<paragraph id="8BC90F13E005FFECFF34D209FBDEF9F8" blockId="25.[151,1437,151,1659]" pageId="25" pageNumber="26">
|
||
<emphasis id="B902D301E005FFECFF34D209FE1DF9A0" bold="true" box="[199,416,1554,1581]" pageId="25" pageNumber="26">Metathoracic leg:</emphasis>
|
||
Tibia slightly dorso-apically extended; slightly more than
|
||
<superScript id="7C03A25BE005FFECFBBAD208FBECF9AC" attach="right" box="[1097,1105,1555,1569]" fontSize="6" pageId="25" pageNumber="26">7</superScript>
|
||
/
|
||
<quantity id="4C8EA2F6E005FFECFBA9D23EFB34F9A0" box="[1114,1161,1555,1587]" metricMagnitude="-1" metricUnit="m" metricValue="2.54" pageId="25" pageNumber="26" unit="in" value="10.0">
|
||
<subScript id="17F20D56E005FFECFBA9D23EFBD7F9BE" attach="left" box="[1114,1130,1573,1587]" fontSize="6" pageId="25" pageNumber="26">10</subScript>
|
||
in
|
||
</quantity>
|
||
length of femur. Tarsomere I–IV combined short; slightly less than
|
||
<superScript id="7C03A25BE005FFECFD59D22DFD0FF9C9" attach="right" box="[682,690,1590,1604]" fontSize="6" pageId="25" pageNumber="26">4</superScript>
|
||
/
|
||
<quantity id="4C8EA2F6E005FFECFD48D252FD5FF9DC" box="[699,738,1591,1623]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="25" pageNumber="26" unit="in" value="5.0">
|
||
<subScript id="17F20D56E005FFECFD48D252FD7EF9DA" attach="left" box="[699,707,1609,1623]" fontSize="6" pageId="25" pageNumber="26">5</subScript>
|
||
in
|
||
</quantity>
|
||
length of tibia. External claw slightly less than
|
||
<superScript id="7C03A25BE005FFECFAEBD22DFA9DF9C9" attach="right" box="[1304,1312,1590,1604]" fontSize="6" pageId="25" pageNumber="26">1</superScript>
|
||
/
|
||
<subScript id="17F20D56E005FFECFADAD252FA84F9DA" attach="left" box="[1321,1337,1609,1623]" fontSize="6" pageId="25" pageNumber="26">10</subScript>
|
||
more in length of pretarsomere. Internal claw slightly less than
|
||
<superScript id="7C03A25BE005FFECFCE3D241FCA5F9E5" attach="right" box="[784,792,1626,1640]" fontSize="6" pageId="25" pageNumber="26">9</superScript>
|
||
/
|
||
<quantity id="4C8EA2F6E005FFECFCD3D276FCF3F9F8" box="[800,846,1627,1659]" metricMagnitude="-1" metricUnit="m" metricValue="2.54" pageId="25" pageNumber="26" unit="in" value="10.0">
|
||
<subScript id="17F20D56E005FFECFCD3D276FC8DF9F6" attach="left" box="[800,816,1645,1659]" fontSize="6" pageId="25" pageNumber="26">10</subScript>
|
||
in
|
||
</quantity>
|
||
length of external claw.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36C5C98E005FFECFF64D2B9FC9DF888" pageId="25" pageNumber="26" type="distribution">
|
||
<paragraph id="8BC90F13E005FFECFF64D2B9FBDAF930" blockId="25.[151,1127,1698,1725]" box="[151,1127,1698,1725]" pageId="25" pageNumber="26">
|
||
<emphasis id="B902D301E005FFECFF64D2B9FE89F931" bold="true" box="[151,308,1698,1724]" pageId="25" pageNumber="26">Distribution.</emphasis>
|
||
<collectingRegion id="49B2C1F1E005FFECFEC8D2B8FDEFF930" box="[315,594,1699,1725]" country="South Africa" name="Northern Cape" pageId="25" pageNumber="26">Northern Cape Province</collectingRegion>
|
||
(
|
||
<figureCitation id="134D1396E005FFECFD92D2B8FD78F930" box="[609,709,1699,1725]" captionStart="FIGURE 140" captionStartId="72.[151,250,1944,1968]" captionTargetBox="[195,1393,188,1910]" captionTargetId="figure-25@72.[185,1402,181,1920]" captionTargetPageId="72" captionText="FIGURE 140. Distributions of 18 Beckhoplia Dombrow, 2005 species, showing the concentration of species around the Northern Cape town of Nieuwoudtville." figureDoi="http://doi.org/10.5281/zenodo.4401964" httpUri="https://zenodo.org/record/4401964/files/figure.png" pageId="25" pageNumber="26">Fig. 140</figureCitation>
|
||
), only known from the
|
||
<typeStatus id="54CDB1B1E005FFECFC20D2B8FBB9F930" box="[979,1028,1699,1725]" pageId="25" pageNumber="26">type</typeStatus>
|
||
locality.
|
||
</paragraph>
|
||
<paragraph id="8BC90F13E005FFECFF64D2F1FC9DF888" blockId="25.[151,800,1770,1797]" box="[151,800,1770,1797]" pageId="25" pageNumber="26">
|
||
<emphasis id="B902D301E005FFECFF64D2F1FEDFF888" bold="true" box="[151,354,1770,1797]" pageId="25" pageNumber="26">
|
||
Vegetation
|
||
<typeStatus id="54CDB1B1E005FFECFEECD2F0FEE0F888" box="[287,349,1771,1797]" pageId="25" pageNumber="26">Type</typeStatus>
|
||
.
|
||
</emphasis>
|
||
Bokkeveld Sandstone Fynbos [FFs 1].
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36C5C98E005FFECFF64D328FD7CF8C0" box="[151,705,1843,1869]" pageId="25" pageNumber="26" type="biology_ecology">
|
||
<paragraph id="8BC90F13E005FFECFF64D328FD7CF8C0" blockId="25.[151,705,1843,1869]" box="[151,705,1843,1869]" pageId="25" pageNumber="26">
|
||
<emphasis id="B902D301E005FFECFF64D328FE96F8C0" bold="true" box="[151,299,1843,1869]" pageId="25" pageNumber="26">Host Plants.</emphasis>
|
||
<taxonomicName id="4C767490E005FFECFEC1D328FE1DF8C0" box="[306,416,1843,1869]" class="Liliopsida" family="Iridaceae" kingdom="Plantae" order="Asparagales" pageId="25" pageNumber="26" phylum="Tracheophyta" rank="family">Iridaceae</taxonomicName>
|
||
:
|
||
<taxonomicName id="4C767490E005FFECFE58D328FD7CF8C0" authority="L. Bolus." authorityName="L. Bolus." box="[427,705,1843,1869]" class="Liliopsida" family="Iridaceae" genus="Moraea" kingdom="Plantae" order="Asparagales" pageId="25" pageNumber="26" phylum="Tracheophyta" rank="species" species="bifida">
|
||
<emphasis id="B902D301E005FFECFE58D328FDEDF8C0" box="[427,592,1843,1869]" italics="true" pageId="25" pageNumber="26">Moraea bifida</emphasis>
|
||
L. Bolus.
|
||
</taxonomicName>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36C5C98E005FFECFF64D360FBB9F818" box="[151,1028,1914,1941]" pageId="25" pageNumber="26" type="etymology">
|
||
<paragraph id="8BC90F13E005FFECFF64D360FBB9F818" blockId="25.[151,1028,1914,1941]" box="[151,1028,1914,1941]" pageId="25" pageNumber="26">
|
||
<emphasis id="B902D301E005FFECFF64D360FE9CF818" bold="true" box="[151,289,1915,1941]" pageId="25" pageNumber="26">Etymology.</emphasis>
|
||
The name derives from the brown elytra [Latin: fusc- = brown].
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |